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<document id="05BB4392DD0EE38D7715B50841F6A4BF" ID-DOI="10.5281/zenodo.5330381" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330381" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630167148711" checkinUser="felipe" docAuthor="Zettel, Herbert, Lane, David J. W., Pangantihon, Clister V., Freitag, Hendrik &amp; Be, Jalan Tungku Link" docDate="2012" docId="196E4D54D378FFBAF6D9D8D5FC72DF29" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.29-48.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Enithares vicintricata Lansbury 1968" docType="treatment" docVersion="4" lastPageNumber="43" masterDocId="E557352CD376FFB4F742DB7AFFC6D93D" masterDocTitle="Notes on Notonectidae (Hemiptera: Heteroptera) from southeastern Asia, mostly from Brunei and the Philippines" masterLastPageNumber="48" masterPageNumber="29" pageNumber="43" updateTime="1699171418864" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="294B5F1A564407C7DC6A856F4A8CD64F">Notes on Notonectidae (Hemiptera: Heteroptera) from southeastern Asia, mostly from Brunei and the Philippines</mods:title>
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<mods:namePart id="D818D70E0D3DCFFB57F1359045357E59">Zettel, Herbert</mods:namePart>
<mods:affiliation id="C72E8A65ED5BA1248E135D4187969A49">Natural History Museum, Entomological Department, Burgring 7, 1010 Vienna, Austria; e-mail: herbert. zettel @ nhm-wien. ac. at</mods:affiliation>
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<mods:namePart id="2337B308574A36CBDF61EFD69408BC0F">Lane, David J. W.</mods:namePart>
<mods:affiliation id="B1D19053E838F315B41CC2F94476E0D9">Department of Biology, Universiti Brunei Darussalam, Jalan Tungku Link BE 1410, Brunei Darussalam; e-mail: david. lane @ ubd. edu. bn</mods:affiliation>
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<mods:name id="4FB8FA0E9806DAE8DE3F1C38DABE6B3B" type="personal">
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<mods:namePart id="3CECD16C06496C68D3A436535427DE29">Pangantihon, Clister V.</mods:namePart>
<mods:affiliation id="574212B4253978459D56947B346D5BAE">Biology Department, University of San Carlos, Cebu City 6000, The Philippines; e-mail: clisterpangantihon @ yahoo. com</mods:affiliation>
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<mods:namePart id="2D5CFB628A0FECD3835ADDAB047A9F0F">Freitag, Hendrik</mods:namePart>
<mods:affiliation id="608914D1FB325F5675E98EC690C6E21A">Ateneo de Manila University, Biology Department, School of Science &amp; Engineering, Loyola Heights, Quezon City 1101, The Philippines &amp; Senckenberg Museum of Zoology, Königsbrücker Landstrasse 159, 01109 Dresden, Germany; e-mail: hendrik. freitag @ senckenberg. de</mods:affiliation>
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<mods:namePart id="8C9A01FFD73257170E35B4CEA262A783">Be, Jalan Tungku Link</mods:namePart>
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<mods:title id="BF551EF9793E7CD69C5F45BEDF3DF9B9">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="24138EE02237F41486AF174BBFEBA4B7">2012</mods:date>
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<subSubSection id="D9DDAFC9D378FFBAF6D9D8D5FCA5DAF5" box="[411,867,943,969]" pageId="14" pageNumber="43" type="nomenclature">
<paragraph id="9178FC42D378FFBAF6D9D8D5FCA5DAF5" blockId="14.[411,867,943,969]" box="[411,867,943,969]" pageId="14" pageNumber="43">
<heading id="CA304B2ED378FFBAF6D9D8D5FCA5DAF5" bold="true" box="[411,867,943,969]" centered="true" fontSize="8" level="2" pageId="14" pageNumber="43" reason="3">
<taxonomicName id="56C787C1D378FFBAF6D9D8D5FCA5DAF5" ID-CoL="39VBX" authority="Lansbury, 1968" authorityName="Lansbury" authorityYear="1968" box="[411,867,943,969]" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="vicintricata">
<emphasis id="A3B32050D378FFBAF6D9D8D5FCA5DAF5" bold="true" box="[411,867,943,969]" pageId="14" pageNumber="43">
<emphasis id="A3B32050D378FFBAF6D9D8D5FDC1DAF5" bold="true" box="[411,519,943,968]" italics="true" pageId="14" pageNumber="43">Enithares</emphasis>
cf.
<emphasis id="A3B32050D378FFBAF572D8CAFD6ADAF5" bold="true" box="[560,684,944,968]" italics="true" pageId="14" pageNumber="43">vicintricata</emphasis>
<bibRefCitation id="F55681B3D378FFBAF5F1D8CBFCA5DAF5" author="LANSBURY I." box="[691,867,944,969]" pageId="14" pageNumber="43" pagination="353 - 442" refId="ref15337" refString="LANSBURY I. 1968: The Enithares (Hemiptera-Heteroptera: Notonectidae) of the Oriental Region. Pacific Insects 10: 353 - 442." type="journal article" year="1968">Lansbury, 1968</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9DDAFC9D378FFBAF7D5D8ADFDCBDAD4" box="[151,525,982,1001]" pageId="14" pageNumber="43" type="reference_group">
<paragraph id="9178FC42D378FFBAF7D5D8ADFDCBDAD4" blockId="14.[151,525,982,1001]" box="[151,525,982,1001]" pageId="14" pageNumber="43">
<treatmentCitationGroup id="B1D7DB6CD378FFBAF7D5D8ADFDCBDAD4" box="[151,525,982,1001]" pageId="14" pageNumber="43">
<treatmentCitation id="1066DA53D378FFBAF7D5D8ADFDCFDAD4" author="LANSBURY I." box="[151,521,982,1001]" page="430" pageId="14" pageNumber="43" year="1968">
<taxonomicName id="56C787C1D378FFBAF7D5D8ADFDCFDAD4" ID-CoL="39VBX" authority="Lansbury, 1968: 430" authorityName="Lansbury" authorityPageNumber="430" authorityYear="1968" box="[151,521,982,1001]" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="vicintricata">
<emphasis id="A3B32050D378FFBAF7D5D8ADFE97DAD4" box="[151,337,983,1001]" italics="true" pageId="14" pageNumber="43">Enithares vicintricata</emphasis>
<bibRefCitation id="F55681B3D378FFBAF614D8ACFDCFDAD4" author="LANSBURY I." box="[342,521,982,1001]" pageId="14" pageNumber="43" pagination="353 - 442" refId="ref15337" refString="LANSBURY I. 1968: The Enithares (Hemiptera-Heteroptera: Notonectidae) of the Oriental Region. Pacific Insects 10: 353 - 442." type="journal article" year="1968">Lansbury, 1968: 430</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
</subSubSection>
<subSubSection id="D9DDAFC9D378FFBAF7D5D885FC72DF29" pageId="14" pageNumber="43" type="materials_examined">
<paragraph id="9178FC42D378FFBAF7D5D885FC5EDD5E" blockId="14.[151,1127,1023,1124]" pageId="14" pageNumber="43">
<emphasis id="A3B32050D378FFBAF7D5D885FDFBDD2E" bold="true" box="[151,573,1023,1043]" pageId="14" pageNumber="43">
Material examined.
<collectingCountry id="E9D0BCD2D378FFBAF609DF7AFE67DD2E" box="[331,417,1024,1043]" name="Brunei" pageId="14" pageNumber="43">BRUNEI</collectingCountry>
:
<collectingRegion id="530332A0D378FFBAF6EBDF7AFE23DD2F" box="[425,485,1024,1043]" country="Brunei" name="Belait" pageId="14" pageNumber="43">BELAIT</collectingRegion>
DISTRICT:
</emphasis>
Labi, Rampayar, Sungai Menderam,
<geoCoordinate id="F4F39A85D378FFBAF434DF7AFC26DD29" box="[886,992,1023,1044]" degrees="4" direction="north" minutes="20" orientation="latitude" pageId="14" pageNumber="43" precision="1" seconds="30.7" value="4.3418612">4°2030.7″N</geoCoordinate>
<geoCoordinate id="F4F39A85D378FFBAF4A6DF7AFBA5DD29" box="[996,1123,1023,1044]" degrees="114" direction="east" minutes="26" orientation="longitude" pageId="14" pageNumber="43" precision="1" seconds="55.2" value="114.44867">114°2655.2″E</geoCoordinate>
,
<date id="E579DA82D378FFBAF7D5DF61FF2BDD13" box="[151,237,1051,1070]" pageId="14" pageNumber="43" value="1996-05-19">19.v.1996</date>
, leg. H. H. Tan, Y. Y. Goh, H. H. Ng &amp; D. Yeo (THH9634, ZRC 6.18843), 1 J
<specimenCount id="87C137CBD378FFBAF4C1DF61FC67DD13" box="[899,929,1045,1070]" pageId="14" pageNumber="43" type="female">1 ♀</specimenCount>
(
<collectionCode id="F7D66487D378FFBAF4EEDF61FC20DD13" box="[940,998,1051,1070]" pageId="14" pageNumber="43">ZRCS</collectionCode>
). Peat-swampy area draining to Sungai Ingei,
<date id="E579DA82D378FFBAF6DEDF4FFE28DD75" box="[412,494,1077,1096]" pageId="14" pageNumber="43" value="2010-06-06">6.vi.2010</date>
, leg. D. J. W. Lane (15), 6 JJ
<specimenCount id="87C137CBD378FFBAF5BFDF4CFCEBDD75" box="[765,813,1071,1097]" pageId="14" pageNumber="43" type="female">4 ♀♀</specimenCount>
(
<collectionCode id="F7D66487D378FFBAF47ADF4CFCBEDD74" box="[824,888,1078,1097]" pageId="14" pageNumber="43">UBDB</collectionCode>
). Sungai Ingei, forest pool,
<geoCoordinate id="F4F39A85D378FFBAF7D5DF2AFF3FDD59" box="[151,249,1103,1124]" degrees="4" direction="north" minutes="08.921" orientation="latitude" pageId="14" pageNumber="43" precision="1" value="4.1486835">4°08.921N</geoCoordinate>
<geoCoordinate id="F4F39A85D378FFBAF7BCDF2AFEB3DD59" box="[254,373,1103,1124]" degrees="114" direction="east" minutes="43.364" orientation="longitude" pageId="14" pageNumber="43" precision="1" value="114.72273">114°43.364E</geoCoordinate>
,
<date id="E579DA82D378FFBAF63FDF2AFE16DD5E" box="[381,464,1104,1123]" pageId="14" pageNumber="43" value="2010-06-08">8.vi.2010</date>
, leg. D. J. W. Lane (#22), 1 J
<specimenCount id="87C137CBD378FFBAF597DF2AFD33DD5E" box="[725,757,1098,1123]" pageId="14" pageNumber="43" type="female">1 ♀</specimenCount>
(
<collectionCode id="F7D66487D378FFBAF5BDDF2AFC81DD5E" box="[767,839,1104,1123]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="43" type="Museum">NHMW</collectionCode>
,
<collectionCode id="F7D66487D378FFBAF40DDF2AFC48DD5E" box="[847,910,1104,1123]" pageId="14" pageNumber="43">UBDB</collectionCode>
).
</paragraph>
<paragraph id="9178FC42D378FFBAF7D5DF06FC72DF29" blockId="14.[151,1127,1148,1556]" pageId="14" pageNumber="43">
<emphasis id="A3B32050D378FFBAF7D5DF06FF18DDA9" bold="true" box="[151,222,1148,1172]" pageId="14" pageNumber="43">Notes.</emphasis>
<taxonomicName id="56C787C1D378FFBAF7A7DF07FE08DDA9" authorityName="Lansbury" authorityYear="1968" box="[229,462,1149,1172]" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="vicintricata">
<emphasis id="A3B32050D378FFBAF7A7DF07FE08DDA9" box="[229,462,1149,1172]" italics="true" pageId="14" pageNumber="43">Enithares vicintricata</emphasis>
</taxonomicName>
was described from
<specimenCount id="87C137CBD378FFBAF5EDDF06FCDADDA9" box="[687,796,1148,1172]" pageId="14" pageNumber="43" type="male">two males</specimenCount>
and
<specimenCount id="87C137CBD378FFBAF410DF06FC0FDDA9" box="[850,969,1148,1172]" pageId="14" pageNumber="43" type="female">one female</specimenCount>
from
<collectingCountry id="E9D0BCD2D378FFBAF349DF06FBA1DDA9" box="[1035,1127,1148,1172]" name="Malaysia" pageId="14" pageNumber="43">Sarawak</collectingCountry>
by
<bibRefCitation id="F55681B3D378FFBAF7F8DFE6FEB5DD8E" author="LANSBURY I." box="[186,371,1179,1203]" pageId="14" pageNumber="43" pagination="353 - 442" refId="ref15337" refString="LANSBURY I. 1968: The Enithares (Hemiptera-Heteroptera: Notonectidae) of the Oriental Region. Pacific Insects 10: 353 - 442." type="journal article" year="1968">LANSBURY (1968)</bibRefCitation>
and is regarded as an endemic species of Borneo (
<bibRefCitation id="F55681B3D378FFBAF4E0DFE6FB9ADD8E" author="CHEN P. - P. &amp; NIESER N. &amp; ZETTEL H." box="[930,1116,1179,1204]" pageId="14" pageNumber="43" refId="ref14748" refString="CHEN P. - P., NIESER N. &amp; ZETTEL H. 2005: The aquatic and semi-aquatic bugs (Heteroptera: Nepomorpha &amp; Gerromorpha) of Malesia. Fauna Malesiana Handbooks. Vol. 5. Brill, Leiden - Boston, 546 pp." type="book" year="2005">CHEN et al. 2005</bibRefCitation>
). Similar, closely related species live on Java and Sumatra (
<taxonomicName id="56C787C1D378FFBAF5B9DFC7FBD4DDEE" authority="Breddin, 1905" authorityName="Breddin" authorityYear="1905" box="[763,1042,1211,1236]" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="intricata">
<emphasis id="A3B32050D378FFBAF5B9DFC7FCCDDDE9" box="[763,779,1213,1236]" italics="true" pageId="14" pageNumber="43">E</emphasis>
.
<emphasis id="A3B32050D378FFBAF455DFC7FCB5DDE9" box="[791,883,1213,1236]" italics="true" pageId="14" pageNumber="43">intricata</emphasis>
Breddin, 1905
</taxonomicName>
) and on the Andaman Islands (
<taxonomicName id="56C787C1D378FFBAF6CBDFA7FD48DDC9" authority="Distant, 1910" authorityName="Distant" authorityYear="1910" box="[393,654,1244,1268]" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="rogersi">
<emphasis id="A3B32050D378FFBAF6CBDFA7FE5FDDC9" box="[393,409,1245,1268]" italics="true" pageId="14" pageNumber="43">E</emphasis>
.
<emphasis id="A3B32050D378FFBAF6EADFA7FE32DDC9" box="[424,500,1245,1268]" italics="true" pageId="14" pageNumber="43">rogersi</emphasis>
<bibRefCitation id="F55681B3D378FFBAF6B9DFA6FD48DDC9" author="DISTANT W. L." box="[507,654,1244,1268]" pageId="14" pageNumber="43" refId="ref14841" refString="DISTANT W. L. 1910: Rhynchota V. Heteroptera: Appendix. In: SHIPLEY A. E. &amp; MARSHALL G. A. K. (eds.): The fauna of British India including Ceylon and Burma. Taylor &amp; Francis, London, xii + 362 pp." type="book" year="1910">Distant, 1910</bibRefCitation>
</taxonomicName>
) (see
<bibRefCitation id="F55681B3D378FFBAF58DDFA6FCBEDDCE" author="LANSBURY I." box="[719,888,1243,1267]" pageId="14" pageNumber="43" pagination="353 - 442" refId="ref15337" refString="LANSBURY I. 1968: The Enithares (Hemiptera-Heteroptera: Notonectidae) of the Oriental Region. Pacific Insects 10: 353 - 442." type="journal article" year="1968">LANSBURY 1968</bibRefCitation>
). The specimens from
<collectingCountry id="E9D0BCD2D378FFBAF7D5DF86FF26DC29" box="[151,224,1276,1300]" name="Brunei" pageId="14" pageNumber="43">Brunei</collectingCountry>
differ slightly from the
<typeStatus id="4E7C42E0D378FFBAF69DDF86FDD1DC29" box="[479,535,1276,1300]" pageId="14" pageNumber="43">types</typeStatus>
of
<taxonomicName id="56C787C1D378FFBAF578DF87FD15DC29" authorityName="Lansbury" authorityYear="1968" box="[570,723,1277,1300]" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="vicintricata">
<emphasis id="A3B32050D378FFBAF578DF87FD15DC29" box="[570,723,1277,1300]" italics="true" pageId="14" pageNumber="43">E. vicintricata</emphasis>
</taxonomicName>
, e.g., by smaller size (
<quantity id="563F51A7D378FFBAF482DF86FB87DC2E" box="[960,1089,1276,1300]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="7.9" metricValueMin="7.3" pageId="14" pageNumber="43" unit="mm" value="7.6" valueMax="7.9" valueMin="7.3">7.37.9 mm</quantity>
vs. 9.0 mm in the
<typeStatus id="4E7C42E0D378FFBAF676DE66FEB7DC09" box="[308,369,1308,1332]" pageId="14" pageNumber="43">types</typeStatus>
); male with three stout setae on the first mesotarsomere (vs. two in the
<typeStatus id="4E7C42E0D378FFBAF7D5DE46FF09DC69" box="[151,207,1340,1364]" pageId="14" pageNumber="43">types</typeStatus>
of
<taxonomicName id="56C787C1D378FFBAF7B6DE47FE48DC69" authorityName="Lansbury" authorityYear="1968" box="[244,398,1340,1364]" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="vicintricata">
<emphasis id="A3B32050D378FFBAF7B6DE47FEC2DC69" box="[244,260,1341,1364]" italics="true" pageId="14" pageNumber="43">E</emphasis>
.
<emphasis id="A3B32050D378FFBAF651DE47FE48DC69" box="[275,398,1341,1364]" italics="true" pageId="14" pageNumber="43">vicintricata</emphasis>
</taxonomicName>
and four in
<taxonomicName id="56C787C1D378FFBAF551DE47FD4BDC69" authorityName="Breddin" authorityYear="1905" box="[531,653,1340,1364]" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="intricata">
<emphasis id="A3B32050D378FFBAF551DE47FDE5DC69" box="[531,547,1341,1364]" italics="true" pageId="14" pageNumber="43">E</emphasis>
.
<emphasis id="A3B32050D378FFBAF573DE47FD4BDC69" box="[561,653,1341,1364]" italics="true" pageId="14" pageNumber="43">intricata</emphasis>
</taxonomicName>
; fide
<bibRefCitation id="F55681B3D378FFBAF589DE46FCB4DC6E" author="LANSBURY I." box="[715,882,1339,1363]" pageId="14" pageNumber="43" pagination="353 - 442" refId="ref15337" refString="LANSBURY I. 1968: The Enithares (Hemiptera-Heteroptera: Notonectidae) of the Oriental Region. Pacific Insects 10: 353 - 442." type="journal article" year="1968">LANSBURY 1968</bibRefCitation>
) and basal plate of the phallus bluntly produced posteroapically (more acutely produced in all other forms). Specimens similar to those of the
<collectingCountry id="E9D0BCD2D378FFBAF680DE06FDCDDCA9" box="[450,523,1404,1428]" name="Brunei" pageId="14" pageNumber="43">Brunei</collectingCountry>
population are known from several locations in
<collectingCountry id="E9D0BCD2D378FFBAF349DE06FBA1DCA9" box="[1035,1127,1404,1428]" name="Malaysia" pageId="14" pageNumber="43">Sarawak</collectingCountry>
(specimens in
<collectionCode id="F7D66487D378FFBAF66CDEE6FE40DC8E" box="[302,390,1436,1459]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="43" type="Museum">NHMW</collectionCode>
, unpublished), but it seems that there is more than one species of the
<taxonomicName id="56C787C1D378FFBAF30DDEE7FED4DCE9" authorityName="Lansbury" authorityYear="1968" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="vicintricata">
<emphasis id="A3B32050D378FFBAF30DDEE7FED4DCE9" italics="true" pageId="14" pageNumber="43">E. vicintricata</emphasis>
</taxonomicName>
complex in western Borneo. A careful analysis on the population structure of
<taxonomicName id="56C787C1D378FFBAF30DDEC7FED4DCC9" authorityName="Lansbury" authorityYear="1968" class="Insecta" family="Notonectidae" genus="Enithares" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="vicintricata">
<emphasis id="A3B32050D378FFBAF30DDEC7FED4DCC9" italics="true" pageId="14" pageNumber="43">E. vicintricata</emphasis>
</taxonomicName>
will be required before drawing further taxonomic conclusions. For that reason, the identification of the
<collectingCountry id="E9D0BCD2D378FFBAF6D4DE86FE19DF29" box="[406,479,1532,1556]" name="Brunei" pageId="14" pageNumber="43">Brunei</collectingCountry>
material should be regarded as preliminary.
</paragraph>
</subSubSection>
</treatment>
</document>