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<mods:title id="318656633037F79E5CDE8D55A6131626">Redescription of Semisulcospira reticulata (Mollusca, Semisulcospiridae) with description of a new species from Lake Biwa, Japan</mods:title>
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<mods:namePart id="8DCC6EE7DD526515DBC6591F42A82182">Sawada, Naoto</mods:namePart>
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<mods:affiliation id="4AEF5EE3A55972518A6A2DE040227F5E">Department of Zoology, Graduate School of Science, Kyoto University, Kitashirakawa-oiwakecho, Sakyo-ku, Kyoto, 606 - 8502, Japan</mods:affiliation>
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<mods:namePart id="36D42DF95C7B332A9C59A6A70AEDAE09">Fuke, Yusuke</mods:namePart>
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<mods:affiliation id="6770B5102EA79B153D0DEB290CFF8704">Ecological Genetics Laboratory, National Institute of Genetics, 1111 Yata, Mishima, Shizuoka, 411 - 8540, Japan</mods:affiliation>
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<mods:namePart id="DEDF135E1A1E80A22D8314B74FF11322">Miura, Osamu</mods:namePart>
<mods:affiliation id="73597519322BBC804AAE7DFB92E31C70">Faculty of Agriculture and Marine Science, Kochi University, 200 Monobe, Nankoku, Kochi, 783 - 8502, Japan</mods:affiliation>
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<mods:namePart id="791549B0D41FA99D7CC27C0371FD5343">Toyohara, Haruhiko</mods:namePart>
<mods:affiliation id="6DB9064B097C55ADDBE6A0B37C41D80C">Faculty of Agriculture, Setsunan University, 45 - 1 Nagaotoge-cho, Hirakata, Osaka, 573 - 0101, Japan</mods:affiliation>
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<taxonomicName id="6E6FA3C3B7DDA501AADB9DE5667CE036" ID-CoL="4WHDP" authority="Kajiyama &amp; Habe, 1961" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" order="Sorbeoconcha" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="FCF713D9F423D329FCBADF6B019101B8" italics="true">Semisulcospira reticulata</emphasis>
Kajiyama &amp; Habe, 1961
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<tableCitation id="9CD81164A4C34F90AFBE6EB15119F8C2" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphometric characters of Semisulcospira reticulata and S. nishimurai sp. nov. Measurements and counts: minimum maximum value (mean ± SD)." httpUri="http://table.plazi.org/id/739C4F23DED6447AA0BA4D6A8D9BC1EC" tableUuid="739C4F23DED6447AA0BA4D6A8D9BC1EC">Table 4</tableCitation>
,
<figureCitation id="8EA123D7312AEF6B3097D321D6180E6B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells and opercula of Semisulcospira reticulata (A Y) and S. nishimurai sp. nov. (Z BF). A F. Holotype of S. reticulata from Oki-shima Island (offshore), NSMT - Mo 39770; G J. Hannoura, KUZ Z 4926, Z 4927; K O. Kitafunaki, KUZ Z 4930, Z 4931; P T. Oki-shima Island (offshore), KUZ Z 4925; U Y. Mano, KUZ Z 4229, Z 4286; Z AF. Holotype of S. nishimurai sp. nov. from Hannoura, KUZ Z 4935; AG AQ. Paratypes of S. nishimurai sp. nov. from Hannoura, KUZ Z 4936 Z 4939; AR AT. Oura Port, KUZ Z 4945; AU AY. Minamihama, KUZ Z 4946, Z 4947; AZ BD. Kitafunaki, KUZ Z 4952, Z 4953; BE BG. Oki-shima Island (coast), KUZ Z 4957. Scale bars: 10 mm (1), female teleoconch (A C, G, K, P, S, U, Z AB, AG, AJ, AM, AR, AU, AZ, BE), male teleoconch (J, N, AP, AX, BC), operculum (H, L, O, Q, T, V, Y, AC, AH, AK, AN, AQ, AS, AV, AY, BA, BD, BF); 2 mm (2), protoconch (D F, I, M, R, W); 1 mm (2), protoconch (AD AF, AI, AL, AO, AT, AW, BB, BG)." httpUri="https://binary.pensoft.net/fig/1065979">Figs 7 A Y</figureCitation>
,
<figureCitation id="03BE1E29DB5D7689F84D5AE291ADE016" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Radulae of Semisulcospira reticulata (A D) and S. nishimurai sp. nov. (E I). A. Hannoura, KUZ Z 4926; B. Kitafunaki, KUZ Z 4930; C. Oki-shima Island (offshore), KUZ Z 4925; D. Mano, KUZ Z 4229; E. Holotype of S. nishimurai sp. nov. from Hannoura, KUZ Z 4935; F. Oura Port, KUZ Z 4945; G. Minamihama, KUZ Z 4946; H. Kitafunaki, KUZ Z 4952; I. Oki-shima Island (coast), KUZ Z 4957. Scale bars: 100 μm." httpUri="https://binary.pensoft.net/fig/1065980">8 A D</figureCitation>
,
<figureCitation id="7D26356040569FEAFF25DEAD08850EDF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Schematic drawings indicating generalised external features of the soft body and reproductive organs of Semisulcospira reticulata and S. nishimurai sp. nov.: A, B. Dorsal and ventral views of organs in visceral mass of mature female of S. nishimurai sp. nov. without operculum; C. External, right lateral view of pallial oviduct of S. reticulata; D. External, view of pallial oviduct of S. nishimurai sp. nov.; E. External, right upper lateral view of the prostate of S. nishimurai sp. nov.; F Internal folding structures of the prostate of S. nishimurai sp. nov. Abbreviations: bp, brood pouch; ct, ctenidium; dg, digestive gland; eg, egg; em, embryo; f, foot; inn, inner wall of brood pouch; int, intestine; kd, main kidney chamber; os, osphradium; ov, ovary; ovi, renal oviduct; pr, prostate; rcs, seminal receptacle; sg, sperm gutter; sn, snout; spb, spermatophore bursa; ss, style sac; sto, stomach; t, cephalic tentacle; vd, vas deferens. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1065981">9 C Japanes name: Kagome-kawanina (Kajiyama and Habe 1961</figureCitation>
)
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<treatmentCitation id="6F1E400FC7B834E13E480E48E363C28A" author="Kajiyama &amp; Habe" page="171" year="1961">
<taxonomicName id="E653E66AEF934C2D26A82B9F281780ED" ID-CoL="4WHDP" authority="Kajiyama &amp; Habe, 1961: 171 - 173" authorityName="Kajiyama &amp; Habe" authorityPageNumber="171 - 173" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" order="Sorbeoconcha" phylum="Mollusca" rank="subSpecies" species="decipiens" subSpecies="reticulata">
<emphasis id="3DDAB7665536C16477451CBC4EA2F21E" italics="true">Semisulcospira decipiens reticulata</emphasis>
<comment id="81A6EF6AB1D256A16C1D0547CEED5FE1">
Kajiyama &amp; Habe, 1961: 171173, 175176, figs 6, 6 a;
<treatmentCitation id="F114F57FB99342B9D6363C332B15CEF8" author="Habe T" page="57" year="1965">
<bibRefCitation id="633A87403FBE222B9553C2A872EB0A02" author="Habe T" firstAuthor="Habe" journalOrPublisher="Ltd., Chiyoda" page="57" refId="B13" refString="Habe T (1965) Semisulcospira decipiens. In: Okada Y (Ed.) New Illuscrated Encyclopedia of the Fauna of Japan. Hukuryu-kan Publishing Co. Ltd., Chiyoda, 57 pp." title="Semisulcospira decipiens. In: Okada Y (Ed.) New Illuscrated Encyclopedia of the Fauna of Japan. Hukuryu-kan Publishing Co." year="1965">Habe 1965: 57</bibRefCitation>
</treatmentCitation>
, unnumbered figure;
<treatmentCitation id="5AE0A950D5F42676A18D7874E746A5EB" author="Kawase M &amp; Murase F &amp; Hayase Y &amp; Ichihara T &amp; Morikawa A &amp; Ieyama H" page="37" year="2012">
<bibRefCitation id="FD61EAFC6411D73B344CD0C75E03DA04" author="Kawase M &amp; Murase F &amp; Hayase Y &amp; Ichihara T &amp; Morikawa A &amp; Ieyama H" etAl="et al." firstAuthor="Kawase" journalOrPublisher="Limnology in Tokai Region of Japan" page="37" pagination="33-42" refId="B17" refString="Kawase M, Murase F, Hayase Y, Ichihara T, Morikawa A, Ieyama H (2012) Freshwater mollusks in Gifu city, Gifu prefecture, Japan. Limnology in Tokai Region of Japan 54: 3342." title="Freshwater mollusks in Gifu city, Gifu prefecture, Japan." volume="54" year="2012">Kawase et al. 2012: 37</bibRefCitation>
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, figs 211.
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<treatmentCitation id="B70BA819D486F236A7945B23FB835143" author="Burch JB &amp; Davis GM" page="36 - 37" year="1967">
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<emphasis id="47D8F18FFF4A66A4DFF644FB17F69895" italics="true">Semisulcospira reticulata</emphasis>
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<comment id="F9EC65E1C68A798AE2DFAD8DD1C0A4F4">
<bibRefCitation id="29441839848BBED89CEEE9BE80E7A0D8" author="Burch JB &amp; Davis GM" firstAuthor="Burch" journalOrPublisher="Annual reports for" page="36 - 37" pagination="36-38" refId="B7" refString="Burch JB, Davis GM (1967) A taxonomic study of some species of the freshwater snail genus Semisulcospira in Japan. Annual reports for 1967: 3638." title="A taxonomic study of some species of the freshwater snail genus Semisulcospira in Japan." volume="1967" year="1967">Burch and Davis 1967: 3637</bibRefCitation>
;
<treatmentCitation id="E3B04E1DCF732387CE6DD85672CC5DC2" author="Burch JB" page="6" year="1968">
<bibRefCitation id="410E28C58B0D28385CE12827780C989B" author="Burch JB" firstAuthor="Burch" journalOrPublisher="Journal de Conchyliologie" page="6" pagination="3-51" refId="B6" refString="Burch JB (1968) Cytotaxonomy of some Japanese Semisulcospira (Streptoneura: Pleuroceridae). Journal de Conchyliologie 107: 351." title="Cytotaxonomy of some Japanese Semisulcospira (Streptoneura: Pleuroceridae)." volume="107" year="1968">Burch 1968: 6</bibRefCitation>
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, 2629, fig. 1 D, pl. 4, figs 4, 5, pl. 5, fig. 8;
<treatmentCitation id="C7969511F8B23F2DF6335CAA8518F6D5" author="Davis GM" page="249" year="1969">
<bibRefCitation id="670EC375E45D9E06E751A04C11CE4067" author="Davis GM" journalOrPublisher="Malacologia" pagination="211-294" refId="B10" refString="Davis GM (1969) A taxonomic study of some species of Semisulcospira in Japan (Mesogastropoda: Pleuroceridae). Malacologia 7: 211 - 294." title="A taxonomic study of some species of Semisulcospira in Japan (Mesogastropoda: Pleuroceridae)." volume="7" year="1969">Davis, 1969: 249</bibRefCitation>
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, 255, pl. 5, figs 1, 2, pl. 6, figs 1, 2, pl. 10, figs 10, 11, pl. 11., figs 14;
<treatmentCitation id="73A981DA69997FDB79DB4FE91C4AA4EE" author="Watanabe NC" page="17" year="">
<bibRefCitation id="D55FD364E653B1A660E5C35920469CEC" DOI="10.18941/venusjjm.29.1_13" author="Watanabe NC" journalOrPublisher="Venus" pagination="13-30" refId="B57" refString="Watanabe NC (1970 a) Studies on three species of Semisulcospira in Lake Biwa: i) comparative studies of shell form and habitat. Venus 29: 1330. https://doi.org/10.18941/venusjjm.29.1_13" title="Studies on three species of Semisulcospira in Lake Biwa: i) comparative studies of shell form and habitat." volume="29">Watanabe 1970 a</bibRefCitation>
: 17
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23, fig. 15-3, 8, 9;
<treatmentCitation id="1E38DBD960915AD34B3104CC26400DBB" author="Habe T" page="322" year="1986">
<bibRefCitation id="9DAA1D7B49F19D935B15F036E41C59D9" author="Habe T" editor="Ueno M" firstAuthor="Habe" journalOrPublisher="4 th edn. Hukuryu-kan Publishing Co. Ltd., Chiyoda" page="322" pagination="309-341" refId="B15" refString="Habe T (1986) Mollusca. In: Ueno M (Ed.) The late Tamiji Kawamura Freshwater Biology of Japan. 4 th edn. Hukuryu-kan Publishing Co. Ltd., Chiyoda, 309341." title="Mollusca." volumeTitle="The late Tamiji Kawamura Freshwater Biology of Japan." year="1986">Habe 1986: 322</bibRefCitation>
</treatmentCitation>
, fig. 17-1 - 17;
<treatmentCitation id="2A9B0C586460AFBA4FB061E61FA77CCE">
<bibRefCitation id="49702C63ED417FC0A6CC1649629832CC" DOI="10.1093/mollus/eyv057" author="Kohler" firstAuthor="Kohler" journalOrPublisher="Journal of Molluscan Studies" pagination="268-281" refId="B21" refString="Köhler F (2016) Rampant taxonomic incongruence in a mitochondrial phylogeny of Semisulcospira freshwater snails from Japan (Cerithioidea: Semisulcospiridae). Journal of Molluscan Studies 82: 268281. https://doi.org/10.1093/mollus/eyv057" title="Rampant taxonomic incongruence in a mitochondrial phylogeny of Semisulcospira freshwater snails from Japan (Cerithioidea: Semisulcospiridae)." volume="82" year="2016">Köhler 2016</bibRefCitation>
</treatmentCitation>
: fig 4 L N;
<treatmentCitation id="3964FFCD3F6DF16D2EF5A3A26CE3C277">
<bibRefCitation id="E0C0837B208D6961A325D098E61FF043" DOI="10.1071/IS22042" author="Sawada and Fuke" firstAuthor="Sawada" journalOrPublisher="Invertebrate Systematics" pagination="1139-1177" refId="B46" refString="Sawada N, Fuke Y (2022) Systematic revision of the Japanese freshwater snail Semisulcospira decipiens (Mollusca: Semisulcospiridae): implications for diversification in the ancient Lake Biwa. Invertebrate Systematics 36: 11391177. https://doi.org/10.1071/IS22042" title="Systematic revision of the Japanese freshwater snail Semisulcospira decipiens (Mollusca: Semisulcospiridae): implications for diversification in the ancient Lake Biwa." volume="36" year="2022">Sawada and Fuke 2022</bibRefCitation>
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: fig. 11 bq bu.
</comment>
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</paragraph>
<paragraph id="2E7AEB9F1269D2F69B8BFD0A7084B037">
<treatmentCitationGroup id="F879D25332BFD768759B174B0C26B7FC">
<treatmentCitation id="D8DD7F4D39049E384EA9E71438DB8D06" author="Habe T" page="94" year="1978">
<taxonomicName id="E9EB61AD40660AE06BDCBB93F3FC1391" ID-CoL="4WHCB" baseAuthorityName="Westerlund" baseAuthorityYear="1883" class="Gastropoda" family="Semisulcospiridae" genus="Biwamelania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Sorbeoconcha" phylum="Mollusca" rank="species" species="decipiens">
<emphasis id="251E55C56D3B0C6E873905E6A780AAB4" italics="true">Biwamelania decipiens</emphasis>
</taxonomicName>
<comment id="64491BD4A298AEAA79F297B4453BE231">
<bibRefCitation id="5CD0CB36C9D320536ECE6F770A910B5E" author="Habe T" firstAuthor="Habe" journalOrPublisher="Yonsei Reports on Tropical Medicine" page="94" pagination="91-96" refId="B14" refString="Habe T (1978) Fresh water molluscs in Korea and Japan as the intermediate hosts of trematodes. Yonsei Reports on Tropical Medicine 9: 9196." title="Fresh water molluscs in Korea and Japan as the intermediate hosts of trematodes." volume="9" year="1978">Habe 1978: 94</bibRefCitation>
;
<treatmentCitation id="463E652A886E9AC2403796A7877B863F" author="Minato H" page="79" year="1991">
<bibRefCitation id="CC1C24FF94FDCD54DB7D0787474AACDB" author="Minato H" firstAuthor="Minato" page="79" refId="B27" refString="Minato H (1991) Biwamelania decipiens. In: Okutani T (Ed.) Illustrations of Animals and Plants. Mollusca. Third Edition. Sekaibunkasha Com., Chiyoda, 391 pp." year="1991">Minato 1991: 79</bibRefCitation>
</treatmentCitation>
, unnumbered fig.
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A7EA1C5818BB08FB78B68F77459A9775">
<treatmentCitationGroup id="432B9EE195A7CA7F2EBD56EF864C5958">
<treatmentCitation id="F26DD642A7B62A5DA60266C8197F52D9" author="Matsuoka K" page="190" year="1985">
<taxonomicName id="A0063E8188E032935112947A1E29187C" ID-CoL="4WHDP" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" order="Sorbeoconcha" phylum="Mollusca" rank="species" species="reticulata" subGenus="Biwamelania">
<emphasis id="5D47AEE101FD43DF606A64AD789CAE98" italics="true">Semisulcospira</emphasis>
(
<emphasis id="649BCE7760E76FB801710C7880792187" italics="true">Biwamelania</emphasis>
)
<emphasis id="8CA1B72BD3C67F4B386EB8F12EB4698E" italics="true">reticulata</emphasis>
</taxonomicName>
<comment id="FB0160A09B8CD7C307282DBD020CCAE7">
<bibRefCitation id="361BEA488FCBD3641217C98F8821C928" author="Matsuoka K" firstAuthor="Matsuoka" journalOrPublisher="Transactions and proceedings of the Paleontological Society of Japan" page="190" pagination="180-195" refId="B26" refString="Matsuoka K (1985) Pliocene freshwater gastropods from the Iga formation of the Kobiwako group, Mie Prefecture, central Japan. Transactions and proceedings of the Paleontological Society of Japan 139: 180195." title="Pliocene freshwater gastropods from the Iga formation of the Kobiwako group, Mie Prefecture, central Japan." volume="139" year="1985">Matsuoka 1985: 190</bibRefCitation>
;
<treatmentCitation id="D5316E69D4461E7107277B954A7452D0">
<bibRefCitation id="4C4B9806D4C50ACB3AB3946B9F8AFFD4" author="Watanabe and Nishino" firstAuthor="Watanabe" journalOrPublisher="Lake Biwa Study Monographs" pagination="1-36" refId="B60" refString="Watanabe NC, Nishino M (1995) A study on taxonomy and distribution of the freshwater snail, genus Semisulcospira in Lake Biwa, with descriptions of eight new species. Lake Biwa Study Monographs 6: 136." title="A study on taxonomy and distribution of the freshwater snail, genus Semisulcospira in Lake Biwa, with descriptions of eight new species." volume="6" year="1995">Watanabe and Nishino 1995</bibRefCitation>
</treatmentCitation>
: app. pl. 1, fig. 8, app. pl. 2, figs 22, 23;
<treatmentCitation id="8C546905BEE3C3F6EF4FCCF6FE260E7C">
<bibRefCitation id="2F820E7F12839492AD076F8CF9375CD4" DOI="10.1016/s0065-2504(00)31011-x" author="Nishino and Watanabe" firstAuthor="Nishino" journalOrPublisher="Advances in Ecological Research" pagination="151-180" refId="B36" refString="Nishino M, Watanabe NC (2000) Evolution and endmism in Lake Biwa, with special reference to its gastropod mollusc fauna. Advances in Ecological Research 31: 151180. https://doi.org/10.1016/s0065-2504 (00) 31011 - x" title="Evolution and endmism in Lake Biwa, with special reference to its gastropod mollusc fauna." volume="31" year="2000">Nishino and Watanabe 2000</bibRefCitation>
</treatmentCitation>
: fig. 212;
<treatmentCitation id="A917269FDFF0C1AF172AF09BA7AD027D" author="Matsuda M" page="8 - 9" year="2000">
<bibRefCitation id="6738199141FD929DFA68AA93E6390EF1" author="Matsuda M" editor="Japan" firstAuthor="Matsuda" journalOrPublisher="Japan Fisheries Resource Conservation Association, Chuo" page="8 - 9" pagination="8-9" refId="B24" refString="Matsuda M (2000) Semisulcospira (Biwamelania) reticulata. In: Japan (Ed.) Data Book on Rare Wild Aquatic Organism in Japan. Japan Fisheries Resource Conservation Association, Chuo, 89." title="Semisulcospira (Biwamelania) reticulata." volumeTitle="Data Book on Rare Wild Aquatic Organism in Japan." year="2000">Matsuda 2000: 89</bibRefCitation>
</treatmentCitation>
, unnumbered fig.;
<treatmentCitation id="5E260B5F7D60D66D13EC36E04D1E2117" author="Kihira H &amp; Matsuda M &amp; Uchiyama R" page="26 - 27" year="2009">
<bibRefCitation id="ABBD1817DDBDFE31B6DE90680AE59DD1" author="Kihira H &amp; Matsuda M &amp; Uchiyama R" etAl="et al." firstAuthor="Kihira" journalOrPublisher="Pieces Publishers, Yokohama" page="26 - 27" refId="B19" refString="Kihira H, Matsuda M, Uchiyama R (2009) Guide of Japanese Freshwater Molluscs (1). Freshwater Molluscs in Lake Biwa and Yodo River, Revised Edition. Pieces Publishers, Yokohama, 159 pp." title="Guide of Japanese Freshwater Molluscs (1). Freshwater Molluscs in Lake Biwa and Yodo River, Revised Edition." year="2009">Kihira et al. 2009: 2627</bibRefCitation>
</treatmentCitation>
, 33, unnumbered figs;
<treatmentCitation id="87C2D6CC235B49B34C80635D9C00CEF1">
<bibRefCitation id="D743A38F87BA8FD83D262432EB698A3F" DOI="10.1002/evl3.92" author="Miura" etAl="et al." firstAuthor="Miura" journalOrPublisher="Evolution Letters" pagination="43-54" refId="B28" refString="Miura O, Urabe M, Nishimura T, Nakai K, Chiba S (2019) Recent lake expansion triggered the adaptive radiation of freshwater snails in the ancient Lake Biwa. Evolution Letters 3: 4354. https://doi.org/10.1002/evl3.92" title="Recent lake expansion triggered the adaptive radiation of freshwater snails in the ancient Lake Biwa." volume="3" year="2019">Miura et al. 2019</bibRefCitation>
</treatmentCitation>
: fig. S 1 p r;
<treatmentCitation id="19682769F6E99DF607FCF3C818A77026" author="Nishino M" page="618" year="2021">
<bibRefCitation id="958FEA5D74B96CCD5E261B071529A8F7" author="Nishino M" firstAuthor="Nishino" journalOrPublisher=", Ltd., Hikone" page="618" refId="B35" refString="Nishino M (2021) Semisulcospira (Biwamelania) reticulata. In: Scientific Committee for Research into the Wildlife in Shiga Prefecture (Ed.) Red Data Book of Shiga Prefecture 2020. Sunrise Publishing Co., Ltd., Hikone, 618 pp." title="Semisulcospira (Biwamelania) reticulata. In: Scientific Committee for Research into the Wildlife in Shiga Prefecture (Ed.) Red Data Book of Shiga Prefecture 2020. Sunrise Publishing Co." year="2021">Nishino 2021: 618</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="2024A4C00662B51CFA85B3EAD807E609">
<treatmentCitationGroup id="645280A9A3D1653F5ECC6B39C85AE5BC">
<treatmentCitation id="3F0991AB5F86CD432FF8B3B644D87892" author="Kihira H &amp; Matsuda M" page="22 - 23" year="1990">
<taxonomicName id="8F47C0FBA5A41C85B1F634DE7BB7FADF" ID-CoL="4WHCR" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Biwamelania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Sorbeoconcha" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="D49122AFB22627F8B2525E5E4155107F" italics="true">Biwamelania reticulata</emphasis>
</taxonomicName>
<comment id="2E765A9647DAC2F3E812F3D450BE3E1B">
<bibRefCitation id="26D00A43911DE47566220ACF6F2FC555" author="Kihira H &amp; Matsuda M" firstAuthor="Kihira" journalOrPublisher="Tatara Shobo Publishers, Yonago" page="22 - 23" refId="B18" refString="Kihira H, Matsuda M (1990) Freshwater Molluscs of Lake Biwa and Yodo River. Tatara Shobo Publishers, Yonago, 131 pp." title="Freshwater Molluscs of Lake Biwa and Yodo River." year="1990">Kihira and Matsuda 1990: 2223</bibRefCitation>
, 25, unnumbered figs;
<treatmentCitation id="B44D8960B56B02B3AD98C3CCA1675265" author="Nishino M &amp; Tanida K" page="60" year="2018">
<bibRefCitation id="5CAF10903314EC0ED75B0DE77D0317F9" author="Nishino M &amp; Tanida K" firstAuthor="Nishino" journalOrPublisher="Incorporated Administrative Agency Japan Water Agency Lake Biwa Development Integrated Operation &amp; Maintenance Office, Otsu" page="60" refId="B37" refString="Nishino M, Tanida K (2018) Illustration of Lake Biwa Benthic Fauna. Second Edition. Incorporated Administrative Agency Japan Water Agency Lake Biwa Development Integrated Operation &amp; Maintenance Office, Otsu, 335 pp." title="Illustration of Lake Biwa Benthic Fauna. Second Edition." year="2018">Nishino and Tanida 2018: 60</bibRefCitation>
</treatmentCitation>
, 253, unnumbered figs.
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="3E992E69AEB7917EA2710064AB987A25">
<treatmentCitationGroup id="EDEF024477EDF684B2F759466B44D7B9">
<treatmentCitation id="EA3DF6995774552DE63521D88DF93E60" author="Kawase M &amp; Murase F &amp; Hayase Y &amp; Ichihara T &amp; Morikawa A &amp; Ieyama H" page="37" year="2012">
<taxonomicName id="EB68E09FBC7A82F61BB3D976B0EB0CBD" ID-CoL="4WHDD" baseAuthorityName="Böttger" baseAuthorityYear="1886" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" order="Sorbeoconcha" phylum="Mollusca" rank="subSpecies" species="decipiens" subSpecies="multigranosa">
<emphasis id="134347BDCD99FEBEBE87EE9574A408C4" italics="true">Semisulcospira decipiens multigranosa</emphasis>
</taxonomicName>
<comment id="DE4CD6B7C5FEE2827789D59E3E1B6476">
<bibRefCitation id="446E54C905F40FE6B396FDE5070E0260" author="Kawase M &amp; Murase F &amp; Hayase Y &amp; Ichihara T &amp; Morikawa A &amp; Ieyama H" etAl="et al." firstAuthor="Kawase" journalOrPublisher="Limnology in Tokai Region of Japan" page="37" pagination="33-42" refId="B17" refString="Kawase M, Murase F, Hayase Y, Ichihara T, Morikawa A, Ieyama H (2012) Freshwater mollusks in Gifu city, Gifu prefecture, Japan. Limnology in Tokai Region of Japan 54: 3342." title="Freshwater mollusks in Gifu city, Gifu prefecture, Japan." volume="54" year="2012">Kawase et al. 2012: 37</bibRefCitation>
, figs 210.
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EO3AI" type="material">
<paragraph id="0D015EDA65B64A380BBA0DA801A4F3D0">
<heading id="1E3668E2BCD62A0E78A767621F192389" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="380181A76F91D50CB2653AEE0E71375A">
<materialsCitation id="A471466A13876B13C95178BE00A91A78" country="Japan" county="Omi-hachiman City" location="Mo" municipality="Okishima-cho" specimenCount="1" stateProvince="Shiga" typeStatus="holotype">
<emphasis id="E5BAA54F66FFF75C3D1717E5DC1BC62F" bold="true" italics="true">
<typeStatus id="E2EABE1FD953030F7122F22F7E902A67" type="holotype">Holotype</typeStatus>
.
</emphasis>
<named-content id="2D99960C668B9DAD975EA6002506683F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-science-museum-natural-history" xlink_title="National Museum of Nature and Science, Tokyo">NSMT</named-content>
-
<location id="87BF8610ECCFB92134D96A9931C950CF" LSID="urn:lsid:plazi:treatment:23DD9564B0C05B38A302C10F7EBAD581:87BF8610ECCFB92134D96A9931C950CF" country="Japan" county="Omi-hachiman City" municipality="Okishima-cho" name="Mo" stateProvince="Shiga">Mo</location>
39770, adult female, collected from “ Off Okinoshima in Lake Biwa ” (
<location id="76D031FE5C990426FD915F9EC1F00A3E" LSID="urn:lsid:plazi:treatment:23DD9564B0C05B38A302C10F7EBAD581:76D031FE5C990426FD915F9EC1F00A3E" country="Japan" county="Omi-hachiman City" municipality="Okishima-cho" name="Lake Biwa" stateProvince="Shiga">Lake Biwa</location>
, offshore of
<location id="A6169623DDF8F8F7E4235A2F4C3802CE" LSID="urn:lsid:plazi:treatment:23DD9564B0C05B38A302C10F7EBAD581:A6169623DDF8F8F7E4235A2F4C3802CE" country="Japan" county="Omi-hachiman City" municipality="Okishima-cho" name="Oki-shima Island" stateProvince="Shiga">Oki-shima Island</location>
,
<collectingMunicipality id="F05FEB98A0309A396EC26E09D63B2A4D">Okishima-cho</collectingMunicipality>
,
<collectingCounty id="AA3E2E100C3157EE8EECCDE0C72382B5">Omi-hachiman City</collectingCounty>
,
<collectingRegion id="A037F50EEC7FBDBA67DAA6527CF4C772" country="Japan" name="Shiga">Shiga Prefecture</collectingRegion>
,
<collectingCountry id="4F0E0D0ABD636F1B04679DF44DC296F0" name="Japan">Japan</collectingCountry>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E53AI" type="material">
<paragraph id="9743B223C39246A64A80678449D9B462">
<heading id="12278CFCB8FB3745703DF9B55C4DFA1E" reason="title">Additional materials.</heading>
</paragraph>
<paragraph id="AE1D2513ED87A2632A4B8F307B6674B3">
<materialsCitation id="0A6F53DAD36430FE4FBA25DE05EE2A83" collectingDate="2022-06-26" collectionCode="KUZ" elevation="7" location="Hannoura" specimenCode="KUZ Z 4926" specimenCount="24" specimenCount-female="15" specimenCount-male="9">
<specimenCode id="7D82DBF5BD5BCFAAC3A9641E1B104D68">
<named-content id="991EAA9A5D1B661174B15D6D8D0AA478" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4926
</specimenCode>
,
<specimenCount id="5C357E5761CB1052953A7F1A362C02E2" count="15" type="female">15 females</specimenCount>
, Z 4927,
<specimenCount id="0C2992F24CBD26B2123682E53CEBCE7C" count="9" type="male">9 males</specimenCount>
, collected from
<quantity id="AFC1013062DF6C6029A7BEFE3FB923BD" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">
<elevation id="8131B2593FDFA270B521E024B7917553" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">68 m</elevation>
</quantity>
water depths at
<location id="9AE346D22834825F77AC016E2B12F504" LSID="urn:lsid:plazi:treatment:23DD9564B0C05B38A302C10F7EBAD581:9AE346D22834825F77AC016E2B12F504" name="Hannoura">Hannoura</location>
on
<date id="670D2940A59F6FB0B71A5944C19AF950" value="2022-06-26">
<collectingDate id="1E1CB7E5E94EBCB7CAB782D16C3A74D7" value="2022-06-26">26 June 2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C94567E9A13B034CB909A4EDFBE6644F" collectingDate="2022-07-10" collectionCode="KUZ" elevation="7" location="Kitafunaki" specimenCode="KUZ ZZ 4930" specimenCount="20" specimenCount-female="15" specimenCount-male="5">
<specimenCode id="989A7FC19489875D0949FAF6A15964DC">
<named-content id="15CDAD5081B277C43D03B7D4089665CB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
ZZ 4930
</specimenCode>
,
<specimenCount id="6F59657F89BC770934CBCABBFB6B9D75" count="15" type="female">15 females</specimenCount>
, Z 4931,
<specimenCount id="F1703124D4AB4AA60E4A3B046CF27B35" count="5" type="male">5 males</specimenCount>
, from
<quantity id="CB38DE2BB5F99BB6EAD25B75F287605B" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">
<elevation id="8C3AB5D76FD049C0B5EAA21EFA62CE1B" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">58 m</elevation>
</quantity>
depths at
<location id="FFCE89FCD5309D58C0C817051B167232" LSID="urn:lsid:plazi:treatment:23DD9564B0C05B38A302C10F7EBAD581:FFCE89FCD5309D58C0C817051B167232" name="Kitafunaki">Kitafunaki</location>
on
<date id="3D364225F69C28C249E1129C52B4E0F1" value="2022-07-10">
<collectingDate id="80682AE209459EE8221EC52039C60866" value="2022-07-10">10 July 2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="98794128E898558E05775C99EF976B01" collectingDate="2019-04-28" collectionCode="KUZ" elevation="13" location="offshore at Oki-shima Island" specimenCode="KUZ Z 4925" specimenCount="15" specimenCount-female="15">
<specimenCode id="606D990F52684CE74B1810EA5AAE10F8">
<named-content id="8929972AF019BCC7A40080A688924907" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4925
</specimenCode>
,
<specimenCount id="3229C237DAAC256CB615C15EA25C1728" count="15" type="female">15 females</specimenCount>
, from
<quantity id="C2FD992AB2CB388070BFBA30BC1D22A9" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">
<elevation id="5B5FBFB790F0400CDFA16FDC9611DE93" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">1015 m</elevation>
</quantity>
depths
<location id="17ED410ED25D9A8D16A0E32ECFF3269D" LSID="urn:lsid:plazi:treatment:23DD9564B0C05B38A302C10F7EBAD581:17ED410ED25D9A8D16A0E32ECFF3269D" name="offshore at Oki-shima Island">offshore at Oki-shima Island</location>
on
<date id="5B738385AE40225AAB838E8632A74531" value="2019-04-28">
<collectingDate id="44372A936C9A35A45CF75BAD0DC507A3" value="2019-04-28">28 April 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="2B8F13056ECF30040BBF12D6204B167F" collectingDate="2021-10-12" collectionCode="KUZ" elevation="9" location="Mano" specimenCode="KUZ Z 4229" specimenCount="8" specimenCount-female="4" specimenCount-male="4">
<specimenCode id="51273EE810D8B26921A9C62C14B5D756">
<named-content id="45DF85BA845844D65E9A1A4AE4CB0DDE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4229
</specimenCode>
,
<specimenCount id="54B45A63E009DA4B2289D54F8E1D4ECE" count="4" type="female">4 females</specimenCount>
, Z 4286,
<specimenCount id="12E69A43CE56762F087F59FAD36C683B" count="4" type="male">4 males</specimenCount>
, from
<quantity id="643B608A45629C180034254BE2ECD1F1" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" unit="m" value="9.0" valueMax="12.0" valueMin="6.0">
<elevation id="A9D3901F2D2708CB303F14850E990AF7" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" unit="m" value="9.0" valueMax="12.0" valueMin="6.0">612 m</elevation>
</quantity>
depths at
<location id="F4EABCE6BEC827EBD8B2E287F5CF88BF" LSID="urn:lsid:plazi:treatment:23DD9564B0C05B38A302C10F7EBAD581:F4EABCE6BEC827EBD8B2E287F5CF88BF" name="Mano">Mano</location>
on
<date id="73F71989C25B82079CB0470ABCACA940" value="2021-10-12">
<collectingDate id="A645F364449BF19DE169B6391366318F" value="2021-10-12">12 October 2021</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EY4AI" type="Amended diagnosis">
<paragraph id="020FEA03C6965FC7488FB9092CD972A2">
<heading id="D950163AD26285AB41339A5A15470360" reason="title">Amended diagnosis.</heading>
</paragraph>
<paragraph id="B115B935C03039D13F0421DDB6AEFDB8">
Viviparous. Teleoconch elongated, large in the genus [SH 36.4 ± 3.7 (mean ± SD) (female), 27.2 ± 3.1 (male) mm;
<abbrev id="ABBRID0E54AI" xlink_title="body whorl length">BWL</abbrev>
18.4 ± 1.8, 14.2 ±
<quantity id="4E250F68DEBE28100927652AFAC559A5" metricMagnitude="-4" metricUnit="m" metricValue="8.0" unit="mm" value="0.8">0.8 mm</quantity>
]; spires laterally broadened (
<abbrev id="ABBRID0EC5AI" xlink_title="spire angle">SA</abbrev>
18.0 ± 2.3, 18.8 ± 3.3 degrees), axially compressed (
<abbrev id="ABBRID0EG5AI" xlink_title="whorl elongation ratio">WER</abbrev>
2.4 ± 0.2, 2.7 ± 0.6); outer lip of aperture simple, smooth; aperture rounded (
<abbrev id="ABBRID0EK5AI" xlink_title="aperture slenderness ratio">ASR</abbrev>
1.6 ± 0.1, 1.6 ± 0.1); basal cords few (
<abbrev id="ABBRID0EO5AI" xlink_title="basal cord number">BCN</abbrev>
3.3 ± 0.7, 3.4 ± 0.9); longitudinal ribs distinct, granulated, in large number on penultimate whorl, (
<abbrev id="ABBRID0ES5AI" xlink_title="longitudinal rib number of penultimate whorl">RN</abbrev>
26.4 ± 3.2, 21.6 ± 2.2); spiral cords in medium number (
<abbrev id="ABBRID0EW5AI" xlink_title="spiral cord number of penultimate whorl">SCN</abbrev>
6.2 ± 0.8, 5.3 ± 0.8); color in light brown background. Protoconch large in the genus (
<abbrev id="ABBRID0E15AI" xlink_title="shell length of the largest protoconch">SLP</abbrev>
5.0 ±
<quantity id="887FA82039602AF8F4CF87978A848079" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
,
<abbrev id="ABBRID0E55AI" xlink_title="whorl number of the largest protoconch">WNP</abbrev>
3.8 ± 0.4); longitudinal ribs distinct; spiral cord weak or absent; color in yellowish-brown, with or without 13 dark brown bands.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EC6AI" type="description">
<paragraph id="6AE5867C54E619BB9728ADAA7C72E5BC">
<heading id="24DA3D289D3EC82BE4380C522D796D88" reason="title">
Description of
<typeStatus id="3B04D67EBB32EBAC9430A2422FFC1D01" type="holotype">holotype</typeStatus>
</heading>
</paragraph>
<paragraph id="FDBCB1F080B0E33395AC1950082CB828">
<emphasis id="667575E3EC6FED65C044DA846E42D97D" bold="true">
(
<named-content id="8ADEB52AE110F32E943189E607F307C6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-science-museum-natural-history" xlink_title="National Museum of Nature and Science, Tokyo">
<collectionCode id="B6E75C1EAA5DCF376C25D2DD7CE87E69" country="Japan" name="National Science Museum (Natural History)">NSMT</collectionCode>
</named-content>
- Mo 39770; Fig.
<figureCitation id="3AE7C07B3DE6968BCF8A506D377C69CA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells and opercula of Semisulcospira reticulata (A Y) and S. nishimurai sp. nov. (Z BF). A F. Holotype of S. reticulata from Oki-shima Island (offshore), NSMT - Mo 39770; G J. Hannoura, KUZ Z 4926, Z 4927; K O. Kitafunaki, KUZ Z 4930, Z 4931; P T. Oki-shima Island (offshore), KUZ Z 4925; U Y. Mano, KUZ Z 4229, Z 4286; Z AF. Holotype of S. nishimurai sp. nov. from Hannoura, KUZ Z 4935; AG AQ. Paratypes of S. nishimurai sp. nov. from Hannoura, KUZ Z 4936 Z 4939; AR AT. Oura Port, KUZ Z 4945; AU AY. Minamihama, KUZ Z 4946, Z 4947; AZ BD. Kitafunaki, KUZ Z 4952, Z 4953; BE BG. Oki-shima Island (coast), KUZ Z 4957. Scale bars: 10 mm (1), female teleoconch (A C, G, K, P, S, U, Z AB, AG, AJ, AM, AR, AU, AZ, BE), male teleoconch (J, N, AP, AX, BC), operculum (H, L, O, Q, T, V, Y, AC, AH, AK, AN, AQ, AS, AV, AY, BA, BD, BF); 2 mm (2), protoconch (D F, I, M, R, W); 1 mm (2), protoconch (AD AF, AI, AL, AO, AT, AW, BB, BG)." httpUri="https://binary.pensoft.net/fig/1065979">7 A F</figureCitation>
).
<emphasis id="30CAB9EDCFC5327A4A6723231687D51F" bold="true" italics="true">Teleoconch.</emphasis>
</emphasis>
AH
<quantity id="05849C2269CC12DBB61A5F0D68A48591" metricMagnitude="-3" metricUnit="m" metricValue="9.9" unit="mm" value="9.9">9.9 mm</quantity>
, AL
<quantity id="37225E6C1AC6C619A5F48F7E1838904C" metricMagnitude="-2" metricUnit="m" metricValue="1.04" unit="mm" value="10.4">10.4 mm</quantity>
,
<abbrev id="ABBRID0EV6AI" xlink_title="aperture slenderness ratio">ASR</abbrev>
1.54, AW
<quantity id="B6D174423CA51DFBD8A8BB0D9C0F434D" metricMagnitude="-3" metricUnit="m" metricValue="6.8" unit="mm" value="6.8">6.8 mm</quantity>
,
<abbrev id="ABBRID0EZ6AI" xlink_title="basal cord number">BCN</abbrev>
4,
<abbrev id="ABBRID0E46AI" xlink_title="body whorl length">BWL</abbrev>
<quantity id="62E1C6E28D86BFCFCCFEF6B951297FE7" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" unit="mm" value="17.9">17.9 mm</quantity>
, FWL
<quantity id="8D015297C4CE1C777034288B317203B2" metricMagnitude="-3" metricUnit="m" metricValue="3.8" unit="mm" value="3.8">3.8 mm</quantity>
, PWL
<quantity id="4A98C2C5B9E3E1B44238257E1B46FE65" metricMagnitude="-3" metricUnit="m" metricValue="7.1" unit="mm" value="7.1">7.1 mm</quantity>
,
<abbrev id="ABBRID0EBABI" xlink_title="longitudinal rib number of penultimate whorl">RN</abbrev>
24,
<abbrev id="ABBRID0EFABI" xlink_title="spire angle">SA</abbrev>
19.5 degrees,
<abbrev id="ABBRID0EJABI" xlink_title="spiral cord number of penultimate whorl">SCN</abbrev>
6, SH
<quantity id="14CC306824E6FDF8A5FA2E935F249070" metricMagnitude="-2" metricUnit="m" metricValue="3.57" unit="mm" value="35.7">35.7 mm</quantity>
, SW
<quantity id="3A06863705D03DA6AF7C1E98147BF56C" metricMagnitude="-2" metricUnit="m" metricValue="1.28" unit="mm" value="12.8">12.8 mm</quantity>
, TWL
<quantity id="BC672C04BD037A34D00C6E087BD810A4" metricMagnitude="-3" metricUnit="m" metricValue="4.6" unit="mm" value="4.6">4.6 mm</quantity>
,
<abbrev id="ABBRID0ENABI" xlink_title="whorl elongation ratio">WER</abbrev>
2.60,
<abbrev id="ABBRID0ERABI" xlink_title="whorl number">WN</abbrev>
7.00; shell elongated; spires moderately broadened laterally, slightly convex; suture almost straight; outer lip of aperture simple, smooth; aperture rounded; apex of shell eroded; longitudinal ribs distinct, oblique, opisthocyrt, prominently granulated at intersection with spiral cords; shell surface brown to blackish-brown with deposits.
