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<document id="D7AA48B94236AA93C07B5DD2131FBCE5" ID-DOI="10.5281/zenodo.5331202" ID-ISSN="0374-1036" ID-Zenodo-Dep="5331202" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630169194360" checkinUser="diego" docAuthor="Švec, Zdeněk" docDate="2012" docId="297B87BCA279FFCFFE3B61A6FECDABBB" docLanguage="en" docName="ActaEntMusNatPra.2012.52.2.411-424.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Agathidium (Macroceble) armigerum Švec 2012, sp. nov." docType="treatment" docVersion="4" lastPageNumber="417" masterDocId="D542FFC4A27DFFC9FFB9651CFFE7A90F" masterDocTitle="New Leiodinae (Coleoptera: Leiodidae) from India and Papua New Guinea" masterLastPageNumber="424" masterPageNumber="411" pageNumber="415" updateTime="1699172137447" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2296FD3F4E7DD4DD9DDA5853F070D58B">New Leiodinae (Coleoptera: Leiodidae) from India and Papua New Guinea</mods:title>
</mods:titleInfo>
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<mods:namePart id="8FEB6A56374B9A7746ED0C4AA9D8A36C">Švec, Zdeněk</mods:namePart>
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<mods:title id="E9019214D1438D10C7DE217642FCD95C">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="0C2015E5E592A7AC0A07BC649420313C">2012</mods:date>
<mods:detail id="2103AFFF710519FA2DC960514AB1B46A" type="pubDate">
<mods:number id="48C2C97231579DB6FDDF4D4FBBBC9F1B">2012-12-17</mods:number>
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<mods:classification id="D794329935CA04AEC02A9841C1A19E86">journal article</mods:classification>
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<treatment id="297B87BCA279FFCFFE3B61A6FECDABBB" ID-DOI="http://doi.org/10.5281/zenodo.5398903" ID-Zenodo-Dep="5398903" LSID="urn:lsid:plazi:treatment:297B87BCA279FFCFFE3B61A6FECDABBB" httpUri="http://treatment.plazi.org/id/297B87BCA279FFCFFE3B61A6FECDABBB" lastPageId="6" lastPageNumber="417" pageId="4" pageNumber="415">
<subSubSection id="E9C86521A279FFCDFE3B61A6FC9BADDB" box="[386,892,1210,1236]" pageId="4" pageNumber="415" type="nomenclature">
<paragraph id="A16D36AAA279FFCDFE3B61A6FC9BADDB" blockId="4.[386,892,1210,1236]" box="[386,892,1210,1236]" pageId="4" pageNumber="415">
<heading id="FA2581C6A279FFCDFE3B61A6FC9BADDB" bold="true" box="[386,892,1210,1236]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="415" reason="3">
<emphasis id="93A6EAB8A279FFCDFE3B61A6FC9BADDB" bold="true" box="[386,892,1210,1236]" pageId="4" pageNumber="415">
<taxonomicName id="66D24D29A279FFCDFE3B61A6FCC7ADDC" ID-CoL="65PFG" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[386,800,1210,1235]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="armigerum" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A279FFCDFE3B61A6FDE3ADDC" bold="true" box="[386,516,1210,1235]" italics="true" pageId="4" pageNumber="415">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A279FFCDFDAD61A6FD72ADDC" bold="true" box="[532,661,1210,1235]" italics="true" pageId="4" pageNumber="415">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A279FFCDFD1C61A7FCC7ADDC" bold="true" box="[677,800,1211,1235]" italics="true" pageId="4" pageNumber="415">armigerum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A279FFCDFC9F61A0FC9BADDB" box="[806,892,1212,1236]" pageId="4" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A279FFCDFD8961C6FD2AADE2" box="[560,717,1242,1261]" pageId="4" pageNumber="415" type="description">
<paragraph id="A16D36AAA279FFCDFD8961C6FD2AADE2" blockId="4.[560,717,1242,1261]" box="[560,717,1242,1261]" pageId="4" pageNumber="415">
(
<figureCitation id="39E92A2FA279FFCDFD8F61C6FD6FADE2" box="[566,648,1242,1261]" captionStart="Figs" captionStartId="5.[113,152,880,899]" captionTargetBox="[158,1041,219,855]" captionTargetId="figure-221@5.[153,1048,219,862]" captionTargetPageId="5" captionText="Figs. 6–10. Genitalia of Agathidium species., 6–8 – Agathidium (Macroceble) armigerum sp. nov. (6 – aedeagus in lateral view; 7 – apex of aedeagus, dorsal view; 8 – spermatheca). 9–10 – aedeagus of A. (Macroceble) unicorne sp. nov. (9 – lateral view; 10 – apex of aedeagus, dorsal view). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331208" httpUri="https://zenodo.org/record/5331208/files/figure.png" pageId="4" pageNumber="415">Figs. 6–8</figureCitation>
,
<figureCitation id="39E92A2FA279FFCDFD2861C6FD20ADE2" box="[657,711,1242,1261]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="4" pageNumber="415">22, 23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A279FFCDFF2E6018FE04AC93" pageId="4" pageNumber="415" type="materials_examined">
<paragraph id="A16D36AAA279FFCDFF2E6018FEE2AC3E" blockId="4.[151,1127,1283,1436]" pageId="4" pageNumber="415">
<emphasis id="93A6EAB8A279FFCDFF2E6018FEEFAC18" bold="true" box="[151,264,1283,1303]" pageId="4" pageNumber="415">
<typeStatus id="7E698808A279FFCDFF2E6018FF24AC18" box="[151,195,1284,1303]" pageId="4" pageNumber="415">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="D9C5763AA279FFCDFEA86018FED9AC18" box="[273,318,1284,1303]" name="India" pageId="4" pageNumber="415">India</collectingCountry>
,
<collectingRegion id="6316F848A279FFCDFEFD6018FE03AC18" box="[324,484,1284,1303]" country="India" name="Arunachal Pradesh" pageId="4" pageNumber="415">Arunachal Pradesh</collectingRegion>
,
<quantity id="662A9B4FA279FFCDFE536018FDFAAC18" box="[490,541,1284,1303]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="415" unit="km" value="11.0">11 km</quantity>
SSE of Tenga, Eagles Nest wildlife sanctuary,
<geoCoordinate id="C4E6506DA279FFCDFC256018FC19AC18" box="[924,1022,1283,1303]" degrees="27" direction="north" minutes="06" orientation="latitude" pageId="4" pageNumber="415" precision="15" seconds="23" value="27.10639">27º06′23″N</geoCoordinate>
<geoCoordinate id="C4E6506DA279FFCDFBBB6018FB84AC18" box="[1026,1123,1283,1303]" degrees="92" direction="east" minutes="26" orientation="longitude" pageId="4" pageNumber="415" precision="15" seconds="02" value="92.43389">92º26′02″E</geoCoordinate>
,
<quantity id="662A9B4FA279FFCDFF2E6002FF3FAC3D" box="[151,216,1310,1330]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="4" pageNumber="415" unit="m" value="2510.0">2510 m</quantity>
a.s.l.
</paragraph>
<paragraph id="A16D36AAA279FFCDFF2E6025FE04AC93" blockId="4.[151,1127,1283,1436]" pageId="4" pageNumber="415">
<emphasis id="93A6EAB8A279FFCDFF2E6025FEFBAC43" bold="true" box="[151,284,1336,1356]" pageId="4" pageNumber="415">Type material.</emphasis>
<materialsCitation id="11BA3CF7A279FFCDFE986025FCA3AC8E" collectingDate="2008-05-04" collectionCode="NMPC" collectorName="Nest WL" country="India" elevation="2510" latitude="27.10639" location="Eagles" longLatPrecision="20" longitude="92.43389" pageId="4" pageNumber="415" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<typeStatus id="7E698808A279FFCDFE986025FE93AC44" box="[289,372,1337,1356]" pageId="4" pageNumber="415" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="C7C3AE6FA279FFCDFE236025FE3BAC43" box="[410,476,1337,1356]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="415">NMPC</collectionCode>
): ‘
<collectingCountry id="D9C5763AA279FFCDFE4A6025FDD1AC43" box="[499,566,1337,1356]" name="India" pageId="4" pageNumber="415">INDIA</collectingCountry>
:
<collectingRegion id="6316F848A279FFCDFD846025FD38AC43" box="[573,735,1337,1356]" country="India" name="Arunachal Pradesh" pageId="4" pageNumber="415">Arunachal Pradesh</collectingRegion>
(15) /
<locationDeviation id="B0DF2F88A279FFCDFCAE6025FC2FAC43" box="[791,968,1337,1356]" location="Tenga" pageId="4" pageNumber="415">
<quantity id="662A9B4FA279FFCDFCAE6025FCABAC43" box="[791,844,1337,1356]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="415" unit="km" value="11.0">11 km</quantity>
SSE of Tenga
</locationDeviation>
,
<location id="A40D6071A279FFCDFC686025FBEDAC43" LSID="urn:lsid:plazi:treatment:297B87BCA279FFCFFE3B61A6FECDABBB:A40D6071A279FFCDFC686025FBEDAC43" box="[977,1034,1337,1356]" country="India" latitude="27.10639" longLatPrecision="20" longitude="92.43389" name="Eagles" pageId="4" pageNumber="415" stateProvince="Arunachal Pradesh">Eagles</location>
/
<collectorName id="0C27537CA279FFCDFBA36025FB81AC43" box="[1050,1126,1337,1356]" pageId="4" pageNumber="415">Nest WL</collectorName>
sanctuary,
<date id="D56C106AA279FFCDFF496048FEDFAC68" box="[240,312,1364,1383]" pageId="4" pageNumber="415" value="2008-05-04">
<collectingDate id="C528E982A279FFCDFF496048FEDFAC68" box="[240,312,1364,1383]" pageId="4" pageNumber="415" value="2008-05-04">4.v.2008</collectingDate>
</date>
/
<geoCoordinate id="C4E6506DA279FFCDFEFF6048FE4FAC68" box="[326,424,1363,1383]" degrees="27" direction="north" minutes="06" orientation="latitude" pageId="4" pageNumber="415" precision="15" seconds="23" value="27.10639">27º06′23″N</geoCoordinate>
<geoCoordinate id="C4E6506DA279FFCDFE156048FDE8AC68" box="[428,527,1363,1383]" degrees="92" direction="east" minutes="26" orientation="longitude" pageId="4" pageNumber="415" precision="15" seconds="02" value="92.43389">92º26′02″E</geoCoordinate>
,
<quantity id="662A9B4FA279FFCDFDAF604FFDB2AC68" box="[534,597,1363,1383]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="4" pageNumber="415" unit="m" value="2510.0">
<elevation id="2AFFD199A279FFCDFDAF604FFDB2AC68" box="[534,597,1363,1383]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="4" pageNumber="415" unit="m" value="2510.0">2510 m</elevation>
</quantity>
/ Fikáček, Podskalská, Šípek lgt. // primary evergreen mountain / cloud forest with very sparse / understory vegetation, / sifting: wet leaf litter’
</materialsCitation>
.
<typeStatus id="7E698808A279FFCDFCF46073FC42AC8F" box="[845,933,1391,1410]" pageId="4" pageNumber="415" type="paratype">PARATYPES</typeStatus>
: 5 JJ
<specimenCount id="B7D4FD23A279FFCDFC5E6072FBFFAC8E" box="[999,1048,1384,1409]" pageId="4" pageNumber="415" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="C7C3AE6FA279FFCDFB9A6072FB84AC8E" box="[1059,1123,1390,1409]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="415">NMPC</collectionCode>
,
<collectionCode id="C7C3AE6FA279FFCDFF2E6095FF29AC93" box="[151,206,1417,1436]" pageId="4" pageNumber="415">ZSPC</collectionCode>
): the same data as the
<typeStatus id="7E698808A279FFCDFE2B6095FE38AC93" box="[402,479,1417,1436]" pageId="4" pageNumber="415" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A279FFCFFF2E60A8FDB9A85C" lastPageId="6" lastPageNumber="417" pageId="4" pageNumber="415" type="description">
<paragraph id="A16D36AAA279FFCDFF2E60A8FCB7AF03" blockId="4.[151,1127,1460,1612]" pageId="4" pageNumber="415">
<emphasis id="93A6EAB8A279FFCDFF2E60A8FEC5ACC3" bold="true" box="[151,290,1460,1484]" pageId="4" pageNumber="415">Description.</emphasis>
Body length
<quantity id="662A9B4FA279FFCDFE0960A8FDC9ACC3" box="[432,558,1460,1484]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="4" pageNumber="415" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
(
<typeStatus id="7E698808A279FFCDFD8560A8FD78ACC3" box="[572,671,1460,1484]" pageId="4" pageNumber="415" type="holotype">holotype</typeStatus>
:
<quantity id="662A9B4FA279FFCDFD1160A8FD1AACC3" box="[680,765,1460,1484]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="415" unit="mm" value="1.9">1.9 mm</quantity>
). Length of body parts (
<typeStatus id="7E698808A279FFCDFC4160A8FBBBACC3" box="[1016,1116,1460,1484]" pageId="4" pageNumber="415" type="holotype">holotype</typeStatus>
): head
<quantity id="662A9B4FA279FFCDFF6960C8FEC2ACE3" box="[208,293,1492,1516]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="415" unit="mm" value="0.3">0.3 mm</quantity>
, pronotum
<quantity id="662A9B4FA279FFCDFE2660C8FE14ACE3" box="[415,499,1492,1516]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="415" unit="mm" value="0.7">0.7 mm</quantity>
, elytra
<quantity id="662A9B4FA279FFCDFDFB60C8FD71ACE3" box="[578,662,1492,1516]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="415" unit="mm" value="0.9">0.9 mm</quantity>
, antenna
<quantity id="662A9B4FA279FFCDFD4260C8FCB7ACE3" box="[763,848,1492,1516]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="415" unit="mm" value="0.7">0.7 mm</quantity>
. Maximum width of body parts (
<typeStatus id="7E698808A279FFCDFF6060E8FEDAAF03" box="[217,317,1524,1548]" pageId="4" pageNumber="415" type="holotype">holotype</typeStatus>
): head
<quantity id="662A9B4FA279FFCDFE3160E8FE3BAF03" box="[392,476,1524,1548]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="415" unit="mm" value="0.9">0.9 mm</quantity>
, pronotum
<quantity id="662A9B4FA279FFCDFDEC60E8FD4EAF03" box="[597,681,1524,1548]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="415" unit="mm" value="1.2">1.2 mm</quantity>
, elytra
<quantity id="662A9B4FA279FFCDFD4E60E8FCABAF03" box="[759,844,1524,1548]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="415" unit="mm" value="1.2">1.2 mm</quantity>
.
</paragraph>
<paragraph id="A16D36AAA279FFCCFF0C6308FC5FAD06" blockId="4.[151,1127,1460,1612]" lastBlockId="5.[113,1090,1009,1578]" lastPageId="5" lastPageNumber="416" pageId="4" pageNumber="415">
Body shortly oval (
<figureCitation id="39E92A2FA279FFCDFE3B6308FE35AF23" box="[386,466,1556,1580]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="4" pageNumber="415">Fig. 22</figureCitation>
), red-brown with lighter anterior part of head, margins of pronotum and margins of elytra; antennae and legs reddish. Venter light yellow. Dorsum without microreticulation; feebly punctate, each puncture with very short and fine seta.
</paragraph>
<caption id="F5AD6622A278FFCCFFC8666CFCC3AAB7" ID-DOI="http://doi.org/10.5281/zenodo.5331208" ID-Zenodo-Dep="5331208" httpUri="https://zenodo.org/record/5331208/files/figure.png" pageId="5" pageNumber="416" startId="5.[113,152,880,899]" targetBox="[158,1041,219,855]" targetPageId="5">
<paragraph id="A16D36AAA278FFCCFFC8666CFCC3AAB7" blockId="5.[113,1090,880,952]" pageId="5" pageNumber="416">
Figs. 6–10. Genitalia of
<taxonomicName id="66D24D29A278FFCCFEF9666DFE08AA8C" box="[320,495,880,899]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="93A6EAB8A278FFCCFEF9666DFE43AA8C" box="[320,420,881,899]" italics="true" pageId="5" pageNumber="416">Agathidium</emphasis>
species.
</taxonomicName>
, 6–8 –
<taxonomicName id="66D24D29A278FFCCFD92666DFC8CAA8C" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[555,875,880,899]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="armigerum" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A278FFCCFD92666DFD68AA8C" box="[555,655,881,899]" italics="true" pageId="5" pageNumber="416">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A278FFCCFD23666DFD18AA8C" box="[666,767,881,899]" italics="true" pageId="5" pageNumber="416">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A278FFCCFCB5666DFC8CAA8C" box="[780,875,881,899]" italics="true" pageId="5" pageNumber="416">armigerum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A278FFCCFCC9666CFC55AA8C" box="[880,946,880,899]" pageId="5" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
(6 – aedeagus in lateral view; 7 – apex of aedeagus, dorsal view; 8 – spermatheca). 9–10 – aedeagus of
<taxonomicName id="66D24D29A278FFCCFCF06697FBC3AA92" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[841,1060,906,925]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="unicorne" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A278FFCCFCF06697FCBBAA92" box="[841,860,907,925]" italics="true" pageId="5" pageNumber="416">A.</emphasis>
(
<emphasis id="93A6EAB8A278FFCCFCDE6697FC2BAA92" box="[871,972,907,925]" italics="true" pageId="5" pageNumber="416">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A278FFCCFC616697FBC3AA92" box="[984,1060,907,925]" italics="true" pageId="5" pageNumber="416">unicorne</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A278FFCCFB916697FF70AAB7" pageId="5" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
(9 – lateral view; 10 – apex of aedeagus, dorsal view). Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="A16D36AAA278FFCCFF29610DFBCEAD86" blockId="5.[113,1090,1009,1578]" pageId="5" pageNumber="416">Head. Maximum width of head just before posterior margin of eyes. Eyes flat, narrow, 11× as long as wide in dorsal view. Clypeus feebly emarginate, clypeal line lacking. Dorsal surface with very sparse, scattered fine punctures separated at least by ca. 10× their diameter. Mandibles without striking characters. Relative length of antennal segments III: II = 1.6.</paragraph>
<paragraph id="A16D36AAA278FFCCFF29618EFD28ADE6" blockId="5.[113,1090,1009,1578]" pageId="5" pageNumber="416">
Pronotum. Widest just before base. Lateral sides roundly tapered anteriorly in dorsal view; rounded in lateral view (
<figureCitation id="39E92A2FA278FFCCFEC161ADFE2FADC5" box="[376,456,1201,1226]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="5" pageNumber="416">Fig. 23</figureCitation>
). Puncturation sparser and finer than that of head, with few slightly longer setae close to lateral margin of pronotum.
</paragraph>
<paragraph id="A16D36AAA278FFCCFF2961EDFD36AC26" blockId="5.[113,1090,1009,1578]" pageId="5" pageNumber="416">Elytra. Broadest just before midlength. Punctures as on pronotum. Surface with few slightly longer setae close to lateral margins. Sutural stria absent.</paragraph>
<paragraph id="A16D36AAA278FFCCFF29602EFCB8AC66" blockId="5.[113,1090,1009,1578]" pageId="5" pageNumber="416">
Legs. Protarsomeres I–III and mesotarsomere I a little enlarged in male. Tarsi slender in female. Femora simple. Tarsal formula 5-5-
<quantity id="662A9B4FA278FFCCFDF8604DFD8CAC66" box="[577,619,1361,1385]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="416" unit="in" value="4.0">4 in</quantity>
male, 4-4-
<quantity id="662A9B4FA278FFCCFD67604EFCEFAC66" box="[734,776,1361,1386]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="416" unit="in" value="4.0">4 in</quantity>
female.
</paragraph>
<paragraph id="A16D36AAA278FFCCFF29606DFE87ACA6" blockId="5.[113,1090,1009,1578]" pageId="5" pageNumber="416">Mesoventrite. Longitudinal mesoventral carina present; lateral lines not developed. Metathoracic wings absent.</paragraph>
<paragraph id="A16D36AAA278FFCCFF2960ADFD0BACE6" blockId="5.[113,1090,1009,1578]" pageId="5" pageNumber="416">Metaventrite. Femoral lines absent. Male and female with high hook-shaped ventro-caudally orientated process at posterior margin between coxae.</paragraph>
<paragraph id="A16D36AAA278FFCCFF2960EDFDCAAF25" blockId="5.[113,1090,1009,1578]" pageId="5" pageNumber="416">
Genitalia. Aedeagus as in
<figureCitation id="39E92A2FA278FFCCFE0A60EDFDFAAF06" box="[435,541,1521,1545]" captionStart="Figs" captionStartId="5.[113,152,880,899]" captionTargetBox="[158,1041,219,855]" captionTargetId="figure-221@5.[153,1048,219,862]" captionTargetPageId="5" captionText="Figs. 6–10. Genitalia of Agathidium species., 6–8 – Agathidium (Macroceble) armigerum sp. nov. (6 – aedeagus in lateral view; 7 – apex of aedeagus, dorsal view; 8 – spermatheca). 9–10 – aedeagus of A. (Macroceble) unicorne sp. nov. (9 – lateral view; 10 – apex of aedeagus, dorsal view). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331208" httpUri="https://zenodo.org/record/5331208/files/figure.png" pageId="5" pageNumber="416">Figs. 6–7</figureCitation>
; operculum T-shaped with slight emargination on anterior margin; spermatheca as in
<figureCitation id="39E92A2FA278FFCCFE51630DFDCEAF25" box="[488,553,1553,1578]" captionStart="Figs" captionStartId="5.[113,152,880,899]" captionTargetBox="[158,1041,219,855]" captionTargetId="figure-221@5.[153,1048,219,862]" captionTargetPageId="5" captionText="Figs. 6–10. Genitalia of Agathidium species., 6–8 – Agathidium (Macroceble) armigerum sp. nov. (6 – aedeagus in lateral view; 7 – apex of aedeagus, dorsal view; 8 – spermatheca). 9–10 – aedeagus of A. (Macroceble) unicorne sp. nov. (9 – lateral view; 10 – apex of aedeagus, dorsal view). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331208" httpUri="https://zenodo.org/record/5331208/files/figure.png" pageId="5" pageNumber="416">Fig. 8</figureCitation>
.
</paragraph>
<paragraph id="A16D36AAA27BFFCFFF2E65C7FDB9A85C" blockId="6.[151,1127,219,692]" pageId="6" pageNumber="417">
<emphasis id="93A6EAB8A27BFFCFFF2E65C7FEF1A9FC" bold="true" box="[151,278,219,243]" pageId="6" pageNumber="417">Variability.</emphasis>
The length ratio of antennal segments III: II varies between
<quantity id="662A9B4FA27BFFCFFC1965C7FBECA9FC" box="[928,1035,219,243]" metricMagnitude="-2" metricUnit="m" metricValue="4.191" metricValueMax="4.572" metricValueMin="3.81" pageId="6" pageNumber="417" unit="in" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 in</quantity>
the
<typeStatus id="7E698808A27BFFCFFB8065C0FB80A9FB" box="[1081,1127,220,244]" pageId="6" pageNumber="417">type</typeStatus>
specimens; the ratio of length:width of the eyes in dorsal view is between 8–11. The coloration of the dorsum varies from uniformly light chestnut to a bicoloured dorsum with a reddish head and pronotum and dark brown elytra.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27BFFCFFF2E6447FD9CAB3C" pageId="6" pageNumber="417" type="diagnosis">
<paragraph id="A16D36AAA27BFFCFFF2E6447FD9CAB3C" blockId="6.[151,1127,219,692]" pageId="6" pageNumber="417">
<emphasis id="93A6EAB8A27BFFCFFF2E6447FE77A87C" bold="true" box="[151,400,347,371]" pageId="6" pageNumber="417">Differential diagnosis.</emphasis>
<taxonomicName id="66D24D29A27BFFCFFE2D6441FCC6A87B" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[404,801,348,372]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="armigerum" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A27BFFCFFE2D6441FDF7A87B" box="[404,528,349,372]" italics="true" pageId="6" pageNumber="417">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A27BFFCFFDA66441FD7AA87B" box="[543,669,349,372]" italics="true" pageId="6" pageNumber="417">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A27BFFCFFD126441FCC6A87B" box="[683,801,349,372]" italics="true" pageId="6" pageNumber="417">armigerum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A27BFFCFFC9F6440FC90A87C" box="[806,887,348,371]" pageId="6" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<emphasis id="93A6EAB8A27BFFCFFC426441FBF4A87B" box="[1019,1043,349,372]" italics="true" pageId="6" pageNumber="417">A.</emphasis>
(
<emphasis id="93A6EAB8A27BFFCFFB996441FBB9A87B" box="[1056,1118,349,372]" italics="true" pageId="6" pageNumber="417">Macr.</emphasis>
)
<taxonomicName id="66D24D29A27BFFCFFF2E6461FDF6A89B" authority="Angelini &amp; DeMarzo, 1981" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[151,529,379,404]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="93A6EAB8A27BFFCFFF2E6461FF00A89B" box="[151,231,381,404]" italics="true" pageId="6" pageNumber="417">caelebs</emphasis>
Angelini &amp; DeMarzo,
<quantity id="662A9B4FA27BFFCFFE656460FDF6A89B" box="[476,529,380,404]" metricMagnitude="1" metricUnit="m" metricValue="5.03174" pageId="6" pageNumber="417" unit="in" value="1981.0">1981</quantity>
</taxonomicName>
in the size of body and the coloration of the dorsum, the shape of the pronotum having the lateral margin broadly rounded, and also in the shape of the spermatheca. It differs from the latter in having a longer antennomere III with the length ratio of antennomeres III: II = 1.5–1.8 (compared to
<quantity id="662A9B4FA27BFFCFFD0664C0FD1DA8FC" box="[703,762,475,500]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="6" pageNumber="417" unit="in" value="1.4">1.4 in</quantity>
<taxonomicName id="66D24D29A27BFFCFFD4664C1FC81A8FB" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[767,870,477,500]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="93A6EAB8A27BFFCFFD4664C1FC81A8FB" box="[767,870,477,500]" italics="true" pageId="6" pageNumber="417">A.caelebs</emphasis>
</taxonomicName>
), tarsal formula 4-4-
<quantity id="662A9B4FA27BFFCFFB8764C0FB80A8FC" box="[1086,1127,475,500]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="417" unit="in" value="4.0">4 in</quantity>
female (5-4-
<quantity id="662A9B4FA27BFFCFFEA364E7FEA4AB1C" box="[282,323,507,531]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="417" unit="in" value="4.0">4 in</quantity>
female of
<taxonomicName id="66D24D29A27BFFCFFE0D64E1FDC6AB1B" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[436,545,509,532]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="93A6EAB8A27BFFCFFE0D64E1FDC6AB1B" box="[436,545,509,532]" italics="true" pageId="6" pageNumber="417">A. caelebs</emphasis>
</taxonomicName>
), and by the shape of the tegmen that is not constricted laterally before the apex as it is in
<taxonomicName id="66D24D29A27BFFCFFDBF6701FD93AB3B" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[518,628,541,564]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="93A6EAB8A27BFFCFFDBF6701FD93AB3B" box="[518,628,541,564]" italics="true" pageId="6" pageNumber="417">A. caelebs</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27BFFCFFF2E6720FECDABBB" pageId="6" pageNumber="417" type="etymology">
<paragraph id="A16D36AAA27BFFCFFF2E6720FBE6AB5C" blockId="6.[151,1127,219,692]" box="[151,1025,571,596]" pageId="6" pageNumber="417">
<emphasis id="93A6EAB8A27BFFCFFF2E6720FEFFAB5B" bold="true" box="[151,280,572,596]" pageId="6" pageNumber="417">Etymology.</emphasis>
The species name refers to the hook-like process on the metaventrite.
</paragraph>
<paragraph id="A16D36AAA27BFFCFFF2E6747FECDABBB" blockId="6.[151,1127,219,692]" pageId="6" pageNumber="417">
<emphasis id="93A6EAB8A27BFFCFFF2E6747FEF2AB7C" bold="true" box="[151,277,603,627]" pageId="6" pageNumber="417">Bionomics.</emphasis>
<typeStatus id="7E698808A27BFFCFFEA36740FEA8AB7B" box="[282,335,604,628]" pageId="6" pageNumber="417">Type</typeStatus>
specimens were found by the sifting of thick layers of wet leaf litter in the primary evergreen mountain cloud forest with very sparse understory vegetation (M. Fikáček, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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