283 lines
33 KiB
XML
283 lines
33 KiB
XML
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<mods:title id="CE3DD25B2D305918EA7418241D580D4B">Four new species of the genus Trichodromeus Luze, 1903 from Nepal and Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="808F9C08D96E5EB4E0214807AF9B0F0F">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="B403CF40A3AA772F28681B6ABAE54D4B">Zootaxa</mods:title>
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<mods:date id="82591BA37FAA21433447289D8A00214B">2021</mods:date>
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<mods:number id="1164B23E1C7D176EE9EA4C528387D7FB">4991</mods:number>
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<treatment id="0579878DFB2B9278FF23F8E9E07BD3DB" ID-DOI="http://doi.org/10.5281/zenodo.5916679" ID-GBIF-Taxon="192745370" ID-Zenodo-Dep="5916679" LSID="urn:lsid:plazi:treatment:0579878DFB2B9278FF23F8E9E07BD3DB" httpUri="http://treatment.plazi.org/id/0579878DFB2B9278FF23F8E9E07BD3DB" lastPageId="6" lastPageNumber="359" pageId="4" pageNumber="357">
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<subSubSection id="C5CA6510FB2B927AFF23F8E9E76AD72B" box="[151,607,1878,1905]" pageId="4" pageNumber="357" type="nomenclature">
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<paragraph id="8D6F369BFB2B927AFF23F8E9E76AD72B" blockId="4.[151,607,1878,1941]" box="[151,607,1878,1905]" pageId="4" pageNumber="357">
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<heading id="D62781F7FB2B927AFF23F8E9E76AD72B" bold="true" box="[151,607,1878,1905]" fontSize="11" level="1" pageId="4" pageNumber="357" reason="1">
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<emphasis id="BFA4EA89FB2B927AFF23F8E9E76AD72B" bold="true" box="[151,607,1878,1905]" pageId="4" pageNumber="357">
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<taxonomicName id="4AD04D18FB2B927AFF23F8E9E722D72A" ID-CoL="8YBYM" authority="Shavrin" authorityName="Shavrin" authorityYear="2021" box="[151,535,1878,1905]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="357" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
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<emphasis id="BFA4EA89FB2B927AFF23F8E9E49FD72B" bold="true" box="[151,426,1878,1905]" italics="true" pageId="4" pageNumber="357">Trichodromeus jaegeri</emphasis>
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Shavrin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A49757F2FB2B927AFD90F8E8E76AD72B" box="[548,607,1879,1905]" pageId="4" pageNumber="357" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C5CA6510FB2B927AFF23F8C4E462D7CF" box="[151,343,1915,1941]" pageId="4" pageNumber="357" type="description">
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<paragraph id="8D6F369BFB2B927AFF23F8C4E462D7CF" blockId="4.[151,607,1878,1941]" box="[151,343,1915,1941]" pageId="4" pageNumber="357">
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(
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<figureCitation id="15EB2A1EFB2B927AFF2BF8C4E5DAD7CF" box="[159,239,1915,1941]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[168,1419,192,1909]" captionTargetId="figure-28@2.[152,1436,179,1920]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Trichodromeus: 1—T. acutipenis (holotype), 2—T. jaegeri (paratype), 3—T. kaliyuga (holotype), 4—T. trilobatus (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5030540" httpUri="https://zenodo.org/record/5030540/files/figure.png" pageId="4" pageNumber="357">Figs. 2</figureCitation>
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,
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<figureCitation id="15EB2A1EFB2B927AFF4FF8C4E412D7CF" box="[251,295,1915,1941]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[173,1420,193,1912]" captionTargetId="figure-16@3.[155,1432,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Trichodromeus (5–6: T. acutipenis (holotype), 7–8: T. jaegeri (holotype), 9–10: T. kaliyuga (holotype): 5, 7, 9–parameral view, 6, 8, 10 lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5030542" httpUri="https://zenodo.org/record/5030542/files/figure.png" pageId="4" pageNumber="357">7–8</figureCitation>
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,
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<figureCitation id="15EB2A1EFB2B927AFE87F8C4E47BD7CF" box="[307,334,1915,1941]" captionStart="FIGURE 11" captionStartId="5.[151,250,1679,1703]" captionTargetBox="[151,1436,704,1656]" captionTargetId="figure-216@5.[151,1436,704,1656]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of Trichodromeus in Nepal: T. acutipenis (circles), T. jaegeri (square), T. kaliyuga (triangle)." figureDoi="http://doi.org/10.5281/zenodo.5030544" httpUri="https://zenodo.org/record/5030544/files/figure.png" pageId="4" pageNumber="357">11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C5CA6510FB2B927BFF23F87CE77ED13E" lastPageId="5" lastPageNumber="358" pageId="4" pageNumber="357" type="materials_examined">
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<paragraph id="8D6F369BFB2B927AFF23F87CE0A8D787" blockId="4.[151,1437,1986,2013]" box="[151,1437,1986,2013]" pageId="4" pageNumber="357">
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<emphasis id="BFA4EA89FB2B927AFF23F87CE4F1D786" bold="true" box="[151,452,1986,2013]" pageId="4" pageNumber="357">Type material examined:</emphasis>
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<materialsCitation id="3DB83CC6FB2B927AFE79F87CE0A8D787" ID-GBIF-Occurrence="3459583301" box="[461,1437,1986,2013]" country="Nepal" location="Dhaulagiri-Himal" municipality="Yak Kharkra" pageId="4" pageNumber="357" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="526B8839FB2B927AFE79F87CE70DD787" box="[461,568,1987,2013]" pageId="4" pageNumber="357" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="9BD6FD12FB2B927AFDF0F87CE76CD787" box="[580,601,1987,2013]" pageId="4" pageNumber="357" type="male">♂</specimenCount>
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[dissected]: ‘
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<collectingCountry id="F5C7760BFB2B927AFD44F87CE665D787" box="[752,848,1987,2013]" name="Nepal" pageId="4" pageNumber="357">NEPAL</collectingCountry>
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,
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<location id="880F6040FB2B927AFCE8F87CE116D787" LSID="urn:lsid:plazi:treatment:0579878DFB2B9278FF23F8E9E07BD3DB:880F6040FB2B927AFCE8F87CE116D787" box="[860,1059,1987,2013]" country="Nepal" municipality="Yak Kharkra" name="Dhaulagiri-Himal" pageId="4" pageNumber="357">Dhaulagiri-Himal</location>
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| Kali Gand. vall.,
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<collectingMunicipality id="6D0BACE1FB2B927AFB4AF87CE0BBD787" box="[1278,1422,1987,2013]" pageId="4" pageNumber="357">Yak Kharkra</collectingMunicipality>
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</materialsCitation>
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</paragraph>
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<paragraph id="8D6F369BFB2A927BFF23FF29E784D08E" blockId="5.[151,1437,150,681]" pageId="5" pageNumber="358">
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(upp. Marpha),
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<quantity id="4A289B7EFB2A927BFEF3FF29E4F7D0EA" box="[327,450,150,176]" metricMagnitude="3" metricUnit="m" metricValue="2.3205" metricValueMax="4.6" metricValueMin="0.040999999999999995" pageId="5" pageNumber="358" unit="m" value="2320.5" valueMax="4600.0" valueMin="41.0">41–4600m</quantity>
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, | 12./
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<date id="F96E105BFB2A927BFDB0FF29E7A5D0EA" box="[516,656,150,176]" pageId="5" pageNumber="358" value="1998-07-13">13.VII.1998</date>
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, leg. Jäger’ <printed>, ‘
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<typeStatus id="526B8839FB2A927BFC2CFF29E104D0EA" box="[920,1073,150,176]" pageId="5" pageNumber="358" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4AD04D18FB2A927BFBF1FF29E1DBD0EA" authorityName="Luze" authorityYear="1903" box="[1093,1262,150,176]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="genus">
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<emphasis id="BFA4EA89FB2A927BFBF1FF29E1DBD0EA" box="[1093,1262,150,176]" italics="true" pageId="5" pageNumber="358">Trichodromeus</emphasis>
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</taxonomicName>
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|
||
<emphasis id="BFA4EA89FB2A927BFAB5FF28E065D0EA" box="[1281,1360,151,176]" italics="true" pageId="5" pageNumber="358">jaegeri</emphasis>
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</taxonomicName>
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<emphasis id="BFA4EA89FB2A927BFAE3FF29E0BAD0EA" bold="true" box="[1367,1423,150,176]" pageId="5" pageNumber="358">
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<taxonomicNameLabel id="A49757F2FB2A927BFAE3FF29E0BAD0EA" box="[1367,1423,150,176]" pageId="5" pageNumber="358" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.
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<collectionCode id="EBC1AE5EFB2A927BFEB9FF05E416D08E" box="[269,291,186,212]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="358" type="Museum">V</collectionCode>
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. des. 2021’ <red, printed> (
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).
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<paragraph id="8D6F369BFB2A927BFF73FF60E77ED13E" blockId="5.[151,1437,150,681]" pageId="5" pageNumber="358">
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<typeStatus id="526B8839FB2A927BFF73FF60E418D0A3" box="[199,301,223,249]" pageId="5" pageNumber="358" type="paratype">Paratype</typeStatus>
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:
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,
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: same data as the holotype (
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<specimenCount id="9BD6FD12FB2A927BFD50FF61E624D0A3" box="[740,785,222,249]" pageId="5" pageNumber="358" type="male">1 ♂</specimenCount>
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:
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<collectionCode id="EBC1AE5EFB2A927BFCA9FF61E677D0A2" box="[797,834,222,248]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="5" pageNumber="358">CS</collectionCode>
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,
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<specimenCount id="9BD6FD12FB2A927BFCFBFF61E649D0A3" box="[847,892,222,249]" pageId="5" pageNumber="358" type="female">1 ♀</specimenCount>
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:
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<collectionCode id="EBC1AE5EFB2A927BFC3DFF61E6E0D0A2" box="[905,981,222,248]" pageId="5" pageNumber="358">SNSD</collectionCode>
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);
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<specimenCount id="9BD6FD12FB2A927BFC5EFF61E119D0A3" box="[1002,1068,222,249]" pageId="5" pageNumber="358" type="female">3 ♀♀</specimenCount>
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: ‘
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<collectingCountry id="F5C7760BFB2A927BFB8AFF61E1AFD0A2" box="[1086,1178,222,248]" name="Nepal" pageId="5" pageNumber="358">NEPAL</collectingCountry>
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<collectingRegion id="4F14F879FB2A927BFB16FF61E02FD0A2" box="[1186,1306,222,248]" country="Nepal" name="Dhawalagiri" pageId="5" pageNumber="358">Dhaulagiri</collectingRegion>
|
||
| upp. Yak- kharka |
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<quantity id="4A289B7EFB2A927BFF43FEBDE45FD147" box="[247,362,258,285]" metricMagnitude="3" metricUnit="m" metricValue="2.2705" metricValueMax="4.5" metricValueMin="0.040999999999999995" pageId="5" pageNumber="358" unit="m" value="2270.5" valueMax="4500.0" valueMin="41.0">41-4500m</quantity>
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12.7.[19]98 | lg.Berndt & Schmidt’ <printed>, ‘Ankauf A. Dobbertin | Rostock, 2001 | Museum Dresden’ <printed> (
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<specimenCount id="9BD6FD12FB2A927BFE31FE99E485D11B" box="[389,432,294,321]" pageId="5" pageNumber="358" type="female">1 ♀</specimenCount>
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:
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<collectionCode id="EBC1AE5EFB2A927BFE0AFE99E4D6D11A" box="[446,483,294,320]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="5" pageNumber="358">CS</collectionCode>
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;
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<specimenCount id="9BD6FD12FB2A927BFE5BFE99E704D11B" box="[495,561,294,321]" pageId="5" pageNumber="358" type="female">2 ♀♀</specimenCount>
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:
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<collectionCode id="EBC1AE5EFB2A927BFD8AFE99E7BFD11A" box="[574,650,294,320]" pageId="5" pageNumber="358">SNSD</collectionCode>
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). All
|
||
<typeStatus id="526B8839FB2A927BFD7CFE98E606D11B" box="[712,819,295,321]" pageId="5" pageNumber="358" type="paratype">paratypes</typeStatus>
|
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with red printed label: ‘
|
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<typeStatus id="526B8839FB2A927BFBF5FE99E1E4D11A" box="[1089,1233,294,320]" pageId="5" pageNumber="358" type="paratype">PARATYPE</typeStatus>
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|
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<taxonomicName id="4AD04D18FB2A927BFB52FE99E0BAD11A" authorityName="Luze" authorityYear="1903" box="[1254,1423,294,320]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BFA4EA89FB2A927BFB52FE99E0BAD11A" box="[1254,1423,294,320]" italics="true" pageId="5" pageNumber="358">Trichodromeus</emphasis>
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||
</taxonomicName>
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||
|
|
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<taxonomicName id="4AD04D18FB2A927BFF23FEF4E5D3D13E" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[151,230,331,356]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
|
||
<emphasis id="BFA4EA89FB2A927BFF23FEF4E5D3D13E" box="[151,230,331,356]" italics="true" pageId="5" pageNumber="358">jaegeri</emphasis>
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</taxonomicName>
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<emphasis id="BFA4EA89FB2A927BFF59FEF5E410D13E" bold="true" box="[237,293,330,356]" pageId="5" pageNumber="358">
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<taxonomicNameLabel id="A49757F2FB2A927BFF59FEF5E410D13E" box="[237,293,330,356]" pageId="5" pageNumber="358" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
|
||
| Shavrin A.
|
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<collectionCode id="EBC1AE5EFB2A927BFE1BFEF5E4F0D13E" box="[431,453,330,356]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="358" type="Museum">V</collectionCode>
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. des. 2021’.
|
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</paragraph>
|
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</subSubSection>
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|
||
<paragraph id="8D6F369BFB2A927BFF73FED1E4BED1AF" blockId="5.[151,1437,150,681]" pageId="5" pageNumber="358">
|
||
<emphasis id="BFA4EA89FB2A927BFF73FED1E46CD1D2" bold="true" box="[199,345,366,392]" pageId="5" pageNumber="358">Description.</emphasis>
|
||
Measurements (min-max; n=6): HW: 1.10–1.22; HL: 0.77–0.82; OL: 0.25–0.27; LT: 0.24–0.25; AL (
|
||
<typeStatus id="526B8839FB2A927BFF7AFE2DE402D1F7" box="[206,311,402,429]" pageId="5" pageNumber="358" type="holotype">holotype</typeStatus>
|
||
): 3.40; PL: 1.00–1.10; PWmax: 1.12–1.37; PWmin: 0.98–1.18; ESL: 1.67–2.21; EW: 1.75–2.28; MTbL (
|
||
<typeStatus id="526B8839FB2A927BFF5AFE09E462D18B" box="[238,343,438,465]" pageId="5" pageNumber="358" type="holotype">holotype</typeStatus>
|
||
): 1.46; MTrL (
|
||
<typeStatus id="526B8839FB2A927BFE48FE09E750D18B" box="[508,613,438,465]" pageId="5" pageNumber="358" type="holotype">holotype</typeStatus>
|
||
): 0.55 (MTrL 1–4: 0.28; MTrL 5: 0.27); AW: 1.75–2.01; AedL: 1.05; BL: 5.35 (
|
||
<typeStatus id="526B8839FB2A927BFF63FE65E408D1AF" box="[215,317,474,501]" pageId="5" pageNumber="358" type="holotype">holotype</typeStatus>
|
||
)–7.60.
|
||
</paragraph>
|
||
<paragraph id="8D6F369BFB2A927BFF73FE41E7ADD2F2" blockId="5.[151,1437,150,681]" pageId="5" pageNumber="358">
|
||
Body and antennomeres brown to black; mouthparts and legs brown; tarsi and sometimes apical segment of maxillary palpi and tibia (
|
||
<specimenCount id="9BD6FD12FB2A927BFE0FFD9DE77BD267" box="[443,590,546,573]" pageId="5" pageNumber="358" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
) yellow-brown. Head with distinct microreticulation: fine and isodiametric in middle, transverse on infraorbital ridges; neck with dense and coarse isodiametric microsculpture; pronotum with very dense and fine, transverse microreticulation, usually denser and coarser in mediobasal portion; scutellum with dense transverse meshes. Habitus as in
|
||
<figureCitation id="15EB2A1EFB2A927BFDE4FD31E7A1D2F2" box="[592,660,654,680]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[168,1419,192,1909]" captionTargetId="figure-28@2.[152,1436,179,1920]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Trichodromeus: 1—T. acutipenis (holotype), 2—T. jaegeri (paratype), 3—T. kaliyuga (holotype), 4—T. trilobatus (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5030540" httpUri="https://zenodo.org/record/5030540/files/figure.png" pageId="5" pageNumber="358">Fig. 2</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="D9AF6613FB2A927BFF23F930E078D6F2" ID-DOI="http://doi.org/10.5281/zenodo.5030544" ID-Zenodo-Dep="5030544" box="[151,1357,1679,1704]" httpUri="https://zenodo.org/record/5030544/files/figure.png" pageId="5" pageNumber="358" startId="5.[151,250,1679,1703]" targetBox="[151,1436,704,1656]" targetPageId="5">
|
||
<paragraph id="8D6F369BFB2A927BFF23F930E078D6F2" blockId="5.[151,1357,1679,1704]" box="[151,1357,1679,1704]" pageId="5" pageNumber="358">
|
||
<emphasis id="BFA4EA89FB2A927BFF23F930E42BD6FD" bold="true" box="[151,286,1679,1703]" pageId="5" pageNumber="358">FIGURE 11.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4AD04D18FB2A927BFE75F92FE76CD6F2" authorityName="Luze" authorityYear="1903" box="[449,601,1680,1704]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BFA4EA89FB2A927BFE75F92FE76CD6F2" box="[449,601,1680,1704]" italics="true" pageId="5" pageNumber="358">Trichodromeus</emphasis>
|
||
</taxonomicName>
|
||
in Nepal:
|
||
<taxonomicName id="4AD04D18FB2A927BFD70F92FE671D6FD" authorityName="Shavrin" authorityYear="2021" box="[708,836,1680,1703]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="species" species="acutipenis">
|
||
<emphasis id="BFA4EA89FB2A927BFD70F92FE671D6FD" box="[708,836,1680,1703]" italics="true" pageId="5" pageNumber="358">T. acutipenis</emphasis>
|
||
</taxonomicName>
|
||
(circles),
|
||
<taxonomicName id="4AD04D18FB2A927BFC1FF92FE13ED6FD" authorityName="Shavrin" authorityYear="2021" box="[939,1035,1680,1703]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="species" species="jaegeri">
|
||
<emphasis id="BFA4EA89FB2A927BFC1FF92FE13ED6FD" box="[939,1035,1680,1703]" italics="true" pageId="5" pageNumber="358">T. jaegeri</emphasis>
|
||
</taxonomicName>
|
||
(square),
|
||
<taxonomicName id="4AD04D18FB2A927BFBC5F92FE1D4D6F2" authorityName="Shavrin" authorityYear="2021" box="[1137,1249,1680,1704]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="species" species="kaliyuga">
|
||
<emphasis id="BFA4EA89FB2A927BFBC5F92FE1D4D6F2" box="[1137,1249,1680,1704]" italics="true" pageId="5" pageNumber="358">T. kaliyuga</emphasis>
|
||
</taxonomicName>
|
||
(triangle).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8D6F369BFB2A9278FF73F969E63FD08F" blockId="5.[151,1437,1750,2028]" lastBlockId="6.[151,1437,151,898]" lastPageId="6" lastPageNumber="359" pageId="5" pageNumber="358">Head 1.4 times as broad as long, with distinctly elevated middle part and portions between ocelli and eyes; anterio-median depression deep, gradually narrowed and extended basad and narrowly connected with interocellar depression; temples convex, about as long as longitudinal length of eyes; interocellar depression deep, somewhat subrectangular, separated from infraorbital ridges by deep, distinctly convergent latero-anteriad anteocellar foveae, reaching level of middle or apical third of eyes. Ocelli small, located slightly below level of posterior margin of eyes, distance between ocelli about as long as distance between ocellus and posterior margin of eye or slightly shorter. Punctation moderately large and deep, irregular, finer in middle and denser in infraorbital ridges. Antennae exceeding middle of apical third of elytra when reclined; basal antennomere wide, more than twice as long as broad, antennomere 2 significantly narrower and shorter than basal antennomere, 3 distinctly longer and narrower than 2, 4–8 slightly broader than 3, 9–10 slightly broder than 8.</paragraph>
|
||
<paragraph id="8D6F369BFB299278FF73FF60E418D1D3" blockId="6.[151,1437,151,898]" pageId="6" pageNumber="359">Pronotum convex, 1.1–1.2 times broader than long and slightly broader than head, with widely rounded anterior angles not protruded apicad; widest anterior portion gradually narrowed toward obtuse posterior angles; anterior margin rounded; posterior margin straight; middle portion with moderately wide logitudinal depression, connected with indistinct transverse mediobasal depression in some specimens. Punctation regular, about as that on head, but slightly finer.</paragraph>
|
||
<paragraph id="8D6F369BFB299278FF73FE2CE4E3D18B" blockId="6.[151,1437,151,898]" pageId="6" pageNumber="359">Elytra convex, about as broad as long, slightly widened posteriad. Punctation about as that on pronotum, but somewhat larger and denser.</paragraph>
|
||
<paragraph id="8D6F369BFB299278FF73FE64E604D1AF" blockId="6.[151,1437,151,898]" box="[199,817,475,501]" pageId="6" pageNumber="359">Abdomen about as broad as elytra or slightly narrower.</paragraph>
|
||
<paragraph id="8D6F369BFB299278FF73FE40E03BD2DF" blockId="6.[151,1437,151,898]" pageId="6" pageNumber="359">
|
||
Male. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII slightly concaved. Median lobe wide, from widest apical third gradually narrowed toward moderately narrow subtriangular apical portion with small, rounded apex; parameres slightly exceeding apex of median lobe, with two moderately long apical and two short preapical setae (
|
||
<figureCitation id="15EB2A1EFB299278FD47FDD4E60DD2DF" box="[755,824,619,645]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[173,1420,193,1912]" captionTargetId="figure-16@3.[155,1432,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Trichodromeus (5–6: T. acutipenis (holotype), 7–8: T. jaegeri (holotype), 9–10: T. kaliyuga (holotype): 5, 7, 9–parameral view, 6, 8, 10 lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5030542" httpUri="https://zenodo.org/record/5030542/files/figure.png" pageId="6" pageNumber="359">Fig. 9</figureCitation>
|
||
). Lateral aspect of aedeagus as in
|
||
<figureCitation id="15EB2A1EFB299278FB03FDD4E03CD2DF" box="[1207,1289,619,645]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[173,1420,193,1912]" captionTargetId="figure-16@3.[155,1432,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Trichodromeus (5–6: T. acutipenis (holotype), 7–8: T. jaegeri (holotype), 9–10: T. kaliyuga (holotype): 5, 7, 9–parameral view, 6, 8, 10 lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5030542" httpUri="https://zenodo.org/record/5030542/files/figure.png" pageId="6" pageNumber="359">Fig. 10</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8D6F369BFB299278FF73FD30E129D2F3" blockId="6.[151,1437,151,898]" box="[199,1052,655,681]" pageId="6" pageNumber="359">Female. Apical margins of abdominal tergite VIII and sternite VIII rounded.</paragraph>
|
||
<paragraph id="8D6F369BFB299278FF73FD0DE7F9D363" blockId="6.[151,1437,151,898]" pageId="6" pageNumber="359">
|
||
<emphasis id="BFA4EA89FB299278FF73FD0DE487D297" bold="true" box="[199,434,690,717]" pageId="6" pageNumber="359">Comparative notes.</emphasis>
|
||
Based on the general shape of the body,
|
||
<taxonomicName id="4AD04D18FB299278FC3DFD0CE6C2D296" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[905,1015,691,717]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
|
||
<emphasis id="BFA4EA89FB299278FC3DFD0CE6ACD296" box="[905,921,691,716]" italics="true" pageId="6" pageNumber="359">T</emphasis>
|
||
.
|
||
<emphasis id="BFA4EA89FB299278FC1CFD0CE6C2D296" box="[936,1015,691,716]" italics="true" pageId="6" pageNumber="359">jaegeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFA4EA89FB299278FBB4FD0CE10DD297" bold="true" box="[1024,1080,691,717]" pageId="6" pageNumber="359">
|
||
<taxonomicNameLabel id="A49757F2FB299278FBB4FD0CE10DD297" box="[1024,1080,691,717]" pageId="6" pageNumber="359" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4AD04D18FB299278FB64FD0CE069D297" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[1232,1372,691,717]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="trilobatus" status="sp. nov.">
|
||
<emphasis id="BFA4EA89FB299278FB64FD0CE1D5D296" box="[1232,1248,691,716]" italics="true" pageId="6" pageNumber="359">T</emphasis>
|
||
.
|
||
<emphasis id="BFA4EA89FB299278FB5BFD0CE069D297" box="[1263,1372,691,717]" italics="true" pageId="6" pageNumber="359">trilobatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFA4EA89FB299278FAD0FD0CE0A9D297" bold="true" box="[1380,1436,691,717]" pageId="6" pageNumber="359">
|
||
<taxonomicNameLabel id="A49757F2FB299278FAD0FD0CE0A9D297" box="[1380,1436,691,717]" pageId="6" pageNumber="359" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4AD04D18FB299278FF73FD68E471D2AB" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[199,324,727,753]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="kaliyuga" status="sp. nov.">
|
||
<emphasis id="BFA4EA89FB299278FF73FD68E5E2D2AA" box="[199,215,727,752]" italics="true" pageId="6" pageNumber="359">T</emphasis>
|
||
.
|
||
<emphasis id="BFA4EA89FB299278FF50FD68E471D2AB" box="[228,324,727,753]" italics="true" pageId="6" pageNumber="359">kaliyuga</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFA4EA89FB299278FEF8FD68E4B1D2AB" bold="true" box="[332,388,727,753]" pageId="6" pageNumber="359">
|
||
<taxonomicNameLabel id="A49757F2FB299278FEF8FD68E4B1D2AB" box="[332,388,727,753]" pageId="6" pageNumber="359" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from which it can be distinguished by the different shape of the apical portion of the median lobe. Additionally, it differs from
|
||
<taxonomicName id="4AD04D18FB299278FDBBFD44E7ACD34F" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[527,665,763,789]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="trilobatus" status="sp. nov.">
|
||
<emphasis id="BFA4EA89FB299278FDBBFD44E72AD34E" box="[527,543,763,788]" italics="true" pageId="6" pageNumber="359">T</emphasis>
|
||
.
|
||
<emphasis id="BFA4EA89FB299278FD98FD44E7ACD34F" box="[556,665,763,789]" italics="true" pageId="6" pageNumber="359">trilobatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFA4EA89FB299278FD14FD44E7EDD34F" bold="true" box="[672,728,763,789]" pageId="6" pageNumber="359">
|
||
<taxonomicNameLabel id="A49757F2FB299278FD14FD44E7EDD34F" box="[672,728,763,789]" pageId="6" pageNumber="359" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the paler coloration and from
|
||
<taxonomicName id="4AD04D18FB299278FBF9FD44E1FFD34F" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[1101,1226,763,789]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="kaliyuga" status="sp. nov.">
|
||
<emphasis id="BFA4EA89FB299278FBF9FD44E168D34E" box="[1101,1117,763,788]" italics="true" pageId="6" pageNumber="359">T</emphasis>
|
||
.
|
||
<emphasis id="BFA4EA89FB299278FBDEFD44E1FFD34F" box="[1130,1226,763,789]" italics="true" pageId="6" pageNumber="359">kaliyuga</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A49757F2FB299278FB65FD44E03CD34F" box="[1233,1289,763,789]" pageId="6" pageNumber="359" rank="species">
|
||
<emphasis id="BFA4EA89FB299278FB65FD44E037D34F" bold="true" box="[1233,1282,763,789]" pageId="6" pageNumber="359">sp.n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
by the larger and somewhat wider elytra and shorter parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C5CA6510FB299278FF73FCFDE1EED307" box="[199,1243,834,861]" pageId="6" pageNumber="359" type="distribution">
|
||
<paragraph id="8D6F369BFB299278FF73FCFDE1EED307" blockId="6.[151,1437,151,898]" box="[199,1243,834,861]" pageId="6" pageNumber="359">
|
||
<emphasis id="BFA4EA89FB299278FF73FCFDE454D306" bold="true" box="[199,353,834,860]" pageId="6" pageNumber="359">Distribution.</emphasis>
|
||
The species is known only from the
|
||
<typeStatus id="526B8839FB299278FD4AFCFCE61BD307" box="[766,814,835,861]" pageId="6" pageNumber="359">type</typeStatus>
|
||
locality in the central
|
||
<collectingCountry id="F5C7760BFB299278FB9DFCFCE159D307" box="[1065,1132,835,861]" name="Nepal" pageId="6" pageNumber="359">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="15EB2A1EFB299278FBCFFCFCE1F8D307" box="[1147,1229,835,861]" captionStart="FIGURE 11" captionStartId="5.[151,250,1679,1703]" captionTargetBox="[151,1436,704,1656]" captionTargetId="figure-216@5.[151,1436,704,1656]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of Trichodromeus in Nepal: T. acutipenis (circles), T. jaegeri (square), T. kaliyuga (triangle)." figureDoi="http://doi.org/10.5281/zenodo.5030544" httpUri="https://zenodo.org/record/5030544/files/figure.png" pageId="6" pageNumber="359">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C5CA6510FB299278FF73FCD8E07BD3DB" box="[199,1358,871,898]" pageId="6" pageNumber="359" type="etymology">
|
||
<paragraph id="8D6F369BFB299278FF73FCD8E07BD3DB" blockId="6.[151,1437,151,898]" box="[199,1358,871,898]" pageId="6" pageNumber="359">
|
||
<emphasis id="BFA4EA89FB299278FF73FCD8E47BD3DB" bold="true" box="[199,334,871,897]" pageId="6" pageNumber="359">Etymology.</emphasis>
|
||
The species is dedicated to Olaf Jäger (Dresden), one of the collectors of the
|
||
<typeStatus id="526B8839FB299278FB07FCD8E1D6D3DB" box="[1203,1251,871,897]" pageId="6" pageNumber="359">type</typeStatus>
|
||
material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |