280 lines
31 KiB
XML
280 lines
31 KiB
XML
<document id="296567CFD67EA017F0009D7C5D18E97A" ID-CLB-Dataset="20262" ID-DOI="10.1080/00222933.2022.2025941" ID-GBIF-Dataset="78c47ef9-5408-4811-9c49-6e5839dec636" ID-ISSN="1464-5262" ID-Zenodo-Dep="6351834" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647255337178" checkinUser="plazi" docAuthor="Maldaner, Maria Eduarda & Vaz-de-Mello, Fernando Z." docDate="2022" docId="2C226743CF05FF95FF45FBEE424DF9D2" docLanguage="en" docName="JNATHIST.55.47-48.2999-3006.pdf" docOrigin="Journal of Natural History 55 (47 - 48)" docSource="http://dx.doi.org/10.1080/00222933.2022.2025941" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Dichotomius (Selenocopris) speciosus" docType="key" docVersion="5" lastPageNumber="3001" masterDocId="D01B1F3BCF04FF97FFE5FFB24647FF8A" masterDocTitle="New data and species for the Dichotomius speciosus (Waterhouse, 1891) species group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="3006" masterPageNumber="2999" pageNumber="3000" updateTime="1699317907125" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="206EC1FD3E94B6E9581051E02F9068B3">New data and species for the Dichotomius speciosus (Waterhouse, 1891) species group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="D08232F365E9C1EC8659C37431C1B7FF" type="personal">
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<mods:namePart id="8BC2AD3B5DDFEA27B2C500F83DC5B116">Maldaner, Maria Eduarda</mods:namePart>
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<mods:namePart id="02995D799D32C744AC2E9FB476FB144D">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="5824E7918673D41A8B015EFF256DC124">Journal of Natural History</mods:title>
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<mods:part id="01A74D47994288AF14935BF16B0BE983">
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<mods:date id="B61F0054924BBD8D24176E016B4E78F5">2022</mods:date>
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<mods:number id="8C2F771761F4B2D149319F15E0C8EC6D">2022-03-08</mods:number>
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<mods:number id="F5B74424E99588547030876BD8F904A4">55</mods:number>
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<mods:number id="931F4335534013499A0B17FC66222F0A">47 - 48</mods:number>
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<mods:url id="AF89F89B7B28FA05D3605688D5C28248">http://dx.doi.org/10.1080/00222933.2022.2025941</mods:url>
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<treatment id="2C226743CF05FF95FF45FBEE424DF9D2" ID-DOI="http://doi.org/10.5281/zenodo.6353835" ID-GBIF-Taxon="193990288" ID-Zenodo-Dep="6353835" LSID="urn:lsid:plazi:treatment:2C226743CF05FF95FF45FBEE424DF9D2" httpUri="http://treatment.plazi.org/id/2C226743CF05FF95FF45FBEE424DF9D2" lastPageId="2" lastPageNumber="3001" pageId="1" pageNumber="3000">
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<subSubSection id="EC9185DECF05FF96FF45FBEE42C3FBFF" box="[160,1156,1116,1141]" pageId="1" pageNumber="3000" type="nomenclature">
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<paragraph id="A434D655CF05FF96FF45FBEE42C3FBFF" blockId="1.[160,1156,1116,1176]" box="[160,1156,1116,1141]" pageId="1" pageNumber="3000">
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Identification key to species of the
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<taxonomicName id="638BADD6CF05FF96FDD9FBEE4593FBFF" ID-CoL="35ND3" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[572,980,1116,1141]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="3000" phylum="Arthropoda" rank="species" species="speciosus" subGenus="Selenocopris">
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<emphasis id="96FF0A47CF05FF96FDD9FBEE4482FBFF" box="[572,709,1116,1141]" italics="true" pageId="1" pageNumber="3000">Dichotomius</emphasis>
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(
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<emphasis id="96FF0A47CF05FF96FD33FBEE451AFBFF" box="[726,861,1116,1141]" italics="true" pageId="1" pageNumber="3000">Selenocopris</emphasis>
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)
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<emphasis id="96FF0A47CF05FF96FC8AFBEE4593FBFF" box="[879,980,1116,1141]" italics="true" pageId="1" pageNumber="3000">speciosus</emphasis>
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</taxonomicName>
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species group.
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</paragraph>
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</subSubSection>
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<subSubSection id="EC9185DECF05FF96FF45FBCD452BFB12" box="[160,876,1151,1176]" pageId="1" pageNumber="3000" type="multiple">
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<paragraph id="A434D655CF05FF96FF45FBCD452BFB12" blockId="1.[160,1156,1116,1176]" box="[160,876,1151,1176]" pageId="1" pageNumber="3000">
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Figure numbers in italics refer to figures in
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<bibRefCitation id="C01AABA4CF05FF96FD62FBCD452BFB12" author="Maldaner ME & Nunes RV & Vaz-de-Mello FZ" box="[647,876,1151,1176]" pageId="1" pageNumber="3000" pagination="549 - 560" refId="ref3351" refString="Maldaner ME, Nunes RV, Vaz-de-Mello FZ. 2015. Taxonomic revision of the Dichotomius speciosus (Waterhouse, 1891) species group (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa. 3986 (5): 549 - 560. doi: 10.11646 / zootaxa. 3986.5.2." type="journal article" year="2015">Maldaner et al. 2015</bibRefCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EC9185DECF05FF96FF5AFB7045A2FA02" pageId="1" pageNumber="3000" type="discussion">
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<paragraph id="A434D655CF05FF96FF5AFB7045A2FA02" blockId="1.[191,1155,1218,1728]" pageId="1" pageNumber="3000">
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The following key allows the identification of the five species currently included in the
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<taxonomicName id="638BADD6CF05FF96FF3AFB574703FB74" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[223,324,1253,1278]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="3000" phylum="Arthropoda" rank="species" species="speciosus">
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<emphasis id="96FF0A47CF05FF96FF3AFB574703FB74" box="[223,324,1253,1278]" italics="true" pageId="1" pageNumber="3000">speciosus</emphasis>
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</taxonomicName>
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species group. Identification of females can be confusing if no males or localities are associated with them (e.g. green females of
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<taxonomicName id="638BADD6CF05FF96FCB2FAB5459EFAAA" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[855,985,1287,1312]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="3000" phylum="Arthropoda" rank="species" species="speciosus">
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<emphasis id="96FF0A47CF05FF96FCB2FAB5459EFAAA" box="[855,985,1287,1312]" italics="true" pageId="1" pageNumber="3000">D. speciosus</emphasis>
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</taxonomicName>
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and females of
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<taxonomicName id="638BADD6CF05FF96FF3AFA984736FAC9" authorityName="Maldaner, Nunes and Vaz-de-Mello" authorityYear="2015" box="[223,369,1322,1347]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="3000" phylum="Arthropoda" rank="species" species="alvarengai">
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<emphasis id="96FF0A47CF05FF96FF3AFA984736FAC9" box="[223,369,1322,1347]" italics="true" pageId="1" pageNumber="3000">D. alvarengai</emphasis>
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</taxonomicName>
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;
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<emphasis id="96FF0A47CF05FF96FE9BFA984798FAC9" box="[382,479,1322,1347]" italics="true" pageId="1" pageNumber="3000">Figs. 4, 6</emphasis>
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). When not specified, the characteristics fit both males and females. This key is modified from the one presented by
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<bibRefCitation id="C01AABA4CF05FF96FC81FAFF420CFAEC" author="Maldaner ME & Nunes RV & Vaz-de-Mello FZ" box="[868,1099,1357,1382]" pageId="1" pageNumber="3000" pagination="549 - 560" refId="ref3351" refString="Maldaner ME, Nunes RV, Vaz-de-Mello FZ. 2015. Taxonomic revision of the Dichotomius speciosus (Waterhouse, 1891) species group (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa. 3986 (5): 549 - 560. doi: 10.11646 / zootaxa. 3986.5.2." type="journal article" year="2015">Maldaner et al. 2015</bibRefCitation>
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, and most characters referring to steps 3 to 5 are illustrated in that paper.
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</paragraph>
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</subSubSection>
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<subSubSection id="EC9185DECF05FF95FF5AFA20424DF9D2" lastPageId="2" lastPageNumber="3001" pageId="1" pageNumber="3000" type="key">
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<paragraph id="A434D655CF05FF96FF5AFA2042C4F9F1" blockId="1.[191,1155,1218,1728]" pageId="1" pageNumber="3000">
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0. Clypeus with two well-defined central teeth, each tooth margined and with rounded to obtuse tips; clypeal border straight or uniformly rounded laterally, with no angles from clypeal teeth to lateral parts of genae (these latter characters identify species of the subgenus
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<taxonomicName id="638BADD6CF05FF96FE9AFA484444F999" ID-CoL="8CYB" box="[383,515,1530,1555]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="3000" phylum="Arthropoda" rank="subGenus" subGenus="Selenocopris">
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<emphasis id="96FF0A47CF05FF96FE9AFA484444F999" box="[383,515,1530,1555]" italics="true" pageId="1" pageNumber="3000">Selenocopris</emphasis>
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</taxonomicName>
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). Metaventrite disc posteriorly widely concave, discrimen visible in the bottom of concavity. Size small, less than
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<quantity id="63737BB0CF05FF96FCD7F9AF45C6F9BC" box="[818,897,1565,1590]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="3000" unit="mm" value="15.0">15 mm</quantity>
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. At least some metallic/ sericeous sheen on elytra, pronotum and metaventrite. High-elevation areas in southeastern and southern
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<collectingCountry id="DC9C96C5CF05FF96FE36F9D04449F9F1" box="[467,526,1634,1659]" name="Brazil" pageId="1" pageNumber="3000">Brazil</collectingCountry>
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...
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<emphasis id="96FF0A47CF05FF96FDDDF9D045AFF9F1" bold="true" box="[568,1000,1634,1659]" pageId="1" pageNumber="3000">
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<taxonomicName id="638BADD6CF05FF96FDDDF9D0457CF9F1" ID-CoL="35ND3" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[568,827,1634,1659]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="3000" phylum="Arthropoda" rank="species" species="speciosus">
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<emphasis id="96FF0A47CF05FF96FDDDF9D0457CF9F1" bold="true" box="[568,827,1634,1659]" italics="true" pageId="1" pageNumber="3000">Dichotomius speciosus</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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........................ 1
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</paragraph>
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<paragraph id="A434D655CF05FF96FF5AF9374435F94A" blockId="1.[191,1155,1218,1728]" pageId="1" pageNumber="3000">
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0’. Different combination of characters...
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<emphasis id="96FF0A47CF05FF96FD45F9374275F914" bold="true" box="[672,1074,1669,1694]" pageId="1" pageNumber="3000">
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other
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<taxonomicName id="638BADD6CF05FF96FD0FF9374591F914" box="[746,982,1669,1694]" pageId="1" pageNumber="3000">
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<emphasis id="96FF0A47CF05FF96FD0FF9374530F914" bold="true" box="[746,887,1669,1694]" italics="true" pageId="1" pageNumber="3000">Dichotomius</emphasis>
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species
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</taxonomicName>
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groups
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</emphasis>
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(check
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<bibRefCitation id="C01AABA4CF05FF96FF3BF915446EF94A" author="Nunes RV & Vaz-de-Mello FZ" box="[222,553,1703,1728]" pageId="1" pageNumber="3000" pagination="411 - 420" refId="ref3424" refString="Nunes RV, Vaz-de-Mello FZ. 2013. New brachypterous species of Dichotomius Hope, with taxonomic notes in the subgenus Luederwaldtinia Martinez (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa. 3609 (4): 411 - 420. doi: 10.11646 / zootaxa. 3609.4.3." type="journal article" year="2013">Nunes and Vaz-de-Mello (2013</bibRefCitation>
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,
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<bibRefCitation id="C01AABA4CF05FF96FDD1F915442EF94A" author="Nunes RV & Vaz-de-Mello FZ" box="[564,617,1703,1728]" pageId="1" pageNumber="3000" pagination="2231 - 2351" refId="ref3474" refString="Nunes RV, Vaz-de-Mello FZ. 2019. Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini). Journal of Natural History. 53 (37 - 38): 2231 - 2351. doi: 10.1080 / 00222933.2019.1692088." type="journal article" year="2019">2019</bibRefCitation>
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)
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</paragraph>
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<key id="D01A8164CF06FF95FF5AFF3D424DF9D2" pageId="2" pageNumber="3001">
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<keyStep id="1F7FCEF0CF06FF95FF5AFF3D42C4FE6A" pageId="2" pageNumber="3001">
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<paragraph id="A434D655CF06FF95FF5AFF3D42C4FEB9" blockId="2.[191,1163,143,1624]" pageId="2" pageNumber="3001">
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<keyLead id="1F7A7560CF06FF95FF5AFF3D42C4FEB9" pageId="2" pageNumber="3001">
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1. Clypeofrontal region with rounded knob, or tubercle, or transversally carinate (female only: carina with two close tubercles and narrower than one-fifth of interocular distance). Dorsum and venter dark, with strong to faint blue metallic reflections. Known only from
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<collectingRegion id="664F18B7CF06FF95FE0DFF4544F6FE9A" box="[488,689,247,272]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="3001">Minas Gerais state</collectingRegion>
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(one doubtful report from
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<collectingRegion id="664F18B7CF06FF95FC3BFF45420BFE9A" box="[990,1100,247,272]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="3001">São Paulo</collectingRegion>
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with no locality data)................................................................................................................................... 2
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</keyLead>
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</paragraph>
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<paragraph id="A434D655CF06FF95FF5AFE8E42C4FE6A" blockId="2.[191,1163,143,1624]" pageId="2" pageNumber="3001">
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<keyLead id="1F7A7560CF06FF95FF5AFE8E42C4FE6A" pageId="2" pageNumber="3001">
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1’. Clypeofrontal region with transverse carina, carina wider than one-third of interocular distance, trilobed, trituberculated (tubercles poorly indicated in females) or with horns or lobe at both extremities (males). Dorsum and venter coloured, with very strong green, red or blue metallic sheen.
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<collectingRegion id="664F18B7CF06FF95FD00FE1645F7FE37" box="[741,944,420,445]" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="3001">Rio Grande do Sul</collectingRegion>
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to southern
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<collectingRegion id="664F18B7CF06FF95FBA4FE164763FE6A" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="3001">Minas Gerais</collectingRegion>
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and
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<collectingRegion id="664F18B7CF06FF95FEB8FE7547BBFE6A" box="[349,508,455,480]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="3001">Rio de Janeiro</collectingRegion>
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............................................................................................................... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="1F7FCEF0CF06FF95FF5AFE5842C4FC92" pageId="2" pageNumber="3001">
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<paragraph id="A434D655CF06FF95FF5AFE5842C3FD07" blockId="2.[191,1163,143,1624]" pageId="2" pageNumber="3001">
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<keyLead id="1F7A7560CF06FF95FF5AFE5842C3FD07" pageId="2" pageNumber="3001">
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2. Males with pronotal disc anteriorly bearing an anteromedial protuberance with a depression at each side; clypeus with a few feeble punctures in the posterior half; clypeofrontal tubercle laterally compressed. Females with clypeofrontal transverse bilobate elevation. Southern Espinhaço range in Central
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<collectingRegion id="664F18B7CF06FF95FCBEFDE045ADFDE1" box="[859,1002,594,619]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="3001">Minas Gerais</collectingRegion>
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........................... ................................................
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<emphasis id="96FF0A47CF06FF95FE12FDC64257FD07" bold="true" box="[503,1040,628,653]" pageId="2" pageNumber="3001">
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<taxonomicName id="638BADD6CF06FF95FE12FDC645ECFD07" ID-CoL="8VQZH" authority="Maldaner & Vaz-de-Mello, 2022" authorityName="Maldaner & Vaz-de-Mello" authorityYear="2022" box="[503,939,628,653]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3001" phylum="Arthropoda" rank="species" species="geraldimi" status="sp. nov." subGenus="Selenocopris">
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<emphasis id="96FF0A47CF06FF95FE12FDC645ECFD07" bold="true" box="[503,939,628,653]" italics="true" pageId="2" pageNumber="3001">Dichotomius (Selenocopris) geraldimi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8DCCB73CCF06FF95FC56FDC64257FD07" box="[947,1040,628,653]" pageId="2" pageNumber="3001" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
|
||
<figureCitation id="3CB0CAD0CF06FF95FBC5FDC6423BFD07" box="[1056,1148,628,653]" captionStart="Figure 2" captionStartId="4.[160,225,1219,1242]" captionTargetBox="[168,1154,143,1185]" captionTargetId="figure-112@4.[167,1157,142,1187]" captionTargetPageId="4" captionText="Figure 2. Dichotomius (Selenocopris) geraldimi: (a) male dorsal habitus; (b) female dorsal habitus; (c) aedeagus, dorsal view; (d) male lateral view." figureDoi="http://doi.org/10.5281/zenodo.6351838" httpUri="https://zenodo.org/record/6351838/files/figure.png" pageId="2" pageNumber="3001">Figure 2</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A434D655CF06FF95FF5AFD2542C4FC92" blockId="2.[191,1163,143,1624]" pageId="2" pageNumber="3001">
|
||
<keyLead id="1F7A7560CF06FF95FF5AFD2542C4FC92" pageId="2" pageNumber="3001">
|
||
2’. Males without anteromedial pronotal projection; clypeus densely punctured posteriorly; frontoclypeal tubercle conical. Females with a single conical frontoclypeal elevation, sometimes reduced to a rounded tumosity. Campos Altos,
|
||
<collectingRegion id="664F18B7CF06FF95FCB8FD6E45AFFD7F" box="[861,1000,732,757]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="3001">Minas Gerais</collectingRegion>
|
||
,
|
||
<collectingCountry id="DC9C96C5CF06FF95FC17FD6E4262FD7F" box="[1010,1061,732,757]" name="Brazil" pageId="2" pageNumber="3001">Brazil</collectingCountry>
|
||
................. .....
|
||
<taxonomicName id="638BADD6CF06FF95FEE6FD4D4272FC92" ID-CoL="6CS37" authority="Maldaner, Nunes and Vaz-de-Mello 2015" authorityName="Maldaner, Nunes, and Vaz-de-Mello" authorityYear="2015" box="[259,1077,767,792]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3001" phylum="Arthropoda" rank="species" species="malyi" subGenus="Selenocopris">
|
||
<emphasis id="96FF0A47CF06FF95FEE6FD4D443FFC92" bold="true" box="[259,632,767,792]" italics="true" pageId="2" pageNumber="3001">Dichotomius (Selenocopris) malyi</emphasis>
|
||
<bibRefCitation id="C01AABA4CF06FF95FD9AFD4D4272FC92" author="Maldaner ME & Nunes RV & Vaz-de-Mello FZ" box="[639,1077,767,792]" pageId="2" pageNumber="3001" pagination="549 - 560" refId="ref3351" refString="Maldaner ME, Nunes RV, Vaz-de-Mello FZ. 2015. Taxonomic revision of the Dichotomius speciosus (Waterhouse, 1891) species group (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa. 3986 (5): 549 - 560. doi: 10.11646 / zootaxa. 3986.5.2." type="journal article" year="2015">Maldaner, Nunes and Vaz-de-Mello 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="96FF0A47CF06FF95FBD9FD4D42C4FC92" box="[1084,1155,767,792]" italics="true" pageId="2" pageNumber="3001">(Fig. 5)</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="1F7FCEF0CF06FF95FF5AFC9042C4FB51" pageId="2" pageNumber="3001">
|
||
<paragraph id="A434D655CF06FF95FF5AFC9042C4FB81" blockId="2.[191,1163,143,1624]" pageId="2" pageNumber="3001">
|
||
<keyLead id="1F7A7560CF06FF95FF5AFC9042C4FB81" pageId="2" pageNumber="3001">
|
||
3. Pronotum of males with anteromedial projection triangular. In laterofrontal view, male pronotum strongly excavated at anterior declivity. Clypeofrontal carina straight to broadly curved
|
||
<emphasis id="96FF0A47CF06FF95FDEBFCD544D7FC0A" box="[526,656,871,896]" italics="true" pageId="2" pageNumber="3001">
|
||
(
|
||
<figureCitation id="3CB0CAD0CF06FF95FDF0FCD5440CFC0A" box="[533,587,871,896]" captionStart="Figure 1" captionStartId="3.[160,225,937,960]" captionTargetBox="[162,1153,142,904]" captionTargetId="figure-164@3.[162,1154,142,905]" captionTargetPageId="3" captionText="Figure 1. Distribution map of Dichotomius speciosus species group." figureDoi="http://doi.org/10.5281/zenodo.6351836" httpUri="https://zenodo.org/record/6351836/files/figure.png" pageId="2" pageNumber="3001">Fig. 1</figureCitation>
|
||
(d–g))
|
||
</emphasis>
|
||
, with one horn at each end
|
||
<emphasis id="96FF0A47CF06FF95FC30FCD5427AFC0A" box="[981,1085,871,896]" italics="true" pageId="2" pageNumber="3001">
|
||
(
|
||
<figureCitation id="3CB0CAD0CF06FF95FC38FCD54253FC0A" box="[989,1044,871,896]" captionStart="Figure 1" captionStartId="3.[160,225,937,960]" captionTargetBox="[162,1153,142,904]" captionTargetId="figure-164@3.[162,1154,142,905]" captionTargetPageId="3" captionText="Figure 1. Distribution map of Dichotomius speciosus species group." figureDoi="http://doi.org/10.5281/zenodo.6351836" httpUri="https://zenodo.org/record/6351836/files/figure.png" pageId="2" pageNumber="3001">Fig. 1</figureCitation>
|
||
(h))
|
||
</emphasis>
|
||
. Body surface red or with red reflections (may be very weak on green specimens)
|
||
<emphasis id="96FF0A47CF06FF95FBFCFC3846B4FC4F" italics="true" pageId="2" pageNumber="3001">
|
||
(
|
||
<figureCitation id="3CB0CAD0CF06FF95FBC5FC384238FC29" box="[1056,1151,906,931]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="3.[160,225,937,960]" captionStartId-1="4.[160,225,1219,1242]" captionTargetBox-0="[162,1153,142,904]" captionTargetBox-1="[168,1154,143,1185]" captionTargetId-0="figure-164@3.[162,1154,142,905]" captionTargetId-1="figure-112@4.[167,1157,142,1187]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figure 1. Distribution map of Dichotomius speciosus species group." captionText-1="Figure 2. Dichotomius (Selenocopris) geraldimi: (a) male dorsal habitus; (b) female dorsal habitus; (c) aedeagus, dorsal view; (d) male lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.6351836" figureDoi-1="http://doi.org/10.5281/zenodo.6351838" httpUri-0="https://zenodo.org/record/6351836/files/figure.png" httpUri-1="https://zenodo.org/record/6351838/files/figure.png" pageId="2" pageNumber="3001">Figs. 1–2</figureCitation>
|
||
, 6)
|
||
</emphasis>
|
||
. Serra da Mantiqueira mountain range:
|
||
<collectingRegion id="664F18B7CF06FF95FD2FFC1E457AFC4F" box="[714,829,940,965]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="3001">São Paulo</collectingRegion>
|
||
(Campos do Jordão),
|
||
<collectingRegion id="664F18B7CF06FF95FBA4FC1E4763FC62" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="3001">Minas Gerais</collectingRegion>
|
||
(Aiuruoca), and
|
||
<collectingRegion id="664F18B7CF06FF95FE38FC7D443BFC62" box="[477,636,975,1000]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="3001">Rio de Janeiro</collectingRegion>
|
||
(Itatiaia),
|
||
<collectingCountry id="DC9C96C5CF06FF95FD0EFC7D4560FC62" box="[747,807,975,1000]" name="Brazil" pageId="2" pageNumber="3001">Brazil</collectingCountry>
|
||
............................................................... ............................................
|
||
<taxonomicName id="638BADD6CF06FF95FE3BFC4042C4FB81" ID-CoL="35ND3" authority="(Waterhouse, 1891)" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[478,1155,1010,1035]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3001" phylum="Arthropoda" rank="species" species="speciosus" subGenus="Selenocopris">
|
||
<emphasis id="96FF0A47CF06FF95FE3BFC4042C4FB81" bold="true" box="[478,1155,1010,1035]" pageId="2" pageNumber="3001">
|
||
<emphasis id="96FF0A47CF06FF95FE3BFC4045D6FB81" bold="true" box="[478,913,1010,1035]" italics="true" pageId="2" pageNumber="3001">Dichotomius (Selenocopris) speciosus</emphasis>
|
||
(
|
||
<bibRefCitation id="C01AABA4CF06FF95FC45FC40423CFB81" author="Waterhouse" box="[928,1147,1010,1035]" pageId="2" pageNumber="3001" pagination="348 - 363" refId="ref3689" refString="Waterhouse, C. O. 1891. New Scarabaeidae in the British Museum: a Fourth Contribution. The Annals and Magazine of Natural History, including Zoology, Botany and Geology 6, 348 - 363." type="journal article" year="1891">Waterhouse, 1891</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A434D655CF06FF95FF5AFBA642C4FB51" blockId="2.[191,1163,143,1624]" pageId="2" pageNumber="3001">
|
||
<keyLead id="1F7A7560CF06FF95FF5AFBA642C4FB51" pageId="2" pageNumber="3001">
|
||
3’. Pronotum of males with anteromedial projection rectangular or rounded (
|
||
<emphasis id="96FF0A47CF06FF95FBCCFBA646BCFBDA" italics="true" pageId="2" pageNumber="3001">Figs. 3A, 4A</emphasis>
|
||
), apically slightly emarginate and bearing a pair of tubercles. Pronotum excavated below anteromedial projection. Clypeofrontal carina straight, with one tubercle on extremities. Body surface blue or dark with blue reflections. Serra da
|
||
<taxonomicName id="638BADD6CF06FF95FBCEFBCE47DFFB32" ID-CoL="B4B53" authority="Mountain Range" authorityName="Mountain Range" class="Arachnida" family="Gonyleptidae" genus="Bocaina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3001" phylum="Arthropoda" rank="genus">Bocaina Mountain Range</taxonomicName>
|
||
(state of
|
||
<collectingRegion id="664F18B7CF06FF95FDE6FB2D4433FB32" box="[515,628,1183,1208]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="3001">São Paulo</collectingRegion>
|
||
,
|
||
<collectingCountry id="DC9C96C5CF06FF95FD9AFB2D44FAFB32" box="[639,701,1183,1208]" name="Brazil" pageId="2" pageNumber="3001">Brazil</collectingCountry>
|
||
) and Serra Geral mountain range (states of
|
||
<collectingRegion id="664F18B7CF06FF95FF18FB7047D8FB51" box="[253,415,1218,1243]" country="Brazil" name="Santa Catarina" pageId="2" pageNumber="3001">Santa Catarina</collectingRegion>
|
||
and
|
||
<collectingRegion id="664F18B7CF06FF95FE3CFB7044E0FB51" box="[473,679,1218,1243]" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="3001">Rio Grande do Sul</collectingRegion>
|
||
,
|
||
<collectingCountry id="DC9C96C5CF06FF95FD54FB7044A8FB51" box="[689,751,1218,1243]" name="Brazil" pageId="2" pageNumber="3001">Brazil</collectingCountry>
|
||
)................................................................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="1F7FCEF0CF06FF95FF5AFB56424DF9D2" pageId="2" pageNumber="3001">
|
||
<paragraph id="A434D655CF06FF95FF5AFB5642C4FA02" blockId="2.[191,1163,143,1624]" pageId="2" pageNumber="3001">
|
||
<keyLead id="1F7A7560CF06FF95FF5AFB5642C4FA02" pageId="2" pageNumber="3001">
|
||
4. Pronotum of male medially and longitudinally sulcate, anteromedial projection emarginate apically. Clypeofrontal carina with one acute tubercle at each end. Elytral striae feebly marked, and defined by fine punctures. Dark blue – some individuals with light brown elytra. Serra Geral mountain range (states of
|
||
<collectingRegion id="664F18B7CF06FF95FCF7FAFE45F7FAEF" box="[786,944,1356,1381]" country="Brazil" name="Santa Catarina" pageId="2" pageNumber="3001">Santa Catarina</collectingRegion>
|
||
and
|
||
<collectingRegion id="664F18B7CF06FF95FC03FAFE4745FA02" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="3001">Rio Grande do Sul</collectingRegion>
|
||
,
|
||
<collectingCountry id="DC9C96C5CF06FF95FEE9FADD470FFA02" box="[268,328,1391,1416]" name="Brazil" pageId="2" pageNumber="3001">Brazil</collectingCountry>
|
||
) ..................................
|
||
<taxonomicName id="638BADD6CF06FF95FDF7FADD42C4FA02" ID-CoL="35NCN" authority="(Felsche, 1910)" baseAuthorityName="Felsche" baseAuthorityYear="1910" box="[530,1155,1391,1416]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3001" phylum="Arthropoda" rank="species" species="opalescens" subGenus="Selenocopris">
|
||
<emphasis id="96FF0A47CF06FF95FDF7FADD42C4FA02" bold="true" box="[530,1155,1391,1416]" pageId="2" pageNumber="3001">
|
||
<emphasis id="96FF0A47CF06FF95FDF7FADD458AFA02" bold="true" box="[530,973,1391,1416]" italics="true" pageId="2" pageNumber="3001">Dichotomius (Selenocopris) opalescens</emphasis>
|
||
(
|
||
<bibRefCitation id="C01AABA4CF06FF95FC3FFADD423DFA02" author="Felsche" box="[986,1146,1391,1416]" pageId="2" pageNumber="3001" pagination="339 - 352" refId="ref3227" refString="Felsche, C. 1910. Ueber coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift 4, 339 - 352." type="journal article" year="1910">Felsche, 1910</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A434D655CF06FF95FF5AFA20424DF9D2" blockId="2.[191,1163,143,1624]" pageId="2" pageNumber="3001">
|
||
<keyLead id="1F7A7560CF06FF95FF5AFA20424DF9D2" pageId="2" pageNumber="3001">
|
||
4’. Pronotum of male with weak, medial and longitudinal sulcus, anteromedial projection weakly emarginated. Male clypeofrontal carina bearing a tubercle on each end, while female cephalic carina lacks them. Elytral striae deeply impressed, and defined by distinct punctures. Green to blue. Restricted to the Serra da
|
||
<taxonomicName id="638BADD6CF06FF95FEABF9AE443AF9BF" ID-CoL="B4B53" authority="Mountain Range" authorityName="Mountain Range" box="[334,637,1564,1589]" class="Arachnida" family="Gonyleptidae" genus="Bocaina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3001" phylum="Arthropoda" rank="genus">Bocaina Mountain Range</taxonomicName>
|
||
, state of
|
||
<collectingRegion id="664F18B7CF06FF95FD17F9AE452CF9BF" box="[754,875,1564,1589]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="3001">São Paulo</collectingRegion>
|
||
,
|
||
<collectingCountry id="DC9C96C5CF06FF95FC9FF9AE45FEF9BF" box="[890,953,1564,1589]" name="Brazil" pageId="2" pageNumber="3001">Brazil</collectingCountry>
|
||
....
|
||
<taxonomicName id="638BADD6CF06FF95FC09F9AE424DF9D2" ID-CoL="6D3R6" authority="Maldaner, Nunes and Vaz-de-Mello 2015" authorityName="Maldaner, Nunes and Vaz-de-Mello" authorityYear="2015" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3001" phylum="Arthropoda" rank="species" species="alvarengai" subGenus="Selenocopris">
|
||
<emphasis id="96FF0A47CF06FF95FC09F9AE4450F9D2" bold="true" italics="true" pageId="2" pageNumber="3001">Dichotomius (Selenocopris) alvarengai</emphasis>
|
||
<bibRefCitation id="C01AABA4CF06FF95FDC6F98D424DF9D2" author="Maldaner ME & Nunes RV & Vaz-de-Mello FZ" box="[547,1034,1599,1624]" pageId="2" pageNumber="3001" pagination="549 - 560" refId="ref3351" refString="Maldaner ME, Nunes RV, Vaz-de-Mello FZ. 2015. Taxonomic revision of the Dichotomius speciosus (Waterhouse, 1891) species group (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa. 3986 (5): 549 - 560. doi: 10.11646 / zootaxa. 3986.5.2." type="journal article" year="2015">Maldaner, Nunes and Vaz-de-Mello 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |