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<document id="ACDACBA034E80D0F215B1E3157A9B809" ID-DOI="10.11646/zootaxa.5039.4.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="5515923" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632051570394" checkinUser="plazi" docAuthor="Pham, Phu Van &amp; Pham, Nhi Thi" docDate="2021" docId="241587CCFFCC0A5EE0D9F9A3FE120316" docLanguage="en" docName="zootaxa.5039.4.10.pdf" docOrigin="Zootaxa 5039 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macronotops flavomedius Pham &amp; Pham 2021, new species" docType="treatment" docVersion="3" lastPageNumber="594" masterDocId="D82CFFB4FFCC0A5FE04EFF9EFFA70455" masterDocTitle="A remarkable new species of the genus Macronotops Krikken, 1977 (Coleoptera: Scarabaeidae: Cetoniinae) from Vietnam" masterLastPageNumber="598" masterPageNumber="593" pageNumber="593" updateTime="1699414812410" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="8144C21398FF63DA14FF5D0389A4AA15" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="57730B1948DBC9A6A5B43FCAE9FD76DD">
<mods:title id="A21315130D49B3542F0F98F0ECB4C3B4">A remarkable new species of the genus Macronotops Krikken, 1977 (Coleoptera: Scarabaeidae: Cetoniinae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="922AEF2B801874B42C201E6EFE59C14D" type="personal">
<mods:role id="95D6840FE014E550178720AC769478CE">
<mods:roleTerm id="81080144579964FBB041125E400DDA15">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3CC45E4A9D1D1A641EC19A05E13917C5">Pham, Phu Van</mods:namePart>
<mods:nameIdentifier id="F2EF8E6BE894E2FF0E584B767BF49521" type="ORCID">0000-0001-9369-6649</mods:nameIdentifier>
<mods:affiliation id="AAD7A94A00B42F09716DBD2E66216108">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi. phupham. iebr @ gmail. com; https: // orcid. org / 0000 - 0001 - 9369 - 6649</mods:affiliation>
<mods:nameIdentifier id="D116FFA0B36BEF78510DAA6B51BF9893" type="email">phupham.iebr@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="3375A29D6520CF16BAC938449728131C" type="personal">
<mods:role id="8FE35DF1828F2B2499F74136186968C7">
<mods:roleTerm id="F3BF85758945A3320BC1BFE960E5BBCA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CEED40E00B29A31787950F1918B9ABFF">Pham, Nhi Thi</mods:namePart>
<mods:nameIdentifier id="CC86311D8987FD3807F2B9F09F8D8390" type="ORCID">0000-0001-9369-6649</mods:nameIdentifier>
<mods:affiliation id="391D641E4483867839990535A6C8FAC6">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi. phupham. iebr @ gmail. com; https: // orcid. org / 0000 - 0001 - 9369 - 6649 &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay Hanoi. ptnhi 2 @ yahoo. com, https: // orcid. org / 0000 - 0001 - 9304 - 9863</mods:affiliation>
<mods:nameIdentifier id="F5F76198AF350AB354ABA0BAE388D20C" type="email">phupham.iebr@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="CE4DC8EDA0D7C387AD3EF10CA1739BBE">text</mods:typeOfResource>
<mods:relatedItem id="3D0A4CDD051A0323A67FA7A02F4B7BF3" type="host">
<mods:titleInfo id="03C7D1AAF825C678B69D20F7FE58FC32">
<mods:title id="DC792CD20B06F12D4B59D50496580449">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FC119F120DBC1E1BFF0284D93ED7DA16">
<mods:date id="C3B790BF7D3F91CC75F18207C07D8DC3">2021</mods:date>
<mods:detail id="B8F7307A1C7EFDA1FEDC5769E47E81BA" type="pubDate">
<mods:number id="E357BDAC4FBF2D983ABE984F2CA49CE2">2021-09-17</mods:number>
</mods:detail>
<mods:detail id="3C135955A9D84DD24310804B5A983CE4" type="volume">
<mods:number id="F30F3114BB31303EFFA05B15C1722D90">5039</mods:number>
</mods:detail>
<mods:detail id="EA37B9759A84DE239F3B614408A8C1A2" type="issue">
<mods:number id="EED375EDCFBFCC6670F4D3CBFBE5A871">4</mods:number>
</mods:detail>
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<mods:start id="5C1D89048695B1171D08EF4E44F90CE6">593</mods:start>
<mods:end id="A85349A78B35158B16F498BB374BE4C5">598</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BDA5D92F60FD5F6E986E066127CE3B98">journal article</mods:classification>
<mods:identifier id="E8D35673923456CD235393684552E7C6" type="DOI">10.11646/zootaxa.5039.4.10</mods:identifier>
<mods:identifier id="C91FDFC66DAFEFAE257070B03467ECAB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C5BF073372ABC58A6D3CFFD573C1E201" type="Zenodo-Dep">5515923</mods:identifier>
</mods:mods>
<treatment id="241587CCFFCC0A5EE0D9F9A3FE120316" LSID="urn:lsid:zoobank.org:act:32115B46-E3F5-45AD-BBD8-77E28605464F" httpUri="http://treatment.plazi.org/id/241587CCFFCC0A5EE0D9F9A3FE120316" lastPageId="1" lastPageNumber="594" pageId="0" pageNumber="593">
<subSubSection id="E4A66551FFCC0A5FE0D9F9A3FC30022F" pageId="0" pageNumber="593" type="nomenclature">
<paragraph id="AC0336DAFFCC0A5FE0D9F9A3FC1C0203" blockId="0.[151,1437,1596,2019]" box="[151,955,1596,1623]" pageId="0" pageNumber="593">
<heading id="F74B81B6FFCC0A5FE0D9F9A3FC1C0203" bold="true" box="[151,955,1596,1623]" fontSize="11" level="1" pageId="0" pageNumber="593" reason="1">
<emphasis id="9EC8EAC8FFCC0A5FE0D9F9A3FC1C0203" bold="true" box="[151,955,1596,1623]" pageId="0" pageNumber="593">
<taxonomicName id="6BBC4D59FFCC0A5FE0D9F9A3FD330202" authority="Pham &amp; Pham" authorityName="Pham &amp; Pham" authorityYear="2021" box="[151,660,1596,1623]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="593" phylum="Arthropoda" rank="species" species="flavomedius" status="sp. nov.">
<emphasis id="9EC8EAC8FFCC0A5FE0D9F9A3FE760202" bold="true" box="[151,465,1596,1623]" italics="true" pageId="0" pageNumber="593">Macronotops flavomedius</emphasis>
Pham &amp; Pham
</taxonomicName>
,
<taxonomicNameLabel id="85FB57B3FFCC0A5FE2EFF9A3FC960203" box="[673,817,1596,1623]" pageId="0" pageNumber="593" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="34872A5FFFCC0A5FE30EF9A2FC160203" box="[832,945,1596,1622]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1943,1967]" captionStartId-2="4.[151,250,1650,1674]" captionTargetBox-0="[192,1386,200,1899]" captionTargetBox-1="[186,1400,195,1919]" captionTargetBox-2="[151,1435,181,1626]" captionTargetId-0="figure-18@2.[176,1412,180,1919]" captionTargetId-1="figure-17@3.[186,1402,181,1919]" captionTargetId-2="figure-18@4.[151,1436,181,1626]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Habitus of Macronotop flavomedius Pham &amp; Pham, new species. a, male dorsal view; b, male ventral view; c, female dorsal view; d, female ventral view. (Figs 1c, d were taken by Dr. Hao Xu)." captionText-1="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." captionText-2="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi-0="http://doi.org/10.5281/zenodo.5515925" figureDoi-1="http://doi.org/10.5281/zenodo.5515927" figureDoi-2="http://doi.org/10.5281/zenodo.5515929" httpUri-0="https://zenodo.org/record/5515925/files/figure.png" httpUri-1="https://zenodo.org/record/5515927/files/figure.png" httpUri-2="https://zenodo.org/record/5515929/files/figure.png" pageId="0" pageNumber="593">Figs. 1–3</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="AC0336DAFFCC0A5FE0D9F9FCFC30022F" blockId="0.[151,1437,1596,2019]" box="[151,919,1634,1658]" pageId="0" pageNumber="593">
<uri id="D82D3AD8FFCC0A5FE0D9F9FCFC30022F" box="[151,919,1634,1658]" pageId="0" pageNumber="593">
urn:lsid:zoobank.org:act:
<uuid id="D81A0C0FFFCC0A5FE1D5F9FCFC30022F" box="[411,919,1634,1658]" pageId="0" pageNumber="593">32115B46-E3F5-45AD-BBD8-77E28605464F</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="E4A66551FFCC0A5FE0D9F919FD8702B3" pageId="0" pageNumber="593" type="materials_examined">
<paragraph id="AC0336DAFFCC0A5FE0D9F919FD8702B3" blockId="0.[151,1437,1596,2019]" pageId="0" pageNumber="593">
Type material.
<materialsCitation id="1CD43C87FFCC0A5FE10AF919FAF602CB" box="[324,1361,1670,1697]" collectingDate="2002-06-02" collectingDateMax="2002-06-03" collectingDateMin="2002-06-02" collectionCode="IEBR" location="Langbian Moutain" municipality="Lac Duong" pageId="0" pageNumber="593" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="holotype">
<emphasis id="9EC8EAC8FFCC0A5FE10AF919FE6002F4" bold="true" box="[324,455,1671,1697]" pageId="0" pageNumber="593">
<typeStatus id="73078878FFCC0A5FE10AF919FE0B02CA" box="[324,428,1671,1695]" pageId="0" pageNumber="593" type="holotype">Holotype</typeStatus>
<specimenCount id="BABAFD53FFCC0A5FE1FDF919FE6002F4" box="[435,455,1671,1697]" pageId="0" pageNumber="593" type="male">♂</specimenCount>
</emphasis>
(
<collectionCode id="CAADAE1FFFCC0A5FE19BF919FDB002CA" box="[469,535,1671,1695]" pageId="0" pageNumber="593">IEBR</collectionCode>
),
<collectingRegion id="6E78F838FFCC0A5FE264F919FCA102CB" box="[554,774,1670,1695]" country="Vietnam" name="Lam Dong" pageId="0" pageNumber="593">Lam Dong Province</collectingRegion>
,
<collectingMunicipality id="4C67ACA0FFCC0A5FE35FF919FC2C02CA" box="[785,907,1671,1695]" pageId="0" pageNumber="593">Lac Duong</collectingMunicipality>
,
<location id="A9636001FFCC0A5FE3D8F918FBF802CB" LSID="urn:lsid:plazi:treatment:241587CCFFCC0A5EE0D9F9A3FE120316:A9636001FFCC0A5FE3D8F918FBF802CB" box="[918,1119,1670,1694]" municipality="Lac Duong" name="Langbian Moutain" pageId="0" pageNumber="593" stateProvince="Lam Dong">Langbian Moutain</location>
,
<collectingMethod id="75FD4ECDFFCC0A5FE424F918FB6F02CB" box="[1130,1224,1670,1694]" pageId="0" pageNumber="593">hand net</collectingMethod>
,
<date id="D802101AFFCC0A5FE49DF918FAF602CB" box="[1235,1361,1670,1694]" pageId="0" pageNumber="593" value="2002-06-02" valueMax="2002-06-03" valueMin="2002-06-02">
<collectingDate id="C846E9F2FFCC0A5FE49DF918FAF602CB" box="[1235,1361,1670,1694]" pageId="0" pageNumber="593" value="2002-06-02" valueMax="2002-06-03" valueMin="2002-06-02">2-3.vi.2002</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="1CD43C87FFCC0A5FE513F919FD4F0297" collectingDate="2002-06-02" collectingDateMax="2002-06-03" collectingDateMin="2002-06-02" collectionCode="IEBR" location="Langbian Moutain" municipality="Lac Duong" pageId="0" pageNumber="593" specimenCount="2" specimenCount-female="2" stateProvince="Lam Dong" typeStatus="paratype">
<emphasis id="9EC8EAC8FFCC0A5FE513F919FF7C0296" bold="true" pageId="0" pageNumber="593">
<typeStatus id="73078878FFCC0A5FE513F919FF710296" pageId="0" pageNumber="593" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="BABAFD53FFCC0A5FE0ACF935FEB80297" box="[226,287,1707,1731]" pageId="0" pageNumber="593" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="CAADAE1FFFCC0A5FE160F935FED70296" box="[302,368,1707,1731]" pageId="0" pageNumber="593">IEBR</collectionCode>
), the same data as for the holotype
</materialsCitation>
;
<materialsCitation id="1CD43C87FFCC0A5FE2BDF935FD8702B3" collectingDate="2002-06-03" collectionCode="NSMT, S-VI-, ETNAM" collectorName="Da Lat &amp; S. Nomura" location="Da Lat" municipality="Mt. Lang Biang" pageId="0" pageNumber="593" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCount id="BABAFD53FFCC0A5FE2BDF935FCB30296" box="[755,788,1707,1731]" pageId="0" pageNumber="593" type="female">1♀</specimenCount>
(
<collectionCode id="CAADAE1FFFCC0A5FE36DF935FCD50296" box="[803,882,1707,1731]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="0" pageNumber="593">NSMT</collectionCode>
),
<collectingMunicipality id="4C67ACA0FFCC0A5FE3CBF935FB950297" box="[901,1074,1706,1731]" pageId="0" pageNumber="593">Mt. Lang Biang</collectingMunicipality>
, nr.
<collectorName id="0149530CFFCC0A5FE42EF935FB0A0297" box="[1120,1197,1707,1730]" pageId="0" pageNumber="593">
<location id="A9636001FFCC0A5FE42EF935FB0A0297" LSID="urn:lsid:plazi:treatment:241587CCFFCC0A5EE0D9F9A3FE120316:A9636001FFCC0A5FE42EF935FB0A0297" box="[1120,1197,1707,1730]" municipality="Mt. Lang Biang" name="Da Lat" pageId="0" pageNumber="593" stateProvince="Lam Dong">Da Lat</location>
</collectorName>
,
<collectingRegion id="6E78F838FFCC0A5FE4F6F935FA8C0296" box="[1208,1323,1707,1731]" country="Vietnam" name="Lam Dong" pageId="0" pageNumber="593">Lam Dong</collectingRegion>
Pr. [
<collectionCode id="CAADAE1FFFCC0A5FE513F935FF5C02B2" pageId="0" pageNumber="593">S-VI- ETNAM</collectionCode>
],
<date id="D802101AFFCC0A5FE142F950FED402B3" box="[268,371,1742,1766]" pageId="0" pageNumber="593" value="2002-06-03">
<collectingDate id="C846E9F2FFCC0A5FE142F950FED402B3" box="[268,371,1742,1766]" pageId="0" pageNumber="593" value="2002-06-03">3.vi.2002</collectingDate>
</date>
,
<collectorName id="0149530CFFCC0A5FE130F951FE5602B3" box="[382,497,1743,1767]" pageId="0" pageNumber="593">S. Nomura</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E4A66551FFCC0A5EE089F96CFB9D00CA" lastPageId="1" lastPageNumber="594" pageId="0" pageNumber="593" type="description">
<paragraph id="AC0336DAFFCC0A5EE089F96CFD9E0792" blockId="0.[151,1437,1596,2019]" lastBlockId="1.[151,1437,151,1256]" lastPageId="1" lastPageNumber="594" pageId="0" pageNumber="593">
<emphasis id="9EC8EAC8FFCC0A5FE089F96CFD61035F" bold="true" box="[199,710,1778,1803]" pageId="0" pageNumber="593">
<emphasis id="9EC8EAC8FFCC0A5FE089F96CFDFE035F" box="[199,601,1778,1803]" inLineHeading="true" pageId="0" pageNumber="593" reason="1">
Description of the
<typeStatus id="73078878FFCC0A5FE1D1F96DFDA6035E" box="[415,513,1779,1803]" pageId="0" pageNumber="593" type="holotype">holotype</typeStatus>
(male).
</emphasis>
General:
</emphasis>
Body length
<quantity id="6B449B3FFFCC0A5FE312F96CFC66035F" box="[860,961,1778,1802]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="0" pageNumber="593" unit="mm" value="14.5">14.5 mm</quantity>
; width
<quantity id="6B449B3FFFCC0A5FE45DF96CFBCE035F" box="[1043,1129,1778,1802]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="0" pageNumber="593" unit="mm" value="6.3">6.3 mm</quantity>
, widest at humeral umbone, gradually narrowed backward. Body brown. Surface with fulvous, brown, light yellow and black setae, and yellow tomentous maculae.
<emphasis id="9EC8EAC8FFCC0A5FE110F8A5FE030306" bold="true" box="[350,420,1851,1875]" pageId="0" pageNumber="593">Head:</emphasis>
Dorsal surface with dense, setiferous punctures; setae fulvous, long. Clypeus with weak brown metalic reflections; anterior margin slightly notched, and slightly raised; clypeolateral ridge not distinct; sides with yellow maculae (
<figureCitation id="34872A5FFFCC0A5FE0B1F81CFEE903CE" box="[255,334,1922,1947]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[186,1400,195,1919]" captionTargetId="figure-17@3.[186,1402,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." figureDoi="http://doi.org/10.5281/zenodo.5515927" httpUri="https://zenodo.org/record/5515927/files/figure.png" pageId="0" pageNumber="593">Fig. 2e</figureCitation>
). Frons with a morderately raised longitudinal ridge. Maxillary palpus and labial palpus dark brown to light brown. Antenna brown, antennal club long, approximately 2.1 times length of antennomeres 2–7 combined; inner side of antennomere 8 with short, dense, fulvous satae. Ventral surface clad with long, yellowish setae (
<figureCitation id="34872A5FFFCC0A5FE4A2F854FA9D03B6" box="[1260,1338,1994,2019]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1435,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi="http://doi.org/10.5281/zenodo.5515929" httpUri="https://zenodo.org/record/5515929/files/figure.png" pageId="0" pageNumber="593">Fig. 3a</figureCitation>
).
<emphasis id="9EC8EAC8FFCC0A5EE500F855FF6204E5" bold="true" lastPageId="1" lastPageNumber="594" pageId="0" pageNumber="593">Pronotum</emphasis>
: Widest at base, basomedian area depressed. Lateral margin curved; posterior margin distinctly protruded. Surface clad with long, yellow, and black setae except for basolateral area; median area with a longitudinal yellow tomentous maculae (
<figureCitation id="34872A5FFFCD0A5EE0B1FF41FEE904AD" box="[255,334,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[186,1400,195,1919]" captionTargetId="figure-17@3.[186,1402,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." figureDoi="http://doi.org/10.5281/zenodo.5515927" httpUri="https://zenodo.org/record/5515927/files/figure.png" pageId="1" pageNumber="594">
Fig.
<quantity id="6B449B3FFFCD0A5EE17EFF7EFEE904AD" box="[304,334,224,248]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="1" pageNumber="594" unit="g" value="2.0">2g</quantity>
</figureCitation>
).
<emphasis id="9EC8EAC8FFCD0A5EE12FFF41FE7B04A2" bold="true" box="[353,476,223,247]" pageId="1" pageNumber="594">Scutellum:</emphasis>
Black, surface clad with long, white yellow setae; median area with a longitudinal yellow tomentous maculae gradually narrow at apex.
<emphasis id="9EC8EAC8FFCD0A5EE2C8FE9AFD410549" bold="true" box="[646,742,260,284]" pageId="1" pageNumber="594">Elytron:</emphasis>
With posthumeral macula, lateral macula, small median macula, and distal macula (
<figureCitation id="34872A5FFFCD0A5EE111FEB9FE120515" box="[351,437,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1386,200,1899]" captionTargetId="figure-18@2.[176,1412,180,1919]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Macronotop flavomedius Pham &amp; Pham, new species. a, male dorsal view; b, male ventral view; c, female dorsal view; d, female ventral view. (Figs 1c, d were taken by Dr. Hao Xu)." figureDoi="http://doi.org/10.5281/zenodo.5515925" httpUri="https://zenodo.org/record/5515925/files/figure.png" pageId="1" pageNumber="594">Figs. 1a</figureCitation>
,
<figureCitation id="34872A5FFFCD0A5EE18FFEB6FE7B0515" box="[449,476,296,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[186,1400,195,1919]" captionTargetId="figure-17@3.[186,1402,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." figureDoi="http://doi.org/10.5281/zenodo.5515927" httpUri="https://zenodo.org/record/5515927/files/figure.png" pageId="1" pageNumber="594">2c</figureCitation>
,
<figureCitation id="34872A5FFFCD0A5EE1A9FEB6FDA50515" box="[487,514,296,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1435,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi="http://doi.org/10.5281/zenodo.5515929" httpUri="https://zenodo.org/record/5515929/files/figure.png" pageId="1" pageNumber="594">3c</figureCitation>
); median macula small and divided into two small parts (
<figureCitation id="34872A5FFFCD0A5EE42EFEB9FB090515" box="[1120,1198,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1435,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi="http://doi.org/10.5281/zenodo.5515929" httpUri="https://zenodo.org/record/5515929/files/figure.png" pageId="1" pageNumber="594">Fig. 3c</figureCitation>
). Surface densely clad with setae; setae on maculae and disc long and light yellow; setae black behind humeral umbone and on lateral and distal declivity; setae dark brown on juxtascutellar area; setae fulvous along lateral margins; humeral umbone glabrous (
<figureCitation id="34872A5FFFCD0A5EE53FFEF1FF1505F9" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1386,200,1899]" captionTargetId="figure-18@2.[176,1412,180,1919]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Macronotop flavomedius Pham &amp; Pham, new species. a, male dorsal view; b, male ventral view; c, female dorsal view; d, female ventral view. (Figs 1c, d were taken by Dr. Hao Xu)." figureDoi="http://doi.org/10.5281/zenodo.5515925" httpUri="https://zenodo.org/record/5515925/files/figure.png" pageId="1" pageNumber="594">Fig. 1a</figureCitation>
).
<emphasis id="9EC8EAC8FFCD0A5EE086FE0DFECB05FE" bold="true" box="[200,364,403,427]" pageId="1" pageNumber="594">Mesepimeron:</emphasis>
Dark brown, clad with yellowish setae, and yellow tomentous maculae.
<emphasis id="9EC8EAC8FFCD0A5EE431FE0DFEAB059A" bold="true" pageId="1" pageNumber="594">Metepisternum and metepimeron:</emphasis>
dark brown, clad with yellowish setae.
<emphasis id="9EC8EAC8FFCD0A5EE2FBFE29FC86059A" bold="true" box="[693,801,439,463]" pageId="1" pageNumber="594">Sternum:</emphasis>
Preprosternum dark brown, almost glabrous. Mesosternum black, clad with long, sparse, light yellow setae. Mesometasternal process short, glabrous, brown, apex rounded; mesometasternal suture depressed, almost without setae. Metasternum dark brown, with long, sparse, yellow setae; middle portion glabrous (
<figureCitation id="34872A5FFFCD0A5EE14CFDBDFEF70669" box="[258,336,547,572]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1435,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi="http://doi.org/10.5281/zenodo.5515929" httpUri="https://zenodo.org/record/5515929/files/figure.png" pageId="1" pageNumber="594">Fig. 3e</figureCitation>
).
<emphasis id="9EC8EAC8FFCD0A5EE12AFDBDFE7D066E" bold="true" box="[356,474,547,571]" pageId="1" pageNumber="594">Pygidium:</emphasis>
Brown; with a yellow macula. Sides clad moderate setae, dense, fulvous and dark brown; macula clad light yellow setae (
<figureCitation id="34872A5FFFCD0A5EE1A4FDD9FD9F0635" box="[490,568,583,608]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[186,1400,195,1919]" captionTargetId="figure-17@3.[186,1402,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." figureDoi="http://doi.org/10.5281/zenodo.5515927" httpUri="https://zenodo.org/record/5515927/files/figure.png" pageId="1" pageNumber="594">Fig. 2c</figureCitation>
).
<emphasis id="9EC8EAC8FFCD0A5EE202FDD9FD63060A" bold="true" box="[588,708,583,607]" pageId="1" pageNumber="594">Abdomen:</emphasis>
Without median groove; median portion almost glabrous, with only some short, yellow setae. Six abdominal sternites visible; sternites II–V clad with yellow setae; setae on sternite II long, dense; setae on sternites III–V short, sparse, but shorter on sides; sternite VII with sparse, long, fulvous setae on sides. Sternites II–VI with a distinct, yellow macula close to posterior margin on each side; surface densely clad with fulvous, dark brown, and yellow setae in dorsal and lateral portion (
<figureCitation id="34872A5FFFCD0A5EE346FD49FCF106BA" box="[776,854,727,751]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1386,200,1899]" captionTargetId="figure-18@2.[176,1412,180,1919]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Macronotop flavomedius Pham &amp; Pham, new species. a, male dorsal view; b, male ventral view; c, female dorsal view; d, female ventral view. (Figs 1c, d were taken by Dr. Hao Xu)." figureDoi="http://doi.org/10.5281/zenodo.5515925" httpUri="https://zenodo.org/record/5515925/files/figure.png" pageId="1" pageNumber="594">Fig. 1b</figureCitation>
).
<emphasis id="9EC8EAC8FFCD0A5EE324FD46FC3806A5" bold="true" box="[874,927,728,752]" pageId="1" pageNumber="594">Legs</emphasis>
: Slender, simple. Metacoxa dark brown; almost glabrous. Femora and tibia reddish-brown; clad with long, light yellow setae. Protibia without tiny teeth (
<figureCitation id="34872A5FFFCD0A5EE54AFD65FAE80746" box="[1284,1359,763,787]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1435,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi="http://doi.org/10.5281/zenodo.5515929" httpUri="https://zenodo.org/record/5515929/files/figure.png" pageId="1" pageNumber="594">Fig. 3f</figureCitation>
); spine near the middle of outer margin of mesotibia sharp, but absent in metatibia (
<figureCitation id="34872A5FFFCD0A5EE3AEFC81FB96076D" box="[992,1073,799,824]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1435,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi="http://doi.org/10.5281/zenodo.5515929" httpUri="https://zenodo.org/record/5515929/files/figure.png" pageId="1" pageNumber="594">
Fig.
<quantity id="6B449B3FFFCD0A5EE45AFCBEFB96076D" box="[1044,1073,800,824]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="1" pageNumber="594" unit="g" value="3.0">3g</quantity>
</figureCitation>
); metatibia with a row of sparse, light yellow setae along inner margin; dorsal tooth of metatibia longer than basitarsus of metatarsus; outer tooth absent. Tarsi dark brown; basitarsus of metatarsus with a cluster of sparse, long setae.
<emphasis id="9EC8EAC8FFCD0A5EE395FCF9FBEA072A" bold="true" box="[987,1101,871,895]" pageId="1" pageNumber="594">Genitalia:</emphasis>
Parameres moderate broad and long; apex transverse. Interparameral split narow, gradually constricted to ¾ lenght of parameres; base with membrane, distinctly divided in medial (
<figureCitation id="34872A5FFFCD0A5EE184FC31FD8A0792" box="[458,557,943,967]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[186,1400,195,1919]" captionTargetId="figure-17@3.[186,1402,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." figureDoi="http://doi.org/10.5281/zenodo.5515927" httpUri="https://zenodo.org/record/5515927/files/figure.png" pageId="1" pageNumber="594">Fig. 2a,b</figureCitation>
).
</paragraph>
<paragraph id="AC0336DAFFCD0A5EE089FC4AFB9D00CA" blockId="1.[151,1437,151,1256]" pageId="1" pageNumber="594">
<emphasis id="9EC8EAC8FFCD0A5EE089FC4AFDB807BE" bold="true" box="[199,543,979,1004]" inLineHeading="true" pageId="1" pageNumber="594" reason="1">
Female (
<figureCitation id="34872A5FFFCD0A5EE167FC4DFE3407BE" box="[297,403,979,1003]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1386,200,1899]" captionTargetId="figure-18@2.[176,1412,180,1919]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Macronotop flavomedius Pham &amp; Pham, new species. a, male dorsal view; b, male ventral view; c, female dorsal view; d, female ventral view. (Figs 1c, d were taken by Dr. Hao Xu)." figureDoi="http://doi.org/10.5281/zenodo.5515925" httpUri="https://zenodo.org/record/5515925/files/figure.png" pageId="1" pageNumber="594">Figs 1c,d</figureCitation>
;
<figureCitation id="34872A5FFFCD0A5EE1ECFC4DFE4107BE" box="[418,486,979,1003]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[186,1400,195,1919]" captionTargetId="figure-17@3.[186,1402,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." figureDoi="http://doi.org/10.5281/zenodo.5515927" httpUri="https://zenodo.org/record/5515927/files/figure.png" pageId="1" pageNumber="594">2d,f,h</figureCitation>
;
<figureCitation id="34872A5FFFCD0A5EE1BAFC4DFDB507BE" box="[500,530,979,1003]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1435,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi="http://doi.org/10.5281/zenodo.5515929" httpUri="https://zenodo.org/record/5515929/files/figure.png" pageId="1" pageNumber="594">3b</figureCitation>
).
</emphasis>
Langer than the male; body length 15.3–16.0 mm; width 6.8–7.0 mm. Body color darker. Surface of head with more dense, setiferous punctures; with long, sparse fulvous and yellowish setae; midline of frons convex, glabrous. Pronotum with a longitudinal yellow tomentous maculae, but smaller and shorter than the male (
<figureCitation id="34872A5FFFCD0A5EE0D1FBA1FF540002" box="[159,243,1087,1111]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[186,1400,195,1919]" captionTargetId="figure-17@3.[186,1402,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." figureDoi="http://doi.org/10.5281/zenodo.5515927" httpUri="https://zenodo.org/record/5515927/files/figure.png" pageId="1" pageNumber="594">Fig. 2h</figureCitation>
). Clypeus, scutellum, mesepimeron, sternites II–VI, and pygidium without yellow tomentous maculae (
<figureCitation id="34872A5FFFCD0A5EE53FFBA1FF62002E" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[186,1400,195,1919]" captionTargetId="figure-17@3.[186,1402,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." figureDoi="http://doi.org/10.5281/zenodo.5515927" httpUri="https://zenodo.org/record/5515927/files/figure.png" pageId="1" pageNumber="594">Fig. 2f,h</figureCitation>
). Posthumeral macula, lateral macula, and median macula on elytron smaller than male (
<figureCitation id="34872A5FFFCD0A5EE43FFBFDFB6B002E" box="[1137,1228,1123,1147]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1386,200,1899]" captionTargetId="figure-18@2.[176,1412,180,1919]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Macronotop flavomedius Pham &amp; Pham, new species. a, male dorsal view; b, male ventral view; c, female dorsal view; d, female ventral view. (Figs 1c, d were taken by Dr. Hao Xu)." figureDoi="http://doi.org/10.5281/zenodo.5515925" httpUri="https://zenodo.org/record/5515925/files/figure.png" pageId="1" pageNumber="594">Fig 1c,d</figureCitation>
;
<figureCitation id="34872A5FFFCD0A5EE499FBFDFB52002E" box="[1239,1269,1123,1147]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1435,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi="http://doi.org/10.5281/zenodo.5515929" httpUri="https://zenodo.org/record/5515929/files/figure.png" pageId="1" pageNumber="594">3d</figureCitation>
). Antennal club shorter than the male, nearly 1.5 times length of antennomeres 2–7 combined (
<figureCitation id="34872A5FFFCD0A5EE390FB19FB8A00CA" box="[990,1069,1159,1183]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1435,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Macronotop flavomedius Pham &amp; Pham, new species. a, male antenna; b, female antenna; c, male median maculae; d, female median maculae; e, male meso- and metasternum; f, male protibia; g, male metatibia." figureDoi="http://doi.org/10.5281/zenodo.5515929" httpUri="https://zenodo.org/record/5515929/files/figure.png" pageId="1" pageNumber="594">Fig. 3b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E4A66551FFCD0A5EE089FB35FD3300BD" pageId="1" pageNumber="594" type="diagnosis">
<paragraph id="AC0336DAFFCD0A5EE089FB35FD3300BD" blockId="1.[151,1437,151,1256]" pageId="1" pageNumber="594">
<emphasis id="9EC8EAC8FFCD0A5EE089FB35FE660096" bold="true" box="[199,449,1195,1219]" inLineHeading="true" pageId="1" pageNumber="594" reason="1">Differential diagnosis.</emphasis>
Male of
<taxonomicName id="6BBC4D59FFCD0A5EE251FB32FC940096" authority="Pham &amp; Pham, 2021" authorityName="Pham &amp; Pham" authorityYear="2021" box="[543,819,1195,1219]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="flavomedius" status="sp. nov.">
<emphasis id="9EC8EAC8FFCD0A5EE251FB32FC940096" box="[543,819,1195,1219]" italics="true" pageId="1" pageNumber="594">Macronotops flavomedius</emphasis>
</taxonomicName>
<emphasis id="9EC8EAC8FFCD0A5EE377FB32FC1C0096" bold="true" box="[825,955,1195,1220]" pageId="1" pageNumber="594">
<taxonomicNameLabel id="85FB57B3FFCD0A5EE377FB32FC1C0096" box="[825,955,1195,1220]" pageId="1" pageNumber="594" rank="species">new species</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="6BBC4D59FFCD0A5EE40DFB32FB170096" baseAuthorityName="Bourgoin" baseAuthorityYear="1916" box="[1091,1200,1195,1219]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="9EC8EAC8FFCD0A5EE40DFB32FB170096" box="[1091,1200,1195,1219]" italics="true" pageId="1" pageNumber="594">M. vuilleti</emphasis>
</taxonomicName>
and
<taxonomicName id="6BBC4D59FFCD0A5EE4ACFB32FACE0096" authorityName="Qui, Xu &amp; Chen" authorityYear="2019" box="[1250,1385,1195,1219]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="dianensis">
<emphasis id="9EC8EAC8FFCD0A5EE4ACFB32FACE0096" box="[1250,1385,1195,1219]" italics="true" pageId="1" pageNumber="594">M. dianensis</emphasis>
</taxonomicName>
, and their differences are summarized in the
<tableCitation id="E13E0361FFCD0A5EE270FB51FD2A00BD" box="[574,653,1231,1256]" captionStart="TABLE 1" captionStartId="1.[151,239,1303,1327]" captionText="TABLE 1. The differences between Macronotops flavomedius new species, M. vuilleti, and M. dianensis." pageId="1" pageNumber="594">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="F8C36652FFCD0A5EE0D9FA89FAA7017A" box="[151,1280,1303,1328]" pageId="1" pageNumber="594" startId="1.[151,239,1303,1327]">
<paragraph id="AC0336DAFFCD0A5EE0D9FA89FAA7017A" blockId="1.[151,1280,1303,1328]" box="[151,1280,1303,1328]" pageId="1" pageNumber="594">
<emphasis id="9EC8EAC8FFCD0A5EE0D9FA89FEAD017A" bold="true" box="[151,266,1303,1327]" pageId="1" pageNumber="594">TABLE 1.</emphasis>
The differences between
<taxonomicName id="6BBC4D59FFCD0A5EE253FA86FC95017A" authority="Pham &amp; Pham, 2021" authorityName="Pham &amp; Pham" authorityYear="2021" box="[541,818,1303,1327]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="flavomedius" status="sp. nov.">
<emphasis id="9EC8EAC8FFCD0A5EE253FA86FC95017A" box="[541,818,1303,1327]" italics="true" pageId="1" pageNumber="594">Macronotops flavomedius</emphasis>
</taxonomicName>
<emphasis id="9EC8EAC8FFCD0A5EE376FA86FC1B017A" bold="true" box="[824,956,1303,1328]" pageId="1" pageNumber="594">
<taxonomicNameLabel id="85FB57B3FFCD0A5EE376FA86FC1B017A" box="[824,956,1303,1328]" pageId="1" pageNumber="594" rank="species">new species</taxonomicNameLabel>
</emphasis>
,
<emphasis id="9EC8EAC8FFCD0A5EE387FA86FB9A017A" box="[969,1085,1303,1327]" italics="true" pageId="1" pageNumber="594">
<taxonomicName id="6BBC4D59FFCD0A5EE387FA86FB9E017A" baseAuthorityName="Bourgoin" baseAuthorityYear="1916" box="[969,1081,1303,1327]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="vuilleti">M. vuilleti</taxonomicName>
,
</emphasis>
and
<taxonomicName id="6BBC4D59FFCD0A5EE43FFA86FB5B017A" authorityName="Qui, Xu &amp; Chen" authorityYear="2019" box="[1137,1276,1303,1327]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="dianensis">M. dianensis</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="AC0336DAFFCD0A5EE0E8FADEFB6A02E4" pageId="1" pageNumber="594">
<table id="DEBCC47AFFCDF5A0E0E8FADEFA2C02E4" box="[166,1419,1344,1713]" gridcols="4" gridrows="8" pageId="1" pageNumber="594">
<tr id="128C3498FFCDF5A0E0E8FADEFA2C0103" box="[166,1419,1344,1366]" gridrow="0" pageId="1" pageNumber="594">
<th id="515D5DE4FFCDF5A0E0E8FADEFE6F0103" box="[166,456,1344,1366]" gridcol="0" gridrow="0" pageId="1" pageNumber="594">
<emphasis id="9EC8EAC8FFCD0A5EE0E8FADEFEBE0100" bold="true" box="[166,281,1344,1365]" pageId="1" pageNumber="594">Characters</emphasis>
</th>
<th id="515D5DE4FFCDF5A0E1A7FADEFD580103" box="[489,767,1344,1366]" gridcol="1" gridrow="0" pageId="1" pageNumber="594">
<emphasis id="9EC8EAC8FFCD0A5EE1A7FADFFD580100" bold="true" box="[489,767,1344,1366]" pageId="1" pageNumber="594">
<taxonomicName id="6BBC4D59FFCD0A5EE1A7FADFFD250103" authority="Pham &amp; Pham, 2021" authorityName="Pham &amp; Pham" authorityYear="2021" box="[489,642,1344,1366]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="flavomedius" status="sp. nov.">
<emphasis id="9EC8EAC8FFCD0A5EE1A7FADFFD250103" bold="true" box="[489,642,1344,1366]" italics="true" pageId="1" pageNumber="594">M. flavomedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="85FB57B3FFCD0A5EE2C6FADEFD580100" box="[648,767,1344,1365]" pageId="1" pageNumber="594" rank="species">new species</taxonomicNameLabel>
</emphasis>
</th>
<th id="515D5DE4FFCDF5A0E360FADEFBE00103" box="[814,1095,1344,1366]" gridcol="2" gridrow="0" pageId="1" pageNumber="594">
<taxonomicName id="6BBC4D59FFCD0A5EE360FADFFC340103" baseAuthorityName="Bourgoin" baseAuthorityYear="1916" box="[814,915,1344,1366]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="9EC8EAC8FFCD0A5EE360FADFFC340103" bold="true" box="[814,915,1344,1366]" italics="true" pageId="1" pageNumber="594">M. vuilleti</emphasis>
</taxonomicName>
</th>
<th id="515D5DE4FFCDF5A0E43CFADEFA2C0103" box="[1138,1419,1344,1366]" gridcol="3" gridrow="0" pageId="1" pageNumber="594">
<taxonomicName id="6BBC4D59FFCD0A5EE43CFADFFB570103" authorityName="Qui, Xu &amp; Chen" authorityYear="2019" box="[1138,1264,1344,1366]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="dianensis">
<emphasis id="9EC8EAC8FFCD0A5EE43CFADFFB570103" bold="true" box="[1138,1264,1344,1366]" italics="true" pageId="1" pageNumber="594">M. dianensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="128C3498FFCDF5A0E0E8FAF6FA2C012B" box="[166,1419,1384,1406]" gridrow="1" pageId="1" pageNumber="594">
<th id="515D5DE4FFCDF5A0E0E8FAF6FE6F012B" box="[166,456,1384,1406]" gridcol="0" gridrow="1" pageId="1" pageNumber="594">Anterior margin of clypeus</th>
<td id="515D5DE4FFCDF5A0E1A7FAF6FD58012B" box="[489,767,1384,1406]" gridcol="1" gridrow="1" pageId="1" pageNumber="594">Slightly notched</td>
<td id="515D5DE4FFCDF5A0E360FAF6FBE0012B" box="[814,1095,1384,1406]" gridcol="2" gridrow="1" pageId="1" pageNumber="594">Nearly straight</td>
<td id="515D5DE4FFCDF5A0E43CFAF6FA2C012B" box="[1138,1419,1384,1406]" gridcol="3" gridrow="1" pageId="1" pageNumber="594">Nearly straight</td>
</tr>
<tr id="128C3498FFCDF5A0E0E8FA11FA2C019C" box="[166,1419,1423,1481]" gridrow="2" pageId="1" pageNumber="594" rowspan-0="1">
<td id="515D5DE4FFCDF5A0E1A7FA11FD58019C" box="[489,767,1423,1481]" gridcol="1" gridrow="2" pageId="1" pageNumber="594">Length greater than width (2/1.5)</td>
<td id="515D5DE4FFCDF5A0E360FA11FBE0019C" box="[814,1095,1423,1481]" gridcol="2" gridrow="2" pageId="1" pageNumber="594">Length almost equal to width</td>
<td id="515D5DE4FFCDF5A0E43CFA11FA2C019C" box="[1138,1419,1423,1481]" gridcol="3" gridrow="2" pageId="1" pageNumber="594">Length almost equal to width</td>
</tr>
<tr id="128C3498FFCDF5A0E0E8FA44FA2C01A5" box="[166,1419,1498,1520]" gridrow="3" pageId="1" pageNumber="594">
<th id="515D5DE4FFCDF5A0E0E8FA44FE6F01A5" box="[166,456,1498,1520]" gridcol="0" gridrow="3" pageId="1" pageNumber="594">Protibia</th>
<td id="515D5DE4FFCDF5A0E1A7FA44FD5801A5" box="[489,767,1498,1520]" gridcol="1" gridrow="3" pageId="1" pageNumber="594">Without tiny teeth</td>
<td id="515D5DE4FFCDF5A0E360FA44FBE001A5" box="[814,1095,1498,1520]" gridcol="2" gridrow="3" pageId="1" pageNumber="594">Without tiny teeth</td>
<td id="515D5DE4FFCDF5A0E43CFA44FA2C01A5" box="[1138,1419,1498,1520]" gridcol="3" gridrow="3" pageId="1" pageNumber="594">Tiny teeth indistinct</td>
</tr>
<tr id="128C3498FFCDF5A0E0E8F99FFA2C026E" box="[166,1419,1537,1595]" gridrow="4" pageId="1" pageNumber="594">
<th id="515D5DE4FFCDF5A0E0E8F99FFE6F026E" box="[166,456,1537,1595]" gridcol="0" gridrow="4" pageId="1" pageNumber="594">The middle of outer margin of metatibia</th>
<td id="515D5DE4FFCDF5A0E1A7F99FFD58026E" box="[489,767,1537,1595]" gridcol="1" gridrow="4" pageId="1" pageNumber="594">Without spine</td>
<td id="515D5DE4FFCDF5A0E360F99FFBE0026E" box="[814,1095,1537,1595]" gridcol="2" gridrow="4" pageId="1" pageNumber="594">With a sharp spine</td>
<td id="515D5DE4FFCDF5A0E43CF99FFA2C026E" box="[1138,1419,1537,1595]" gridcol="3" gridrow="4" pageId="1" pageNumber="594">With a blunt spine</td>
</tr>
<tr id="128C3498FFCDF5A0E0E8F9D3FA2C0236" box="[166,1419,1613,1635]" gridrow="5" pageId="1" pageNumber="594">
<th id="515D5DE4FFCDF5A0E0E8F9D3FE6F0236" box="[166,456,1613,1635]" gridcol="0" gridrow="5" pageId="1" pageNumber="594">Yellow macula on pronotum</th>
<td id="515D5DE4FFCDF5A0E1A7F9D3FD580236" box="[489,767,1613,1635]" gridcol="1" gridrow="5" pageId="1" pageNumber="594">Present</td>
<td id="515D5DE4FFCDF5A0E360F9D3FBE00236" box="[814,1095,1613,1635]" gridcol="2" gridrow="5" pageId="1" pageNumber="594">Absent</td>
<td id="515D5DE4FFCDF5A0E43CF9D3FA2C0236" box="[1138,1419,1613,1635]" gridcol="3" gridrow="5" pageId="1" pageNumber="594">Absent</td>
</tr>
<tr id="128C3498FFCDF5A0E0E8F9EAFA2C02DF" box="[166,1419,1652,1674]" gridrow="6" pageId="1" pageNumber="594">
<th id="515D5DE4FFCDF5A0E0E8F9EAFE6F02DF" box="[166,456,1652,1674]" gridcol="0" gridrow="6" pageId="1" pageNumber="594">Yellow macula on pygidium</th>
<td id="515D5DE4FFCDF5A0E1A7F9EAFD5802DF" box="[489,767,1652,1674]" gridcol="1" gridrow="6" pageId="1" pageNumber="594">Present</td>
<td id="515D5DE4FFCDF5A0E360F9EAFBE002DF" box="[814,1095,1652,1674]" gridcol="2" gridrow="6" pageId="1" pageNumber="594">Present/Absent</td>
<td id="515D5DE4FFCDF5A0E43CF9EAFA2C02DF" box="[1138,1419,1652,1674]" gridcol="3" gridrow="6" pageId="1" pageNumber="594">Absent</td>
</tr>
<tr id="128C3498FFCDF5A0E0E8F905FA2C02E4" box="[166,1419,1691,1713]" gridrow="7" pageId="1" pageNumber="594">
<th id="515D5DE4FFCDF5A0E0E8F905FE6F02E4" box="[166,456,1691,1713]" gridcol="0" gridrow="7" pageId="1" pageNumber="594">Longitudinal ridge on frons</th>
<td id="515D5DE4FFCDF5A0E1A7F905FD5802E4" box="[489,767,1691,1713]" gridcol="1" gridrow="7" pageId="1" pageNumber="594">Morderately raised</td>
<td id="515D5DE4FFCDF5A0E360F905FBE002E4" box="[814,1095,1691,1713]" gridcol="2" gridrow="7" pageId="1" pageNumber="594">Absent or indistinct</td>
<td id="515D5DE4FFCDF5A0E43CF905FA2C02E4" box="[1138,1419,1691,1713]" gridcol="3" gridrow="7" pageId="1" pageNumber="594">Indistinct</td>
</tr>
</table>
</paragraph>
<subSubSection id="E4A66551FFCD0A5EE089F97DFB4902AE" box="[199,1262,1763,1787]" pageId="1" pageNumber="594" type="etymology">
<paragraph id="AC0336DAFFCD0A5EE089F97DFB4902AE" blockId="1.[151,1436,1763,1859]" box="[199,1262,1763,1787]" pageId="1" pageNumber="594">
<emphasis id="9EC8EAC8FFCD0A5EE089F97DFEEF02AE" bold="true" box="[199,328,1763,1787]" inLineHeading="true" pageId="1" pageNumber="594" reason="1">Etymology.</emphasis>
The new species is named after its yellow macula on the middle of pronotum (
<figureCitation id="34872A5FFFCD0A5EE4DFF97DFB4602AE" box="[1169,1249,1763,1787]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[186,1400,195,1919]" captionTargetId="figure-17@3.[186,1402,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Macronotop flavomedius Pham &amp; Pham, new species. a, parameres, dorsal view; b, parameres, lateral view; c, male distal macula and pygidium; d, female distal macula and pygidium; e, male clypeus; f, female clypeus; g, male pronotum; h, female pronotum." figureDoi="http://doi.org/10.5281/zenodo.5515927" httpUri="https://zenodo.org/record/5515927/files/figure.png" pageId="1" pageNumber="594">
Fig.
<quantity id="6B449B3FFFCD0A5EE48DF97DFB4602AE" box="[1219,1249,1763,1787]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="1" pageNumber="594" unit="g" value="2.0">2g</quantity>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E4A66551FFCD0A5EE089F899FE120316" pageId="1" pageNumber="594" type="distribution">
<paragraph id="AC0336DAFFCD0A5EE089F899FE120316" blockId="1.[151,1436,1763,1859]" pageId="1" pageNumber="594">
<emphasis id="9EC8EAC8FFCD0A5EE089F899FEFD034A" bold="true" box="[199,346,1799,1823]" inLineHeading="true" pageId="1" pageNumber="594" reason="1">Distribution.</emphasis>
<collectingCountry id="D4AB764AFFCD0A5EE12FF899FE1C034A" box="[353,443,1799,1823]" name="Vietnam" pageId="1" pageNumber="594">Vietnam</collectingCountry>
(
<collectingRegion id="6E78F838FFCD0A5EE185F899FD0A034A" box="[459,685,1799,1823]" country="Vietnam" name="Lam Dong" pageId="1" pageNumber="594">Lam Dong Province</collectingRegion>
).
<taxonomicName id="6BBC4D59FFCD0A5EE28EF896FC71034A" authority="Pham &amp; Pham, 2021" authorityName="Pham &amp; Pham" authorityYear="2021" box="[704,982,1799,1823]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="flavomedius" status="sp. nov.">
<emphasis id="9EC8EAC8FFCD0A5EE28EF896FC71034A" box="[704,982,1799,1823]" italics="true" pageId="1" pageNumber="594">Macronotops flavomedius</emphasis>
</taxonomicName>
<emphasis id="9EC8EAC8FFCD0A5EE390F899FBC3034A" bold="true" box="[990,1124,1799,1823]" pageId="1" pageNumber="594">
<taxonomicNameLabel id="85FB57B3FFCD0A5EE390F899FBC3034A" box="[990,1124,1799,1823]" pageId="1" pageNumber="594" rank="species">new species</taxonomicNameLabel>
</emphasis>
is the southern-most species of the genus
<emphasis id="9EC8EAC8FFCD0A5EE16EF8B2FE120316" box="[288,437,1836,1859]" italics="true" pageId="1" pageNumber="594">
<taxonomicName id="6BBC4D59FFCD0A5EE16EF8B2FE160316" authorityName="Krikken" authorityYear="1977" box="[288,433,1836,1859]" class="Insecta" family="Cetoniidae" genus="Macronotops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus">Macronotops</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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