</paragraph>
<caption id="E60D9EB9F464561C81E27B25FDE4B208" ID-arpha="C47FF4CF-6A48-5C45-9097-FF1A497FE4D0" httpUri="https://binary.pensoft.net/fig/1065979" startId="F7">
<paragraph id="0070C669B0CB6A91ADEE0708EBC1EA41">
<label id="5C420DA3AB678180FCF9EC87FC0194FB">Figure 7.</label>
</paragraph>
<paragraph id="8E780E13900E0642BACE5A51F5268370">
Shells and opercula of
<taxonomicName id="E7C464CC277B66813E9B21A3F0CDEE0B" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="F3B6DCEE48C5F305E603086B74DFDF44" italics="true">Semisulcospira reticulata</emphasis>
</taxonomicName>
(
<emphasis id="DF9F24F54BC408409591ED1402D7C774" bold="true">A Y</emphasis>
) and
<taxonomicName id="C79752059638C0C6939C474DED0170DD" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="0735D5557370D1404F8BD99453634FB5" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0B7625793298AE9C945FA08BBDEB21D2" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="F034D568A7FD37F642251886A115C392" bold="true">Z BF</emphasis>
).
<emphasis id="A7088D3CEDF5EF40297C9FC947EF23AC" bold="true">A F.</emphasis>
Holotype of
<taxonomicName id="B67B3FEC3F7BB4C06B754D4115EC7970" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="F1AAFBC5E7761B9140C88E7BDFB8B694" italics="true">S. reticulata</emphasis>
</taxonomicName>
from Oki-shima Island (offshore),
<named-content id="97932BD5CAD7EF8E18EE51AEBE4A91B1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-science-museum-natural-history" xlink_title="National Museum of Nature and Science, Tokyo">NSMT</named-content>
- Mo 39770;
<emphasis id="319DBD34B86ECAC262C25F4A21CA1F9C" bold="true">G J.</emphasis>
Hannoura,
<specimenCode id="E6472C4C3760C038608E19F28E8F1A09">
<named-content id="ACD245F86499816E88352C34CE34CB07" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4926
</specimenCode>
, Z 4927;
<emphasis id="1E4D7A742301DCBC87B687A1C8C8927C" bold="true">K O.</emphasis>
Kitafunaki,
<specimenCode id="C2B7725286229311B18A7164C476DDF1">
<named-content id="8908762E3C53C5C4808E911EE5435F25" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4930
</specimenCode>
, Z 4931;
<emphasis id="37F8714F1DF5091E7D999005D7D7C11C" bold="true">P T.</emphasis>
Oki-shima Island (offshore),
<specimenCode id="9CE14A34C58150F3176CA8C2B2C54F01">
<named-content id="561846E455A3F2C1D231A4461D94C3B8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4925
</specimenCode>
;
<emphasis id="3E7899FA00D236417F46AC43364FAD4C" bold="true">U Y.</emphasis>
Mano,
<specimenCode id="59A043E019F3F6CEA2671E8A704C125C">
<named-content id="6FEB15DB9C7973DC1F89AEC87CD9954B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4229
</specimenCode>
, Z 4286;
<emphasis id="481BED5F48D3D9D54DEB162547B683CD" bold="true">Z AF.</emphasis>
Holotype of
<taxonomicName id="4F5633467E05039B34EC5AE0FF073DFB" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="8B8F357123D913CAA3B269A9B38A539F" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DCF77ECBD505CDAA068CD81334EA0FAC" rank="species">sp. nov.</taxonomicNameLabel>
from Hannoura,
<specimenCode id="B239208F846D76EE4A4587674C809945">
<named-content id="160B10DB18787D518320C10137FDC84A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4935
</specimenCode>
;
<emphasis id="67A7B6D2FE843EF3CC7BBDD879B0A684" bold="true">AG AQ.</emphasis>
Paratypes of
<taxonomicName id="FC67616491F958233B0F12F5856112F2" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="2FADB1937C97CBDB0ED5C99ADFAB75DC" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7246889FA8DD56C3FFC11552F43C24DB" rank="species">sp. nov.</taxonomicNameLabel>
from Hannoura,
<specimenCode id="08AF9ABD2A5A82377CB44FBEECDA23FF">
<named-content id="24D346C1EC4EECC4128B32741D290AA6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4936
</specimenCode>
Z 4939;
<emphasis id="C06B5777E87D57D1DA385B9BA8B0C4A1" bold="true">AR AT.</emphasis>
Oura Port,
<specimenCode id="61DC1BB0CCAFC9C5103F3AD3404C815D">
<named-content id="83E1FA39173B1F8617BA5F6458905DE2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4945
</specimenCode>
;
<emphasis id="2A3FF28890090A591CFEEA0793523F19" bold="true">AU AY.</emphasis>
Minamihama,
<specimenCode id="B348D8B44E89212C2CF9D2859CA6EFA9">
<named-content id="9C600F28E04B2291098BD594B2769593" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4946
</specimenCode>
, Z 4947;
<emphasis id="8C431E8C9099B9221497C2496DC2733E" bold="true">AZ BD.</emphasis>
Kitafunaki,
<specimenCode id="98D01C029FEFD9706ABAC424A82CBF37">
<named-content id="E15293959882C54BF8BE56DB59FD451F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4952
</specimenCode>
, Z 4953;
<emphasis id="D7990D81C12BD79C5E5B1CFD0A98B05B" bold="true">BE BG.</emphasis>
Oki-shima Island (coast),
<specimenCode id="63357F81AAAF0A3FE09D1A02C72B3741">
<named-content id="2955CCE7A2FD43F47C3852512B6E0457" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4957
</specimenCode>
. Scale bars: 10 mm (
<emphasis id="652E9733AE13606E3E0534889167034B" bold="true">1</emphasis>
), female teleoconch (
<emphasis id="26D3AD42EB58208F13B69A056B45A286" bold="true">A C, G, K, P, S, U, Z AB, AG, AJ, AM, AR, AU, AZ, BE</emphasis>
), male teleoconch (
<emphasis id="AEAD5A8C7C84B5FEDE3AE266736350C0" bold="true">J, N, AP, AX, BC</emphasis>
), operculum (
<emphasis id="C31CC10BBFB9132B2A2CF3BFB015599A" bold="true">H, L, O, Q, T, V, Y, AC, AH, AK, AN, AQ, AS, AV, AY, BA, BD, BF</emphasis>
); 2 mm (
<emphasis id="90AC2AB963327A950870C691392B4327" bold="true">2</emphasis>
), protoconch (
<emphasis id="56A5F722BEACD2C933384791E715EB9D" bold="true">D F, I, M, R, W</emphasis>
); 1 mm (
<emphasis id="6960A735506ACA4E832E529DC8A99096" bold="true">2</emphasis>
), protoconch (
<emphasis id="E12897B809BEDF4CD66E6CB39338C371" bold="true">AD AF, AI, AL, AO, AT, AW, BB, BG</emphasis>
).
</paragraph>
</caption>
<paragraph id="7032310C7738C719A9285D36E2A2274F">
<tp_taxon-name id="D2AE9D791A1D1E9C40E336AB7620039B">
<tp_taxon-name-part id="B7EEDE16EEED2BF70BB2761029E4C62E" reg="Protoconchs" taxon-name-part-type="genus">
<emphasis id="7DAEAB8721F8C562FB303AFD9F606AEA" bold="true" italics="true">Protoconchs.</emphasis>
</tp_taxon-name-part>
</tp_taxon-name>
<abbrev id="ABBRID0EGGBI" xlink_title="longitudinal rib number on body whorl of the largest protoconch">RNP</abbrev>
14,
<abbrev id="ABBRID0EKGBI" xlink_title="shell length of the largest protoconch">SLP</abbrev>
<quantity id="6B7CE42E5C70F1A5093F08F6151B902F" metricMagnitude="-3" metricUnit="m" metricValue="5.8" unit="mm" value="5.8">5.8 mm</quantity>
, SWP
<quantity id="AEEACE6FA409F2B11568862837E8DB06" metricMagnitude="-3" metricUnit="m" metricValue="4.1" unit="mm" value="4.1">4.1 mm</quantity>
,
<abbrev id="ABBRID0EOGBI" xlink_title="whorl number of the largest protoconch">WNP</abbrev>
4.00; shell globose; suture slightly undulating; longitudinal ribs remarkable, with rounded nodes in 2 rows; spiral cord weak on lower part of whorls; shell colored light beige in background, without color band.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESGBI" type="variation">
<paragraph id="5A599BD13A5DE2FBCC59B40F498FED75">
<heading id="12EC1973D1D91A29735987E0B6AFAC69" reason="title">Variation.</heading>
</paragraph>
<paragraph id="75D302FE38E6E255D84521688585466C">
<emphasis id="0246BACBDF56A304170A0E989A23DEED" bold="true" italics="true">Teleoconchs</emphasis>
(Fig.
<figureCitation id="9AC672993DDCE9DEF1E1CC3EB3852835" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells and opercula of Semisulcospira reticulata (A Y) and S. nishimurai sp. nov. (Z BF). A F. Holotype of S. reticulata from Oki-shima Island (offshore), NSMT - Mo 39770; G J. Hannoura, KUZ Z 4926, Z 4927; K O. Kitafunaki, KUZ Z 4930, Z 4931; P T. Oki-shima Island (offshore), KUZ Z 4925; U Y. Mano, KUZ Z 4229, Z 4286; Z AF. Holotype of S. nishimurai sp. nov. from Hannoura, KUZ Z 4935; AG AQ. Paratypes of S. nishimurai sp. nov. from Hannoura, KUZ Z 4936 Z 4939; AR AT. Oura Port, KUZ Z 4945; AU AY. Minamihama, KUZ Z 4946, Z 4947; AZ BD. Kitafunaki, KUZ Z 4952, Z 4953; BE BG. Oki-shima Island (coast), KUZ Z 4957. Scale bars: 10 mm (1), female teleoconch (A C, G, K, P, S, U, Z AB, AG, AJ, AM, AR, AU, AZ, BE), male teleoconch (J, N, AP, AX, BC), operculum (H, L, O, Q, T, V, Y, AC, AH, AK, AN, AQ, AS, AV, AY, BA, BD, BF); 2 mm (2), protoconch (D F, I, M, R, W); 1 mm (2), protoconch (AD AF, AI, AL, AO, AT, AW, BB, BG)." httpUri="https://binary.pensoft.net/fig/1065979">7 G, J, K, N, P, S, U, X</figureCitation>
). Measurements and counts shown in Table
<tableCitation id="CAD1D596256226BFA5BEA68E131259D3" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphometric characters of Semisulcospira reticulata and S. nishimurai sp. nov. Measurements and counts: minimum maximum value (mean ± SD)." httpUri="http://table.plazi.org/id/739C4F23DED6447AA0BA4D6A8D9BC1EC" tableUuid="739C4F23DED6447AA0BA4D6A8D9BC1EC">4</tableCitation>
. Spires slightly to moderately broadened, slightly to strongly convex; suture typically slightly undulating, sometimes almost straight; longitudinal ribs distinct, oblique, typically orthocline to opisthocyrt on upper whorls, opisthocyrt on lower whorls, sometimes opisthocline on upper and lower whorls, rarely fade in body whorl; spiral cords rarely distinct; shell colored yellowish-brown to light brown, with dark brown longitudinal stripes; color bands dark brown or dark olive, sometimes present on central part of upper whorls.
</paragraph>
<caption id="739C4F23DED6447AA0BA4D6A8D9BC1EC" ID-Table-UUID="739C4F23DED6447AA0BA4D6A8D9BC1EC" httpUri="http://table.plazi.org/id/739C4F23DED6447AA0BA4D6A8D9BC1EC" startId="T4">
<paragraph id="8DE24AA0FA40F6E2C11ADD7092ED66E1">
<label id="C6A7D7302D88C716A2A1AD3B1FB83231">Table 4.</label>
</paragraph>
<paragraph id="B732EBCA73E58D846447D9105DE91386">
Morphometric characters of
<taxonomicName id="438F3DC52BED708BC376CFCB8FC3378F" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="B5EDC5B45FBFF3969522D4B2456AB837" italics="true">Semisulcospira reticulata</emphasis>
</taxonomicName>
and
<taxonomicName id="2578CABE56BAE9A427C5BF4D617924A1" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="229B9EFEC6F9A576C12C3D14CC092704" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36260271E049AEA5EE9DFBB320776245" rank="species">sp. nov.</taxonomicNameLabel>
Measurements and counts: minimum maximum value (mean ± SD).
</paragraph>
</caption>
<paragraph id="16459B00E7AA5BACDF08C0058253401D">
<table id="TID0EOVBI" rules="all">
<tbody id="87ED51EF1E5EF4FE6C47499508E8CB9B">
<tr id="3BE6BAA2519190D5B4D1CEA013B71192">
<th id="F94D0DBC7A7AD274E11C8CD9AA5641BF" colspan="1" rowspan="1" style="color: #231f20">Species</th>
<emphasis id="FD0B3F0A26216FA5812E54937DEE8654" italics="true">
<th id="8B9EBD1586919FB85B9855922660D1D1" colspan="4" rowspan="1" style="color: #231f20">
<taxonomicName id="C06C1D048DAD80C45BC1F5AF1BBA1894" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">Semisulcospira reticulata</taxonomicName>
</th>
<th id="D465B4FEE9F59ED9CFABD9D713FF524A" colspan="5" rowspan="1" style="color: #231f20">
<taxonomicName id="C8D4566307C1DD24B3C493BE1F17F884" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">Semisulcospira nishimurai</taxonomicName>
<taxonomicNameLabel id="57C4365EA3ADAF17545B8D3755E8B30E" rank="species">sp. nov.</taxonomicNameLabel>
</th>
</emphasis>
</tr>
<tr id="27CB0F1DD36938EE66359846997C63F5">
<td id="B1089F832445C99D375D8B1793E26504" colspan="1" rowspan="1" style="color: #231f20">Locality</td>
<td id="4564967A4308E4D26833AE9494562B0E" colspan="1" rowspan="1" style="color: #231f20">Hannoura</td>
<td id="C15B66AB51B84A4BF158F702C2FA032F" colspan="1" rowspan="1" style="color: #231f20">Kitafunaki</td>
<td id="CF2AE565BE59100FEBC30155E863D827" colspan="1" rowspan="1" style="color: #231f20">Oki shima Island</td>
<td id="4B9A33625A18550BC991D406A2993278" colspan="1" rowspan="1" style="color: #231f20">Mano</td>
<td id="19FA2688688E8303D0F53BD650F1C0A0" colspan="1" rowspan="1" style="color: #231f20">Hannoura</td>
<td id="F32ECD1D2A7643AEB3C6F70CFB173ED7" colspan="1" rowspan="1" style="color: #231f20">Oura Port</td>
<td id="8590ACB2229C66653591AA7C686E0C2D" colspan="1" rowspan="1" style="color: #231f20">Minamihama</td>
<td id="DDED433115C66322BAE554EF19F7729F" colspan="1" rowspan="1" style="color: #231f20">Kitahunaki</td>
<td id="CCFFE15E4DF9C3591CCACB053A4E92EE" colspan="1" rowspan="1" style="color: #231f20">Oki shima Island</td>
</tr>
<tr id="D8296D0F4DAAF863B846347257CCB955">
<td id="92B668721E2BB43939C9790174BAF7D5" colspan="1" rowspan="1" style="color: #231f20">specimen number of shells (mature male / mature female / operculum / protoconch)</td>
<td id="75A5FBB7F8AA3C63A2FD1F202B98279D" colspan="1" rowspan="1" style="color: #231f20">3 / 15 / 5 / 9</td>
<td id="07C4DE04A85DB421FBD94124C382D73F" colspan="1" rowspan="1" style="color: #231f20">5 / 15 / 8 / 15</td>
<td id="2038671561F37C9A90003FAD1AB463CF" colspan="1" rowspan="1" style="color: #231f20">0 / 15 / 7 / 10</td>
<td id="3E8F3AFE8E16774A3FC5F036386165BB" colspan="1" rowspan="1" style="color: #231f20">4 / 4 / 2 / 3</td>
<td id="D88704DC90FC51E69F1934E6577CBC09" colspan="1" rowspan="1" style="color: #231f20">12 / 15 / 11 / 14</td>
<td id="A31DE3C1C3D760C50DA5F19924D998A5" colspan="1" rowspan="1" style="color: #231f20">0 / 2 / 2 / 2</td>
<td id="0204B2AE0B6F4F2D382F80492172D472" colspan="1" rowspan="1" style="color: #231f20">13 / 15 / 7 / 12</td>
<td id="B1E9407EB318878A8F5F2A9DB8BBD475" colspan="1" rowspan="1" style="color: #231f20">5 / 14 / 5 / 11</td>
<td id="52C09116BC8EA2EA9C35982E28634D23" colspan="1" rowspan="1" style="color: #231f20">0 / 3 / 3 / 2</td>
</tr>
<tr id="9F2C01663358FA20580AF1872CF922CB">
<td id="8589C29DC6DD3EECE981E7DE635704B5" colspan="1" rowspan="1" style="color: #231f20">specimen number of radulae</td>
<td id="9CC4E4BDFCC7A6261929272C2E659006" colspan="1" rowspan="1" style="color: #231f20">5</td>
<td id="E47FB7A718A0260BF6EAE297EED4820E" colspan="1" rowspan="1" style="color: #231f20">5</td>
<td id="1F10BB499BA9A8967E005240A03E0DA1" colspan="1" rowspan="1" style="color: #231f20">4</td>
<td id="842D4762D44916183217F29AB30F7126" colspan="1" rowspan="1" style="color: #231f20">3</td>
<td id="850B6668EE33AB72B7F21BAB7A9F3CB7" colspan="1" rowspan="1" style="color: #231f20">5</td>
<td id="FF3BC5563CB34CC9435D5B7E9DC53B20" colspan="1" rowspan="1" style="color: #231f20">2</td>
<td id="CEB5F23D2F3BED60F2346F60107B8F7B" colspan="1" rowspan="1" style="color: #231f20">5</td>
<td id="096BA1CA935BCA5F0A7F01852177EA8E" colspan="1" rowspan="1" style="color: #231f20">5</td>
<td id="C9D5434026C5286012466806465BF646" colspan="1" rowspan="1" style="color: #231f20">2</td>
</tr>
<tr id="271724044400767C52D8691D6AF11C8E">
<td id="6F9DDD1C7C514143F7F245E0452A6606" colspan="1" rowspan="1" style="color: #231f20">specimen number of reproductive organs (male / female)</td>
<td id="C437F6AF6F920CA8B6CEFC4D504802C2" colspan="1" rowspan="1" style="color: #231f20">3 / 8</td>
<td id="35DA375C13397EEE1D4B224966D1104F" colspan="1" rowspan="1" style="color: #231f20">3 / 8</td>
<td id="942C9694CD2AC7717D1C5C9D2880F1A5" colspan="1" rowspan="1" style="color: #231f20">0 / 10</td>
<td id="F156B95491604C5BA632D26432636D0E" colspan="1" rowspan="1" style="color: #231f20">5 / 4</td>
<td id="5949EB959283927A23F0FA8534FFCDE4" colspan="1" rowspan="1" style="color: #231f20">6 / 8</td>
<td id="8213409C6C3BF9F84C1191B47A47B8E3" colspan="1" rowspan="1" style="color: #231f20">0 / 0</td>
<td id="FAD1620BBD3BD763FF44016E4066F6D2" colspan="1" rowspan="1" style="color: #231f20">5 / 7</td>
<td id="02E5CFD224FE69723D7E5C189D45B86F" colspan="1" rowspan="1" style="color: #231f20">5 / 7</td>
<td id="F88E26AD18BC2E49B9601B0A5C073E3C" colspan="1" rowspan="1" style="color: #231f20">0 / 1</td>
</tr>
<tr id="40581F5E3915CA5D9B49DC2CDD23154E">
<td id="3915497AA54CC125B35466E0CA165677" colspan="10" rowspan="1" style="color: #231f20">
<emphasis id="DF2B47424C903B52C8522B6CA3FF4C79" bold="true">Morphological characters of mature female teleoconchs</emphasis>
</td>
</tr>
<tr id="37E07F1DE862324FF83EA517CBB27D7B">
<td id="3CEB2F9B17A398287634EBB2AFBA0454" colspan="1" rowspan="1" style="color: #231f20">Aperture height (AH) (mm)</td>
<td id="CB05F4CF0DF5DF96EE6EC191F7A910CB" colspan="1" rowspan="1" style="color: #231f20">10.213.1 (11.6 ± 0.8)</td>
<td id="F2C47279D519799D17527B4B114E226B" colspan="1" rowspan="1" style="color: #231f20">9.512.0 (10.1 ± 0.6)</td>
<td id="FE89354E975CFA1E24383652883DF1D8" colspan="1" rowspan="1" style="color: #231f20">8.912.0 (10.2 ± 0.9)</td>
<td id="489073289D44B964639F345000704578" colspan="1" rowspan="1" style="color: #231f20">8.012.0 (10.3 ± 1.7)</td>
<td id="68AA40DCE23485864A35424503D23DC6" colspan="1" rowspan="1" style="color: #231f20">8.112.5 (9.9 ± 1.0)</td>
<td id="68100F6A12BE8C222FFDB8DDEB3C694A" colspan="1" rowspan="1" style="color: #231f20">10.010.1 (10.0 ± 0.1)</td>
<td id="E8824D9BD10F5FA1B53A137F3D2D163B" colspan="1" rowspan="1" style="color: #231f20">7.49.7 (8.8 ± 0.5)</td>
<td id="F34811AF013DFB3D12BEB7412E43C264" colspan="1" rowspan="1" style="color: #231f20">8.011.9 (9.0 ± 0.9)</td>
<td id="8610F55EAEE255013AD10B1F28DC26FB" colspan="1" rowspan="1" style="color: #231f20">8.18.2 (8.1 ± 0.1)</td>
</tr>
<tr id="C7C5DFAB74574D7CB008C92BD693B45F">
<td id="68107F766263B399DDA68B4DB1286563" colspan="1" rowspan="1" style="color: #231f20">Aperture length (AL) (mm)</td>
<td id="1817BACBF9F1A13EBF131B48AD2C44A7" colspan="1" rowspan="1" style="color: #231f20">9.013.6 (11.9 ± 1.1)</td>
<td id="070D38C44900AA25699C3E3A442F1F6C" colspan="1" rowspan="1" style="color: #231f20">9.712.9 (10.5 ± 0.8)</td>
<td id="6C842A3F6A2E3BD93B21F2DDB4104A59" colspan="1" rowspan="1" style="color: #231f20">9.512.5 (10.8 ± 0.8)</td>
<td id="9BA1A0144FC033E2F602B86C70272C56" colspan="1" rowspan="1" style="color: #231f20">8.712.1 (10.7 ± 1.4)</td>
<td id="81E7529F29E71CAC79A196A0F58712E4" colspan="1" rowspan="1" style="color: #231f20">8.713.1 (10.2 ± 1.1)</td>
<td id="986CF64B645947AC1EAD80B0222B9FE5" colspan="1" rowspan="1" style="color: #231f20">10.210.3 (10.3 ± 0.1)</td>
<td id="238D3A13D763701C017D8F5E8ECA9F6D" colspan="1" rowspan="1" style="color: #231f20">7.49.8 (8.9 ± 0.6)</td>
<td id="A7813FD08DDF1289837122FECADF121B" colspan="1" rowspan="1" style="color: #231f20">7.812.0 (9.0 ± 1.0)</td>
<td id="74F174DB92741AB79D412F379EFBFEE1" colspan="1" rowspan="1" style="color: #231f20">7.98.4 (8.1 ± 0.2)</td>
</tr>
<tr id="C078C02C79E467CC8A2D031CEFED92F5">
<td id="76EDE57F0BDAF6C3760EF892CACB29A4" colspan="1" rowspan="1" style="color: #231f20">
Aperture slenderness ratio (
<abbrev id="ABBRID0EWRBI" xlink_title="aperture slenderness ratio">ASR</abbrev>
)
</td>
<td id="87136E4438AFF58E88B60A951E5E7D1A" colspan="1" rowspan="1" style="color: #231f20">1.311.68 (1.5 ± 0.1)</td>
<td id="106373B035896C61998F88927E657A2A" colspan="1" rowspan="1" style="color: #231f20">1.521.89 (1.7 ± 0.11)</td>
<td id="7C6C5BEA23B24998F29F40E8C2F21EEC" colspan="1" rowspan="1" style="color: #231f20">1.471.91 (1.6 ± 0.1)</td>
<td id="A281D3C87B469BAEB8768AD256533397" colspan="1" rowspan="1" style="color: #231f20">1.301.50 (1.4 ± 0.1)</td>
<td id="95BE4C62D9EA164541228DD9B7CA6538" colspan="1" rowspan="1" style="color: #231f20">1.421.68 (1.6 ± 0.1)</td>
<td id="58964B915C8AD0EFE5108105AA81E572" colspan="1" rowspan="1" style="color: #231f20">1.621.64 (1.6 ± 0.1)</td>
<td id="D861FA71F1A50F263E714A408F170F77" colspan="1" rowspan="1" style="color: #231f20">1.461.77 (1.6 ± 0.1)</td>
<td id="6AC559DC6EC4A3AC2D31DCC7BD69DCDA" colspan="1" rowspan="1" style="color: #231f20">1.421.81 (1.6 ± 0.1)</td>
<td id="117CDFD4CDFAA549EEA3F57450ED3C83" colspan="1" rowspan="1" style="color: #231f20">1.491.64 (1.6 ± 0.1)</td>
</tr>
<tr id="2D3F5D14E644EC30C3EF3FC954406FD2">
<td id="1A7A50BCE9F37117D06F23C643ABAF9D" colspan="1" rowspan="1" style="color: #231f20">Aperture width (AW) (mm)</td>
<td id="8BD1E26B236A6367A9158D922DD40BD4" colspan="1" rowspan="1" style="color: #231f20">6.98.5 (7.7 ± 0.5)</td>
<td id="EF591CC121E25E46D94F89CEF2D32539" colspan="1" rowspan="1" style="color: #231f20">5.77.2 (6.2 ± 0.4)</td>
<td id="A760C46A75D4483E966EAE180279DF5C" colspan="1" rowspan="1" style="color: #231f20">5.67.8 (6.6 ± 0.5)</td>
<td id="414BD04CF72E58B20B9EFD4DD734C9A4" colspan="1" rowspan="1" style="color: #231f20">6.18.4 (7.6 ± 1.0)</td>
<td id="A4E474C6C3D16DDD740338422E25FF30" colspan="1" rowspan="1" style="color: #231f20">5.67.8 (6.4 ± 0.5)</td>
<td id="32ADBD557C121EB2998D65916CCE12F1" colspan="1" rowspan="1" style="color: #231f20">6.3</td>
<td id="44808C908D5550F94348DD92A19FEFED" colspan="1" rowspan="1" style="color: #231f20">4.56.1 (5.6 ± 0.4)</td>
<td id="1E69D97E9DD4384498F80056BB3C5556" colspan="1" rowspan="1" style="color: #231f20">4.67.2 (5.6 ± 0.7)</td>
<td id="5D77336E9145A61F837A59F04FE4EECA" colspan="1" rowspan="1" style="color: #231f20">5.15.3 (5.2 ± 0.1)</td>
</tr>
<tr id="04DF58ABB51927BC54A9A51E187D5CA1">
<td id="4F45BBBA43C9977EB8BB0E5DDCF58707" colspan="1" rowspan="1" style="color: #231f20">
Basal cord number (
<abbrev id="ABBRID0ENUBI" xlink_title="basal cord number">BCN</abbrev>
)
</td>
<td id="4F9073FAC2B31C64B4CC714F90ACA006" colspan="1" rowspan="1" style="color: #231f20">25 (3.7 ± 0.7)</td>
<td id="8E44890F8F2F753AA772F127C5707C8B" colspan="1" rowspan="1" style="color: #231f20">34 (3.3 ± 0.5)</td>
<td id="16AE77976FE9D7BB814FCE490C40D630" colspan="1" rowspan="1" style="color: #231f20">24 (2.9 ± 0.5)</td>
<td id="4222CEEA1061D129A8107E35AE7331A1" colspan="1" rowspan="1" style="color: #231f20">34 (3.5 ± 0.6)</td>
<td id="9409D1A6BEE3FAA572D979A79AE02453" colspan="1" rowspan="1" style="color: #231f20">24 (3.2 ± 0.7)</td>
<td id="B544409CEB341D299FF3E734FA76E0F3" colspan="1" rowspan="1" style="color: #231f20">34 (3.5 ± 0.7)</td>
<td id="43B00D047E6AEF6EA95A5BF44030069C" colspan="1" rowspan="1" style="color: #231f20">24 (2.7 ± 0.6)</td>
<td id="71011B5C2650B1DE8F92A33F163A3C75" colspan="1" rowspan="1" style="color: #231f20">24 (2.9 ± 0.5)</td>
<td id="F0A859C7C057B337EEC96F62A080B076" colspan="1" rowspan="1" style="color: #231f20">23 (2.7 ± 0.6)</td>
</tr>
<tr id="B5E45F3C14DA2384C2DFA70D5EA7A67A">
<td id="87A6A1EC116FDD3DE3C94EF03C807C12" colspan="1" rowspan="1" style="color: #231f20">
Body whorl length (
<abbrev id="ABBRID0E2VBI" xlink_title="body whorl length">BWL</abbrev>
) (mm)
</td>
<td id="89F5D6EBE2CF222D5E38F04B5CB31AD0" colspan="1" rowspan="1" style="color: #231f20">17.922.4 (20.1 ± 1.2)</td>
<td id="0BB01D074ECE3BB593636411D6C9E364" colspan="1" rowspan="1" style="color: #231f20">16.320.6 (17.3 ± 1.1)</td>
<td id="BC2E8C895B064685D737377FA178886D" colspan="1" rowspan="1" style="color: #231f20">16.220.7 (18.3 ± 1.4)</td>
<td id="C393397BA3983A85390F283995DBDD8F" colspan="1" rowspan="1" style="color: #231f20">13.719.9 (17.6 ± 2.8)</td>
<td id="0CCA4B1BF490CB4531237434B4E7A762" colspan="1" rowspan="1" style="color: #231f20">15.022.0 (17.2 ± 1.7)</td>
<td id="66C12981D5888AC2BFBE23CBE1A4CE5C" colspan="1" rowspan="1" style="color: #231f20">17.017.9 (17.5 ± 0.6)</td>
<td id="E858264D8FC8BEF3D9BBB57FF07E3AEC" colspan="1" rowspan="1" style="color: #231f20">12.816.5 (15.1 ± 0.8)</td>
<td id="44CB68FE704FBD09B128264CC0452032" colspan="1" rowspan="1" style="color: #231f20">14.019.4 (15.5 ± 1.4)</td>
<td id="B69FC4F655A0715233CBB2ED0114275D" colspan="1" rowspan="1" style="color: #231f20">13.714.2 (13.9 ± 0.2)</td>
</tr>
<tr id="9581CBCE2230215A65D31409DDD140A4">
<td id="EFD920CE22AEE0C1D0B352C48A630672" colspan="1" rowspan="1" style="color: #231f20">Fourth whorl length (FWL) (mm)</td>
<td id="EF42574B79E91D017346EE3740F88134" colspan="1" rowspan="1" style="color: #231f20">4.05.2 (4.7 ± 0.3)</td>
<td id="793D2D4ADEF30D5C6E466AC360D6BA59" colspan="1" rowspan="1" style="color: #231f20">3.84.6 (4.1 ± 0.3)</td>
<td id="5C4397F5467C59DEA9F13E27C8EBE13A" colspan="1" rowspan="1" style="color: #231f20">3.95.2 (4.4 ± 0.4)</td>
<td id="E69920A7A2506327F0D160DD584427E4" colspan="1" rowspan="1" style="color: #231f20">3.44.4 (4.1 ± 0.5)</td>
<td id="B04A15B24701E8ECAF14C1E072491748" colspan="1" rowspan="1" style="color: #231f20">3.35.3 (4.1 ± 0.5)</td>
<td id="AD0366E4DC97312E08FA55F1703EC239" colspan="1" rowspan="1" style="color: #231f20">3.64.4 (4.0 ± 0.6)</td>
<td id="8D0232178DED3A62DE75795CE023A982" colspan="1" rowspan="1" style="color: #231f20">3.14.2 (3.7 ± 0.3)</td>
<td id="EB7C1B325C6E25D620FE8C3724421DFA" colspan="1" rowspan="1" style="color: #231f20">3.34.4 (3.8 ± 0.3)</td>
<td id="C8D3D3DA4BE8DFB3CC0B9863AA3A3791" colspan="1" rowspan="1" style="color: #231f20">3.23.3 (3.3 ± 0.1)</td>
</tr>
<tr id="90565477D8D4AD448F000BC21BF5F37E">
<td id="2AD0F50E25CDA23AC8CE47332B980DEC" colspan="1" rowspan="1" style="color: #231f20">Penultimate whorl length (PWL) (mm)</td>
<td id="A4E2367BA76BE3FF7494E8BC4D9A52B1" colspan="1" rowspan="1" style="color: #231f20">6.88.3 (7.7 ± 0.5)</td>
<td id="D21D7BE3EF557858B7FF41FBC9239DD1" colspan="1" rowspan="1" style="color: #231f20">5.87.7 (6.5 ± 0.5)</td>
<td id="FB972B1C6EFC15576E478E88FA36D003" colspan="1" rowspan="1" style="color: #231f20">6.18.7 (7.4 ± 0.8)</td>
<td id="6B839CAF86E2638728126158C837AB35" colspan="1" rowspan="1" style="color: #231f20">5.17.4 (6.7 ± 1.1)</td>
<td id="451289B34C3EB14727749AAFEF7476FB" colspan="1" rowspan="1" style="color: #231f20">6.08.5 (6.6 ± 0.7)</td>
<td id="B27BD23D2FF385AF95DF91D0E4045135" colspan="1" rowspan="1" style="color: #231f20">6.47.4 (6.9 ± 0.7)</td>
<td id="81FB840056CE99B8F6C2A1F6D74F2C08" colspan="1" rowspan="1" style="color: #231f20">4.86.3 (5.8 ± 0.4)</td>
<td id="116FFCB88B9E284115240A3C46DC83E1" colspan="1" rowspan="1" style="color: #231f20">4.97.4 (6.1 ± 0.7)</td>
<td id="DB1873F2D1548D8A7FE9024DD7E88616" colspan="1" rowspan="1" style="color: #231f20">5.25.5 (5.3 ± 0.2)</td>
</tr>
<tr id="3591A98D2FA8A5CEC5AAEA72EC4AA719">
<td id="4F972EB72BFDBF9139D36CA40D67CFC4" colspan="1" rowspan="1" style="color: #231f20">
Longitudinal rib number of penultimate whorl (
<abbrev id="ABBRID0E2ZBI" xlink_title="longitudinal rib number of penultimate whorl">RN</abbrev>
)
</td>
<td id="ACA00B8093D29AABF18FFB56EC3E169A" colspan="1" rowspan="1" style="color: #231f20">2031 (26.5 ± 2.9)</td>
<td id="7DAECB37D5A5E9024FA3B0E3DCE1A774" colspan="1" rowspan="1" style="color: #231f20">2233 (27.4 ± 3.0)</td>
<td id="C429DCC2B3CA7CAE9A56FE87B4151B38" colspan="1" rowspan="1" style="color: #231f20">2030 (24.7 ± 2.8)</td>
<td id="15284C5FB779BFC47FD05E52EA981229" colspan="1" rowspan="1" style="color: #231f20">2534 (29.3 ± 4.4)</td>
<td id="30E9BE07CBE22F53520828F7999D57D2" colspan="1" rowspan="1" style="color: #231f20">1628 (22.8 ± 3.4)</td>
<td id="510C559E75CC24A3E97CA27B37A742D4" colspan="1" rowspan="1" style="color: #231f20">2125 (23.0 ± 2.8)</td>
<td id="DFD75014D1A9DEA55B380F35C3739F61" colspan="1" rowspan="1" style="color: #231f20">1421 (18.6 ± 1.8)</td>
<td id="078F97CCC2BB600064DD4FB747E7CC3D" colspan="1" rowspan="1" style="color: #231f20">1731 (20.5 ± 4.0)</td>
<td id="7FA3969891F14DDAB3FEE1D9E9FA83B3" colspan="1" rowspan="1" style="color: #231f20">1517 (15.7 ± 1.2)</td>
</tr>
<tr id="2CF30DB423FC04FE92D6393774763D96">
<td id="43A485E688BFB8DFD01F7B2A36CFE0A6" colspan="1" rowspan="1" style="color: #231f20">
Spire angle (
<abbrev id="ABBRID0EJ2BI" xlink_title="spire angle">SA</abbrev>
) (degrees)
</td>
<td id="A06E01486F128720EAF09D3BFCEEACB6" colspan="1" rowspan="1" style="color: #231f20">13.722.3 (19.0 ± 2.4)</td>
<td id="3B7ED70F1B9983E76AC924B44E369FA2" colspan="1" rowspan="1" style="color: #231f20">14.120.4 (17.4 ± 2.2)</td>
<td id="4B78C3524EC8478306784FF8503569B9" colspan="1" rowspan="1" style="color: #231f20">14.422.4 (17.4 ± 2.1)</td>
<td id="337CD170C81671BAA5EAF23787809AB4" colspan="1" rowspan="1" style="color: #231f20">18.520.3 (19.5 ± 0.8)</td>
<td id="00532116DEE201A1BCFB0AA5304B57F6" colspan="1" rowspan="1" style="color: #231f20">15.020.5 (17.9 ± 1.8)</td>
<td id="752E858079F27EBFEEAC969346275581" colspan="1" rowspan="1" style="color: #231f20">17.718.6 (18.1 ± 0.6)</td>
<td id="52C4B28A96928F26592A037895033802" colspan="1" rowspan="1" style="color: #231f20">13.823.2 (18.0 ± 2.9)</td>
<td id="DC6F6CAD785DA868D520568228136BBA" colspan="1" rowspan="1" style="color: #231f20">13.018.1 (15.3 ± 1.6)</td>
<td id="9C729F390FDD3F56127A836FE4F6860F" colspan="1" rowspan="1" style="color: #231f20">18.220.4 (19.3 ± 1.1)</td>
</tr>
<tr id="BC2A6C717055D103A37A2F555B6C0BB3">
<td id="2AE2933998F37809796C5BA9D392636C" colspan="1" rowspan="1" style="color: #231f20">
Spiral cord number of penultimate whorl (
<abbrev id="ABBRID0EX3BI" xlink_title="spiral cord number of penultimate whorl">SCN</abbrev>
)
</td>
<td id="FC9CEBFBEEC13BA531C707C0A4C6192C" colspan="1" rowspan="1" style="color: #231f20">68 (6.5 ± 0.7)</td>
<td id="BE533F44B76C86437402EC205B0AEB24" colspan="1" rowspan="1" style="color: #231f20">57 (6.2 ± 0.9)</td>
<td id="C23F4068D770030C5FCC08B84A68725E" colspan="1" rowspan="1" style="color: #231f20">57 (6.0 ± 0.7)</td>
<td id="F95ED2782F7ED95EEE8671652BB2070A" colspan="1" rowspan="1" style="color: #231f20">56 (5.5 ± 0.6)</td>
<td id="1217D7D12BB530E7DFBB4CD332E55612" colspan="1" rowspan="1" style="color: #231f20">57 (5.9 ± 0.7)</td>
<td id="34AF77532B93BB0FD6F5889DB66037AD" colspan="1" rowspan="1" style="color: #231f20">67 (6.5 ± 0.7)</td>
<td id="4903A5106D21F43E06123C4F7A43CB02" colspan="1" rowspan="1" style="color: #231f20">67 (6.4 ± 0.5)</td>
<td id="1C3673E07608549B248AB931A6BC2B37" colspan="1" rowspan="1" style="color: #231f20">59 (6.5 ± 1.1)</td>
<td id="81122EF666E26245397E669D7C9A5BD7" colspan="1" rowspan="1" style="color: #231f20">67 (6.5 ± 0.7)</td>
</tr>
<tr id="7BD84B7C992C621D627DF2A75C45ADD4">
<td id="A55FC0D2368BC4B1BF69CB195A4F6BBF" colspan="1" rowspan="1" style="color: #231f20">Shell length (SL) (mm)</td>
<td id="B9521AFF449E07A15150B058FE170EB7" colspan="1" rowspan="1" style="color: #231f20">32.141.7 (38.0 ± 2.9)</td>
<td id="73550E4864F79624A507AEA0957FE179" colspan="1" rowspan="1" style="color: #231f20">30.139.5 (33.4 ± 2.5)</td>
<td id="54654A2DD627F74A864EC27D442CD256" colspan="1" rowspan="1" style="color: #231f20">31.943.8 (37.7 ± 3.4)</td>
<td id="29915C70CBC8CA0240019F6E988B03DD" colspan="1" rowspan="1" style="color: #231f20">28.241.2 (36.6 ± 5.7)</td>
<td id="08B91206268A7A0EB50A75BB78BD8100" colspan="1" rowspan="1" style="color: #231f20">29.340.4 (33.8 ± 2.7)</td>
<td id="4488888B3DD31523D57CF09186330230" colspan="1" rowspan="1" style="color: #231f20">30.331.9 (31.1 ± 1.1)</td>
<td id="73224C69CBB0C603DB7AE37D19D36DF5" colspan="1" rowspan="1" style="color: #231f20">27.334.1 (29.5 ± 1.8)</td>
<td id="C783A4F81982FA94BE1936F6BA4F0352" colspan="1" rowspan="1" style="color: #231f20">26.233.9 (29.6 ± 2.7)</td>
<td id="19867E8AF442070758AE4965D9877D4A" colspan="1" rowspan="1" style="color: #231f20">23.328.3 (25.6 ± 2.5)</td>
</tr>
<tr id="5759F432C9435103F1E11DB018AA105D">
<td id="33604EE881C80AA440AC0EFED34DC59C" colspan="1" rowspan="1" style="color: #231f20">Shell width (SW) (mm)</td>
<td id="E4FC6156B1B74D690F1E38FA46218B6A" colspan="1" rowspan="1" style="color: #231f20">12.615.9 (14.5 ± 1.0)</td>
<td id="1013B9CD13367334ECA362E9C7088E46" colspan="1" rowspan="1" style="color: #231f20">11.413.9 (12.1 ± 0.7)</td>
<td id="4542BC92C683EB4DB9F86E050EF51327" colspan="1" rowspan="1" style="color: #231f20">11.414.3 (13.1 ± 0.7)</td>
<td id="2ED8FE906A01FD2835BE66FDCFE525AE" colspan="1" rowspan="1" style="color: #231f20">10.814.6 (13.3 ± 1.7)</td>
<td id="93871F7E69D73F58FA62B5E4B1650B1F" colspan="1" rowspan="1" style="color: #231f20">10.514.2 (11.7 ± 1.0)</td>
<td id="F6D4524D505AC0E0DEC964B1050B845A" colspan="1" rowspan="1" style="color: #231f20">11.211.3 (11.3 ± 0.1)</td>
<td id="65C9E3BBB10AFDFA940EB3DFCDDD6937" colspan="1" rowspan="1" style="color: #231f20">8.411.1 (10.1 ± 0.6)</td>
<td id="7C0599E9C780109F415BF723A210BECA" colspan="1" rowspan="1" style="color: #231f20">8.812.2 (10.0 ± 0.9)</td>
<td id="841B5A415FFFA2A9AFD3F51D745F23BD" colspan="1" rowspan="1" style="color: #231f20">9.39.7 (9.5 ± 0.2)</td>
</tr>
<tr id="8554B135C3A80A22C559E293AF62A14D">
<td id="2569685D30F30355ECF00AE11BCDEE7C" colspan="1" rowspan="1" style="color: #231f20">Third whorl length (TWL) (mm)</td>
<td id="621725661BF6A86FF0A8669BF992F52F" colspan="1" rowspan="1" style="color: #231f20">5.16.8 (6.0 ± 0.5)</td>
<td id="5A28C2FAEEA5D3D0714FEE7C1B2AD209" colspan="1" rowspan="1" style="color: #231f20">4.45.9 (5.1 ± 0.5)</td>
<td id="4798A0950DC5ABE22820C9821DC0E580" colspan="1" rowspan="1" style="color: #231f20">5.06.8 (5.8 ± 0.5)</td>
<td id="447C3B72946984AB83FB9C6A3E656808" colspan="1" rowspan="1" style="color: #231f20">4.25.9 (5.4 ± 0.8)</td>
<td id="5B40F6A69DB98B4EB968738EDC9EEAD7" colspan="1" rowspan="1" style="color: #231f20">4.56.6 (5.2 ± 0.6)</td>
<td id="F4092579A842504D1819505A341815A0" colspan="1" rowspan="1" style="color: #231f20">4.8</td>
<td id="6574C091298D886E6771918F96C3DEE4" colspan="1" rowspan="1" style="color: #231f20">3.65.2 (4.5 ± 0.4)</td>
<td id="A0C8350FAFB156B78614161666FE40C7" colspan="1" rowspan="1" style="color: #231f20">4.36.0 (4.9 ± 0.4)</td>
<td id="546C6DE8826A410C0DBCE670ECBB2326" colspan="1" rowspan="1" style="color: #231f20">4.04.3 (4.2 ± 0.2)</td>
</tr>
<tr id="08E6905C77F6DEF3EAC075C3DD0BC94D">
<td id="31BFC69867C36B9B1DB08069E9E343C1" colspan="1" rowspan="1" style="color: #231f20">
Whorl elongation ratio (
<abbrev id="ABBRID0EACCI" xlink_title="whorl elongation ratio">WER</abbrev>
)
</td>
<td id="F4786368A25DA63EB55648A2D623FDF6" colspan="1" rowspan="1" style="color: #231f20">2.212.78 (2.5 ± 0.2)</td>
<td id="4253D802951DDFFCE326EDDF608E486B" colspan="1" rowspan="1" style="color: #231f20">2.192.75 (2.5 ± 0.2)</td>
<td id="B2102853ECEE755D1A9343243FA14A91" colspan="1" rowspan="1" style="color: #231f20">1.972.59 (2.3 ± 0.2)</td>
<td id="9B0CC9A63E4578A79455465029813611" colspan="1" rowspan="1" style="color: #231f20">2.372.73 (2.5 ± 0.2)</td>
<td id="F9A99B838F6619729C91CDE1833FBD23" colspan="1" rowspan="1" style="color: #231f20">2.212.98 (2.4 ± 0.2)</td>
<td id="E49DF8273A1F8942A626860D0ECDA9AC" colspan="1" rowspan="1" style="color: #231f20">2.282.79 (2.5 ± 0.4)</td>
<td id="45612A802B7F555D60AB478C5CF5E73D" colspan="1" rowspan="1" style="color: #231f20">2.092.76 (2.4 ± 0.2)</td>
<td id="353AAFF8FF9E5097D9219463041560C4" colspan="1" rowspan="1" style="color: #231f20">2.132.68 (2.4 ± 0.1)</td>
<td id="6A2AAFBA02615ACDC50B6486A6B0CB1A" colspan="1" rowspan="1" style="color: #231f20">2.442.51 (2.5 ± 0.1)</td>
</tr>
<tr id="136C5933E4964C89452018A9EB675F34">
<td id="F493ACE0127D59937E3B820A2AE97D1D" colspan="1" rowspan="1" style="color: #231f20">
Whorl number (
<abbrev id="ABBRID0EODCI" xlink_title="whorl number">WN</abbrev>
)
</td>
<td id="E92B640F0DEF630B0CC3A8938DF6C234" colspan="1" rowspan="1" style="color: #231f20">4.507.25 (5.3 ± 0.8)</td>
<td id="1EC1F429FFA7759D74469B45475C9F21" colspan="1" rowspan="1" style="color: #231f20">4.507.00 (5.6 ± 0.7)</td>
<td id="4CDD8A846EBA0FA7D09FF03C4823C067" colspan="1" rowspan="1" style="color: #231f20">5.007.50 (6.7 ± 0.9)</td>
<td id="F0A56417F23B37A71BC649F1E5DB8ACD" colspan="1" rowspan="1" style="color: #231f20">6.257.00 (6.7 ± 0.4)</td>
<td id="4347984044EC6F2876C02DB7876F8EFD" colspan="1" rowspan="1" style="color: #231f20">4.257.00 (5.9 ± 0.7)</td>
<td id="F984813FE4604CD91765B57E78AEB599" colspan="1" rowspan="1" style="color: #231f20">5</td>
<td id="D8BBE17E240825B9BBB378EFC9F6ACEE" colspan="1" rowspan="1" style="color: #231f20">3.508.00 (5.8 ± 1.0)</td>
<td id="18FB8BD2637DA0559C2CE05D3B8C99A4" colspan="1" rowspan="1" style="color: #231f20">3.508.75 (5.5 ± 1.5)</td>
<td id="D3E2093FE3CE488C6FBE874DFAC7E88F" colspan="1" rowspan="1" style="color: #231f20">4.005.75 (4.9 ± 0.1)</td>
</tr>
<tr id="D5662CD6AC6FAD3DC3283A7EB8A821A9">
<td id="4F34C42596656FDC18E23F3C57B6B5B6" colspan="1" rowspan="1" style="color: #231f20">Sculpture Type (node / granulated rib / smooth rib / spiral cord / smooth) (%)</td>
<td id="E7555BB7DA734BBF4FBB3242935478CE" colspan="1" rowspan="1" style="color: #231f20">87 / 0 / 0 / 13 / 0</td>
<td id="FDE9D19C979C5AEFDC1EFDA4D7572D67" colspan="1" rowspan="1" style="color: #231f20">47 / 53 / 0 / 0 / 0</td>
<td id="82B5FF24C1E117293C0D733D501F6612" colspan="1" rowspan="1" style="color: #231f20">80 / 20 / 0 / 0 / 0</td>
<td id="1063FF7F6C945FF737D174E7282CAC79" colspan="1" rowspan="1" style="color: #231f20">100 / 0 / 0 / 0 / 0</td>
<td id="323B551A40F91754CE48C990BE5670E0" colspan="1" rowspan="1" style="color: #231f20">40 / 60 / 0 / 0 / 0</td>
<td id="8C8E74CBB7C5DD6CD4A048910A5BD9D4" colspan="1" rowspan="1" style="color: #231f20">100 / 0 / 0 / 0 / 0</td>
<td id="09A9B0FF401E52B72D0C450B20EA4A76" colspan="1" rowspan="1" style="color: #231f20">27 / 47 / 27 / 0 / 0</td>
<td id="96CDB6701FEEC814E4D164B13C1F8AC8" colspan="1" rowspan="1" style="color: #231f20">64 / 29 / 7 / 0 / 0</td>
<td id="AACFB5482F1AE324A3CD3D06231AED4F" colspan="1" rowspan="1" style="color: #231f20">0 / 67 / 33 / 0 / 0</td>
</tr>
<tr id="28ED4EFF27DCDED88D6694AF9F488189">
<td id="0D9D20D66CB4D73B6A12B0DF274A4AC8" colspan="10" rowspan="1" style="color: #231f20">
<emphasis id="5B109E62FD52C014E9CE6F59E81507B0" bold="true">Morphological characters of mature male teleoconchs</emphasis>
</td>
</tr>
<tr id="1B81A324E154075089CA1CC60CE15F5E">
<td id="FBB5698D207F44BC05A22D6D605F9D82" colspan="1" rowspan="1" style="color: #231f20">Aperture height (AH) (mm)</td>
<td id="BE6794786373744600BEBE8BE0BC2E02" colspan="1" rowspan="1" style="color: #231f20">7.69.2 (8.4 ± 0.8)</td>
<td id="F421053C4FB54CEB7F08AC01BC63E3F1" colspan="1" rowspan="1" style="color: #231f20">7.79.1 (8.2 ± 0.5)</td>
<td id="F09C59BD27EEA80F5C4E2B74257820F2" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="AE7A270BB9A5F3C26542F3D839ECE5C6" colspan="1" rowspan="1" style="color: #231f20">8.08.6 (8.2 ± 0.3)</td>
<td id="1D67CF16EAD69550E061CD22A2E0A257" colspan="1" rowspan="1" style="color: #231f20">7.510.6 (9.1 ± 0.9)</td>
<td id="AB5BADC20DD7FDEB03F6AD2753095C2C" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="ECA255F73D6B95B08A1D8340B4173750" colspan="1" rowspan="1" style="color: #231f20">6.99.5 (8.3 ± 0.7)</td>
<td id="E10EDDE61C05B9437B06DA5A60A813E9" colspan="1" rowspan="1" style="color: #231f20">7.99.2 (8.3 ± 0.6)</td>
<td id="E78B688260C7ACA0D909D661A6F50A54" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="3CDCA6019FF78C41C6E9C41E235595A4">
<td id="36C5D28A9D4CC1D16649E97B1A647CFA" colspan="1" rowspan="1" style="color: #231f20">Aperture length (AL) (mm)</td>
<td id="E134122410FE2C51D65D0DB06A65B805" colspan="1" rowspan="1" style="color: #231f20">8.59.8 (9.0 ± 0.6)</td>
<td id="458AA48036C0279189B4194D583CF002" colspan="1" rowspan="1" style="color: #231f20">8.09.3 (8.5 ± 0.5)</td>
<td id="1017F2E2CE18BEE739BCA11A7543C170" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="BE2B20B91F3EAB439309CF9FA4F0A68C" colspan="1" rowspan="1" style="color: #231f20">8.38.9 (8.6 ± 0.3)</td>
<td id="B6768CF164536DE47339B370FCFC5E55" colspan="1" rowspan="1" style="color: #231f20">7.810.7 (9.2 ± 0.8)</td>
<td id="3EB388A39045B19FD3645990BB004CDE" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="E5C80E44B0B5D4CD25E6472A11466489" colspan="1" rowspan="1" style="color: #231f20">6.89.7 (8.4 ± 0.8)</td>
<td id="642A85AD4D8FBE5D3A89F3B4F8106266" colspan="1" rowspan="1" style="color: #231f20">7.79.3 (8.5 ± 0.6)</td>
<td id="83EA1691D6ED5B5CC49B629EB6EE2453" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="8EC1C13C7E001E1E6EA9BF959B077298">
<td id="95F8B461632CEB03FF93E88DEBDE9D25" colspan="1" rowspan="1" style="color: #231f20">
Aperture slenderness ratio (
<abbrev id="ABBRID0E6ICI" xlink_title="aperture slenderness ratio">ASR</abbrev>
)
</td>
<td id="CBBB91AED181A28AC7CD1BA084DF98C3" colspan="1" rowspan="1" style="color: #231f20">1.551.68 (1.6 ± 0.1)</td>
<td id="1CD74426D640D82E10D1F541424F6508" colspan="1" rowspan="1" style="color: #231f20">1.341.76 (1.6 ± 0.2)</td>
<td id="F48E56E27EE73C1ED4BD1172AFABA85C" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="B710816CCFE18BC419020DA6161A0FB0" colspan="1" rowspan="1" style="color: #231f20">1.471.69 (1.6 ± 0.1)</td>
<td id="816C1D9E06209B267B63EC8964A61860" colspan="1" rowspan="1" style="color: #231f20">1.451.69 (1.6 ± 0.1)</td>
<td id="7839FFC4814FD4C587EA7FB93B457111" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="D1E922C674214F18F8FA7A99BB28F515" colspan="1" rowspan="1" style="color: #231f20">1.451.91 (1.7 ± 0.1)</td>
<td id="B94A2D68A8D49B047AB9DE42704D41B9" colspan="1" rowspan="1" style="color: #231f20">1.741.81 (1.8 ± 0.1)</td>
<td id="F735F9CB06D191232E943D35C7069D9D" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="615FC77606A214061D2FE2F6AB5BDA79">
<td id="53C34D3299F79B6179F99A02242FE622" colspan="1" rowspan="1" style="color: #231f20">Aperture width (AW) (mm)</td>
<td id="50257BC0C5077DE1F1A7918CD449593A" colspan="1" rowspan="1" style="color: #231f20">5.35.8 (5.5 ± 0.3)</td>
<td id="0F927FD4B228F7A0E24D287CD0F53245" colspan="1" rowspan="1" style="color: #231f20">4.66.0 (5.2 ± 0.6)</td>
<td id="5F94082D38DB91007E615009D0A152A6" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="2433FD6D0D96D708761E855126D8329E" colspan="1" rowspan="1" style="color: #231f20">5.35.7 (5.5 ± 0.2)</td>
<td id="9FF20178882DA715689C84113A8FBAEF" colspan="1" rowspan="1" style="color: #231f20">4.66.6 (5.8 ± 0.6)</td>
<td id="665DDBA12E914C6851CC6A400F7EB850" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="5760BCBEB16E07A4C4786D7E82091110" colspan="1" rowspan="1" style="color: #231f20">4.35.9 (5.0 ± 0.5)</td>
<td id="9794D123D02F7A1A495C20FEDD2739FA" colspan="1" rowspan="1" style="color: #231f20">4.45.3 (4.8 ± 0.4)</td>
<td id="59A3A3E1D07B5CE79BCB3166CDCDA2AC" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="32F294825B67BB52D2F0F0D169FE6AC9">
<td id="A3785CF8C615694BFA7F16EEBDD78936" colspan="1" rowspan="1" style="color: #231f20">
Basal cord number (
<abbrev id="ABBRID0EWLCI" xlink_title="basal cord number">BCN</abbrev>
)
</td>
<td id="5F8DD027F6B1FCF5732D3E1D8037FEF9" colspan="1" rowspan="1" style="color: #231f20">45 (4.3 ± 0.6)</td>
<td id="27CD5E8D298E6DE8026235F1A2D96EDE" colspan="1" rowspan="1" style="color: #231f20">24 (3.4 ± 0.9)</td>
<td id="CA7730896CBBA15180E728E4D8FC97CC" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="8121BB8928C86A698CC0C9619F0D31E2" colspan="1" rowspan="1" style="color: #231f20">23 (2.8 ± 0.5)</td>
<td id="FF83366050856794F69312740EFD195E" colspan="1" rowspan="1" style="color: #231f20">24 (2.9 ± 0.7)</td>
<td id="32789FB21EFC74D7D455808266AE53BA" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="B15FA90D6CD71701C0B5E0CE3DE7E6F5" colspan="1" rowspan="1" style="color: #231f20">26 (3.4 ± 1.0)</td>
<td id="F1FF6DF6B55FE88A635F1B816C3B9A70" colspan="1" rowspan="1" style="color: #231f20">3</td>
<td id="1627D3C49C256C8AE7762221798CFCE6" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="7C4C9EF90055BFE06939DD6120790C91">
<td id="0E847B2F19A3C714DD5AA45A41577D96" colspan="1" rowspan="1" style="color: #231f20">
Body whorl length (
<abbrev id="ABBRID0EENCI" xlink_title="body whorl length">BWL</abbrev>
) (mm)
</td>
<td id="17BB6C4265DFEFA8C5CBD10E33856F26" colspan="1" rowspan="1" style="color: #231f20">13.816.2 (14.6 ± 1.4)</td>
<td id="513B803DAD607408C7ECEC1993AB13E3" colspan="1" rowspan="1" style="color: #231f20">13.015.3 (14.0 ± 0.9)</td>
<td id="0BB76561BCE94EAFCD5DB76B66B09BCD" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="185B4C42C21D123CF2078C017FF3631D" colspan="1" rowspan="1" style="color: #231f20">13.914.4 (14.1 ± 0.2)</td>
<td id="6EB637917679B903E6B6FADCBF466C96" colspan="1" rowspan="1" style="color: #231f20">12.918.3 (15.5 ± 1.5)</td>
<td id="C55F7CFD42900751C7DD33EBCAEC5E29" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="6DBCF15C7F9F5F7EFA9CD0E62D19A0CC" colspan="1" rowspan="1" style="color: #231f20">11.116.1 (13.7 ± 1.2)</td>
<td id="17786DDCD7332DAFF851F858EE54C66C" colspan="1" rowspan="1" style="color: #231f20">1315.7 (14.1 ± 1.0)</td>
<td id="430EA388DD857D8F96BF04C345B1ADC4" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="37BD25654B628A3F4A7475207CC6752E">
<td id="612B015D85536B65EE030A51805BE35D" colspan="1" rowspan="1" style="color: #231f20">Fourth whorl length (FWL) (mm)</td>
<td id="676D08A8E3EE8E3CD4250E650152474D" colspan="1" rowspan="1" style="color: #231f20">2.73.0 (2.9 ± 0.2)</td>
<td id="EA73B74F8CD988BF664C7B652A26615A" colspan="1" rowspan="1" style="color: #231f20">2.03.7 (3.0 ± 0.6)</td>
<td id="3B62BA3A649C541E0B2096C8E83CACC5" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="A9BF5B59BB4F1BFE77B4F8C54831B891" colspan="1" rowspan="1" style="color: #231f20">3.63.8 (3.7 ± 0.1)</td>
<td id="562BD4CD7364543A3F4976D82065A4C4" colspan="1" rowspan="1" style="color: #231f20">3.34.2 (3.7 ± 0.3)</td>
<td id="147CBC72132866EE555C8A2EB09ED587" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="0D4FC35DF1A206CA87F1FDADCEB99AB6" colspan="1" rowspan="1" style="color: #231f20">2.53.8 (3.2 ± 0.4)</td>
<td id="80BB7A9FD44FAC61BA353285B1A51C52" colspan="1" rowspan="1" style="color: #231f20">3.13.9 (3.4 ± 0.3)</td>
<td id="B4A48E1826EAC9DA502D01292AD5E555" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="8093E9764DDC8EBFB2512CE625231A08">
<td id="6D6BC547D86A2E935D6D41DF9A973E9D" colspan="1" rowspan="1" style="color: #231f20">Penultimate whorl length (PWL) (mm)</td>
<td id="8E1BB391073B839D02385FB783ECF96B" colspan="1" rowspan="1" style="color: #231f20">4.66.2 (5.4 ± 0.8)</td>
<td id="2CB0BB7EC7A260645188313828760A43" colspan="1" rowspan="1" style="color: #231f20">4.75.9 (5.3 ± 0.6)</td>
<td id="6E44F2046B112BDCA352F915419C9AFA" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="07F91AB9DF49267BC500D97F8E8DE305" colspan="1" rowspan="1" style="color: #231f20">5.05.8 (5.5 ± 0.4)</td>
<td id="38D2B99D446B936FDDEFD47B0CEFAC2F" colspan="1" rowspan="1" style="color: #231f20">4.97.1 (6.0 ± 0.7)</td>
<td id="3D7808A669F71E9DA10A98A65308F957" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="26408DC69BFE31ADE7DF439835FA66EA" colspan="1" rowspan="1" style="color: #231f20">4.06.1 (4.9 ± 0.6)</td>
<td id="969A8275C75894184A2C5CF090ACF18B" colspan="1" rowspan="1" style="color: #231f20">4.75.9 (5.2 ± 0.4)</td>
<td id="61EF96261DCA074593C10BFC651B2B73" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="E12F682CCD863F7977D88D9D4E889030">
<td id="DF84BBECF0ACABB83D107FDAD3F98193" colspan="1" rowspan="1" style="color: #231f20">
Longitudinal rib number of penultimate whorl (
<abbrev id="ABBRID0EERCI" xlink_title="longitudinal rib number of penultimate whorl">RN</abbrev>
)
</td>
<td id="742CFA21E79A5C89E4004A07867A6E17" colspan="1" rowspan="1" style="color: #231f20">2025 (23.0 ± 2.6)</td>
<td id="ADCC3E55B323481442B2D455CD0E5FEA" colspan="1" rowspan="1" style="color: #231f20">1924 (21.4 ± 2.5)</td>
<td id="D6E5D2CF231F516E2D83F0FD4F1CBBF8" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="2F3212DBE0019034F997237DB1785278" colspan="1" rowspan="1" style="color: #231f20">2021 (20.7 ± 0.6)</td>
<td id="EF57E8C2802245345D03B131A87907D8" colspan="1" rowspan="1" style="color: #231f20">1623 (18.8 ± 2.3)</td>
<td id="119DD3415F1A44193AA834F9197BE463" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="1764C5BB49FF5A6431B1DE36F0015A00" colspan="1" rowspan="1" style="color: #231f20">1523 (18.3 ± 2.3)</td>
<td id="7A69C267356B3A6F2F72381090DE872A" colspan="1" rowspan="1" style="color: #231f20">1421 (16.6 ± 2.7)</td>
<td id="D28A53A36C7069B5CC6FF0CD9114616F" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="24EE4345F814FFDA18535277940E034E">
<td id="7C156E0FE76488C3F0E25AEE3386AFC6" colspan="1" rowspan="1" style="color: #231f20">
Spire angle (
<abbrev id="ABBRID0ESSCI" xlink_title="spire angle">SA</abbrev>
) (degrees)
</td>
<td id="484CC1B10EA1E61DDA8399C446B0E05F" colspan="1" rowspan="1" style="color: #231f20">21.524.8 (23.5 ± 1.7)</td>
<td id="24E9C139747CA283B0AE0C93EDFB511D" colspan="1" rowspan="1" style="color: #231f20">16.219.3 (17.7 ± 1.5)</td>
<td id="63563F4037C93F13EEDA487CB49E8DCB" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="D7DB4F728B22701C959A81C31E137DB7" colspan="1" rowspan="1" style="color: #231f20">14.819.5 (16.7 ± 2.0)</td>
<td id="0B33267368CAD6208D405EB9F7DCD741" colspan="1" rowspan="1" style="color: #231f20">12.118.4 (16.1 ± 1.9)</td>
<td id="DC2A14653D18376603C1AE24574C29D8" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="99BD29EA060816E1B19D17E2899F4050" colspan="1" rowspan="1" style="color: #231f20">14.623.1 (19.1 ± 2.2)</td>
<td id="562E55264ADC82C57FEBF612F85890C7" colspan="1" rowspan="1" style="color: #231f20">15.420.3 (17.4 ± 2.1)</td>
<td id="52BA8F9FAA080EA037BEE638249A4672" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="FBDAF8951840FF41E8841783423A906C">
<td id="EBF72241847C32433A1CB67D9AB02643" colspan="1" rowspan="1" style="color: #231f20">
Spiral cord number of penultimate whorl (
<abbrev id="ABBRID0EAUCI" xlink_title="spiral cord number of penultimate whorl">SCN</abbrev>
)
</td>
<td id="8308753B13758E6B1B20F5C0C8DA5FDA" colspan="1" rowspan="1" style="color: #231f20">56 (5.7 ± 0.6)</td>
<td id="F5811C12F1A368BCAFD6736E08A1ED78" colspan="1" rowspan="1" style="color: #231f20">56 (5.4 ± 0.5)</td>
<td id="B546338779AECBF754014609F2AB7176" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="B8591C3A1CD3C751446E52E62E4A97D9" colspan="1" rowspan="1" style="color: #231f20">46 (4.8 ± 1.0)</td>
<td id="6F184B8A0A1C9277AA35E5A1F9E43E56" colspan="1" rowspan="1" style="color: #231f20">56 (5.8 ± 0.4)</td>
<td id="50B1DC0B1F0971DB9B5A8E4BFB34E1E6" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="C8EDA0565D7DC692CC753AEE20F2CC5F" colspan="1" rowspan="1" style="color: #231f20">57 (5.9 ± 0.6)</td>
<td id="02CE29500DA1BDEF3D97B6F74FFA34FD" colspan="1" rowspan="1" style="color: #231f20">56 (5.8 ± 0.5)</td>
<td id="7B69E91570417EC3BD8C1D3EB7D8165B" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="38E9213E1FDC85D1846CDD3B85061066">
<td id="4727496EE0CEFD32F75717A4B8DBBB61" colspan="1" rowspan="1" style="color: #231f20">Shell length (SL) (mm)</td>
<td id="3DD9BF78DB6335319968962E138E2939" colspan="1" rowspan="1" style="color: #231f20">23.227.2 (24.8 ± 2.1)</td>
<td id="25435B72A5D6AC017914ABD48161D0FE" colspan="1" rowspan="1" style="color: #231f20">22.731.6 (26.6 ± 3.4)</td>
<td id="0381CA0D288A8DFD4ACFF829B8B82616" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="240ABD0416100F7B0AE2EEB0CA4C8F65" colspan="1" rowspan="1" style="color: #231f20">28.731.1 (29.8 ± 1.1)</td>
<td id="892F0F33260A8BFB824862E6D59728C7" colspan="1" rowspan="1" style="color: #231f20">25.334.0 (29.7 ± 3.2)</td>
<td id="87A1D107D682E1C9D8EEB809E3599EE4" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="41A7BDF2198D6B8E3E580D56ABB63A3A" colspan="1" rowspan="1" style="color: #231f20">20.530.2 (25.9 ± 2.7)</td>
<td id="B4657330AA050C8CB97BCDB69C588D5D" colspan="1" rowspan="1" style="color: #231f20">24.128.6 (25.8 ± 1.8)</td>
<td id="BBC0A4FA790419713A9CF8ADA7183A56" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="74412A8BA4375299A7BAE6C3AE51A328">
<td id="720898031CB39F842FBE7A7E4F3129F1" colspan="1" rowspan="1" style="color: #231f20">Shell width (SW) (mm)</td>
<td id="680FC8C6FCD4590D3904C12705655B6B" colspan="1" rowspan="1" style="color: #231f20">10.111.2 (10.6 ± 0.5)</td>
<td id="FCB0D8E01698E437EB2B5B6E8173BAD6" colspan="1" rowspan="1" style="color: #231f20">8.610.2 (9.5 ± 0.7)</td>
<td id="D3D9E3F9CAE067038980872D28447241" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="75F4A779F81AC574B99534904ADDC58F" colspan="1" rowspan="1" style="color: #231f20">10.011.0 (10.6 ± 0.5)</td>
<td id="4A01FB675A1603492E7510E63294D913" colspan="1" rowspan="1" style="color: #231f20">8.511.3 (10.2 ± 0.9)</td>
<td id="E13070CDFB47D0D7E85E64AAAAA0FE23" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="0A3EC90C51385056584A81EC8D1B2421" colspan="1" rowspan="1" style="color: #231f20">7.810.2 (8.8 ± 0.7)</td>
<td id="7F43CD90B906D0F8E704AD03594AE446" colspan="1" rowspan="1" style="color: #231f20">8.210.2 (9.2 ± 0.8)</td>
<td id="9DF9F44C5FBE1FAA35AC5219DCFA96B1" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="A227A4DC02C4044F24DC6050F127C185">
<td id="981C18C669B9276D8017AB011D213819" colspan="1" rowspan="1" style="color: #231f20">Third whorl length (TWL) (mm)</td>
<td id="F8130AE3660779CB26223428FCDF8057" colspan="1" rowspan="1" style="color: #231f20">3.24.3 (3.7 ± 0.5)</td>
<td id="6AEDE723C2FB7906BCB2459F167451D5" colspan="1" rowspan="1" style="color: #231f20">3.64.1 (3.9 ± 0.2)</td>
<td id="4D0FC139C0548ECE267BDDF17493E7FD" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="95F37FCAAEF47BFE41BEB982F0382443" colspan="1" rowspan="1" style="color: #231f20">4.35.1 (4.5 ± 0.4)</td>
<td id="C4CC8BEE7F7B5BEC1D05E8079D53095A" colspan="1" rowspan="1" style="color: #231f20">4.15.4 (4.7 ± 0.4)</td>
<td id="6356BC326FE1153A86AF187C33B8BD51" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="7468BC0389A33CC4C1522989A40A1505" colspan="1" rowspan="1" style="color: #231f20">3.44.5 (4.0 ± 0.4)</td>
<td id="85EBA8D5E3B9E87367AF37BBC5BE35CF" colspan="1" rowspan="1" style="color: #231f20">3.64.5 (4.0 ± 0.3)</td>
<td id="AA7624D4D89049C5EE9DDC6698956182" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="E7C1B0C32A55C2D405664EAD053F590C">
<td id="02E1894455A7C21E6635E1202CF2237F" colspan="1" rowspan="1" style="color: #231f20">
Whorl elongation ratio (
<abbrev id="ABBRID0EJZCI" xlink_title="whorl elongation ratio">WER</abbrev>
)
</td>
<td id="FF2EBE0D8B1D540DDE82C09CAF6BBB8E" colspan="1" rowspan="1" style="color: #231f20">2.503.15 (2.9 ± 0.4)</td>
<td id="E54503913F939962AF7E1C353DD2236E" colspan="1" rowspan="1" style="color: #231f20">2.184.05 (2.8 ± 0.7)</td>
<td id="907234393F1DE0892FF2EAA4821778B8" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="25B7B58B071BF1BAA0DAB52AB955E56E" colspan="1" rowspan="1" style="color: #231f20">2.112.41 (2.2 ± 0.1)</td>
<td id="A566D8A2A1E68F1944A2B318E0F51356" colspan="1" rowspan="1" style="color: #231f20">2.282.65 (2.5 ± 0.1)</td>
<td id="F5024D1295D97E1DB4B4E041BFA21199" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="F58F1D21018A7B167AC85E67517D9B22" colspan="1" rowspan="1" style="color: #231f20">2.183.05 (2.6 ± 0.3)</td>
<td id="BDBEA28010667B8F405824CD4D630E80" colspan="1" rowspan="1" style="color: #231f20">2.192.66 (2.5 ± 0.2)</td>
<td id="C6B32C579A09B08761057DE740AE130E" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="4BAAE54E8AC5B8A923D39552E5B00000">
<td id="AD2672CEA653643C46F03A9893B01C94" colspan="1" rowspan="1" style="color: #231f20">
Whorl number (
<abbrev id="ABBRID0EX1CI" xlink_title="whorl number">WN</abbrev>
)
</td>
<td id="70E0C26A2F0949D87AA6A984D4DE1466" colspan="1" rowspan="1" style="color: #231f20">5.005.25 (5.1 ± 0.1)</td>
<td id="A7C3AB6E200DB8ADEEB83068A4E4FDC9" colspan="1" rowspan="1" style="color: #231f20">4.508.00 (6.0 ± 1.4)</td>
<td id="50673CF8933C57A7C78465B2CD54A4C9" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="E5B2624F8FD4E4A8E9ABC23D97F58F00" colspan="1" rowspan="1" style="color: #231f20">6.008.00 (6.8 ± 0.9)</td>
<td id="E0E82FE72CC8E0EA379EF52DDEE29142" colspan="1" rowspan="1" style="color: #231f20">4.506.25 (5.2 ± 0.6)</td>
<td id="0E6327BDEF7B16676AA74418CA2BD71B" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="F2754116079E37DD49F9957C989EE619" colspan="1" rowspan="1" style="color: #231f20">4.258.00 (5.8 ± 1.2)</td>
<td id="46CB0EA894764CCEB3942ACDE3412BA4" colspan="1" rowspan="1" style="color: #231f20">4.255.25 (4.8 ± 0.4)</td>
<td id="CE2C793BA2F1506CA3274C37274C2599" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="660B4C86F8E1CD3743EFF09CD674EA5D">
<td id="2B3DB56CFF61B0038ABFA830A0AEB4BF" colspan="1" rowspan="1" style="color: #231f20">Sculpture Type (node / granulated rib / smooth rib / spiral cord / smooth) (%)</td>
<td id="345CDC48AEB9AC4100920CEB3AF00A45" colspan="1" rowspan="1" style="color: #231f20">33 / 67 / 0 / 0 / 0</td>
<td id="FA11DAC90ED596D1ED67BA888F2BA5B0" colspan="1" rowspan="1" style="color: #231f20">60 / 40 / 0 / 0 / 0</td>
<td id="7038B89CCF02FE4833C171F2E277045A" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="E55A765391D56E727329012ADCF544D3" colspan="1" rowspan="1" style="color: #231f20">100 / 0 / 0 / 0 / 0</td>
<td id="BA29EC8FFAE6D80785C8677C3D64B7BA" colspan="1" rowspan="1" style="color: #231f20">50 / 42 / 8 / 0 / 0</td>
<td id="95CED46994CCF2CDB064EAE3EB7E13B1" colspan="1" rowspan="1" style="color: #231f20"></td>
<td id="4AD1AA01EE3A585ADDE90455BEA71DF8" colspan="1" rowspan="1" style="color: #231f20">23 / 62 / 15 / 0 / 0</td>
<td id="152BC0B8CB7224EAC7E115130864F41C" colspan="1" rowspan="1" style="color: #231f20">60 / 20 / 20 / 0 / 0</td>
<td id="CCCE469B0B5288F7937171CF057A676F" colspan="1" rowspan="1" style="color: #231f20"></td>
</tr>
<tr id="597E7D3634604FE29858E872578D2B73">
<td id="512D7F8D5BBF0C565E5808CF49493F3C" colspan="1" rowspan="1" style="color: #231f20">Morphological characters of protoconchs</td>
<td id="A0F72E68C6A64FC272E378566F6B7BBF" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="03ABFC3F6C9077F3C73014A23BD19B7F" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="FB47B6659F408F79D70E6F9D3FA78EE1" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="E36202A6F04ABCAE0F743C54FF39D7CA" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="75D99ACB6010B1B577BDC71344868FE4" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="3EB0F5F3A9D08B39A6D3944BE9FF8475" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="87604E6BD8EC47DCB41D3FE3ECDDCD26" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="8149E317117D945DB9104D009B3DAC3E" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="FC977AAE294438E0999A8EC01C0719B1" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="19C272A76E896D74E6C8CAB6BE269CD9">
<td id="5832C11D68D0D6B96FA917BA42B560E8" colspan="1" rowspan="1" style="color: #231f20">
Number of protoconchs (
<abbrev id="ABBRID0EO5CI" xlink_title="number of protoconchs">PN</abbrev>
)
</td>
<td id="5EF00486DEFF8C504EFBF074367F2071" colspan="1" rowspan="1" style="color: #231f20">417 (9.3 ± 4.8)</td>
<td id="38B7A62EB5CC6AE41F9332AF3FB32880" colspan="1" rowspan="1" style="color: #231f20">114 (7.3 ± 4.2)</td>
<td id="4EF97AFBC9C83457BB81E8E6F5A1056E" colspan="1" rowspan="1" style="color: #231f20">314 (7.7 ± 3.8)</td>
<td id="045A38A7683142340AF3724FE352ADD2" colspan="1" rowspan="1" style="color: #231f20">24 (3.0 ± 1.0)</td>
<td id="B94AB78E5049B40228462BDD1609B682" colspan="1" rowspan="1" style="color: #231f20">26102 (57.4 ± 18.9)</td>
<td id="4E17C5EC93246B27BC7B07DA7D721D54" colspan="1" rowspan="1" style="color: #231f20">5697 (76.5 ± 29.0)</td>
<td id="4B7127295D0FFAB2FACB2FB72A134F3B" colspan="1" rowspan="1" style="color: #231f20">552 (23.3 ± 13.8)</td>
<td id="D4356BCB905357BB61D79A9F9A71DEB9" colspan="1" rowspan="1" style="color: #231f20">543 (21.3 ± 9.4)</td>
<td id="8D73056FF9FDF7CE2706CDF929018642" colspan="1" rowspan="1" style="color: #231f20">424 (14.0 ± 14.1)</td>
</tr>
<tr id="3F4F0B59144A9CF820731812D0BE19B8">
<td id="27B4363B35D1DC054AC962078D847239" colspan="1" rowspan="1" style="color: #231f20">
Longitudinal rib number on body whorl of the largest protoconch (
<abbrev id="ABBRID0E36CI" xlink_title="longitudinal rib number on body whorl of the largest protoconch">RNP</abbrev>
)
</td>
<td id="7B0A008747C3E86B811E0DE5F4F43EF4" colspan="1" rowspan="1" style="color: #231f20">1115 (13.3 ± 1.6)</td>
<td id="C346FAAC53AC7B5013F4A6B8EEEB061A" colspan="1" rowspan="1" style="color: #231f20">1219 (15.1 ± 2.4)</td>
<td id="111B28B8DC4C3CD05CB0547DEE7C621B" colspan="1" rowspan="1" style="color: #231f20">1215 (13.6 ± 1.0)</td>
<td id="61E028578E8799D49B6318956A62FF2B" colspan="1" rowspan="1" style="color: #231f20">1015 (12.7 ± 2.5)</td>
<td id="9FC4BBB0DDDBD84CAA0C747CD62091C9" colspan="1" rowspan="1" style="color: #231f20">1116 (12.9 ± 1.4)</td>
<td id="D049AD6435782B5EFC3A374A87D1B5D8" colspan="1" rowspan="1" style="color: #231f20">1113 (12.0 ± 1.4)</td>
<td id="E37FEE1E7A1290B98A1BF80A6BB16493" colspan="1" rowspan="1" style="color: #231f20">1116 (12.6 ± 1.4)</td>
<td id="19D9BAE04C4EB14B5ECF079A8FBBA1DB" colspan="1" rowspan="1" style="color: #231f20">915 (11.7 ± 1.7)</td>
<td id="3EBBBBEE3052272E225B43262795BD4D" colspan="1" rowspan="1" style="color: #231f20">1112 (11.5 ± 0.7)</td>
</tr>
<tr id="793CE96D52B0326D4986BC3C51389F05">
<td id="C2A060F1315AB009271157A2F57B0C20" colspan="1" rowspan="1" style="color: #231f20">
Shell length of the largest protoconch (
<abbrev id="ABBRID0EKBDI" xlink_title="shell length of the largest protoconch">SLP</abbrev>
) (mm)
</td>
<td id="4BEB8BE1D50AE28841AD96FB7402EACB" colspan="1" rowspan="1" style="color: #231f20">4.05.9 (5.2 ± 0.5)</td>
<td id="D82BFE7115419B95F8D170DD7B44A5C4" colspan="1" rowspan="1" style="color: #231f20">2.56.6 (4.6 ± 1.1)</td>
<td id="C4132634A2C3F9741843E5F5266624CB" colspan="1" rowspan="1" style="color: #231f20">4.77.5 (5.9 ± 1.0)</td>
<td id="8FB149E63F2366A5B9A0C9DA8545926F" colspan="1" rowspan="1" style="color: #231f20">2.74.5 (3.9 ± 1.0)</td>
<td id="F001BB6999E9AA9142C90D4210616C69" colspan="1" rowspan="1" style="color: #231f20">2.12.9 (2.5 ± 0.2)</td>
<td id="BC4C0C2BFD8D212FAC7F5003DC910B14" colspan="1" rowspan="1" style="color: #231f20">2.63.1 (2.8 ± 0.4)</td>
<td id="4DA115146A1925938F66D5F703B5DD42" colspan="1" rowspan="1" style="color: #231f20">1.82.5 (2.3 ± 0.2)</td>
<td id="AB43B0A777716D12709D07E534962D46" colspan="1" rowspan="1" style="color: #231f20">2.02.7 (2.3 ± 0.2)</td>
<td id="BE08CD8CE86DCF2CCDB1F5E3D2C6A32F" colspan="1" rowspan="1" style="color: #231f20">2.32.9 (2.6 ± 0.4)</td>
</tr>
<tr id="18304AB388EAD77074B945F9EDF55F63">
<td id="2DC92B6B8177EB7EF1B4FC3C30192499" colspan="1" rowspan="1" style="color: #231f20">Shell width of the largest protoconch (SWP) (mm)</td>
<td id="ADA6BAA3FD1581AD5839FA99123C0DBA" colspan="1" rowspan="1" style="color: #231f20">3.44.2 (3.8 ± 0.2)</td>
<td id="06DA94A516BC7790F6814D17EFDB953C" colspan="1" rowspan="1" style="color: #231f20">2.13.9 (3.2 ± 0.5)</td>
<td id="9CA6FBCD60F774C2EF364BD17C1B8E74" colspan="1" rowspan="1" style="color: #231f20">3.44.3 (3.8 ± 0.3)</td>
<td id="2A93C4DD634E119F47BE551255EDEF9E" colspan="1" rowspan="1" style="color: #231f20">2.83.7 (3.3 ± 0.5)</td>
<td id="D083B02A5A112F981EA0937685185461" colspan="1" rowspan="1" style="color: #231f20">1.52.1 (1.8 ± 0.2)</td>
<td id="23E1E32BBD38AF03720C6919927988B3" colspan="1" rowspan="1" style="color: #231f20">1.81.9 (1.9 ± 0.1)</td>
<td id="E88C2AB2D34640ADC6C5F6D9C383E4D9" colspan="1" rowspan="1" style="color: #231f20">1.61.9 (1.8 ± 0.1)</td>
<td id="17531ADC3727BA7EFB3D2785D89D6CF9" colspan="1" rowspan="1" style="color: #231f20">1.51.9 (1.7 ± 0.1)</td>
<td id="6E65FC2B17042D21EBFB34035A489115" colspan="1" rowspan="1" style="color: #231f20">1.8</td>
</tr>
<tr id="7558A4CA6E003FC20A2701BC1F1C948D">
<td id="F55FD1FBEB7156133B872F8C50F12795" colspan="1" rowspan="1" style="color: #231f20">
Whorl number of the largest protoconch (
<abbrev id="ABBRID0EBEDI" xlink_title="whorl number of the largest protoconch">WNP</abbrev>
)
</td>
<td id="ED252EA6AEDD798714665088A8DB5939" colspan="1" rowspan="1" style="color: #231f20">3.004.15 (3.8 ± 0.4)</td>
<td id="0362DF1816326E7376F5AE27EECCB1E4" colspan="1" rowspan="1" style="color: #231f20">2.504.50 (3.8 ± 0.5)</td>
<td id="E9DAF1D0915BC045A7B68066FA8B15BC" colspan="1" rowspan="1" style="color: #231f20">3.505.00 (4.1 ± 0.5)</td>
<td id="E97FD6094D9EB3218892C0A8BCEE24B0" colspan="1" rowspan="1" style="color: #231f20">2.503.50 (3.1 ± 0.5)</td>
<td id="58E16D7546EA09CA765AEDDE0C67179C" colspan="1" rowspan="1" style="color: #231f20">2.754.00 (3.3 ± 0.3)</td>
<td id="209CE5A4B5E2AE0D1BEDA07222F4EDD9" colspan="1" rowspan="1" style="color: #231f20">3.503.75 (3.6 ± 0.2)</td>
<td id="CE1F3192EF95521F964C11454AC3A1A9" colspan="1" rowspan="1" style="color: #231f20">2.253.50 (3.1 ± 0.3)</td>
<td id="9804F0542E151C666B83D022D706303A" colspan="1" rowspan="1" style="color: #231f20">2.753.50 (3.1 ± 0.2)</td>
<td id="D056D7543254CBD63FE17A4A83FC961E" colspan="1" rowspan="1" style="color: #231f20">2.753.50 (3.1 ± 0.5)</td>
</tr>
<tr id="402FC2BD9673A57030DB810A4118D856">
<td id="60D13335AB6C9EF817FA00BC07D503F8" colspan="1" rowspan="1" style="color: #231f20">Number of nodes on body whorl of the largest protoconch (Node Number) (0 / 1 / 2 or 3) (%)</td>
<td id="32D9037C91105430180B25C050C3C676" colspan="1" rowspan="1" style="color: #231f20">100 / 0 / 0</td>
<td id="D1251EDFE19D33852EA07A64327CF3AF" colspan="1" rowspan="1" style="color: #231f20">100 / 0 / 0</td>
<td id="B6E23AF0AA4F8232195561AFD25C1E24" colspan="1" rowspan="1" style="color: #231f20">93 / 7 / 0</td>
<td id="231669BBBD3EB17FDF75CB7BD06F6539" colspan="1" rowspan="1" style="color: #231f20">0 / 33 / 67</td>
<td id="7CDD6557C34A9B9B60588FF24BF07012" colspan="1" rowspan="1" style="color: #231f20">7 / 93 / 0</td>
<td id="A0ECEE15458A741124B4D9580D2A465F" colspan="1" rowspan="1" style="color: #231f20">50 / 50 / 0</td>
<td id="BDCFC4A043A5159FE45C91A7BD2E6744" colspan="1" rowspan="1" style="color: #231f20">20 / 73 / 7</td>
<td id="813914F2426605DF7EF947C44D45415B" colspan="1" rowspan="1" style="color: #231f20">0 / 36 / 64</td>
<td id="D1D2275B95496EE2A3B0FED6C5A79006" colspan="1" rowspan="1" style="color: #231f20">50 / 50 / 0</td>
</tr>
<tr id="32205E0A04BDF0F4FCA9CAC8C3E19028">
<td id="7E4118CE47DF5097EA02B06BBF9D62A5" colspan="1" rowspan="1" style="color: #231f20">Keel type on body whorl of the largest protoconch (Keel Type) (absent / weak / prominent) (%)</td>
<td id="EBF720E330B30373E615F6095EE85AD0" colspan="1" rowspan="1" style="color: #231f20">73 / 27 / 0</td>
<td id="59096A61B2123CF2CF943ACF6B5A355B" colspan="1" rowspan="1" style="color: #231f20">80 / 20 / 0</td>
<td id="C5C9D8C5CFDADF4C626101E5AD971323" colspan="1" rowspan="1" style="color: #231f20">50 / 50 / 0</td>
<td id="5FC02A12E1A6315B3C7E3A898A34A704" colspan="1" rowspan="1" style="color: #231f20">100 / 0 / 0</td>
<td id="6FB79ECD883B0C64E7F38A7F017B4886" colspan="1" rowspan="1" style="color: #231f20">0 / 20 / 80</td>
<td id="7DA621938811BA2CF5E05FCA5B65A2AB" colspan="1" rowspan="1" style="color: #231f20">0 / 50 / 50</td>
<td id="6F1B28CAA137A0A8A9E223D87AECD288" colspan="1" rowspan="1" style="color: #231f20">0 / 27 / 73</td>
<td id="63DD2DEA750EB3813CD7D3153B94A3DC" colspan="1" rowspan="1" style="color: #231f20">0 / 29 / 71</td>
<td id="78945412C2F538FCF1A0DEBA99DA806F" colspan="1" rowspan="1" style="color: #231f20">0 / 50 / 50</td>
</tr>
<tr id="AA56084D2059D798A66CBA4B17020C0E">
<td id="1421EE6A02847624442AF33EA8494F71" colspan="1" rowspan="1" style="color: #231f20">Morphological characters of radulae</td>
<td id="7FE32E1723F2806823D38CACC88CC611" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="C1FD4E78B60CE505A7EAC52B60295C34" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="DF872705A764DBAAD5C34895A1DCE766" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="03C7607CE60301DB6076C5841C47BC8B" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="A51A5261B5434E944FF9314887EF1C90" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="59C57315035BB848701403E5AED89274" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="AAA675507F956AD3F8ECB7A9CA355BD5" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="98CCE28CD1CC61A4AA127794C313F74E" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="9446E245C1F0A49F8F519FF335AAB468" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="8385DD177434D1A6A3FE6DC0EDC6B0CD">
<td id="FD917C457439FB3E73CB11A45CB447F7" colspan="1" rowspan="1" style="color: #231f20">Cusp number of rachidian</td>
<td id="C4F33503EC5E82A48AED4C72D33186D0" colspan="1" rowspan="1" style="color: #231f20">57</td>
<td id="0A66F539E7CAE58B060A631D927F3043" colspan="1" rowspan="1" style="color: #231f20">58</td>
<td id="EF20788B50EB8A81E69D41745A022F9D" colspan="1" rowspan="1" style="color: #231f20">56</td>
<td id="51E8FEF5A9370AC5C3C7EBD2210A565A" colspan="1" rowspan="1" style="color: #231f20">58</td>
<td id="338887E3E57BDE638181616FCFE7DF3E" colspan="1" rowspan="1" style="color: #231f20">58</td>
<td id="1D39E9F514A66F00B345271E6928A5B1" colspan="1" rowspan="1" style="color: #231f20">57</td>
<td id="CA6EE2E7A17C19859BEAB26187952318" colspan="1" rowspan="1" style="color: #231f20">57</td>
<td id="42D3A3D5C827E7B4133A462CE5AF96DF" colspan="1" rowspan="1" style="color: #231f20">58</td>
<td id="ECD0661FB61AB07FCD84EE65B0DF1342" colspan="1" rowspan="1" style="color: #231f20">56</td>
</tr>
<tr id="5F3E8BAB4624C7B1C5694ACD45BD4A8A">
<td id="4369DFFEE636631A17DEE692E74EEFD8" colspan="1" rowspan="1" style="color: #231f20">Cusp number of laterial teeth</td>
<td id="6B0129AC52E8880D3AA9A740D36D18C1" colspan="1" rowspan="1" style="color: #231f20">48</td>
<td id="33D61E8F2D427F363B605EA476BA57A3" colspan="1" rowspan="1" style="color: #231f20">57</td>
<td id="C544FCDABE435D6D3CD03F31A7E5DBC5" colspan="1" rowspan="1" style="color: #231f20">46</td>
<td id="5BDD45912B17B905827EA163A9CFD4DC" colspan="1" rowspan="1" style="color: #231f20">67</td>
<td id="D869BB6F0E87F3C08BD1F60CEC7C86C0" colspan="1" rowspan="1" style="color: #231f20">57</td>
<td id="EE7DA958A048283CC247B535773AB815" colspan="1" rowspan="1" style="color: #231f20">56</td>
<td id="0C6EBCF548FE3C734B3B20228EF03B0C" colspan="1" rowspan="1" style="color: #231f20">57</td>
<td id="268DF177CEB51F13881BA532048712D3" colspan="1" rowspan="1" style="color: #231f20">46</td>
<td id="9314814C35FDF245881B3CB587BB25BF" colspan="1" rowspan="1" style="color: #231f20">46</td>
</tr>
<tr id="4664E5BF3D1EEB3D208D6247E2BAD494">
<td id="B46910CEB627F55D15BFF7A8E8F36A31" colspan="1" rowspan="1" style="color: #231f20">Cusp number of interior marginal teeth</td>
<td id="0ECC6D08CF243862E24DB8E9FAC5E5F4" colspan="1" rowspan="1" style="color: #231f20">45</td>
<td id="952D33351F68C10CF091EE5410BEA38F" colspan="1" rowspan="1" style="color: #231f20">56</td>
<td id="DF0895819C81A3C9D77B35C63101B783" colspan="1" rowspan="1" style="color: #231f20">45</td>
<td id="DFC618CF2663D472F90EC5F625190A0E" colspan="1" rowspan="1" style="color: #231f20">36</td>
<td id="130A8E4D940A70D9E9D1A8533DE405C7" colspan="1" rowspan="1" style="color: #231f20">45</td>
<td id="049BA4461BEC0B460318FF27A72BBCAC" colspan="1" rowspan="1" style="color: #231f20">45</td>
<td id="1EF3F3596EB360147A88DC6E81493A41" colspan="1" rowspan="1" style="color: #231f20">45</td>
<td id="EC8EE30F69AFD69248CC1DEE11CAAA10" colspan="1" rowspan="1" style="color: #231f20">46</td>
<td id="2F29855270C046A7135B7E9CFBF62C4C" colspan="1" rowspan="1" style="color: #231f20">57</td>
</tr>
<tr id="33909C59B37601DBA239561F2DAD10E4">
<td id="1AEC80A68139FCB9AD2DD97644522467" colspan="1" rowspan="1" style="color: #231f20">Cusp number of exterior marginal teeth</td>
<td id="DFE134ADEA08B9627D0D7A8CC0F64E80" colspan="1" rowspan="1" style="color: #231f20">34</td>
<td id="8226DBBF64433D294245C51EDAAC15D0" colspan="1" rowspan="1" style="color: #231f20">45</td>
<td id="2B51DD0F4FA3159C7F1362CA16648CE1" colspan="1" rowspan="1" style="color: #231f20">4</td>
<td id="1D7223862FA61EEC9342AE8E2F081E00" colspan="1" rowspan="1" style="color: #231f20">35</td>
<td id="0B30CE1B385E43149ABAFEE5F7636692" colspan="1" rowspan="1" style="color: #231f20">34</td>
<td id="456C695C598BE95C476C626EE55609BC" colspan="1" rowspan="1" style="color: #231f20">34</td>
<td id="A4E28A2678C12BA4BADE89963BDB2B69" colspan="1" rowspan="1" style="color: #231f20">34</td>
<td id="2C9847AEC3EE8AE7C58F55AF8CB12F77" colspan="1" rowspan="1" style="color: #231f20">35</td>
<td id="51D0C968E876F8BF8E7829D74C219B99" colspan="1" rowspan="1" style="color: #231f20">45</td>
</tr>
<tr id="D1184F9F539E5F85FC917704FFA5009F">
<td id="8A2CE0F2C568D04F207D1AA880E24AF8" colspan="1" rowspan="1" style="color: #231f20">Shape of the largest cusp of rachidian tooth</td>
<td id="97293D042F2E36BD50163B5052051085" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="DC9E98044ACCF65C60FC202161B9184E" colspan="1" rowspan="1" style="color: #231f20">very narrow, pointed</td>
<td id="D56F835704285DE5FE717B37A3CA887C" colspan="1" rowspan="1" style="color: #231f20">very narrow, pointed</td>
<td id="11F309E47D49F6B2ED41F3F40267C899" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="F7F50F7DE90D155BD676009F585F7C4C" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="68FE905C38A4D14D00A7F8B31621C31A" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="5422414B9C8F7D77B1AD96E9B1E23D3B" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="C0B7D7EC2FB7E13A5E20F798021E1551" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="8A5C8A48AAB494EEDAE14BA9A1D969D7" colspan="1" rowspan="1" style="color: #231f20">slightly narrow, pointed</td>
</tr>
<tr id="317105CA872665A7250C5187F06F2197">
<td id="7F5AC85E7549BE17807D3DDEDD523B96" colspan="1" rowspan="1" style="color: #231f20">Shape of the largest cusp of lateral tooth</td>
<td id="71BF36D36C4C250D4B438FCF155EB818" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="6EBF7F063445FBF0A39F19951F0715AC" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="3B6CF22DD610DC3F50DC67D6535C4B70" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="3EC988AFDAA2FDB55B2E4EDE1D2DB692" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="63CA3C99E222D67900B73B9BDE30CDE7" colspan="1" rowspan="1" style="color: #231f20">narrow, pointed</td>
<td id="361A2B61B5A9C000850442990D548E81" colspan="1" rowspan="1" style="color: #231f20">slightly narrow, pointed</td>
<td id="8AE268A6BD9F80C7B605476C7E9C8746" colspan="1" rowspan="1" style="color: #231f20">slightly narrow, pointed</td>
<td id="9736B1EB12EAA2F22C11E4231C152678" colspan="1" rowspan="1" style="color: #231f20">slightly narrow, pointed</td>
<td id="D0D0E4BEFF254BD73BF68DC526A06BAB" colspan="1" rowspan="1" style="color: #231f20">wide, rounded</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="AEA9D45F6DDB523B3364239330C210AC">
<emphasis id="5142480458AC658EEF8130AA3FC9DF1E" bold="true" italics="true">Opercula</emphasis>
(Fig.
<figureCitation id="E75266443A9F23B88EDFB9338D9FF966" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells and opercula of Semisulcospira reticulata (A Y) and S. nishimurai sp. nov. (Z BF). A F. Holotype of S. reticulata from Oki-shima Island (offshore), NSMT - Mo 39770; G J. Hannoura, KUZ Z 4926, Z 4927; K O. Kitafunaki, KUZ Z 4930, Z 4931; P T. Oki-shima Island (offshore), KUZ Z 4925; U Y. Mano, KUZ Z 4229, Z 4286; Z AF. Holotype of S. nishimurai sp. nov. from Hannoura, KUZ Z 4935; AG AQ. Paratypes of S. nishimurai sp. nov. from Hannoura, KUZ Z 4936 Z 4939; AR AT. Oura Port, KUZ Z 4945; AU AY. Minamihama, KUZ Z 4946, Z 4947; AZ BD. Kitafunaki, KUZ Z 4952, Z 4953; BE BG. Oki-shima Island (coast), KUZ Z 4957. Scale bars: 10 mm (1), female teleoconch (A C, G, K, P, S, U, Z AB, AG, AJ, AM, AR, AU, AZ, BE), male teleoconch (J, N, AP, AX, BC), operculum (H, L, O, Q, T, V, Y, AC, AH, AK, AN, AQ, AS, AV, AY, BA, BD, BF); 2 mm (2), protoconch (D F, I, M, R, W); 1 mm (2), protoconch (AD AF, AI, AL, AO, AT, AW, BB, BG)." httpUri="https://binary.pensoft.net/fig/1065979">7 H, L, O, Q, T, V, Y</figureCitation>
).
<quantity id="018A6543CC11D4A6F4F45066A2ADC8F9" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="8.9" metricValueMin="5.8" unit="mm" value="7.35" valueMax="8.9" valueMin="5.8">5.88.9 mm</quantity>
in long diameter; nearly egg-shaped subcircular, paucispiral, comprising around 3 whorls; nucleus subcentral.
</paragraph>
<paragraph id="5D4ED80DFC8F9001409280EF0952060E">
<emphasis id="E9FC44C9AAA248D23579F4FDEC053A1A" bold="true" italics="true">Protoconchs</emphasis>
(Fig.
<figureCitation id="48DDE81FADBCF5680651CFAFCAE591CF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells and opercula of Semisulcospira reticulata (A Y) and S. nishimurai sp. nov. (Z BF). A F. Holotype of S. reticulata from Oki-shima Island (offshore), NSMT - Mo 39770; G J. Hannoura, KUZ Z 4926, Z 4927; K O. Kitafunaki, KUZ Z 4930, Z 4931; P T. Oki-shima Island (offshore), KUZ Z 4925; U Y. Mano, KUZ Z 4229, Z 4286; Z AF. Holotype of S. nishimurai sp. nov. from Hannoura, KUZ Z 4935; AG AQ. Paratypes of S. nishimurai sp. nov. from Hannoura, KUZ Z 4936 Z 4939; AR AT. Oura Port, KUZ Z 4945; AU AY. Minamihama, KUZ Z 4946, Z 4947; AZ BD. Kitafunaki, KUZ Z 4952, Z 4953; BE BG. Oki-shima Island (coast), KUZ Z 4957. Scale bars: 10 mm (1), female teleoconch (A C, G, K, P, S, U, Z AB, AG, AJ, AM, AR, AU, AZ, BE), male teleoconch (J, N, AP, AX, BC), operculum (H, L, O, Q, T, V, Y, AC, AH, AK, AN, AQ, AS, AV, AY, BA, BD, BF); 2 mm (2), protoconch (D F, I, M, R, W); 1 mm (2), protoconch (AD AF, AI, AL, AO, AT, AW, BB, BG)." httpUri="https://binary.pensoft.net/fig/1065979">7 I, M, R, W</figureCitation>
). Measurements and counts shown in Table
<tableCitation id="3DE72077F174A23E5ED5F2DC6CBDEAA6" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphometric characters of Semisulcospira reticulata and S. nishimurai sp. nov. Measurements and counts: minimum maximum value (mean ± SD)." httpUri="http://table.plazi.org/id/739C4F23DED6447AA0BA4D6A8D9BC1EC" tableUuid="739C4F23DED6447AA0BA4D6A8D9BC1EC">4</tableCitation>
. Shell globose or slightly elongated; suture slightly to moderately undulating; longitudinal ribs distinct, smooth, rarely granulated on central part of whorls in 12 rows; spiral cords weak or absent on lower part of whorls; shell colored beige to light beige in background; color bands olive or dark brown, thick, sometimes present on upper and lower parts of whorls and basal part.
</paragraph>
<paragraph id="82F976E83CE81077CFDBC1E46ACD0C5B">
<emphasis id="AF89C82E1071170FE00433BCCC902DAD" bold="true" italics="true">Radulae</emphasis>
(Fig.
<figureCitation id="2BFB0B863B6D9A1824350A573830CF46" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Radulae of Semisulcospira reticulata (A D) and S. nishimurai sp. nov. (E I). A. Hannoura, KUZ Z 4926; B. Kitafunaki, KUZ Z 4930; C. Oki-shima Island (offshore), KUZ Z 4925; D. Mano, KUZ Z 4229; E. Holotype of S. nishimurai sp. nov. from Hannoura, KUZ Z 4935; F. Oura Port, KUZ Z 4945; G. Minamihama, KUZ Z 4946; H. Kitafunaki, KUZ Z 4952; I. Oki-shima Island (coast), KUZ Z 4957. Scale bars: 100 μm." httpUri="https://binary.pensoft.net/fig/1065980">8 A D</figureCitation>
). Taenioglossa. Rachidian teeth consisting of central denticle and 24 small pointed triangular cusps on each side; tip of rachidian central denticle narrow, pointed, triangular, 2.03.5 times longer than other triangular cusps. Lateral teeth consisting of large central denticle and 24 inner and outer pointed cusps; central cusp of lateral teeth narrow, pointed, triangular, 2.03.0 times longer than other triangular cusps. Interior and exterior marginal teeth spoon-shaped, with 46 rounded denticles.
</paragraph>
<caption id="038C63BA27CCCD30F0370A4060ADFDFB" ID-arpha="5B9F6E2A-D46B-5C2F-BB6C-536A03CB0D01" httpUri="https://binary.pensoft.net/fig/1065980" startId="F8">
<paragraph id="5B28FABC53A520C9D7144D2ADC98DFF0">
<label id="6098159CE456767FF7D50EF1072F3EDD">Figure 8.</label>
</paragraph>
<paragraph id="641C4851B60956510000A3BC1273F534">
Radulae of
<taxonomicName id="560DCF2B2CFAEBAA23FF8953BFB5CF77" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="B99D0263CCC0E79F50CE1EE52A47C5E3" italics="true">Semisulcospira reticulata</emphasis>
</taxonomicName>
(
<emphasis id="48B56E761DCB5E8611761A7530DD41DD" bold="true">A D</emphasis>
) and
<taxonomicName id="2584E1F29546FF17D2F7A4E6D8742634" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="8E6DED98E12193867ECF041EF61615EB" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3EFB1D2B043EC49A423EB92A1141FA0" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="2B223DB9445436D986477124E9E69697" bold="true">E I</emphasis>
).
<emphasis id="355BBE426208BF64F977B57C21FC0BBB" bold="true">A.</emphasis>
Hannoura,
<specimenCode id="56D7CFF842C298594D6D563BAFB034BD">
<named-content id="DD682B1FE9ECF8DD5393A9E6114754D9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4926
</specimenCode>
;
<emphasis id="3C8B679EEF4A2075A825E484B79289B8" bold="true">B.</emphasis>
Kitafunaki,
<specimenCode id="A0641BE168695B2AF2B0BB099396042C">
<named-content id="A26DF9F4A213BD0DDB99808C92545BDA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4930
</specimenCode>
;
<emphasis id="54E95B86121E6354BBF178277F9726E9" bold="true">C.</emphasis>
Oki-shima Island (offshore),
<specimenCode id="C43393A18725B3E73204E3695C4720F0">
<named-content id="999900F4879D21D65AF0FD5B68AFF169" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4925
</specimenCode>
;
<emphasis id="B9BC2E56210A5C1CF766269C1B48385B" bold="true">D.</emphasis>
Mano,
<specimenCode id="EAC3F5446C81D833313741B92C8C2A1F">
<named-content id="7240545AF1F666E329B1E6008D065010" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4229
</specimenCode>
;
<emphasis id="F28EE8B9351568EBD53FF2FDD356A549" bold="true">E.</emphasis>
Holotype of
<taxonomicName id="432F96D6B9B2810669C2C7CB5E39E64C" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="90E5E297D969ABF27A1ADA92C2F2891A" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8F10FFFF84AE4EAB2ABCE544C5F4613F" rank="species">sp. nov.</taxonomicNameLabel>
from Hannoura,
<specimenCode id="A404F65194CB02C15DCF7CE317DEA00D">
<named-content id="D2433D2CF5C09A850F371B39A75B9013" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4935
</specimenCode>
;
<emphasis id="5066D82DC38E41D1297A73996FC007D5" bold="true">F.</emphasis>
Oura Port,
<specimenCode id="06BBA91747ED19B412824A49C4A4CE54">
<named-content id="D4DFDB8717CE88902B1531C61ACBD633" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4945
</specimenCode>
;
<emphasis id="A3A3900F62405B50EAEA86BC0165767C" bold="true">G.</emphasis>
Minamihama,
<specimenCode id="C2ED9197AB2CADC8CABEE4E5F00904A0">
<named-content id="E74B9CBC593ED307DAB513EEBD646E70" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4946
</specimenCode>
;
<emphasis id="7E83149A440088E277FF77E26D4CB052" bold="true">H.</emphasis>
Kitafunaki,
<specimenCode id="06C22A977C680134D36220193195048A">
<named-content id="EBC9FA1F032020244F88FB5B86408CE2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4952
</specimenCode>
;
<emphasis id="AD3FD5FCBCED4E666F5DF321EC6C13A6" bold="true">I.</emphasis>
Oki-shima Island (coast),
<specimenCode id="2FB5B6A81507D6D89551EAFE63B4C3C7">
<named-content id="A0B87B05A3540B410E959C33664BFCEC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoological-collection-kyoto-university" xlink_title="Zoological Collection of Kyoto University">KUZ</named-content>
Z 4957
</specimenCode>
. Scale bars: 100 μm.
</paragraph>
</caption>
<paragraph id="751C30C8B9C457471B3F9D691D6E755C">
<emphasis id="A387AC700138CBF8E33B437D808CECA5" bold="true" italics="true">Reproductive organs: Female</emphasis>
(Fig.
<figureCitation id="2DEADBDCD8AA6600AAC835E28EB6EA55" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Schematic drawings indicating generalised external features of the soft body and reproductive organs of Semisulcospira reticulata and S. nishimurai sp. nov.: A, B. Dorsal and ventral views of organs in visceral mass of mature female of S. nishimurai sp. nov. without operculum; C. External, right lateral view of pallial oviduct of S. reticulata; D. External, view of pallial oviduct of S. nishimurai sp. nov.; E. External, right upper lateral view of the prostate of S. nishimurai sp. nov.; F Internal folding structures of the prostate of S. nishimurai sp. nov. Abbreviations: bp, brood pouch; ct, ctenidium; dg, digestive gland; eg, egg; em, embryo; f, foot; inn, inner wall of brood pouch; int, intestine; kd, main kidney chamber; os, osphradium; ov, ovary; ovi, renal oviduct; pr, prostate; rcs, seminal receptacle; sg, sperm gutter; sn, snout; spb, spermatophore bursa; ss, style sac; sto, stomach; t, cephalic tentacle; vd, vas deferens. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1065981">9 C</figureCitation>
). Oviduct long, narrow, emerging from ovary entering near seminal receptacle on ventral side of soft body; protrusions of seminal receptacle long or short. Sperm gutter extending from spermatophore bursa towards mantle cavity, curved inward along whorls. Brood pouch extending along dorsal side of spermatophore bursa and sperm gutter, dorsally inflated, separated into many chambers, including eggs and embryos; eggs colored beige to orange, radially developing from base of brood pouch near seminal receptacle.
</paragraph>
<caption id="282F61939D31FAFCBB0D1FEAC4EC66F6" ID-arpha="A0108918-826C-5D1A-8C0C-CFD99FF678C3" httpUri="https://binary.pensoft.net/fig/1065981" startId="F9">
<paragraph id="5CB4ACCBAA547FEC60A7201E86E33B0D">
<label id="EF9007EC4F39A3A647E27C353E400D2A">Figure 9.</label>
</paragraph>
<paragraph id="C229776B918B91F40EE02B38CC36FA19">
Schematic drawings indicating generalised external features of the soft body and reproductive organs of
<taxonomicName id="4345F76766FF82B700E9CCA292C0B2C5" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="412EDE684A9EF4B38797781253080D5A" italics="true">Semisulcospira reticulata</emphasis>
</taxonomicName>
and
<taxonomicName id="3F5303ECDBF4432A81C6DCDB53C4AC36" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="E461478FDF72C520CF123D64FCCE3BE7" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="856F67D0B380D2FAC8D5C49A87E9CBF1" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="59958DE92E50AE7C15B6FA35D46E8285" bold="true">A, B.</emphasis>
Dorsal and ventral views of organs in visceral mass of mature female of
<taxonomicName id="4C990C6243A454E9D15C7580CD50F6B8" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="26C5036EC7F6B1FC6C96B118D4EC170D" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DCF4DF8D0DDCCE89200CC41264D92451" rank="species">sp. nov.</taxonomicNameLabel>
without operculum;
<emphasis id="86ED266636929A63F656459D7573CD85" bold="true">C.</emphasis>
External, right lateral view of pallial oviduct of
<taxonomicName id="B925CF2223A3298591D1A8EE140DA8B4" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="5DE1926453F81FD3DED87DF30704BCDF" italics="true">S. reticulata</emphasis>
</taxonomicName>
;
<emphasis id="7D128A32CC17A0E40521F70063D83661" bold="true">D.</emphasis>
External, view of pallial oviduct of
<taxonomicName id="94DEC3CAFFEB90B5D8BC4398730FB9E8" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="96C0E6D410F479931ED69D1C9A97487B" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84BAE60EF3CA3D2FFE9DEB6DFD5A3691" rank="species">sp. nov.</taxonomicNameLabel>
;
<emphasis id="82036926E62C675156F990360FD71420" bold="true">E.</emphasis>
External, right upper lateral view of the prostate of
<taxonomicName id="838DCE265EE5E4F094D5623B449743E9" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="3113E5E1DE36AE0B110E7029884B9148" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="606E83A2E1CB8B0CEA41AA1A89475E42" rank="species">sp. nov.</taxonomicNameLabel>
;
<emphasis id="0D6AB782AFF9F1A81FFA91953AED3BA8" bold="true">F</emphasis>
Internal folding structures of the prostate of
<taxonomicName id="F928BFF379C0F9F2FFF51DD2BDE3A4BC" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="C91FAD5517298C9E40B304877B70492E" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B465EA53C371AACAC5AEEA0E42C1615" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations: bp, brood pouch; ct, ctenidium; dg, digestive gland; eg, egg; em, embryo; f, foot; inn, inner wall of brood pouch; int, intestine; kd, main kidney chamber; os, osphradium; ov, ovary; ovi, renal oviduct; pr, prostate; rcs, seminal receptacle; sg, sperm gutter; sn, snout; spb, spermatophore bursa; ss, style sac; sto, stomach; t, cephalic tentacle; vd, vas deferens. Scale bars: 5 mm.
</paragraph>
</caption>
<paragraph id="22402D0A6CA14A9C4AA54CE89CA0DC55">
<emphasis id="BD88EC153AA84DA98EEBCE667960BF4D" bold="true" italics="true">Male.</emphasis>
Vas deferens long, narrow, emerging from testes, entering posterior end of prostate. Prostate without penis elongated, inflated in posterior ventral side, with deep groove, forming U-shape in transverse section, anterior narrowly opening to mantle cavity.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESZDI" type="distribution">
<paragraph id="C919182E3176F8AE9DEF5D73C0AF1121">
<heading id="1F95C9128C06E5287E5FCDF3AFC78B68" reason="title">Distribution and ecology.</heading>
</paragraph>
<paragraph id="1F97FD9FF2FC1BDC2A8B8B18512EA349">
<taxonomicName id="C23D75BDC344AE683519EE30A186E5B2" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="32406B99D22E72F1F310ABE38B7394B8" italics="true">Semisulcospira reticulata</emphasis>
</taxonomicName>
has been recorded from the northern and southern basins of Lake Biwa (
<bibRefCitation id="FF25409AB4673D01E42DFDBF36D4B391" author="Watanabe and Nishino" firstAuthor="Watanabe" journalOrPublisher="Lake Biwa Study Monographs" pagination="1-36" refId="B60" refString="Watanabe NC, Nishino M (1995) A study on taxonomy and distribution of the freshwater snail, genus Semisulcospira in Lake Biwa, with descriptions of eight new species. Lake Biwa Study Monographs 6: 136." title="A study on taxonomy and distribution of the freshwater snail, genus Semisulcospira in Lake Biwa, with descriptions of eight new species." volume="6" year="1995">Watanabe and Nishino 1995</bibRefCitation>
: fig. 5 e; Fig.
<figureCitation id="89D68E81E14142D35C94589BD0AB7580" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map representing seven localities where Semisulcospira reticulata (purple) and S. nishimurai sp. nov. (red) were collected in Lake Biwa, Japan. The detailed localities are shown in Table 1." httpUri="https://binary.pensoft.net/fig/1065973">1</figureCitation>
). However, this species has not been collected from the southern basin of the lake in recent years (
<bibRefCitation id="D85727A40547BDC290CF81735C4D444C" author="Nishino and Tanida" firstAuthor="Nishino" journalOrPublisher="Incorporated Administrative Agency Japan Water Agency Lake Biwa Development Integrated Operation &amp; Maintenance Office, Otsu" refId="B37" refString="Nishino M, Tanida K (2018) Illustration of Lake Biwa Benthic Fauna. Second Edition. Incorporated Administrative Agency Japan Water Agency Lake Biwa Development Integrated Operation &amp; Maintenance Office, Otsu, 335 pp." title="Illustration of Lake Biwa Benthic Fauna. Second Edition." year="2018">Nishino and Tanida 2018</bibRefCitation>
).
<taxonomicName id="FC15509FB1A48D7E694D3B677F588116" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="CE31ECBF1950139CAE49B52CCFF91C9D" italics="true">Semisulcospira reticulata</emphasis>
</taxonomicName>
was found on sandy mud to mud bottoms at depths of
<quantity id="903E1DB4BCF8A688A7166FB76EDBA06C" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="3.0" metricValueMin="0.4" unit="m" value="17.0" valueMax="30.0" valueMin="4.0">430 m</quantity>
(Watanabe 1970,
<bibRefCitation id="0C6856883A1D7819CD72EAADB4560517" DOI="10.3739/rikusui.41.212" author="Watanabe" firstAuthor="Watanabe" journalOrPublisher="Japanese Journal of Limnology" pagination="212-224" refId="B59" refString="Watanabe NC (1980) Some factors affecting the distribution and abundance of the two melanian snails, Semisulcospira decipiens and S. reticulata, in Lake Biwa. Japanese Journal of Limnology 41: 212224. https://doi.org/10.3739/rikusui.41.212" title="Some factors affecting the distribution and abundance of the two melanian snails, Semisulcospira decipiens and S. reticulata, in Lake Biwa." volume="41" year="1980">1980</bibRefCitation>
; Table
<tableCitation id="CEB10CC5E097E1FD0C95AC83688A76E9" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Specimen list of presently examined Semisulcospira niponica - group species with voucher numbers, collection localities, and DDBJ Sequence Read Archive (DRA) accession numbers for the specimens used for the genetic analysis. The locality numbers correspond to Fig. 1. Acronym: KUZ, Zoological Collection of Kyoto University; NSMT, National Museum of Nature and Science, Japan." httpUri="http://table.plazi.org/id/86DEDE3B0696CA956A00C4FE52957AA5" tableUuid="86DEDE3B0696CA956A00C4FE52957AA5">1</tableCitation>
). The species was collected with other congeners:
<taxonomicName id="0BA74489D0A3207DE5868522BD403D27" baseAuthorityName="Smith" baseAuthorityYear="1876" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="niponica">
<emphasis id="086844CD11E9F43E4BBF9517510E1DB6" italics="true">S. niponica</emphasis>
</taxonomicName>
- group,
<taxonomicName id="8997FD39477B0365D8B7F9487558FABB" baseAuthorityName="Westerlund" baseAuthorityYear="1883" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="decipiens">
<emphasis id="D38CF8FF502EE1CAB84EC9C266E1148D" italics="true">S. decipiens</emphasis>
</taxonomicName>
at Hannoura, Kitafunaki, and Mano,
<taxonomicName id="08E61A242641336653DD6A3A911601C6" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="56DA42F09A1CB580B55313561792E3AD" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FD3E3CBEC413ACFA177C3351C8372C7" rank="species">sp. nov.</taxonomicNameLabel>
at Hannoura and Kitafunaki;
<taxonomicName id="F5B839CDB032C778924AF8A3AB54D867" baseAuthorityName="Kuroda" baseAuthorityYear="1929" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nakasekoae">
<emphasis id="3827BAE95A622191FDACA2A69432C3DB" italics="true">S. nakasekoae</emphasis>
</taxonomicName>
- group,
<taxonomicName id="1ADB8373ED2800DD76A60C29B011B27A" authorityName="Sawada" authorityYear="2022" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="cryptica">
<emphasis id="F6AEA7502D71155A783030614E338545" italics="true">S. cryptica</emphasis>
</taxonomicName>
and
<taxonomicName id="0AC406E66FD628B486EBC2E915AF9579" authorityName="N. Watanabe &amp; Nishino" authorityYear="1995" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="ourensis">
<emphasis id="E1BB90DF7914E9C8721EB5A80645096D" italics="true">S. ourensis</emphasis>
</taxonomicName>
at Hannoura,
<taxonomicName id="2F160DA152B44C0A3EEF1B6AE0D0741B" authorityName="Watanabe &amp; Nishino" authorityYear="1995" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="arenicola">
<emphasis id="466F4ABF58990A3E8149D8D0650E425E" italics="true">S. arenicola</emphasis>
</taxonomicName>
at Kitafunaki,
<taxonomicName id="CCF3C38E01B2EA733D85F2F1ECEED6F9" authorityName="Sawada" authorityYear="2022" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="elongata">
<emphasis id="53434D7B5125C085006B642C462E6FEF" italics="true">S. elongata</emphasis>
</taxonomicName>
at Kitafunaki, Oki-shima Island, and Mano,
<taxonomicName id="ECBB6E97A7B8539FD5B4F1262681B324" authorityName="Sawada &amp; Nakano" authorityYear="2021" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="davisi">
<emphasis id="CDC1ADEB76104BD62F8E072AA32284F7" italics="true">S. davisi</emphasis>
</taxonomicName>
at Hannoura, Kitafunaki, and Oki-shima Island.
<taxonomicName id="3252A6D6723209CE502F9409D47AB4C7" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="C43218B07315681457897D4A5B17AC41" italics="true">Semisulcospira reticulata</emphasis>
</taxonomicName>
was found with
<taxonomicName id="D7B2AB78D0F3A593A51E470FB5DE3B6D" baseAuthorityName="Brot" baseAuthorityYear="1876" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reiniana">
<emphasis id="2721C409D3641841CAE8C73A8503E611" italics="true">S. reiniana</emphasis>
</taxonomicName>
at Kitafunaki.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E25DI" type="remarks">
<paragraph id="59440BEFD50328B6F2F7AD1D6D9E21A0">
<heading id="F421B79AD731A0D2AA6F7F3F23AB07ED" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="DAA8995764523D4DCED51CCE466ADB66">
Genetic differentiation among specimens from Kitafunaki and the other localities were clarified in this species (Figs
<figureCitation id="E5ACDE5CAD28621FC8EFA14A33D8A5D6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The maximum likelihood tree of 11 Semisulcospira niponica - group species calculated based on 19,767 SNPs. The numbers associated with the nodes represent values of SH-aLRT (left) / ultrafast bootstrap (right). Values of SH-aLRT higher than 80.0 and that of ultrafast bootstrap greater than 95 are indicated." httpUri="https://binary.pensoft.net/fig/1065974">2</figureCitation>
,
<figureCitation id="713C68D4D30396C7C67B0233366509F8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SVDquartets tree of Semisulcospira niponica - group populations calculated based on 19,767 SNPs. The numbers associated with the nodes represent bootstrap values." httpUri="https://binary.pensoft.net/fig/1065975">3</figureCitation>
) whereas morphological distinctiveness were not observed in the Kitafunaki specimens (Table
<tableCitation id="77A0B22A2B301C6E732ED20C5BCAA20E" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphometric characters of Semisulcospira reticulata and S. nishimurai sp. nov. Measurements and counts: minimum maximum value (mean ± SD)." httpUri="http://table.plazi.org/id/739C4F23DED6447AA0BA4D6A8D9BC1EC" tableUuid="739C4F23DED6447AA0BA4D6A8D9BC1EC">4</tableCitation>
). The teleoconch of
<taxonomicName id="B3CFD655A42D0373B40D322703EAF800" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="8031EE119F044E3D1F430F68507F6325" italics="true">S. reticulata</emphasis>
</taxonomicName>
resembles that of
<taxonomicName id="392702EB66DFC3C6288A0045AFD31FFF" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="B3DF9233C79D5C627386F6A34C0EC9A9" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6649E018ACFE1083E68BBFE946F3D77" rank="species">sp. nov.</taxonomicNameLabel>
However,
<taxonomicName id="97A71BE53D62F637CFECEE59F4B811B2" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="39E8CA7B02470BF5BCDFCA5E9D25B281" italics="true">S. reticulata</emphasis>
</taxonomicName>
represents a greater number of axial ribs on the teleoconch, and the protoconch size of
<taxonomicName id="0185934729AE436398B9409E1980C9D8" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="B2F1E01871E83A810E40916BC806D376" italics="true">S. reticulata</emphasis>
</taxonomicName>
is prominently larger than that of
<taxonomicName id="E10DAA3BBC478C7001EACD92520E940A" authority="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano, 2024" authorityName="Sawada &amp; Fuke &amp; Miura &amp; Toyohara &amp; Nakano" authorityYear="2024" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="nishimurai" status="sp. nov.">
<emphasis id="A996385B6A6A078CAE6B10BC402A05D5" italics="true">S. nishimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3FD3D49835334D9655B00CF6020DCC12" rank="species">sp. nov.</taxonomicNameLabel>
Although the protoconch size and shape of
<taxonomicName id="96AEB1C28A2585E6A36C1336C7772DC8" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="4810E7A6F13C3B120471A1ABB91FE073" italics="true">S. reticulata</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="4F97AB3C65B901F426747C7D0E61A5DC" authorityName="Sawada &amp; Nakano" authorityYear="2021" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="davisi">
<emphasis id="D4A3ED415BCF2408544C80DE9F6B5132" italics="true">S. davisi</emphasis>
</taxonomicName>
, larger SH,
<abbrev id="ABBRID0E2BAK" xlink_title="body whorl length">BWL</abbrev>
, and
<abbrev id="ABBRID0E6BAK" xlink_title="spire angle">SA</abbrev>
values of
<taxonomicName id="82CE48776FFAEF8969D992D8C028D07D" authorityName="Kajiyama &amp; Habe" authorityYear="1961" class="Gastropoda" family="Semisulcospiridae" genus="Semisulcospira" kingdom="Animalia" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="B12E166BB3434E6FADEAFDDE90FD138D" italics="true">S. reticulata</emphasis>
</taxonomicName>
discriminate the two species.
</paragraph>
</subSubSection>
</treatment>
</document>