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<document id="9DCD3ED4683ED2DA45ECCE80566CD564" ID-CLB-Dataset="5188" ID-DOI="10.1649/0010-065X-71.4.639" ID-GBIF-Dataset="5e2cbcdd-1887-48ab-9c07-8763aa32c869" ID-ISSN="1938-4394" ID-Zenodo-Dep="4788749" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621946355242" checkinUser="carolina" docAuthor="Erwin, Terry L." docDate="2017" docId="2F1787E8FFD2827CFCE4FE868FE9F54F" docLanguage="en" docName="TheColeopBull.71.4.639-651.pdf" docOrigin="The Coleopterists Bulletin 71 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-71.4.639" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Agra truquii Chaudoir 1866" docType="treatment" docVersion="8" lastPageNumber="649" masterDocId="D32EFF90FFDA8276FFAFFFC88E23F705" masterDocTitle="Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life" masterLastPageNumber="651" masterPageNumber="639" pageNumber="647" updateTime="1698989650016" updateUser="ExternalLinkService">
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<mods:title id="0A264D40EB5C23BE8D109650B83609F4">Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life</mods:title>
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<mods:namePart id="C5E665A9AEFA625DB3E62427105F629F">Erwin, Terry L.</mods:namePart>
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<mods:title id="4DB4EEF64144BDE10D313B2D2C7EB04E">The Coleopterists Bulletin</mods:title>
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<mods:date id="D1F3398E59806FBB110300E8897881B5">2017</mods:date>
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<mods:number id="8B9F0073213C513173AD3299F7E6A49C">2017-12-22</mods:number>
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<mods:identifier id="B01CC000F1F32D442FD902F9C1440D25" type="CLB-Dataset">5188</mods:identifier>
<mods:identifier id="995F7E40E3EE8D732C395E6F29049721" type="DOI">10.1649/0010-065X-71.4.639</mods:identifier>
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<treatment id="2F1787E8FFD2827CFCE4FE868FE9F54F" ID-DOI="http://doi.org/10.5281/zenodo.5157174" ID-GBIF-Taxon="183910543" ID-Zenodo-Dep="5157174" LSID="urn:lsid:plazi:treatment:2F1787E8FFD2827CFCE4FE868FE9F54F" httpUri="http://treatment.plazi.org/id/2F1787E8FFD2827CFCE4FE868FE9F54F" lastPageId="10" lastPageNumber="649" pageId="8" pageNumber="647">
<subSubSection id="EFA46575FFD2827EFCE4FE868A02F661" box="[843,1057,334,356]" pageId="8" pageNumber="647" type="nomenclature">
<paragraph id="A70136FEFFD2827EFCE4FE868A02F661" blockId="8.[843,1057,334,356]" box="[843,1057,334,356]" pageId="8" pageNumber="647">
<heading id="FC498192FFD2827EFCE4FE868A02F661" box="[843,1057,334,356]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="647" reason="2">
<emphasis id="95CAEAECFFD2827EFCE4FE868A02F661" bold="true" box="[843,1057,334,356]" pageId="8" pageNumber="647">
<taxonomicName id="60BE4D7DFFD2827EFCE4FE868DACF661" ID-CoL="65SKX" authorityName="Chaudoir" authorityYear="1866" box="[843,911,334,356]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="truquii">
<emphasis id="95CAEAECFFD2827EFCE4FE868DACF661" bold="true" box="[843,911,334,356]" italics="true" pageId="8" pageNumber="647">truquii</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EFA46575FFD2827EFD74FEB18DABF309" pageId="8" pageNumber="647" type="description">
<paragraph id="A70136FEFFD2827EFD74FEB18DABF309" blockId="8.[707,1193,377,1089]" pageId="8" pageNumber="647">
Members of this rather uniform group are of small to medium size and blackish; the nominate species has a subtle bluish tint to the elytra. A diagnostic combination of the adults of the
<taxonomicName id="60BE4D7DFFD2827EFBBDFE018A71F6DB" authorityName="Chaudoir" authorityYear="1866" box="[1042,1106,457,478]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="truquii">
<emphasis id="95CAEAECFFD2827EFBBDFE018A71F6DB" box="[1042,1106,457,478]" italics="true" pageId="8" pageNumber="647">truquii</emphasis>
</taxonomicName>
speciesgroup includes the following: labrum slightly depressed medially with rounded anterior lateral margin and slightly emarginate apex; antennae of males with antennomeres more or less subequal in length (except short pedicel); that of females with antennomere 8 short, slightly less than half the length of 7, antennomeres 9–11 half the length of 3-7. Antenna of normal length, reaching level of middle coxa, not markedly short and robust nor long and markedly thin. Head behind eyes moderately elongate, broadly rounded posteriorly in female, slightly tapered to neck in male; elytral interneurs each a series of large, coarse, variously separated punctures; elytral apex nearly perfectly truncated, laterally with a small, broadly acute tooth, sutural corner rounded; legs and tarsi normal, not markedly modified; male metasternum moderately setigerous, abdominal sterna III–V bilaterally setigerous, setae long and numerous, simply sparsely setiferous on female; male phallus (
<figureCitation id="3F852A7BFFD2827EFC02FC148DDDF4F4" box="[941,1022,988,1009]" captionStart="Fig" captionStartId="5.[175,206,1344,1363]" captionTargetBox="[167,1140,221,1288]" captionTargetId="figure-157@5.[151,1173,194,1328]" captionTargetPageId="5" captionText="Fig. 3. Agra species. Male genitalia: A) A. rileyi (ADP109124), median lobe, right lateral, ventral, and left lateral views, B) A. truquii (ADP058747), ventral view. Female stylomere 2, dorsal and ventral aspects: C) A. rileyi (ADP124942), D) A. wickhami (ADP056075)." figureDoi="http://doi.org/10.5281/zenodo.4788785" httpUri="https://zenodo.org/record/4788785/files/figure.png" pageId="8" pageNumber="647">Fig. 3B</figureCitation>
) broadly arrowshaped, tip rounded.
</paragraph>
</subSubSection>
<subSubSection id="EFA46575FFD2827EFD74FBD98D6DF344" pageId="8" pageNumber="647" type="description">
<paragraph id="A70136FEFFD2827EFD74FBD98D6DF344" blockId="8.[707,1193,377,1089]" pageId="8" pageNumber="647">Three species are currently recognized in the species-group:</paragraph>
</subSubSection>
<subSubSection id="EFA46575FFD2827EFD6CFB9E8A3AF3A5" pageId="8" pageNumber="647" type="reference_group">
<paragraph id="A70136FEFFD2827EFD6CFB9E8A30F36E" blockId="8.[707,1049,1110,1184]" box="[707,1043,1110,1132]" pageId="8" pageNumber="647">
<taxonomicName id="60BE4D7DFFD2827EFD6CFB9E8A30F36E" ID-CoL="B2XV" authority="Chaudoir (1866)" authorityName="Chaudoir" authorityYear="1866" box="[707,1043,1110,1132]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="smaragdina">
<emphasis id="95CAEAECFFD2827EFD6CFB9E8D4FF369" box="[707,876,1110,1132]" italics="true" pageId="8" pageNumber="647">Agra smaragdina</emphasis>
<bibRefCitation id="C32F4B0FFFD2827EFCDCFB9E8A30F36E" author="Chaudoir, M. de" box="[883,1043,1110,1131]" pageId="8" pageNumber="647" pagination="77 - 104" refId="ref5403" refString="Chaudoir, M. de. 1866. Corrections et additions a la Revision du Genre Agra. Annales de la Societe Entomologique de France 6: 77 - 104." type="journal article" year="1866">Chaudoir (1866)</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A70136FEFFD2827EFD6CFBB88DC2F383" blockId="8.[707,1049,1110,1184]" box="[707,993,1136,1158]" pageId="8" pageNumber="647">
<taxonomicName id="60BE4D7DFFD2827EFD6CFBB88DC2F383" ID-CoL="65SKX" authority="Chaudoir (1866)" authorityName="Chaudoir" authorityYear="1866" box="[707,993,1136,1158]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="truquii">
<emphasis id="95CAEAECFFD2827EFD6CFBB88D19F383" box="[707,826,1136,1158]" italics="true" pageId="8" pageNumber="647">Agra truquii</emphasis>
<bibRefCitation id="C32F4B0FFFD2827EFCEDFBB98DC2F383" author="Chaudoir, M. de" box="[834,993,1137,1158]" pageId="8" pageNumber="647" pagination="77 - 104" refId="ref5403" refString="Chaudoir, M. de. 1866. Corrections et additions a la Revision du Genre Agra. Annales de la Societe Entomologique de France 6: 77 - 104." type="journal article" year="1866">Chaudoir (1866)</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A70136FEFFD2827EFD6CFB428A3AF3A5" blockId="8.[707,1049,1110,1184]" box="[707,1049,1162,1184]" pageId="8" pageNumber="647">
<taxonomicName id="60BE4D7DFFD2827EFD6CFB428DBBF3A5" ID-CoL="8MK8M" authority="Erwin" authorityName="Erwin" authorityYear="2017" box="[707,920,1162,1184]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="wickhami" status="sp. nov.">
<emphasis id="95CAEAECFFD2827EFD6CFB428D76F3A5" box="[707,853,1162,1184]" italics="true" pageId="8" pageNumber="647">Agra wickhami</emphasis>
Erwin
</taxonomicName>
,
<emphasis id="95CAEAECFFD2827EFC0BFB438A3AF3A5" bold="true" box="[932,1049,1163,1184]" pageId="8" pageNumber="647">
<taxonomicNameLabel id="8EF95797FFD2827EFC0BFB438A3AF3A5" box="[932,1049,1163,1184]" pageId="8" pageNumber="647" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="EFA46575FFD2827EFCABFB1C8DDAF226" pageId="8" pageNumber="647" type="description">
<paragraph id="A70136FEFFD2827EFCABFB1C8DDAF226" blockId="8.[772,1127,1236,1315]" pageId="8" pageNumber="647">
<emphasis id="95CAEAECFFD2827EFCABFB1C8A44F3EC" bold="true" box="[772,1127,1236,1258]" pageId="8" pageNumber="647">
<taxonomicName id="60BE4D7DFFD2827EFCABFB1C8DC6F3EC" authority="Erwin" authorityName="Erwin" authorityYear="2017" box="[772,997,1236,1258]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="wickhami" status="sp. nov.">
<emphasis id="95CAEAECFFD2827EFCABFB1C8DBEF3EF" bold="true" box="[772,925,1236,1258]" italics="true" pageId="8" pageNumber="647">Agra wickhami</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="8EF95797FFD2827EFC5EFB1C8A44F3EC" box="[1009,1127,1236,1257]" pageId="8" pageNumber="647" rank="species">new species</taxonomicNameLabel>
</emphasis>
Wickham’ s elegant canopy beetle (
<figureCitation id="3F852A7BFFD2827EFCD5FAC68D9CF226" box="[890,959,1294,1315]" captionStart="Fig" captionStartId="4.[197,228,1739,1758]" captionTargetBox="[212,1118,204,1707]" captionTargetId="figure-0@4.[205,1160,194,1723]" captionTargetPageId="4" captionText="Fig. 2. Agra wickhami, dorsal habitus illustration. Mexico: Yucatán, ADP056075, female." figureDoi="http://doi.org/10.5281/zenodo.4788773" httpUri="https://zenodo.org/record/4788773/files/figure.png" pageId="8" pageNumber="647">Figs. 2</figureCitation>
,
<figureCitation id="3F852A7BFFD2827EFC64FAC68DFBF226" box="[971,984,1294,1315]" captionStart="Fig" captionStartId="5.[175,206,1344,1363]" captionTargetBox="[167,1140,221,1288]" captionTargetId="figure-157@5.[151,1173,194,1328]" captionTargetPageId="5" captionText="Fig. 3. Agra species. Male genitalia: A) A. rileyi (ADP109124), median lobe, right lateral, ventral, and left lateral views, B) A. truquii (ADP058747), ventral view. Female stylomere 2, dorsal and ventral aspects: C) A. rileyi (ADP124942), D) A. wickhami (ADP056075)." figureDoi="http://doi.org/10.5281/zenodo.4788785" httpUri="https://zenodo.org/record/4788785/files/figure.png" pageId="8" pageNumber="647">3</figureCitation>
,
<figureCitation id="3F852A7BFFD2827EFC4AFAC68DD2F226" box="[997,1009,1294,1315]" captionStart="Fig" captionStartId="7.[175,206,1334,1353]" captionTargetBox="[151,1173,800,1317]" captionTargetId="figure-155@7.[151,1173,800,1317]" captionTargetPageId="7" captionText="Fig. 5. Distribution map of known localities for Agra wickhami." figureDoi="http://doi.org/10.5281/zenodo.4788807" httpUri="https://zenodo.org/record/4788807/files/figure.png" pageId="8" pageNumber="647">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EFA46575FFD2827EFD74FAF18DB3F15D" pageId="8" pageNumber="647" type="materials_examined">
<paragraph id="A70136FEFFD2827EFD74FAF18A66F286" blockId="8.[707,1193,1337,1730]" pageId="8" pageNumber="647">
<materialsCitation id="17D63CA3FFD2827EFD74FAF18A66F286" ID-GBIF-Occurrence="3330927304" collectorName="HF Wickham" country="United States of America" county="Cameron County" elevation="9" latitude="25.86" location="Brownsville" longLatPrecision="746" longitude="-97.43" pageId="8" pageNumber="647" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
<emphasis id="95CAEAECFFD2827EFD74FAF18DAAF24B" bold="true" box="[731,905,1337,1358]" pageId="8" pageNumber="647">
<typeStatus id="7805885CFFD2827EFD74FAF18D18F24B" box="[731,827,1337,1358]" pageId="8" pageNumber="647" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="DFA9766EFFD2827EFCFDFAF18DA6F24B" box="[850,901,1337,1358]" name="United States of America" pageId="8" pageNumber="647">USA</collectingCountry>
:
</emphasis>
<collectingRegion id="657AF81CFFD2827EFC31FAF18DFAF249" box="[926,985,1337,1358]" country="United States of America" name="Texas" pageId="8" pageNumber="647">TEXAS</collectingRegion>
:
<collectingCounty id="4E604E72FFD2827EFC5CFAF18A86F24B" box="[1011,1189,1337,1358]" pageId="8" pageNumber="647">Cameron County</collectingCounty>
,
<location id="A2616025FFD2827EFD6CFA9C8D1AF26C" LSID="urn:lsid:plazi:treatment:2F1787E8FFD2827CFCE4FE868FE9F54F:A2616025FFD2827EFD6CFA9C8D1AF26C" box="[707,825,1364,1385]" country="United States of America" county="Cameron County" latitude="25.86" longLatPrecision="746" longitude="-97.43" name="Brownsville" pageId="8" pageNumber="647" stateProvince="Texas">Brownsville</location>
,
<quantity id="60469B1BFFD2827EFCEBFA9C8D4FF26C" box="[836,876,1364,1385]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="647" unit="m" value="9.0">
<elevation id="2C93D1CDFFD2827EFCEBFA9C8D4FF26C" box="[836,876,1364,1385]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="647" unit="m" value="9.0">9 m</elevation>
</quantity>
,
<geoCoordinate id="C28A5039FFD2827EFCD8FA9C8DEAF26C" box="[887,969,1364,1385]" degrees="25.86" direction="north" orientation="latitude" pageId="8" pageNumber="647" precision="555" value="25.86">25.86°N</geoCoordinate>
,
<geoCoordinate id="C28A5039FFD2827EFC7BFA9C8A08F26C" box="[980,1067,1364,1385]" degrees="97.43" direction="west" orientation="longitude" pageId="8" pageNumber="647" precision="555" value="-97.43">97.43°W</geoCoordinate>
, no date (
<collectorName id="0A4B5328FFD2827EFB26FA9C8D03F286" pageId="8" pageNumber="647">HF Wickham</collectorName>
) (
<collectingRegion id="657AF81CFFD2827EFC9AFAA68D44F286" box="[821,871,1390,1411]" country="United States of America" name="CAS" pageId="8" pageNumber="647">CAS</collectingRegion>
: ADP091173, female).
</materialsCitation>
</paragraph>
<paragraph id="A70136FEFFD2827EFD74FA418A78F127" blockId="8.[707,1193,1337,1730]" pageId="8" pageNumber="647">
<emphasis id="95CAEAECFFD2827EFD74FA418A05F29B" bold="true" box="[731,1062,1417,1438]" pageId="8" pageNumber="647">Derivation of Specific Epithet.</emphasis>
The epithet “
<taxonomicName id="60BE4D7DFFD2827EFD62FA6A8D0BF2BD" authorityName="Erwin" authorityYear="2017" box="[717,808,1442,1464]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="wickhami">
<emphasis id="95CAEAECFFD2827EFD62FA6A8D0BF2BD" box="[717,808,1442,1464]" italics="true" pageId="8" pageNumber="647">wickhami</emphasis>
</taxonomicName>
” is an eponym based on the family name of H. F. Wickham, Coleopterist and Paleoentomologist of the late 19
<superScript id="50CB9BB6FFD2827EFC66FA1B8DF4F2E4" attach="left" box="[969,983,1491,1505]" fontSize="6" pageId="8" pageNumber="647">th</superScript>
and early 20
<superScript id="50CB9BB6FFD2827EFBF7FA1B8A45F2E4" attach="left" box="[1112,1126,1491,1505]" fontSize="6" pageId="8" pageNumber="647">th</superScript>
centuries, who collected the adult
<typeStatus id="7805885CFFD2827EFC62FA3B8A03F10D" box="[973,1056,1523,1544]" pageId="8" pageNumber="647" type="holotype">holotype</typeStatus>
of this species somewhere in or near Brownsville,
<collectingRegion id="657AF81CFFD2827EFBB2F9C58A74F127" box="[1053,1111,1549,1570]" country="United States of America" name="Texas" pageId="8" pageNumber="647">Texas</collectingRegion>
.
</paragraph>
<paragraph id="A70136FEFFD2827EFD74F9E08DB3F15D" blockId="8.[707,1193,1337,1730]" pageId="8" pageNumber="647">
<emphasis id="95CAEAECFFD2827EFD74F9E08A1FF138" bold="true" box="[731,1084,1576,1597]" pageId="8" pageNumber="647">Proposed English Vernacular Name.</emphasis>
Wickham’ s elegant canopy beetle.
</paragraph>
</subSubSection>
<subSubSection id="EFA46575FFD2827FFD74F9958C79F2E8" lastPageId="9" lastPageNumber="648" pageId="8" pageNumber="647" type="diagnosis">
<paragraph id="A70136FEFFD2827FFD74F9958C79F2E8" blockId="8.[707,1193,1337,1730]" lastBlockId="9.[151,638,1336,1730]" lastPageId="9" lastPageNumber="648" pageId="8" pageNumber="647">
<emphasis id="95CAEAECFFD2827EFD74F9958D61F177" bold="true" box="[731,834,1629,1650]" pageId="8" pageNumber="647">Diagnosis.</emphasis>
<taxonomicName id="60BE4D7DFFD2827EFCE9F9948DF6F177" authorityName="Erwin" authorityYear="2017" box="[838,981,1628,1650]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="wickhami">
<emphasis id="95CAEAECFFD2827EFCE9F9948DF6F177" box="[838,981,1628,1650]" italics="true" pageId="8" pageNumber="647">Agra wickhami</emphasis>
</taxonomicName>
has all the attributes of the genus and species-group as described above and is large for the
<taxonomicName id="60BE4D7DFFD2827EFCF3F95B8DBFF1AD" authorityName="Chaudoir" authorityYear="1866" box="[860,924,1683,1704]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="truquii">
<emphasis id="95CAEAECFFD2827EFCF3F95B8DBFF1AD" box="[860,924,1683,1704]" italics="true" pageId="8" pageNumber="647">truquii</emphasis>
</taxonomicName>
species-group. Adults with black integument; appendages darkly infuscated; antennomeres 6–11 bicolored. Frons and occiput moderately domed, smooth, mostly glabrous, with less than 10 scattered setae in addition to fixed supraorbital setae. Pronotum with linear rows of large, coarse punctures from apex to base. Elytral interneurs each with large coarse puncture; apex truncate with small, obtuse dent apicolaterally.
</paragraph>
</subSubSection>
<caption id="F3C16676FFD3827FFF00FF0B8EF8F600" pageId="9" pageNumber="648" startId="9.[175,225,195,214]" targetBox="[313,1014,303,1253]" targetPageId="9">
<paragraph id="A70136FEFFD3827FFF00FF0B8EF8F600" blockId="9.[151,1172,194,262]" pageId="9" pageNumber="648">
<emphasis id="95CAEAECFFD3827FFF00FF0B8ED7F7D3" bold="true" box="[175,244,195,214]" pageId="9" pageNumber="648">Table 2.</emphasis>
Adult measurements and ratios for female
<taxonomicName id="60BE4D7DFFD3827FFDC9FF0A8CC6F7D0" authorityName="Erwin" authorityYear="2017" box="[614,741,194,213]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="648" phylum="Arthropoda" rank="species" species="wickhami">
<emphasis id="95CAEAECFFD3827FFDC9FF0A8CC6F7D0" box="[614,741,194,213]" italics="true" pageId="9" pageNumber="648">Agra wickhami</emphasis>
</taxonomicName>
. All values are in millimeters.Apparent body length (ABL) and standardize body length (SBL) are also provided in the descriptions. Means provided for ratios are “harmonic means.”
</paragraph>
</caption>
<subSubSection id="EFA46575FFD3827CFF00FA3B8FE9F54F" lastPageId="10" lastPageNumber="649" pageId="9" pageNumber="648" type="description">
<paragraph id="A70136FEFFD3827FFF00FA3B8A5AF177" blockId="9.[151,638,1336,1730]" lastBlockId="9.[685,1173,1337,1730]" pageId="9" pageNumber="648">
<emphasis id="95CAEAECFFD3827FFF00FA3B8F49F10D" bold="true" box="[175,362,1523,1544]" pageId="9" pageNumber="648">Description. Size:</emphasis>
Large for species group, ABL =
<quantity id="60469B1BFFD3827FFF45F9C58F5AF127" box="[234,377,1549,1571]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" metricValueMax="1.63" metricValueMin="1.55" pageId="9" pageNumber="648" unit="mm" value="15.9" valueMax="16.3" valueMin="15.5">15.5–16.3 mm</quantity>
, SBL =
<quantity id="60469B1BFFD3827FFE7CF9C58C59F127" box="[467,634,1549,1571]" metricMagnitude="-2" metricUnit="m" metricValue="1.6195" metricValueMax="1.7469999999999999" metricValueMin="1.492" pageId="9" pageNumber="648" unit="mm" value="16.195" valueMax="17.47" valueMin="14.92">14.92–17.47 mm</quantity>
, TW =
<quantity id="60469B1BFFD3827FFF49F9E08F55F138" box="[230,374,1576,1598]" metricMagnitude="-3" metricUnit="m" metricValue="4.395" metricValueMax="4.45" metricValueMin="4.34" pageId="9" pageNumber="648" unit="mm" value="4.395" valueMax="4.45" valueMin="4.34">4.34–4.45 mm</quantity>
(
<tableCitation id="EA3C0345FFD3827FFE28F9E08FF5F138" box="[391,470,1576,1597]" captionStart="Table 2" captionStartId="9.[175,225,195,214]" captionTargetId="figure-265@9.[311,1016,301,1255]" captionTargetPageId="9" captionText="Table 2. Adult measurements and ratios for female Agra wickhami. All values are in millimeters.Apparent body length (ABL) and standardize body length (SBL) are also provided in the descriptions. Means provided for ratios are “harmonic means.”" pageId="9" pageNumber="648">Table 2</tableCitation>
).
<emphasis id="95CAEAECFFD3827FFE42F9E08C0DF138" bold="true" box="[493,558,1576,1597]" pageId="9" pageNumber="648">Color:</emphasis>
As described above and antennomers 1–5 infuscated, 6–11 bicolored with lateral black stripes; mouthparts infuscated, especially maxillary palpomeres, legs and tarsi dark brownish.
<emphasis id="95CAEAECFFD3827FFE6FF95A8C28F1A2" bold="true" box="[448,523,1682,1703]" pageId="9" pageNumber="648">Luster:</emphasis>
Head, pronotum, and legs shiny, elytra matte black.
<emphasis id="95CAEAECFFD3827FFDEFF9658C5EF1C7" bold="true" box="[576,637,1709,1730]" pageId="9" pageNumber="648">Head:</emphasis>
As described above.
<emphasis id="95CAEAECFFD3827FFCD2FAF18DCFF24B" bold="true" box="[893,1004,1337,1358]" pageId="9" pageNumber="648">Prothorax:</emphasis>
Short, about the length of head, moderately constricted near base, devoid of hind angles, narrowed anteriorly to about width of neck; surface of disc as described above (
<figureCitation id="3F852A7BFFD3827FFD1BFA6B8CCCF2BD" box="[692,751,1443,1464]" captionStart="Fig" captionStartId="4.[197,228,1739,1758]" captionTargetBox="[212,1118,204,1707]" captionTargetId="figure-0@4.[205,1160,194,1723]" captionTargetPageId="4" captionText="Fig. 2. Agra wickhami, dorsal habitus illustration. Mexico: Yucatán, ADP056075, female." figureDoi="http://doi.org/10.5281/zenodo.4788773" httpUri="https://zenodo.org/record/4788773/files/figure.png" pageId="9" pageNumber="648">Fig. 2</figureCitation>
).
<emphasis id="95CAEAECFFD3827FFCADFA6B8DA0F2BD" bold="true" box="[770,899,1443,1464]" pageId="9" pageNumber="648">Pterothorax:</emphasis>
Elytron moderately convex, narrow in anterior third, slightly flared from middle to apical third and rounded to lateral hind angle; intervals moderately convex, quite regular in width.
<emphasis id="95CAEAECFFD3827FFD59F9C58D0EF127" bold="true" box="[758,813,1549,1570]" pageId="9" pageNumber="648">Legs:</emphasis>
Simple in females.
<emphasis id="95CAEAECFFD3827FFBADF9C58A4EF127" bold="true" box="[1026,1133,1549,1570]" pageId="9" pageNumber="648">Abdomen:</emphasis>
As described above.
<emphasis id="95CAEAECFFD3827FFCC0F9E08A3AF138" bold="true" box="[879,1049,1576,1597]" pageId="9" pageNumber="648">Male genitalia:</emphasis>
Unknown.
<emphasis id="95CAEAECFFD3827FFD02F98B8D5AF15D" bold="true" box="[685,889,1603,1624]" pageId="9" pageNumber="648">Female ovipositor:</emphasis>
Female internal parts not investigated. Female stylomere 2 as in
<figureCitation id="3F852A7BFFD3827FFB97F9958A56F177" box="[1080,1141,1629,1650]" captionStart="Fig" captionStartId="5.[175,206,1344,1363]" captionTargetBox="[167,1140,221,1288]" captionTargetId="figure-157@5.[151,1173,194,1328]" captionTargetPageId="5" captionText="Fig. 3. Agra species. Male genitalia: A) A. rileyi (ADP109124), median lobe, right lateral, ventral, and left lateral views, B) A. truquii (ADP058747), ventral view. Female stylomere 2, dorsal and ventral aspects: C) A. rileyi (ADP124942), D) A. wickhami (ADP056075)." figureDoi="http://doi.org/10.5281/zenodo.4788785" httpUri="https://zenodo.org/record/4788785/files/figure.png" pageId="9" pageNumber="648">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="A70136FEFFD3827FFD6AF9B08D78F1C7" blockId="9.[685,1173,1337,1730]" pageId="9" pageNumber="648">
<emphasis id="95CAEAECFFD3827FFD6AF9B08DAEF188" bold="true" box="[709,909,1656,1677]" pageId="9" pageNumber="648">Dispersal Potential.</emphasis>
These beetles are macropterous and probably capable of flight; they are swift and agile runners.
</paragraph>
<paragraph id="A70136FEFFD0827CFF6AFF0A8F26F6AE" blockId="10.[173,659,193,586]" pageId="10" pageNumber="649">
<emphasis id="95CAEAECFFD0827CFF6AFF0A8F6FF7D2" bold="true" box="[197,332,194,215]" pageId="10" pageNumber="649">Way of Life.</emphasis>
See
<taxonomicName id="60BE4D7DFFD0827CFE2AFF098FF5F7D2" authorityName="Erwin" authorityYear="2017" box="[389,470,193,215]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="rileyi">
<emphasis id="95CAEAECFFD0827CFE2AFF098FF5F7D2" box="[389,470,193,215]" italics="true" pageId="10" pageNumber="649">A. rileyi</emphasis>
</taxonomicName>
above for general information. Specimens of
<taxonomicName id="60BE4D7DFFD0827CFE7CFF138C77F7F4" authorityName="Erwin" authorityYear="2017" box="[467,596,219,241]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="wickhami">
<emphasis id="95CAEAECFFD0827CFE7CFF138C77F7F4" box="[467,596,219,241]" italics="true" pageId="10" pageNumber="649">A. wickhami</emphasis>
</taxonomicName>
were taken at lowland elevations along the Rio Grande Valley in southeastern
<collectingRegion id="657AF81CFFD0827CFE3DFED98FEDF623" box="[402,462,273,294]" country="United States of America" name="Texas" pageId="10" pageNumber="649">Texas</collectingRegion>
, where sabal palms and sugar hackberry grow. Adults are active in the late rainy season in the
<collectingRegion id="657AF81CFFD0827CFE0EFE8E8FD3F65E" box="[417,496,326,347]" country="Mexico" name="Yucatan" pageId="10" pageNumber="649">Yucatán</collectingRegion>
of
<collectingRegion id="657AF81CFFD0827CFDB9FE8E8C47F65E" box="[534,612,326,347]" country="Mexico" name="Mexico" pageId="10" pageNumber="649">Mexico</collectingRegion>
. See
<bibRefCitation id="C32F4B0FFFD0827CFF02FEA98F77F673" author="Wickham, H. F." box="[173,340,353,374]" pageId="10" pageNumber="649" pagination="96 - 115" refId="ref6457" refString="Wickham, H. F. 1897. The Coleoptera of the Lower Rio Grande Valley. Bulletin from the Laboratories of Natural History of the State University of Iowa 4 (2): 96 - 115." type="journal article" year="1897">Wickham (1897)</bibRefCitation>
for descriptions of the vegetation at Brownsville at the time of his collecting the
<typeStatus id="7805885CFFD0827CFF02FE5E8F22F6AE" box="[173,257,406,427]" pageId="10" pageNumber="649" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="A70136FEFFD0827CFF6AFE798C45F6FE" blockId="10.[173,659,193,586]" pageId="10" pageNumber="649">
<materialsCitation id="17D63CA3FFD0827CFF6AFE798F3FF6E5" ID-GBIF-Occurrence="3330927301" country="Mexico" elevation="16" location="Arco" pageId="10" pageNumber="649" specimenCount="1" stateProvince="Yucatan">
<emphasis id="95CAEAECFFD0827CFF6AFE798C11F6C3" bold="true" box="[197,562,433,454]" pageId="10" pageNumber="649">
Other Specimens Examined.
<collectingCountry id="DFA9766EFFD0827CFE4DFE798C0DF6C3" box="[482,558,433,454]" name="Mexico" pageId="10" pageNumber="649">Mexico</collectingCountry>
:
</emphasis>
<collectingRegion id="657AF81CFFD0827CFD98FE798CAEF6C1" box="[567,653,433,454]" country="Mexico" name="Yucatan" pageId="10" pageNumber="649">YUCATÁN</collectingRegion>
,
<location id="A2616025FFD0827CFF02FE038EFDF6E5" LSID="urn:lsid:plazi:treatment:2F1787E8FFD2827CFCE4FE868FE9F54F:A2616025FFD0827CFF02FE038EFDF6E5" box="[173,222,459,480]" country="Mexico" name="Arco" pageId="10" pageNumber="649" stateProvince="Yucatan">Arco</location>
,
<quantity id="60469B1BFFD0827CFF47FE038F3FF6E5" box="[232,284,459,480]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="649" unit="m" value="16.0">
<elevation id="2C93D1CDFFD0827CFF47FE038F3FF6E5" box="[232,284,459,480]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="649" unit="m" value="16.0">16 m</elevation>
</quantity>
</materialsCitation>
,
<geoCoordinate id="C28A5039FFD0827CFE88FE038FB3F6E5" box="[295,400,459,480]" degrees="21.0918" direction="north" orientation="latitude" pageId="10" pageNumber="649" precision="5" value="21.0918">
21.0918°
<collectingRegion id="657AF81CFFD0827CFED1FE038FB3F6E5" box="[382,400,459,480]" country="Mexico" name="N" pageId="10" pageNumber="649">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="C28A5039FFD0827CFE35FE038C25F6E5" box="[410,518,459,480]" degrees="88.6654" direction="west" orientation="longitude" pageId="10" pageNumber="649" precision="5" value="-88.6654">
88.6654°
<collectingRegion id="657AF81CFFD0827CFE40FE038C25F6E5" box="[495,518,459,480]" country="Mexico" name="W" pageId="10" pageNumber="649">W</collectingRegion>
</geoCoordinate>
,
<materialsCitation id="17D63CA3FFD0827CFDBEFE038C42F6FE" ID-GBIF-Occurrence="3330927302" collectingDate="1965-09-23" collectionCode="AMNH" country="Mexico" pageId="10" pageNumber="649" specimenCode="ADP056075" specimenCount="1" stateProvince="W" typeStatus="paratype">
<date id="D300103EFFD0827CFDBEFE038EFFF6FE" pageId="10" pageNumber="649" value="1965-09-23">
<collectingDate id="C344E9D6FFD0827CFDBEFE038EFFF6FE" pageId="10" pageNumber="649" value="1965-09-23">23 September 1965</collectingDate>
</date>
(
<collectionCode id="C1AFAE3BFFD0827CFF45FE2E8F15F6FE" box="[234,310,486,507]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="10" pageNumber="649">AMNH</collectionCode>
:
<specimenCode id="F7189E85FFD0827CFEEFFE2E8F94F6FE" box="[320,439,486,507]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="10" pageNumber="649">ADP056075</specimenCode>
, female
<typeStatus id="7805885CFFD0827CFDA7FE2E8C7FF6FE" box="[520,604,486,507]" pageId="10" pageNumber="649" type="paratype">paratype</typeStatus>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A70136FEFFD0827CFF6AFDC88FE9F54F" blockId="10.[173,659,193,586]" pageId="10" pageNumber="649">
<emphasis id="95CAEAECFFD0827CFF6AFDC88FE5F510" bold="true" box="[197,454,512,533]" pageId="10" pageNumber="649">Geographic Distribution.</emphasis>
<materialsCitation id="17D63CA3FFD0827CFE7EFDC88FE6F54F" ID-GBIF-Occurrence="3330927303" country="Mexico" location="Yucatan" pageId="10" pageNumber="649" specimenCount="1" stateProvince="Yucatan">
This species is currently known from the
<typeStatus id="7805885CFFD0827CFE2CFDD38F8FF535" box="[387,428,539,560]" pageId="10" pageNumber="649">type</typeStatus>
locality and the
<collectingRegion id="657AF81CFFD0827CFDE9FDD38CB0F535" box="[582,659,539,560]" country="Mexico" name="Yucatan" pageId="10" pageNumber="649">Yucatán</collectingRegion>
peninsula of
<collectingCountry id="DFA9766EFFD0827CFE84FDFD8F50F54F" box="[299,371,565,586]" name="Mexico" pageId="10" pageNumber="649">Mexico</collectingCountry>
(
<figureCitation id="3F852A7BFFD0827CFE2DFDFD8F9EF54F" box="[386,445,565,586]" captionStart="Fig" captionStartId="7.[175,206,1334,1353]" captionTargetBox="[151,1173,800,1317]" captionTargetId="figure-155@7.[151,1173,800,1317]" captionTargetPageId="7" captionText="Fig. 5. Distribution map of known localities for Agra wickhami." figureDoi="http://doi.org/10.5281/zenodo.4788807" httpUri="https://zenodo.org/record/4788807/files/figure.png" pageId="10" pageNumber="649">Fig. 5</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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