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<document id="11DC2E6EB760F8D679C859E01E3A347F" ID-DOI="10.5281/zenodo.5331202" ID-ISSN="0374-1036" ID-Zenodo-Dep="5331202" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630169194360" checkinUser="diego" docAuthor="Švec, Zdeněk" docDate="2012" docId="297B87BCA27AFFC1FEE66611FECDAD9B" docLanguage="en" docName="ActaEntMusNatPra.2012.52.2.411-424.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Agathidium (Macroceble) guttiferum Švec 2012, sp. nov." docType="treatment" docVersion="4" lastPageNumber="419" masterDocId="D542FFC4A27DFFC9FFB9651CFFE7A90F" masterDocTitle="New Leiodinae (Coleoptera: Leiodidae) from India and Papua New Guinea" masterLastPageNumber="424" masterPageNumber="411" pageNumber="418" updateTime="1699172137447" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B423B754F0B9F2CAAD66E0E8CE8B8D32" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="65CD2E3BAF67575F5635B7EFF0BD5E29">
<mods:title id="F6398E0F00413A2E7CD6F0F5FE927AE3">New Leiodinae (Coleoptera: Leiodidae) from India and Papua New Guinea</mods:title>
</mods:titleInfo>
<mods:name id="D96DF9E6FD88FB049171C05270C76B36" type="personal">
<mods:role id="B500CF8DF34AEA391B04D6764D959D01">
<mods:roleTerm id="01CC4B76452BCB0C945E3BD2F9FC1674">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8C73598FB11169A1A6584F60D7F138B5">Švec, Zdeněk</mods:namePart>
</mods:name>
<mods:typeOfResource id="23E96AA8B18FB0C4882F48BE96DE7D23">text</mods:typeOfResource>
<mods:relatedItem id="B27CAA539210699DD00C26CC9F2B7986" type="host">
<mods:titleInfo id="5317BD7E3E0715E26720443BAC9FE1CE">
<mods:title id="B1BE4637F3B92342A5E6E3897EE0B04B">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="2B70C540CA690D16D66FE45182691782">
<mods:date id="AEF9166F6AE0DF847BC17BED77A462F3">2012</mods:date>
<mods:detail id="5560B10E70B8F5B8C6BA3BC9691A767B" type="pubDate">
<mods:number id="05F909A9A9D9BAD191F4E7F3B4D72E3C">2012-12-17</mods:number>
</mods:detail>
<mods:detail id="3F44EEF3D0FFAB9D17A8A0C088757B1B" type="volume">
<mods:number id="4B237FC96F53D55C5582AA92F271BA48">52</mods:number>
</mods:detail>
<mods:detail id="BA1A6474485C940DBBE5AF0D96BF7060" type="issue">
<mods:number id="FE8D499D12D861085722AB6027BA431D">2</mods:number>
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<mods:start id="7DAD459984113DF25621F97A9C0375FB">411</mods:start>
<mods:end id="1D71520042F662C9AA706C1C40E3F7E6">424</mods:end>
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<mods:classification id="00A2E297D1B116A6FC0E50A5CA43A4A8">journal article</mods:classification>
<mods:identifier id="C670050AE231639E05C87D247D001A35" type="DOI">10.5281/zenodo.5331202</mods:identifier>
<mods:identifier id="B00711988D1B2CF3264279C1B8B88F91" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="404A7B797D0181FCF0D4D323910881D6" type="Zenodo-Dep">5331202</mods:identifier>
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<treatment id="297B87BCA27AFFC1FEE66611FECDAD9B" ID-DOI="http://doi.org/10.5281/zenodo.5398905" ID-Zenodo-Dep="5398905" LSID="urn:lsid:plazi:treatment:297B87BCA27AFFC1FEE66611FECDAD9B" httpUri="http://treatment.plazi.org/id/297B87BCA27AFFC1FEE66611FECDAD9B" lastPageId="8" lastPageNumber="419" pageId="7" pageNumber="418">
<subSubSection id="E9C86521A27AFFCEFEE66611FCB4AA28" box="[351,851,781,807]" pageId="7" pageNumber="418" type="nomenclature">
<paragraph id="A16D36AAA27AFFCEFEE66611FCB4AA28" blockId="7.[351,851,781,807]" box="[351,851,781,807]" pageId="7" pageNumber="418">
<heading id="FA2581C6A27AFFCEFEE66611FCB4AA28" bold="true" box="[351,851,781,807]" centered="true" fontSize="8" level="2" pageId="7" pageNumber="418" reason="3">
<emphasis id="93A6EAB8A27AFFCEFEE66611FCB4AA28" bold="true" box="[351,851,781,807]" pageId="7" pageNumber="418">
<taxonomicName id="66D24D29A27AFFCEFEE66611FD10AA29" ID-CoL="5TPG2" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[351,759,781,806]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="guttiferum" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A27AFFCEFEE66611FE06AA29" bold="true" box="[351,481,781,806]" italics="true" pageId="7" pageNumber="418">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A27AFFCEFE486611FD95AA29" bold="true" box="[497,626,781,806]" italics="true" pageId="7" pageNumber="418">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A27AFFCEFD3B6611FD10AA29" bold="true" box="[642,759,781,806]" italics="true" pageId="7" pageNumber="418">guttiferum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A27AFFCEFD446613FCB4AA28" box="[765,851,783,807]" pageId="7" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27AFFCEFDB96631FD55AA4F" box="[512,690,813,832]" pageId="7" pageNumber="418" type="description">
<paragraph id="A16D36AAA27AFFCEFDB96631FD55AA4F" blockId="7.[512,690,813,832]" box="[512,690,813,832]" pageId="7" pageNumber="418">
(
<figureCitation id="39E92A2FA27AFFCEFDBF6631FD8BAA4F" box="[518,620,813,832]" captionStart="Figs" captionStartId="9.[113,152,885,904]" captionTargetBox="[138,1064,224,854]" captionTargetId="figure-227@9.[131,1071,219,865]" captionTargetPageId="9" captionText="Figs. 11–17. Genitalia of Agathidium and Zeadolopus. 11–13 – Agathidium (Macroceble) guttigerum sp nov. (11 – aedeagus, lateral view; 12 – apex of aedeagus, dorsal view; 13 – spermatheca). 14–16 – A. (Macroceble) acutangulum sp. nov.(14 – aedeagus, lateral view; 15 – apex of aedeagus, dorsal view; 16 – spermatheca). 17 – Zeadolopus punctiventris sp. nov., aedeagus, dorsal view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331212" httpUri="https://zenodo.org/record/5331212/files/figure.png" pageId="7" pageNumber="418">Figs. 11–13</figureCitation>
,
<figureCitation id="39E92A2FA27AFFCEFDCF6631FD4BAA4F" box="[630,684,813,832]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="7" pageNumber="418">26, 27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27AFFCEFFC8664BFE7EAAE0" pageId="7" pageNumber="418" type="materials_examined">
<paragraph id="A16D36AAA27AFFCEFFC8664BFF38AA8B" blockId="7.[113,1089,854,1007]" pageId="7" pageNumber="418">
<emphasis id="93A6EAB8A27AFFCEFFC8664BFF00AA65" bold="true" box="[113,231,854,874]" pageId="7" pageNumber="418">
<typeStatus id="7E698808A27AFFCEFFC8664BFF7AAA65" box="[113,157,855,874]" pageId="7" pageNumber="418">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D9C5763AA27AFFCEFF53664AFEF0AA66" box="[234,279,854,873]" name="India" pageId="7" pageNumber="418">India</collectingCountry>
,
<collectingRegion id="6316F848A27AFFCEFEA4664AFE5AAA66" box="[285,445,854,873]" country="India" name="Arunachal Pradesh" pageId="7" pageNumber="418">Arunachal Pradesh</collectingRegion>
,
<quantity id="662A9B4FA27AFFCEFE7A664BFE11AA66" box="[451,502,854,874]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="418" unit="km" value="11.0">11 km</quantity>
SSE of Tenga, Eagles Nest wildlife sanctuary,
<geoCoordinate id="C4E6506DA27AFFCEFCCF664AFC3FAA65" box="[886,984,854,874]" degrees="27" direction="north" minutes="06" orientation="latitude" pageId="7" pageNumber="418" precision="15" seconds="23" value="27.10639">27º06′23″N</geoCoordinate>
<geoCoordinate id="C4E6506DA27AFFCEFC65664AFBDAAA65" box="[988,1085,854,874]" degrees="92" direction="east" minutes="26" orientation="longitude" pageId="7" pageNumber="418" precision="15" seconds="02" value="92.43389">92º26′02″E</geoCoordinate>
,
<quantity id="662A9B4FA27AFFCEFFC8666DFF55AA8A" box="[113,178,881,901]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="7" pageNumber="418" unit="m" value="2510.0">2510 m</quantity>
a.s.l.
</paragraph>
<paragraph id="A16D36AAA27AFFCEFFC86690FE7EAAE0" blockId="7.[113,1089,854,1007]" pageId="7" pageNumber="418">
<emphasis id="93A6EAB8A27AFFCEFFC86690FF11AA90" bold="true" box="[113,246,907,927]" pageId="7" pageNumber="418">Type material.</emphasis>
<materialsCitation id="11BA3CF7A27AFFCEFF426690FCD4AADB" collectingDate="2008-05-04" collectionCode="NMPC" collectorName="Nest WL Sanctuary" country="India" elevation="2510" latitude="27.10639" location="Eagles" longLatPrecision="20" longitude="92.43389" pageId="7" pageNumber="418" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<typeStatus id="7E698808A27AFFCEFF426690FEA8AA91" box="[251,335,908,927]" pageId="7" pageNumber="418" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="C7C3AE6FA27AFFCEFECC6690FE50AA90" box="[373,439,908,927]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="418">NMPC</collectionCode>
): ‘
<collectingCountry id="D9C5763AA27AFFCEFE706690FDEBAA90" box="[457,524,908,927]" name="India" pageId="7" pageNumber="418">INDIA</collectingCountry>
:
<collectingRegion id="6316F848A27AFFCEFDAA6690FD51AA90" box="[531,694,908,927]" country="India" name="Arunachal Pradesh" pageId="7" pageNumber="418">Arunachal Pradesh</collectingRegion>
(15) /
<locationDeviation id="B0DF2F88A27AFFCEFD566690FC46AA90" box="[751,929,908,927]" location="Tenga" pageId="7" pageNumber="418">
<quantity id="662A9B4FA27AFFCEFD566690FCC3AA90" box="[751,804,908,927]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="418" unit="km" value="11.0">11 km</quantity>
SSE of Tenga
</locationDeviation>
,
<location id="A40D6071A27AFFCEFC136690FC04AA90" LSID="urn:lsid:plazi:treatment:297B87BCA27AFFC1FEE66611FECDAD9B:A40D6071A27AFFCEFC136690FC04AA90" box="[938,995,908,927]" country="India" latitude="27.10639" longLatPrecision="20" longitude="92.43389" name="Eagles" pageId="7" pageNumber="418" stateProvince="Arunachal Pradesh">Eagles</location>
/
<collectorName id="0C27537CA27AFFCEFC4D6690FF20AAB5" pageId="7" pageNumber="418">Nest WL Sanctuary</collectorName>
,
<date id="D56C106AA27AFFCEFF7666BBFEFEAAB5" box="[207,281,935,954]" pageId="7" pageNumber="418" value="2008-05-04">
<collectingDate id="C528E982A27AFFCEFF7666BBFEFEAAB5" box="[207,281,935,954]" pageId="7" pageNumber="418" value="2008-05-04">4.v.2008</collectingDate>
</date>
/
<geoCoordinate id="C4E6506DA27AFFCEFE9166BAFE6CAAB5" box="[296,395,934,954]" degrees="27" direction="north" minutes="06" orientation="latitude" pageId="7" pageNumber="418" precision="15" seconds="23" value="27.10639">27º06′23″N</geoCoordinate>
<geoCoordinate id="C4E6506DA27AFFCEFE2966BAFE14AAB5" box="[400,499,934,954]" degrees="92" direction="east" minutes="26" orientation="longitude" pageId="7" pageNumber="418" precision="15" seconds="02" value="92.43389">92º26′02″E</geoCoordinate>
,
<quantity id="662A9B4FA27AFFCEFE4266BAFDDCAAB5" box="[507,571,934,954]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="7" pageNumber="418" unit="m" value="2510.0">
<elevation id="2AFFD199A27AFFCEFE4266BAFDDCAAB5" box="[507,571,934,954]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="7" pageNumber="418" unit="m" value="2510.0">2510 m</elevation>
</quantity>
/ Fikáček, Podskalská, Šípek lgt. // Primary evergreen mountain / cloud forest with very sparse / understory vegetation / sifting: wet leaf litter’
</materialsCitation>
.
<typeStatus id="7E698808A27AFFCEFC8266DEFC74AADC" box="[827,915,962,981]" pageId="7" pageNumber="418" type="paratype">PARATYPES</typeStatus>
: 1 J
<specimenCount id="B7D4FD23A27AFFCEFC7A66DDFC15AADB" box="[963,1010,955,980]" pageId="7" pageNumber="418" type="female">9 ♀♀</specimenCount>
(
<collectionCode id="C7C3AE6FA27AFFCEFC4466DDFBDAAADB" box="[1021,1085,961,980]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="418">NMPC</collectionCode>
,
<collectionCode id="C7C3AE6FA27AFFCEFFC866C0FF4FAAE0" box="[113,168,988,1007]" pageId="7" pageNumber="418">ZSPC</collectionCode>
): same data as the
<typeStatus id="7E698808A27AFFCEFEFE66C0FE73AAE0" box="[327,404,988,1007]" pageId="7" pageNumber="418" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27AFFC1FFC8611BFC40A8FC" lastPageId="8" lastPageNumber="419" pageId="7" pageNumber="418" type="description">
<paragraph id="A16D36AAA27AFFCEFFC8611BFCCDAD50" blockId="7.[113,1089,1031,1631]" pageId="7" pageNumber="418">
<emphasis id="93A6EAB8A27AFFCEFFC8611BFF1BAD10" bold="true" box="[113,252,1031,1055]" pageId="7" pageNumber="418">Description.</emphasis>
Body length 1.8–2.0 mm (
<typeStatus id="7E698808A27AFFCEFDAF611BFD9EAD10" box="[534,633,1031,1055]" pageId="7" pageNumber="418" type="holotype">holotype</typeStatus>
: 2.0 mm). Length of body parts (
<typeStatus id="7E698808A27AFFCEFC6B611BFBD1AD10" box="[978,1078,1031,1055]" pageId="7" pageNumber="418" type="holotype">holotype</typeStatus>
): head
<quantity id="662A9B4FA27AFFCEFF12613BFF18AD30" box="[171,255,1063,1087]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="418" unit="mm" value="0.3">0.3 mm</quantity>
, pronotum
<quantity id="662A9B4FA27AFFCEFEC0613BFE2AAD30" box="[377,461,1063,1087]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="418" unit="mm" value="0.8">0.8 mm</quantity>
, elytra
<quantity id="662A9B4FA27AFFCEFDA5613BFD97AD30" box="[540,624,1063,1087]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="418" unit="mm" value="0.9">0.9 mm</quantity>
, antenna
<quantity id="662A9B4FA27AFFCEFD6C613BFCCDAD30" box="[725,810,1063,1087]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="418" unit="mm" value="0.6">0.6 mm</quantity>
. Maximum width of body parts (
<typeStatus id="7E698808A27AFFCEFF0A615BFEF0AD50" box="[179,279,1095,1119]" pageId="7" pageNumber="418" type="holotype">holotype</typeStatus>
): head
<quantity id="662A9B4FA27AFFCEFEDB615BFE51AD50" box="[354,438,1095,1119]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="418" unit="mm" value="0.9">0.9 mm</quantity>
, pronotum
<quantity id="662A9B4FA27AFFCEFD96615BFD64AD50" box="[559,643,1095,1119]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="418" unit="mm" value="1.2">1.2 mm</quantity>
, elytra
<quantity id="662A9B4FA27AFFCEFD68615BFCC1AD50" box="[721,806,1095,1119]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="418" unit="mm" value="1.1">1.1 mm</quantity>
.
</paragraph>
<paragraph id="A16D36AAA27AFFCEFF29617BFE97ADD0" blockId="7.[113,1089,1031,1631]" pageId="7" pageNumber="418">
Body shortly oval (
<figureCitation id="39E92A2FA27AFFCEFED9617BFE56AD70" box="[352,433,1127,1151]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="7" pageNumber="418">Fig. 26</figureCitation>
), red-brown with posterior half of head, majority of pronotal surface except margins and large triangular ill-defined patch on base of elytra dark brown. Venter yellow-reddish. Dorsum without microreticulation, punctuate, each puncture with very short and fine seta.
</paragraph>
<paragraph id="A16D36AAA27AFFCEFF2961FBFF10AC70" blockId="7.[113,1089,1031,1631]" pageId="7" pageNumber="418">Head. Maximum width of head at eyes shortly before their posterior margin. Eyes very flat, narrow, 7× as long as wide in dorsal view. Clypeus feebly emarginate, clypeal line absent. Dorsal surface with very sparsely scattered fine punctures separated by more than 10× their diameter. Mandibles without striking characters. Relative length of antennomeres III: II = 1.7.</paragraph>
<paragraph id="A16D36AAA27AFFCEFF296094FDAFACD0" blockId="7.[113,1089,1031,1631]" pageId="7" pageNumber="418">
Pronotum. Widest shortly before base. Lateral sides in basal half of pronotum roundly tapered, then conically narrowed to anterior corners in dorsal view; closely rounded in lateral view (
<figureCitation id="39E92A2FA27AFFCEFF0D60DBFEE3ACD0" box="[180,260,1479,1503]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="7" pageNumber="418">Fig. 27</figureCitation>
). Puncturation as that of head.
</paragraph>
<paragraph id="A16D36AAA27AFFCEFF2960FBFF15AF10" blockId="7.[113,1089,1031,1631]" pageId="7" pageNumber="418">Elytra. Broadest approximately at midlength. Punctures as on head and pronotum. Sutural stria absent.</paragraph>
<paragraph id="A16D36AAA27AFFCEFF296334FDBAAF50" blockId="7.[113,1089,1031,1631]" pageId="7" pageNumber="418">
Legs.Anterior tarsomere I a little enlarged in male. Tarsi slender in female. Femora simple. Tarsal formula 5-5-
<quantity id="662A9B4FA27AFFCEFE86635BFE8EAF50" box="[319,361,1607,1631]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="418" unit="in" value="4.0">4 in</quantity>
male, 4-4-
<quantity id="662A9B4FA27AFFCEFE65635BFDE1AF50" box="[476,518,1607,1631]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="418" unit="in" value="4.0">4 in</quantity>
female.
</paragraph>
<paragraph id="A16D36AAA275FFC1FF0C65C7FBBCA9FC" blockId="8.[151,1127,219,1172]" box="[181,1115,219,243]" pageId="8" pageNumber="419">Mesoventrite. Low longitudinal mesoventral carina present; lateral lines not developed.</paragraph>
<paragraph id="A16D36AAA275FFC1FF0C65E7FE3CA81C" blockId="8.[151,1127,219,1172]" box="[181,475,251,275]" pageId="8" pageNumber="419">Metathoracic wings absent.</paragraph>
<paragraph id="A16D36AAA275FFC1FF0C6407FD83A85C" blockId="8.[151,1127,219,1172]" pageId="8" pageNumber="419">Metaventrite. Femoral lines absent. Both sexes with horn-shaped ventro-caudally oriented process at posterior margin between coxae.</paragraph>
<paragraph id="A16D36AAA275FFC1FF0C6447FF0EA89B" blockId="8.[151,1127,219,1172]" pageId="8" pageNumber="419">
Genitalia. Aedeagus as in
<figureCitation id="39E92A2FA275FFC1FE6A6447FDB2A87B" box="[467,597,347,372]" captionStart="Figs" captionStartId="9.[113,152,885,904]" captionTargetBox="[138,1064,224,854]" captionTargetId="figure-227@9.[131,1071,219,865]" captionTargetPageId="9" captionText="Figs. 11–17. Genitalia of Agathidium and Zeadolopus. 11–13 – Agathidium (Macroceble) guttigerum sp nov. (11 – aedeagus, lateral view; 12 – apex of aedeagus, dorsal view; 13 – spermatheca). 14–16 – A. (Macroceble) acutangulum sp. nov.(14 – aedeagus, lateral view; 15 – apex of aedeagus, dorsal view; 16 – spermatheca). 17 – Zeadolopus punctiventris sp. nov., aedeagus, dorsal view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331212" httpUri="https://zenodo.org/record/5331212/files/figure.png" pageId="8" pageNumber="419">Figs. 11–12</figureCitation>
. Operculum small, triangular. Spermatheca as in
<figureCitation id="39E92A2FA275FFC1FF2E6467FF02A89B" box="[151,229,379,404]" captionStart="Figs" captionStartId="9.[113,152,885,904]" captionTargetBox="[138,1064,224,854]" captionTargetId="figure-227@9.[131,1071,219,865]" captionTargetPageId="9" captionText="Figs. 11–17. Genitalia of Agathidium and Zeadolopus. 11–13 – Agathidium (Macroceble) guttigerum sp nov. (11 – aedeagus, lateral view; 12 – apex of aedeagus, dorsal view; 13 – spermatheca). 14–16 – A. (Macroceble) acutangulum sp. nov.(14 – aedeagus, lateral view; 15 – apex of aedeagus, dorsal view; 16 – spermatheca). 17 – Zeadolopus punctiventris sp. nov., aedeagus, dorsal view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331212" httpUri="https://zenodo.org/record/5331212/files/figure.png" pageId="8" pageNumber="419">Fig. 13</figureCitation>
.
</paragraph>
<paragraph id="A16D36AAA275FFC1FF2E6487FC40A8FC" blockId="8.[151,1127,219,1172]" pageId="8" pageNumber="419">
<emphasis id="93A6EAB8A275FFC1FF2E6487FEF1A8BC" bold="true" box="[151,278,411,435]" pageId="8" pageNumber="419">Variability.</emphasis>
The length ratio of antennomeres III: II varies between
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the type specimens; the ratio of length: width of the eyes in the dorsal view being between 6–8. The majority of the
<typeStatus id="7E698808A275FFC1FE8564C0FE45A8FB" box="[316,418,476,500]" pageId="8" pageNumber="419" type="paratype">paratypes</typeStatus>
are reddish with a dark vertex and elytral patch.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A275FFC1FF2E64E7FC16AD1B" pageId="8" pageNumber="419" type="diagnosis">
<paragraph id="A16D36AAA275FFC1FF2E64E7FC16AD1B" blockId="8.[151,1127,219,1172]" pageId="8" pageNumber="419">
<emphasis id="93A6EAB8A275FFC1FF2E64E7FE75AB1C" bold="true" box="[151,402,507,531]" pageId="8" pageNumber="419">Differential diagnosis.</emphasis>
<taxonomicName id="66D24D29A275FFC1FE2364E1FCC3AB1B" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[410,804,508,532]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="guttiferum" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A275FFC1FE2364E1FDF1AB1B" box="[410,534,509,532]" italics="true" pageId="8" pageNumber="419">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A275FFC1FD9F64E1FD43AB1B" box="[550,676,509,532]" italics="true" pageId="8" pageNumber="419">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A275FFC1FD0C64E1FCC3AB1B" box="[693,804,509,532]" italics="true" pageId="8" pageNumber="419">guttiferum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A275FFC1FC9564E0FC67AB1C" box="[812,896,508,531]" pageId="8" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<emphasis id="93A6EAB8A275FFC1FC4064E1FBF7AB1B" box="[1017,1040,509,532]" italics="true" pageId="8" pageNumber="419">A.</emphasis>
(
<emphasis id="93A6EAB8A275FFC1FB9964E1FBB9AB1B" box="[1056,1118,509,532]" italics="true" pageId="8" pageNumber="419">Macr.</emphasis>
)
<taxonomicName id="66D24D29A275FFC1FF2E6701FDDCAB3B" authority="Angelini &amp; DeMarzo, 1981" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[151,571,539,564]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="93A6EAB8A275FFC1FF2E6701FEEEAB3B" box="[151,265,541,564]" italics="true" pageId="8" pageNumber="419">brancuccii</emphasis>
Angelini &amp; DeMarzo, 1981
</taxonomicName>
and
<emphasis id="93A6EAB8A275FFC1FDC86701FD6FAB3B" box="[625,648,541,564]" italics="true" pageId="8" pageNumber="419">A.</emphasis>
(
<emphasis id="93A6EAB8A275FFC1FD206701FD30AB3B" box="[665,727,541,564]" italics="true" pageId="8" pageNumber="419">Macr.</emphasis>
)
<taxonomicName id="66D24D29A275FFC1FD5E6701FC6DAB3B" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[743,906,541,564]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="cochleariforme" status="sp. nov.">
<emphasis id="93A6EAB8A275FFC1FD5E6701FC6DAB3B" box="[743,906,541,564]" italics="true" pageId="8" pageNumber="419">cochleariforme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A275FFC1FC286700FC02AB3C" box="[913,997,540,563]" pageId="8" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
in the body size, coloration of the antennae, the presence of dorsal puncturation, absence of microsculpture on the dorsum, absence of sutural striae and by the presence of a semivertical process on the metaventrite. It also resembles
<taxonomicName id="66D24D29A275FFC1FE5C6761FD92AB9B" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[485,629,637,660]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="93A6EAB8A275FFC1FE5C6761FD92AB9B" box="[485,629,637,660]" italics="true" pageId="8" pageNumber="419">A. brancuccii</emphasis>
</taxonomicName>
in the shape of the aedeagus. The new species differs from
<taxonomicName id="66D24D29A275FFC1FEA46781FE4BABBB" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[285,428,669,692]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="93A6EAB8A275FFC1FEA46781FE4BABBB" box="[285,428,669,692]" italics="true" pageId="8" pageNumber="419">A. brancuccii</emphasis>
</taxonomicName>
and
<taxonomicName id="66D24D29A275FFC1FE596781FD47ABBB" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[480,672,669,692]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="cochleariforme" status="sp. nov.">
<emphasis id="93A6EAB8A275FFC1FE596781FD47ABBB" box="[480,672,669,692]" italics="true" pageId="8" pageNumber="419">A. cochleariforme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A275FFC1FD1E6780FD1EABBC" box="[679,761,668,691]" pageId="8" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
in the longer antennomere III (the length ratio antennomeres III: II =
<quantity id="662A9B4FA275FFC1FDA867A0FD9BABDC" box="[529,636,699,724]" metricMagnitude="-2" metricUnit="m" metricValue="4.445" metricValueMax="4.572" metricValueMin="4.318" pageId="8" pageNumber="419" unit="in" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 in</quantity>
<taxonomicName id="66D24D29A275FFC1FD3A67A1FCE8ABDB" authorityName="Švec" authorityYear="2012" box="[643,783,701,724]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="guttiferum">
<emphasis id="93A6EAB8A275FFC1FD3A67A1FCE8ABDB" box="[643,783,701,724]" italics="true" pageId="8" pageNumber="419">A. guttiferum</emphasis>
</taxonomicName>
, while it is
<quantity id="662A9B4FA275FFC1FC3467A0FC2DABDC" box="[909,970,699,724]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="8" pageNumber="419" unit="in" value="1.4">1.4 in</quantity>
<emphasis id="93A6EAB8A275FFC1FC6967A1FB81ABDB" box="[976,1126,701,724]" italics="true" pageId="8" pageNumber="419">
<taxonomicName id="66D24D29A275FFC1FC6967A1FB84ABDB" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[976,1123,701,724]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="brancuccii">A. brancuccii</taxonomicName>
,
</emphasis>
and
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<taxonomicName id="66D24D29A275FFC1FEB067C1FE2DABFB" authorityName="Švec" authorityYear="2012" box="[265,458,733,756]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="cochleariforme">
<emphasis id="93A6EAB8A275FFC1FEB067C1FE2DABFB" box="[265,458,733,756]" italics="true" pageId="8" pageNumber="419">A. cochleariforme</emphasis>
</taxonomicName>
), the presence of the triangular darker patch on the elytra, and finally by the slender basal part of the spermatheca (in contrast to the subhaemisphaerical basal portion of spermatheca in both
<emphasis id="93A6EAB8A275FFC1FDA46601FC78AA3B" box="[541,927,797,820]" italics="true" pageId="8" pageNumber="419">
<taxonomicName id="66D24D29A275FFC1FDA46601FD4BAA3B" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[541,684,797,820]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="brancuccii">A. brancuccii</taxonomicName>
and
<taxonomicName id="66D24D29A275FFC1FD666601FC78AA3B" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[735,927,797,820]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="cochleariforme" status="sp. nov.">A. cochleariforme</taxonomicName>
</emphasis>
<taxonomicNameLabel id="889557C3A275FFC1FC1C6600FC1DAA3B" box="[933,1018,796,820]" pageId="8" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
). The new species also differs from
<taxonomicName id="66D24D29A275FFC1FE276621FDCAAA5B" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[414,557,829,852]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="93A6EAB8A275FFC1FE276621FDCAAA5B" box="[414,557,829,852]" italics="true" pageId="8" pageNumber="419">A. brancuccii</emphasis>
</taxonomicName>
in the distinctly sparser dorsal puncturation, the tarsal formula in females 4-4-4 (in contrast to 5-4-
<quantity id="662A9B4FA275FFC1FDE56647FD63AA7C" box="[604,644,859,883]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="419" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="66D24D29A275FFC1FD316641FCF2AA7B" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[648,789,861,884]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="93A6EAB8A275FFC1FD316641FCF2AA7B" box="[648,789,861,884]" italics="true" pageId="8" pageNumber="419">A. brancuccii</emphasis>
</taxonomicName>
), and by the shape of the median lobe that ends in a small pointed process (in contrast to the simply pointed apex of the median lobe in
<taxonomicName id="66D24D29A275FFC1FF506681FE9DAABB" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[233,378,925,948]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="93A6EAB8A275FFC1FF506681FE9DAABB" box="[233,378,925,948]" italics="true" pageId="8" pageNumber="419">A. brancuccii</emphasis>
</taxonomicName>
). The new species also differs from
<taxonomicName id="66D24D29A275FFC1FCBC6681FC20AABB" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[773,967,925,948]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="cochleariforme" status="sp. nov.">
<emphasis id="93A6EAB8A275FFC1FCBC6681FC20AABB" box="[773,967,925,948]" italics="true" pageId="8" pageNumber="419">A. cochleariforme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A275FFC1FC766680FBC5AABC" box="[975,1058,924,947]" pageId="8" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
in the very flat and narrow eyes (in contrast to larger dorsal portion of eyes in
<taxonomicName id="66D24D29A275FFC1FC1C66A1FB81AADB" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[933,1126,957,980]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="cochleariforme" status="sp. nov.">
<emphasis id="93A6EAB8A275FFC1FC1C66A1FB81AADB" box="[933,1126,957,980]" italics="true" pageId="8" pageNumber="419">A. cochleariforme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A275FFC1FF2E66C0FF0AAAFC" box="[151,237,988,1011]" pageId="8" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
, see the description of that species), and by the simple distal part of the parameres (in contrast to the spoon-shaped parameral apices in
<taxonomicName id="66D24D29A275FFC1FD7166E1FC6EAD1B" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[712,905,1021,1044]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="cochleariforme" status="sp. nov.">
<emphasis id="93A6EAB8A275FFC1FD7166E1FC6EAD1B" box="[712,905,1021,1044]" italics="true" pageId="8" pageNumber="419">A. cochleariforme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A275FFC1FC3666E0FC04AD1B" box="[911,995,1020,1044]" pageId="8" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A275FFC1FF2E6100FECDAD9B" pageId="8" pageNumber="419" type="etymology">
<paragraph id="A16D36AAA275FFC1FF2E6100FC58AD3C" blockId="8.[151,1127,219,1172]" box="[151,959,1051,1076]" pageId="8" pageNumber="419">
<emphasis id="93A6EAB8A275FFC1FF2E6100FEFFAD3B" bold="true" box="[151,280,1052,1076]" pageId="8" pageNumber="419">Etymology.</emphasis>
The species name refers to the drop-shaped patch on the elytra.
</paragraph>
<paragraph id="A16D36AAA275FFC1FF2E6127FECDAD9B" blockId="8.[151,1127,219,1172]" pageId="8" pageNumber="419">
<emphasis id="93A6EAB8A275FFC1FF2E6127FEF2AD5C" bold="true" box="[151,277,1083,1107]" pageId="8" pageNumber="419">Bionomics.</emphasis>
<typeStatus id="7E698808A275FFC1FEA36120FEA8AD5B" box="[282,335,1084,1108]" pageId="8" pageNumber="419">Type</typeStatus>
specimens were found by the sifting of thick layers of wet leaf litter in the primary evergreen mountain cloud forest with very sparse understory vegetation (M. Fikáček, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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