236 lines
33 KiB
XML
236 lines
33 KiB
XML
<document id="66E3F6F1737AAF6A73B318ED3770FCB1" ID-DOI="10.5281/zenodo.169435" ID-GBIF-Dataset="dff6ff7c-0fd1-4149-b3ae-4e3c66ca7ed1" ID-ISSN="11755326" ID-Zenodo-Dep="169435" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461031555560" checkinUser="plazi" docAuthor="Vilkamaa, Pekka & Hippa, Heikki" docDate="2004" docId="2C48879DFFDCFFC5FEB2B4F8FD2D023F" docLanguage="en" docName="zt00699.pdf" docOrigin="Zootaxa 699" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Xenosciara Vilkamaa & Hippa, 2004, gen. n." docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="D071FFE5FFD5FFCBFFBAB67CFF9B0129" masterDocTitle="The genus Xenosciara gen. n. and the phylogeny of the Sciaridae (Diptera)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="10" updateTime="1698573401539" updateUser="plazi">
|
||
<mods:mods id="F606B82F71319A095F8661AD62BD4E0E" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="D2BFF22975BAC13F298774DB34AF5D85">
|
||
<mods:title id="8A7010D44BFD4C3CD55AA0B9C472F23C">The genus Xenosciara gen. n. and the phylogeny of the Sciaridae (Diptera)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="8C3ACB4B59D739F2136EC384510AD800" type="personal">
|
||
<mods:role id="E2E0421DF9F2049E485B33BE7B44308D">
|
||
<mods:roleTerm id="4F6F0127151074A1DB3A27943DA6CE28">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C56D19CBB6E45DED2B9183AC6A5C8AB5">Vilkamaa, Pekka</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="EAAC1D46C5B517C789CA3578D00973FF" type="personal">
|
||
<mods:role id="9836195FF13902CD5C1EFB36A7862E63">
|
||
<mods:roleTerm id="DCF21BC242703D67114DC22D55F23F13">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="ADF52AB27374BB2CA1B391ECDFAE0794">Hippa, Heikki</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="43A213F05B734FD3266F5182A2181C4D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="1E07ECF54856E2B27DBF91F90B294824" type="host">
|
||
<mods:titleInfo id="CE21A1EE14C6EBFB0AC6EF900F44FF8F">
|
||
<mods:title id="ECEB7B0E269291285E5189840ED93DA7">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1C6C121A46A53240FD67864DB6B8C605">
|
||
<mods:date id="8A29CC5F7114FDF689A698B8271F8E87">2004</mods:date>
|
||
<mods:detail id="2DA08F6250EF2EC746B9003E56F3A44A" type="volume">
|
||
<mods:number id="E8BE10F118DCE79357788D6D39A63766">699</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="C378D585CEAE42561472C95CA7BF994A" unit="page">
|
||
<mods:start id="B6EEA9FEED7108038CF67D766AFFE1BE">1</mods:start>
|
||
<mods:end id="DFC2283276B0B4AB9AD47F0E189C9EE6">24</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="7A929FE328E1DF59F077D09295116CCE">journal article</mods:classification>
|
||
<mods:identifier id="C0BC90069637D967300FE04D3676915F" type="DOI">10.5281/zenodo.169435</mods:identifier>
|
||
<mods:identifier id="96D4B676445B5D3569DDEB057B60C1C3" type="GBIF-Dataset">dff6ff7c-0fd1-4149-b3ae-4e3c66ca7ed1</mods:identifier>
|
||
<mods:identifier id="01779E79D2B6706B56E74EF135BA0689" type="ISSN">11755326</mods:identifier>
|
||
<mods:identifier id="806880BD625A325044ACE9CFDA234C2C" type="Zenodo-Dep">169435</mods:identifier>
|
||
<mods:identifier id="C31D39A51FAD2AC8ED8373165DB18DB7" type="ZooBank">327C8C4A-B3BB-40C1-8229-A9B342731D11</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="2C48879DFFDCFFC5FEB2B4F8FD2D023F" ID-DOI="http://doi.org/10.5281/zenodo.6270940" ID-GBIF-Taxon="119608144" ID-Zenodo-Dep="6270940" LSID="urn:lsid:plazi:treatment:2C48879DFFDCFFC5FEB2B4F8FD2D023F" httpUri="http://treatment.plazi.org/id/2C48879DFFDCFFC5FEB2B4F8FD2D023F" lastPageId="14" lastPageNumber="14" pageId="9" pageNumber="10">
|
||
<subSubSection id="ECFB6500FFDCFFC2FEB2B4F8FDE903B7" box="[264,626,644,670]" pageId="9" pageNumber="10" type="nomenclature">
|
||
<paragraph id="A45E368BFFDCFFC2FEB2B4F8FDE903B7" blockId="9.[264,626,644,670]" box="[264,626,644,670]" pageId="9" pageNumber="10">
|
||
<heading id="FF1681E7FFDCFFC2FEB2B4F8FDE903B7" bold="true" box="[264,626,644,670]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
|
||
<emphasis id="9695EA99FFDCFFC2FEB2B4F8FDE903B7" bold="true" box="[264,626,644,670]" pageId="9" pageNumber="10">
|
||
The genus
|
||
<taxonomicName id="63E14D08FFDCFFC2FE35B4F8FD8303B7" ID-CoL="88Y7" box="[399,536,644,670]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="9695EA99FFDCFFC2FE35B4F8FD8303B7" bold="true" box="[399,536,644,670]" italics="true" pageId="9" pageNumber="10">Xenosciara</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8DA657E2FFDCFFC2FDA5B4F8FDE903B7" box="[543,626,644,670]" pageId="9" pageNumber="10" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ECFB6500FFDCFFC5FEB2B4B3FD2D023F" lastPageId="14" lastPageNumber="15" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="A45E368BFFDCFFC2FEB2B4B3FD5A03C0" blockId="9.[264,705,719,745]" box="[264,705,719,745]" pageId="9" pageNumber="10">
|
||
Typespecies
|
||
<taxonomicName id="63E14D08FFDCFFC2FE12B4B3FDE003C1" box="[424,635,719,744]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="invisa" status="sp. nov.">
|
||
<emphasis id="9695EA99FFDCFFC2FE12B4B3FDE003C1" box="[424,635,719,744]" italics="true" pageId="9" pageNumber="10">Xenosciara invisa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8DA657E2FFDCFFC2FD38B4B3FD5A03C0" box="[642,705,719,745]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="A45E368BFFDCFFC2FEB2B563FE170211" blockId="9.[264,1323,799,1905]" box="[264,396,799,824]" pageId="9" pageNumber="10">
|
||
<emphasis id="9695EA99FFDCFFC2FEB2B563FE170211" box="[264,396,799,824]" italics="true" pageId="9" pageNumber="10">Characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A45E368BFFDCFFC2FE82B53BFCC20248" blockId="9.[264,1323,799,1905]" box="[312,857,839,865]" pageId="9" pageNumber="10">
|
||
Mediumsized
|
||
<taxonomicName id="63E14D08FFDCFFC2FE53B53BFDC20248" box="[489,601,839,865]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
|
||
, wing length
|
||
<quantity id="63199B6EFFDCFFC2FD4DB53BFCCE0248" box="[759,853,839,865]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="10" unit="mm" value="3.2">3.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A45E368BFFDCFFC2FE82B512FB8C04C8" blockId="9.[264,1323,799,1905]" pageId="9" pageNumber="10">
|
||
<emphasis id="9695EA99FFDCFFC2FE82B512FEE002A1" bold="true" box="[312,379,878,904]" pageId="9" pageNumber="10">Head</emphasis>
|
||
(
|
||
<figureCitation id="3CDA2A0EFFDCFFC2FE37B513FE4D02A0" box="[397,470,879,905]" captionStart="FIGURE 6" captionStartId="10.[264,368,1464,1488]" captionTargetBox="[354,1206,449,1463]" captionTargetId="figure@10.[354,1233,437,1471]" captionTargetPageId="10" captionText="FIGURE 6. Xenosciara invisa sp. n. (holotype). — A: Head, frontal view, — B: Maxillary palp, basal palpomeres, dorsal view, apical palpomere, mesioventral view, — C: antennal flagellomere 4, ventral view, — D: apex of flagellomere 14, ventral view. Scale for A 0.50 mm, for B and C 0.10 mm, for D 0.05 mm." httpUri="https://zenodo.org/record/169441/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
|
||
). Eye bridge present, 4 facets wide. Eyes without interfacetal microtrichia, with interfacetal setae which extend well beyond the curvature of the facets. Ocelli three, in an equilateral triangle. Anterior vertex short, setose. Prefrons setose, the setae unequal in size. Clypeus distinctly separated from prefrons, setose. Antennal scape and pedicel (
|
||
<figureCitation id="3CDA2A0EFFDCFFC2FED4B273FE2F0500" box="[366,436,1039,1065]" captionStart="FIGURE 6" captionStartId="10.[264,368,1464,1488]" captionTargetBox="[354,1206,449,1463]" captionTargetId="figure@10.[354,1233,437,1471]" captionTargetPageId="10" captionText="FIGURE 6. Xenosciara invisa sp. n. (holotype). — A: Head, frontal view, — B: Maxillary palp, basal palpomeres, dorsal view, apical palpomere, mesioventral view, — C: antennal flagellomere 4, ventral view, — D: apex of flagellomere 14, ventral view. Scale for A 0.50 mm, for B and C 0.10 mm, for D 0.05 mm." httpUri="https://zenodo.org/record/169441/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
|
||
A) subglobular, with several ventral setae, flagellum with 14 flagellomeres which are subequal in size, the body about two and a half times as long as wide, the neck short, about onethird the width of the body, the vestiture of the flagellomeral body (
|
||
<figureCitation id="3CDA2A0EFFDCFFC2FB41B223FE8C0588" captionStart="FIGURE 6" captionStartId="10.[264,368,1464,1488]" captionTargetBox="[354,1206,449,1463]" captionTargetId="figure@10.[354,1233,437,1471]" captionTargetPageId="10" captionText="FIGURE 6. Xenosciara invisa sp. n. (holotype). — A: Head, frontal view, — B: Maxillary palp, basal palpomeres, dorsal view, apical palpomere, mesioventral view, — C: antennal flagellomere 4, ventral view, — D: apex of flagellomere 14, ventral view. Scale for A 0.50 mm, for B and C 0.10 mm, for D 0.05 mm." httpUri="https://zenodo.org/record/169441/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
|
||
C) with long setalike sensillae arising from weak platelike depressions, with a few short setalike sensillae, without socketed setae, without hyaline sensillae, the ultimate flagellomere (
|
||
<figureCitation id="3CDA2A0EFFDCFFC2FE17B2ABFE6F05D8" box="[429,500,1239,1265]" captionStart="FIGURE 6" captionStartId="10.[264,368,1464,1488]" captionTargetBox="[354,1206,449,1463]" captionTargetId="figure@10.[354,1233,437,1471]" captionTargetPageId="10" captionText="FIGURE 6. Xenosciara invisa sp. n. (holotype). — A: Head, frontal view, — B: Maxillary palp, basal palpomeres, dorsal view, apical palpomere, mesioventral view, — C: antennal flagellomere 4, ventral view, — D: apex of flagellomere 14, ventral view. Scale for A 0.50 mm, for B and C 0.10 mm, for D 0.05 mm." httpUri="https://zenodo.org/record/169441/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
|
||
D) apically with two sharp pegs, microtrichia present only basally on flagellomere 1. Maxillary palp with palpifer and three palpomeres; galea long, about as long as palpomere 1, apically branched; palpomere 1 unusually long, about four times as long as broad, palpomere 2 onethird of the length of palpomere 1, palpomere 3 about the length of palpomere 2, only slightly constricted basally; palpifer nonsetose, palpomere 1 with one large lateral seta and several weaker dorsal setae, with scattered hyaline sensillae dorsally, palpomeres 2–3 with numerous setae, all with blunt apex.
|
||
</paragraph>
|
||
<paragraph id="A45E368BFFDCFFC1FE82B392FA810077" blockId="9.[264,1323,799,1905]" lastBlockId="10.[264,1322,284,350]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
<emphasis id="9695EA99FFDCFFC2FE82B392FE0E0721" bold="true" box="[312,405,1518,1544]" pageId="9" pageNumber="10">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="3CDA2A0EFFDCFFC2FE1EB393FE770720" box="[420,492,1519,1545]" captionStart="FIGURE 7" captionStartId="11.[264,368,1674,1698]" captionTargetBox="[414,1174,685,1640]" captionTargetId="figure@11.[354,1233,643,1654]" captionTargetPageId="11" captionText="FIGURE 7. Xenosciara invisa sp. n. (holotype): Thorax, lateral view. Scale 0.50 mm. 1 = scutum, 2 = scutellum, 3 = metanotum (phragma), 4 = anterior pronotum, 5 = episternum 1, 6 = anterior anepisternum, 7 = anterior anepimeron, 8 = pleurotergite, 9 = katepisternum, 10 = pleural pit, 11 = episternum 3, 12 = halter, 13 = coxa 2, 14 = anterior spiracle, 15 = posterior spiracle." httpUri="https://zenodo.org/record/169442/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
|
||
). Dark brown, setae dark. Scutum broadly setose laterally, with shorter and longer setae, dorsocentral setae in three rows, with longer and shorter setae, acrostichal setae in one row which is present only on the anterior half. Scutellum with numerous setae of unequal length, with a few long curved setae posteromedially. Anterior pronotum with a few setae, posterior pronotum nonsetose, episternum 1 setose. Anepisternum with a deep vshaped cleft dorsally, nonsetose. Anepimeron with the anterior margin interrupted by the pleural pit, the posteroventral process reaching metepisternum and posteroventral margin of katepisternum, not shortened, the anterior margin ending ventrally at anterodorsal corner of episternum 3, nonsetose. Katepisternum high, the dorsal margin slightly curved, not angulated, nonsetose. Pleural pit at the middle of the dorsal margin of katepisternum. Laterotergite with anterior margin strongly sigmoid, nonsetose. Metepisternum nonsetose. Metanotum nonsetose, phragma intruding into the base of abdomen.
|
||
</paragraph>
|
||
<caption id="F09E6603FFDFFFC1FEB2B3C4FE7E071E" httpUri="https://zenodo.org/record/169441/files/figure.png" pageId="10" pageNumber="11" targetBox="[354,1206,449,1463]" targetPageId="10">
|
||
<paragraph id="A45E368BFFDFFFC1FEB2B3C4FE7E071E" blockId="10.[264,1323,1464,1591]" pageId="10" pageNumber="11">
|
||
<emphasis id="9695EA99FFDFFFC1FEB2B3C4FE1604F9" bold="true" box="[264,397,1464,1488]" pageId="10" pageNumber="11">FIGURE 6.</emphasis>
|
||
<taxonomicName id="63E14D08FFDFFFC1FE2FB3C5FDCC04F9" box="[405,599,1465,1488]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="invisa" status="sp. nov.">
|
||
<emphasis id="9695EA99FFDFFFC1FE2FB3C5FDCC04F9" box="[405,599,1465,1488]" italics="true" pageId="10" pageNumber="11">Xenosciara invisa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8DA657E2FFDFFFC1FDE5B3C4FD0104E6" box="[607,666,1464,1487]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
(holotype). — A: Head, frontal view, — B: Maxillary palp, basal palpomeres, dorsal view, apical palpomere, mesioventral view, — C: antennal flagellomere 4, ventral view, — D: apex of flagellomere 14, ventral view. Scale for A 0.50 mm, for B and C 0.10 mm, for D 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A45E368BFFDFFFC1FE82B0F2FBED0609" blockId="10.[264,1323,1678,1904]" pageId="10" pageNumber="11">
|
||
<emphasis id="9695EA99FFDFFFC1FE82B0F2FEE70781" bold="true" box="[312,380,1678,1704]" pageId="10" pageNumber="11">Wing</emphasis>
|
||
(
|
||
<figureCitation id="3CDA2A0EFFDFFFC1FE37B0F2FE550781" box="[397,462,1678,1704]" captionStart="FIGURE 8" captionStartId="12.[264,368,1168,1192]" captionTargetBox="[401,1224,306,1111]" captionTargetId="figure@12.[354,1233,282,1144]" captionTargetPageId="12" captionText="FIGURE 8. Xenosciara invisa sp. n. (holotype). — A: Apical part of front tibia, prolateral view, — B: Apical part of hind tibia, retrolateral view, — C: Wing, dorsal view, — D: Abdominal segment 5, lateral view. Scale for A, B and D 0.20 mm, for C 2.0 mm. 1 = tergite, 2 = sternite, 3 = spiracle, 4 = plaque." httpUri="https://zenodo.org/record/169443/files/figure.png" pageId="10" pageNumber="11">Fig.8</figureCitation>
|
||
). Hyaline, tinged with brown. R1 slightly shorter than R. rm unusually long. stCu present, C, R, R1, R5 setose, other veins dorsally nonsetose, all except C ventrally nonsetose; wing membrane nonsetose, the setae on posterior wing margin only on dorsal side. Haltere as long as coxae, the knob with a dorsal stripe of setae.
|
||
</paragraph>
|
||
<paragraph id="A45E368BFFDFFFC0FE82B152FD9B0367" blockId="10.[264,1323,1678,1904]" lastBlockId="11.[264,1323,284,590]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
<emphasis id="9695EA99FFDFFFC1FE82B152FEE80661" bold="true" box="[312,371,1838,1864]" pageId="10" pageNumber="11">Legs</emphasis>
|
||
(
|
||
<figureCitation id="3CDA2A0EFFDFFFC1FE38B152FE530661" box="[386,456,1838,1864]" captionStart="FIGURE 8" captionStartId="12.[264,368,1168,1192]" captionTargetBox="[401,1224,306,1111]" captionTargetId="figure@12.[354,1233,282,1144]" captionTargetPageId="12" captionText="FIGURE 8. Xenosciara invisa sp. n. (holotype). — A: Apical part of front tibia, prolateral view, — B: Apical part of hind tibia, retrolateral view, — C: Wing, dorsal view, — D: Abdominal segment 5, lateral view. Scale for A, B and D 0.20 mm, for C 2.0 mm. 1 = tergite, 2 = sternite, 3 = spiracle, 4 = plaque." httpUri="https://zenodo.org/record/169443/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
|
||
). Unicolorous, slightly paler brown than thorax; slender. Coxae long, several times as long as broad, metacoxa slightly shorter than the others. Tibiae with both socketed setae and nonsocketed setae (enlarged microtrichia), both subequal in size, middle and hind tibia with a few enlarged dorsal and lateral setae, on front tibia only apically, front tibia prolaterally with a subapical patch of setae, hind tibia with a retrolateral subapical transverse row of strong setae, without a dorsal row of strong setae. Tibial spurs 1+2+2, all subequal in size, longer than the apical width of tibia. Tarsomeres short, basitarsomere less than half the length of tibia, the two basal tarsomeres with enlarged setae ventrally/lateroventrally, tarsomeres 3 and 4 only apically; tarsal claws simple, pulvilli and empodium branched.
|
||
</paragraph>
|
||
<caption id="F09E6603FFDEFFC0FEB2B0F6FB080623" httpUri="https://zenodo.org/record/169442/files/figure.png" pageId="11" pageNumber="12" targetBox="[414,1174,685,1640]" targetPageId="11">
|
||
<paragraph id="A45E368BFFDEFFC0FEB2B0F6FB080623" blockId="11.[264,1322,1674,1802]" pageId="11" pageNumber="12">
|
||
<emphasis id="9695EA99FFDEFFC0FEB2B0F6FE10078B" bold="true" box="[264,395,1674,1698]" pageId="11" pageNumber="12">FIGURE 7.</emphasis>
|
||
<taxonomicName id="63E14D08FFDEFFC0FE28B0F7FDC8078B" box="[402,595,1675,1698]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="invisa" status="sp. nov.">
|
||
<emphasis id="9695EA99FFDEFFC0FE28B0F7FDC8078B" box="[402,595,1675,1698]" italics="true" pageId="11" pageNumber="12">Xenosciara invisa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8DA657E2FFDEFFC0FDE0B0F7FD08078B" box="[602,659,1675,1698]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
|
||
(holotype): Thorax, lateral view. Scale 0.50 mm. 1 = scutum, 2 = scutellum, 3 = metanotum (phragma), 4 = anterior pronotum, 5 = episternum 1, 6 = anterior anepisternum, 7 = anterior anepimeron, 8 = pleurotergite, 9 = katepisternum, 10 = pleural pit, 11 = episternum 3, 12 = halter, 13 = coxa 2, 14 = anterior spiracle, 15 = posterior spiracle.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F09E6603FFD9FFC7FEB2B2ECFECC0426" httpUri="https://zenodo.org/record/169443/files/figure.png" pageId="12" pageNumber="13" targetBox="[401,1224,306,1111]" targetPageId="12">
|
||
<paragraph id="A45E368BFFD9FFC7FEB2B2ECFECC0426" blockId="12.[264,1323,1168,1295]" pageId="12" pageNumber="13">
|
||
<emphasis id="9695EA99FFD9FFC7FEB2B2ECFE100581" bold="true" box="[264,395,1168,1192]" pageId="12" pageNumber="13">FIGURE 8.</emphasis>
|
||
<taxonomicName id="63E14D08FFD9FFC7FE2BB2EDFDCB0581" box="[401,592,1169,1192]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="invisa" status="sp. nov.">
|
||
<emphasis id="9695EA99FFD9FFC7FE2BB2EDFDCB0581" box="[401,592,1169,1192]" italics="true" pageId="12" pageNumber="13">Xenosciara invisa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8DA657E2FFD9FFC7FDECB2ECFD14058E" box="[598,655,1168,1191]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
|
||
(holotype). — A: Apical part of front tibia, prolateral view, — B: Apical part of hind tibia, retrolateral view, — C: Wing, dorsal view, — D: Abdominal segment 5, lateral view. Scale for A, B and D 0.20 mm, for C 2.0 mm. 1 = tergite, 2 = sternite, 3 = spiracle, 4 = plaque.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A45E368BFFD9FFC7FE82B31FFD1A061C" blockId="12.[264,1323,1379,1885]" pageId="12" pageNumber="13">
|
||
<emphasis id="9695EA99FFD9FFC7FE82B31FFE290454" bold="true" box="[312,434,1379,1405]" pageId="12" pageNumber="13">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="3CDA2A0EFFD9FFC7FE7CB31FFD8B0454" box="[454,528,1379,1405]" captionStart="FIGURE 8" captionStartId="12.[264,368,1168,1192]" captionTargetBox="[401,1224,306,1111]" captionTargetId="figure@12.[354,1233,282,1144]" captionTargetPageId="12" captionText="FIGURE 8. Xenosciara invisa sp. n. (holotype). — A: Apical part of front tibia, prolateral view, — B: Apical part of hind tibia, retrolateral view, — C: Wing, dorsal view, — D: Abdominal segment 5, lateral view. Scale for A, B and D 0.20 mm, for C 2.0 mm. 1 = tergite, 2 = sternite, 3 = spiracle, 4 = plaque." httpUri="https://zenodo.org/record/169443/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
|
||
). Brown. Tergites 2–7 with one pair of plaques. Male sternum 9 united with hypopygium.
|
||
<emphasis id="9695EA99FFD9FFC7FDF9B3F7FD7D048C" bold="true" box="[579,742,1419,1445]" pageId="12" pageNumber="13">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="3CDA2A0EFFD9FFC7FD40B3F7FCDD048C" box="[762,838,1419,1445]" captionStart="FIGURE 9" captionStartId="13.[264,368,1360,1384]" captionTargetBox="[364,1231,300,1314]" captionTargetId="figure@13.[354,1233,278,1352]" captionTargetPageId="13" captionText="FIGURE 9. Xenosciara invisa sp. n. (holotype). — A: Hypopygium, lateral view, — B: Hypopygium, dorsal view, — C: Hypopygium, ventral view, — D: Hypopygium, posterior view. Scale 0.20 mm. 1 = gonocoxite, 2 = gonostylus, 3 = basolateral apophysis of gonostylus, 4 = apical megasetae of gonostylus, 5 = tergite 9, 6 = cercus, 7 = tegmen, 8 = apodeme of tegmen (parameral apodeme), 9 = sternite 10, 10 = apicodorsal flagellate seta of gonocoxite." httpUri="https://zenodo.org/record/169444/files/figure.png" pageId="12" pageNumber="13">Fig. 9</figureCitation>
|
||
). Tergite 9 large, apically rather evenly curved. Gonocoxites ventrally completely fused, gonocoxal apodemes short, weakly sclerotized, ending far posterior from the anterior margin of hypopygium, the apodemes of both sides medially united only by membrane. Gonostylus flexible in vertical plane, without apical tooth, with two apical megasetae, without flagellate setae, with the basomesial structures simple, with the basolateral apophysis strong, basoventral in position. Parameres weak, united to form a tegmen, parameral apodemes transverse, free from gonocoxal apodemes. Aedeagus without teeth or microtrichia, aedeagal apodeme/genital rod absent; a setose membrane extending from the ventral apicomesial part of gonocoxites to aedeagus. Tergite 10 membraneous, bilobed, with several short setae. Cercus flattened, setose only on posterior margin.
|
||
</paragraph>
|
||
<paragraph id="A45E368BFFD9FFC7FE82B13FFD130674" blockId="12.[264,1323,1379,1885]" box="[312,648,1859,1885]" pageId="12" pageNumber="13">Female characters unknown.</paragraph>
|
||
<caption id="F09E6603FFD8FFC6FEB2B32CFC0704D4" httpUri="https://zenodo.org/record/169444/files/figure.png" pageId="13" pageNumber="14" targetBox="[364,1231,300,1314]" targetPageId="13">
|
||
<paragraph id="A45E368BFFD8FFC6FEB2B32CFC0704D4" blockId="13.[264,1323,1360,1533]" pageId="13" pageNumber="14">
|
||
<emphasis id="9695EA99FFD8FFC6FEB2B32CFE170441" bold="true" box="[264,396,1360,1384]" pageId="13" pageNumber="14">FIGURE 9.</emphasis>
|
||
<taxonomicName id="63E14D08FFD8FFC6FE2EB32CFDCE044E" box="[404,597,1360,1383]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="invisa" status="sp. nov.">
|
||
<emphasis id="9695EA99FFD8FFC6FE2EB32CFDCE044E" box="[404,597,1360,1383]" italics="true" pageId="13" pageNumber="14">Xenosciara invisa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8DA657E2FFD8FFC6FDE6B32CFD0C044E" box="[604,663,1360,1383]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
|
||
(holotype). — A: Hypopygium, lateral view, — B: Hypopygium, dorsal view, — C: Hypopygium, ventral view, — D: Hypopygium, posterior view. Scale 0.20 mm. 1 = gonocoxite, 2 = gonostylus, 3 = basolateral apophysis of gonostylus, 4 = apical megasetae of gonostylus, 5 = tergite 9, 6 = cercus, 7 = tegmen, 8 = apodeme of tegmen (parameral apodeme), 9 = sternite 10, 10 = apicodorsal flagellate seta of gonocoxite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A45E368BFFD8FFC6FEB2B020FD96075C" blockId="13.[264,1323,1628,1894]" box="[264,525,1628,1653]" pageId="13" pageNumber="14">
|
||
<emphasis id="9695EA99FFD8FFC6FEB2B020FD96075C" box="[264,525,1628,1653]" italics="true" pageId="13" pageNumber="14">Diagnostic characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A45E368BFFD8FFC6FE82B0F8FA9F063F" blockId="13.[264,1323,1628,1894]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="63E14D08FFD8FFC6FE82B0F8FE2607B4" box="[312,445,1668,1693]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFD8FFC6FE82B0F8FE2607B4" box="[312,445,1668,1693]" italics="true" pageId="13" pageNumber="14">Xenosciara</emphasis>
|
||
</taxonomicName>
|
||
differs from all known
|
||
<taxonomicName id="63E14D08FFD8FFC6FD5AB0F8FCD607B7" box="[736,845,1668,1694]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Sciarinae">Sciarinae</taxonomicName>
|
||
by each of the following characters: 1) the very long palpomere 1, 2) ventrally fused male gonocoxites, 3) gonostyli movable in a vertical direction, 4) vertical tegmen, 5) setae only on the margin of the cercus, and 6) a setose membrane from the fused ventral part of the gonocoxites covering the aedeagus.
|
||
</paragraph>
|
||
<paragraph id="A45E368BFFD8FFC5FE82B158FD2D023F" blockId="13.[264,1323,1628,1894]" lastBlockId="14.[264,1323,284,790]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
In its habitus,
|
||
<taxonomicName id="63E14D08FFD8FFC6FE67B158FDF80614" box="[477,611,1828,1853]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFD8FFC6FE67B158FDF80614" box="[477,611,1828,1853]" italics="true" pageId="13" pageNumber="14">Xenosciara</emphasis>
|
||
</taxonomicName>
|
||
is a normal, unspecialized member of the
|
||
<taxonomicName id="63E14D08FFD8FFC6FBE9B158FB5F0617" box="[1107,1220,1828,1854]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
|
||
. With its long maxillary palp,
|
||
<taxonomicName id="63E14D08FFD8FFC6FDB0B130FD0B064C" box="[522,656,1868,1893]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFD8FFC6FDB0B130FD0B064C" box="[522,656,1868,1893]" italics="true" pageId="13" pageNumber="14">Xenosciara</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="63E14D08FFD8FFC6FCA6B130FBFB064F" authority="Tuomikoski" authorityName="Tuomikoski" box="[796,1120,1868,1894]" class="Insecta" family="Sciaridae" genus="Dolichosciara" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFD8FFC6FCA6B130FC5F064C" box="[796,964,1868,1893]" italics="true" pageId="13" pageNumber="14">Dolichosciara</emphasis>
|
||
Tuomikoski
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="63E14D08FFD8FFC6FBD5B130FABD064F" authority="Frey" authorityName="Frey" box="[1135,1318,1868,1894]" class="Insecta" family="Sciaridae" genus="Prosciara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFD8FFC6FBD5B130FB78064C" box="[1135,1251,1868,1893]" italics="true" pageId="13" pageNumber="14">Prosciara</emphasis>
|
||
Frey
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="63E14D08FFDBFFC5FE80B760FD86001F" authority="Mohrig" authorityName="Mohrig" box="[314,541,284,310]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFDBFFC5FE80B760FE21001C" box="[314,442,284,309]" italics="true" pageId="14" pageNumber="15">Keilbachia</emphasis>
|
||
Mohrig
|
||
</taxonomicName>
|
||
, but in these groups palpomere 3 is elongated and distinctly longer than the other palpomeres, and only
|
||
<taxonomicName id="63E14D08FFDBFFC5FD00B738FCA40074" box="[698,831,324,349]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFDBFFC5FD00B738FCA40074" box="[698,831,324,349]" italics="true" pageId="14" pageNumber="15">Xenosciara</emphasis>
|
||
</taxonomicName>
|
||
has palpomere 1 strongly elongated. The basoventrally fused gonocoxites is not a rare character in the
|
||
<taxonomicName id="63E14D08FFDBFFC5FC6EB710FBDA00AF" box="[980,1089,364,390]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
|
||
and the fused basal part can take up to half the length of the coxites in some groups (
|
||
<taxonomicName id="63E14D08FFDBFFC5FB88B7E8FE6600FF" authority="Hippa, Vilkamaa & Mohrig" authorityName="Hippa, Vilkamaa & Mohrig" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFDBFFC5FB88B7E8FB4B0084" box="[1074,1232,404,429]" italics="true" pageId="14" pageNumber="15">Claustropyga</emphasis>
|
||
Hippa, Vilkamaa & Mohrig
|
||
</taxonomicName>
|
||
), but there are no other taxa in which the ventral aspect of hypopygium resembles that of
|
||
<taxonomicName id="63E14D08FFDBFFC5FDA6B798FD3A00D4" box="[540,673,484,509]" class="Insecta" family="Sciaridae" genus="Xenosciara" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFDBFFC5FDA6B798FD3A00D4" box="[540,673,484,509]" italics="true" pageId="14" pageNumber="15">Xenosciara</emphasis>
|
||
</taxonomicName>
|
||
. A reminiscent structural
|
||
<typeStatus id="7B5A8829FFDBFFC5FC6EB798FB9D00D7" box="[980,1030,484,510]" pageId="14" pageNumber="15">type</typeStatus>
|
||
is found in several other Sciaroidea, however. The gonostyli that work in a vertical direction are quite exceptional in the
|
||
<taxonomicName id="63E14D08FFDBFFC5FEEFB448FE5E0367" box="[341,453,564,590]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
|
||
, but occur in some taxa of the other Sciaroidea (e.g.
|
||
<taxonomicName id="63E14D08FFDBFFC5FBF9B448FDF6035F" authority="Loew, Rhynchoheterotricha Freeman" authorityName="Loew, Rhynchoheterotricha Freeman" class="Insecta" family="Diadocidiidae" genus="Heterotricha" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFDBFFC5FBF9B448FB420364" box="[1091,1241,564,589]" italics="true" pageId="14" pageNumber="15">Heterotricha</emphasis>
|
||
Loew,
|
||
<emphasis id="9695EA99FFDBFFC5FEB2B420FE65035C" box="[264,510,604,629]" italics="true" pageId="14" pageNumber="15">Rhynchoheterotricha</emphasis>
|
||
Freeman
|
||
</taxonomicName>
|
||
and a few groups of the
|
||
<taxonomicName id="63E14D08FFDBFFC5FC2EB420FBD7035F" box="[916,1100,604,630]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="family">Cecidomyiidae</taxonomicName>
|
||
). The
|
||
<typeStatus id="7B5A8829FFDBFFC5FB2EB420FB5C035F" box="[1172,1223,604,630]" pageId="14" pageNumber="15">type</typeStatus>
|
||
of association of the gonocoxites and the aedeagus/tegmen complex does not resemble that of any other
|
||
<taxonomicName id="63E14D08FFDBFFC5FEF6B4D0FE2203EF" box="[332,441,684,710]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subFamily" subFamily="Sciarinae">Sciarinae</taxonomicName>
|
||
but is reminiscent of
|
||
<taxonomicName id="63E14D08FFDBFFC5FD06B4D0FCD503EC" box="[700,846,684,709]" class="Insecta" family="Sciaridae" genus="Sciarotricha" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9695EA99FFDBFFC5FD06B4D0FCD503EC" box="[700,846,684,709]" italics="true" pageId="14" pageNumber="15">Sciarotricha</emphasis>
|
||
</taxonomicName>
|
||
in terms of the membranous connection between the aedeagus and gonocoxites.
|
||
<emphasis id="9695EA99FFDBFFC5FD61B4A8FCE803C4" box="[731,883,724,749]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="63E14D08FFDBFFC5FD61B4A8FCF403C4" box="[731,879,724,749]" class="Insecta" family="Sciaridae" genus="Sciarotricha" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Sciarotricha</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
however, has separate parameres and the membrane is only microtrichose.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |