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<document id="C1D77A7CF208915D1A88EB3641DA9C9D" ID-DOI="10.1080/00222930600628416" ID-ISSN="1464-5262" ID-Zenodo-Dep="4669378" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617793766532" checkinUser="carolina" docAuthor="Sanamyan, K. E. &amp; Sanamyan, N. P." docDate="2006" docId="1925951FFFEAE84B704DFC7AFB9B9B94" docLanguage="en" docName="JNATHIST.40.5-6.307-344.pdf" docOrigin="Journal of Natural History 40 (5 - 6)" docSource="http://dx.doi.org/10.1080/00222930600628416" docStyle="DocumentStyle:5A9EB2ED5589A523F0236C66DE2A6BA8.4:JNatHist.2005-2007.journal_article.1cover" docStyleId="5A9EB2ED5589A523F0236C66DE2A6BA8" docStyleName="JNatHist.2005-2007.journal_article.1cover" docStyleVersion="4" docTitle="Chelyosoma columbianum Huntsman 1912" docType="treatment" docVersion="3" lastPageNumber="309" masterDocId="E51CED67FFE9E8487132FFDAFFD29D01" masterDocTitle="Deep-water ascidians (Tunicata: Ascidiacea) from the northern and western Pacific" masterLastPageNumber="344" masterPageNumber="307" pageNumber="309" updateTime="1698902202717" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="898979D2389C9024E84AC605B885F1F0">Deep-water ascidians (Tunicata: Ascidiacea) from the northern and western Pacific</mods:title>
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<mods:namePart id="AD3F6679C24DBD8C802F435D245AEE79">Sanamyan, K. E.</mods:namePart>
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<mods:title id="A8C825D97F02ACD3D495F07A96FF5273">Journal of Natural History</mods:title>
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<mods:date id="EEB663447942B3831462634BE5F8C238">2006</mods:date>
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<paragraph id="91332409FFEAE84B704DFC7AFC7E9EB9" blockId="3.[138,1185,928,1015]" box="[383,940,928,952]" pageId="3" pageNumber="309">
<heading id="CA7B9365FFEAE84B704DFC7AFC7E9EB9" box="[383,940,928,952]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="309" reason="2">
<taxonomicName id="568C5F8AFFEAE84B704DFC7AFC7E9EB9" authority="Huntsman, 1912" authorityName="Huntsman" authorityYear="1912" box="[383,940,928,952]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="columbianum">
<emphasis id="A3F8F81BFFEAE84B704DFC7AFD0E9EB9" bold="true" box="[383,732,928,952]" italics="true" pageId="3" pageNumber="309">Chelyosoma columbianum</emphasis>
<bibRefCitation id="F51D59F8FFEAE84B73D7FC7AFC7E9EB9" author="Huntsman AG" box="[741,940,928,952]" pageId="3" pageNumber="309" pagination="111 - 148" refId="ref13034" refString="Huntsman AG. 1912. Ascidians from the coasts of Canada. Transactions of the Royal Canadian Institute 9: 111 - 148." type="journal article" year="1912">Huntsman, 1912</bibRefCitation>
</taxonomicName>
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<paragraph id="91332409FFEAE84B71B8FC1AFDDB9EF6" blockId="3.[138,1185,928,1015]" pageId="3" pageNumber="309">
<taxonomicName id="568C5F8AFFEAE84B71B8FC1AFD1C9ED9" authority="Huntsman 1912, p 124" authorityName="Huntsman" authorityPageNumber="124" authorityYear="1912" box="[138,718,960,984]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="columbianum">
<emphasis id="A3F8F81BFFEAE84B71B8FC1AFE789ED9" box="[138,426,960,984]" italics="true" pageId="3" pageNumber="309">Chelyosoma columbianum</emphasis>
<bibRefCitation id="F51D59F8FFEAE84B7087FC1AFDA99ED9" author="Huntsman AG" box="[437,635,960,984]" pageId="3" pageNumber="309" pagination="111 - 148" refId="ref13034" refString="Huntsman AG. 1912. Ascidians from the coasts of Canada. Transactions of the Royal Canadian Institute 9: 111 - 148." type="journal article" year="1912">Huntsman 1912</bibRefCitation>
, p 124
</taxonomicName>
;
<bibRefCitation id="F51D59F8FFEAE84B73D2FC1AFCBE9ED9" author="Ritter WE" box="[736,876,960,984]" pageId="3" pageNumber="309" pagination="427 - 509" refId="ref14440" refString="Ritter WE. 1913. The simple ascidians from the North-eastern Pacific in the collection of the United States National Museum. Proceedings of the United States National Museum 45 (1989): 427 - 509." type="journal article" year="1913">Ritter 1913</bibRefCitation>
, p 486;
<bibRefCitation id="F51D59F8FFEAE84B72E2FC1AFB4E9ED9" author="Van Name WG" box="[976,1180,960,984]" pageId="3" pageNumber="309" pagination="1 - 476" refId="ref14863" refString="Van Name WG. 1945. The North and South American ascidians. Bulletin of the American Museum of Natural History 84: 1 - 476." type="journal article" year="1945">Van Name 1945</bibRefCitation>
, p 208;
<bibRefCitation id="F51D59F8FFEAE84B71C7FC05FE669EF6" author="Sanamyan K &amp; Sanamyan N" box="[245,436,991,1015]" pageId="3" pageNumber="309" pagination="209 - 214" refId="ref14575" refString="Sanamyan K, Sanamyan N. 1998. Some deep-water ascidians from the NW Pacific (Tunicata: Ascidiacea). Zoosystematica Rossica 7: 209 - 214." type="journal article" year="1998">Sanamyan 1998</bibRefCitation>
, p 110.
</paragraph>
</subSubSection>
<subSubSection id="D9967782FFEAE84B71B8FBF5FB9B9B94" pageId="3" pageNumber="309" type="materials_examined">
<paragraph id="91332409FFEAE84B71B8FBF5FE8A9946" blockId="3.[138,344,1071,1095]" box="[138,344,1071,1095]" pageId="3" pageNumber="309">
<emphasis id="A3F8F81BFFEAE84B71B8FBF5FE8A9946" box="[138,344,1071,1095]" italics="true" pageId="3" pageNumber="309">Material examined</emphasis>
</paragraph>
<paragraph id="91332409FFEAE84B71B8FB85FE2F9976" blockId="3.[138,509,1119,1143]" box="[138,509,1119,1143]" pageId="3" pageNumber="309">
St.
<quantity id="567489ECFFEAE84B7181FB85FE9B9976" box="[179,329,1119,1143]" metricMagnitude="6" metricUnit="m" metricValue="6.08917" pageId="3" pageNumber="309" unit="m" value="6089170.0">6089, 170 m</quantity>
,
<specimenCount id="878AEF80FFEAE84B7065FB85FE2A9976" box="[343,504,1119,1143]" pageId="3" pageNumber="309" type="generic">25 specimens</specimenCount>
.
</paragraph>
<paragraph id="91332409FFEAE84B71B8FB75FF3B99C6" blockId="3.[138,233,1199,1223]" box="[138,233,1199,1223]" pageId="3" pageNumber="309">
<emphasis id="A3F8F81BFFEAE84B71B8FB75FF3B99C6" box="[138,233,1199,1223]" italics="true" pageId="3" pageNumber="309">Remarks</emphasis>
</paragraph>
<paragraph id="91332409FFEAE84B71B8FB05FB9B9B94" blockId="3.[138,1185,1247,1685]" pageId="3" pageNumber="309">
All newly recorded specimens are identical to the one from the Commander Islands (
<bibRefCitation id="F51D59F8FFEAE84B71A0FB24FE879817" author="Sanamyan K &amp; Sanamyan N" box="[146,341,1278,1302]" pageId="3" pageNumber="309" pagination="209 - 214" refId="ref14575" refString="Sanamyan K, Sanamyan N. 1998. Some deep-water ascidians from the NW Pacific (Tunicata: Ascidiacea). Zoosystematica Rossica 7: 209 - 214." type="journal article" year="1998">Sanamyan 1998</bibRefCitation>
). They all have two central plates connected by short muscles, and 811 marginal plates. None of the 25 examined specimens had intermediate plates that
<bibRefCitation id="F51D59F8FFEAE84B71B8FAE4FE8C9857" author="Huntsman AG" box="[138,350,1342,1366]" pageId="3" pageNumber="309" pagination="111 - 148" refId="ref13034" refString="Huntsman AG. 1912. Ascidians from the coasts of Canada. Transactions of the Royal Canadian Institute 9: 111 - 148." type="journal article" year="1912">Huntsman (1912)</bibRefCitation>
recorded for this species.
<bibRefCitation id="F51D59F8FFEAE84B73BDFAE4FCB09857" author="Huntsman AG" box="[655,866,1342,1366]" pageId="3" pageNumber="309" pagination="111 - 148" refId="ref13034" refString="Huntsman AG. 1912. Ascidians from the coasts of Canada. Transactions of the Royal Canadian Institute 9: 111 - 148." type="journal article" year="1912">Huntsman (1912)</bibRefCitation>
stated that
<taxonomicName id="568C5F8AFFEAE84B72DEFAE4FB729857" authorityName="Huntsman" authorityYear="1912" box="[1004,1184,1342,1366]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="columbianum">
<emphasis id="A3F8F81BFFEAE84B72DEFAE4FB729857" box="[1004,1184,1342,1366]" italics="true" pageId="3" pageNumber="309">C. columbianum</emphasis>
</taxonomicName>
may be readily distinguished from
<taxonomicName id="568C5F8AFFEAE84B7311FA84FCAF9877" authority="Stimpson, 1864" authorityName="Stimpson" authorityYear="1864" box="[547,893,1374,1398]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="productum">
<emphasis id="A3F8F81BFFEAE84B7311FA84FD6B9877" box="[547,697,1374,1398]" italics="true" pageId="3" pageNumber="309">C. productum</emphasis>
<bibRefCitation id="F51D59F8FFEAE84B73F1FA84FCAF9877" author="Stimpson W" box="[707,893,1374,1398]" pageId="3" pageNumber="309" pagination="153 - 161" refId="ref14745" refString="Stimpson W. 1864. Descriptions of new species of marine invertebrata from Puget Sound collected by the naturalists of the Northwest Boundary Commission. Proceedings of the Academy of Natural Sciences of Philadelphia 16: 153 - 161." type="journal article" year="1864">Stimpson, 1864</bibRefCitation>
</taxonomicName>
by the presence of short muscles connecting the two central plates.
<bibRefCitation id="F51D59F8FFEAE84B7344FAA4FCDD9897" author="Ritter WE" box="[630,783,1406,1430]" pageId="3" pageNumber="309" pagination="427 - 509" refId="ref14440" refString="Ritter WE. 1913. The simple ascidians from the North-eastern Pacific in the collection of the United States National Museum. Proceedings of the United States National Museum 45 (1989): 427 - 509." type="journal article" year="1913">Ritter (1913)</bibRefCitation>
based his assignation of specimens from the Strait of Juan de Fuca to
<taxonomicName id="568C5F8AFFEAE84B7302FA44FD3598B7" authorityName="Huntsman" authorityYear="1912" box="[560,743,1438,1462]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="columbianum">
<emphasis id="A3F8F81BFFEAE84B7302FA44FD3598B7" box="[560,743,1438,1462]" italics="true" pageId="3" pageNumber="309">C. columbianum</emphasis>
</taxonomicName>
solely on this feature. However, it is possible that the muscles between the central plates have been overlooked in those specimens assigned to
<taxonomicName id="568C5F8AFFEAE84B7083FA04FD8398F7" authorityName="Stimpson" authorityYear="1864" box="[433,593,1502,1526]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="productum">
<emphasis id="A3F8F81BFFEAE84B7083FA04FD8398F7" box="[433,593,1502,1526]" italics="true" pageId="3" pageNumber="309">C. productum</emphasis>
</taxonomicName>
and it may not differ from
<taxonomicName id="568C5F8AFFEAE84B72E8FA04FB4B98F7" authorityName="Huntsman" authorityYear="1912" box="[986,1177,1502,1526]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="columbianum">
<emphasis id="A3F8F81BFFEAE84B72E8FA04FB4B98F7" box="[986,1177,1502,1526]" italics="true" pageId="3" pageNumber="309">C. columbianum</emphasis>
</taxonomicName>
. Unfortunately no specimens clearly referable to
<taxonomicName id="568C5F8AFFEAE84B73F9FA27FCB19B14" authorityName="Stimpson" authorityYear="1864" box="[715,867,1533,1557]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="productum">
<emphasis id="A3F8F81BFFEAE84B73F9FA27FCB19B14" box="[715,867,1533,1557]" italics="true" pageId="3" pageNumber="309">C. productum</emphasis>
</taxonomicName>
(with elongated body and more numerous marginal plates) were available to confirm this view. The present species cannot be confused with the other north Pacific species of
<taxonomicName id="568C5F8AFFEAE84B7203F9E7FC609B54" authorityName="Broderip &amp; Sowerby I" authorityYear="1830" box="[817,946,1597,1621]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis id="A3F8F81BFFEAE84B7203F9E7FC609B54" box="[817,946,1597,1621]" italics="true" pageId="3" pageNumber="309">Chelyosoma</emphasis>
</taxonomicName>
, which either have a single central plate (
<taxonomicName id="568C5F8AFFEAE84B70A2F987FC1C9B74" authority="Broderip and Sowerby, 1830" authorityName="Broderip and Sowerby" authorityYear="1830" box="[400,974,1629,1653]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="macleayanum">
<emphasis id="A3F8F81BFFEAE84B70A2F987FD809B74" box="[400,594,1629,1653]" italics="true" pageId="3" pageNumber="309">C. macleayanum</emphasis>
Broderip and Sowerby, 1830
</taxonomicName>
and
<taxonomicName id="568C5F8AFFEAE84B7528F987FE899B94" authority="Redikorzev, 1911" authorityName="Redikorzev" authorityYear="1911" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="orientale">
<emphasis id="A3F8F81BFFEAE84B7528F987FB739B74" box="[1050,1185,1629,1653]" italics="true" pageId="3" pageNumber="309">C. orientale</emphasis>
Redikorzev, 1911
</taxonomicName>
), or lack the plates on the disk (
<taxonomicName id="568C5F8AFFEAE84B73EAF9A7FBEE9B94" authority="Redikorzev, 1913" authorityName="Redikorzev" authorityYear="1913" box="[728,1084,1661,1685]" class="Ascidiacea" family="Corellidae" genus="Chelyosoma" kingdom="Animalia" order="Enterogona" pageId="3" pageNumber="309" phylum="Chordata" rank="species" species="inaequale">
<emphasis id="A3F8F81BFFEAE84B73EAF9A7FCB09B94" box="[728,866,1661,1685]" italics="true" pageId="3" pageNumber="309">C. inaequale</emphasis>
Redikorzev, 1913
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
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