283 lines
22 KiB
XML
283 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit & Jochum, Adrienne" docDate="2023" docId="2FDE4A26983B52388379B1950E652C7A" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila erawanica Pall-Gergely & Dumrongrojwattana 2023, sp. nov." docType="treatment" docUuid="0C2132C3-73C5-4330-8ADE-CBF861481089" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunyadi, Andras</mods:namePart>
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<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grego, Jozef</mods:namePart>
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<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reischuetz, Alexander</mods:namePart>
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<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
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<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Oerstan, Aydin</mods:namePart>
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<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fekete, Judit</mods:namePart>
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<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1147</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>177</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
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<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:0C2132C3-73C5-4330-8ADE-CBF861481089" httpUri="http://treatment.plazi.org/id/2FDE4A26983B52388379B1950E652C7A" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="https://zoobank.org/0C2132C3-73C5-4330-8ADE-CBF861481089" authority="Páll-Gergely & Dumrongrojwattana" authorityName="Pall-Gergely & Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila erawanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="erawanica" status="sp. nov.">
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Angustopila erawanica
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Dumrongrojwattana
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Angustopila erawanica Pall-Gergely & Dumrongrojwattana, sp. nov. (holotype, CUMZ 7437). Apertural (A), ventral (B), lateral (C) and apical (D) sides of the shell; aperture (F); sculpture of the protoconch (E), ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure37" httpUri="https://binary.pensoft.net/fig/808846" pageId="0" pageNumber="1">Figs 37</figureCitation>
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<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Conchological variability of Angustopila erawanica Pall-Gergely & Dumrongrojwattana, sp. nov. A and C paratypes [HNHM 103474] B holotype [CUMZ 7437]." figureDoi="10.3897/zookeys.1147.93824.figure38" httpUri="https://binary.pensoft.net/fig/808847" pageId="0" pageNumber="1">, 38</figureCitation>
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<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Anatomy of Angustopila erawanica Pall-Gergely & Dumrongrojwattana, sp. nov. (HNHM 103472) A photo of the left lateral side of the body for which the genitalia are dissected out and specified in B B distal genitalia. Abbreviations: a: atrium; e: epiphallus; g: gut; h: head; p: penis; pc: penial caecum; rm: retractor muscle; s: sole; v: vagina; vd: vas deferens. White areas in B indicate parts of the genitalia that were lighter in colour (i. e., less translucent). Scale bars: 200 μm." figureDoi="10.3897/zookeys.1147.93824.figure39" httpUri="https://binary.pensoft.net/fig/808848" pageId="0" pageNumber="1">, 39</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: Thailand • 1 empty shell (H: 1.11 mm, D: 0.97 mm); Phrae Province, Long District, Tham Erawan (cave) near Maharat Rock Garden;
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<geoCoordinate degrees="18" direction="north" minutes="09.21" orientation="latitude" precision="9" value="18.1535">18°09.21'N</geoCoordinate>
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,
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<geoCoordinate degrees="99" direction="east" minutes="59.24" orientation="longitude" precision="9" value="99.987335">99°59.24'E</geoCoordinate>
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; 18 Jun. 2016; P. Dumrongrojwattana, P. Juangsantad, K. Khwantong, N. Namisa & P. Panthong leg.; CUMZ 7437.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: Thailand • 2 figured shells; same data as for holotype; HNHM 103474 • 2 dissected paratypes in ethanol; same data as for holotype; HNHM 103472 • 2 shells; same data as for holotype; NMBE 551275 • 7 specimens in ethanol; same data as for holotype; coll. PGB • 8 specimens in ethanol; same data as for holotype; coll. PD • 10 shells; same data as for holotype; coll. PD • 2 shells; same data as for holotype; coll. HA • 34 shells; same data as for holotype; coll. PGB.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<collectingCountry name="Thailand">Thailand</collectingCountry>
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• 18 j/b shells; same data as for holotype; coll. PD.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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A medium-sized to large
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<taxonomicName authorityName="Jochum, Slapnik & Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
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</taxonomicName>
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species with a narrow umbilicus and an elliptical or ovoid, slightly protruding aperture with wide sinulus and impressed at the position of the strong parietal tooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell medium- to large-sized for the genus, slightly higher than wide or slightly wider than high; pale grey, conical-ovoid, last or penultimate whorl widest from standard apertural view; protoconch consists of 1.25 whorls, with very slight indication of spiral striae preceding the first teleoconch whorl; teleoconch sculpture overall weak; teleoconch finely ornamented with irregularly spaced radial growth lines crossed by fine rows of regularly spaced spiral striae (ca. 16-18 spiral striae on body whorl in apertural view, the denser striation on the holotype is probably due to duplication of spiral striation, other shells have fewer striae); on both frontal and ventral surfaces of body whorl spiral lines dominant; whorls 4.5, slightly shouldered; aperture strongly oblique to shell axis from lateral view; umbilicus very narrow, less than 1/4 of shell width; aperture slightly protruding (visible from ventral and lateral views); aperture elliptical or ovoid, impressed at position of parietal tooth, sinulus wide; peristome slightly expanded, not reflected; parietal callus strongly elevated, sharp, detached from penultimate whorl; parietal tooth strongly developed; parietal side and parietal tooth align at an angle less than 90 degrees.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
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<paragraph pageId="0" pageNumber="1">
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H = 0.94-1.11; D = 0.97-1.05, H/D*100 = 95.9-114.4 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 4), RUD = 20.4-22.4 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 3).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The aperture shape, in combination with the large size and narrow umbilicus, distinguishes this species from its congeners.
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<taxonomicName authorityName="Pall-Gergely & Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila erawanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="erawanica">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila erawanica</emphasis>
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</taxonomicName>
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sp. nov. is most similar to
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<taxonomicName lsidName="A. bidentata" pageId="0" pageNumber="1" rank="species" species="bidentata">
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<emphasis italics="true" pageId="0" pageNumber="1">A. bidentata</emphasis>
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</taxonomicName>
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sp. nov. by its narrow umbilical form and the shape of the aperture, but that species has a strong palatal tooth, a less protruding and more oblique aperture, and stronger spiral striation on the protoconch.
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<taxonomicName authorityName="Pall-Gergely & Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila dominikae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dominikae">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila dominikae</emphasis>
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</taxonomicName>
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also possesses a strong palatal tooth and has a more globular shell.
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<taxonomicName baseAuthorityName="Thompson & Upatham" baseAuthorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila concava" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="concava">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila concava</emphasis>
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</taxonomicName>
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has a wider body whorl, a wider umbilicus, and a more oblique aperture. See also under
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<taxonomicName lsidName="A. huoyani" pageId="0" pageNumber="1" rank="species" species="huoyani">
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<emphasis italics="true" pageId="0" pageNumber="1">A. huoyani</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="anatomy">
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<paragraph pageId="0" pageNumber="1">Anatomy.</paragraph>
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<paragraph pageId="0" pageNumber="1">Two specimens were dissected. Due to the small size of the animals, some traits could not be examined in detail. The traits of the distal genitalia could be relatively clearly seen, but proximal to the vagina, the animals consisted of a very soft, gelatinous material. No bursa copulatrix or any other female glands found, but this can be due to the difficulties to separate these organs. Penis well-developed but short, with triangular, thick penial caecum; retractor muscle inserts at end of caecum; epiphallus starts from base of caecum, thickened at its proximal end; vas deferens slender, but could not be followed (got torn just after its distal end); the lumen of the penis is visible under light microscope as a whiter (= less translucent) area, this can be followed from the distal penis throughout the penial caecum to the proximal end of epiphallus; vagina thick, longer than penis, although its proximal end could not be identified, inside vagina was a thick whiter area.</paragraph>
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<caption doi="10.3897/zookeys.1147.93824.figure37" httpUri="https://binary.pensoft.net/fig/808846" pageId="0" pageNumber="1" start="Figure 37" startId="F37">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 37.</emphasis>
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<taxonomicName authorityName="Pall-Gergely & Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila erawanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="erawanica">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila erawanica</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Dumrongrojwattana, sp. nov. (holotype, CUMZ 7437). Apertural (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), lateral (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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) and apical (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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) sides of the shell; aperture (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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); sculpture of the protoconch (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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), ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
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) and frontal (
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<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
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) surface of the body whorl.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.1147.93824.figure38" httpUri="https://binary.pensoft.net/fig/808847" pageId="0" pageNumber="1" start="Figure 38" startId="F38">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 38.</emphasis>
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Conchological variability of
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<taxonomicName authorityName="Pall-Gergely & Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila erawanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="erawanica">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila erawanica</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Dumrongrojwattana, sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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|
and
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
paratypes [HNHM 103474]
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
holotype [CUMZ 7437].
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|
</paragraph>
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|
</caption>
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|
<caption doi="10.3897/zookeys.1147.93824.figure39" httpUri="https://binary.pensoft.net/fig/808848" pageId="0" pageNumber="1" start="Figure 39" startId="F39">
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|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 39.</emphasis>
|
|
Anatomy of
|
|
<taxonomicName authorityName="Pall-Gergely & Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila erawanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="erawanica">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila erawanica</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
|
& Dumrongrojwattana, sp. nov. (HNHM 103472)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
photo of the left lateral side of the body for which the genitalia are dissected out and specified in B
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
distal genitalia. Abbreviations: a: atrium; e: epiphallus; g: gut; h: head; p: penis; pc: penial caecum; rm: retractor muscle; s: sole; v: vagina; vd: vas deferens. White areas in
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
indicate parts of the genitalia that were lighter in colour (i.e., less translucent). Scale bars: 200
|
|
<normalizedToken originalValue="μm">μm</normalizedToken>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">The new species is named after its type locality (Erawan cave). Specific epithet is used as a noun in apposition. Erawan is the Thai form of the mythical three- or thirty-three-headed, white elephant ridden by Indra (Sanskrit), the diety of rain, thunder, weather, and river flows.</paragraph>
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|
</subSubSection>
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|
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
This species is known from the type locality only (Fig.
|
|
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure31" httpUri="https://binary.pensoft.net/fig/808840" pageId="0" pageNumber="1">31</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
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|
</document> |