315 lines
34 KiB
XML
315 lines
34 KiB
XML
<document ID-CLB-Dataset="8199" ID-DOI="http://dx.doi.org/10.3897/phytokeys.172.62511" ID-GBIF-Dataset="e3a1f979-8f8b-4095-8027-1959bcd2096e" ID-PMC="PMC7895818" ID-Pensoft-Pub="1314-2003-172-75" ID-Pensoft-UUID="FDAE8D4147695752AAF24B45EA7067B0" ID-PubMed="33633487" ModsDocID="1314-2003-172-75" checkinTime="1613177897904" checkinUser="pensoft" docAuthor="Xu, Ke-Wang, Wang, Lu-Lu & Zhang, Li-Bing" docDate="2021" docId="230A5D191A5F56518C542771D1A5F30F" docLanguage="en" docName="PhytoKeys 172: 75-91" docOrigin="PhytoKeys 172" docPubDate="2021-02-12" docSource="http://dx.doi.org/10.3897/phytokeys.172.62511" docTitle="Asplenium alatulum Ching, Acta Phytotax. Sin. 9 (4): 359. 1964." docType="treatment" docVersion="6" id="FDAE8D4147695752AAF24B45EA7067B0" lastPageNumber="75" masterDocId="FDAE8D4147695752AAF24B45EA7067B0" masterDocTitle="Taxonomic revision of the Asplenium wrightii complex (Aspleniaceae) with reinstatement of A. alatulum and A. subcrenatum" masterLastPageNumber="91" masterPageNumber="75" pageNumber="75" updateTime="1732828702414" updateUser="ExternalLinkService">
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<mods:mods id="F171F67D7126D2EA55FABA8FCB32D58E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="973B5B3140DFFFCCB09F1B3406740048">
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<mods:title id="7FF33A366EAC8CFD76FF2967A77C9475">Taxonomic revision of the Asplenium wrightii complex (Aspleniaceae) with reinstatement of A. alatulum and A. subcrenatum</mods:title>
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</mods:titleInfo>
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<mods:name id="D3A1F1B5B1824BC1291D6CCA80DB377D" type="personal">
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<mods:roleTerm id="8104A263A9376373EF2A72A72ADACE2E">Author</mods:roleTerm>
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<mods:namePart id="F460C88E5895D929CBB9E383757C593E">Xu, Ke-Wang</mods:namePart>
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<mods:nameIdentifier id="51C6BB1563B998E6A5CA51052861BF06" type="ORCID">https://orcid.org/0000-0003-2239-0487</mods:nameIdentifier>
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<mods:affiliation id="255368C98936975DCCE7C5993CB6A86D">Co-Innovation Center for Sustainable Forestry in Southern China, College of Biology and the Environment, Nanjing Forestry University, Nanjing; 510275, China</mods:affiliation>
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<mods:nameIdentifier id="34A0060903EAAE0FD0A5253B5AAE9718" type="email">xukw10@njfu.edu.cn</mods:nameIdentifier>
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<mods:name id="9FD10C457D94709356A51E95BDF9F7BA" type="personal">
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<mods:roleTerm id="624CE1D39E140573C78E3E4A005C6420">Author</mods:roleTerm>
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<mods:namePart id="D0A3736762E85934D3FE188C58D52304">Wang, Lu-Lu</mods:namePart>
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<mods:affiliation id="6A95A6184E542EEC16A6DB3AAF6F4A05">Co-Innovation Center for Sustainable Forestry in Southern China, College of Biology and the Environment, Nanjing Forestry University, Nanjing; 510275, China</mods:affiliation>
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<mods:name id="5690FD719CC8445C2F0809592975106E" type="personal">
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<mods:roleTerm id="B0A217946F8502C8DCFCB2A049CD9B25">Author</mods:roleTerm>
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<mods:namePart id="86F0A5B9B02D6F0E073490C0733E25B3">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="886DF4F20C66FD4DDB7DB079D3A3D274">Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166 - 0299, USA</mods:affiliation>
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<mods:nameIdentifier id="EE38DE8ADF55B07A5B88D40D84BBB0EA" type="email">libing.zhang@mobot.org</mods:nameIdentifier>
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<mods:typeOfResource id="856998CC4AF7D0575D941DD4DA4888DE">text</mods:typeOfResource>
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<mods:titleInfo id="5BC0603FD7C94EEA8DE6D7EF4969945F">
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<mods:title id="8FE0BFF5492DD31CD2C32E72EA3A84E6">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="6E26BCB9D7DB87759036C9C25ECED9BA">
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<mods:date id="1AF96023A557ECA511397CFDAC283756">2021</mods:date>
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<mods:detail id="CBADB42371A0E77A1B21245C3FE90FDF" type="pubDate">
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<mods:number id="5AC2A458A3595AF966B08091BB7A11C3">2021-02-12</mods:number>
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</mods:detail>
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<mods:detail id="92A1FBBAECB2DB0F4325A9326C626867" type="volume">
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<mods:number id="1E0142E79C78DE23D3B4952EB76B00EA">172</mods:number>
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<mods:start id="BD92154439C9AB1B6FC5A8D162FDD45E">75</mods:start>
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<mods:end id="BE6C812CEF6A6DCCB4C4ACFB6A248AB0">91</mods:end>
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<mods:location id="8738CD83F6D2CBB190EAE8857019FC91">
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<mods:url id="1FF161C99A1042F2859CBD12EC95D449">http://dx.doi.org/10.3897/phytokeys.172.62511</mods:url>
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</mods:location>
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<mods:classification id="A4DB480BDE5F97D1340018E883042375">journal article</mods:classification>
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<mods:identifier id="7D1318BB59DB096600ECFBF69BEF2475" type="DOI">http://dx.doi.org/10.3897/phytokeys.172.62511</mods:identifier>
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<mods:identifier id="A960EE7D526A45185ED25F2B4F466CB1" type="Pensoft-Pub">1314-2003-172-75</mods:identifier>
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<mods:identifier id="D0AF3F7375BA5A6208206A42E96210F4" type="Pensoft-UUID">FDAE8D4147695752AAF24B45EA7067B0</mods:identifier>
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<treatment id="230A5D191A5F56518C542771D1A5F30F" ID-GBIF-Taxon="177383159" LSID="urn:lsid:plazi:treatment:230A5D191A5F56518C542771D1A5F30F" httpUri="http://treatment.plazi.org/id/230A5D191A5F56518C542771D1A5F30F" lastPageNumber="75" pageId="0" pageNumber="75" scope_family="Aspleniaceae">
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<subSubSection id="548D187BCB89EF6103CB88BB43C63CBD" pageId="0" pageNumber="75" type="nomenclature">
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<paragraph id="5EF6AEA905F9BF335694B26F890D2389" pageId="0" pageNumber="75">
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<taxonomicName id="3CEF134D0F0366DEC4FA817C8758DFBC" ID-CoL="7RZZ5" LSID="230A5D19-1A5F-5651-8C54-2771D1A5F30F" authority="Ching, Acta Phytotax. Sin. 9 (4): 359. 1964." authorityName="Ching, Acta Phytotax. Sin. 9 (4): 359. 1964." class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alatulum">Asplenium alatulum Ching, Acta Phytotax. Sin. 9(4): 359. 1964.</taxonomicName>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="610F8066F118C8E27E91D3E8A1126C2D" pageId="0" pageNumber="75" type="materials_examined">
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<paragraph id="FFE0C1F7AFC6B44AB4203A77FC73C4FD" pageId="0" pageNumber="75">Type.</paragraph>
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<paragraph id="26AECC96BFE8E738961E08AFBA8DFA16" pageId="0" pageNumber="75">
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China.
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<emphasis id="D32C7F40F8F646BE880F2C6A999EEDAF" bold="true" pageId="0" pageNumber="75">Hainan</emphasis>
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: Five-Finger Mountain, 19 May 1922,
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<emphasis id="4B18CAED58121406CA15008038CE47B5" italics="true" pageId="0" pageNumber="75">F.A.McClure 9713</emphasis>
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(holotype: PE (PE00059412 [image!])). Fig.
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<figureCitation id="C1AA96C8B32C73ACCE16B309F6B035CD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Type of Asplenium alatulum Ching (A barcode PE 00059412), A. subcrenatum Ching ex S. H. Wu (B barcode PE 00059494), A. wrightii Eaton ex Hooker (C barcode NY 00128031) and isotype of A. wrightii GH (D barcode GH 00020612)." figureDoi="10.3897/phytokeys.172.62511.figure3" httpUri="https://binary.pensoft.net/fig/509068" pageId="0" pageNumber="75">3A</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="2CFEE2C81CD8915E82C5B7B31A523F00" pageId="0" pageNumber="75" type="description">
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<paragraph id="A56FE0BA51073FE85BC034B4A120B2D8" pageId="0" pageNumber="75">Description.</paragraph>
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<paragraph id="351AF30489281EFBEDF4C4633659B8DA" pageId="0" pageNumber="75">
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Plants up to 50 cm tall. Rhizomes erect to decumbent, densely scaly; scales dark brown, narrowly lanceolate or lanceolate-ovate, 5-9
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<normalizedToken id="B6D4BE82966D113E3CFBF353094BC3D1" originalValue="×">x</normalizedToken>
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0.4-0.8 mm, denticulate glandular margin or long fibrillose (Fig.
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<figureCitation id="D1F65FB4A47E614DCFED788A0AE0EAA4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Scales of A. alatulum, A. subcrenatum and A. wrightii. A, D, G A. wrightii B, E, H A. subcrenatum C, F, I A. alatulum." figureDoi="10.3897/phytokeys.172.62511.figure2" httpUri="https://binary.pensoft.net/fig/509067" pageId="0" pageNumber="75">2C, F, I</figureCitation>
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). Fronds tufted; stipe dull to semi-shiny, greyish-green to brown or stramineous-green, 12-25 cm, sparsely scaly, scales similar to those on rhizome; lamina triangular-ovate to elliptic, (15-)18-25(-30)
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<normalizedToken id="7019B9439B17405DEE4D3EB6718CE517" originalValue="×">x</normalizedToken>
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(10-)12-18(-22) cm, base truncate, apex acute, 1-pinnate (Fig.
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<figureCitation id="592C0D95A79946F20DE4C7E77E98B020" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Asplenium alatulum A Habitat and habit B Portion of lamina showing adaxial view of pinna C Portion of lamina showing abaxial view of pinna D Adaxial view of upper portion of lamina E Lower portion of rachis showing wide lateral wings." figureDoi="10.3897/phytokeys.172.62511.figure4" httpUri="https://binary.pensoft.net/fig/509069" pageId="0" pageNumber="75">4A</figureCitation>
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); pinnae 10-15 pairs, basal pinnae subopposite, others alternate, at an angle of ca. 60°-70° to rachis, with stalks (1-)2-4 mm, lower pinnae not reduced, suprabasal pinnae falcate-lanceolate, (5-)6-10(-12)
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<normalizedToken id="82967AD62C86F3AC9EC8C8C664E3BCFA" originalValue="×">x</normalizedToken>
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(0.8-)1.2-1.8(-2.0) cm, base asymmetrical, acroscopic side truncate at an angle of (55°-)65°-75(-85°) to costa, basiscopic side cuneate, becoming decurrent on rachis in apical part of lamina, margin serrate to dentate, apex acuminate (Fig.
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<figureCitation id="C6BBD065344B40FAE992F181CEA64A8E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Asplenium alatulum A Habitat and habit B Portion of lamina showing adaxial view of pinna C Portion of lamina showing abaxial view of pinna D Adaxial view of upper portion of lamina E Lower portion of rachis showing wide lateral wings." figureDoi="10.3897/phytokeys.172.62511.figure4" httpUri="https://binary.pensoft.net/fig/509069" pageId="0" pageNumber="75">4B, C, D</figureCitation>
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). Veins (1 or) 2-forked, with terminal hydathode. Fronds papery, brownish-green when dry, subglabrous; rachis brown to greyish-green or stramineous-green, subglabrous, terete abaxially, with greyish-green lateral wings (Fig.
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<figureCitation id="9073A065C1887CBCCE1D6CBD46479851" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Asplenium alatulum A Habitat and habit B Portion of lamina showing adaxial view of pinna C Portion of lamina showing abaxial view of pinna D Adaxial view of upper portion of lamina E Lower portion of rachis showing wide lateral wings." figureDoi="10.3897/phytokeys.172.62511.figure4" httpUri="https://binary.pensoft.net/fig/509069" pageId="0" pageNumber="75">4E</figureCitation>
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). Sori linear, (2-)5-9(-12) mm, on acroscopic veinlets, medial to supramedial (Fig.
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<figureCitation id="FFDE98DF08FD466358C33062C20720CD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Asplenium alatulum A Habitat and habit B Portion of lamina showing adaxial view of pinna C Portion of lamina showing abaxial view of pinna D Adaxial view of upper portion of lamina E Lower portion of rachis showing wide lateral wings." figureDoi="10.3897/phytokeys.172.62511.figure4" httpUri="https://binary.pensoft.net/fig/509069" pageId="0" pageNumber="75">4C</figureCitation>
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); indusia greyish-brown to dark brown, linear, papery, margin entire, opening towards costa, persistent. Spores with average exospore length 40-45
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<normalizedToken id="240309BB7A2E5AD5546C5804B6F5BC87" originalValue="μm">μm</normalizedToken>
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, perispore cristato-alate (Fig.
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<figureCitation id="A2B0C05B565F4B9116F1F5E8AF95E4BF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scanning electron micrographs of spore and stomata of A. alatulum, A. subcrenatum and A. wrightii A-F A. wrightii G-I A. subcrenatum J-L A. alatulum." figureDoi="10.3897/phytokeys.172.62511.figure1" httpUri="https://binary.pensoft.net/fig/509066" pageId="0" pageNumber="75">1G</figureCitation>
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).
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</paragraph>
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<caption id="FAAD8B97ABFE111018D0A4C3A0860D61" doi="10.3897/phytokeys.172.62511.figure1" httpUri="https://binary.pensoft.net/fig/509066" pageId="0" pageNumber="75" start="Figure 1" startId="F1">
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<paragraph id="6DC06C9922B04839230221CA1680C4D1" pageId="0" pageNumber="75">
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<emphasis id="5CB13A270864401CF3CDFD130A64DBBB" bold="true" pageId="0" pageNumber="75">Figure 1.</emphasis>
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Scanning electron micrographs of spore and stomata of
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<taxonomicName id="D59837A821FF9E2DF2750935EAA7E5E7" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="alatulum">
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<emphasis id="6546D7E98943014C48587A4AE02C9E32" italics="true" pageId="0" pageNumber="75">A. alatulum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="52FC0A29635E19FB3184FCFF7667165A" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. subcrenatum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="subcrenatum">
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<emphasis id="09E4F587C981A3C11D90CB4ADF4DDA9E" italics="true" pageId="0" pageNumber="75">A. subcrenatum</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="EA75F241D1D49AB93052ADE652FBD7EA" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
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<emphasis id="10AD270251BD98355A2B7B72A25CCCA8" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
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</taxonomicName>
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<emphasis id="FBE05BB5DF43D2AF71E8B545A88F3836" bold="true" pageId="0" pageNumber="75">A-F</emphasis>
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<taxonomicName id="068CC2D8F66BC8268695398FAA7F2227" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
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<emphasis id="E78C7FED09C635D24DD9AF1535353F91" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
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</taxonomicName>
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<emphasis id="F501FE4D397F9A562141BA0D1419F473" bold="true" pageId="0" pageNumber="75">G-I</emphasis>
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<taxonomicName id="07DF02462801655265731DD01E761860" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. subcrenatum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="subcrenatum">
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<emphasis id="47D8D4664397D301D2511841BACA1679" italics="true" pageId="0" pageNumber="75">A. subcrenatum</emphasis>
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</taxonomicName>
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<emphasis id="C80663BA12666AF514715F833B05A8DE" bold="true" pageId="0" pageNumber="75">J-L</emphasis>
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<taxonomicName id="2FFAA3C6DADEA2EA998B7E733C8124CA" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="alatulum">
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<emphasis id="B263AC5A82EFEC368A8D17C3BF5A09B0" italics="true" pageId="0" pageNumber="75">A. alatulum</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<caption id="44A9B655ABC10F59594FA7CE1A423BA1" doi="10.3897/phytokeys.172.62511.figure2" httpUri="https://binary.pensoft.net/fig/509067" pageId="0" pageNumber="75" start="Figure 2" startId="F2">
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<paragraph id="CAF091AC56EB94340A3C47940CFEE68E" pageId="0" pageNumber="75">
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<emphasis id="D717A2FFD9394A70509DD1DF8B6E7143" bold="true" pageId="0" pageNumber="75">Figure 2.</emphasis>
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Scales of
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<taxonomicName id="7171742CF7CD20C7961803707451ACE7" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="alatulum">
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<emphasis id="6DE0B179326CEBDB382F82E99A6732E1" italics="true" pageId="0" pageNumber="75">A. alatulum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="D649AF9888F60604FC35C45B73AD5E19" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. subcrenatum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="subcrenatum">
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<emphasis id="55C8EA1234624637ACB2B027D20ECF33" italics="true" pageId="0" pageNumber="75">A. subcrenatum</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="0800BA4A4F571F7BDDF6A993CAFF0970" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
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<emphasis id="82EBBAE447FC76B3798BDB7D3D696873" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
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</taxonomicName>
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.
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<emphasis id="C237ACC1D0004D4BF9FC3738AF9F0130" bold="true" pageId="0" pageNumber="75">A, D, G</emphasis>
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<taxonomicName id="4C48A36FEB9C7368C8BAAEDEA1CC9C1E" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
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<emphasis id="81F8D2FD2C71DC0E37E1F8FC64E69BA1" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
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</taxonomicName>
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<emphasis id="FC1B5B99C2585ECC3A6D1C8C78D7F343" bold="true" pageId="0" pageNumber="75">B, E, H</emphasis>
|
||
<taxonomicName id="FE1B1D05B773D91E5D3AD07D503B9BBB" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. subcrenatum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="subcrenatum">
|
||
<emphasis id="7211D1D243A69BDCCC761532CF4FE852" italics="true" pageId="0" pageNumber="75">A. subcrenatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="59FCF60CCFF8C765950F19F84FA65037" bold="true" pageId="0" pageNumber="75">C, F, I</emphasis>
|
||
<taxonomicName id="EED123EE1AC3B407CF261D91D4A369D0" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="alatulum">
|
||
<emphasis id="CE40A4A7C22C30EAB87580BDABEFCA0A" italics="true" pageId="0" pageNumber="75">A. alatulum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="E4436B3C9E26949D400AF24B0A45CAFC" pageId="0" pageNumber="75" type="distribution">
|
||
<paragraph id="5724A1788A1B55BB5320AB1FC437117A" pageId="0" pageNumber="75">Distribution and habitat.</paragraph>
|
||
<paragraph id="4C80D641E65BF716CAF7367044A2B2E5" pageId="0" pageNumber="75">
|
||
<taxonomicName id="DD01D1736DF6A0C61A2AACA8DE55602D" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium alatulum" order="Filicales" pageId="0" pageNumber="75" rank="species" species="alatulum">
|
||
<emphasis id="2B40402E837C4DC8D53C21027B6C211D" italics="true" pageId="0" pageNumber="75">Asplenium alatulum</emphasis>
|
||
</taxonomicName>
|
||
is apparently restricted to China. It grows on rocks in ravines of broad-leaved forests at an elevation of ca. 500-1600 m. (Fig.
|
||
<figureCitation id="BAB4E6421757EEB39FA2B9DCF19816CA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Asplenium alatulum A Habitat and habit B Portion of lamina showing adaxial view of pinna C Portion of lamina showing abaxial view of pinna D Adaxial view of upper portion of lamina E Lower portion of rachis showing wide lateral wings." figureDoi="10.3897/phytokeys.172.62511.figure4" httpUri="https://binary.pensoft.net/fig/509069" pageId="0" pageNumber="75">4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="24E0D8966A2959D6C30922A58B035723" pageId="0" pageNumber="75" type="materials_examined">
|
||
<paragraph id="CF2D628D6B5766E2ED7974100B623259" pageId="0" pageNumber="75">Additional specimens examined.</paragraph>
|
||
<paragraph id="A02F3015AF8BBD7C26BADA0528C27B56" pageId="0" pageNumber="75">
|
||
<emphasis id="E9660B990E50200B0FAFA4ABF9D4A1ED" bold="true" pageId="0" pageNumber="75">China. Hainan</emphasis>
|
||
: Wuzhishan, Shuiman Village, elev. 870 m,
|
||
<geoCoordinate id="1B6F91A10C61F27F08CE6BF351D95F2C" degrees="18" direction="north" minutes="54" orientation="latitude" precision="1" seconds="10.6" value="18.902945">18°54'10.6"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A853814A0935FE991DADD54DA25204AC" degrees="109" direction="east" minutes="41" orientation="longitude" precision="1" seconds="15.6" value="109.68767">109°41'15.6"E</geoCoordinate>
|
||
, 7 Apr 2016,
|
||
<emphasis id="6D94468A0E8298D99ED856DB1F78BEAC" italics="true" pageId="0" pageNumber="75">K.W.Xu 107</emphasis>
|
||
(SYS!); the same locality, elev. 1550 m, 20 Dec 2010,
|
||
<emphasis id="A5AC0DAA1ADB8678E75D517F5CA69F5F" italics="true" pageId="0" pageNumber="75">X.P.Wei & R.Wei WXP113</emphasis>
|
||
(PE-2286681!).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22BB3B8FCC895F55DC3BF33E24A6C843" pageId="0" pageNumber="75" type="notes">
|
||
<paragraph id="C1886C8561EC9AF4D6D28FC3E7A2E0C9" pageId="0" pageNumber="75">Note.</paragraph>
|
||
<paragraph id="B7D0D2D3BC1A7107413733EDCA326EA5" pageId="0" pageNumber="75">
|
||
Though
|
||
<taxonomicName id="0B571BAE14EC3278F57071FB0A869466" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium alatulum" order="Filicales" pageId="0" pageNumber="75" rank="species" species="alatulum">
|
||
<emphasis id="CA5E95CA302C816EE9632574EB2765AF" italics="true" pageId="0" pageNumber="75">Asplenium alatulum</emphasis>
|
||
</taxonomicName>
|
||
was thought to be an endemic species to the Hainan Island before its synonymisation with
|
||
<taxonomicName id="DD0BFDB8336F71D76DD3108B9841A788" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="57902031DE6CE97872621B48F7267513" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="0D5741E34E2F668858487E29A88ABF83" author="Lin, YX" editor="Wu, ZY" journalOrPublisher="2 - 3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis" pageId="0" pageNumber="75" pagination="267 - 316" refId="B12" refString="Lin, YX, Viane, R, 2013. Aspleniaceae. In: Wu, ZY, Raven, PH, Hong, DY, Eds., Flora of China (Vol. 2-3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis: 267 - 316" title="Aspleniaceae." volumeTitle="Flora of China (Vol." year="2013">Lin and Viane (2013)</bibRefCitation>
|
||
, the morphological distinction between
|
||
<taxonomicName id="6423E5C5794D841D5D1A37EDA10C5FAA" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="alatulum">
|
||
<emphasis id="A3CC3EB07054A769545EE788345B302E" italics="true" pageId="0" pageNumber="75">A. alatulum</emphasis>
|
||
</taxonomicName>
|
||
and its closely-related species was obscure due to the insufficient field investigations. In the protologue,
|
||
<bibRefCitation id="DBA1F2E64C6D02D6FF8E5B1E3D73C5E9" author="Ching, RC" journalOrPublisher="Zhiwu Fenlei Xuebao" pageId="0" pageNumber="75" pagination="345 - 373" refId="B2" refString="Ching, RC, Wang, CH, 1964. Additional Materials for the Pteridophytic Flora of Hainan. Zhiwu Fenlei Xuebao 9: 345 - 373" title="Additional Materials for the Pteridophytic Flora of Hainan." volume="9" year="1964">Ching and Wang (1964)</bibRefCitation>
|
||
emphasised its small size, the short pinnae, the crenate-serrate pinna margins and whole rachis with lateral wings and stated that
|
||
<taxonomicName id="1F2B3475C561A4EF833F7CC0B8C43ED1" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="alatulum">
|
||
<emphasis id="9E7135A9F68E7B8159EFB8F5BAFC0A9B" italics="true" pageId="0" pageNumber="75">A. alatulum</emphasis>
|
||
</taxonomicName>
|
||
is markedly different from
|
||
<taxonomicName id="94FC3345A950FBDB5300171F1EC835AB" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="846326A8D0445F166446589AFB767736" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<bibRefCitation id="8DC8D546E91FB3021FFBF9029A94755C" author="Lin, YX" editor="Wu, ZY" journalOrPublisher="2 - 3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis" pageId="0" pageNumber="75" pagination="267 - 316" refId="B12" refString="Lin, YX, Viane, R, 2013. Aspleniaceae. In: Wu, ZY, Raven, PH, Hong, DY, Eds., Flora of China (Vol. 2-3). Science Press, Beijing & Missouri Botanical Garden Press, St. Louis: 267 - 316" title="Aspleniaceae." volumeTitle="Flora of China (Vol." year="2013">Lin and Viane (2013)</bibRefCitation>
|
||
recognised that the plant and pinna size and margin shape are variable in the
|
||
<taxonomicName id="7505EEEAB5B618EE7CEB01AF15805CE2" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="7F92795584973888524833EB49BBC5ED" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
complex and included
|
||
<taxonomicName id="33BBD8561BA83739F53AA738B1A4A07F" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="alatulum">
|
||
<emphasis id="FA7715FFB97216FEC37CDA90BFC5125D" italics="true" pageId="0" pageNumber="75">A. alatulum</emphasis>
|
||
</taxonomicName>
|
||
in the synonymy of
|
||
<taxonomicName id="BF4C7834B77A657F68E583B309963E46" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="D06D56F31137DF49A0B728EBED437D70" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
. Recently, our study, based on specimen examination and recent field observations of the two species, supports their distinction.
|
||
</paragraph>
|
||
<paragraph id="1EDBCBC53A153D14C45BC57A7CA42088" pageId="0" pageNumber="75">
|
||
<taxonomicName id="448498E297596A7815C07D3B3A1AC7E9" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium alatulum" order="Filicales" pageId="0" pageNumber="75" rank="species" species="alatulum">
|
||
<emphasis id="1EDF0E9405776D68A7235271F42403B9" italics="true" pageId="0" pageNumber="75">Asplenium alatulum</emphasis>
|
||
</taxonomicName>
|
||
is distinct from
|
||
<taxonomicName id="8981E79E9FC985DA90EEC13049FF49B4" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="B9FBEB1D279DE8420212FBDE6A32C958" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
in having erect to decumbent rhizome (vs. erect rhizome), oblong cells of scales (Fig.
|
||
<figureCitation id="3A3271D96BA019DD5985DCEB2ADEE119" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Scales of A. alatulum, A. subcrenatum and A. wrightii. A, D, G A. wrightii B, E, H A. subcrenatum C, F, I A. alatulum." figureDoi="10.3897/phytokeys.172.62511.figure2" httpUri="https://binary.pensoft.net/fig/509067" pageId="0" pageNumber="75">2C, F, I</figureCitation>
|
||
) (vs. quadrangle cells of scales, Fig.
|
||
<figureCitation id="F5F1F9739B000F196D91062797488C08" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Scales of A. alatulum, A. subcrenatum and A. wrightii. A, D, G A. wrightii B, E, H A. subcrenatum C, F, I A. alatulum." figureDoi="10.3897/phytokeys.172.62511.figure2" httpUri="https://binary.pensoft.net/fig/509067" pageId="0" pageNumber="75">2A, D, G</figureCitation>
|
||
), winged rachis (vs. winged towards apex) and fewer pinna pairs. Phylogenetically, our earlier molecular work resolved
|
||
<taxonomicName id="DF108FD91EC75DFF31A57BF6B32BE34A" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="alatulum">
|
||
<emphasis id="964BFA647AFC57F376A275CB512CF46A" italics="true" pageId="0" pageNumber="75">A. alatulum</emphasis>
|
||
</taxonomicName>
|
||
in a distinct clade, sister to the clade containing
|
||
<taxonomicName id="3AF4E9866469A0D7C0327250C7ED1396" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="AE63188EC5C3527873CEF76CC950A943" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A1AE6E2715CBD3AB0E79B0E116079BA5" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. shikokianum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="shikokianum">
|
||
A.
|
||
<normalizedToken id="163E8DC0CFA7E97D0039C4BCEBE25342" originalValue="×">x</normalizedToken>
|
||
shikokianum
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="0FBE34F6C4095FC0701DACFBEAB28FD1" DOI="https://doi.org/10.1111/cla.12384" author="Xu, KW" journalOrPublisher="Cladistics" pageId="0" pageNumber="75" pagination="22 - 71" refId="B24" refString="Xu, KW, Zhang, L, Rothfels, CJ, Smith, AR, Viane, R, Lorence, D, Wood, KR, Chen, CW, Knapp, R, Zhou, L, Lu, NT, Zhou, X-M, Wei, H-J, Fan, Q, Chen, S-F, Cicuzza, D, Gao, X-F, Liao, W-B, Zhang, L-B, 2020. A global plastid phylogeny of the fern genus Asplenium (Aspleniaceae). Cladistics 36 (1): 22 - 71, DOI: https://doi.org/10.1111/cla.12384" title="A global plastid phylogeny of the fern genus Asplenium (Aspleniaceae)." url="https://doi.org/10.1111/cla.12384" volume="36" year="2020">Xu et al. 2020</bibRefCitation>
|
||
). One accession from the Taiwan Island was nested within
|
||
<taxonomicName id="28A59D049ECFE33A62FD6179BB2E465D" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. alatulum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="alatulum">
|
||
<emphasis id="65CB91DF1CCDE9F4E684BB9C575143C6" italics="true" pageId="0" pageNumber="75">A. alatulum</emphasis>
|
||
</taxonomicName>
|
||
, indicating that this species should also be distributed in Taiwan.
|
||
</paragraph>
|
||
<caption id="CEC091D9E6365CE7F9E0C5DB2CB08C9A" doi="10.3897/phytokeys.172.62511.figure3" httpUri="https://binary.pensoft.net/fig/509068" pageId="0" pageNumber="75" start="Figure 3" startId="F3">
|
||
<paragraph id="F6D021A3891F170F2692019CCC1CCE79" pageId="0" pageNumber="75">
|
||
<emphasis id="F6B47FD34C65EE179438336F3B64854A" bold="true" pageId="0" pageNumber="75">Figure 3.</emphasis>
|
||
Type of
|
||
<taxonomicName id="B2541081F396DB66386F65A61D0BAA87" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium alatulum" order="Filicales" pageId="0" pageNumber="75" rank="species" species="alatulum">
|
||
<emphasis id="2FF2939A390C07D01A72CB26A1F8D2F4" italics="true" pageId="0" pageNumber="75">Asplenium alatulum</emphasis>
|
||
</taxonomicName>
|
||
Ching (
|
||
<emphasis id="1D4FE72A7B5B2AAA5C47674D75D1C1BC" bold="true" pageId="0" pageNumber="75">A</emphasis>
|
||
barcode PE00059412),
|
||
<taxonomicName id="8A905A1A2A6AD0EFF0BE3AC8DA3D7649" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. subcrenatum" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="subcrenatum">
|
||
<emphasis id="1DA1BF49FE2492450FA05E2932FF2DC7" italics="true" pageId="0" pageNumber="75">A. subcrenatum</emphasis>
|
||
</taxonomicName>
|
||
Ching ex S.H.Wu (
|
||
<emphasis id="8A1ABF138828A9989CA5DAD602BD8043" bold="true" pageId="0" pageNumber="75">B</emphasis>
|
||
barcode PE00059494),
|
||
<taxonomicName id="2DF61C7FA56205D9831B07ADEFA3BA62" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="0ACB35CF7ED042F29E9A4F7AA655760D" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
Eaton ex Hooker (
|
||
<emphasis id="A9A6EE5E05C7101728F395E5CDF122EA" bold="true" pageId="0" pageNumber="75">C</emphasis>
|
||
barcode NY00128031) and isotype of
|
||
<taxonomicName id="18636456EA510148EDE1261528A3F67D" class="Filicopsida" family="Aspleniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. wrightii" order="Polypodiales" pageId="0" pageNumber="75" phylum="Pteridophyta" rank="species" species="wrightii">
|
||
<emphasis id="5B527C25380194140EED9507B2A2340E" italics="true" pageId="0" pageNumber="75">A. wrightii</emphasis>
|
||
</taxonomicName>
|
||
GH (
|
||
<emphasis id="A9C775E5BEE8C69A5620A247C7E6CB87" bold="true" pageId="0" pageNumber="75">D</emphasis>
|
||
barcode GH00020612).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="905603DFA519CAB1205D351702A5E202" doi="10.3897/phytokeys.172.62511.figure4" httpUri="https://binary.pensoft.net/fig/509069" pageId="0" pageNumber="75" start="Figure 4" startId="F4">
|
||
<paragraph id="FB7EE354B8D4035261AAB09F0F6475CE" pageId="0" pageNumber="75">
|
||
<emphasis id="57846610503AEE551A352F90A7EAE9B4" bold="true" pageId="0" pageNumber="75">Figure 4.</emphasis>
|
||
<taxonomicName id="CA866500BE3270FF9D465A72AF354649" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" lsidName="Asplenium alatulum" order="Filicales" pageId="0" pageNumber="75" rank="species" species="alatulum">
|
||
<emphasis id="9267F21F687CDE5C47501403A6A73252" italics="true" pageId="0" pageNumber="75">Asplenium alatulum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E73F2B4532DF0E03A80499AD9FDD878D" bold="true" pageId="0" pageNumber="75">A</emphasis>
|
||
Habitat and habit
|
||
<emphasis id="F2A386CA00C2D40D40BF5A197DFDC3AC" bold="true" pageId="0" pageNumber="75">B</emphasis>
|
||
Portion of lamina showing adaxial view of pinna
|
||
<emphasis id="99F5EAA89BDC189369212CA77C4757EB" bold="true" pageId="0" pageNumber="75">C</emphasis>
|
||
Portion of lamina showing abaxial view of pinna
|
||
<emphasis id="32FB7195267AA58C300E494F2B30751D" bold="true" pageId="0" pageNumber="75">D</emphasis>
|
||
Adaxial view of upper portion of lamina
|
||
<emphasis id="E6482AE82528270624C0B14029A47CA0" bold="true" pageId="0" pageNumber="75">E</emphasis>
|
||
Lower portion of rachis showing wide lateral wings.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |