638 lines
70 KiB
XML
638 lines
70 KiB
XML
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<mods:title id="A162E0DC60BF104C8EAD90C18DF43379">Seven new Alpine species of the genus Alpiscorpius Gantenbein et al., 1999, stat. n. (Scorpiones: Euscorpiidae)</mods:title>
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<mods:namePart id="7E0F5E3714DEEDA6D5E129E01324C699">Kovařík, František</mods:namePart>
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<mods:namePart id="C5F67FB2BA5FE29849CF644A13CD9D8E">Štundlová, Jana</mods:namePart>
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<mods:namePart id="C3582B4AF13F548EE1CFB48266DE4C02">Fet, Victor</mods:namePart>
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<mods:namePart id="E90F29D6A7F3C8D0DFCB8B8C94987EC0">Šťáhlavský, František</mods:namePart>
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<mods:title id="0119C06442B64E6324DEEBE8E0FEDA3A">Euscorpius</mods:title>
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<mods:date id="C3C07438F09805CCD560729F823F573C">2019</mods:date>
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<mods:number id="C413FE90D3E024FC7EF7038A774CE616">287</mods:number>
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<heading id="D630FF0B4745861EFF2DF93291D64096" centered="true" fontSize="11" level="2" pageId="8" pageNumber="7" reason="2">
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<taxonomicName id="4AC733E44745861EFF2DF932925F40F1" ID-CoL="C73M" authority="(Di Caporiacco, 1950)" authorityName="Kovařík & Štundlová & Fet & Šťáhlavský" authorityYear="2019" baseAuthorityName="Di Caporiacco" baseAuthorityYear="1950" box="[138,621,1686,1714]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="beta" status="comb. nov.">
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<emphasis id="BFB394754745861EFF2DF932916E40F2" bold="true" box="[138,348,1686,1713]" italics="true" pageId="8" pageNumber="7">Alpiscorpius beta</emphasis>
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(
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<bibRefCitation id="E95635964745861EFECCF933925640F1" author="DI CAPORIACCO" box="[363,612,1687,1714]" pageId="8" pageNumber="7" pagination="159 - 230" refId="ref26659" refString="DI CAPORIACCO, L. 1950. Le specie e sottospecie del genere " Euscorpius " viventi in Italia ed in alcune zone confinanti. Memorie / Accademia nazionale dei Lincei (ser. 8), 2: 159 - 230." type="journal article" year="1950">Di Caporiacco, 1950</bibRefCitation>
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)
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</taxonomicName>
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,
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<emphasis id="BFB394754745861EFDDFF933928E40F2" bold="true" box="[632,700,1687,1713]" pageId="8" pageNumber="7">comb</emphasis>
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.
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<emphasis id="BFB394754745861EFD6BF93392EE40F2" bold="true" box="[716,732,1687,1713]" pageId="8" pageNumber="7">n</emphasis>
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.
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</taxonomicNameLabel>
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,
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<emphasis id="BFB394754745861EFE37F91E918C4097" bold="true" box="[400,446,1722,1748]" pageId="8" pageNumber="7">stat</emphasis>
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.
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<emphasis id="BFB394754745861EFE6AF91E91EF4097" bold="true" box="[461,477,1722,1748]" pageId="8" pageNumber="7">n</emphasis>
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.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C5DD1BEC4745861EFEA4F97892B04177" pageId="8" pageNumber="7" type="description">
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<paragraph id="8D7848674745861EFEA4F97893314157" blockId="8.[113,771,1686,1844]" pageId="8" pageNumber="7">
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(
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<figureCitation id="15FC54E24745861EFEADF978919A40B7" box="[266,424,1756,1780]" captionStart="Figures 13–22" captionStartId="9.[117,194,1967,1989]" captionTargetBox="[127,1463,238,1949]" captionTargetId="figure-90@9.[120,1469,225,1961]" captionTargetPageId="9" captionText="Figures 13–22: Alpiscorpius beta comb. n., stat. n., male (13–17, No. S102) from Switzerland, Gondo, 46.195°N 8.137°E and Alpiscorpius delta sp. n., male holotype (18–22, No. S277). Figures 13–15, 18–20. Pedipalp chela and patella dorsal (13, 18), eXternal (14, 19), and vental (15, 20) views. Figures 16–17, 21–22. Metasoma III–V and telson, ventral (16, 21), and lateral (17, 22) views." pageId="8" pageNumber="7">Figures 13–17</figureCitation>
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,
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<figureCitation id="15FC54E24745861EFE14F97891E240B7" box="[435,464,1756,1780]" captionStart="Figure 71" captionStartId="28.[115,183,1204,1226]" captionTargetBox="[113,1474,234,1186]" captionTargetId="figure-33@28.[113,1474,231,1186]" captionTargetPageId="28" captionText="Figure 71. Map showing confirmed distribution of studied species of Alpiscorpius, stat. n." pageId="8" pageNumber="7">71</figureCitation>
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,
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<tableCitation id="C0457DDC4745861EFE7DF978920140B7" box="[474,563,1756,1780]" captionStart="Table 1" captionStartId="5.[111,167,1484,1505]" captionText="Table 1. The currently valid Alpiscorpius taXa, their karyotypes, and distribution." pageId="8" pageNumber="7">Tables 1</tableCitation>
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,
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||
<tableCitation id="C0457DDC4745861EFDE6F978926240B7" box="[577,592,1756,1780]" captionStart="Table 2" captionStartId="8.[116,172,1002,1023]" captionText="Table 2. Comparative measurements of Alpiscorpius beta comb. n., male (No. S102) from Switzerland, Gondo, 46.195°N 8.137°E, A. delta sp. n., A. kappa sp. n., and A. lambda sp. n., male holotypes. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="8" pageNumber="7">2</tableCitation>
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,
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<tableCitation id="C0457DDC4745861EFDFCF978925B40B7" box="[603,617,1756,1780]" captionStart="Table 5" captionStartId="26.[113,169,1192,1213]" captionText="Table 5. Diagnostic table of studied Alpiscorpius spp. (based on eXamined material; see also Notes under species)." pageId="8" pageNumber="7">5</tableCitation>
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)
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<emphasis id="BFB394754745861EFFD6F95893314157" bold="true" box="[113,771,1788,1812]" pageId="8" pageNumber="7">http://zoobank.org/urn:lsid:zoobank.org:act:02256D81-</emphasis>
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<paragraph id="8D7848674745861EFF54F8B892B04177" blockId="8.[113,771,1686,1844]" box="[243,642,1820,1844]" pageId="8" pageNumber="7">
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<emphasis id="BFB394754745861EFF54F8B892B04177" bold="true" box="[243,642,1820,1844]" pageId="8" pageNumber="7">6FBF-46C0-BB92-5B23BD676CA5</emphasis>
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</paragraph>
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<subSubSection id="C5DD1BEC4745861EFFD6F8FD94B54312" pageId="8" pageNumber="7" type="reference_group">
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<paragraph id="8D7848674745861EFFD6F8FD92AC4132" blockId="8.[113,772,1881,2033]" box="[113,670,1881,1905]" pageId="8" pageNumber="7">
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<taxonomicName id="4AC733E44745861EFFD6F8FD916B4132" authorityName="Kovařík & Štundlová & Fet & Šťáhlavský" authorityYear="2019" baseAuthorityName="Kovařík & Štundlová & Fet & Šťáhlavský" baseAuthorityYear="1837" box="[113,345,1881,1905]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="germanus">
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||
<emphasis id="BFB394754745861EFFD6F8FD916B4132" box="[113,345,1881,1905]" italics="true" pageId="8" pageNumber="7">Euscorpius germanus</emphasis>
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</taxonomicName>
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“forma B”:
|
||
<treatmentCitation id="0C666E764745861EFE79F8FD92AB4132" author="CAPRA" box="[478,665,1881,1905]" page="204" pageId="8" pageNumber="7" year="1939">
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<bibRefCitation id="E95635964745861EFE79F8FD92AB4132" author="CAPRA" box="[478,665,1881,1905]" pageId="8" pageNumber="7" pagination="199 - 213" refId="ref26586" refString="CAPRA, F. 1939. L' Euscorpius germanus (C. L. Koch) in Italia (Arachnida, Scorpiones). Memorie della Societa Entomologica Italiana, 18 (2): 199 - 213." type="journal article" year="1939">Capra, 1939: 204</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8D7848674745861EFFD6F8DD929C41D2" blockId="8.[113,772,1881,2033]" box="[113,686,1913,1937]" pageId="8" pageNumber="7">
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<treatmentCitationGroup id="ADD76F494745861EFFD6F8DD929C41D2" box="[113,686,1913,1937]" pageId="8" pageNumber="7">
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<treatmentCitation id="0C666E764745861EFFD6F8DD929841D2" author="DI CAPORIACCO" box="[113,682,1913,1937]" page="211" pageId="8" pageNumber="7" year="1950">
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||
<taxonomicName id="4AC733E44745861EFFD6F8DD929841D2" authority="Di Caporiacco, 1950: 211" authorityName="Di Caporiacco" authorityPageNumber="211" authorityYear="1950" box="[113,682,1913,1937]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="germanus" subSpecies="beta">
|
||
<emphasis id="BFB394754745861EFFD6F8DD91BC41D2" box="[113,398,1913,1937]" italics="true" pageId="8" pageNumber="7">Euscorpius germanus beta</emphasis>
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<bibRefCitation id="E95635964745861EFE33F8DD929841D2" author="DI CAPORIACCO" box="[404,682,1913,1937]" pageId="8" pageNumber="7" pagination="159 - 230" refId="ref26659" refString="DI CAPORIACCO, L. 1950. Le specie e sottospecie del genere " Euscorpius " viventi in Italia ed in alcune zone confinanti. Memorie / Accademia nazionale dei Lincei (ser. 8), 2: 159 - 230." type="journal article" year="1950">Di Caporiacco, 1950: 211</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8D7848674745861EFFD6F83D933641F2" blockId="8.[113,772,1881,2033]" box="[113,772,1945,1969]" pageId="8" pageNumber="7">
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<treatmentCitationGroup id="ADD76F494745861EFFD6F83D933641F2" box="[113,772,1945,1969]" pageId="8" pageNumber="7">
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<treatmentCitation id="0C666E764745861EFFD6F83D929041F2" author="BONACINA" box="[113,674,1945,1969]" page="58" pageId="8" pageNumber="7" year="1980">
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<taxonomicName id="4AC733E44745861EFFD6F83D929041F2" authority="germanus: Bonacina, 1980: 58" authorityName="germanus: Bonacina" authorityPageNumber="58" authorityYear="1980" box="[113,674,1945,1969]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="germanus" subSpecies="germanus">
|
||
<emphasis id="BFB394754745861EFFD6F83D91F541F2" box="[113,455,1945,1969]" italics="true" pageId="8" pageNumber="7">Euscorpius germanus germanus</emphasis>
|
||
:
|
||
<bibRefCitation id="E95635964745861EFE73F83D929041F2" author="BONACINA" box="[468,674,1945,1969]" pageId="8" pageNumber="7" pagination="47 - 100" refId="ref26484" refString="BONACINA, A. 1980. Sistematica specifica e sottospecifica del complesso " Euscorpius germanus " (Scorpiones, Chactidae). Rivista del Museo Civico di Scienze Naturali " Enrico Caffi " (Bergamo), 2: 47 - 100." type="journal article" year="1980">Bonacina, 1980: 58</bibRefCitation>
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</taxonomicName>
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(in part).
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</paragraph>
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<paragraph id="8D7848674745861EFFD6F81D919541B2" blockId="8.[113,772,1881,2033]" pageId="8" pageNumber="7">
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<taxonomicName id="4AC733E44745861EFFD6F81D917041B2" authority=": Fet & Sissom, 2000: 366 - 368" authorityName=": Fet & Sissom" authorityPageNumber="366 - 368" authorityYear="2000" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="germanus" subGenus="Euscorpius" subSpecies="alpha">
|
||
<emphasis id="BFB394754745861EFFD6F81D90DB4193" box="[113,233,1977,2000]" italics="true" pageId="8" pageNumber="7">Euscorpius</emphasis>
|
||
(
|
||
<emphasis id="BFB394754745861EFEA6F81D914B4193" box="[257,377,1977,2000]" italics="true" pageId="8" pageNumber="7">Euscorpius</emphasis>
|
||
)
|
||
<emphasis id="BFB394754745861EFE37F81E92774192" box="[400,581,1977,2001]" italics="true" pageId="8" pageNumber="7">germanus alpha</emphasis>
|
||
: Fet & Sissom, 2000: 366–368
|
||
</taxonomicName>
|
||
(in part).
|
||
</paragraph>
|
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<paragraph id="8D7848674745861EFC97FBFD950B42F2" blockId="8.[816,1474,1113,1363]" pageId="8" pageNumber="7">
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<treatmentCitationGroup id="ADD76F494745861EFC97FBFD950B42F2" pageId="8" pageNumber="7">
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<treatmentCitation id="0C666E764745861EFC97FBFD95F04232" author="GANTENBEIN" box="[816,1474,1113,1137]" page="858" pageId="8" pageNumber="7" year="2000">
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<taxonomicName id="4AC733E44745861EFC97FBFD95F04232" authority=": Gantenbein et al., 2000: 858" authorityName="Gantenbein" authorityPageNumber="858" authorityYear="2000" box="[816,1474,1113,1137]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="alpha" subGenus="Alpiscorpius">
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<emphasis id="BFB394754745861EFC97FBFD939A4233" box="[816,936,1113,1136]" italics="true" pageId="8" pageNumber="7">Euscorpius</emphasis>
|
||
(
|
||
<emphasis id="BFB394754745861EFC10FBFD940C4232" box="[951,1086,1113,1137]" italics="true" pageId="8" pageNumber="7">Alpiscorpius</emphasis>
|
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)
|
||
<emphasis id="BFB394754745861EFBEAFBFD94B84232" box="[1101,1162,1113,1137]" italics="true" pageId="8" pageNumber="7">alpha</emphasis>
|
||
:
|
||
<bibRefCitation id="E95635964745861EFB30FBFD95F04232" author="GANTENBEIN" box="[1175,1474,1113,1137]" pageId="8" pageNumber="7" pagination="843 - 869" refId="ref27133" refString="GANTENBEIN, B., V. FET, M. BARKER & A. SCHOLL. 2000. Nuclear and mitochondrial markers reveal the eXistence of two parapatric scorpion species in the Alps: Euscorpius germanus (C. L. Koch, 1837) and E. alpha Caporiacco, 1950, stat. nov. (Scorpiones, Euscorpiidae). Revue Suisse de Zoologie, 107 (4): 843 - 869." type="journal article" year="2000">Gantenbein et al., 2000: 858</bibRefCitation>
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</taxonomicName>
|
||
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|
||
(in part; Gondo, Fountainemore); Fet & Braunwalder, 2005: 34 (in part; Gondo, Zwischbergental).
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</paragraph>
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<treatmentCitation id="0C666E764745861EFC97FB1D95354292" author="BRAUNWALDER" box="[816,1287,1209,1233]" page="281" pageId="8" pageNumber="7" year="2001">
|
||
<taxonomicName id="4AC733E44745861EFC97FB1D95354292" authority=": Braunwalder, 2001: 281" authorityName="Braunwalder" authorityPageNumber="281" authorityYear="2001" box="[816,1287,1209,1233]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="alpha">
|
||
<emphasis id="BFB394754745861EFC97FB1D93DD4292" box="[816,1007,1209,1233]" italics="true" pageId="8" pageNumber="7">Euscorpius alpha</emphasis>
|
||
:
|
||
<bibRefCitation id="E95635964745861EFBA7FB1D95354292" author="BRAUNWALDER" box="[1024,1287,1209,1233]" pageId="8" pageNumber="7" pagination="279 - 286" refId="ref26527" refString="BRAUNWALDER, M. E. 2001. Scorpions of Switzerland: summary of a faunistic survey. Pp. 279 - 286 in: Fet, V. & P. A. Selden (eds.). Scorpions 2001. In Memoriam Gary A. Polis. Burnham Beeches, Bucks: British Arachnological Society." type="book chapter" year="2001">Braunwalder, 2001: 281</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(in part; Gondo, Zwischbergental); Vignoli & Salomone, 2008: 206 (in part);
|
||
<treatmentCitation id="0C666E764745861EFC06FB5D94134352" author="FET" box="[929,1057,1273,1297]" page="6" pageId="8" pageNumber="7" year="2010">
|
||
<bibRefCitation id="E95635964745861EFC06FB5D94134352" author="FET" box="[929,1057,1273,1297]" pageId="8" pageNumber="7" pagination="3 - 12" refId="ref26702" refString="FET, V. 2010. Scorpions of Europe. Acta Zoologica Bulgarica, 62 (1): 3 - 12." type="journal article" year="2010">Fet, 2010: 6</bibRefCitation>
|
||
</treatmentCitation>
|
||
(in part).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8D7848674745861EFC97FABD94B54312" blockId="8.[816,1474,1113,1363]" pageId="8" pageNumber="7">
|
||
<treatmentCitationGroup id="ADD76F494745861EFC97FABD94B54312" pageId="8" pageNumber="7">
|
||
<taxonomicName id="4AC733E44745861EFC97FABD95234372" authority="Karyotypic" authorityName="Karyotypic" box="[816,1297,1305,1329]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="alpha" subGenus="Alpiscorpius">
|
||
<emphasis id="BFB394754745861EFC97FABD939A4373" box="[816,936,1305,1328]" italics="true" pageId="8" pageNumber="7">Euscorpius</emphasis>
|
||
(
|
||
<emphasis id="BFB394754745861EFC1DFABD94734372" box="[954,1089,1305,1329]" italics="true" pageId="8" pageNumber="7">Alpiscorpius</emphasis>
|
||
)
|
||
<emphasis id="BFB394754745861EFBF4FABD94A24372" box="[1107,1168,1305,1329]" italics="true" pageId="8" pageNumber="7">alpha</emphasis>
|
||
Karyotypic
|
||
</taxonomicName>
|
||
race I (Eal 60):
|
||
<treatmentCitation id="0C666E764745861EFCC7FA9D94B14312" author="STUNDLOVA" box="[864,1155,1337,1363]" page="155" pageId="8" pageNumber="7" year="2019">
|
||
<bibRefCitation id="E95635964745861EFCC7FA9D94B14312" author="STUNDLOVA" box="[864,1155,1337,1363]" pageId="8" pageNumber="7" pagination="152 - 163" refId="ref28276" refString="STUNDLOVA, J., J. SMID, P. NGUYEN & F. STAHLAVSKY. 2019. Cryptic diversity and dynamic chromosome evolution in Alpine scorpions (Euscorpiidae: Euscorpius). Molecular Phylogenetics and Evolution, 134: 152 - 163." type="journal article" year="2019">ŠtundlovÁ et al., 2019: 155</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C5DD1BEC4745861EFC97FADD957D40B2" pageId="8" pageNumber="7" type="materials_examined">
|
||
<paragraph id="8D7848674745861EFC97FADD94A44073" blockId="8.[816,1475,1401,1585]" pageId="8" pageNumber="7">
|
||
TYPE LOCALITY AND TYPE DEPOSITORY.
|
||
<emphasis id="BFB394754745861EFAA1FADD950943D2" bold="true" box="[1286,1339,1401,1425]" pageId="8" pageNumber="7">
|
||
<collectingCountry id="F5D008F74745861EFAA1FADD950943D2" box="[1286,1339,1401,1425]" name="Italy" pageId="8" pageNumber="7">Italy</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="BFB394754745861EFAF3FADD958843D2" box="[1364,1466,1401,1425]" italics="true" pageId="8" pageNumber="7">
|
||
<collectingRegion id="4F0386854745861EFAF3FADD958843D2" box="[1364,1466,1401,1425]" country="Italy" name="Piemonte" pageId="8" pageNumber="7">Piedmont</collectingRegion>
|
||
</emphasis>
|
||
: Novara, Monte Massone, [
|
||
<geoCoordinate id="E8F32EA04745861EFBEFFA3D948743F2" box="[1096,1205,1433,1457]" degrees="45.947" direction="north" orientation="latitude" pageId="8" pageNumber="7" precision="55" value="45.947">45.947°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E8F32EA04745861EFB18FA3D952A43F2" box="[1215,1304,1433,1457]" degrees="8.337" direction="east" orientation="longitude" pageId="8" pageNumber="7" precision="55" value="8.337">8.337°E</geoCoordinate>
|
||
] (
|
||
<collectionCode id="EBD6D0A24745861EFA8CFA3D954543F2" box="[1323,1399,1433,1457]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="8" pageNumber="7">MZUF</collectionCode>
|
||
5588),
|
||
<typeStatus id="527CF6C54745861EFC97FA1D93A04392" box="[816,914,1465,1489]" pageId="8" pageNumber="7" type="lectotype">lectotype</typeStatus>
|
||
<specimenCount id="9BC183EE4745861EFC3EFA1D939F4393" box="[921,941,1465,1488]" pageId="8" pageNumber="7" type="male">♂</specimenCount>
|
||
designated by
|
||
<bibRefCitation id="E95635964745861EFBEAFA1D954D4392" author="GANTENBEIN" box="[1101,1407,1465,1489]" pageId="8" pageNumber="7" pagination="843 - 869" refId="ref27133" refString="GANTENBEIN, B., V. FET, M. BARKER & A. SCHOLL. 2000. Nuclear and mitochondrial markers reveal the eXistence of two parapatric scorpion species in the Alps: Euscorpius germanus (C. L. Koch, 1837) and E. alpha Caporiacco, 1950, stat. nov. (Scorpiones, Euscorpiidae). Revue Suisse de Zoologie, 107 (4): 843 - 869." type="journal article" year="2000">Gantenbein et al. (2000: 856</bibRefCitation>
|
||
, 858).
|
||
<materialsCitation id="3DAF423A4745861EFC97FA7D958C43B2" box="[816,1470,1497,1521]" collectionCode="MZUF" pageId="8" pageNumber="7" specimenCode="MZUF 5589-5593" specimenCount="10" specimenCount-female="5" specimenCount-male="5" typeStatus="paralectotype">
|
||
<typeStatus id="527CF6C54745861EFC97FA7D93E243B2" box="[816,976,1497,1521]" pageId="8" pageNumber="7" type="paralectotype">Paralectotypes</typeStatus>
|
||
:
|
||
<specimenCount id="9BC183EE4745861EFC44FA7D943A43B2" box="[995,1032,1497,1521]" pageId="8" pageNumber="7" type="male">5♂</specimenCount>
|
||
<specimenCount id="9BC183EE4745861EFBAFFA7D941443B2" box="[1032,1062,1497,1521]" pageId="8" pageNumber="7" type="female">5♀</specimenCount>
|
||
(
|
||
<specimenCode id="DD61E01C4745861EFB9AFA7D952343B2" box="[1085,1297,1497,1521]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="8" pageNumber="7">MZUF 5589-5593</specimenCode>
|
||
), type locality
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3DAF423A4745861EFC97FA5D93B64073" collectionCode="MZUF" elevation="2000" location="Colle della Piccola Mologna" pageId="8" pageNumber="7" specimenCode="MZUF 5584-5585" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paralectotype">
|
||
<specimenCount id="9BC183EE4745861EFC97FA5D93644052" box="[816,854,1529,1553]" pageId="8" pageNumber="7" type="male">1♂</specimenCount>
|
||
<specimenCount id="9BC183EE4745861EFCF1FA5D93414052" box="[854,883,1529,1553]" pageId="8" pageNumber="7" type="female">1♀</specimenCount>
|
||
(
|
||
<specimenCode id="DD61E01C4745861EFC2FFA5D94684052" box="[904,1114,1529,1553]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="8" pageNumber="7">MZUF 5584-5585</specimenCode>
|
||
),
|
||
<location id="88181EBC4745861EFBD3FA5D958C4052" LSID="urn:lsid:plazi:treatment:056EF9714745861CFF2DF93294274286:88181EBC4745861EFBD3FA5D958C4052" box="[1140,1470,1529,1553]" name="Colle della Piccola Mologna" pageId="8" pageNumber="7">Colle della Piccola Mologna</location>
|
||
,
|
||
<quantity id="4A3FE5824745861EFC97F9BD93B64073" box="[816,900,1561,1585]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="7" unit="m" value="2000.0">
|
||
<elevation id="06EAAF544745861EFC97F9BD93B64073" box="[816,900,1561,1585]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="7" unit="m" value="2000.0">2000 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3DAF423A4745861EFC28F9BD94A34073" box="[911,1169,1561,1585]" collectionCode="MZUF" location="Lamorano" pageId="8" pageNumber="7" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
|
||
<specimenCount id="9BC183EE4745861EFC28F9BD93824072" box="[911,944,1561,1585]" pageId="8" pageNumber="7" type="female">2♀</specimenCount>
|
||
(
|
||
<collectionCode id="EBD6D0A24745861EFC18F9BD943D4072" box="[959,1039,1561,1585]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="8" pageNumber="7">MZUF</collectionCode>
|
||
),
|
||
<location id="88181EBC4745861EFB87F9BD94A34073" LSID="urn:lsid:plazi:treatment:056EF9714745861CFF2DF93294274286:88181EBC4745861EFB87F9BD94A34073" box="[1056,1169,1561,1584]" name="Lamorano" pageId="8" pageNumber="7">Lamorano</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8D7848674745861EFC97F9FD957D40B2" blockId="8.[816,1474,1625,1777]" pageId="8" pageNumber="7">
|
||
<materialsCitation id="3DAF423A4745861EFC97F9FD94DE4032" box="[816,1260,1625,1649]" collectionCode="NMPC" pageId="8" pageNumber="7" specimenCount="12" specimenCount-female="8" specimenCount-male="4">
|
||
MATERIAL EXAMINED (
|
||
<collectionCode id="EBD6D0A24745861EFB92F9FD94B54032" box="[1077,1159,1625,1649]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="7">NMPC</collectionCode>
|
||
):
|
||
<specimenCount id="9BC183EE4745861EFB00F9FD94FC4032" box="[1191,1230,1625,1649]" pageId="8" pageNumber="7" type="male">4♂</specimenCount>
|
||
<specimenCount id="9BC183EE4745861EFB69F9FD94DE4032" box="[1230,1260,1625,1649]" pageId="8" pageNumber="7" type="female">8♀</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3DAF423A4745861EFAA3F9FD94DE40D2" country="Italy" latitude="45.662" location="Tavigliano" longLatPrecision="67" longitude="8.072" pageId="8" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Piemonte">
|
||
<emphasis id="BFB394754745861EFAA3F9FD950B4032" bold="true" box="[1284,1337,1625,1649]" pageId="8" pageNumber="7">
|
||
<collectingCountry id="F5D008F74745861EFAA3F9FD950B4032" box="[1284,1337,1625,1649]" name="Italy" pageId="8" pageNumber="7">Italy</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="BFB394754745861EFAF3F9FD95884032" box="[1364,1466,1625,1649]" italics="true" pageId="8" pageNumber="7">
|
||
<collectingRegion id="4F0386854745861EFAF3F9FD95884032" box="[1364,1466,1625,1649]" country="Italy" name="Piemonte" pageId="8" pageNumber="7">Piedmont</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<location id="88181EBC4745861EFC97F9DD939640D2" LSID="urn:lsid:plazi:treatment:056EF9714745861CFF2DF93294274286:88181EBC4745861EFC97F9DD939640D2" box="[816,932,1657,1681]" country="Italy" latitude="45.662" longLatPrecision="67" longitude="8.072" name="Tavigliano" pageId="8" pageNumber="7" stateProvince="Piemonte">Tavigliano</location>
|
||
,
|
||
<geoCoordinate id="E8F32EA04745861EFC0AF9DD942640D2" box="[941,1044,1657,1681]" degrees="45.662" direction="north" orientation="latitude" pageId="8" pageNumber="7" precision="55" value="45.662">45.662°N</geoCoordinate>
|
||
<geoCoordinate id="E8F32EA04745861EFBBEF9DD944040D2" box="[1049,1138,1657,1681]" degrees="8.072" direction="east" orientation="longitude" pageId="8" pageNumber="7" precision="55" value="8.072">8.072°E</geoCoordinate>
|
||
,
|
||
<specimenCount id="9BC183EE4745861EFBDCF9DD94AE40D2" box="[1147,1180,1657,1681]" pageId="8" pageNumber="7" type="female">1♀</specimenCount>
|
||
(S647)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3DAF423A4745861EFB52F9DD940F40F2" country="Italy" latitude="45.902" location="Loreglia" longLatPrecision="67" longitude="8.37" pageId="8" pageNumber="7" specimenCount="2" specimenCount-male="2" stateProvince="Piemonte">
|
||
<location id="88181EBC4745861EFB52F9DD956040D2" LSID="urn:lsid:plazi:treatment:056EF9714745861CFF2DF93294274286:88181EBC4745861EFB52F9DD956040D2" box="[1269,1362,1657,1681]" country="Italy" latitude="45.902" longLatPrecision="67" longitude="8.37" name="Loreglia" pageId="8" pageNumber="7" stateProvince="Piemonte">Loreglia</location>
|
||
,
|
||
<geoCoordinate id="E8F32EA04745861EFAFCF9DD95F040D2" box="[1371,1474,1657,1681]" degrees="45.902" direction="north" orientation="latitude" pageId="8" pageNumber="7" precision="55" value="45.902">45.902°N</geoCoordinate>
|
||
<geoCoordinate id="E8F32EA04745861EFC97F93D93B840F2" box="[816,906,1689,1713]" degrees="8.370" direction="east" orientation="longitude" pageId="8" pageNumber="7" precision="55" value="8.37">8.370°E</geoCoordinate>
|
||
,
|
||
<specimenCount id="9BC183EE4745861EFC34F93D938140F2" box="[915,947,1689,1713]" pageId="8" pageNumber="7" type="male">2♂</specimenCount>
|
||
(S589-S590)
|
||
</materialsCitation>
|
||
<materialsCitation id="3DAF423A4745861EFBE6F93D94B440F2" box="[1089,1158,1689,1713]" country="Italy" isTailEnumeration="true" latitude="45.902" location="Loreglia" longLatPrecision="67" longitude="8.37" pageId="8" pageNumber="7" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Piemonte">
|
||
<specimenCount id="9BC183EE4745861EFBE6F93D945440F2" box="[1089,1126,1689,1713]" pageId="8" pageNumber="7" type="male">1♂</specimenCount>
|
||
<specimenCount id="9BC183EE4745861EFBC2F93D94B440F2" box="[1125,1158,1689,1713]" pageId="8" pageNumber="7" type="female">3♀</specimenCount>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3DAF423A4745861EFB37F93D944F4092" country="Italy" latitude="45.788" location="Andre" longLatPrecision="67" longitude="7.589" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Valle d'Aosta">
|
||
<emphasis id="BFB394754745861EFB37F93D953840F2" box="[1168,1290,1689,1713]" italics="true" pageId="8" pageNumber="7">
|
||
<collectingRegion id="4F0386854745861EFB37F93D953840F2" box="[1168,1290,1689,1713]" country="Italy" name="Valle d'Aosta" pageId="8" pageNumber="7">Val d’Aosta</collectingRegion>
|
||
</emphasis>
|
||
: Antey-St
|
||
<location id="88181EBC4745861EFADEF93D958C40F2" LSID="urn:lsid:plazi:treatment:056EF9714745861CFF2DF93294274286:88181EBC4745861EFADEF93D958C40F2" box="[1401,1470,1689,1713]" country="Italy" latitude="45.788" longLatPrecision="67" longitude="7.589" name="Andre" pageId="8" pageNumber="7" stateProvince="Valle d'Aosta">André</location>
|
||
,
|
||
<geoCoordinate id="E8F32EA04745861EFC97F91D93AA4092" box="[816,920,1721,1745]" degrees="45.788" direction="north" orientation="latitude" pageId="8" pageNumber="7" precision="55" value="45.788">45.788°N</geoCoordinate>
|
||
<geoCoordinate id="E8F32EA04745861EFC06F91D93C84092" box="[929,1018,1721,1745]" degrees="7.589" direction="east" orientation="longitude" pageId="8" pageNumber="7" precision="55" value="7.589">7.589°E</geoCoordinate>
|
||
,
|
||
<specimenCount id="9BC183EE4745861EFBA0F91D941A4092" box="[1031,1064,1721,1745]" pageId="8" pageNumber="7" type="male">1♂</specimenCount>
|
||
(S098)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3DAF423A4745861EFB2EF91C949A40B2" country="Switzerland" latitude="46.195" location="Gondo" longLatPrecision="67" longitude="8.137" pageId="8" pageNumber="7" specimenCount="2" specimenCount-male="2" stateProvince="Valais">
|
||
<emphasis id="BFB394754745861EFB2EF91C95234093" bold="true" box="[1161,1297,1720,1744]" pageId="8" pageNumber="7">
|
||
<collectingCountry id="F5D008F74745861EFB2EF91C95234093" box="[1161,1297,1720,1744]" name="Switzerland" pageId="8" pageNumber="7">Switzerland</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="BFB394754745861EFA86F91D95504092" box="[1313,1378,1721,1745]" italics="true" pageId="8" pageNumber="7">
|
||
<collectingRegion id="4F0386854745861EFA86F91D95504092" box="[1313,1378,1721,1745]" country="Switzerland" name="Valais" pageId="8" pageNumber="7">Valais</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<location id="88181EBC4745861EFAD4F91D958C4092" LSID="urn:lsid:plazi:treatment:056EF9714745861CFF2DF93294274286:88181EBC4745861EFAD4F91D958C4092" box="[1395,1470,1721,1745]" country="Switzerland" latitude="46.195" longLatPrecision="67" longitude="8.137" name="Gondo" pageId="8" pageNumber="7" stateProvince="Valais">Gondo</location>
|
||
,
|
||
<geoCoordinate id="E8F32EA04745861EFC97F97D93AA40B2" box="[816,920,1753,1777]" degrees="46.195" direction="north" orientation="latitude" pageId="8" pageNumber="7" precision="55" value="46.195">46.195°N</geoCoordinate>
|
||
<geoCoordinate id="E8F32EA04745861EFC39F97D93C540B2" box="[926,1015,1753,1777]" degrees="8.137" direction="east" orientation="longitude" pageId="8" pageNumber="7" precision="55" value="8.137">8.137°E</geoCoordinate>
|
||
,
|
||
<specimenCount id="9BC183EE4745861EFBA4F97D941140B2" box="[1027,1059,1753,1777]" pageId="8" pageNumber="7" type="male">2♂</specimenCount>
|
||
(S547, 589)
|
||
</materialsCitation>
|
||
<materialsCitation id="3DAF423A4745861EFB08F97D951240B2" box="[1199,1312,1753,1777]" country="Switzerland" isTailEnumeration="true" latitude="46.195" location="Gondo" longLatPrecision="67" longitude="8.137" pageId="8" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Valais">
|
||
<specimenCount id="9BC183EE4745861EFB08F97D94E240B2" box="[1199,1232,1753,1777]" pageId="8" pageNumber="7" type="female">1♀</specimenCount>
|
||
(S637)
|
||
</materialsCitation>
|
||
<specimenCount id="9BC183EE4745861EFA81F97D957840B2" box="[1318,1354,1753,1777]" pageId="8" pageNumber="7" type="female">3♀</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C5DD1BEC4745861CFC97F8BD914347E7" lastPageId="10" lastPageNumber="9" pageId="8" pageNumber="7" type="diagnosis">
|
||
<paragraph id="8D7848674745861CFC97F8BD914347E7" blockId="8.[816,1475,1817,2033]" lastBlockId="10.[113,771,237,421]" lastPageId="10" lastPageNumber="9" pageId="8" pageNumber="7">
|
||
DIAGNOSIS.
|
||
<taxonomicName id="4AC733E44745861EFC0AF8BD94554172" authorityName="Kovařík & Štundlová & Fet & Šťáhlavský" authorityYear="2019" baseAuthorityName="Di Caporiacco" baseAuthorityYear="1950" box="[941,1127,1817,1841]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="beta">
|
||
<emphasis id="BFB394754745861EFC0AF8BD94554172" box="[941,1127,1817,1841]" italics="true" pageId="8" pageNumber="7">Alpiscorpius beta</emphasis>
|
||
</taxonomicName>
|
||
is a cryptic species of the
|
||
<emphasis id="BFB394754745861EFAD1F8BD95F04172" box="[1398,1474,1817,1841]" italics="true" pageId="8" pageNumber="7">
|
||
“
|
||
<taxonomicName id="4AC733E44745861EFAD9F8BD95F04172" box="[1406,1474,1817,1841]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="alpha">alpha</taxonomicName>
|
||
</emphasis>
|
||
group” of the “
|
||
<taxonomicName id="4AC733E44745861EFC69F89E94054112" authorityName="Kovařík & Štundlová & Fet & Šťáhlavský" authorityYear="2019" baseAuthorityName="Kovařík & Štundlová & Fet & Šťáhlavský" baseAuthorityYear="1837" box="[974,1079,1850,1873]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="germanus">
|
||
<emphasis id="BFB394754745861EFC69F89E94054112" box="[974,1079,1850,1873]" italics="true" pageId="8" pageNumber="7">germanus</emphasis>
|
||
</taxonomicName>
|
||
compleX” defined by
|
||
<bibRefCitation id="E95635964745861EFAB9F89D93454132" author="STUNDLOVA" pageId="8" pageNumber="7" pagination="152 - 163" refId="ref28276" refString="STUNDLOVA, J., J. SMID, P. NGUYEN & F. STAHLAVSKY. 2019. Cryptic diversity and dynamic chromosome evolution in Alpine scorpions (Euscorpiidae: Euscorpius). Molecular Phylogenetics and Evolution, 134: 152 - 163." type="journal article" year="2019">ŠtundlovÁ et al. (2019)</bibRefCitation>
|
||
as “Karyotypic race I (Eal 60)”, which has 2n = 60 chromosomes. The karyotype is composed of 34 metacentric, 14 submetacentric, 2 subtelocentric, and 10 telocentric chromosomes (fig. S1A in
|
||
<bibRefCitation id="E95635964745861EFBE8F81D950D4192" author="STUNDLOVA" box="[1103,1343,1977,2003]" pageId="8" pageNumber="7" pagination="152 - 163" refId="ref28276" refString="STUNDLOVA, J., J. SMID, P. NGUYEN & F. STAHLAVSKY. 2019. Cryptic diversity and dynamic chromosome evolution in Alpine scorpions (Euscorpiidae: Euscorpius). Molecular Phylogenetics and Evolution, 134: 152 - 163." type="journal article" year="2019">ŠtundlovÁ et al., 2019</bibRefCitation>
|
||
). The rDNA clusters are localized in the subterminal region of the long arms of the metacentric chromosome pair 4 (fig. 2A and fig. S1A in
|
||
<bibRefCitation id="E95635964747861CFFD6FEA9916F4766" author="STUNDLOVA" box="[113,349,269,295]" pageId="10" pageNumber="9" pagination="152 - 163" refId="ref28276" refString="STUNDLOVA, J., J. SMID, P. NGUYEN & F. STAHLAVSKY. 2019. Cryptic diversity and dynamic chromosome evolution in Alpine scorpions (Euscorpiidae: Euscorpius). Molecular Phylogenetics and Evolution, 134: 152 - 163." type="journal article" year="2019">ŠtundlovÁ et al., 2019</bibRefCitation>
|
||
). Number of pectinal teeth (Dp) in male usually 8. Number of patellar ventral trichobothria (
|
||
<emphasis id="BFB394754747861CFD3FFE8992864707" box="[664,692,301,324]" italics="true" pageId="10" pageNumber="9">Pv</emphasis>
|
||
) varies from 5 to 6. Reduction of patellar eXternal trichobothria from 4 to 3 rare in series
|
||
<emphasis id="BFB394754747861CFEE4FEC9916447C7" box="[323,342,365,388]" italics="true" pageId="10" pageNumber="9">et</emphasis>
|
||
(<10%) and more common in series
|
||
<emphasis id="BFB394754747861CFD45FEC9933147C8" box="[738,771,365,395]" italics="true" pageId="10" pageNumber="9">
|
||
eb
|
||
<subScript id="11434A224747861CFD5CFED9933147C8" attach="left" box="[763,771,381,395]" fontSize="6" pageId="10" pageNumber="9">a</subScript>
|
||
</emphasis>
|
||
(>10%); see also Notes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="D9B818EF4744861FFFD2F80B94AF41BC" pageId="9" pageNumber="8" startId="9.[117,194,1967,1989]" targetBox="[127,1463,238,1949]" targetPageId="9">
|
||
<paragraph id="8D7848674744861FFFD2F80B94AF41BC" blockId="9.[117,1476,1967,2047]" pageId="9" pageNumber="8">
|
||
<emphasis id="BFB394754744861FFFD2F80B91374187" bold="true" box="[117,261,1967,1989]" pageId="9" pageNumber="8">Figures 13–22</emphasis>
|
||
:
|
||
<taxonomicName id="4AC733E44744861FFEB5F81491894186" authorityName="Kovařík & Štundlová & Fet & Šťáhlavský" authorityYear="2019" baseAuthorityName="Di Caporiacco" baseAuthorityYear="1950" box="[274,443,1968,1989]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="beta" status="comb. nov.">
|
||
<emphasis id="BFB394754744861FFEB5F81491894186" box="[274,443,1968,1989]" italics="true" pageId="9" pageNumber="8">Alpiscorpius beta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A480290E4744861FFE66F814922A4186" box="[449,536,1968,1989]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="BFB394754744861FFE66F81491CB4186" bold="true" box="[449,505,1968,1989]" pageId="9" pageNumber="8">comb</emphasis>
|
||
.
|
||
<emphasis id="BFB394754744861FFDA1F81492214186" bold="true" box="[518,531,1968,1989]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="BFB394754744861FFD81F81492794186" bold="true" box="[550,587,1968,1989]" pageId="9" pageNumber="8">stat</emphasis>
|
||
.
|
||
<emphasis id="BFB394754744861FFDF0F81492564186" bold="true" box="[599,612,1968,1989]" pageId="9" pageNumber="8">n</emphasis>
|
||
., male (13–17, No. S102) from Switzerland, Gondo, 46.195°N 8.137°E and
|
||
<taxonomicName id="4AC733E44744861FFAEDF814909741A2" authority="Kovařík & Štundlová & Fet & Šťáhlavský, 2019" authorityName="Kovařík & Štundlová & Fet & Šťáhlavský" authorityYear="2019" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="delta" status="sp. nov.">
|
||
<emphasis id="BFB394754744861FFAEDF814909741A2" italics="true" pageId="9" pageNumber="8">Alpiscorpius delta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A480290E4744861FFF0CF86990D241A1" box="[171,224,1997,2018]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="BFB394754744861FFF0CF86990F041A1" bold="true" box="[171,194,1997,2018]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="BFB394754744861FFF69F86990E941A1" bold="true" box="[206,219,1997,2018]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotype (18–22, No. S277).
|
||
<emphasis id="BFB394754744861FFD9DF86892FA41A2" bold="true" box="[570,712,1996,2018]" pageId="9" pageNumber="8">Figures 13–15</emphasis>
|
||
,
|
||
<emphasis id="BFB394754744861FFD73F868932241A2" bold="true" box="[724,784,1996,2018]" pageId="9" pageNumber="8">18–20</emphasis>
|
||
. Pedipalp chela and patella dorsal (13, 18), eXternal (14, 19), and vental (15, 20) views.
|
||
<emphasis id="BFB394754744861FFEACF84D91A841BD" bold="true" box="[267,410,2025,2047]" pageId="9" pageNumber="8">Figures 16–17</emphasis>
|
||
,
|
||
<emphasis id="BFB394754744861FFE01F84D91D041BD" bold="true" box="[422,482,2025,2047]" pageId="9" pageNumber="8">21–22</emphasis>
|
||
. Metasoma III–V and telson, ventral (16, 21), and lateral (17, 22) views.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C5DD1BEC4747861CFFD6FE6992394047" pageId="10" pageNumber="9" type="description">
|
||
<paragraph id="8D7848674747861CFFD6FE6991814226" blockId="10.[113,772,461,1125]" pageId="10" pageNumber="9">
|
||
DESCRIPTION OF THE MALE. The following description is based primarily on a male from Gondo,
|
||
<collectingCountry id="F5D008F74747861CFE7CFE49926C4446" box="[475,606,493,517]" name="Switzerland" pageId="10" pageNumber="9">Switzerland</collectingCountry>
|
||
. Measurements of this specimen are presented in
|
||
<tableCitation id="C0457DDC4747861CFE60FDA992264466" box="[455,532,525,549]" captionStart="Table 1" captionStartId="5.[111,167,1484,1505]" captionText="Table 1. The currently valid Alpiscorpius taXa, their karyotypes, and distribution." pageId="10" pageNumber="9">Table 1</tableCitation>
|
||
. Carapace, tergites, and pedipalps reddish-brown; carapace with some darker mottling; metasoma dark reddish-brown with dark mottling; telson, legs, and sternites dark yellow to reddish-brown; chelicerae light, slightly reticulated. Anterior edge of carapace essentially straight; smooth and lustrous, lacking any indication of carinae. There are two lateral eyes. Median eyes and tubercle are small in size. Tergites I–VII essentially smooth; tergite VII lacking lateral and median carinal pairs. Sternites III–VII smooth and lustrous; VII lacking lateral and median carinae. Stigmata are small, narrow elliptical. Metasomal segment V smooth without granulation and carinae. Vesicle of telson swollen and elongated, with short highly curved aculeus. Vesicle essentially void of granules, lustrous. Pectinal teeth number 8/7. Pedipalp carinae well-developed (
|
||
<figureCitation id="15FC54E24747861CFDD5FC6992C545A7" box="[626,759,972,997]" captionStart="Figures 13–22" captionStartId="9.[117,194,1967,1989]" captionTargetBox="[127,1463,238,1949]" captionTargetId="figure-90@9.[120,1469,225,1961]" captionTargetPageId="9" captionText="Figures 13–22: Alpiscorpius beta comb. n., stat. n., male (13–17, No. S102) from Switzerland, Gondo, 46.195°N 8.137°E and Alpiscorpius delta sp. n., male holotype (18–22, No. S277). Figures 13–15, 18–20. Pedipalp chela and patella dorsal (13, 18), eXternal (14, 19), and vental (15, 20) views. Figures 16–17, 21–22. Metasoma III–V and telson, ventral (16, 21), and lateral (17, 22) views." pageId="10" pageNumber="9">Figs. 13–15</figureCitation>
|
||
). Trichobothrial pattern
|
||
<typeStatus id="527CF6C54747861CFEC7FC4991BC4246" box="[352,398,1005,1029]" pageId="10" pageNumber="9">type</typeStatus>
|
||
C, neobothriotaXic: chela ventral = 4/4; patellar
|
||
<emphasis id="BFB394754747861CFF5EFBA991204266" box="[249,274,1037,1061]" italics="true" pageId="10" pageNumber="9">eb</emphasis>
|
||
= 4/5,
|
||
<emphasis id="BFB394754747861CFEF9FBA9914D4268" box="[350,383,1037,1067]" italics="true" pageId="10" pageNumber="9">
|
||
eb
|
||
<subScript id="11434A224747861CFED0FBB9914D4268" attach="left" box="[375,383,1053,1067]" fontSize="6" pageId="10" pageNumber="9">a</subScript>
|
||
</emphasis>
|
||
=4/4,
|
||
<emphasis id="BFB394754747861CFE62FBA991DB4266" box="[453,489,1037,1061]" italics="true" pageId="10" pageNumber="9">esb</emphasis>
|
||
= 2/2,
|
||
<emphasis id="BFB394754747861CFD93FBA992614267" box="[564,595,1037,1060]" italics="true" pageId="10" pageNumber="9">em</emphasis>
|
||
= 3/3,
|
||
<emphasis id="BFB394754747861CFD07FBA9928C4267" box="[672,702,1037,1060]" italics="true" pageId="10" pageNumber="9">est</emphasis>
|
||
= 3/3,
|
||
<emphasis id="BFB394754747861CFFD6FB8990B64207" box="[113,132,1069,1092]" italics="true" pageId="10" pageNumber="9">et</emphasis>
|
||
= 4/5; patellar ventral = 5/5. Tarsus of legs with single row of spinules on ventral surface.
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFFD6FB2990E842A6" blockId="10.[113,771,1165,1541]" pageId="10" pageNumber="9">
|
||
VARIABILITY. We scored standard phenotypic markers for 7 eXamined specimens (
|
||
<specimenCount id="9BC183EE4747861CFEC3FB0891B54287" box="[356,391,1196,1220]" pageId="10" pageNumber="9" type="male">5♂</specimenCount>
|
||
,
|
||
<specimenCount id="9BC183EE4747861CFE31FB09918A4286" box="[406,440,1197,1221]" pageId="10" pageNumber="9" type="female">2♀</specimenCount>
|
||
), with the following variation observed:
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFFD6FB4992C04367" blockId="10.[113,771,1165,1541]" pageId="10" pageNumber="9">
|
||
<emphasis id="BFB394754747861CFFD6FB4990C04346" box="[113,242,1261,1285]" italics="true" pageId="10" pageNumber="9">Dp in males</emphasis>
|
||
(n=5): 7/7 (1), 8/8 (3), 8/9 (1); in total,
|
||
<quantity id="4A3FE5824747861CFD3CFB4992F64346" box="[667,708,1261,1285]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="10" pageNumber="9" unit="in" value="7.0">7 in</quantity>
|
||
20% (2),
|
||
<quantity id="4A3FE5824747861CFF39FAA990F54366" box="[158,199,1293,1317]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="10" pageNumber="9" unit="in" value="8.0">8 in</quantity>
|
||
70% (7) and
|
||
<quantity id="4A3FE5824747861CFEFFFAA991B34366" box="[344,385,1293,1317]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="10" pageNumber="9" unit="in" value="9.0">9 in</quantity>
|
||
10% (1); mean = 7.90,
|
||
<collectingRegion id="4F0386854747861CFDD9FAA992924366" box="[638,672,1293,1317]" country="Serbia" name="Sid" pageId="10" pageNumber="9">SD</collectingRegion>
|
||
= 0.57.
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFFD6FA8991C84306" blockId="10.[113,771,1165,1541]" box="[113,506,1324,1349]" pageId="10" pageNumber="9">
|
||
<emphasis id="BFB394754747861CFFD6FA8991374306" box="[113,261,1325,1349]" italics="true" pageId="10" pageNumber="9">Dp in females</emphasis>
|
||
(n=2): 6/4 (1), 6/6 (1).
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFFD6FAE992D243C7" blockId="10.[113,771,1165,1541]" pageId="10" pageNumber="9">
|
||
<emphasis id="BFB394754747861CFFD6FAE990BF4327" box="[113,141,1357,1380]" italics="true" pageId="10" pageNumber="9">Pv</emphasis>
|
||
(n=7): 5/5 (3), 5/6 (1), 6/5 (1), 6/6 (2); in total,
|
||
<quantity id="4A3FE5824747861CFD2CFAE892864326" box="[651,692,1356,1381]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="9" unit="in" value="5.0">5 in</quantity>
|
||
57.14% (8), and
|
||
<quantity id="4A3FE5824747861CFE83FAC8917F43C6" box="[292,333,1388,1413]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="9" unit="in" value="6.0">6 in</quantity>
|
||
42.86% (6); mean = 5.43,
|
||
<collectingRegion id="4F0386854747861CFDCCFAC992BF43C6" box="[619,653,1389,1413]" country="Serbia" name="Sid" pageId="10" pageNumber="9">SD</collectingRegion>
|
||
= 0.51.
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFFD6FA2992394386" blockId="10.[113,771,1165,1541]" pageId="10" pageNumber="9">
|
||
<emphasis id="BFB394754747861CFFD6FA2990B643E7" box="[113,132,1421,1444]" italics="true" pageId="10" pageNumber="9">et</emphasis>
|
||
(n=7): 4/3 (1), 4/4 (13); in total,
|
||
<quantity id="4A3FE5824747861CFE5FFA29921143E6" box="[504,547,1421,1445]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="9" unit="in" value="3.0">3 in</quantity>
|
||
7.14% (1), and
|
||
<quantity id="4A3FE5824747861CFD7FFA29933143E6" box="[728,771,1421,1445]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="9" unit="in" value="4.0">4 in</quantity>
|
||
92.86% (13); mean = 3.93,
|
||
<collectingRegion id="4F0386854747861CFE30FA09918B4386" box="[407,441,1453,1477]" country="Serbia" name="Sid" pageId="10" pageNumber="9">SD</collectingRegion>
|
||
= 0.27.
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFFD6FA6992394047" blockId="10.[113,771,1165,1541]" pageId="10" pageNumber="9">
|
||
<emphasis id="BFB394754747861CFFD6FA6990A143A8" box="[113,147,1485,1515]" italics="true" pageId="10" pageNumber="9">
|
||
eb
|
||
<subScript id="11434A224747861CFF2CFA7990A143A8" attach="left" box="[139,147,1501,1515]" fontSize="6" pageId="10" pageNumber="9">a</subScript>
|
||
</emphasis>
|
||
(n=7): 4/3 (2), 4/4 (12); in total,
|
||
<quantity id="4A3FE5824747861CFE5DFA69921643A6" box="[506,548,1485,1509]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="9" unit="in" value="3.0">3 in</quantity>
|
||
14.29% (2) and
|
||
<quantity id="4A3FE5824747861CFD7EFA69933143A6" box="[729,771,1485,1509]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="9" unit="in" value="4.0">4 in</quantity>
|
||
85.71% (12); mean = 3.86,
|
||
<collectingRegion id="4F0386854747861CFE30FA49918B4046" box="[407,441,1517,1541]" country="Serbia" name="Sid" pageId="10" pageNumber="9">SD</collectingRegion>
|
||
= 0.36.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C5DD1BEC4747861CFFD6F98994274286" pageId="10" pageNumber="9" type="distribution">
|
||
<paragraph id="8D7848674747861CFFD6F98991DD4026" blockId="10.[113,771,1581,1637]" pageId="10" pageNumber="9">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="F5D008F74747861CFEBDF98991794006" box="[282,331,1581,1605]" name="Italy" pageId="10" pageNumber="9">Italy</collectingCountry>
|
||
(Piedmont, Val d’Aosta),
|
||
<collectingCountry id="F5D008F74747861CFD23F98993314006" box="[644,771,1581,1605]" name="Switzerland" pageId="10" pageNumber="9">Switzerland</collectingCountry>
|
||
(
|
||
<collectingRegion id="4F0386854747861CFFDEF9E990F24026" box="[121,192,1613,1637]" country="Switzerland" name="Valais" pageId="10" pageNumber="9">Valais</collectingRegion>
|
||
: Zwischbergental) (
|
||
<figureCitation id="15FC54E24747861CFE35F9E991D04026" box="[402,482,1613,1637]" captionStart="Figure 71" captionStartId="28.[115,183,1204,1226]" captionTargetBox="[113,1474,234,1186]" captionTargetId="figure-33@28.[113,1474,231,1186]" captionTargetPageId="28" captionText="Figure 71. Map showing confirmed distribution of studied species of Alpiscorpius, stat. n." pageId="10" pageNumber="9">Fig. 71</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFFD6F92992F541A6" blockId="10.[113,772,1677,2021]" pageId="10" pageNumber="9">
|
||
NOTES. This taXon was first informally delineated by
|
||
<bibRefCitation id="E95635964747861CFD64F929908B4086" author="CAPRA" pageId="10" pageNumber="9" pagination="199 - 213" refId="ref26586" refString="CAPRA, F. 1939. L' Euscorpius germanus (C. L. Koch) in Italia (Arachnida, Scorpiones). Memorie della Societa Entomologica Italiana, 18 (2): 199 - 213." type="journal article" year="1939">Capra (1939)</bibRefCitation>
|
||
as “Form B” from Val d’ Aosta, including the marginal, westernmost populations of “
|
||
<taxonomicName id="4AC733E44747861CFE51F96992C540A7" authorityName="Kovařík & Štundlová & Fet & Šťáhlavský" authorityYear="2019" baseAuthorityName="Kovařík & Štundlová & Fet & Šťáhlavský" baseAuthorityYear="1837" box="[502,759,1741,1764]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="germanus">
|
||
<emphasis id="BFB394754747861CFE51F96992C540A7" box="[502,759,1741,1764]" italics="true" pageId="10" pageNumber="9">Euscorpius germanus</emphasis>
|
||
</taxonomicName>
|
||
” (sensu lato). It was formally described as a subspecies by
|
||
<bibRefCitation id="E95635964747861CFFD6F8A9915D4167" author="DI CAPORIACCO" box="[113,367,1804,1829]" pageId="10" pageNumber="9" pagination="159 - 230" refId="ref26659" refString="DI CAPORIACCO, L. 1950. Le specie e sottospecie del genere " Euscorpius " viventi in Italia ed in alcune zone confinanti. Memorie / Accademia nazionale dei Lincei (ser. 8), 2: 159 - 230." type="journal article" year="1950">Di Caporiacco (1950)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="E95635964747861CFE24F8A9920D4166" author="BONACINA" box="[387,575,1805,1829]" pageId="10" pageNumber="9" pagination="47 - 100" refId="ref26484" refString="BONACINA, A. 1980. Sistematica specifica e sottospecifica del complesso " Euscorpius germanus " (Scorpiones, Chactidae). Rivista del Museo Civico di Scienze Naturali " Enrico Caffi " (Bergamo), 2: 47 - 100." type="journal article" year="1980">Bonacina (1980)</bibRefCitation>
|
||
synonymized
|
||
<taxonomicName id="4AC733E44747861CFD4BF8A9908D4106" authorityName="Di Caporiacco" authorityYear="1950" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="germanus" subSpecies="beta">
|
||
<emphasis id="BFB394754747861CFD4BF8A9908D4106" italics="true" pageId="10" pageNumber="9">E. g. beta</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4AC733E44747861CFEA1F88991834107" authorityName="germanus: Bonacina" authorityYear="1980" box="[262,433,1837,1860]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="germanus" subSpecies="germanus">
|
||
<emphasis id="BFB394754747861CFEA1F88991834107" box="[262,433,1837,1860]" italics="true" pageId="10" pageNumber="9">E. g. germanus</emphasis>
|
||
</taxonomicName>
|
||
and noted that both taXa had five trichobothria on ventral aspect of patella (
|
||
<emphasis id="BFB394754747861CFD26F8E992AF4127" box="[641,669,1869,1892]" italics="true" pageId="10" pageNumber="9">Pv</emphasis>
|
||
=5). This synonymy, however, created a disjunct distribution for
|
||
<taxonomicName id="4AC733E44747861CFD68F8C991E941E6" authority="sensu Bonacina (1980)" authorityName="Bonacina" authorityYear="1980" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="germanus" subSpecies="germanus">
|
||
<emphasis id="BFB394754747861CFD68F8C990E841E7" italics="true" pageId="10" pageNumber="9">E. g. germanus</emphasis>
|
||
sensu
|
||
<bibRefCitation id="E95635964747861CFE82F82991E941E6" author="BONACINA" box="[293,475,1933,1957]" pageId="10" pageNumber="9" pagination="47 - 100" refId="ref26484" refString="BONACINA, A. 1980. Sistematica specifica e sottospecifica del complesso " Euscorpius germanus " (Scorpiones, Chactidae). Rivista del Museo Civico di Scienze Naturali " Enrico Caffi " (Bergamo), 2: 47 - 100." type="journal article" year="1980">Bonacina (1980)</bibRefCitation>
|
||
</taxonomicName>
|
||
. Italian populations studied by
|
||
<bibRefCitation id="E95635964747861CFF36F80991704186" author="BONACINA" box="[145,322,1965,1989]" pageId="10" pageNumber="9" pagination="47 - 100" refId="ref26484" refString="BONACINA, A. 1980. Sistematica specifica e sottospecifica del complesso " Euscorpius germanus " (Scorpiones, Chactidae). Rivista del Museo Civico di Scienze Naturali " Enrico Caffi " (Bergamo), 2: 47 - 100." type="journal article" year="1980">Bonacina (1980)</bibRefCitation>
|
||
should be reassessed, and their phenotypic markers scored from the material deposited in MCSNB.
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFCC7FF4994B247E7" blockId="10.[816,1475,237,1221]" pageId="10" pageNumber="9">
|
||
Populations from Fontainemore (
|
||
<collectingRegion id="4F0386854747861CFB44FF4995404746" box="[1251,1394,237,261]" country="Italy" name="Valle d'Aosta" pageId="10" pageNumber="9">Val d’Aosta</collectingRegion>
|
||
,
|
||
<collectingCountry id="F5D008F74747861CFA2FFF4995884746" box="[1416,1466,237,261]" name="Italy" pageId="10" pageNumber="9">Italy</collectingCountry>
|
||
) and Gondo, on the Simplon Pass road (
|
||
<collectingCountry id="F5D008F74747861CFB40FEA995594766" box="[1255,1387,269,293]" name="Switzerland" pageId="10" pageNumber="9">Switzerland</collectingCountry>
|
||
,
|
||
<collectingRegion id="4F0386854747861CFADFFEA995894766" box="[1400,1467,269,293]" country="Switzerland" name="Valais" pageId="10" pageNumber="9">Valais</collectingRegion>
|
||
) were studied by
|
||
<bibRefCitation id="E95635964747861CFC4FFE8994C14706" author="GANTENBEIN" box="[1000,1267,301,325]" pageId="10" pageNumber="9" pagination="843 - 869" refId="ref27133" refString="GANTENBEIN, B., V. FET, M. BARKER & A. SCHOLL. 2000. Nuclear and mitochondrial markers reveal the eXistence of two parapatric scorpion species in the Alps: Euscorpius germanus (C. L. Koch, 1837) and E. alpha Caporiacco, 1950, stat. nov. (Scorpiones, Euscorpiidae). Revue Suisse de Zoologie, 107 (4): 843 - 869." type="journal article" year="2000">Gantenbein et al. (2000)</bibRefCitation>
|
||
who synonymized
|
||
<taxonomicName id="4AC733E44747861CFC97FEE993C74726" authorityName="Di Caporiacco" authorityYear="1950" box="[816,1013,333,357]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="germanus" subSpecies="beta">
|
||
<emphasis id="BFB394754747861CFC97FEE993C74726" box="[816,1013,333,357]" italics="true" pageId="10" pageNumber="9">E. germanus beta</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<emphasis id="BFB394754747861CFB9CFEE994934726" box="[1083,1185,333,357]" italics="true" pageId="10" pageNumber="9">
|
||
<taxonomicName id="4AC733E44747861CFB9CFEE994AF4726" box="[1083,1181,333,357]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="alpha">E. alpha</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The Gondo (or Simplon) population was studied in more detail by
|
||
<bibRefCitation id="E95635964747861CFB4DFEC995F047C6" author="BRAUNWALDER" box="[1258,1474,365,389]" pageId="10" pageNumber="9" pagination="279 - 286" refId="ref26527" refString="BRAUNWALDER, M. E. 2001. Scorpions of Switzerland: summary of a faunistic survey. Pp. 279 - 286 in: Fet, V. & P. A. Selden (eds.). Scorpions 2001. In Memoriam Gary A. Polis. Burnham Beeches, Bucks: British Arachnological Society." type="book chapter" year="2001">Braunwalder (2001)</bibRefCitation>
|
||
and Fet & Braunwalder (2005).
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFCC7FE09957C4466" blockId="10.[816,1475,237,1221]" pageId="10" pageNumber="9">
|
||
<bibRefCitation id="E95635964747861CFCC7FE0994B54786" author="GANTENBEIN" box="[864,1159,429,453]" pageId="10" pageNumber="9" pagination="843 - 869" refId="ref27133" refString="GANTENBEIN, B., V. FET, M. BARKER & A. SCHOLL. 2000. Nuclear and mitochondrial markers reveal the eXistence of two parapatric scorpion species in the Alps: Euscorpius germanus (C. L. Koch, 1837) and E. alpha Caporiacco, 1950, stat. nov. (Scorpiones, Euscorpiidae). Revue Suisse de Zoologie, 107 (4): 843 - 869." type="journal article" year="2000">Gantenbein et al. (2000)</bibRefCitation>
|
||
published a
|
||
<emphasis id="BFB394754747861CFA94FE0895694787" box="[1331,1371,428,452]" italics="true" pageId="10" pageNumber="9">16S</emphasis>
|
||
mtDNA sequence (
|
||
<accessionNumber id="9294D5844747861CFC06FE69942647A6" box="[929,1044,461,485]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AJ389379" pageId="10" pageNumber="9">AJ389379</accessionNumber>
|
||
), identical for Gondo and Fontainemore populations, which closely matches those of Eal 60 (=
|
||
<taxonomicName id="4AC733E44747861CFAC8FE49958B4446" authorityName="Kovařík & Štundlová & Fet & Šťáhlavský" authorityYear="2019" baseAuthorityName="Di Caporiacco" baseAuthorityYear="1950" box="[1391,1465,493,517]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="beta">
|
||
<emphasis id="BFB394754747861CFAC8FE49958B4446" box="[1391,1465,493,517]" italics="true" pageId="10" pageNumber="9">A. beta</emphasis>
|
||
</taxonomicName>
|
||
) obtained independently by
|
||
<bibRefCitation id="E95635964747861CFBF5FDA995784466" author="STUNDLOVA" box="[1106,1354,525,551]" pageId="10" pageNumber="9" pagination="152 - 163" refId="ref28276" refString="STUNDLOVA, J., J. SMID, P. NGUYEN & F. STAHLAVSKY. 2019. Cryptic diversity and dynamic chromosome evolution in Alpine scorpions (Euscorpiidae: Euscorpius). Molecular Phylogenetics and Evolution, 134: 152 - 163." type="journal article" year="2019">ŠtundlovÁ et al. (2019)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFCC7FD8994904547" blockId="10.[816,1475,237,1221]" pageId="10" pageNumber="9">
|
||
Following
|
||
<bibRefCitation id="E95635964747861CFC7DFD8994DC4406" author="GANTENBEIN" box="[986,1262,557,581]" pageId="10" pageNumber="9" pagination="843 - 869" refId="ref27133" refString="GANTENBEIN, B., V. FET, M. BARKER & A. SCHOLL. 2000. Nuclear and mitochondrial markers reveal the eXistence of two parapatric scorpion species in the Alps: Euscorpius germanus (C. L. Koch, 1837) and E. alpha Caporiacco, 1950, stat. nov. (Scorpiones, Euscorpiidae). Revue Suisse de Zoologie, 107 (4): 843 - 869." type="journal article" year="2000">Gantenbein et al. (2000)</bibRefCitation>
|
||
, both
|
||
<bibRefCitation id="E95635964747861CFA9FFD89934A4426" author="BRAUNWALDER" pageId="10" pageNumber="9" pagination="279 - 286" refId="ref26527" refString="BRAUNWALDER, M. E. 2001. Scorpions of Switzerland: summary of a faunistic survey. Pp. 279 - 286 in: Fet, V. & P. A. Selden (eds.). Scorpions 2001. In Memoriam Gary A. Polis. Burnham Beeches, Bucks: British Arachnological Society." type="book chapter" year="2001">Braunwalder (2001)</bibRefCitation>
|
||
and Fet & Braunwalder (2005) addressed the Swiss population from Simplon as
|
||
<taxonomicName id="4AC733E44747861CFB31FDC994CF44C6" box="[1174,1277,621,645]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="alpha">
|
||
<emphasis id="BFB394754747861CFB31FDC994CF44C6" box="[1174,1277,621,645]" italics="true" pageId="10" pageNumber="9">E. alpha</emphasis>
|
||
</taxonomicName>
|
||
. However, both
|
||
<bibRefCitation id="E95635964747861CFC97FD29940044E6" author="GANTENBEIN" box="[816,1074,653,677]" pageId="10" pageNumber="9" pagination="843 - 869" refId="ref27133" refString="GANTENBEIN, B., V. FET, M. BARKER & A. SCHOLL. 2000. Nuclear and mitochondrial markers reveal the eXistence of two parapatric scorpion species in the Alps: Euscorpius germanus (C. L. Koch, 1837) and E. alpha Caporiacco, 1950, stat. nov. (Scorpiones, Euscorpiidae). Revue Suisse de Zoologie, 107 (4): 843 - 869." type="journal article" year="2000">Gantenbein et al. (2000)</bibRefCitation>
|
||
and Fet & Braunwalder (2005) noted morphological features (Pv=5 to 6, mean 5.40) distinguishing it from the more eastern Swiss populations of
|
||
<taxonomicName id="4AC733E44747861CFAB7FD69955B44A6" box="[1296,1385,717,741]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="alpha">
|
||
<emphasis id="BFB394754747861CFAB7FD69955B44A6" box="[1296,1385,717,741]" italics="true" pageId="10" pageNumber="9">E. alpha</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingRegion id="4F0386854747861CFAD2FD69958C44A6" box="[1397,1470,717,741]" country="Switzerland" name="Ticino" pageId="10" pageNumber="9">Ticino</collectingRegion>
|
||
,
|
||
<collectingRegion id="4F0386854747861CFC97FD4993B54546" box="[816,903,749,773]" country="Switzerland" name="Graubuenden" pageId="10" pageNumber="9">Grisons</collectingRegion>
|
||
), which usually had Pv=6.
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFCC7FCA994F14526" blockId="10.[816,1475,237,1221]" pageId="10" pageNumber="9">
|
||
<materialsCitation id="3DAF423A4747861CFCC7FCA994F24526" collectorName="Fet & Braunwalder" country="Switzerland" location="Switzerland" pageId="10" pageNumber="9" specimenCount="25">
|
||
<collectorName id="20322DB14747861CFCC7FCA993B04567" box="[864,898,781,804]" pageId="10" pageNumber="9">Fet</collectorName>
|
||
&
|
||
<collectorName id="20322DB14747861CFC06FCA994194566" box="[929,1067,781,805]" pageId="10" pageNumber="9">Braunwalder</collectorName>
|
||
(2005) scored the standard phenotypic markers for
|
||
<specimenCount id="9BC183EE4747861CFC1CFC88947D4506" box="[955,1103,812,837]" pageId="10" pageNumber="9" type="generic">25 specimens</specimenCount>
|
||
of this species from
|
||
<collectingCountry id="F5D008F74747861CFAE4FC8995F04506" box="[1347,1474,813,837]" name="Switzerland" pageId="10" pageNumber="9">Switzerland</collectingCountry>
|
||
(Simplon), with the following results
|
||
</materialsCitation>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFC97FCC994C945C7" blockId="10.[816,1475,237,1221]" box="[816,1275,876,901]" pageId="10" pageNumber="9">
|
||
<emphasis id="BFB394754747861CFC97FCC9938345C6" box="[816,945,877,901]" italics="true" pageId="10" pageNumber="9">Dp in males</emphasis>
|
||
(n=6): mean= 8.17,
|
||
<collectingRegion id="4F0386854747861CFB2AFCC9949D45C6" box="[1165,1199,877,901]" country="Serbia" name="Sid" pageId="10" pageNumber="9">SD</collectingRegion>
|
||
= 0.58.
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFC97FC29951345E7" blockId="10.[816,1475,237,1221]" box="[816,1313,908,933]" pageId="10" pageNumber="9">
|
||
<emphasis id="BFB394754747861CFC97FC2993F645E6" box="[816,964,909,933]" italics="true" pageId="10" pageNumber="9">Dp in females</emphasis>
|
||
(n=17): mean = 6.71,
|
||
<collectingRegion id="4F0386854747861CFB13FC2994E445E6" box="[1204,1238,909,933]" country="Serbia" name="Sid" pageId="10" pageNumber="9">SD</collectingRegion>
|
||
= 0.52,
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFC97FC09949B4587" blockId="10.[816,1475,237,1221]" box="[816,1193,940,965]" pageId="10" pageNumber="9">
|
||
<emphasis id="BFB394754747861CFC97FC09937E4587" box="[816,844,941,964]" italics="true" pageId="10" pageNumber="9">Pv</emphasis>
|
||
(n=25): mean = 5.40,
|
||
<collectingRegion id="4F0386854747861CFB9BFC09946C4586" box="[1084,1118,941,965]" country="Serbia" name="Sid" pageId="10" pageNumber="9">SD</collectingRegion>
|
||
= 0.50,
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFC97FC6993974247" blockId="10.[816,1475,237,1221]" pageId="10" pageNumber="9">
|
||
<emphasis id="BFB394754747861CFC97FC69937145A7" box="[816,835,973,996]" italics="true" pageId="10" pageNumber="9">et</emphasis>
|
||
(n=23):
|
||
<quantity id="4A3FE5824747861CFC06FC6993F845A6" box="[929,970,973,997]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="9" unit="in" value="3.0">3 in</quantity>
|
||
2.17% (1), and
|
||
<quantity id="4A3FE5824747861CFBD2FC6994AF45A6" box="[1141,1181,973,997]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="9" unit="in" value="4.0">4 in</quantity>
|
||
97.83% (45); mean = 3.98,
|
||
<collectingRegion id="4F0386854747861CFC97FC4993604246" box="[816,850,1005,1029]" country="Serbia" name="Sid" pageId="10" pageNumber="9">SD</collectingRegion>
|
||
= 0.15.
|
||
</paragraph>
|
||
<paragraph id="8D7848674747861CFCC7FBA994274286" blockId="10.[816,1475,237,1221]" pageId="10" pageNumber="9">
|
||
These data are consistent with the 7 eXamined specimens used for karyotype and DNA phylogeny study by
|
||
<bibRefCitation id="E95635964747861CFAF1FB8993854226" author="STUNDLOVA" pageId="10" pageNumber="9" pagination="152 - 163" refId="ref28276" refString="STUNDLOVA, J., J. SMID, P. NGUYEN & F. STAHLAVSKY. 2019. Cryptic diversity and dynamic chromosome evolution in Alpine scorpions (Euscorpiidae: Euscorpius). Molecular Phylogenetics and Evolution, 134: 152 - 163." type="journal article" year="2019">ŠtundlovÁ et al. (2019)</bibRefCitation>
|
||
. We confirm that males indeed have a higher Dp value than
|
||
<taxonomicName id="4AC733E44747861CFC00FBC9943142C6" box="[935,1027,1133,1157]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="alpha">
|
||
<emphasis id="BFB394754747861CFC00FBC9943142C6" box="[935,1027,1133,1157]" italics="true" pageId="10" pageNumber="9">A. alpha</emphasis>
|
||
</taxonomicName>
|
||
(8 vs. 7),
|
||
<emphasis id="BFB394754747861CFBD5FBC994BC42C7" box="[1138,1166,1133,1156]" italics="true" pageId="10" pageNumber="9">Pv</emphasis>
|
||
varies between 5 and 6, and reduction of
|
||
<emphasis id="BFB394754747861CFC10FB2993F842E7" box="[951,970,1165,1188]" italics="true" pageId="10" pageNumber="9">et</emphasis>
|
||
from 4 to 3 is rare (ca. 2%, only one case out of 46 scored pedipalps).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |