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<document ID-CLB-Dataset="171017" ID-DOI="http://dx.doi.org/10.3897/evolsyst.6.91418" ID-GBIF-Dataset="5f3155b2-beda-4a2b-a7d0-a187d505dbec" ID-Pensoft-Pub="2535-0730-2-175" ID-Pensoft-UUID="16C7BE0236B45BAA847AF82C613E4244" ID-ZooBank="34F513BA207A4A4B952120F9F3BE046C" ModsDocID="2535-0730-6-2-175" checkinTime="1666388770642" checkinUser="pensoft" docAuthor="Castanheira, Pedro de Souza, Baptista, Renner Luiz Cerqueira &amp; Oliveira, Francisca Samia Martins" docDate="2022" docId="056537D414465376857865C28E957B14" docLanguage="en" docName="EvolutSyst 6(2): 175-210" docOrigin="Evolutionary Systematics 6 (2)" docPubDate="2022-10-21" docSource="http://dx.doi.org/10.3897/evolsyst.6.91418" docTitle="Tetragnatha nitens" docType="treatment" docVersion="5" id="16C7BE0236B45BAA847AF82C613E4244" lastPageNumber="175" masterDocId="16C7BE0236B45BAA847AF82C613E4244" masterDocTitle="Five new species of the long-jawed orb-weaving spider genus Tetragnatha (Araneae, Tetragnathidae) in South America, with a key to the species from Argentina and Brazil" masterLastPageNumber="210" masterPageNumber="175" pageNumber="175" updateTime="1732741109323" updateUser="ExternalLinkService">
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<mods:title id="EE2421E3D8B8928B17E649E0222B56FF">Five new species of the long-jawed orb-weaving spider genus Tetragnatha (Araneae, Tetragnathidae) in South America, with a key to the species from Argentina and Brazil</mods:title>
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<mods:name id="504DA23E835722BF84D000CD35D3CA30" type="personal">
<mods:role id="EC37B1C799D5A5A4D51BF54BC9A44C8F">
<mods:roleTerm id="C5B04D2747D0920D9EAD0A571300E73B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="07A9FE3F256655C44CE90AEEE009D6A3">Castanheira, Pedro de Souza</mods:namePart>
<mods:nameIdentifier id="516E7F1EE09EC7D11C73E0EE2EFBECA7" type="ORCID">https://orcid.org/0000-0002-0623-1622</mods:nameIdentifier>
<mods:affiliation id="6BB7A580DA44895F2C15A603F9266F92">Harry Butler Institute, Murdoch University, Murdoch, Western Australia 6150, Australia &amp; Laboratorio de Diversidade de Aracnideos, Universidade do Brasil / Universidade Federal do Rio de Janeiro. Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundao, Rio de Janeiro, Brazil</mods:affiliation>
<mods:nameIdentifier id="A44334976135F114DAABFEF24AB6B578" type="email">pedrocastanheira.bio@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="72E56231D556FDF3120D150702A07095" type="personal">
<mods:role id="B74DB37C21C0982B4F598C7AE060BECC">
<mods:roleTerm id="D2DC54FCABE5175F31350A00D75D9359">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E5638740BA29F5214C134F3FA22FF52">Baptista, Renner Luiz Cerqueira</mods:namePart>
<mods:nameIdentifier id="BF4F12D19E946502D271A6B4B1C8329C" type="ORCID">https://orcid.org/0000-0002-2372-5034</mods:nameIdentifier>
<mods:affiliation id="AD92CB624F1F5799FBF9E3366031CC53">Laboratorio de Diversidade de Aracnideos, Universidade do Brasil / Universidade Federal do Rio de Janeiro. Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundao, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:roleTerm id="9D1A274179DAF3AE836B423C4CD7DC54">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8AD2283CF052C1D7B3F5572DA8AA9F4C">Oliveira, Francisca Samia Martins</mods:namePart>
<mods:affiliation id="9C40935B3411D0EA5723CCC368E5BE70">Laboratorio de Diversidade de Aracnideos, Universidade do Brasil / Universidade Federal do Rio de Janeiro. Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundao, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:title id="484C88FCA80BCA9ADD7C9BE106495DDB">Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part id="293E72D6ADEB1DCEC949628E58CB7178">
<mods:date id="E170BD0250C9F3FC0667AD26A83FB35A">2022</mods:date>
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<mods:number id="8BFAF670B515AD8EDBB5C2C5FFCDC832">2022-10-21</mods:number>
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<mods:number id="B7CD346651D0B9589BF901049A6AA9DF">6</mods:number>
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<mods:number id="05888755A3B1DA730C2E2CD23CA92424">2</mods:number>
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<mods:start id="7E93114E979FEF783AE7C2880064A376">175</mods:start>
<mods:end id="154C18BF05B16D2A320627F1E137A24C">210</mods:end>
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<mods:classification id="738B7366A6E666746656046319734494">journal article</mods:classification>
<mods:identifier id="245CC4558EFF83700F0FEECEA03E0408" type="DOI">http://dx.doi.org/10.3897/evolsyst.6.91418</mods:identifier>
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<treatment id="056537D414465376857865C28E957B14" ID-GBIF-Taxon="203124944" LSID="urn:lsid:plazi:treatment:056537D414465376857865C28E957B14" httpUri="http://treatment.plazi.org/id/056537D414465376857865C28E957B14" lastPageNumber="175" pageId="0" pageNumber="175" scope_class="Arachnida" scope_family="Tetragnathidae" scope_order="Araneae">
<subSubSection id="0F1E8A980212250189A19ECD685778D8" pageId="0" pageNumber="175" type="nomenclature">
<paragraph id="97722C8D9ED89DF0692982B0BDBB36FF" pageId="0" pageNumber="175">
<taxonomicName id="8E6B29B538CD1A25577BCAD6C3043C25" ID-CoL="55MFS" LSID="056537D4-1446-5376-8578-65C28E957B14" authority="(Audouin, 1826)" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">Tetragnatha nitens (Audouin, 1826)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="21DBBA09E51C905F325D535B5D946A99" pageId="0" pageNumber="175" type="description">
<paragraph id="A3301763AA550BEEB1638B99FB3CCCBA" pageId="0" pageNumber="175">
<figureCitation id="D6B8F125A85997F552A6DF00F4004F0E" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tetragnatha nitens under T. americana, male. A. Left chelicera, inner view; B. Chelicerae, slanted lower view." httpUri="https://binary.pensoft.net/fig/760564" pageId="0" pageNumber="175">Figs 12</figureCitation>
<figureCitation id="4D23B29029D1F46F4AB51050C0F663A7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tetragnatha nitens under T. labialis, female lectotype (MNHN 4209). A. Left chelicera, upper view; B. Left chelicera, inner view; C. Left chelicera, lower view; D. Original label." httpUri="https://binary.pensoft.net/fig/760565" pageId="0" pageNumber="175">, 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6D2FB9C6A43FCD46ED41C471447112FD" pageId="0" pageNumber="175" type="reference_group">
<paragraph id="2CC7A046367D271899077A20FAE56158" pageId="0" pageNumber="175">
<taxonomicName id="E9A4F546ECEE76AA70720E486AD2DC00" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha extensa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="extensa">Tetragnatha extensa</taxonomicName>
(Linnaeus, 1758): Nicolet, 1849: 516, plate 4, figs 5, 5a-d (female, misidentified).
</paragraph>
<paragraph id="02BEB72F523D45CAC029290943B5691D" pageId="0" pageNumber="175">
<taxonomicName id="FCFA5F53A10B6B5977963732C9F76693" authorityName="Audouin" authorityYear="1826" class="Arachnida" family="Tetragnathidae" genus="Eugnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eugnatha nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">Eugnatha nitens</taxonomicName>
<bibRefCitation id="938172F8F008B37E1AFF9BBD68569140" author="Audouin, V" journalOrPublisher="Histoire Naturelle" pageId="0" pageNumber="175" pagination="1 - 339" refId="B5" refString="Audouin, V, 1826. Explication sommaire des planches d'arachnides de l'Egypte et de la Syrie. In: Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee franҫaise, publie par les ordres de sa Majeste l'Empereur Napoleon le Grand. Paris: Ed. Imprimerie Imperiale. Histoire Naturelle 1 (4): 1 - 339" title="Explication sommaire des planches d'arachnides de l'Egypte et de la Syrie. In: Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee franҫaise, publie par les ordres de sa Majeste l'Empereur Napoleon le Grand. Paris: Ed. Imprimerie Imperiale." volume="1" year="1826">Audouin 1826</bibRefCitation>
: 118, plate 2, fig. 2 (female).
</paragraph>
<paragraph id="A2A19372F20EEB94820A3EC260CDE595" pageId="0" pageNumber="175">
<taxonomicName id="455C500C2B15FB94227B809C7406613F" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">Tetragnatha nitens</taxonomicName>
:
<bibRefCitation id="9B461E0D691B1AEB61CEF5CFD1F4CC16" DOI="https://doi.org/10.5962/bhl.title.61095" author="Walckenaer, CA" journalOrPublisher="Apteres" pageId="0" pageNumber="175" pagination="1 - 549" refId="B72" refString="Walckenaer, CA, 1841. Histoire naturelle des Insects. Paris. Apteres 2: 1 - 549, DOI: https://doi.org/10.5962/bhl.title.61095" title="Histoire naturelle des Insects. Paris." url="https://doi.org/10.5962/bhl.title.61095" volume="2" year="1841">Walckenaer 1841</bibRefCitation>
: 209.
</paragraph>
<paragraph id="72934C616D633F7E414E3635352DD652" pageId="0" pageNumber="175">
<taxonomicName id="34998B83F29BEC707F40AB6771F8769D" authorityName="Nicolet" authorityYear="1849" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">Tetragnatha labialis</taxonomicName>
<bibRefCitation id="0327E368E7DD4401908458DF5A7ED955" author="Gay, C" journalOrPublisher="Zoologia" pageId="0" pageNumber="175" pagination="319 - 543" refId="B52" refString="Gay, C, 1849. Aracnidos. In: Historia fisica y politica de Chile. Paris, Ed. Maude y Renou. Zoologia 3: 319 - 543" title="Aracnidos." volume="3" volumeTitle="Historia fisica y politica de Chile. Paris, Ed. Maude y Renou." year="1849">Nicolet 1849</bibRefCitation>
: 520 (female). New Synonymy.
</paragraph>
<paragraph id="0E40DC5764893E1BD5DF58D4FCC5C442" pageId="0" pageNumber="175">
<taxonomicName id="E4295AFBDB9B2CAB86B9D0DCA2B257F2" authorityName="Nicolet" authorityYear="1849" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">Tetragnatha labialis</taxonomicName>
:
<bibRefCitation id="890374E9BC063A9008EC55223ADEFB39" author="Keyserling, E" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="175" pagination="799 - 856" refId="B37" refString="Keyserling, E, 1865. Beitraege zur Kenntniss der Orbitelae Latr. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 15: 799 - 856" title="Beitraege zur Kenntniss der Orbitelae Latr." volume="15" year="1865">Keyserling 1865</bibRefCitation>
: 851, plate 21, figs 11-13 (male and female misidentified).
</paragraph>
<paragraph id="81FEB32D8EAC141BEBB501A2EF417DAB" pageId="0" pageNumber="175">
<taxonomicName id="CE594D4BFAAD2C27061D374A097AB008" authorityName="Simon" authorityYear="1905" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">Tetragnatha americana</taxonomicName>
Simon, 1896: CV-CVI (sex?), New Synonymy.
</paragraph>
<paragraph id="EBFA39689036EF00177729AD9A37237A" pageId="0" pageNumber="175">
<taxonomicName id="29FA0801B93C21AE922C43DCD06FA732" authorityName="Nicolet" authorityYear="1849" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">Tetragnatha labialis</taxonomicName>
Simon, 1896: CVII (sex?); 1902: 25; 1904: 94.
</paragraph>
<paragraph id="3E5BBD22C48D73ED5C8008B33432F6B6" pageId="0" pageNumber="175">
<taxonomicName id="F06F7DFD32C3BE2D26685E23CEDE31E4" authorityName="Simon" authorityYear="1905" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">Tetragnatha americana</taxonomicName>
: Simon, 1905: 10 (male, female, misidentified).
</paragraph>
</subSubSection>
<subSubSection id="DD91378C3582614CD64579CE9B0452A7" pageId="0" pageNumber="175" type="type material">
<paragraph id="69F809B55F781800099FCF7EA21375A2" pageId="0" pageNumber="175">Type-material.</paragraph>
<paragraph id="F6AA97626424B9A795210C2DE302A695" pageId="0" pageNumber="175">
<taxonomicName id="54942081760875559A9BED05B5777E21" authorityName="Simon" authorityYear="1905" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="22E81F475EF6CE56D75FD35793482949" italics="true" pageId="0" pageNumber="175">Tetragnatha americana</emphasis>
</taxonomicName>
: Unspecified number of syntypes from
<normalizedToken id="928E9AB0755943CBFC1BEE26C7DAEDFD" originalValue="Peñaflor">Penaflor</normalizedToken>
, Santiago, Chile, Lataste Coll., MNHN?, MHNS?,
<emphasis id="8C9DA499A27CAB47788A3A0E12253FE2" bold="true" pageId="0" pageNumber="175">presumed lost</emphasis>
.
<taxonomicName id="C5A8F365E9E65740756FCC713C8BDA3D" authorityName="Nicolet" authorityYear="1849" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="C432806D7ED1EB0F8C2B6DDA380ABE19" italics="true" pageId="0" pageNumber="175">Tetragnatha labialis</emphasis>
</taxonomicName>
: Unspecified number of adult female syntypes from Santiago, Chile, Nicolet Coll. (female MNHN 4209, herein designated as lectotype),
<emphasis id="63673F3783F4FC424DE6B6D2F2B7BD44" bold="true" pageId="0" pageNumber="175">examined</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="6E66FA02B4472F4335F130AC78F57AA2" pageId="0" pageNumber="175" type="additional record">
<paragraph id="BF504F432BBBDF0F8C7DD1CD45341DB2" pageId="0" pageNumber="175">Additional Record.</paragraph>
<paragraph id="291C1FE8116936CC26ACAFD29F056DA0" pageId="0" pageNumber="175">
Chile - • 1 male from Punta Arenas (
<taxonomicName id="F989C8F6F3EFDD36D5A33FD7A57949D5" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="90A5302E438EE877D774D1FE27B6C8AE" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
: Simon det., MNHN 22312),
<emphasis id="F4325AEE4D6100EF2AA8C6856B396156" bold="true" pageId="0" pageNumber="175">examined</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="CBD3D26ED79BD7AFA6C970F22CFF05F8" pageId="0" pageNumber="175" type="notes">
<paragraph id="1AF1316C913E49E4DFE356B0B271E237" pageId="0" pageNumber="175">Notes.</paragraph>
<paragraph id="9A1663D95596EDE42C939720B1C00579" pageId="0" pageNumber="175">
<taxonomicName id="61A2E4D0DA6243FB5209A24B2FC63DA6" authorityName="Simon" authorityYear="1905" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="3370014DC37C399EA9F1D3328C35A2C8" italics="true" pageId="0" pageNumber="175">Tetragnatha americana</emphasis>
</taxonomicName>
was a new name
<bibRefCitation id="B535E24713D8BF258FCBB5F80620FEB3" author="Simon, E" journalOrPublisher="Actes de la Societe Scientifique du Chili" pageId="0" pageNumber="175" pagination="63 - 70" refId="B65" refString="Simon, E, 1896. Etude sur les Arachnides du Chili. Premier memoire. Actes de la Societe Scientifique du Chili 6: 63 - 70" title="Etude sur les Arachnides du Chili. Premier memoire." volume="6" year="1896">Simon (1896</bibRefCitation>
, p. CV-CVI) applied to a species he considered as the same one misidentified as
<taxonomicName id="2287C6AE38A9B3C5A3B0DE602F26C3D3" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. extensa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="extensa">
<emphasis id="8B8320EB4F97915B0A1C0844AB6A24D0" italics="true" pageId="0" pageNumber="175">T. extensa</emphasis>
</taxonomicName>
and redescribed by
<bibRefCitation id="5BC34D1E87AC393E8080491FCBE1DE94" author="Gay, C" journalOrPublisher="Zoologia" pageId="0" pageNumber="175" pagination="319 - 543" refId="B52" refString="Gay, C, 1849. Aracnidos. In: Historia fisica y politica de Chile. Paris, Ed. Maude y Renou. Zoologia 3: 319 - 543" title="Aracnidos." volume="3" volumeTitle="Historia fisica y politica de Chile. Paris, Ed. Maude y Renou." year="1849">Nicolet (1849</bibRefCitation>
, p. 516). He only added the following expression in a list of spiders from Chile assembled by F. Lataste: &quot;
<taxonomicName id="D86E1C84A5CC9B93958271071356B733" authorityName="Simon" authorityYear="1905" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="BB96EB10D2EF2EB6E35630C22080FBB9" italics="true" pageId="0" pageNumber="175">Tetragnatha americana</emphasis>
</taxonomicName>
E. Sim. (=
<taxonomicName id="118DEA407009AE456D7A631D849C0347" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha extensa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="extensa">
<emphasis id="4DD41E092119B6DDB42692C47467EEDB" italics="true" pageId="0" pageNumber="175">Tetragnatha extensa</emphasis>
</taxonomicName>
Nicolet)&quot;. Those specimens were collected in
<normalizedToken id="9D34F924672EB743B4273354D59164F1" originalValue="Peñaflor">Penaflor</normalizedToken>
, Santiago, Chile, and deposited at the &quot;Museo Zoolojico de la Escuela de Medicina&quot;, currently the Museo Nacional de Historia Natural, Chile (MHNS, see
<bibRefCitation id="E79BAB32B6A5A37720E2A57EF6DFEBFA" author="Diaz, CF" journalOrPublisher="Chile, Santiago, Ed. Centro de Investigaciones PEIP" pageId="0" pageNumber="175" refId="B23" refString="Diaz, CF, 2019. Naturalistas en el Chile Decimononico. Chile, Santiago, Ed. Centro de Investigaciones PEIP" title="Naturalistas en el Chile Decimononico." year="2019">
<normalizedToken id="CB77BDB2BDCE02DE7A880DDEB9F57C9D" originalValue="Díaz">Diaz</normalizedToken>
(2019)
</bibRefCitation>
for additional information).
<bibRefCitation id="262968C60BC7126F04EE2FD812865A14" DOI="https://doi.org/10.5962/bhl.part.9304" author="Simon, E" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino" pageId="0" pageNumber="175" pagination="1 - 18" refId="B68" refString="Simon, E, 1905. Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino 20 (511): 1 - 18, DOI: https://doi.org/10.5962/bhl.part.9304" title="Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri." url="https://doi.org/10.5962/bhl.part.9304" volume="20" year="1905">Simon (1905</bibRefCitation>
, p. 10) applied again without any justification the name
<taxonomicName id="1D02E3D27CEA77B6E4908B399A773956" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="56BB5EDA23F28496398CC22F78850DD5" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
to the species he considered as
<taxonomicName id="93C236102E30736ED6CE4043B060FD2D" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. extensa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="extensa">
<emphasis id="AAA30362DAB15FC0FEB24D5D09CE80CE" italics="true" pageId="0" pageNumber="175">T. extensa</emphasis>
</taxonomicName>
sensu Nicolet, citing three females from Santa Cruz, Argentina. He listed
<taxonomicName id="C7A0D86DEA382769A6ABD62E47A05DC3" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. extensa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="extensa">
<emphasis id="D85D4982634C7E65A1739D144FE1A935" italics="true" pageId="0" pageNumber="175">T. extensa</emphasis>
</taxonomicName>
sensu Nicolet under the title of
<taxonomicName id="8A679D1B6EB226E548DCBE0A3F81C030" authorityName="Simon" authorityYear="1905" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="9DD3B1C540E7A538292A008BE2F8BC9E" italics="true" pageId="0" pageNumber="175">Tetragnatha americana</emphasis>
</taxonomicName>
, and included the bibliographical reference to the book, again without mentioning the examination of
<normalizedToken id="2982032FDE6A7DD35910CF55C7852EB3" originalValue="Nicolet’s">Nicolet's</normalizedToken>
specimens or any morphological character.
<bibRefCitation id="00F3E52C13A03567D4CD5678D9E11792" DOI="https://doi.org/10.5962/bhl.part.9304" author="Simon, E" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino" pageId="0" pageNumber="175" pagination="1 - 18" refId="B68" refString="Simon, E, 1905. Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino 20 (511): 1 - 18, DOI: https://doi.org/10.5962/bhl.part.9304" title="Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri." url="https://doi.org/10.5962/bhl.part.9304" volume="20" year="1905">Simon (1905)</bibRefCitation>
added that the specimens he had from Santa Cruz &quot;were not different from
<taxonomicName id="7E1B260F93C0169CF418145FBE2E45F2" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="CAA06E00349C5CE6FDB8894F58BA49B0" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
of Chile, where the species is very common&quot;.
</paragraph>
<paragraph id="092487E65D20F46F4DE96DA2CE6CD180" pageId="0" pageNumber="175">
Despite
<bibRefCitation id="6092A42566379650F8AEC57824D02B09" author="Simon, E" journalOrPublisher="Actes de la Societe Scientifique du Chili" pageId="0" pageNumber="175" pagination="63 - 70" refId="B65" refString="Simon, E, 1896. Etude sur les Arachnides du Chili. Premier memoire. Actes de la Societe Scientifique du Chili 6: 63 - 70" title="Etude sur les Arachnides du Chili. Premier memoire." volume="6" year="1896">Simon (1896</bibRefCitation>
,
<bibRefCitation id="17F4C5FACD948B443A08ACBE5C9EEA6E" DOI="https://doi.org/10.5962/bhl.part.9304" author="Simon, E" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino" pageId="0" pageNumber="175" pagination="1 - 18" refId="B68" refString="Simon, E, 1905. Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino 20 (511): 1 - 18, DOI: https://doi.org/10.5962/bhl.part.9304" title="Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri." url="https://doi.org/10.5962/bhl.part.9304" volume="20" year="1905">1905</bibRefCitation>
) not clearly stating that he intended to add a new replacement name for
<taxonomicName id="FFAE929DB778E5B6482275A19828C7B4" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. extensa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="extensa">
<emphasis id="60DCC823829ABBE88D769A21E0BF81AC" italics="true" pageId="0" pageNumber="175">T. extensa</emphasis>
</taxonomicName>
sensu Nicolet, no previous citation to
<taxonomicName id="F870162C5FD2A8677FCEDEAFD4F09112" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="6E6D6C311CB576D27785F4879CFA331D" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
was found after an analysis of
<normalizedToken id="16E4E5CD00A89B6C846FE366FC803722" originalValue="Simon’s">Simon's</normalizedToken>
papers dealing with spiders from Chile. Following
<bibRefCitation id="4ADAC52F79E4C3402574D4C648158AE4" author="Bonnet, P" journalOrPublisher="Memoires Couronnes et Memoires des Savants Etrangers de l'Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique" pageId="0" pageNumber="175" refId="B7" refString="Bonnet, P, 1959. Bibliographia araneorum. Analyse methodique de toute la litterature araneologique jusqu'en 1939. Tome II. Systematique des araignees (Etude par ordre alphabetique) (5eme partie: T-Z). Douladoure, Toulouse, 4231-5058." title="Bibliographia araneorum. Analyse methodique de toute la litterature araneologique jusqu'en 1939. Tome II. Systematique des araignees (Etude par ordre alphabetique) (5 eme partie: T-Z). Douladoure, Toulouse, 4231 - 5058." year="1959">Bonnet (1959</bibRefCitation>
, sub &quot;
<taxonomicName id="031004BE0051A10D40874D1C10BA07F7" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="569FCCF8772E8AD64CC323BCDF35D7EE" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
Simon, 1897&quot;, p. 4317) and against
<bibRefCitation id="D00C52ED74452E3DCBF2350F89E15202" author="Roewer, CF" journalOrPublisher="Natura, Buchhandlung fuer Naturkunde und exakte Wissenschaften Paul Budy Bremen" pageId="0" pageNumber="175" refId="B60" refString="Roewer, CF, 1942. Katalog der Araneae von 1758 bis 1940. 1. Band (Mesothelae, Orthognatha, Labidognatha: Dysderaeformia, Scytodiformia, Pholciformia, Zodariiformia, Hersiliaeformia, Argyopiformia). Natura, Buchhandlung fuer Naturkunde und exakte Wissenschaften Paul Budy Bremen" title="Katalog der Araneae von 1758 bis 1940. 1. Band (Mesothelae, Orthognatha, Labidognatha: Dysderaeformia, Scytodiformia, Pholciformia, Zodariiformia, Hersiliaeformia, Argyopiformia)." year="1942">Roewer (1942</bibRefCitation>
, p. 988) and the
<bibRefCitation id="283560FF03EC69538D7110E390BA2AE9" author="Wiehle, H" journalOrPublisher="Die Tierwelt Deutschlands" pageId="0" pageNumber="175" publicationUrl="http://wsc.nmbe.ch" refId="B75" refString="2022. . http://wsc.nmbe.ch" url="http://wsc.nmbe.ch" year="2022">World Spider Catalog (2022)</bibRefCitation>
, we consider 1896 and not 1905 as the year of the proposition of the new name
<taxonomicName id="CE4CF8D1635D8B15F52DD549351AF47C" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="31087622C97BBBD69717E43185C8598E" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
. The citation by
<bibRefCitation id="6516D23CFCD88A14EC9F841FF87ED6A2" author="Simon, E" journalOrPublisher="Actes de la Societe Scientifique du Chili" pageId="0" pageNumber="175" pagination="63 - 70" refId="B65" refString="Simon, E, 1896. Etude sur les Arachnides du Chili. Premier memoire. Actes de la Societe Scientifique du Chili 6: 63 - 70" title="Etude sur les Arachnides du Chili. Premier memoire." volume="6" year="1896">Simon (1896</bibRefCitation>
, p. CV) was apparently the first appearance of the name and it is available under the International Code of Zoological Nomenclature (ICZN 1999), following its article 12, which establishes that an unambiguous reference to a
<normalizedToken id="18AD68B4D3F0CFC118525007F5ABB38C" originalValue="species’">species'</normalizedToken>
description suffices to make available a name published before 1931.
</paragraph>
<paragraph id="10F8C61FA4AFD40CD8C1B19E8C29465A" pageId="0" pageNumber="175">
The type-series of
<taxonomicName id="389944FD8275A9E57E3498B7BF95B1DA" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B94149A9BB59C7B579875217F35BC5D5" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
Simon, 1896 is therefore composed of the specimens from
<normalizedToken id="411C05C2FC0EB406AC4ECDC166B09267" originalValue="Peñaflor">Penaflor</normalizedToken>
examined by
<bibRefCitation id="E5C5986CD54502C04FA4F52A1550BEB4" author="Simon, E" journalOrPublisher="Actes de la Societe Scientifique du Chili" pageId="0" pageNumber="175" pagination="63 - 70" refId="B65" refString="Simon, E, 1896. Etude sur les Arachnides du Chili. Premier memoire. Actes de la Societe Scientifique du Chili 6: 63 - 70" title="Etude sur les Arachnides du Chili. Premier memoire." volume="6" year="1896">Simon (1896)</bibRefCitation>
when proposing the new name and also the specimens from Valdivia province studied and illustrated by
<bibRefCitation id="BE3811C0E192CB6C5A0C5FCD946AAFC3" author="Gay, C" journalOrPublisher="Zoologia" pageId="0" pageNumber="175" pagination="319 - 543" refId="B52" refString="Gay, C, 1849. Aracnidos. In: Historia fisica y politica de Chile. Paris, Ed. Maude y Renou. Zoologia 3: 319 - 543" title="Aracnidos." volume="3" volumeTitle="Historia fisica y politica de Chile. Paris, Ed. Maude y Renou." year="1849">Nicolet (1849)</bibRefCitation>
, but not by the specimens from Santa Cruz, Argentina (Fig.
<figureCitation id="249D086FA451AF5AE518A703DD3421DA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tetragnatha laboriosa under T. americana, female (MNHN 3140). A. Dorsal habitus; B. Original label citing Santa Cruz, Argentina; C. Ventral habitus; D. Left chelicera, inner view; E. Chelicerae, apical view." httpUri="https://binary.pensoft.net/fig/760563" pageId="0" pageNumber="175">11</figureCitation>
) and unspecified localities of Chile cited by
<bibRefCitation id="DD245B3CE8ED40459F8270B7F3562A3B" DOI="https://doi.org/10.5962/bhl.part.9304" author="Simon, E" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino" pageId="0" pageNumber="175" pagination="1 - 18" refId="B68" refString="Simon, E, 1905. Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino 20 (511): 1 - 18, DOI: https://doi.org/10.5962/bhl.part.9304" title="Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri." url="https://doi.org/10.5962/bhl.part.9304" volume="20" year="1905">Simon (1905)</bibRefCitation>
. According to the article 72.4.1.1 and its example, all the specimens studied by the
<normalizedToken id="74EA01EF96C9D1AB6A026E4103E0CF92" originalValue="species’">species'</normalizedToken>
author or referred unambiguously to in the original description constitute the type series. As stated by
<bibRefCitation id="F1215BA5458D842A6FCC3DD336FEA7BB" author="Levi, HW" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="0" pageNumber="175" pagination="1 - 20" refId="B42" refString="Levi, HW, 1967. The theridiid spider fauna of Chile. Bulletin of the Museum of Comparative Zoology 136: 1 - 20" title="The theridiid spider fauna of Chile." volume="136" year="1967">Levi (1967</bibRefCitation>
,
<bibRefCitation id="C809C0F31AE77758CA9DF976158F14C4" author="Levi, HW" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="0" pageNumber="175" pagination="445 - 475" refId="B45" refString="Levi, HW, 2001. The orbweavers of the genera Molinaranea and Nicolepeira, a new species of Parawixia, and comments on orb weavers of temperate South America (Araneae: Araneidae). Bulletin of the Museum of Comparative Zoology 155: 445 - 475" title="The orbweavers of the genera Molinaranea and Nicolepeira, a new species of Parawixia, and comments on orb weavers of temperate South America (Araneae: Araneidae)." volume="155" year="2001">2001</bibRefCitation>
), Nicolet probably returned to Paris in 1846 and brought with him a large part of the types of Chilean species he described. Levi was able to find at MNHN types of many species of
<taxonomicName id="718693F55EAEEFD66028953F25505D64" authorityName="Sundevall" authorityYear="1833" class="Arachnida" family="Theridiidae" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="family">Theridiidae</taxonomicName>
, but he considered most
<taxonomicName id="C0AD9160D7B7E20E3DF2D2E5B3B1066D" class="Arachnida" family="Araneidae" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="family">Araneidae</taxonomicName>
types as lost. We also have not been able to find most
<taxonomicName id="33494F1F8FC6C33DD312DB95D11C8DDA" authorityName="Menge" authorityYear="1866" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="family">Tetragnathidae</taxonomicName>
types of species described by Nicolet in MNHN, excepting
<taxonomicName id="0176B8F7412CBEC64F21C7F96A5CCD57" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="6AC5D52B1F47F7CABC4DD0452A33A7CC" italics="true" pageId="0" pageNumber="175">T. labialis</emphasis>
</taxonomicName>
. We requested information about
<taxonomicName id="4F6AE434E4538A73F26EA9FCBD33BE0D" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="475C9A45F7A86534D31239B8D8D49939" italics="true" pageId="0" pageNumber="175">Tetragnatha</emphasis>
</taxonomicName>
specimens from the MHNS but did not receive an answer and there is no evidence that any of Nicolet or
<normalizedToken id="8A092EDF60F64011930198030A5174DA" originalValue="Simon’s">Simon's</normalizedToken>
specimens are still available in this institution. Because
<taxonomicName id="B18EC2C59D0A32DA59209AA922679649" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="1E79710A6769E9C93F2B9265FBE1FCAA" italics="true" pageId="0" pageNumber="175">T. labialis</emphasis>
</taxonomicName>
was found in MNHN, we consider that this was probably the same depository institution for the other species described by
<bibRefCitation id="960C1E019AC5F2D5AF2C6E60D106439F" author="Gay, C" journalOrPublisher="Zoologia" pageId="0" pageNumber="175" pagination="319 - 543" refId="B52" refString="Gay, C, 1849. Aracnidos. In: Historia fisica y politica de Chile. Paris, Ed. Maude y Renou. Zoologia 3: 319 - 543" title="Aracnidos." volume="3" volumeTitle="Historia fisica y politica de Chile. Paris, Ed. Maude y Renou." year="1849">Nicolet (1849)</bibRefCitation>
and the specimens from Valdivia province described as
<taxonomicName id="567D96D47EB7D7FF3C1D371B8DA8A7B1" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. extensa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="extensa">
<emphasis id="FA3113C2DE461B385DB852F3BC456BA5" italics="true" pageId="0" pageNumber="175">T. extensa</emphasis>
</taxonomicName>
. However, except for
<taxonomicName id="0DA72DE0F5F986BD41B547497449CDCF" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="0B670682E7BFF8F4332A10AF054ECEDD" italics="true" pageId="0" pageNumber="175">T. labialis</emphasis>
</taxonomicName>
, no other original specimen from
<bibRefCitation id="3D32769F2743DA17BA88F6FA05F02B48" author="Gay, C" journalOrPublisher="Zoologia" pageId="0" pageNumber="175" pagination="319 - 543" refId="B52" refString="Gay, C, 1849. Aracnidos. In: Historia fisica y politica de Chile. Paris, Ed. Maude y Renou. Zoologia 3: 319 - 543" title="Aracnidos." volume="3" volumeTitle="Historia fisica y politica de Chile. Paris, Ed. Maude y Renou." year="1849">Nicolet (1849)</bibRefCitation>
was found at MNHN. Likewise,
<taxonomicName id="E648943CF546492375298AF00F48D96E" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="95807FEDB60470AE66B88D28E5EEDA9E" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
specimens identified by Simon from
<normalizedToken id="2260ED18FE228303F11558EB020B0B5A" originalValue="Peñaflor">Penaflor</normalizedToken>
and Santiago were also not found in the general collection of the MNHN. Therefore, we consider the syntypes of
<taxonomicName id="4D6E920013B700F5BF54C08A39DAA3C0" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="E4A2AB0B182BD5F022D5DB24A69AAA69" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
presumed lost.
</paragraph>
<paragraph id="7FF812876A503B8C6375138094C45EF6" pageId="0" pageNumber="175">
<normalizedToken id="0BF02FBD67C533EAA374F434814DE392" originalValue="Nicolet’s">Nicolet's</normalizedToken>
description focused on the shape and size of structures such as carapace, eyes, abdomen, chelicerae and pedipalps, but did not include precise details on chelicerae and genital morphology to allow species identification. Fortunately, he also provided relatively good drawings (
<bibRefCitation id="627DEAFFF5838CA1C17585917752356C" author="Gay, C" journalOrPublisher="Zoologia" pageId="0" pageNumber="175" pagination="319 - 543" refId="B52" refString="Gay, C, 1849. Aracnidos. In: Historia fisica y politica de Chile. Paris, Ed. Maude y Renou. Zoologia 3: 319 - 543" title="Aracnidos." volume="3" volumeTitle="Historia fisica y politica de Chile. Paris, Ed. Maude y Renou." year="1849">Nicolet 1849</bibRefCitation>
, plate 4, fig. 5), which clearly depict a female specimen of
<taxonomicName id="421A83B2B709C0D8F5ECF23E14E68982" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="67FBC2386E1C251D76348FFC8294D58C" italics="true" pageId="0" pageNumber="175">T. nitens</emphasis>
</taxonomicName>
, based for example on the sinuous cheliceral fang, Gu and U2 large and of similar size and well-spaced, and AXl large but not overreaching the fang basis. So, the description and illustrations by
<bibRefCitation id="95AD1A8BDC6472EF8368EC8EFD02EC47" author="Gay, C" journalOrPublisher="Zoologia" pageId="0" pageNumber="175" pagination="319 - 543" refId="B52" refString="Gay, C, 1849. Aracnidos. In: Historia fisica y politica de Chile. Paris, Ed. Maude y Renou. Zoologia 3: 319 - 543" title="Aracnidos." volume="3" volumeTitle="Historia fisica y politica de Chile. Paris, Ed. Maude y Renou." year="1849">Nicolet (1849)</bibRefCitation>
are the only reliable information on the type series we recovered.
</paragraph>
<paragraph id="C39953CAD528F8A1C84523F57F37C943" pageId="0" pageNumber="175">
We were able to analyse all the specimens currently present in the MNHN collection and identified as
<taxonomicName id="CB0AABB068A32547D67421B0BB2BB1FE" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="F658FD728CBD9680873A96C158583C51" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
by Simon. Some of these are specimens of
<taxonomicName id="54FC05EB6AC4F14EA65DEE18A3881580" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. laboriosa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="laboriosa">
<emphasis id="FE2396650ED092DC015D3BA83F5662E4" italics="true" pageId="0" pageNumber="175">T. laboriosa</emphasis>
</taxonomicName>
, represented by several specimens from Chile (MNHN 12628) and just one of the three females from Santa Cruz, Argentina (MNHN 3140, Fig.
<figureCitation id="5045088DD3B99590F8E5BA0796836CF5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tetragnatha laboriosa under T. americana, female (MNHN 3140). A. Dorsal habitus; B. Original label citing Santa Cruz, Argentina; C. Ventral habitus; D. Left chelicera, inner view; E. Chelicerae, apical view." httpUri="https://binary.pensoft.net/fig/760563" pageId="0" pageNumber="175">11</figureCitation>
) cited by
<bibRefCitation id="0EE2F8CB259A91C16BB43E206EC9E74B" DOI="https://doi.org/10.5962/bhl.part.9304" author="Simon, E" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino" pageId="0" pageNumber="175" pagination="1 - 18" refId="B68" refString="Simon, E, 1905. Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino 20 (511): 1 - 18, DOI: https://doi.org/10.5962/bhl.part.9304" title="Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri." url="https://doi.org/10.5962/bhl.part.9304" volume="20" year="1905">Simon (1905)</bibRefCitation>
. On the other hand, there is a male from Punta Arenas, Chile, that is clearly
<taxonomicName id="A652F2F0FD8F626B1125A34FF59744CC" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="EA1E4FF1D37426A0ECC32AB9C3B8052E" italics="true" pageId="0" pageNumber="175">T. nitens</emphasis>
</taxonomicName>
(MNHN 22312, Fig.
<figureCitation id="E16BAFB838201FAE80E23D9301BC68E9" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tetragnatha nitens under T. americana, male. A. Left chelicera, inner view; B. Chelicerae, slanted lower view." httpUri="https://binary.pensoft.net/fig/760564" pageId="0" pageNumber="175">12</figureCitation>
). Although these specimens are just additional material examined, they were identified by Simon himself, thus helping to establish his concept of
<taxonomicName id="A6D431FFAF39821E5F86A82F55101119" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="5966824A2289AF5E654DD124359DC01B" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
. Although some specimens Simon identified as
<taxonomicName id="B9493741AF6F337B9F4858281A0DEEBA" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="D650A4AC918075E16602F8B88EA631B7" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
are in fact
<taxonomicName id="5ECA0B7C7CFF9A2AD0A3051171F7E227" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. laboriosa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="laboriosa">
<emphasis id="071FE415380B3355272F50C364F12D78" italics="true" pageId="0" pageNumber="175">T. laboriosa</emphasis>
</taxonomicName>
, the original drawings of a female syntype of &quot;
<taxonomicName id="74F12BC231D9F803B11F9262520D23E8" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. extensa" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="extensa">
<emphasis id="0807A2A881E1F5FA4E2BC9B55863F45B" italics="true" pageId="0" pageNumber="175">T. extensa</emphasis>
</taxonomicName>
&quot; from Nicolet and secondarily the male from Punta Arenas (MNHN 22312, Fig.
<figureCitation id="47F524D768C02C2E69665076307FBD5A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tetragnatha nitens under T. americana, male. A. Left chelicera, inner view; B. Chelicerae, slanted lower view." httpUri="https://binary.pensoft.net/fig/760564" pageId="0" pageNumber="175">12</figureCitation>
) identified by Simon allow us to indicate that
<taxonomicName id="8D762DB8270F1CEBCB2A9F1FD1416357" authorityName="Simon" authorityYear="1905" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="F2DB5783FCD50F91FE741ECA1B9CE5FA" italics="true" pageId="0" pageNumber="175">Tetragnatha americana</emphasis>
</taxonomicName>
Simon, 1896 is
<taxonomicName id="0700B1D4B788A12186E46CB29FD31D0D" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="9AFCF2FC44D1A58B337EE33B8F3D32F5" italics="true" pageId="0" pageNumber="175">Tetragnatha nitens</emphasis>
</taxonomicName>
(Audouin, 1826) syn. nov.
</paragraph>
<caption id="B61ECD1DE40343928C5053E164859ED3" httpUri="https://binary.pensoft.net/fig/760564" pageId="0" pageNumber="175" start="Figure 12" startId="F12">
<paragraph id="FDEDE69E04D2474DA6285EF28916621D" pageId="0" pageNumber="175">
<emphasis id="5D0A04451A22B11CEDAB7EC964A9EB3F" bold="true" pageId="0" pageNumber="175">Figure 12.</emphasis>
<taxonomicName id="4E427B77E12AD65486665083061FA080" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B7B7317AE8B38123FB19AAB40AB07A68" italics="true" pageId="0" pageNumber="175">Tetragnatha nitens</emphasis>
</taxonomicName>
under
<taxonomicName id="8557B4A1759558B63E20FBE1F2650F0E" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="EB22D85DC149C446E31C5D4E959A371B" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
, male.
<emphasis id="3AD1AD646BB9B750F375F946B7916663" bold="true" pageId="0" pageNumber="175">A.</emphasis>
Left chelicera, inner view;
<emphasis id="5927A5A75C618E9DB22A3D6B4047F5B5" bold="true" pageId="0" pageNumber="175">B.</emphasis>
Chelicerae, slanted lower view.
</paragraph>
</caption>
<paragraph id="66B12CBBCF4E81CD93E7DBC7E774D78D" pageId="0" pageNumber="175">
As mentioned above, the only species dealt with by
<bibRefCitation id="C1BB7B1DF78A035B09783E48934C93D5" author="Gay, C" journalOrPublisher="Zoologia" pageId="0" pageNumber="175" pagination="319 - 543" refId="B52" refString="Gay, C, 1849. Aracnidos. In: Historia fisica y politica de Chile. Paris, Ed. Maude y Renou. Zoologia 3: 319 - 543" title="Aracnidos." volume="3" volumeTitle="Historia fisica y politica de Chile. Paris, Ed. Maude y Renou." year="1849">Nicolet (1849)</bibRefCitation>
that we were able to locate at MNHN was
<taxonomicName id="831C45EF6E88CCC1E432E9F1C8BB8B7A" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="69F17049FB6421059ACEA9097A109505" italics="true" pageId="0" pageNumber="175">T. labialis</emphasis>
</taxonomicName>
, described based on an unspecified number of female syntypes from Santiago, Chile. A female specimen (MNHN 4209), labelled as &quot;
<taxonomicName id="D7E24128BCDC9BE175B3FBC27492D393" authorityName="Nicolet" authorityYear="1849" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="CE6436383CDBDED9B2CC5308FE460FBC" italics="true" pageId="0" pageNumber="175">Tetragnatha labialis</emphasis>
</taxonomicName>
Nicol. Gay Saint Iago&quot; (Fig.
<figureCitation id="F94F5BFC8DC3DC2CDFBD4482B111D40D" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tetragnatha nitens under T. labialis, female lectotype (MNHN 4209). A. Left chelicera, upper view; B. Left chelicera, inner view; C. Left chelicera, lower view; D. Original label." httpUri="https://binary.pensoft.net/fig/760565" pageId="0" pageNumber="175">13</figureCitation>
) was examined. This female clearly belongs to the syntype series examined by Nicolet and is herein designated as lectotype. Following
<bibRefCitation id="B03544B1C21F9D3F8BE90CB72EB70F80" DOI="https://doi.org/10.3897/zse.95.36762" author="Castanheira, PdS" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="175" pagination="465 - 505" refId="B18" refString="Castanheira, PdS, Baptista, RLC, Pizzeti, DdP, Teixeira, RA, 2019. Contributions to the taxonomy of the long-jawed orb-weaving spider genus Tetragnatha (Araneae, Tetragnathidae) in the Neotropical region, with comments on the morphology of the chelicerae. Zoosystematics and Evolution 95 (2): 465 - 505, DOI: https://doi.org/10.3897/zse.95.36762" title="Contributions to the taxonomy of the long-jawed orb-weaving spider genus Tetragnatha (Araneae, Tetragnathidae) in the Neotropical region, with comments on the morphology of the chelicerae." url="https://doi.org/10.3897/zse.95.36762" volume="95" year="2019">Castanheira et al. (2019)</bibRefCitation>
, this specimen is clearly
<taxonomicName id="51EEB488C6F3AAF83ABA9448C0A92CA9" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="70DE82A5C2460F61A21A6AEB9C2496FC" italics="true" pageId="0" pageNumber="175">T. nitens</emphasis>
</taxonomicName>
, as shown for example by the following diagnostic cheliceral characters: Gl distalward followed by smaller L2 and L3, AXl reduced and point distalward, Gu elongated and not contiguous to U2 and robust basal cusp placed at the lower side of fang (Fig.
<figureCitation id="E230BEE3E69E03AA96B344D238649BA0" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tetragnatha nitens under T. labialis, female lectotype (MNHN 4209). A. Left chelicera, upper view; B. Left chelicera, inner view; C. Left chelicera, lower view; D. Original label." httpUri="https://binary.pensoft.net/fig/760565" pageId="0" pageNumber="175">13A-C</figureCitation>
). Therefore, we propose
<taxonomicName id="59FD5841682FD6C00430F09FA771FBB0" authorityName="Nicolet" authorityYear="1849" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="3C7DDD127AA24998FFE8BC03CFFFFF72" italics="true" pageId="0" pageNumber="175">Tetragnatha labialis</emphasis>
</taxonomicName>
<bibRefCitation id="F28F9A353E8DCC4F57EC6E9F7D2BA323" author="Gay, C" journalOrPublisher="Zoologia" pageId="0" pageNumber="175" pagination="319 - 543" refId="B52" refString="Gay, C, 1849. Aracnidos. In: Historia fisica y politica de Chile. Paris, Ed. Maude y Renou. Zoologia 3: 319 - 543" title="Aracnidos." volume="3" volumeTitle="Historia fisica y politica de Chile. Paris, Ed. Maude y Renou." year="1849">Nicolet 1849</bibRefCitation>
=
<taxonomicName id="F84265F17550D4FA661A0499DB1D57DB" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="7590E7E97755A76424E7BEDD5153ABB7" italics="true" pageId="0" pageNumber="175">Tetragnatha nitens</emphasis>
</taxonomicName>
(Audouin, 1826) syn. nov. Additionally, the male and female specimens from Nova Granada (probably Colombia) named
<taxonomicName id="B4685FBEBBBD2AB2B071127F3CC20193" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="CB643908FFF4CFBE335D0C5DFD6AD375" italics="true" pageId="0" pageNumber="175">T. labialis</emphasis>
</taxonomicName>
by
<bibRefCitation id="A88D75F546558C2C47A3D138BA308035" author="Keyserling, E" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="175" pagination="799 - 856" refId="B37" refString="Keyserling, E, 1865. Beitraege zur Kenntniss der Orbitelae Latr. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 15: 799 - 856" title="Beitraege zur Kenntniss der Orbitelae Latr." volume="15" year="1865">Keyserling (1865)</bibRefCitation>
are misidentified. The original drawings of the chelicerae resemble
<taxonomicName id="ADE4C0C44658DB2105769BFA88A1CA92" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. argentinensis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="E7A5A18EC4EA6338A1ED371377E3E49A" italics="true" pageId="0" pageNumber="175">T. argentinensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="546194981C2088DC611DA42D5B76AC55" author="Keyserling, E" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="175" pagination="799 - 856" refId="B37" refString="Keyserling, E, 1865. Beitraege zur Kenntniss der Orbitelae Latr. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 15: 799 - 856" title="Beitraege zur Kenntniss der Orbitelae Latr." volume="15" year="1865">Keyserling 1865</bibRefCitation>
, fig. 11-13) or perhaps some related, undescribed species from Colombia. Furthermore, the original locality (Nova Granada) is too far north of the currently known
<taxonomicName id="2AFFB152983950505F282EA69E5E8559" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. argentinensis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="3518849BB2B91F245E8B9230A62689D9" italics="true" pageId="0" pageNumber="175">T. argentinensis</emphasis>
</taxonomicName>
distribution range (
<bibRefCitation id="517CBB5A1278391C9D16EBCD3A15D881" DOI="https://doi.org/10.1636/JoA-S-21-006" author="Cargnelutti, F" journalOrPublisher="Journal of Arachnology" pageId="0" pageNumber="175" pagination="67 - 80" refId="B13" refString="Cargnelutti, F, Bollatti, F, Izquierdo, MA, Castanheira, PdS, Baptista, RLC, Barrantes, G, Aisenberg, A, 2022. Together but not intertwined: differences in sexual behavior between two sympatric and synchronic spider species, including one new synonymy (Araneae: Tetragnathidae: Tetragnatha). Journal of Arachnology 50 (1): 67 - 80, DOI: https://doi.org/10.1636/JoA-S-21-006" title="Together but not intertwined: differences in sexual behavior between two sympatric and synchronic spider species, including one new synonymy (Araneae: Tetragnathidae: Tetragnatha)." url="https://doi.org/10.1636/JoA-S-21-006" volume="50" year="2022">Cargnelutti et al. 2022</bibRefCitation>
). The Chilean specimens of
<taxonomicName id="A4C86B09A1C3DA2B40D29C9D2B90752A" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="63BDD8DFAF8D6DC16B4F1386E0E6F823" italics="true" pageId="0" pageNumber="175">T. labialis</emphasis>
</taxonomicName>
cited by
<bibRefCitation id="D6E7FBE36EC3308D7B72C873D40ADEDA" author="Simon, E" journalOrPublisher="Band Arthropoden" pageId="0" pageNumber="175" pagination="1 - 47" refId="B66" refString="Simon, E, 1902. Arachnoideen, excl. Acariden und Gonyleptiden. In: Ergebnisse der Hamburger Magalhaensischen Sammelreise 1892/1893. Hamburg, L. Ed. Friederichsen &amp; Co. Band Arthropoden 6 (4): 1 - 47" title="Arachnoideen, excl. Acariden und Gonyleptiden. In: Ergebnisse der Hamburger Magalhaensischen Sammelreise 1892 / 1893. Hamburg, L. Ed. Friederichsen &amp; Co." volume="6" year="1902">Simon (1902</bibRefCitation>
,
<bibRefCitation id="4BA4322212BF7793FC465F621DD03CCF" author="Simon, E" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="175" pagination="83 - 114" refId="B67" refString="Simon, E, 1904. Etude sur les arachnides du Chili recueillis en 1900, 1901 et 1902, par MM. C. Porter, Dr Delfin, Barcey Wilson et Edwards. Annales de la Societe Entomologique de Belgique 48: 83 - 114" title="Etude sur les arachnides du Chili recueillis en 1900, 1901 et 1902, par MM. C. Porter, Dr Delfin, Barcey Wilson et Edwards." volume="48" year="1904">1904</bibRefCitation>
) from Valdivia and Punta Arenas, respectively, were not found at MNHN. Only a male from
<taxonomicName id="A3B89C074284D2F4502045D9E78B6D43" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. americana" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="A0E2DC7045F05909B5B06565F5ADAA60" italics="true" pageId="0" pageNumber="175">T. americana</emphasis>
</taxonomicName>
from Punta Arenas was located as we explained above (Fig.
<figureCitation id="E263276E25E04036E9CD17BE86B6707D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tetragnatha nitens under T. americana, male. A. Left chelicera, inner view; B. Chelicerae, slanted lower view." httpUri="https://binary.pensoft.net/fig/760564" pageId="0" pageNumber="175">12</figureCitation>
).
</paragraph>
<caption id="D1C2D54D202D3CB68462865E32290878" httpUri="https://binary.pensoft.net/fig/760565" pageId="0" pageNumber="175" start="Figure 13" startId="F13">
<paragraph id="8301903E97837C70B1BC8011515EF806" pageId="0" pageNumber="175">
<emphasis id="1E143F4559DFFCB70EAF6F059C7DFDFF" bold="true" pageId="0" pageNumber="175">Figure 13.</emphasis>
<taxonomicName id="F1810E40DF0A0B8E486A64D5E0372B32" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetragnatha nitens" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="D81C1ECDB73A2635A4CF72B3DFCDA063" italics="true" pageId="0" pageNumber="175">Tetragnatha nitens</emphasis>
</taxonomicName>
under
<taxonomicName id="5E3F703BF9A8E5793EE9E74F52A50934" class="Arachnida" family="Tetragnathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. labialis" order="Araneae" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="labialis">
<emphasis id="A7AB567E3E950B8B18A4B588213B8702" italics="true" pageId="0" pageNumber="175">T. labialis</emphasis>
</taxonomicName>
, female lectotype (MNHN 4209).
<emphasis id="986C418AB939AA6F8943D088ADBFF1E9" bold="true" pageId="0" pageNumber="175">A.</emphasis>
Left chelicera, upper view;
<emphasis id="8B1B738EC24AB435A8653F7E85E49C24" bold="true" pageId="0" pageNumber="175">B.</emphasis>
Left chelicera, inner view;
<emphasis id="1F2A40D97AE869A8B007110844A2D1B0" bold="true" pageId="0" pageNumber="175">C.</emphasis>
Left chelicera, lower view;
<emphasis id="66170364E180D3C44C68E5C97AAA2FA5" bold="true" pageId="0" pageNumber="175">D.</emphasis>
Original label.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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