423 lines
31 KiB
XML
423 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1024.56927" ID-GBIF-Dataset="7a0ded71-ddb1-49e5-9c2c-10c01fd03a1f" ID-PMC="PMC7990858" ID-Pensoft-Pub="1313-2970-1024-157" ID-Pensoft-UUID="086BAB631B8D5B888571137F0CEFF686" ID-PubMed="33786006" ID-ZooBank="FDD37EA991214385B67D51AD313CB49E" ModsDocID="1313-2970-1024-157" checkinTime="1615979483707" checkinUser="pensoft" docAuthor="Nestor-Arriola, Jorge Ismael, Toledo-Hernandez, Victor Hugo, Solis, Angel, Gonzalez, Guillermo & Vetrovec, Jaroslav" docDate="2021" docId="24119BE8686A536E818D44DC8BEFDAC9" docLanguage="en" docName="ZooKeys 1024: 157-196" docOrigin="ZooKeys 1024" docPubDate="2021-03-16" docSource="http://dx.doi.org/10.3897/zookeys.1024.56927" docTitle="Brachiacantha gorhami Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec 2021, comb. nov." docType="treatment" docVersion="5" id="086BAB631B8D5B888571137F0CEFF686" lastPageNumber="157" masterDocId="086BAB631B8D5B888571137F0CEFF686" masterDocTitle="The Brachiacantha Dejean, 1837 (Coleoptera, Coccinellidae) of Central America" masterLastPageNumber="196" masterPageNumber="157" pageNumber="157" updateTime="1668149905347" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Brachiacantha Dejean, 1837 (Coleoptera, Coccinellidae) of Central America</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nestor-Arriola, Jorge Ismael</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2394-1586</mods:nameIdentifier>
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<mods:affiliation>Centro de Investigacion en Biodiversidad y Conservacion (CIByC), Universidad Autonoma del Estado de Morelos. Av. Universidad # 1001, Col. Chamilpa, Cuernavaca, Morelos, C. P. 62209, Mexico</mods:affiliation>
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<mods:nameIdentifier type="email">jorge.nestorarr@uaem.edu.mx</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Toledo-Hernandez, Victor Hugo</mods:namePart>
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<mods:affiliation>Centro de Investigacion en Biodiversidad y Conservacion (CIByC), Universidad Autonoma del Estado de Morelos. Av. Universidad # 1001, Col. Chamilpa, Cuernavaca, Morelos, C. P. 62209, Mexico</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Solis, Angel</mods:namePart>
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<mods:affiliation>Departamento de Historia Natural, Museo Nacional de Costa Rica. Santo Domingo de Heredia, Costa Rica</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gonzalez, Guillermo</mods:namePart>
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<mods:affiliation>La Reina, Santiago, Chile</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vetrovec, Jaroslav</mods:namePart>
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<mods:affiliation>Buzulucka, Hradec Kralove, Ceska Republika</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-03-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1024</mods:number>
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</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>157</mods:start>
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<mods:end>196</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1024.56927</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1024.56927</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1024-157</mods:identifier>
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<mods:identifier type="ZooBank">FDD37EA991214385B67D51AD313CB49E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">086BAB631B8D5B888571137F0CEFF686</mods:identifier>
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||
</mods:mods>
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<subSection pageId="0" pageNumber="157" type="results">
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||
<treatment ID-GBIF-Taxon="180585862" LSID="urn:lsid:plazi:treatment:24119BE8686A536E818D44DC8BEFDAC9" httpUri="http://treatment.plazi.org/id/24119BE8686A536E818D44DC8BEFDAC9" lastPageNumber="157" pageId="0" pageNumber="157">
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<subSubSection pageId="0" pageNumber="157" type="nomenclature">
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<paragraph pageId="0" pageNumber="157">
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<taxonomicName LSID="24119BE8-686A-536E-818D-44DC8BEFDAC9" authority="(Weise)" authorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" authorityYear="2021" baseAuthorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" baseAuthorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha gorhami" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="gorhami" status="comb. nov.">Brachiacantha gorhami (Weise)</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="157">comb. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 9–12" captionStartId="F3" captionText="Figures 9 - 12. Brachiacantha sp. 9 Brachiacantha guatemalensis comb. nov., male 10 Brachiacantha papiliona sp. nov., female 11 Brachiacantha mimica sp. nov., female 12 Brachiacantha gorhami comb. nov., female." figureDoi="10.3897/zookeys.1024.56927.figures9-12" httpUri="https://binary.pensoft.net/fig/520868" pageId="0" pageNumber="157">Figure 12</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="157" type="reference_group">
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<paragraph pageId="0" pageNumber="157">
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<taxonomicName authorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" authorityYear="2021" baseAuthorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" baseAuthorityYear="2021" class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella pantherina" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="pantherina" status="comb. nov.">Coccinella pantherina</taxonomicName>
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Gorham, 1892: 161, tab. IX, fig. 9 (non
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<taxonomicName authorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" authorityYear="2021" baseAuthorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" baseAuthorityYear="2021" class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella pantherina" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="pantherina" status="comb. nov.">Coccinella pantherina</taxonomicName>
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Linnaeus, 1758: 368) (non
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<taxonomicName authorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" authorityYear="2021" baseAuthorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" baseAuthorityYear="2021" class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella pantherina" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="pantherina" status="comb. nov.">Coccinella pantherina</taxonomicName>
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DeGeer, 1773).
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</paragraph>
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<paragraph pageId="0" pageNumber="157">
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<taxonomicName authorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" authorityYear="2021" baseAuthorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" baseAuthorityYear="2021" class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella gorhami" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="gorhami" status="comb. nov.">Coccinella gorhami</taxonomicName>
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Weise, 1904: 357.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="materials_examined">
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<paragraph pageId="0" pageNumber="157">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="157">
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<materialsCitation ID-GBIF-Occurrence="3063616335" collectingDate="1971-05-10" collectorName="H. F. Howden" country="El Salvador" elevation="2300" location="Montecristo" specimenCount="1" specimenCount-female="1">
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<collectingCountry name="El Salvador">El Salvador</collectingCountry>
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•
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<specimenCount type="female">1♀</specimenCount>
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;
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<location LSID="urn:lsid:plazi:treatment:24119BE8686A536E818D44DC8BEFDAC9:9B9E275194BF369722F4A3D0A0A9C662" country="El Salvador" name="Montecristo">Montecristo</location>
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,
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<locationDeviation location="Metapan">
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<quantity metricMagnitude="4" metricUnit="m" metricValue="2.3" unit="km" value="23.0">23 km</quantity>
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N of Metapan
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</locationDeviation>
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;
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<quantity metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">2300 m</elevation>
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</quantity>
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;
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<date value="1971-05-10">
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<collectingDate value="1971-05-10">10 May 1971</collectingDate>
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</date>
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;
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<collectorName>H. F. Howden</collectorName>
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leg. (USNMNH)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="diagnosis">
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<paragraph pageId="0" pageNumber="157">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="157">
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Dorsal color pale yellow except head with brown clypeus and black above the eyes; pronotum with a black macula on the posterior margin and two large convergent subtriangular spots at center; scutellar shield black; each elytron with five black marks: a vitta on the elytral suture extending laterally in the second half of the elytron, an oval spot at the humeral callus, an elongated spot parallel to the sutural margin in the basal half of the elytron, an oval spot at middle and a small spot on the lateral margin in the apical half of the elytron (Fig.
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<figureCitation captionStart="Figures 9–12" captionStartId="F3" captionText="Figures 9 - 12. Brachiacantha sp. 9 Brachiacantha guatemalensis comb. nov., male 10 Brachiacantha papiliona sp. nov., female 11 Brachiacantha mimica sp. nov., female 12 Brachiacantha gorhami comb. nov., female." figureDoi="10.3897/zookeys.1024.56927.figures9-12" httpUri="https://binary.pensoft.net/fig/520868" pageId="0" pageNumber="157">12</figureCitation>
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). Ventral surface brown. Protibial tooth small.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="variation">
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<paragraph pageId="0" pageNumber="157">Variation.</paragraph>
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<paragraph pageId="0" pageNumber="157">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="distribution">
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<paragraph pageId="0" pageNumber="157">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="157">
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<collectingCountry name="El Salvador">El Salvador</collectingCountry>
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and
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<collectingCountry name="Guatemala">Guatemala</collectingCountry>
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. New record to
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<collectingCountry name="El Salvador">El Salvador</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="157" type="discussion">
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<paragraph pageId="0" pageNumber="157">Discussion.</paragraph>
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<paragraph pageId="0" pageNumber="157">
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The material was identified using the descriptions and tables of Gorham. The species was described by Gorham as
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<taxonomicName authorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" authorityYear="2021" baseAuthorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" baseAuthorityYear="2021" class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella pantherina" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="pantherina">
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<emphasis italics="true" pageId="0" pageNumber="157">Coccinella pantherina</emphasis>
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</taxonomicName>
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, but the name was preoccupied by a junior synonym of
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<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Coccinellidae" genus="Adalia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Adalia bipunctata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="bipunctata">
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<emphasis italics="true" pageId="0" pageNumber="157">Adalia bipunctata</emphasis>
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</taxonomicName>
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(Linnaeus, 1758) and a junior synonym of
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<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Coccinellidae" genus="Adalia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Adalia decempunctata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="decempunctata">
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<emphasis italics="true" pageId="0" pageNumber="157">Adalia decempunctata</emphasis>
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</taxonomicName>
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(DeGeer, 1773).
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<bibRefCitation DOI="https://doi.org/10.1002/mmnd.48019030205" author="Weise, J" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="157" refId="B34" refString="Weise, J, 1903. Drei neue Coccinellen. Deutsche Entomologische Zeitschrift 207-208. https://doi.org/10.1002/mmnd.48019030205" title="Drei neue Coccinellen. Deutsche Entomologische Zeitschrift 207 - 208." url="https://doi.org/10.1002/mmnd.48019030205" year="1903">Weise (1903)</bibRefCitation>
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noticed the homonymy and created the name
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<taxonomicName authorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" authorityYear="2021" baseAuthorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" baseAuthorityYear="2021" class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella gorhami" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="gorhami">
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<emphasis italics="true" pageId="0" pageNumber="157">Coccinella gorhami</emphasis>
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</taxonomicName>
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for the species.
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</paragraph>
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<paragraph pageId="0" pageNumber="157">
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Although Gorham placed the species as a member of the genus
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<taxonomicName class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Coccinella</emphasis>
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</taxonomicName>
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, he compared it with the species
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<taxonomicName lsidName="B. lepida" pageId="0" pageNumber="157" rank="species" species="lepida">
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<emphasis italics="true" pageId="0" pageNumber="157">B. lepida</emphasis>
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</taxonomicName>
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, thus indicating the small size of the species. In the last century, several species originally described as
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<taxonomicName class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Coccinella</emphasis>
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</taxonomicName>
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have been revised and assigned to other genera such as
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<taxonomicName class="Insecta" family="Coccinellidae" genus="Mulsantina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Mulsantina" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Mulsantina</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Crotch" authorityYear="1871" class="Insecta" family="Coccinellidae" genus="Cycloneda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Cycloneda" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Cycloneda</emphasis>
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</taxonomicName>
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, but there are still several species which need revision. The specimen examined corresponded to the description and figures of Gorham (1892), but the presence of eye emarginated by the eye canthus, the lack of large abdominal primary pores between fourth and fifth ventrites and the presence of a small tibial tooth identified the species as a member of
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<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha</emphasis>
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</taxonomicName>
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. The dorsal coloration is unique among
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<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha</emphasis>
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</taxonomicName>
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species but is similar to those of other genera like
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<taxonomicName authorityName="Duverger" authorityYear="2001" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dilatitibialis" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Dilatitibialis</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Gordon & Canepari" authorityYear="2013" class="Insecta" family="Coccinellidae" genus="Serratitibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Serratitibia" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Serratitibia</emphasis>
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</taxonomicName>
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, and
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<taxonomicName class="Insecta" family="Coccinellidae" genus="Cyrea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrea" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="157">Cyrea</emphasis>
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</taxonomicName>
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. A revision of the original material may be needed to confirm the taxonomic changes.
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</paragraph>
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<caption doi="10.3897/zookeys.1024.56927.figures21-24" httpUri="https://binary.pensoft.net/fig/520870" pageId="0" pageNumber="157" start="Figures 21–24" startId="F5">
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<paragraph pageId="0" pageNumber="157">
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<emphasis bold="true" pageId="0" pageNumber="157">Figures 21-24.</emphasis>
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<taxonomicName authorityName="Nestor-Arriola, Toledo-Hernandez & Solis" authorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha nubes" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="nubes">
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<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha nubes</emphasis>
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</taxonomicName>
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sp. nov., male
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<emphasis bold="true" pageId="0" pageNumber="157">21</emphasis>
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Dorsal view
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<emphasis bold="true" pageId="0" pageNumber="157">22</emphasis>
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Abdomen
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<emphasis bold="true" pageId="0" pageNumber="157">23</emphasis>
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Tegmen, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">24</emphasis>
|
||
Tegmen, concave side view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures25-28" httpUri="https://binary.pensoft.net/fig/520871" pageId="0" pageNumber="157" start="Figures 25–28" startId="F6">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 25-28.</emphasis>
|
||
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha fenestrata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="fenestrata">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha fenestrata</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia
|
||
<emphasis bold="true" pageId="0" pageNumber="157">25</emphasis>
|
||
Tegmen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">26</emphasis>
|
||
Tegmen lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">27</emphasis>
|
||
Penis
|
||
<emphasis bold="true" pageId="0" pageNumber="157">28</emphasis>
|
||
Penis apex.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures29-32" httpUri="https://binary.pensoft.net/fig/520872" pageId="0" pageNumber="157" start="Figures 29–32" startId="F7">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 29-32.</emphasis>
|
||
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha octostigma" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="octostigma">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha octostigma</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia
|
||
<emphasis bold="true" pageId="0" pageNumber="157">29</emphasis>
|
||
Tegmen, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">30</emphasis>
|
||
Tegmen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">31</emphasis>
|
||
Penis
|
||
<emphasis bold="true" pageId="0" pageNumber="157">32</emphasis>
|
||
Penis apex.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures33-36" httpUri="https://binary.pensoft.net/fig/520873" pageId="0" pageNumber="157" start="Figures 33–36" startId="F8">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 33-36.</emphasis>
|
||
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha bipartita" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="bipartita">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha bipartita</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia
|
||
<emphasis bold="true" pageId="0" pageNumber="157">33</emphasis>
|
||
Tegmen, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">34</emphasis>
|
||
Tegmen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">35</emphasis>
|
||
Penis
|
||
<emphasis bold="true" pageId="0" pageNumber="157">36</emphasis>
|
||
Penis apex.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures37-40" httpUri="https://binary.pensoft.net/fig/520874" pageId="0" pageNumber="157" start="Figures 37–40" startId="F9">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 37-40.</emphasis>
|
||
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha aperta" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="aperta">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha aperta</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia
|
||
<emphasis bold="true" pageId="0" pageNumber="157">37</emphasis>
|
||
Penis
|
||
<emphasis bold="true" pageId="0" pageNumber="157">38</emphasis>
|
||
Penis apex
|
||
<emphasis bold="true" pageId="0" pageNumber="157">39</emphasis>
|
||
Tegmen, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">40</emphasis>
|
||
Tegmen.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures41-45" httpUri="https://binary.pensoft.net/fig/520875" pageId="0" pageNumber="157" start="Figures 41–45" startId="F10">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 41-45.</emphasis>
|
||
<taxonomicName authorityName="Nestor-Arriola, Toledo-Hernandez & Solis" authorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha dentata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="dentata">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha dentata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male
|
||
<emphasis bold="true" pageId="0" pageNumber="157">41</emphasis>
|
||
Dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">42</emphasis>
|
||
Abdomen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">43</emphasis>
|
||
Tegmen, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">44</emphasis>
|
||
Tegmen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">45</emphasis>
|
||
Penis.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures46-51" httpUri="https://binary.pensoft.net/fig/520876" pageId="0" pageNumber="157" start="Figures 46–51" startId="F11">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 46-51.</emphasis>
|
||
<taxonomicName authorityName="Nestor-Arriola, Solis & Toledo-Hernandez" authorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha aurantiapleura" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="aurantiapleura">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha aurantiapleura</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male
|
||
<emphasis bold="true" pageId="0" pageNumber="157">46</emphasis>
|
||
Lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">47</emphasis>
|
||
Protibia
|
||
<emphasis bold="true" pageId="0" pageNumber="157">48</emphasis>
|
||
Abdomen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">49</emphasis>
|
||
Tegmen, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">50</emphasis>
|
||
Tegmen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">51</emphasis>
|
||
Penis.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures52-55" httpUri="https://binary.pensoft.net/fig/520877" pageId="0" pageNumber="157" start="Figures 52–55" startId="F12">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 52-55.</emphasis>
|
||
<taxonomicName authorityName="Nestor-Arriola, Toledo-Hernandez & Solis" authorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha isthmena" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="isthmena">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha isthmena</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male genitalia
|
||
<emphasis bold="true" pageId="0" pageNumber="157">52</emphasis>
|
||
Tegmen, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">53</emphasis>
|
||
Tegmen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">54</emphasis>
|
||
Penis
|
||
<emphasis bold="true" pageId="0" pageNumber="157">55</emphasis>
|
||
Penis apex.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures56-58" httpUri="https://binary.pensoft.net/fig/520878" pageId="0" pageNumber="157" start="Figures 56–58" startId="F13">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 56-58.</emphasis>
|
||
<taxonomicName authorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" authorityYear="2021" baseAuthorityName="Nestor-Arriola & Toledo-Hernández & Solís & González & Větrovec" baseAuthorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha guatemalensis" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="guatemalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha guatemalensis</emphasis>
|
||
</taxonomicName>
|
||
comb. nov., male
|
||
<emphasis bold="true" pageId="0" pageNumber="157">56</emphasis>
|
||
Penis
|
||
<emphasis bold="true" pageId="0" pageNumber="157">57</emphasis>
|
||
Penis apex
|
||
<emphasis bold="true" pageId="0" pageNumber="157">58</emphasis>
|
||
Tegmen.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures59-64" httpUri="https://binary.pensoft.net/fig/520879" pageId="0" pageNumber="157" start="Figures 59–64" startId="F14">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 59-64.</emphasis>
|
||
<taxonomicName authorityName="Nestor-Arriola, Toledo-Hernandez & Solis" authorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha invertita" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="invertita">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha invertita</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male
|
||
<emphasis bold="true" pageId="0" pageNumber="157">59</emphasis>
|
||
Dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">60</emphasis>
|
||
Abdomen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">61</emphasis>
|
||
Tegmen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">62</emphasis>
|
||
Tegmen lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">63</emphasis>
|
||
Penis
|
||
<emphasis bold="true" pageId="0" pageNumber="157">64</emphasis>
|
||
Penis apex.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures65-68" httpUri="https://binary.pensoft.net/fig/520880" pageId="0" pageNumber="157" start="Figures 65–68" startId="F15">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 65-68.</emphasis>
|
||
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha cachensis" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="cachensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha cachensis</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia
|
||
<emphasis bold="true" pageId="0" pageNumber="157">65</emphasis>
|
||
Tegmen
|
||
<emphasis bold="true" pageId="0" pageNumber="157">66</emphasis>
|
||
Tegmen lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">67</emphasis>
|
||
Penis
|
||
<emphasis bold="true" pageId="0" pageNumber="157">68</emphasis>
|
||
Penis apex.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1024.56927.figures69-70" httpUri="https://binary.pensoft.net/fig/520881" pageId="0" pageNumber="157" start="Figures 69, 70" startId="F16">
|
||
<paragraph pageId="0" pageNumber="157">
|
||
<emphasis bold="true" pageId="0" pageNumber="157">Figures 69, 70.</emphasis>
|
||
<taxonomicName authorityName="Nestor-Arriola, Toledo-Hernandez & Solis" authorityYear="2021" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha tica" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="tica">
|
||
<emphasis italics="true" pageId="0" pageNumber="157">Brachiacantha tica</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., female
|
||
<emphasis bold="true" pageId="0" pageNumber="157">69</emphasis>
|
||
Lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="157">70</emphasis>
|
||
Protibia.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |