treatments-xml/data/05/59/87/055987E28B30737FFF7DD645FD4689D6.xml

<document id="3503B35E627CB1B50ACF73FA99C41307" ID-DOI="10.11646/zootaxa.5375.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10184224" ID-ZooBank="F3D0E008-556C-4FAD-BF51-4F1A714325DA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700642529072" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2023" docId="055987E28B30737FFF7DD645FD4689D6" docLanguage="en" docName="zootaxa.5375.2.1.pdf" docOrigin="Zootaxa 5375 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.2.1/52321" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nesamblyopina Sokolov 2023, subtribe nova" docType="treatment" docVersion="4" lastPageNumber="164" masterDocId="F960FF9A8B327372FFEAD113FFB08B1B" masterDocTitle="Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe" masterLastPageNumber="192" masterPageNumber="151" pageNumber="153" updateTime="1700746931339" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C3AD431A50D3E9074BF19E7815D20AE9">Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe</mods:title>
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<mods:namePart id="B1DADC4D920B847538552F7B50146C93">Sokolov, Igor M.</mods:namePart>
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<mods:date id="2EBEFDD1CE7741FDC29431CFA689F26B">2023</mods:date>
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<treatment id="055987E28B30737FFF7DD645FD4689D6" LSID="urn:lsid:plazi:treatment:055987E28B30737FFF7DD645FD4689D6" httpUri="http://treatment.plazi.org/id/055987E28B30737FFF7DD645FD4689D6" lastPageId="13" lastPageNumber="164" pageId="2" pageNumber="153">
<subSubSection id="C5EA657F8B307370FF7DD645FDA38C6A" box="[151,531,1878,1905]" pageId="2" pageNumber="153" type="nomenclature">
<paragraph id="8D4F36F48B307370FF7DD645FDA38C6A" blockId="2.[151,531,1878,1905]" box="[151,531,1878,1905]" pageId="2" pageNumber="153">
<heading id="D60781988B307370FF7DD645FDA38C6A" bold="true" box="[151,531,1878,1905]" fontSize="11" level="1" pageId="2" pageNumber="153" reason="1">
<emphasis id="BF84EAE68B307370FF7DD645FDA38C6A" bold="true" box="[151,531,1878,1905]" pageId="2" pageNumber="153">
<taxonomicName id="4AF04D778B307370FF7DD645FEEB8C6B" authority="Sokolov, 2023" authorityName="Sokolov" authorityYear="2023" box="[151,347,1878,1904]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="subTribe" status="subtrib. nov." subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
, 
<taxonomicNameLabel id="A4B7579D8B307370FE82D645FDA38C6A" box="[360,531,1878,1905]" pageId="2" pageNumber="153" rank="subTribe">subtribe nova</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C5EA657F8B307376FF7DD68FFCB28BAA" lastPageId="4" lastPageNumber="155" pageId="2" pageNumber="153" type="materials_examined">
<paragraph id="8D4F36F48B307370FF7DD68FFD418CA8" blockId="2.[151,753,1947,1972]" box="[151,753,1947,1972]" pageId="2" pageNumber="153">
<typeStatus id="524B88568B307370FF7DD68FFF798CAF" box="[151,201,1948,1972]" pageId="2" pageNumber="153">Type</typeStatus>
genus: 
<taxonomicName id="4AF04D778B307370FEF2D688FD848CA8" authority="Jeannel, 1937" authorityName="Jeannel" authorityYear="1937" box="[280,564,1947,1971]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B307370FEF2D688FE128CA8" bold="true" box="[280,418,1947,1971]" italics="true" pageId="2" pageNumber="153">Nesamblyops</emphasis>
<bibRefCitation id="E9614B058B307370FE43D688FD848CA8" author="Jeannel, R." box="[425,564,1947,1971]" pageId="2" pageNumber="153" pagination="241 - 339" refId="ref26574" refString="Jeannel, R. (1937) Les Bembidiides endoges (Col. Carabidae). Revue francaise d'Entomologie, 3, 241 - 339." type="journal article" year="1937">Jeannel, 1937</bibRefCitation>
</taxonomicName>
(here designated).
</paragraph>
<caption id="D98F667C8B317371FF7CD0FDF90E8913" ID-Table-UUID="D98F667C8B317371FF7CD0FDF90E8913" box="[150,1726,494,520]" httpUri="http://table.plazi.org/id/D98F667C8B317371FF7CD0FDF90E8913" pageId="3" pageNumber="154" startId="3.[150,238,494,520]" targetBox="[160,2030,531,1596]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="8D4F36F48B317371FF7CD0FDF90E8913" blockId="3.[150,1726,494,520]" box="[150,1726,494,520]" pageId="3" pageNumber="154">
<emphasis id="BF84EAE68B317371FF7CD0FDFEB98913" bold="true" box="[150,265,494,520]" pageId="3" pageNumber="154">TABLE 1.</emphasis>
Variation of character states among the representatives of the 
<taxonomicName id="4AF04D778B317371FC4AD0FDFBBA8913" box="[928,1034,494,520]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="subFamily" subFamily="Trechinae">Trechinae</taxonomicName>
(character states of 
<taxonomicName id="4AF04D778B317371FB0BD0FDFA388913" authorityName="Sokolov" authorityYear="2023" box="[1249,1416,494,520]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
are marked with blue color).
</paragraph>
</caption>
<paragraph id="8D4F36F48B317371FD0CD300FE258D27" pageId="3" pageNumber="154">
<table id="FFF0C4548B318C8DFF4AD300F85E8D27" box="[160,2030,531,1596]" gridcols="12" gridrows="34" pageId="3" pageNumber="154">
<tr id="33C034B68B318C8DFF4AD300F85E8931" box="[160,2030,531,554]" gridrow="0" pageId="3" pageNumber="154" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<th id="70115DCA8B318C8DFD0CD300FCD28931" box="[742,866,531,554]" gridcol="3" gridrow="0" pageId="3" pageNumber="154">Stipital *</th>
<th id="70115DCA8B318C8DFC9ED300FC408931" box="[884,1008,531,554]" gridcol="4" gridrow="0" pageId="3" pageNumber="154">Setation on</th>
<th id="70115DCA8B318C8DFBE8D300FBED8931" box="[1026,1117,531,554]" gridcol="5" gridrow="0" pageId="3" pageNumber="154">Mesoepi-</th>
<th id="70115DCA8B318C8DFB9AD300FB588931" box="[1136,1256,531,554]" gridcol="6" gridrow="0" pageId="3" pageNumber="154">Metasternal</th>
<th id="70115DCA8B318C8DFB17D300FA268931" box="[1277,1430,531,554]" gridcol="7" gridrow="0" pageId="3" pageNumber="154">Metaventrite**,</th>
<th id="70115DCA8B318C8DFA4CD300F9A28931" box="[1446,1554,531,554]" gridcol="8" gridrow="0" pageId="3" pageNumber="154">Umbilical</th>
<th id="70115DCA8B318C8DF9CDD300F95C8931" box="[1575,1772,531,554]" gridcol="9" gridrow="0" pageId="3" pageNumber="154">Distance ***</th>
<th id="70115DCA8B318C8DF8E8D300F8DD8931" box="[1794,1901,531,554]" gridcol="10" gridrow="0" pageId="3" pageNumber="154">Posterior</th>
<th id="70115DCA8B318C8DF868D300F85E8931" box="[1922,2030,531,554]" gridcol="11" gridrow="0" pageId="3" pageNumber="154">Ovipositor,</th>
</tr>
<tr id="33C034B68B318C8DFF4AD320F85E8951" box="[160,2030,563,586]" gridrow="1" pageId="3" pageNumber="154" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="70115DCA8B318C8DFD0CD320FCD28951" box="[742,866,563,586]" gridcol="3" gridrow="1" pageId="3" pageNumber="154">chaetotaxy,</td>
<td id="70115DCA8B318C8DFC9ED320FC408951" box="[884,1008,563,586]" gridcol="4" gridrow="1" pageId="3" pageNumber="154">basal margin</td>
<td id="70115DCA8B318C8DFBE8D320FBED8951" box="[1026,1117,563,586]" gridcol="5" gridrow="1" pageId="3" pageNumber="154">sternum,</td>
<td id="70115DCA8B318C8DFB9AD320FB588951" box="[1136,1256,563,586]" gridcol="6" gridrow="1" pageId="3" pageNumber="154">process,</td>
<td id="70115DCA8B318C8DFB17D320FA268951" box="[1277,1430,563,586]" gridcol="7" gridrow="1" pageId="3" pageNumber="154">length</td>
<td id="70115DCA8B318C8DFA4CD320F9A28951" box="[1446,1554,563,586]" gridcol="8" gridrow="1" pageId="3" pageNumber="154">setal pores,</td>
<td id="70115DCA8B318C8DF9CDD320F95C8951" box="[1575,1772,563,586]" gridcol="9" gridrow="1" pageId="3" pageNumber="154">from elytral</td>
<td id="70115DCA8B318C8DF8E8D320F8DD8951" box="[1794,1901,563,586]" gridcol="10" gridrow="1" pageId="3" pageNumber="154">setal row,</td>
<td id="70115DCA8B318C8DF868D320F85E8951" box="[1922,2030,563,586]" gridcol="11" gridrow="1" pageId="3" pageNumber="154">ensiform</td>
</tr>
<tr id="33C034B68B318C8DFF4AD340F85E8977" box="[160,2030,595,620]" gridrow="2" pageId="3" pageNumber="154" rowspan-0="1" rowspan-1="1" rowspan-7="1">
<td id="70115DCA8B318C8DFDA4D340FD7D8977" box="[590,717,595,620]" gridcol="2" gridrow="2" pageId="3" pageNumber="154">Tribe/</td>
<td id="70115DCA8B318C8DFD0CD340FCD28977" box="[742,866,595,620]" gridcol="3" gridrow="2" pageId="3" pageNumber="154">longest setae</td>
<td id="70115DCA8B318C8DFC9ED340FC408977" box="[884,1008,595,620]" gridcol="4" gridrow="2" pageId="3" pageNumber="154">of pronotum</td>
<td id="70115DCA8B318C8DFBE8D340FBED8977" box="[1026,1117,595,620]" gridcol="5" gridrow="2" pageId="3" pageNumber="154">suture</td>
<td id="70115DCA8B318C8DFB9AD340FB588977" box="[1136,1256,595,620]" gridcol="6" gridrow="2" pageId="3" pageNumber="154">margination</td>
<td id="70115DCA8B318C8DFA4CD340F9A28977" box="[1446,1554,595,620]" gridcol="8" gridrow="2" pageId="3" pageNumber="154">number</td>
<td id="70115DCA8B318C8DF9CDD340F95C8977" box="[1575,1772,595,620]" gridcol="9" gridrow="2" pageId="3" pageNumber="154">margin to 5 &amp; 6USP</td>
<td id="70115DCA8B318C8DF8E8D340F8DD8977" box="[1794,1901,595,620]" gridcol="10" gridrow="2" pageId="3" pageNumber="154">psr, n setae</td>
<td id="70115DCA8B318C8DF868D340F85E8977" box="[1922,2030,595,620]" gridcol="11" gridrow="2" pageId="3" pageNumber="154">setae, n</td>
</tr>
<tr id="33C034B68B318C8DFF4AD361F85E8992" box="[160,2030,626,649]" gridrow="3" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD361FF0F8992" box="[160,191,626,649]" gridcol="0" gridrow="3" pageId="3" pageNumber="154">n/n</th>
<td id="70115DCA8B318C8DFF24D361FD838992" box="[206,563,626,649]" gridcol="1" gridrow="3" pageId="3" pageNumber="154">Species</td>
<td id="70115DCA8B318C8DFDA4D361FD7D8992" box="[590,717,626,649]" gridcol="2" gridrow="3" pageId="3" pageNumber="154">Subtribe</td>
<td id="70115DCA8B318C8DFD0CD361FCD28992" box="[742,866,626,649]" gridcol="3" gridrow="3" pageId="3" pageNumber="154">1</td>
<td id="70115DCA8B318C8DFC9ED361FC408992" box="[884,1008,626,649]" gridcol="4" gridrow="3" pageId="3" pageNumber="154">2</td>
<td id="70115DCA8B318C8DFBE8D361FBED8992" box="[1026,1117,626,649]" gridcol="5" gridrow="3" pageId="3" pageNumber="154">3</td>
<td id="70115DCA8B318C8DFB9AD361FB588992" box="[1136,1256,626,649]" gridcol="6" gridrow="3" pageId="3" pageNumber="154">4</td>
<td id="70115DCA8B318C8DFB17D361FA268992" box="[1277,1430,626,649]" gridcol="7" gridrow="3" pageId="3" pageNumber="154">5</td>
<td id="70115DCA8B318C8DFA4CD361F9A28992" box="[1446,1554,626,649]" gridcol="8" gridrow="3" pageId="3" pageNumber="154">6</td>
<td id="70115DCA8B318C8DF9CDD361F95C8992" box="[1575,1772,626,649]" gridcol="9" gridrow="3" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF8E8D361F8DD8992" box="[1794,1901,626,649]" gridcol="10" gridrow="3" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF868D361F85E8992" box="[1922,2030,626,649]" gridcol="11" gridrow="3" pageId="3" pageNumber="154">10</td>
</tr>
<tr id="33C034B68B318C8DFF4AD381F85E89B2" box="[160,2030,658,681]" gridrow="4" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD381FF0F89B2" box="[160,191,658,681]" gridcol="0" gridrow="4" pageId="3" pageNumber="154">1</th>
<td id="70115DCA8B318C8DFF24D381FD8389B2" box="[206,563,658,681]" gridcol="1" gridrow="4" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D381FDB789B2" authority="(Hayward)" baseAuthorityName="Hayward" baseAuthorityYear="1897" box="[206,519,658,681]" class="Insecta" family="Carabidae" genus="Phrypeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="rickseckeri">
<emphasis id="BF84EAE68B317371FF24D381FE2889B2" box="[206,408,658,681]" italics="true" pageId="3" pageNumber="154">Phrypeus rickseckeri</emphasis>
(Hayward)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D381FD7D89B2" box="[590,717,658,681]" gridcol="2" gridrow="4" pageId="3" pageNumber="154">Sinozolini</td>
<td id="70115DCA8B318C8DFD0CD381FCD289B2" box="[742,866,658,681]" gridcol="3" gridrow="4" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED381FC4089B2" box="[884,1008,658,681]" gridcol="4" gridrow="4" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFBE8D381FBED89B2" box="[1026,1117,658,681]" gridcol="5" gridrow="4" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD381FB5889B2" box="[1136,1256,658,681]" gridcol="6" gridrow="4" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D381FA2689B2" box="[1277,1430,658,681]" gridcol="7" gridrow="4" pageId="3" pageNumber="154">&gt; Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD381F9A289B2" box="[1446,1554,658,681]" gridcol="8" gridrow="4" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD381F95C89B2" box="[1575,1772,658,681]" gridcol="9" gridrow="4" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D381F8DD89B2" box="[1794,1901,658,681]" gridcol="10" gridrow="4" pageId="3" pageNumber="154">7</td>
<td id="70115DCA8B318C8DF868D381F85E89B2" box="[1922,2030,658,681]" gridcol="11" gridrow="4" pageId="3" pageNumber="154">0</td>
</tr>
<tr id="33C034B68B318C8DFF4AD3A1F85E89D2" box="[160,2030,690,713]" gridrow="5" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD3A1FF0F89D2" box="[160,191,690,713]" gridcol="0" gridrow="5" pageId="3" pageNumber="154">2</th>
<td id="70115DCA8B318C8DFF24D3A1FD8389D2" box="[206,563,690,713]" gridcol="1" gridrow="5" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D3A1FE4389D2" authority="Solier" authorityName="Solier" authorityYear="1849" box="[206,499,690,713]" class="Insecta" family="Carabidae" genus="Merizodus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="BF84EAE68B317371FF24D3A1FE0589D2" box="[206,437,690,713]" italics="true" pageId="3" pageNumber="154">Merizodus angusticollis</emphasis>
Solier
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D3A1FD7D89D2" box="[590,717,690,713]" gridcol="2" gridrow="5" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D3A1FD3989D2" box="[590,649,690,713]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Zolini">Zolini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD3A1FCD289D2" box="[742,866,690,713]" gridcol="3" gridrow="5" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED3A1FC4089D2" box="[884,1008,690,713]" gridcol="4" gridrow="5" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D3A1FBED89D2" box="[1026,1117,690,713]" gridcol="5" gridrow="5" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD3A1FB5889D2" box="[1136,1256,690,713]" gridcol="6" gridrow="5" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D3A1FA2689D2" box="[1277,1430,690,713]" gridcol="7" gridrow="5" pageId="3" pageNumber="154">&gt; Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD3A1F9A289D2" box="[1446,1554,690,713]" gridcol="8" gridrow="5" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF9CDD3A1F95C89D2" box="[1575,1772,690,713]" gridcol="9" gridrow="5" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D3A1F8DD89D2" box="[1794,1901,690,713]" gridcol="10" gridrow="5" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF868D3A1F85E89D2" box="[1922,2030,690,713]" gridcol="11" gridrow="5" pageId="3" pageNumber="154">0</td>
</tr>
<tr id="33C034B68B318C8DFF4AD3C1F85E89F2" box="[160,2030,722,745]" gridrow="6" pageId="3" pageNumber="154" rowspan-2="1">
<th id="70115DCA8B318C8DFF4AD3C1FF0F89F2" box="[160,191,722,745]" gridcol="0" gridrow="6" pageId="3" pageNumber="154">3</th>
<td id="70115DCA8B318C8DFF24D3C1FD8389F2" box="[206,563,722,745]" gridcol="1" gridrow="6" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D3C1FE1D89F2" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1830" box="[206,429,722,745]" class="Insecta" family="Carabidae" genus="Merizodus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="soledadinus">
<emphasis id="BF84EAE68B317371FF24D3C1FE1D89F2" box="[206,429,722,745]" italics="true" pageId="3" pageNumber="154">Merizodus soledadinus</emphasis>
</taxonomicName>
(Guerin-
</td>
<td id="70115DCA8B318C8DFD0CD3C1FCD289F2" box="[742,866,722,745]" gridcol="3" gridrow="6" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED3C1FC4089F2" box="[884,1008,722,745]" gridcol="4" gridrow="6" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D3C1FBED89F2" box="[1026,1117,722,745]" gridcol="5" gridrow="6" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD3C1FB5889F2" box="[1136,1256,722,745]" gridcol="6" gridrow="6" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D3C1FA2689F2" box="[1277,1430,722,745]" gridcol="7" gridrow="6" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD3C1F9A289F2" box="[1446,1554,722,745]" gridcol="8" gridrow="6" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF9CDD3C1F95C89F2" box="[1575,1772,722,745]" gridcol="9" gridrow="6" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D3C1F8DD89F2" box="[1794,1901,722,745]" gridcol="10" gridrow="6" pageId="3" pageNumber="154">7</td>
<td id="70115DCA8B318C8DF868D3C1F85E89F2" box="[1922,2030,722,745]" gridcol="11" gridrow="6" pageId="3" pageNumber="154">0</td>
</tr>
<tr id="33C034B68B318C8DFF4AD3FCF85E881D" box="[160,2030,751,774]" gridrow="7" pageId="3" pageNumber="154" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td id="70115DCA8B318C8DFF24D3FCFD83881D" box="[206,563,751,774]" gridcol="1" gridrow="7" pageId="3" pageNumber="154">Meneville)</td>
</tr>
<tr id="33C034B68B318C8DFF4AD21CF85E883D" box="[160,2030,783,806]" gridrow="8" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD21CFF0F883D" box="[160,191,783,806]" gridcol="0" gridrow="8" pageId="3" pageNumber="154">4</th>
<td id="70115DCA8B318C8DFF24D21CFD83883D" box="[206,563,783,806]" gridcol="1" gridrow="8" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D21CFD98883D" authority="(Mannerheim)" baseAuthorityName="Mannerheim" baseAuthorityYear="1852" box="[206,552,783,806]" class="Insecta" family="Carabidae" genus="Amerizus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="oblongulus">
<emphasis id="BF84EAE68B317371FF24D21CFE29883D" box="[206,409,783,806]" italics="true" pageId="3" pageNumber="154">Amerizus oblongulus</emphasis>
(Mannerheim)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D21CFD7D883D" box="[590,717,783,806]" gridcol="2" gridrow="8" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D21CFD0A883D" box="[590,698,783,806]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Bembidiini">Bembidiini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD21CFCD2883D" box="[742,866,783,806]" gridcol="3" gridrow="8" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED21CFC40883D" box="[884,1008,783,806]" gridcol="4" gridrow="8" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D21CFBED883D" box="[1026,1117,783,806]" gridcol="5" gridrow="8" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD21CFB58883D" box="[1136,1256,783,806]" gridcol="6" gridrow="8" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D21CFA26883D" box="[1277,1430,783,806]" gridcol="7" gridrow="8" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD21CF9A2883D" box="[1446,1554,783,806]" gridcol="8" gridrow="8" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD21CF95C883D" box="[1575,1772,783,806]" gridcol="9" gridrow="8" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D21CF8DD883D" box="[1794,1901,783,806]" gridcol="10" gridrow="8" pageId="3" pageNumber="154">4</td>
<td id="70115DCA8B318C8DF868D21CF85E883D" box="[1922,2030,783,806]" gridcol="11" gridrow="8" pageId="3" pageNumber="154">n/a</td>
</tr>
<tr id="33C034B68B318C8DFF4AD23CF85E885D" box="[160,2030,815,838]" gridrow="9" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD23CFF0F885D" box="[160,191,815,838]" gridcol="0" gridrow="9" pageId="3" pageNumber="154">5</th>
<td id="70115DCA8B318C8DFF24D23CFD83885D" box="[206,563,815,838]" gridcol="1" gridrow="9" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D23CFE64885D" authority="(Casey)" baseAuthorityName="Casey" baseAuthorityYear="1918" box="[206,468,815,838]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="osculans">
<emphasis id="BF84EAE68B317371FF24D23CFE33885D" box="[206,387,815,838]" italics="true" pageId="3" pageNumber="154">Lionepha osculans</emphasis>
(Casey)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D23CFD7D885D" box="[590,717,815,838]" gridcol="2" gridrow="9" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D23CFD0A885D" box="[590,698,815,838]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Bembidiini">Bembidiini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD23CFCD2885D" box="[742,866,815,838]" gridcol="3" gridrow="9" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED23CFC40885D" box="[884,1008,815,838]" gridcol="4" gridrow="9" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D23CFBED885D" box="[1026,1117,815,838]" gridcol="5" gridrow="9" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD23CFB58885D" box="[1136,1256,815,838]" gridcol="6" gridrow="9" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D23CFA26885D" box="[1277,1430,815,838]" gridcol="7" gridrow="9" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD23CF9A2885D" box="[1446,1554,815,838]" gridcol="8" gridrow="9" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF9CDD23CF95C885D" box="[1575,1772,815,838]" gridcol="9" gridrow="9" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D23CF8DD885D" box="[1794,1901,815,838]" gridcol="10" gridrow="9" pageId="3" pageNumber="154">2</td>
<td id="70115DCA8B318C8DF868D23CF85E885D" box="[1922,2030,815,838]" gridcol="11" gridrow="9" pageId="3" pageNumber="154">0</td>
</tr>
<tr id="33C034B68B318C8DFF4AD25DF85E887E" box="[160,2030,846,869]" gridrow="10" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD25DFF0F887E" box="[160,191,846,869]" gridcol="0" gridrow="10" pageId="3" pageNumber="154">6</th>
<td id="70115DCA8B318C8DFF24D25DFD83887E" box="[206,563,846,869]" gridcol="1" gridrow="10" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D25DFE41887E" authority="(Schaum)" baseAuthorityName="Schaum" baseAuthorityYear="1859" box="[206,497,846,869]" class="Insecta" family="Carabidae" genus="Xystosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="BF84EAE68B317371FF24D25DFE3E887E" box="[206,398,846,869]" italics="true" pageId="3" pageNumber="154">Xystosomus inflatus</emphasis>
(Schaum)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D25DFD7D887E" box="[590,717,846,869]" gridcol="2" gridrow="10" pageId="3" pageNumber="154">Xystosomina</td>
<td id="70115DCA8B318C8DFD0CD25DFCD2887E" box="[742,866,846,869]" gridcol="3" gridrow="10" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED25DFC40887E" box="[884,1008,846,869]" gridcol="4" gridrow="10" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D25DFBED887E" box="[1026,1117,846,869]" gridcol="5" gridrow="10" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD25DFB58887E" box="[1136,1256,846,869]" gridcol="6" gridrow="10" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D25DFA26887E" box="[1277,1430,846,869]" gridcol="7" gridrow="10" pageId="3" pageNumber="154">= ¼ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD25DF9A2887E" box="[1446,1554,846,869]" gridcol="8" gridrow="10" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD25DF95C887E" box="[1575,1772,846,869]" gridcol="9" gridrow="10" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D25DF8DD887E" box="[1794,1901,846,869]" gridcol="10" gridrow="10" pageId="3" pageNumber="154">2</td>
<td id="70115DCA8B318C8DF868D25DF85E887E" box="[1922,2030,846,869]" gridcol="11" gridrow="10" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD27DF85E889E" box="[160,2030,878,901]" gridrow="11" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD27DFF0F889E" box="[160,191,878,901]" gridcol="0" gridrow="11" pageId="3" pageNumber="154">7</th>
<td id="70115DCA8B318C8DFF24D27DFD83889E" box="[206,563,878,901]" gridcol="1" gridrow="11" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D27DFE48889E" authority="(Schaum)" baseAuthorityName="Schaum" baseAuthorityYear="1863" box="[206,504,878,901]" class="Insecta" family="Carabidae" genus="Xystosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="turgidus">
<emphasis id="BF84EAE68B317371FF24D27DFE25889E" box="[206,405,878,901]" italics="true" pageId="3" pageNumber="154">Xystosomus turgidus</emphasis>
(Schaum)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D27DFD7D889E" box="[590,717,878,901]" gridcol="2" gridrow="11" pageId="3" pageNumber="154">Xystosomina</td>
<td id="70115DCA8B318C8DFD0CD27DFCD2889E" box="[742,866,878,901]" gridcol="3" gridrow="11" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED27DFC40889E" box="[884,1008,878,901]" gridcol="4" gridrow="11" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D27DFBED889E" box="[1026,1117,878,901]" gridcol="5" gridrow="11" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD27DFB58889E" box="[1136,1256,878,901]" gridcol="6" gridrow="11" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D27DFA26889E" box="[1277,1430,878,901]" gridcol="7" gridrow="11" pageId="3" pageNumber="154">= ¼ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD27DF9A2889E" box="[1446,1554,878,901]" gridcol="8" gridrow="11" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD27DF95C889E" box="[1575,1772,878,901]" gridcol="9" gridrow="11" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D27DF8DD889E" box="[1794,1901,878,901]" gridcol="10" gridrow="11" pageId="3" pageNumber="154">4</td>
<td id="70115DCA8B318C8DF868D27DF85E889E" box="[1922,2030,878,901]" gridcol="11" gridrow="11" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD29DF85E88BE" box="[160,2030,910,933]" gridrow="12" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD29DFF0F88BE" box="[160,191,910,933]" gridcol="0" gridrow="12" pageId="3" pageNumber="154">8</th>
<td id="70115DCA8B318C8DFF24D29DFD8388BE" box="[206,563,910,933]" gridcol="1" gridrow="12" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D29DFE6188BE" authority="(Erwin)" box="[206,465,910,933]" class="Insecta" family="Carabidae" genus="Xystosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="tholus">
<emphasis id="BF84EAE68B317371FF24D29DFE3188BE" box="[206,385,910,933]" italics="true" pageId="3" pageNumber="154">Xystosomus tholus</emphasis>
(Erwin)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D29DFD7D88BE" box="[590,717,910,933]" gridcol="2" gridrow="12" pageId="3" pageNumber="154">Xystosomina</td>
<td id="70115DCA8B318C8DFD0CD29DFCD288BE" box="[742,866,910,933]" gridcol="3" gridrow="12" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED29DFC4088BE" box="[884,1008,910,933]" gridcol="4" gridrow="12" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D29DFBED88BE" box="[1026,1117,910,933]" gridcol="5" gridrow="12" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD29DFB5888BE" box="[1136,1256,910,933]" gridcol="6" gridrow="12" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D29DFA2688BE" box="[1277,1430,910,933]" gridcol="7" gridrow="12" pageId="3" pageNumber="154">= ¼ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD29DF9A288BE" box="[1446,1554,910,933]" gridcol="8" gridrow="12" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD29DF95C88BE" box="[1575,1772,910,933]" gridcol="9" gridrow="12" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D29DF8DD88BE" box="[1794,1901,910,933]" gridcol="10" gridrow="12" pageId="3" pageNumber="154">4</td>
<td id="70115DCA8B318C8DF868D29DF85E88BE" box="[1922,2030,910,933]" gridcol="11" gridrow="12" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD2BDF85E88DE" box="[160,2030,942,965]" gridrow="13" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD2BDFF0F88DE" box="[160,191,942,965]" gridcol="0" gridrow="13" pageId="3" pageNumber="154">9</th>
<td id="70115DCA8B318C8DFF24D2BDFD8388DE" box="[206,563,942,965]" gridcol="1" gridrow="13" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D2BDFE6E88DE" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1882" box="[206,478,942,965]" class="Insecta" family="Carabidae" genus="Erwiniana" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="sublaevis">
<emphasis id="BF84EAE68B317371FF24D2BDFE2388DE" box="[206,403,942,965]" italics="true" pageId="3" pageNumber="154">Erwiniana sublaevis</emphasis>
(Bates)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D2BDFD7D88DE" box="[590,717,942,965]" gridcol="2" gridrow="13" pageId="3" pageNumber="154">Xystosomina</td>
<td id="70115DCA8B318C8DFD0CD2BDFCD288DE" box="[742,866,942,965]" gridcol="3" gridrow="13" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED2BDFC4088DE" box="[884,1008,942,965]" gridcol="4" gridrow="13" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D2BDFBED88DE" box="[1026,1117,942,965]" gridcol="5" gridrow="13" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD2BDFB5888DE" box="[1136,1256,942,965]" gridcol="6" gridrow="13" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D2BDFA2688DE" box="[1277,1430,942,965]" gridcol="7" gridrow="13" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD2BDF9A288DE" box="[1446,1554,942,965]" gridcol="8" gridrow="13" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD2BDF95C88DE" box="[1575,1772,942,965]" gridcol="9" gridrow="13" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D2BDF8DD88DE" box="[1794,1901,942,965]" gridcol="10" gridrow="13" pageId="3" pageNumber="154">3</td>
<td id="70115DCA8B318C8DF868D2BDF85E88DE" box="[1922,2030,942,965]" gridcol="11" gridrow="13" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD2DDF85E88FE" box="[160,2030,974,997]" gridrow="14" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD2DDFF0F88FE" box="[160,191,974,997]" gridcol="0" gridrow="14" pageId="3" pageNumber="154">10</th>
<td id="70115DCA8B318C8DFF24D2DDFD8388FE" box="[206,563,974,997]" gridcol="1" gridrow="14" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D2DDFE5988FE" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1882" box="[206,489,974,997]" class="Insecta" family="Carabidae" genus="Erwiniana" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="sulcicostis">
<emphasis id="BF84EAE68B317371FF24D2DDFE2E88FE" box="[206,414,974,997]" italics="true" pageId="3" pageNumber="154">Erwiniana sulcicostis</emphasis>
(Bates)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D2DDFD7D88FE" box="[590,717,974,997]" gridcol="2" gridrow="14" pageId="3" pageNumber="154">Xystosomina</td>
<td id="70115DCA8B318C8DFD0CD2DDFCD288FE" box="[742,866,974,997]" gridcol="3" gridrow="14" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED2DDFC4088FE" box="[884,1008,974,997]" gridcol="4" gridrow="14" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D2DDFBED88FE" box="[1026,1117,974,997]" gridcol="5" gridrow="14" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD2DDFB5888FE" box="[1136,1256,974,997]" gridcol="6" gridrow="14" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D2DDFA2688FE" box="[1277,1430,974,997]" gridcol="7" gridrow="14" pageId="3" pageNumber="154">= ⅔ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD2DDF9A288FE" box="[1446,1554,974,997]" gridcol="8" gridrow="14" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD2DDF95C88FE" box="[1575,1772,974,997]" gridcol="9" gridrow="14" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D2DDF8DD88FE" box="[1794,1901,974,997]" gridcol="10" gridrow="14" pageId="3" pageNumber="154">2</td>
<td id="70115DCA8B318C8DF868D2DDF85E88FE" box="[1922,2030,974,997]" gridcol="11" gridrow="14" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD2FEF85E8F1F" box="[160,2030,1005,1028]" gridrow="15" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD2FEFF0F8F1F" box="[160,191,1005,1028]" gridcol="0" gridrow="15" pageId="3" pageNumber="154">11</th>
<td id="70115DCA8B318C8DFF24D2FEFD838F1F" box="[206,563,1005,1028]" gridcol="1" gridrow="15" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D2FEFE5B8F1F" authority="(Erwin)" baseAuthorityName="Erwin" baseAuthorityYear="1973" box="[206,491,1005,1028]" class="Insecta" family="Carabidae" genus="Geballusa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="microtreta">
<emphasis id="BF84EAE68B317371FF24D2FEFE2B8F1F" box="[206,411,1005,1028]" italics="true" pageId="3" pageNumber="154">Geballusa microtreta</emphasis>
(Erwin)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D2FEFD7D8F1F" box="[590,717,1005,1028]" gridcol="2" gridrow="15" pageId="3" pageNumber="154">Xystosomina</td>
<td id="70115DCA8B318C8DFD0CD2FEFCD28F1F" box="[742,866,1005,1028]" gridcol="3" gridrow="15" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED2FEFC408F1F" box="[884,1008,1005,1028]" gridcol="4" gridrow="15" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D2FEFBED8F1F" box="[1026,1117,1005,1028]" gridcol="5" gridrow="15" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD2FEFB588F1F" box="[1136,1256,1005,1028]" gridcol="6" gridrow="15" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D2FEFA268F1F" box="[1277,1430,1005,1028]" gridcol="7" gridrow="15" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD2FEF9A28F1F" box="[1446,1554,1005,1028]" gridcol="8" gridrow="15" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD2FEF95C8F1F" box="[1575,1772,1005,1028]" gridcol="9" gridrow="15" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D2FEF8DD8F1F" box="[1794,1901,1005,1028]" gridcol="10" gridrow="15" pageId="3" pageNumber="154">3</td>
<td id="70115DCA8B318C8DF868D2FEF85E8F1F" box="[1922,2030,1005,1028]" gridcol="11" gridrow="15" pageId="3" pageNumber="154">3 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD51EF85E8F3F" box="[160,2030,1037,1060]" gridrow="16" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD51EFF0F8F3F" box="[160,191,1037,1060]" gridcol="0" gridrow="16" pageId="3" pageNumber="154">12</th>
<td id="70115DCA8B318C8DFF24D51EFD838F3F" box="[206,563,1037,1060]" gridcol="1" gridrow="16" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D51EFE5F8F3F" authority="(Dejean)" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[206,495,1037,1060]" class="Insecta" family="Carabidae" genus="Gouleta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="cayennensis">
<emphasis id="BF84EAE68B317371FF24D51EFE268F3F" box="[206,406,1037,1060]" italics="true" pageId="3" pageNumber="154">Gouleta cayennensis</emphasis>
(Dejean)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D51EFD7D8F3F" box="[590,717,1037,1060]" gridcol="2" gridrow="16" pageId="3" pageNumber="154">Xystosomina</td>
<td id="70115DCA8B318C8DFD0CD51EFCD28F3F" box="[742,866,1037,1060]" gridcol="3" gridrow="16" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED51EFC408F3F" box="[884,1008,1037,1060]" gridcol="4" gridrow="16" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D51EFBED8F3F" box="[1026,1117,1037,1060]" gridcol="5" gridrow="16" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD51EFB588F3F" box="[1136,1256,1037,1060]" gridcol="6" gridrow="16" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D51EFA268F3F" box="[1277,1430,1037,1060]" gridcol="7" gridrow="16" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD51EF9A28F3F" box="[1446,1554,1037,1060]" gridcol="8" gridrow="16" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD51EF95C8F3F" box="[1575,1772,1037,1060]" gridcol="9" gridrow="16" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D51EF8DD8F3F" box="[1794,1901,1037,1060]" gridcol="10" gridrow="16" pageId="3" pageNumber="154">6</td>
<td id="70115DCA8B318C8DF868D51EF85E8F3F" box="[1922,2030,1037,1060]" gridcol="11" gridrow="16" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD53EF85E8F5F" box="[160,2030,1069,1092]" gridrow="17" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD53EFF0F8F5F" box="[160,191,1069,1092]" gridcol="0" gridrow="17" pageId="3" pageNumber="154">13</th>
<td id="70115DCA8B318C8DFF24D53EFD838F5F" box="[206,563,1069,1092]" gridcol="1" gridrow="17" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D53EFE548F5F" authority="(Say)" baseAuthorityName="Say" baseAuthorityYear="1823" box="[206,484,1069,1092]" class="Insecta" family="Carabidae" genus="Mioptachys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="flavicauda">
<emphasis id="BF84EAE68B317371FF24D53EFE198F5F" box="[206,425,1069,1092]" italics="true" pageId="3" pageNumber="154">Mioptachys flavicauda</emphasis>
(Say)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D53EFD7D8F5F" box="[590,717,1069,1092]" gridcol="2" gridrow="17" pageId="3" pageNumber="154">Xystosomina</td>
<td id="70115DCA8B318C8DFD0CD53EFCD28F5F" box="[742,866,1069,1092]" gridcol="3" gridrow="17" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED53EFC408F5F" box="[884,1008,1069,1092]" gridcol="4" gridrow="17" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D53EFBED8F5F" box="[1026,1117,1069,1092]" gridcol="5" gridrow="17" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD53EFB588F5F" box="[1136,1256,1069,1092]" gridcol="6" gridrow="17" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D53EFA268F5F" box="[1277,1430,1069,1092]" gridcol="7" gridrow="17" pageId="3" pageNumber="154">= ⅔ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD53EF9A28F5F" box="[1446,1554,1069,1092]" gridcol="8" gridrow="17" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD53EF95C8F5F" box="[1575,1772,1069,1092]" gridcol="9" gridrow="17" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D53EF8DD8F5F" box="[1794,1901,1069,1092]" gridcol="10" gridrow="17" pageId="3" pageNumber="154">4</td>
<td id="70115DCA8B318C8DF868D53EF85E8F5F" box="[1922,2030,1069,1092]" gridcol="11" gridrow="17" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD55EF85E8F7F" box="[160,2030,1101,1124]" gridrow="18" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD55EFF0F8F7F" box="[160,191,1101,1124]" gridcol="0" gridrow="18" pageId="3" pageNumber="154">14</th>
<td id="70115DCA8B318C8DFF24D55EFD838F7F" box="[206,563,1101,1124]" gridcol="1" gridrow="18" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D55EFE438F7F" authority="(Putzeys)" baseAuthorityName="Putzeys" baseAuthorityYear="1845" box="[206,499,1101,1124]" class="Insecta" family="Carabidae" genus="Pericompsus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="reichei">
<emphasis id="BF84EAE68B317371FF24D55EFE228F7F" box="[206,402,1101,1124]" italics="true" pageId="3" pageNumber="154">Pericompsus reichei</emphasis>
(Putzeys)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D55EFD7D8F7F" box="[590,717,1101,1124]" gridcol="2" gridrow="18" pageId="3" pageNumber="154">Tachyina</td>
<td id="70115DCA8B318C8DFD0CD55EFCD28F7F" box="[742,866,1101,1124]" gridcol="3" gridrow="18" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED55EFC408F7F" box="[884,1008,1101,1124]" gridcol="4" gridrow="18" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D55EFBED8F7F" box="[1026,1117,1101,1124]" gridcol="5" gridrow="18" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD55EFB588F7F" box="[1136,1256,1101,1124]" gridcol="6" gridrow="18" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D55EFA268F7F" box="[1277,1430,1101,1124]" gridcol="7" gridrow="18" pageId="3" pageNumber="154">&gt; Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD55EF9A28F7F" box="[1446,1554,1101,1124]" gridcol="8" gridrow="18" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD55EF95C8F7F" box="[1575,1772,1101,1124]" gridcol="9" gridrow="18" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D55EF8DD8F7F" box="[1794,1901,1101,1124]" gridcol="10" gridrow="18" pageId="3" pageNumber="154">10</td>
<td id="70115DCA8B318C8DF868D55EF85E8F7F" box="[1922,2030,1101,1124]" gridcol="11" gridrow="18" pageId="3" pageNumber="154">1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD57FF85E8F98" box="[160,2030,1132,1155]" gridrow="19" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD57FFF0F8F98" box="[160,191,1132,1155]" gridcol="0" gridrow="19" pageId="3" pageNumber="154">15</th>
<td id="70115DCA8B318C8DFF24D57FFD838F98" box="[206,563,1132,1155]" gridcol="1" gridrow="19" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D57FFDBC8F98" authority="(Dejean)" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[206,524,1132,1155]" class="Insecta" family="Carabidae" genus="Elaphropus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="ferrugineus">
<emphasis id="BF84EAE68B317371FF24D57FFE028F98" box="[206,434,1132,1155]" italics="true" pageId="3" pageNumber="154">Elaphropus ferrugineus</emphasis>
(Dejean)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D57FFD7D8F98" box="[590,717,1132,1155]" gridcol="2" gridrow="19" pageId="3" pageNumber="154">Tachyina</td>
<td id="70115DCA8B318C8DFD0CD57FFCD28F98" box="[742,866,1132,1155]" gridcol="3" gridrow="19" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED57FFC408F98" box="[884,1008,1132,1155]" gridcol="4" gridrow="19" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D57FFBED8F98" box="[1026,1117,1132,1155]" gridcol="5" gridrow="19" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD57FFB588F98" box="[1136,1256,1132,1155]" gridcol="6" gridrow="19" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D57FFA268F98" box="[1277,1430,1132,1155]" gridcol="7" gridrow="19" pageId="3" pageNumber="154">= ½ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD57FF9A28F98" box="[1446,1554,1132,1155]" gridcol="8" gridrow="19" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD57FF95C8F98" box="[1575,1772,1132,1155]" gridcol="9" gridrow="19" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D57FF8DD8F98" box="[1794,1901,1132,1155]" gridcol="10" gridrow="19" pageId="3" pageNumber="154">10</td>
<td id="70115DCA8B318C8DF868D57FF85E8F98" box="[1922,2030,1132,1155]" gridcol="11" gridrow="19" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD59FF85E8FB8" box="[160,2030,1164,1187]" gridrow="20" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD59FFF0F8FB8" box="[160,191,1164,1187]" gridcol="0" gridrow="20" pageId="3" pageNumber="154">16</th>
<td id="70115DCA8B318C8DFF24D59FFD838FB8" box="[206,563,1164,1187]" gridcol="1" gridrow="20" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D59FFE548FB8" authority="LeConte" authorityName="LeConte" authorityYear="1848" box="[206,484,1164,1187]" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="scitulus">
<emphasis id="BF84EAE68B317371FF24D59FFE3B8FB8" box="[206,395,1164,1187]" italics="true" pageId="3" pageNumber="154">Paratachys scitulus</emphasis>
LeConte
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D59FFD7D8FB8" box="[590,717,1164,1187]" gridcol="2" gridrow="20" pageId="3" pageNumber="154">Tachyina</td>
<td id="70115DCA8B318C8DFD0CD59FFCD28FB8" box="[742,866,1164,1187]" gridcol="3" gridrow="20" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED59FFC408FB8" box="[884,1008,1164,1187]" gridcol="4" gridrow="20" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D59FFBED8FB8" box="[1026,1117,1164,1187]" gridcol="5" gridrow="20" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD59FFB588FB8" box="[1136,1256,1164,1187]" gridcol="6" gridrow="20" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D59FFA268FB8" box="[1277,1430,1164,1187]" gridcol="7" gridrow="20" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD59FF9A28FB8" box="[1446,1554,1164,1187]" gridcol="8" gridrow="20" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD59FF95C8FB8" box="[1575,1772,1164,1187]" gridcol="9" gridrow="20" pageId="3" pageNumber="154">5&gt;6</td>
<td id="70115DCA8B318C8DF8E8D59FF8DD8FB8" box="[1794,1901,1164,1187]" gridcol="10" gridrow="20" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF868D59FF85E8FB8" box="[1922,2030,1164,1187]" gridcol="11" gridrow="20" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD5BFF85E8FD8" box="[160,2030,1196,1219]" gridrow="21" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD5BFFF0F8FD8" box="[160,191,1196,1219]" gridcol="0" gridrow="21" pageId="3" pageNumber="154">17</th>
<td id="70115DCA8B318C8DFF24D5BFFD838FD8" box="[206,563,1196,1219]" gridcol="1" gridrow="21" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D5BFFE1C8FD8" authority="(Say)" baseAuthorityName="Say" baseAuthorityYear="1823" box="[206,428,1196,1219]" class="Insecta" family="Carabidae" genus="Tachyta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="inornata">
<emphasis id="BF84EAE68B317371FF24D5BFFEC28FD8" box="[206,370,1196,1219]" italics="true" pageId="3" pageNumber="154">Tachyta inornata</emphasis>
(Say)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D5BFFD7D8FD8" box="[590,717,1196,1219]" gridcol="2" gridrow="21" pageId="3" pageNumber="154">Tachyina</td>
<td id="70115DCA8B318C8DFD0CD5BFFCD28FD8" box="[742,866,1196,1219]" gridcol="3" gridrow="21" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED5BFFC408FD8" box="[884,1008,1196,1219]" gridcol="4" gridrow="21" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D5BFFBED8FD8" box="[1026,1117,1196,1219]" gridcol="5" gridrow="21" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD5BFFB588FD8" box="[1136,1256,1196,1219]" gridcol="6" gridrow="21" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D5BFFA268FD8" box="[1277,1430,1196,1219]" gridcol="7" gridrow="21" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD5BFF9A28FD8" box="[1446,1554,1196,1219]" gridcol="8" gridrow="21" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD5BFF95C8FD8" box="[1575,1772,1196,1219]" gridcol="9" gridrow="21" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D5BFF8DD8FD8" box="[1794,1901,1196,1219]" gridcol="10" gridrow="21" pageId="3" pageNumber="154">11</td>
<td id="70115DCA8B318C8DF868D5BFF85E8FD8" box="[1922,2030,1196,1219]" gridcol="11" gridrow="21" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD5DFF85E8FF8" box="[160,2030,1228,1251]" gridrow="22" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD5DFFF0F8FF8" box="[160,191,1228,1251]" gridcol="0" gridrow="22" pageId="3" pageNumber="154">18</th>
<td id="70115DCA8B318C8DFF24D5DFFD838FF8" box="[206,563,1228,1251]" gridcol="1" gridrow="22" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D5DFFE048FF8" authority="LeConte" authorityName="LeConte" authorityYear="1852" box="[206,436,1228,1251]" class="Insecta" family="Carabidae" genus="Tachys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="vittiger">
<emphasis id="BF84EAE68B317371FF24D5DFFEEB8FF8" box="[206,347,1228,1251]" italics="true" pageId="3" pageNumber="154">Tachys vittiger</emphasis>
LeConte
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D5DFFD7D8FF8" box="[590,717,1228,1251]" gridcol="2" gridrow="22" pageId="3" pageNumber="154">Tachyina</td>
<td id="70115DCA8B318C8DFD0CD5DFFCD28FF8" box="[742,866,1228,1251]" gridcol="3" gridrow="22" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED5DFFC408FF8" box="[884,1008,1228,1251]" gridcol="4" gridrow="22" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D5DFFBED8FF8" box="[1026,1117,1228,1251]" gridcol="5" gridrow="22" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD5DFFB588FF8" box="[1136,1256,1228,1251]" gridcol="6" gridrow="22" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D5DFFA268FF8" box="[1277,1430,1228,1251]" gridcol="7" gridrow="22" pageId="3" pageNumber="154">&gt; Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD5DFF9A28FF8" box="[1446,1554,1228,1251]" gridcol="8" gridrow="22" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD5DFF95C8FF8" box="[1575,1772,1228,1251]" gridcol="9" gridrow="22" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D5DFF8DD8FF8" box="[1794,1901,1228,1251]" gridcol="10" gridrow="22" pageId="3" pageNumber="154">10</td>
<td id="70115DCA8B318C8DF868D5DFF85E8FF8" box="[1922,2030,1228,1251]" gridcol="11" gridrow="22" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD5FFF85E8E18" box="[160,2030,1260,1283]" gridrow="23" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD5FFFF0F8E18" box="[160,191,1260,1283]" gridcol="0" gridrow="23" pageId="3" pageNumber="154">19</th>
<td id="70115DCA8B318C8DFF24D5FFFD838E18" box="[206,563,1260,1283]" gridcol="1" gridrow="23" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D5FFFE4A8E18" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[206,506,1260,1283]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B317371FF24D5FFFE178E18" box="[206,423,1260,1283]" italics="true" pageId="3" pageNumber="154">Nesamblyops oreobius</emphasis>
(Broun)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D5FFFD7D8E18" box="[590,717,1260,1283]" gridcol="2" gridrow="23" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D5FFFD298E18" authorityName="Jeannel" authorityYear="1937" box="[590,665,1260,1283]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD5FFFCD28E18" box="[742,866,1260,1283]" gridcol="3" gridrow="23" pageId="3" pageNumber="154">2Med+1Bas</td>
<td id="70115DCA8B318C8DFC9ED5FFFC408E18" box="[884,1008,1260,1283]" gridcol="4" gridrow="23" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D5FFFBED8E18" box="[1026,1117,1260,1283]" gridcol="5" gridrow="23" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD5FFFB588E18" box="[1136,1256,1260,1283]" gridcol="6" gridrow="23" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D5FFFA268E18" box="[1277,1430,1260,1283]" gridcol="7" gridrow="23" pageId="3" pageNumber="154">= ¼ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD5FFF9A28E18" box="[1446,1554,1260,1283]" gridcol="8" gridrow="23" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD5FFF95C8E18" box="[1575,1772,1260,1283]" gridcol="9" gridrow="23" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D5FFF8DD8E18" box="[1794,1901,1260,1283]" gridcol="10" gridrow="23" pageId="3" pageNumber="154">6</td>
<td id="70115DCA8B318C8DF868D5FFF85E8E18" box="[1922,2030,1260,1283]" gridcol="11" gridrow="23" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD418F85E8E39" box="[160,2030,1291,1314]" gridrow="24" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD418FF0F8E39" box="[160,191,1291,1314]" gridcol="0" gridrow="24" pageId="3" pageNumber="154">20</th>
<td id="70115DCA8B318C8DFF24D418FD838E39" box="[206,563,1291,1314]" gridcol="1" gridrow="24" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D418FDB98E39" authority="(Broun)" baseAuthorityName="Broun" box="[206,521,1291,1314]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B317371FF24D418FE078E39" box="[206,439,1291,1314]" italics="true" pageId="3" pageNumber="154">Nesamblyops subcaecus</emphasis>
(Broun)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D418FD7D8E39" box="[590,717,1291,1314]" gridcol="2" gridrow="24" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D418FD298E39" authorityName="Jeannel" authorityYear="1937" box="[590,665,1291,1314]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD418FCD28E39" box="[742,866,1291,1314]" gridcol="3" gridrow="24" pageId="3" pageNumber="154">2Med+1Bas</td>
<td id="70115DCA8B318C8DFC9ED418FC408E39" box="[884,1008,1291,1314]" gridcol="4" gridrow="24" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D418FBED8E39" box="[1026,1117,1291,1314]" gridcol="5" gridrow="24" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFB9AD418FB588E39" box="[1136,1256,1291,1314]" gridcol="6" gridrow="24" pageId="3" pageNumber="154">Margined</td>
<td id="70115DCA8B318C8DFB17D418FA268E39" box="[1277,1430,1291,1314]" gridcol="7" gridrow="24" pageId="3" pageNumber="154">= ¼ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD418F9A28E39" box="[1446,1554,1291,1314]" gridcol="8" gridrow="24" pageId="3" pageNumber="154">8</td>
<td id="70115DCA8B318C8DF9CDD418F95C8E39" box="[1575,1772,1291,1314]" gridcol="9" gridrow="24" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D418F8DD8E39" box="[1794,1901,1291,1314]" gridcol="10" gridrow="24" pageId="3" pageNumber="154">5</td>
<td id="70115DCA8B318C8DF868D418F85E8E39" box="[1922,2030,1291,1314]" gridcol="11" gridrow="24" pageId="3" pageNumber="154">2 + 1</td>
</tr>
<tr id="33C034B68B318C8DFF4AD438F85E8E59" box="[160,2030,1323,1346]" gridrow="25" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD438FF0F8E59" box="[160,191,1323,1346]" gridcol="0" gridrow="25" pageId="3" pageNumber="154">21</th>
<td id="70115DCA8B318C8DFF24D438FD838E59" box="[206,563,1323,1346]" gridcol="1" gridrow="25" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D438FE7D8E59" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1884" box="[206,461,1323,1346]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="BF84EAE68B317371FF24D438FECB8E59" box="[206,379,1323,1346]" italics="true" pageId="3" pageNumber="154">Zeanillus pallidus</emphasis>
(Broun)
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D438FD7D8E59" box="[590,717,1323,1346]" gridcol="2" gridrow="25" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D438FD298E59" authorityName="Jeannel" authorityYear="1937" box="[590,665,1323,1346]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD438FCD28E59" box="[742,866,1323,1346]" gridcol="3" gridrow="25" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED438FC408E59" box="[884,1008,1323,1346]" gridcol="4" gridrow="25" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFBE8D438FBED8E59" box="[1026,1117,1323,1346]" gridcol="5" gridrow="25" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFB9AD438FB588E59" box="[1136,1256,1323,1346]" gridcol="6" gridrow="25" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D438FA268E59" box="[1277,1430,1323,1346]" gridcol="7" gridrow="25" pageId="3" pageNumber="154">= ⅔ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD438F9A28E59" box="[1446,1554,1323,1346]" gridcol="8" gridrow="25" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF9CDD438F95C8E59" box="[1575,1772,1323,1346]" gridcol="9" gridrow="25" pageId="3" pageNumber="154">5&gt;6</td>
<td id="70115DCA8B318C8DF8E8D438F8DD8E59" box="[1794,1901,1323,1346]" gridcol="10" gridrow="25" pageId="3" pageNumber="154">14</td>
<td id="70115DCA8B318C8DF868D438F85E8E59" box="[1922,2030,1323,1346]" gridcol="11" gridrow="25" pageId="3" pageNumber="154">2</td>
</tr>
<tr id="33C034B68B318C8DFF4AD458F85E8E79" box="[160,2030,1355,1378]" gridrow="26" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD458FF0F8E79" box="[160,191,1355,1378]" gridcol="0" gridrow="26" pageId="3" pageNumber="154">22</th>
<td id="70115DCA8B318C8DFF24D458FD838E79" box="[206,563,1355,1378]" gridcol="1" gridrow="26" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D458FDA28E79" authority="Jeannel" authorityName="Jeannel" box="[206,530,1355,1378]" class="Insecta" family="Carabidae" genus="Pelodiaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="sulcatipennis">
<emphasis id="BF84EAE68B317371FF24D458FE758E79" box="[206,453,1355,1378]" italics="true" pageId="3" pageNumber="154">Pelodiaetus sulcatipennis</emphasis>
Jeannel
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D458FD7D8E79" box="[590,717,1355,1378]" gridcol="2" gridrow="26" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D458FD298E79" authorityName="Jeannel" authorityYear="1937" box="[590,665,1355,1378]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD458FCD28E79" box="[742,866,1355,1378]" gridcol="3" gridrow="26" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED458FC408E79" box="[884,1008,1355,1378]" gridcol="4" gridrow="26" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFBE8D458FBED8E79" box="[1026,1117,1355,1378]" gridcol="5" gridrow="26" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFB9AD458FB588E79" box="[1136,1256,1355,1378]" gridcol="6" gridrow="26" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D458FA268E79" box="[1277,1430,1355,1378]" gridcol="7" gridrow="26" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD458F9A28E79" box="[1446,1554,1355,1378]" gridcol="8" gridrow="26" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF9CDD458F95C8E79" box="[1575,1772,1355,1378]" gridcol="9" gridrow="26" pageId="3" pageNumber="154">5&gt;6</td>
<td id="70115DCA8B318C8DF8E8D458F8DD8E79" box="[1794,1901,1355,1378]" gridcol="10" gridrow="26" pageId="3" pageNumber="154">12</td>
<td id="70115DCA8B318C8DF868D458F85E8E79" box="[1922,2030,1355,1378]" gridcol="11" gridrow="26" pageId="3" pageNumber="154">2</td>
</tr>
<tr id="33C034B68B318C8DFF4AD478F85E8E99" box="[160,2030,1387,1410]" gridrow="27" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD478FF0F8E99" box="[160,191,1387,1410]" gridcol="0" gridrow="27" pageId="3" pageNumber="154">23</th>
<td id="70115DCA8B318C8DFF24D478FD838E99" box="[206,563,1387,1410]" gridcol="1" gridrow="27" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D478FDB78E99" authority="Moore" authorityName="Moore" box="[206,519,1387,1410]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="prominens">
<emphasis id="BF84EAE68B317371FF24D478FE708E99" box="[206,448,1387,1410]" italics="true" pageId="3" pageNumber="154">Pelodiaetodes prominens</emphasis>
Moore
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D478FD7D8E99" box="[590,717,1387,1410]" gridcol="2" gridrow="27" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D478FD298E99" authorityName="Jeannel" authorityYear="1937" box="[590,665,1387,1410]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD478FCD28E99" box="[742,866,1387,1410]" gridcol="3" gridrow="27" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED478FC408E99" box="[884,1008,1387,1410]" gridcol="4" gridrow="27" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFBE8D478FBED8E99" box="[1026,1117,1387,1410]" gridcol="5" gridrow="27" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFB9AD478FB588E99" box="[1136,1256,1387,1410]" gridcol="6" gridrow="27" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D478FA268E99" box="[1277,1430,1387,1410]" gridcol="7" gridrow="27" pageId="3" pageNumber="154">= ⅔ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD478F9A28E99" box="[1446,1554,1387,1410]" gridcol="8" gridrow="27" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF9CDD478F95C8E99" box="[1575,1772,1387,1410]" gridcol="9" gridrow="27" pageId="3" pageNumber="154">5&gt;6</td>
<td id="70115DCA8B318C8DF8E8D478F8DD8E99" box="[1794,1901,1387,1410]" gridcol="10" gridrow="27" pageId="3" pageNumber="154">12</td>
<td id="70115DCA8B318C8DF868D478F85E8E99" box="[1922,2030,1387,1410]" gridcol="11" gridrow="27" pageId="3" pageNumber="154">2</td>
</tr>
<tr id="33C034B68B318C8DFF4AD499F85E8EB9" box="[160,2030,1418,1442]" gridrow="28" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD499FF0F8EB9" box="[160,191,1418,1442]" gridcol="0" gridrow="28" pageId="3" pageNumber="154">24</th>
<td id="70115DCA8B318C8DFF24D499FD838EB9" box="[206,563,1418,1442]" gridcol="1" gridrow="28" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D498FE7F8EB9" authority="Sokolov" authorityName="Sokolov" box="[206,463,1419,1442]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="BF84EAE68B317371FF24D498FECA8EB9" box="[206,378,1419,1442]" italics="true" pageId="3" pageNumber="154">Anillinus lescheni</emphasis>
Sokolov
</taxonomicName>
and
</td>
<td id="70115DCA8B318C8DFDA4D499FD7D8EB9" box="[590,717,1418,1442]" gridcol="2" gridrow="28" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D499FD298EBA" authorityName="Jeannel" authorityYear="1937" box="[590,665,1418,1441]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD499FCD28EB9" box="[742,866,1418,1442]" gridcol="3" gridrow="28" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED499FC408EB9" box="[884,1008,1418,1442]" gridcol="4" gridrow="28" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D499FBED8EB9" box="[1026,1117,1418,1442]" gridcol="5" gridrow="28" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFB9AD499FB588EB9" box="[1136,1256,1418,1442]" gridcol="6" gridrow="28" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D499FA268EB9" box="[1277,1430,1418,1442]" gridcol="7" gridrow="28" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD499F9A28EB9" box="[1446,1554,1418,1442]" gridcol="8" gridrow="28" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF9CDD499F95C8EB9" box="[1575,1772,1418,1442]" gridcol="9" gridrow="28" pageId="3" pageNumber="154">5&gt;6</td>
<td id="70115DCA8B318C8DF8E8D499F8DD8EB9" box="[1794,1901,1418,1442]" gridcol="10" gridrow="28" pageId="3" pageNumber="154">10</td>
<td id="70115DCA8B318C8DF868D499F85E8EB9" box="[1922,2030,1418,1442]" gridcol="11" gridrow="28" pageId="3" pageNumber="154">2</td>
</tr>
<tr id="33C034B68B318C8DFF4AD4BBF85E8EA4" box="[160,2030,1448,1471]" gridrow="29" pageId="3" pageNumber="154" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td id="70115DCA8B318C8DFF24D4BBFD838EA4" box="[206,563,1448,1471]" gridcol="1" gridrow="29" pageId="3" pageNumber="154">Carlton</td>
</tr>
<tr id="33C034B68B318C8DFF4AD4DBF85E8EC4" box="[160,2030,1480,1503]" gridrow="30" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD4DBFF0F8EC4" box="[160,191,1480,1503]" gridcol="0" gridrow="30" pageId="3" pageNumber="154">25</th>
<td id="70115DCA8B318C8DFF24D4DBFD838EC4" box="[206,563,1480,1503]" gridcol="1" gridrow="30" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D4DBFE0D8EC4" authority="Barr" authorityName="Barr" authorityYear="1995" box="[206,445,1480,1503]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="BF84EAE68B317371FF24D4DBFE3C8EC4" box="[206,396,1480,1503]" italics="true" pageId="3" pageNumber="154">Serranillus jeanneli</emphasis>
Barr
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D4DBFD7D8EC4" box="[590,717,1480,1503]" gridcol="2" gridrow="30" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D4DBFD298EC4" authorityName="Jeannel" authorityYear="1937" box="[590,665,1480,1503]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD4DBFCD28EC4" box="[742,866,1480,1503]" gridcol="3" gridrow="30" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED4DBFC408EC4" box="[884,1008,1480,1503]" gridcol="4" gridrow="30" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFBE8D4DBFBED8EC4" box="[1026,1117,1480,1503]" gridcol="5" gridrow="30" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFB9AD4DBFB588EC4" box="[1136,1256,1480,1503]" gridcol="6" gridrow="30" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D4DBFA268EC4" box="[1277,1430,1480,1503]" gridcol="7" gridrow="30" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD4DBF9A28EC4" box="[1446,1554,1480,1503]" gridcol="8" gridrow="30" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF9CDD4DBF95C8EC4" box="[1575,1772,1480,1503]" gridcol="9" gridrow="30" pageId="3" pageNumber="154">5&gt;6</td>
<td id="70115DCA8B318C8DF8E8D4DBF8DD8EC4" box="[1794,1901,1480,1503]" gridcol="10" gridrow="30" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF868D4DBF85E8EC4" box="[1922,2030,1480,1503]" gridcol="11" gridrow="30" pageId="3" pageNumber="154">2</td>
</tr>
<tr id="33C034B68B318C8DFF4AD4F4F85E8EE5" box="[160,2030,1511,1534]" gridrow="31" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD4F4FF0F8EE5" box="[160,191,1511,1534]" gridcol="0" gridrow="31" pageId="3" pageNumber="154">26</th>
<td id="70115DCA8B318C8DFF24D4F4FD838EE5" box="[206,563,1511,1534]" gridcol="1" gridrow="31" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D4F4FD838EE5" authority="Vigna Taglianti" authorityName="Vigna Taglianti" authorityYear="1973" box="[206,563,1511,1534]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="zullinii">
<emphasis id="BF84EAE68B317371FF24D4F4FE2A8EE5" box="[206,410,1511,1534]" italics="true" pageId="3" pageNumber="154">Geocharidius zullinii</emphasis>
Vigna Taglianti
</taxonomicName>
</td>
<td id="70115DCA8B318C8DFDA4D4F4FD7D8EE5" box="[590,717,1511,1534]" gridcol="2" gridrow="31" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D4F4FD298EE5" authorityName="Jeannel" authorityYear="1937" box="[590,665,1511,1534]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD4F4FCD28EE5" box="[742,866,1511,1534]" gridcol="3" gridrow="31" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED4F4FC408EE5" box="[884,1008,1511,1534]" gridcol="4" gridrow="31" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFBE8D4F4FBED8EE5" box="[1026,1117,1511,1534]" gridcol="5" gridrow="31" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFB9AD4F4FB588EE5" box="[1136,1256,1511,1534]" gridcol="6" gridrow="31" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D4F4FA268EE5" box="[1277,1430,1511,1534]" gridcol="7" gridrow="31" pageId="3" pageNumber="154">= ½ Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD4F4F9A28EE5" box="[1446,1554,1511,1534]" gridcol="8" gridrow="31" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF9CDD4F4F95C8EE5" box="[1575,1772,1511,1534]" gridcol="9" gridrow="31" pageId="3" pageNumber="154">5&gt;6</td>
<td id="70115DCA8B318C8DF8E8D4F4F8DD8EE5" box="[1794,1901,1511,1534]" gridcol="10" gridrow="31" pageId="3" pageNumber="154">11</td>
<td id="70115DCA8B318C8DF868D4F4F85E8EE5" box="[1922,2030,1511,1534]" gridcol="11" gridrow="31" pageId="3" pageNumber="154">2</td>
</tr>
<tr id="33C034B68B318C8DFF4AD714F85E8D05" box="[160,2030,1543,1566]" gridrow="32" pageId="3" pageNumber="154">
<th id="70115DCA8B318C8DFF4AD714FF0F8D05" box="[160,191,1543,1566]" gridcol="0" gridrow="32" pageId="3" pageNumber="154">27</th>
<td id="70115DCA8B318C8DFF24D714FD838D05" box="[206,563,1543,1566]" gridcol="1" gridrow="32" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FF24D714FE5C8D05" authority="Andujar" authorityName="Andujar" box="[206,492,1543,1566]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="BF84EAE68B317371FF24D714FE288D05" box="[206,408,1543,1566]" italics="true" pageId="3" pageNumber="154">Typhlocharis martini</emphasis>
Andujar
</taxonomicName>
,
</td>
<td id="70115DCA8B318C8DFDA4D714FD7D8D05" box="[590,717,1543,1566]" gridcol="2" gridrow="32" pageId="3" pageNumber="154">
<taxonomicName id="4AF04D778B317371FDA4D714FD298D05" authorityName="Jeannel" authorityYear="1937" box="[590,665,1543,1566]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</td>
<td id="70115DCA8B318C8DFD0CD714FCD28D05" box="[742,866,1543,1566]" gridcol="3" gridrow="32" pageId="3" pageNumber="154">1MedProx</td>
<td id="70115DCA8B318C8DFC9ED714FC408D05" box="[884,1008,1543,1566]" gridcol="4" gridrow="32" pageId="3" pageNumber="154">Present</td>
<td id="70115DCA8B318C8DFBE8D714FBED8D05" box="[1026,1117,1543,1566]" gridcol="5" gridrow="32" pageId="3" pageNumber="154">Absent</td>
<td id="70115DCA8B318C8DFB9AD714FB588D05" box="[1136,1256,1543,1566]" gridcol="6" gridrow="32" pageId="3" pageNumber="154">Unmargined</td>
<td id="70115DCA8B318C8DFB17D714FA268D05" box="[1277,1430,1543,1566]" gridcol="7" gridrow="32" pageId="3" pageNumber="154">= Ø mscx</td>
<td id="70115DCA8B318C8DFA4CD714F9A28D05" box="[1446,1554,1543,1566]" gridcol="8" gridrow="32" pageId="3" pageNumber="154">7</td>
<td id="70115DCA8B318C8DF9CDD714F95C8D05" box="[1575,1772,1543,1566]" gridcol="9" gridrow="32" pageId="3" pageNumber="154">5=6</td>
<td id="70115DCA8B318C8DF8E8D714F8DD8D05" box="[1794,1901,1543,1566]" gridcol="10" gridrow="32" pageId="3" pageNumber="154">9</td>
<td id="70115DCA8B318C8DF868D714F85E8D05" box="[1922,2030,1543,1566]" gridcol="11" gridrow="32" pageId="3" pageNumber="154">0</td>
</tr>
<tr id="33C034B68B318C8DFF4AD736F85E8D27" box="[160,2030,1573,1596]" gridrow="33" pageId="3" pageNumber="154" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td id="70115DCA8B318C8DFF24D736FD838D27" box="[206,563,1573,1596]" gridcol="1" gridrow="33" pageId="3" pageNumber="154">Lencina and Serrano</td>
</tr>
</table>
</paragraph>
<tableNote id="7016377A8B317371FF7CD755F90B8DBE" pageId="3" pageNumber="154" targetBox="[160,2030,531,1596]" targetPageId="3">
<paragraph id="8D4F36F48B317371FF7CD755FCDE8D46" blockId="3.[150,1723,1606,1701]" box="[150,878,1606,1629]" pageId="3" pageNumber="154">* Med—medial setae, MedProx—medial proximal seta, Bas—basal seta;</paragraph>
<paragraph id="8D4F36F48B317371FF7CD779FE4D8D9A" blockId="3.[150,1723,1606,1701]" box="[150,509,1642,1665]" pageId="3" pageNumber="154">** Ø mscx—diameter of mesocoxa;</paragraph>
<paragraph id="8D4F36F48B317371FF7CD79DF90B8DBE" blockId="3.[150,1723,1606,1701]" box="[150,1723,1678,1701]" pageId="3" pageNumber="154">*** 5 &amp; 6USP—5th and 6th pores of umbilical series; “5=6” means: the distance from 5th pore to elytral margin equals the distance from 6th pore to elytral margin</paragraph>
</tableNote>
<paragraph id="8D4F36F48B367376FF7DD185FCB28BAA" blockId="4.[151,1437,150,1833]" box="[151,770,150,177]" pageId="4" pageNumber="155">
<emphasis id="BF84EAE68B367376FF7DD185FEE88BAA" bold="true" box="[151,344,150,177]" pageId="4" pageNumber="155">Subtribal name.</emphasis>
Based on the name of the 
<typeStatus id="524B88568B367376FD68D184FD028BAA" box="[642,690,151,177]" pageId="4" pageNumber="155">type</typeStatus>
genus.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B36737FFF2DD1A9FD4689D6" lastPageId="13" lastPageNumber="164" pageId="4" pageNumber="155" type="description">
<paragraph id="8D4F36F48B367376FF2DD1A9FBDB8926" blockId="4.[151,1437,150,1833]" pageId="4" pageNumber="155">
<emphasis id="BF84EAE68B367376FF2DD1A9FEED8BCF" bold="true" box="[199,349,186,212]" pageId="4" pageNumber="155">Recognition.</emphasis>
The new subtribe comprises members of 
<taxonomicName id="4AF04D778B367376FCD1D1A8FC228BCE" authorityName="Jeannel" authorityYear="1937" box="[827,914,187,213]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
with the following combination of diagnostic features: (1) basistipes of maxilla with four setae (two basal and two medial), of which outer basal and both medial stipital setae of similar length (
<figureCitation id="15CB2A718B367376FDE1D010FD3E8A06" box="[523,654,259,285]" captionStart="FIGURE 2" captionStartId="6.[151,250,1585,1609]" captionTargetBox="[189,1397,181,1561]" captionTargetId="figure-32@6.[189,1397,181,1561]" captionTargetPageId="6" captionText="FIGURE 2. SEM illustrations of structural features of labial complex of Anillini: A–B—subtribe Nesamblyopina; A— Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Zeanillus punctigerus (Broun); D—Pelodiaetodes insularis Sokolov; E—Anillinus forthoodensis Sokolov and Reddell; F—Geocharidius minimus Sokolov and Kavanaugh. Legend: ca—cardo, bs—basistipes, bss—basal stipital setae, mss—medial stipital setae, pf—palpifer. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184228" httpUri="https://zenodo.org/record/10184228/files/figure.png" pageId="4" pageNumber="155">Figs 2A–B</figureCitation>
, bss, mss); (2) posterior angles of pronotum rounded and without basolateral denticles (
<figureCitation id="15CB2A718B367376FE62D034FDB68A5A" box="[392,518,295,321]" captionStart="FIGURE3" captionStartId="7.[199,298,1291,1315]" captionTargetBox="[151,1436,181,1267]" captionTargetId="figure-15@7.[151,1436,181,1267]" captionTargetPageId="7" captionText="FIGURE3. SEM illustrationsof structuralfeatures of pronota ofAnillini:A–B—subtribe Nesamblyopina;A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus alleni Sokolov and Carlton; D—Anillinus forthoodensis Sokolov andReddell;E—Serranillusjeanneli Barr; F—Geocharidius balini Sokolov and Kavanaugh; G—Zeanillus punctigerus (Broun); H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus nunni Sokolov. Legend: ast—setae of anterior margin of pronotum, bst—setae of basal margin of pronotum. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184231" httpUri="https://zenodo.org/record/10184231/files/figure.png" pageId="4" pageNumber="155">Figs 3A–B</figureCitation>
); (3) basal margin of pronotum lacking marginal setation (
<figureCitation id="15CB2A718B367376FB60D034FAB88A5A" box="[1162,1288,295,321]" captionStart="FIGURE3" captionStartId="7.[199,298,1291,1315]" captionTargetBox="[151,1436,181,1267]" captionTargetId="figure-15@7.[151,1436,181,1267]" captionTargetPageId="7" captionText="FIGURE3. SEM illustrationsof structuralfeatures of pronota ofAnillini:A–B—subtribe Nesamblyopina;A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus alleni Sokolov and Carlton; D—Anillinus forthoodensis Sokolov andReddell;E—Serranillusjeanneli Barr; F—Geocharidius balini Sokolov and Kavanaugh; G—Zeanillus punctigerus (Broun); H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus nunni Sokolov. Legend: ast—setae of anterior margin of pronotum, bst—setae of basal margin of pronotum. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184231" httpUri="https://zenodo.org/record/10184231/files/figure.png" pageId="4" pageNumber="155">Figs 3A–B</figureCitation>
, cf. with 
<figureCitation id="15CB2A718B367376FA86D034FF7F8A7E" captionStart="FIGURE3" captionStartId="7.[199,298,1291,1315]" captionTargetBox="[151,1436,181,1267]" captionTargetId="figure-15@7.[151,1436,181,1267]" captionTargetPageId="7" captionText="FIGURE3. SEM illustrationsof structuralfeatures of pronota ofAnillini:A–B—subtribe Nesamblyopina;A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus alleni Sokolov and Carlton; D—Anillinus forthoodensis Sokolov andReddell;E—Serranillusjeanneli Barr; F—Geocharidius balini Sokolov and Kavanaugh; G—Zeanillus punctigerus (Broun); H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus nunni Sokolov. Legend: ast—setae of anterior margin of pronotum, bst—setae of basal margin of pronotum. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184231" httpUri="https://zenodo.org/record/10184231/files/figure.png" pageId="4" pageNumber="155">Figs 3F–I</figureCitation>
, bst); (4) pterothorax with well-developed mesothoracic anapleural sutures (
<figureCitation id="15CB2A718B367376FBF3D058FB268A7E" box="[1049,1174,331,357]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="4" pageNumber="155">Figs 4A–B</figureCitation>
, msts); (5) metaventrite (
<figureCitation id="15CB2A718B367376FF75D07CFEAE8A92" box="[159,286,367,393]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="4" pageNumber="155">Figs 4A–B</figureCitation>
, mtv) very short with distance between meso- and metacoxae about 0.15–0.20 diameter of mesocoxa; (6) metaventral process (
<figureCitation id="15CB2A718B367376FE43D080FD968AB6" box="[425,550,403,429]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="4" pageNumber="155">Figs 4A–B</figureCitation>
, mtp) narrowly margined; (7) metendosternite of T-shape, without visible lateral arms (
<figureCitation id="15CB2A718B367376FF36D0A4FEEE8ACA" box="[220,350,438,465]" captionStart="FIGURE 5" captionStartId="9.[151,250,1796,1820]" captionTargetBox="[185,1401,181,1771]" captionTargetId="figure-15@9.[185,1401,181,1772]" captionTargetPageId="9" captionText="FIGURE 5. Digital illustrations of structural features of pterothorax of Anillini, ventral aspect: A–D—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Nesamblyops brouni Sokolov; D—Nesamblyops viator n.sp.; E—Anillinus langdoni Sokolov and Carlton; F—Anillinus Cherokee Sokolov and Carlton; G—Geocharidius longinoi Sokolov and Kavanaugh; H—Geocharidius disjunctus Sokolov and Kavanaugh; I—Zapotecanillus longinoi Sokolov; J—Zeanillus pellucidus Sokolov; K—Zeanillus pallidus (Broun); L—Zeanillus nunni Sokolov; M—Pelodiaetodes insularis Sokolov; N—Pelodiaetodes nunni Sokolov; O—Pelodiaetodes moorei Sokolov; P—Pelodiaetus sulcatipennis Jeannel.Legend: mtes—metendosternite. Scales intentionally omitted." figureDoi="http://doi.org/10.5281/zenodo.10184235" httpUri="https://zenodo.org/record/10184235/files/figure.png" pageId="4" pageNumber="155">Figs 5A–D</figureCitation>
); (8) elytra with umbilical series of eight pores situated along virtual line almost parallel to lateral elytral margin (
<figureCitation id="15CB2A718B367376FEACD0C8FE788AEE" box="[326,456,475,501]" captionStart="FIGURE 6" captionStartId="10.[151,250,1867,1891]" captionTargetBox="[189,1398,181,1843]" captionTargetId="figure-32@10.[189,1398,181,1843]" captionTargetPageId="10" captionText="FIGURE 6. SEM illustrations of structural features of elytra of Anillini, left lateral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus carolinae Casey; D—Anillinus alleni Sokolov and Carlton; E—Serranillus donovani (Jeannel); F—Geocharidius jalapensis Sokolov and Kavanaugh; G—Zeanillus punctigerus (Broun); H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus nunni Sokolov. Legend: ed2—scutellar seta; ed6— discal seta, ed8—apical seta; eo1…eo9—setae 1–9 of umbilical series of pores. Scale = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184237" httpUri="https://zenodo.org/record/10184237/files/figure.png" pageId="4" pageNumber="155">Figs 6A–B</figureCitation>
); (9) protibia with short oblique notch and short posterior setal row of 4–6 setae (
<figureCitation id="15CB2A718B367376FA80D0C8FF6C8902" captionStart="FIGURE 7" captionStartId="11.[151,250,1452,1476]" captionTargetBox="[168,1419,201,1429]" captionTargetId="figure-15@11.[168,1419,201,1429]" captionTargetPageId="11" captionText="FIGURE 7. SEM illustrations of structural features of protibia of Anillini, dorsal (E) and ventral (A–D, F–H) aspects: A– B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D—Geocharidius comayaguanus Sokolov and Kavanaugh; E—Geocharidius zullini Vigna Taglianti; F—Zeanillus brouni Sokolov; G—Pelodiaetodes nunni Sokolov; H—Pelodiaetus sulcatipennis Jeannel. Legend: as—articulosetae; psr—posterior setal row. Scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10184239" httpUri="https://zenodo.org/record/10184239/files/figure.png" pageId="4" pageNumber="155">Figs 7A–B</figureCitation>
, psr); (10) basal lobes of median lobe of males asymmetrical (
<figureCitation id="15CB2A718B367376FC49D0ECFB978902" box="[931,1063,511,537]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStartId-0="30.[151,250,1365,1389]" captionStartId-1="32.[151,250,1380,1404]" captionStartId-2="33.[151,250,788,812]" captionTargetBox-0="[157,1433,187,1336]" captionTargetBox-1="[157,1431,188,1353]" captionTargetBox-2="[158,1431,188,760]" captionTargetId-0="figure-32@30.[151,1436,181,1341]" captionTargetId-1="figure-32@32.[151,1436,181,1356]" captionTargetId-2="figure-15@33.[152,1436,181,764]" captionTargetPageId-0="30" captionTargetPageId-1="32" captionTargetPageId-2="33" captionText-0="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." captionText-1="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthur’s Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." captionText-2="FIGURE 16. Line drawings of male genitalia of New Zealand Nesamblyops species. N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. viator (Secretary Island, Grono Bay, Fjordland, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. victoriae (Capleston, Victoria Range, Buller, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10184255" figureDoi-1="http://doi.org/10.5281/zenodo.10184257" figureDoi-2="http://doi.org/10.5281/zenodo.10184259" httpUri-0="https://zenodo.org/record/10184255/files/figure.png" httpUri-1="https://zenodo.org/record/10184257/files/figure.png" httpUri-2="https://zenodo.org/record/10184259/files/figure.png" pageId="4" pageNumber="155">Figs 14–16</figureCitation>
); (11) ovipositor of females with gonocoxite 2 with three (1 medial + 2 lateral) ensiform setae (
<figureCitation id="15CB2A718B367376FCA6D330FC7B8926" box="[844,971,547,573]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[167,1420,200,1667]" captionTargetId="figure-32@12.[167,1420,200,1667]" captionTargetPageId="12" captionText="FIGURE 8. SEM illustrations of ovipositor sclerites of Anillini, lateral (A, C–F) and ventral (B) aspects: A–B—subtribe Nesamblyopina; A—Nesamblyops brouni Sokolov, B—Nesamblyops brouni Sokolov; C—Anillinus lescheni Sokolov and Carlton; D—Geocharidius longinoi Sokolov and Kavanaugh; E—Pelodiaetodes moorei Sokolov; F—Pelodiaetodes nunni Sokolov. Legend: gc1—gonocoxite 1; gc2—gonocoxite 2; les1-2—lateral ensiform setae 1 and 2; mes—medial ensiform seta; ns—nematiform setae. Scale = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.10184243" httpUri="https://zenodo.org/record/10184243/files/figure.png" pageId="4" pageNumber="155">Figs 8A–B</figureCitation>
, mes, les1–2).
</paragraph>
<paragraph id="8D4F36F48B367376FF2DD355FA8E89D6" blockId="4.[151,1437,150,1833]" pageId="4" pageNumber="155">
<emphasis id="BF84EAE68B367376FF2DD355FD93897B" bold="true" box="[199,547,582,608]" pageId="4" pageNumber="155">
Relationships within 
<taxonomicName id="4AF04D778B367376FE28D355FDAE897B" authorityName="Jeannel" authorityYear="1937" box="[450,542,582,608]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
.
</emphasis>
The new subtribe represents a morphologically distinct lineage within the tribe 
<taxonomicName id="4AF04D778B367376FF7DD378FF5E899E" authorityName="Jeannel" authorityYear="1937" box="[151,238,619,645]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
and differs from other tribe representatives in many important characteristics of the labiomaxillary complex, pronotum, pterothorax, elytral and leg chaetotaxy as well as the structural details of the male and female genitalia, traits which did not attract significant attention from researchers who worked on the taxonomy of 
<taxonomicName id="4AF04D778B367376FB35D3A0FA8A89D6" authorityName="Jeannel" authorityYear="1937" box="[1247,1338,691,717]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
.
</paragraph>
<paragraph id="8D4F36F48B367376FF2DD3C4FA9C889A" blockId="4.[151,1437,150,1833]" pageId="4" pageNumber="155">
<emphasis id="BF84EAE68B367376FF2DD3C4FE6E89EA" box="[199,478,727,753]" italics="true" pageId="4" pageNumber="155">Labiomaxillary complex.</emphasis>
The members of 
<taxonomicName id="4AF04D778B367376FD71D3C4FCFB89EA" authorityName="Sokolov" authorityYear="2023" box="[667,843,727,753]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
have setae of undifferentiated length on the basistipes (plesiomorphic state).This distinguishes them from other representatives of 
<taxonomicName id="4AF04D778B367376FC33D3E8FB83880E" authorityName="Jeannel" authorityYear="1937" box="[985,1075,763,789]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, which have the medial proximal seta on basistipes much longer than the others (derived state) either in the basal position (members of 
<taxonomicName id="4AF04D778B367376FB1ED20CFA288822" box="[1268,1432,799,825]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="162" phylum="Arthropoda" rank="subTribe" subTribe="Typhlocharina">Typhlocharina</taxonomicName>
, cf. 
<figureCitation id="15CB2A718B367376FF51D250FF4E8846" box="[187,254,835,861]" captionStart="FIGURE3" captionStartId="7.[199,298,1291,1315]" captionTargetBox="[151,1436,181,1267]" captionTargetId="figure-15@7.[151,1436,181,1267]" captionTargetPageId="7" captionText="FIGURE3. SEM illustrationsof structuralfeatures of pronota ofAnillini:A–B—subtribe Nesamblyopina;A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus alleni Sokolov and Carlton; D—Anillinus forthoodensis Sokolov andReddell;E—Serranillusjeanneli Barr; F—Geocharidius balini Sokolov and Kavanaugh; G—Zeanillus punctigerus (Broun); H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus nunni Sokolov. Legend: ast—setae of anterior margin of pronotum, bst—setae of basal margin of pronotum. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184231" httpUri="https://zenodo.org/record/10184231/files/figure.png" pageId="4" pageNumber="155">Fig. 3</figureCitation>
, p. 
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<figureCitation id="15CB2A718B367376FEC2D250FEF48846" box="[296,324,835,861]" captionStart="FIGURE 12" captionStartId="26.[151,250,1215,1239]" captionTargetBox="[157,1430,181,1192]" captionTargetId="figure-32@26.[157,1430,181,1192]" captionTargetPageId="26" captionText="FIGURE 12. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. magnificus (Allison Reserve, Akatore, Dunedin, SO), G—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), H—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), I—N. ovipennis (Mount Arthur, Nelson, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184251" httpUri="https://zenodo.org/record/10184251/files/figure.png" pageId="4" pageNumber="155">12</figureCitation>
in
</quantity>
<bibRefCitation id="E9614B058B367376FE82D250FD358846" author="Serrano, A. R. &amp; Aguiar, C. A. S." box="[360,645,835,861]" pageId="4" pageNumber="155" pagination="9 - 18" refId="ref27254" refString="Serrano, A. R. &amp; Aguiar, C. A. S. (2006) Two new species of Typhlocharis Dieck, 1869 of the silvanoides group from Portugal (Coleoptera, Carabidae). Animal Biodiversity and Conservation, 29.1, 9 - 18. https: // doi. org / 10.32800 / abc. 2006.29.0009" type="book chapter" year="2006">Serrano &amp; Aguiar (2006)</bibRefCitation>
; 
<figureCitation id="15CB2A718B367376FD7BD250FCB08846" box="[657,768,835,861]" captionStart="FIGURE 2" captionStartId="6.[151,250,1585,1609]" captionTargetBox="[189,1397,181,1561]" captionTargetId="figure-32@6.[189,1397,181,1561]" captionTargetPageId="6" captionText="FIGURE 2. SEM illustrations of structural features of labial complex of Anillini: A–B—subtribe Nesamblyopina; A— Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Zeanillus punctigerus (Broun); D—Pelodiaetodes insularis Sokolov; E—Anillinus forthoodensis Sokolov and Reddell; F—Geocharidius minimus Sokolov and Kavanaugh. Legend: ca—cardo, bs—basistipes, bss—basal stipital setae, mss—medial stipital setae, pf—palpifer. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184228" httpUri="https://zenodo.org/record/10184228/files/figure.png" pageId="4" pageNumber="155">Fig. 2d, p</figureCitation>
. 
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<bibRefCitation id="E9614B058B367376FCA1D250FB8B8846" author="Andujar, C. &amp; Lencina, J. L. &amp; Serrano, J." box="[843,1083,835,861]" pageId="4" pageNumber="155" pagination="35 - 44" refId="ref25847" refString="Andujar, C., Lencina, J. L. &amp; Serrano, J. (2008) Typhlocharis Dieck, 1869 (Coleoptera: Carabidae, Anillini): a new species from the Iberian Peninsula, with notes about its relationships and the evolution of the diecki species group. Zootaxa, 1842 (1), 35 - 44." type="journal article" year="2008">
Andujar 
<emphasis id="BF84EAE68B367376FC44D250FC568846" box="[942,998,835,861]" italics="true" pageId="4" pageNumber="155">et al.</emphasis>
(2008)
</bibRefCitation>
; 
<figureCitation id="15CB2A718B367376FBADD250FB068846" box="[1095,1206,835,861]" captionStart="FIGURE 2" captionStartId="6.[151,250,1585,1609]" captionTargetBox="[189,1397,181,1561]" captionTargetId="figure-32@6.[189,1397,181,1561]" captionTargetPageId="6" captionText="FIGURE 2. SEM illustrations of structural features of labial complex of Anillini: A–B—subtribe Nesamblyopina; A— Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Zeanillus punctigerus (Broun); D—Pelodiaetodes insularis Sokolov; E—Anillinus forthoodensis Sokolov and Reddell; F—Geocharidius minimus Sokolov and Kavanaugh. Legend: ca—cardo, bs—basistipes, bss—basal stipital setae, mss—medial stipital setae, pf—palpifer. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184228" httpUri="https://zenodo.org/record/10184228/files/figure.png" pageId="4" pageNumber="155">Fig. 2b, p</figureCitation>
. 
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<bibRefCitation id="E9614B058B367376FAEBD250FF5B889A" author="Andujar, C. &amp; Grebennikov, V. V. &amp; Lencina, J. L. &amp; Serrano, J." pageId="4" pageNumber="155" pagination="47 - 57" refId="ref25923" refString="Andujar, C., Grebennikov, V. V., Lencina, J. L. &amp; Serrano, J. (2010) A new Typhlocharis (Coleoptera: Carabidae: Anillina) from Spain: combining adult and larval morphological data with DNA information. Zootaxa, 2485 (1), 47 - 57. https: // doi. org / 10.11646 / zootaxa. 2485.1.4" type="journal article" year="2010">
Andujar 
<emphasis id="BF84EAE68B367376FA8ED250FA2D8846" box="[1380,1437,835,861]" italics="true" pageId="4" pageNumber="155">et al.</emphasis>
(2010))
</bibRefCitation>
or closer to the middle of sclerite (members of 
<taxonomicName id="4AF04D778B367376FD15D274FCEB889A" box="[767,859,871,897]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="subTribe" subTribe="Anillina">Anillina</taxonomicName>
and 
<taxonomicName id="4AF04D778B367376FC7BD274FB98889A" box="[913,1064,871,897]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="subTribe" subTribe="Scotodipnina">Scotodipnina</taxonomicName>
, as in 
<figureCitation id="15CB2A718B367376FB84D274FB54889A" box="[1134,1252,871,897]" captionStart="FIGURE 2" captionStartId="6.[151,250,1585,1609]" captionTargetBox="[189,1397,181,1561]" captionTargetId="figure-32@6.[189,1397,181,1561]" captionTargetPageId="6" captionText="FIGURE 2. SEM illustrations of structural features of labial complex of Anillini: A–B—subtribe Nesamblyopina; A— Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Zeanillus punctigerus (Broun); D—Pelodiaetodes insularis Sokolov; E—Anillinus forthoodensis Sokolov and Reddell; F—Geocharidius minimus Sokolov and Kavanaugh. Legend: ca—cardo, bs—basistipes, bss—basal stipital setae, mss—medial stipital setae, pf—palpifer. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184228" httpUri="https://zenodo.org/record/10184228/files/figure.png" pageId="4" pageNumber="155">Figs 2C–F</figureCitation>
, mss).
</paragraph>
<paragraph id="8D4F36F48B367376FF2DD298FABC8E2A" blockId="4.[151,1437,150,1833]" pageId="4" pageNumber="155">
<emphasis id="BF84EAE68B367376FF2DD298FE8E88BF" box="[199,318,907,932]" italics="true" pageId="4" pageNumber="155">Pronotum.</emphasis>
The vast majority of anilline taxa have a pronotum with basolateral denticles and posterior angles with varying degrees of prominence (cf. anilline fauna of 
<collectingCountry id="F5E776648B367376FCC3D2BCFCC888D2" box="[809,888,943,969]" name="Greece" pageId="4" pageNumber="155">Greece</collectingCountry>
in 
<bibRefCitation id="E9614B058B367376FC74D2BCFB0888D2" author="Giachino, P. M. &amp; Vailati, D." box="[926,1208,943,969]" pageId="4" pageNumber="155" pagination="1 - 112" refId="ref26446" refString="Giachino, P. M. &amp; Vailati, D. (2011) Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini). Biodiversity Journal, Monograph 1, 1 - 112. [http: // www. biodiversityjournal. com / pdf / Monogr 1. pdf]" type="journal article" year="2011">Giachino &amp; Vailati 2011</bibRefCitation>
). Rounded posterior angles are rare among anillines and have been reported for several genera only. Within its distributional range, the members of 
<taxonomicName id="4AF04D778B367376FEB9D2E4FDB38F0A" authorityName="Sokolov" authorityYear="2023" box="[339,515,1015,1041]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
share the state of rounded posterior pronotal angles with their geographical and ecological neighbours, the anilline species of the genus 
<taxonomicName id="4AF04D778B367376FCF2D508FC6D8F2E" authority="Jeannel" authorityName="Jeannel" authorityYear="1937" box="[792,989,1051,1077]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FCF2D508FC308F2E" box="[792,896,1051,1077]" italics="true" pageId="4" pageNumber="155">Zeanillus</emphasis>
Jeannel
</taxonomicName>
(cf. 
<figureCitation id="15CB2A718B367376FBFFD508FB5E8F2E" box="[1045,1262,1051,1077]" captionStart="FIGURE3" captionStartId="7.[199,298,1291,1315]" captionTargetBox="[151,1436,181,1267]" captionTargetId="figure-15@7.[151,1436,181,1267]" captionTargetPageId="7" captionText="FIGURE3. SEM illustrationsof structuralfeatures of pronota ofAnillini:A–B—subtribe Nesamblyopina;A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus alleni Sokolov and Carlton; D—Anillinus forthoodensis Sokolov andReddell;E—Serranillusjeanneli Barr; F—Geocharidius balini Sokolov and Kavanaugh; G—Zeanillus punctigerus (Broun); H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus nunni Sokolov. Legend: ast—setae of anterior margin of pronotum, bst—setae of basal margin of pronotum. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184231" httpUri="https://zenodo.org/record/10184231/files/figure.png" pageId="4" pageNumber="155">Figs 3A–B and 3G</figureCitation>
). Outside 
<collectingCountry id="F5E776648B367376FA8DD508FF448F42" name="New Zealand" pageId="4" pageNumber="155">New Zealand</collectingCountry>
, the members of 
<taxonomicName id="4AF04D778B367376FE53D52CFDD98F42" authorityName="Sokolov" authorityYear="2023" box="[441,617,1087,1113]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
share the state of rounded posterior angles with the following genera of the Old World: cavernicolous European 
<taxonomicName id="4AF04D778B367376FD8BD570FABD8F66" authority="Espanol &amp; Comas (Ortuno &amp; Sendra 2007)" authorityName="Espanol &amp; Comas (Ortuno &amp; Sendra" authorityYear="2007" box="[609,1293,1123,1149]" class="Insecta" family="Carabidae" genus="Aphaenotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FD8BD570FCA58F66" box="[609,789,1123,1149]" italics="true" pageId="4" pageNumber="155">Aphaenotyphlus</emphasis>
Español &amp; Comas (
<bibRefCitation id="E9614B058B367376FC15D570FAB58F66" author="Ortuno, V. M. &amp; Sendra, A." box="[1023,1285,1123,1149]" pageId="4" pageNumber="155" pagination="297 - 310" refId="ref26995" refString="Ortuno, V. M. &amp; Sendra, A. (2007) Taxonomie, systematique et biologie d'un Anillini troglobie exceptionnel (Coleoptera: Carbidae: Trechinae): Aphaenotyphlus alegrei Espanol &amp; Comas 1985. Annales de la Societe Entomologique de France, Nouvelle Serie, 43, 297 - 310. https: // doi. org / 10.1080 / 00379271.2007.10697525" type="journal article" year="2007">Ortuno &amp; Sendra 2007</bibRefCitation>
)
</taxonomicName>
and a group of African and Madagascan genera with shortened elytra, such as 
<taxonomicName id="4AF04D778B367376FC4ED594FB3E8FBA" authority="Giachino" authorityName="Giachino" authorityYear="2008" box="[932,1166,1159,1185]" class="Insecta" family="Carabidae" genus="Bulirschia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FC4ED594FBA98FBA" box="[932,1049,1159,1185]" italics="true" pageId="4" pageNumber="155">Bulirschia</emphasis>
Giachino
</taxonomicName>
, 
<taxonomicName id="4AF04D778B367376FB77D594FA288FBA" authority="Jeannel" authorityName="Jeannel" authorityYear="1963" box="[1181,1432,1159,1185]" class="Insecta" family="Carabidae" genus="Caeconannus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FB77D594FA858FBA" box="[1181,1333,1159,1185]" italics="true" pageId="4" pageNumber="155">Caeconannus</emphasis>
Jeannel
</taxonomicName>
, 
<taxonomicName id="4AF04D778B367376FF7DD5B8FEC48FDE" authority="Jeannel" authorityName="Jeannel" authorityYear="1950" box="[151,372,1195,1221]" class="Insecta" family="Carabidae" genus="Cryptorites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FF7DD5B8FEA78FDE" box="[151,279,1195,1221]" italics="true" pageId="4" pageNumber="155">Cryptorites</emphasis>
Jeannel
</taxonomicName>
, and 
<taxonomicName id="4AF04D778B367376FE47D5B8FD148FDE" authority="Jeannel" authorityName="Jeannel" authorityYear="1957" box="[429,676,1195,1221]" class="Insecta" family="Carabidae" genus="Microdipnites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FE47D5B8FDF98FDE" box="[429,585,1195,1221]" italics="true" pageId="4" pageNumber="155">Microdipnites</emphasis>
Jeannel
</taxonomicName>
(
<bibRefCitation id="E9614B058B367376FD5ED5B8FCEC8FDE" author="Giachino, P. M." box="[692,860,1195,1221]" pageId="4" pageNumber="155" pagination="91 - 136" refId="ref26347" refString="Giachino, P. M. (2008) New genera and species of Anillina (Coleoptera: Carabidae: Bembidiini) from Madagascar and Seychelles Islands, with notes about their origin and distributions. Bollettino del Museo Civico di Storia Naturale di Verona, 32 (Botanica Zoologia), 91 - 136." type="journal article" year="2008">Giachino 2008</bibRefCitation>
, 
<bibRefCitation id="E9614B058B367376FC8CD5B9FC138FDE" author="Giachino, P. M." box="[870,931,1194,1221]" pageId="4" pageNumber="155" pagination="15 - 31" refId="ref26403" refString="Giachino, P. M. (2015) New Anillina from South Africa, Tanzania, Madagascar and Seychelles Islands. Fragmenta Entomologica, 47, 15 - 31. https: // doi. org / 10.4081 / fe. 2015.130" type="journal article" year="2015">2015</bibRefCitation>
). Interestingly, none of the species of 
<taxonomicName id="4AF04D778B367376FAAFD5B8FA2C8FDE" authorityName="Jeannel" authorityYear="1937" box="[1349,1436,1195,1221]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
with rounded posterior angles of the pronotum have been documented in the New World to date. It seems that all cases of similarity in the shape of the posterior angles between 
<taxonomicName id="4AF04D778B367376FC48D5E0FBE28E16" authorityName="Sokolov" authorityYear="2023" box="[930,1106,1267,1293]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
and Old World anillines are homoplastic, since this state of character arose in different phyletic lineages and subtribes of the tribe.
</paragraph>
<paragraph id="8D4F36F48B367376FF2DD428FBA48C32" blockId="4.[151,1437,150,1833]" pageId="4" pageNumber="155">
Another pronotal character, the setation of the basal margin of the pronotum, is also widely distributed among the representatives of 
<taxonomicName id="4AF04D778B367376FE7BD44CFE5C8E62" authorityName="Jeannel" authorityYear="1937" box="[401,492,1375,1401]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
. Like the situation with the state of posterior pronotal angles, cases where setation is lacking are not common among the species of the tribe. Noteworthy, the members of 
<taxonomicName id="4AF04D778B367376FB95D490FA9F8E86" authorityName="Sokolov" authorityYear="2023" box="[1151,1327,1411,1437]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
share the lack of setation on the basal margin of the pronotum with the North American lineage of 
<taxonomicName id="4AF04D778B367376FB44D4B4FAB88EDA" authorityName="Jeannel" authorityYear="1937" box="[1198,1288,1447,1473]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, specifically, 
<taxonomicName id="4AF04D778B367376FF7DD4D8FEFB8EFE" authority="Casey" authorityName="Casey" authorityYear="1918" box="[151,331,1483,1509]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FF7DD4D8FF4C8EFE" box="[151,252,1483,1509]" italics="true" pageId="4" pageNumber="155">Anillinus</emphasis>
Casey
</taxonomicName>
, 
<taxonomicName id="4AF04D778B367376FEB3D4D8FDA48EFE" authority="Barr" authorityName="Barr" box="[345,532,1483,1509]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FEB3D4D8FE658EFE" box="[345,469,1483,1509]" italics="true" pageId="4" pageNumber="155">Serranillus</emphasis>
Barr
</taxonomicName>
, 
<taxonomicName id="4AF04D778B367376FDCBD4D8FD5C8EFE" authority="Jeannel" authorityName="Jeannel" authorityYear="1963" box="[545,748,1483,1509]" class="Insecta" family="Carabidae" genus="Anillodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FDCBD4D8FD3B8EFE" box="[545,651,1483,1509]" italics="true" pageId="4" pageNumber="155">Anillodes</emphasis>
Jeannel
</taxonomicName>
, 
<taxonomicName id="4AF04D778B367376FD13D4D8FF558D12" authority="Jeannel, and Medusapyga LaBonte &amp; Maddison (2023)" authorityName="Jeannel, and Medusapyga LaBonte &amp; Maddison" authorityYear="2023" class="Insecta" family="Carabidae" genus="Anillaspis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FD13D4D8FCD98EFE" box="[761,873,1483,1509]" italics="true" pageId="4" pageNumber="155">Anillaspis</emphasis>
Jeannel, and 
<emphasis id="BF84EAE68B367376FBE0D4D8FB298EFE" box="[1034,1177,1483,1509]" italics="true" pageId="4" pageNumber="155">Medusapyga</emphasis>
<bibRefCitation id="E9614B058B367376FB48D4D8FF558D12" author="LaBonte, J. R. &amp; Maddison, D. R." pageId="4" pageNumber="155" pagination="401 - 432" refId="ref26638" refString="LaBonte, J. R. &amp; Maddison, D. R. (2023) Medusapyga LaBonte and Maddison, a new genus of Anillini (Coleopetra: Carabidae: Trechinae) from the Pacific Northwest of the United States. Proceedings of the California Academy of Sciences, Series 4, 67, 401 - 432." type="journal article" year="2023">LaBonte &amp; Maddison (2023)</bibRefCitation>
</taxonomicName>
. Recent results of molecular analysis have placed all these North American genera in a separate clade on the phylogenetic tree of 
<taxonomicName id="4AF04D778B367376FE68D700FE698D36" authorityName="Jeannel" authorityYear="1937" box="[386,473,1555,1581]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
right above the 
<taxonomicName id="4AF04D778B367376FD7DD700FC9C8D36" authorityName="Jeannel" authorityYear="1937" box="[663,812,1555,1581]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FD7DD700FC9C8D36" box="[663,812,1555,1581]" italics="true" pageId="4" pageNumber="155">Nesamblyops</emphasis>
</taxonomicName>
clade, i.e., as a sister clade to all other 
<taxonomicName id="4AF04D778B367376FB19D700FAFA8D36" authorityName="Jeannel" authorityYear="1937" box="[1267,1354,1555,1581]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
except 
<taxonomicName id="4AF04D778B367376FF7DD724FE9C8D4A" authorityName="Jeannel" authorityYear="1937" box="[151,300,1591,1617]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B367376FF7DD724FE9C8D4A" box="[151,300,1591,1617]" italics="true" pageId="4" pageNumber="155">Nesamblyops</emphasis>
</taxonomicName>
(
<bibRefCitation id="E9614B058B367376FED6D724FDAF8D4A" author="Andujar, C. &amp; Faille, A. &amp; Perez-Gonzalez, S. &amp; Zaballos, J. P. &amp; Vogler, A. P. &amp; Ribera, I." box="[316,543,1591,1617]" pageId="4" pageNumber="155" pagination="235 - 246" refId="ref25995" refString="Andujar, C., Faille, A., Perez-Gonzalez, S., Zaballos, J. P., Vogler, A. P. &amp; Ribera, I. (2016) Gondwanian relicts and oceanic dispersal in a cosmopolitan radiation of euedaphic ground beetles. Molecular Phylogenetics and Evolution, 99, 235 - 246. https: // doi. org / 10.1016 / j. ympev. 2016.03.013" type="journal article" year="2016">
Andujar 
<emphasis id="BF84EAE68B367376FE48D724FE6B8D4A" box="[418,475,1591,1617]" italics="true" pageId="4" pageNumber="155">et al.</emphasis>
2016
</bibRefCitation>
; 
<bibRefCitation id="E9614B058B367376FDC1D724FC928D4A" author="Maddison, D. R. &amp; Kanda, K. &amp; Boyd, O. F. &amp; Faille, A. &amp; Porch, N. &amp; Erwin, T. L. &amp; Roig-Junent, S." box="[555,802,1591,1617]" pageId="4" pageNumber="155" pagination="151 - 176" refId="ref26793" refString="Maddison, D. R., Kanda, K., Boyd, O. F., Faille, A., Porch, N., Erwin, T. L. &amp; Roig-Junent, S. (2019) Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): Unexpected clades, isolated lineages, and morphological convergence. Molecular Phylogenetics and Evolution, 132, 151 - 176. https: // doi. org / 10.1016 / j. ympev. 2018.11.006." type="journal article" year="2019">
Maddison 
<emphasis id="BF84EAE68B367376FD49D724FD6D8D4A" box="[675,733,1591,1617]" italics="true" pageId="4" pageNumber="155">et al.</emphasis>
2019
</bibRefCitation>
). To date, the lack of setation of the basal margin of the pronotum is the only known morphological trait allowing researchers to distinguish the members of the North American 
<taxonomicName id="4AF04D778B367376FEE7D76CFED48D82" authorityName="Jeannel" authorityYear="1937" box="[269,356,1663,1689]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
from other members of the tribe. In accordance with the basal positions of both clades on the tree, such state of setation can be treated as a symplesiomorphy for both clades. Unfortunately, the taxon coverage in the abovementioned molecular analyses of 
<taxonomicName id="4AF04D778B367376FD70D7D4FD418DFA" authorityName="Jeannel" authorityYear="1937" box="[666,753,1735,1761]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
is far from perfect, given that only 17 genera were analyzed from almost 100 described to date. Therefore, clade positions on the tree and their composition might be subject to change in the future after additional material is examined in molecular analysis.
</paragraph>
<caption id="D98F667C8B377377FF7DD47FFF768D23" ID-DOI="http://doi.org/10.5281/zenodo.10184226" ID-Zenodo-Dep="10184226" httpUri="https://zenodo.org/record/10184226/files/figure.png" pageId="5" pageNumber="156" startId="5.[151,250,1388,1412]" targetBox="[151,1436,179,1362]" targetPageId="5" targetType="figure">
<paragraph id="8D4F36F48B377377FF7DD47FFF768D23" blockId="5.[151,1437,1388,1592]" pageId="5" pageNumber="156">
<emphasis id="BF84EAE68B377377FF7DD47FFEBF8E9F" bold="true" box="[151,271,1388,1412]" pageId="5" pageNumber="156">FIGURE 1</emphasis>
. SEM illustrations of structural features of heads of 
<taxonomicName id="4AF04D778B377377FCDDD47FFC3A8E9F" authorityName="Jeannel" authorityYear="1937" box="[823,906,1388,1412]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
: 
<emphasis id="BF84EAE68B377377FC7CD47FFC778E9F" bold="true" box="[918,967,1388,1412]" pageId="5" pageNumber="156">A–C</emphasis>
—subtribe 
<taxonomicName id="4AF04D778B377377FBD1D47FFB6D8E9F" authorityName="Sokolov" authorityYear="2023" box="[1083,1245,1388,1412]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
; 
<emphasis id="BF84EAE68B377377FB03D47FFB4B8E9F" bold="true" box="[1257,1275,1388,1412]" pageId="5" pageNumber="156">A</emphasis>
— 
<taxonomicName id="4AF04D778B377377FAFFD47FFEF98EB3" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B377377FAFFD47FFF5F8EB3" italics="true" pageId="5" pageNumber="156">Nesamblyops oreobius</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B377377FEBED483FED58EB3" bold="true" box="[340,357,1424,1448]" pageId="5" pageNumber="156">B</emphasis>
— 
<taxonomicName id="4AF04D778B377377FE95D483FD7B8EB3" authority="(Sharp)" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[383,715,1424,1448]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B377377FE95D483FDC58EB3" box="[383,629,1424,1448]" italics="true" pageId="5" pageNumber="156">Nesamblyops subcaecus</emphasis>
(Sharp)
</taxonomicName>
; 
<emphasis id="BF84EAE68B377377FD3CD483FD588EB3" bold="true" box="[726,744,1424,1448]" pageId="5" pageNumber="156">C</emphasis>
— 
<emphasis id="BF84EAE68B377377FCE8D483FBB68EB3" box="[770,1030,1424,1448]" italics="true" pageId="5" pageNumber="156">
<taxonomicName id="4AF04D778B377377FCE8D483FBB28EB3" authority="Sokolov, 2023" authorityName="Sokolov" authorityYear="2023" box="[770,1026,1424,1448]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="magnificus" status="sp. nov.">Nesamblyops magnificus</taxonomicName>
,
</emphasis>
<emphasis id="BF84EAE68B377377FBE6D483FB8F8EB3" bold="true" box="[1036,1087,1424,1448]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A4B7579D8B377377FBE6D483FB8F8EB3" box="[1036,1087,1424,1448]" pageId="5" pageNumber="156" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
; 
<emphasis id="BF84EAE68B377377FBA7D483FBEF8EB3" bold="true" box="[1101,1119,1424,1448]" pageId="5" pageNumber="156">D</emphasis>
— 
<taxonomicName id="4AF04D778B377377FB93D483FF568ED7" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="alleni">
<emphasis id="BF84EAE68B377377FB93D483FAA58EB3" box="[1145,1301,1424,1448]" italics="true" pageId="5" pageNumber="156">Anillinus alleni</emphasis>
Sokolov and Carlton
</taxonomicName>
; 
<emphasis id="BF84EAE68B377377FF18D4A7FEB38ED7" bold="true" box="[242,259,1460,1484]" pageId="5" pageNumber="156">E</emphasis>
— 
<taxonomicName id="4AF04D778B377377FEF6D4A7FD648ED7" authority="Sokolov and Reddell" authorityName="Sokolov and Reddell" box="[284,724,1460,1484]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="acutipennis">
<emphasis id="BF84EAE68B377377FEF6D4A7FE448ED7" box="[284,500,1460,1484]" italics="true" pageId="5" pageNumber="156">Anillinus acutipennis</emphasis>
Sokolov and Reddell
</taxonomicName>
; 
<emphasis id="BF84EAE68B377377FD35D4A7FD5E8ED7" bold="true" box="[735,750,1460,1484]" pageId="5" pageNumber="156">F</emphasis>
— 
<taxonomicName id="4AF04D778B377377FCE2D4A7FAB68ED7" authority="Sokolov and Kavanaugh" authorityName="Sokolov and Kavanaugh" box="[776,1286,1460,1484]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="jalapensis">
<emphasis id="BF84EAE68B377377FCE2D4A7FBB08ED7" box="[776,1024,1460,1484]" italics="true" pageId="5" pageNumber="156">Geocharidius jalapensis</emphasis>
Sokolov and Kavanaugh
</taxonomicName>
; 
<emphasis id="BF84EAE68B377377FAFBD4A7FA958ED7" bold="true" box="[1297,1317,1460,1484]" pageId="5" pageNumber="156">G</emphasis>
— 
<taxonomicName id="4AF04D778B377377FAD4D4A7FEDB8EEB" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1914" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="BF84EAE68B377377FAD4D4A7FEBF8EEB" italics="true" pageId="5" pageNumber="156">Zeanillus punctigerus</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B377377FE93D4CBFE3D8EEB" bold="true" box="[377,397,1496,1520]" pageId="5" pageNumber="156">H</emphasis>
— 
<taxonomicName id="4AF04D778B377377FE4CD4CBFD6A8EEB" authority="Sokolov" authorityName="Sokolov" box="[422,730,1496,1520]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B377377FE4CD4CBFDC98EEB" box="[422,633,1496,1520]" italics="true" pageId="5" pageNumber="156">Pelodiaetodes nunni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B377377FD0DD4CBFD418EEB" bold="true" box="[743,753,1496,1520]" pageId="5" pageNumber="156">I</emphasis>
— 
<taxonomicName id="4AF04D778B377377FCE0D4CBFB768EEB" authority="Jeannel. Legend" authorityName="Jeannel. Legend" box="[778,1222,1496,1520]" class="Insecta" family="Carabidae" genus="Pelodiaetus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="sulcatipennis">
<emphasis id="BF84EAE68B377377FCE0D4CBFBA18EEB" box="[778,1041,1496,1520]" italics="true" pageId="5" pageNumber="156">Pelodiaetus sulcatipennis</emphasis>
Jeannel. Legend
</taxonomicName>
: cl—clypaeus, lse— lateral clypeal setula, mse—medial clypeal setula, om—ommatidium, trom—trace of disappeared ommatidium. Scale = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8D4F36F48B377377FF2DD799FB808C43" blockId="5.[151,1437,1674,2025]" pageId="5" pageNumber="156">
<emphasis id="BF84EAE68B377377FF2DD799FEE58DBF" box="[199,341,1674,1700]" italics="true" pageId="5" pageNumber="156">Pterothorax.</emphasis>
The structure of the pterothorax of the members of 
<taxonomicName id="4AF04D778B377377FC6DD799FB878DBF" authorityName="Sokolov" authorityYear="2023" box="[903,1079,1674,1700]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
differs significantly from that of other 
<taxonomicName id="4AF04D778B377377FF3CD7BDFE9D8DD3" authorityName="Jeannel" authorityYear="1937" box="[214,301,1710,1736]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
in many aspects. For instance, mesothoracic anapleural sutures can be observed in all representatives of the tribe on microscopic slides in translucent light. However, external traces of sutures are obliterated (apomorphic state) and cannot be seen on specimens in direct light in all anillines in which the mesothorax configuration is documented (cf. 
<figureCitation id="15CB2A718B377377FEB8D609FE718C2F" box="[338,449,1818,1844]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="5" pageNumber="156">Figs 4C–I</figureCitation>
with 
<figureCitation id="15CB2A718B377377FE14D609FDCA8C2F" box="[510,634,1818,1844]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="5" pageNumber="156">Figs 4A–B</figureCitation>
, msts). At present, the members of 
<taxonomicName id="4AF04D778B377377FC15D609FB1F8C2F" authorityName="Sokolov" authorityYear="2023" box="[1023,1199,1818,1844]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
are the only anillines that have externally visible mesothoracic anapleural sutures (plesiomorphic state).
</paragraph>
<paragraph id="8D4F36F48B377377FF2DD671FCE38CF3" blockId="5.[151,1437,1674,2025]" pageId="5" pageNumber="156">
An entirely similar situation exists with the metaventral process. All anilline species in which the metaventrite configuration is documented possess an unmargined metaventral process, whereas the members of 
<taxonomicName id="4AF04D778B377377FB06D695FA2C8CBB" authorityName="Sokolov" authorityYear="2023" box="[1260,1436,1926,1952]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
have a metaventral process with a distinct and narrow margination (cf. 
<figureCitation id="15CB2A718B377377FC78D6B9FBB98CDF" box="[914,1033,1962,1988]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="5" pageNumber="156">Figs 4A–B</figureCitation>
with 
<figureCitation id="15CB2A718B377377FBA9D6B9FB018CDF" box="[1091,1201,1962,1988]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="5" pageNumber="156">Figs 4C–I</figureCitation>
, mtp). Unfortunately, polarity for this character cannot be determined with certainty.
</paragraph>
<caption id="D98F667C8B347374FF7DD722FAF98DAF" ID-DOI="http://doi.org/10.5281/zenodo.10184228" ID-Zenodo-Dep="10184228" httpUri="https://zenodo.org/record/10184228/files/figure.png" pageId="6" pageNumber="157" startId="6.[151,250,1585,1609]" targetBox="[189,1397,181,1561]" targetPageId="6" targetType="figure">
<paragraph id="8D4F36F48B347374FF7DD722FAF98DAF" blockId="6.[151,1437,1585,1717]" pageId="6" pageNumber="157">
<emphasis id="BF84EAE68B347374FF7DD722FEA28D52" bold="true" box="[151,274,1585,1609]" pageId="6" pageNumber="157">FIGURE 2</emphasis>
. SEM illustrations of structural features of labial complex of 
<taxonomicName id="4AF04D778B347374FC5DD722FBBA8D52" authorityName="Jeannel" authorityYear="1937" box="[951,1034,1585,1609]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
: 
<emphasis id="BF84EAE68B347374FBF2D722FBF88D52" bold="true" box="[1048,1096,1585,1609]" pageId="6" pageNumber="157">A–B</emphasis>
—subtribe 
<taxonomicName id="4AF04D778B347374FB55D722FAD18D52" authorityName="Sokolov" authorityYear="2023" box="[1215,1377,1585,1609]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
; 
<emphasis id="BF84EAE68B347374FA9BD722FA338D52" bold="true" box="[1393,1411,1585,1609]" pageId="6" pageNumber="157">A</emphasis>
— 
<taxonomicName id="4AF04D778B347374FF7DD746FE6A8D76" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[151,474,1621,1645]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B347374FF7DD746FECF8D76" box="[151,383,1621,1645]" italics="true" pageId="6" pageNumber="157">Nesamblyops oreobius</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B347374FE0DD746FE488D76" bold="true" box="[487,504,1621,1645]" pageId="6" pageNumber="157">B</emphasis>
— 
<taxonomicName id="4AF04D778B347374FDF8D746FCD18D76" authority="(Sharp)" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[530,865,1621,1645]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B347374FDF8D746FCBA8D76" box="[530,778,1621,1645]" italics="true" pageId="6" pageNumber="157">Nesamblyops subcaecus</emphasis>
(Sharp)
</taxonomicName>
; 
<emphasis id="BF84EAE68B347374FC84D746FC308D76" bold="true" box="[878,896,1621,1645]" pageId="6" pageNumber="157">C</emphasis>
— 
<taxonomicName id="4AF04D778B347374FC70D746FB648D76" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1914" box="[922,1236,1621,1645]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="BF84EAE68B347374FC70D746FBC98D76" box="[922,1145,1621,1645]" italics="true" pageId="6" pageNumber="157">Zeanillus punctigerus</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B347374FB0BD746FB438D76" bold="true" box="[1249,1267,1621,1645]" pageId="6" pageNumber="157">D</emphasis>
— 
<taxonomicName id="4AF04D778B347374FAE7D746FEE38D8A" authority="Sokolov" authorityName="Sokolov" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="BF84EAE68B347374FAE7D746FF408D8A" italics="true" pageId="6" pageNumber="157">Pelodiaetodes insularis</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B347374FE8BD76AFEC28D8A" bold="true" box="[353,370,1657,1681]" pageId="6" pageNumber="157">E</emphasis>
— 
<taxonomicName id="4AF04D778B347374FE61D76AFCD78D8A" authority="Sokolov and Reddell" authorityName="Sokolov and Reddell" box="[395,871,1657,1681]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="forthoodensis">
<emphasis id="BF84EAE68B347374FE61D76AFDCC8D8A" box="[395,636,1657,1681]" italics="true" pageId="6" pageNumber="157">Anillinus forthoodensis</emphasis>
Sokolov and Reddell
</taxonomicName>
; 
<emphasis id="BF84EAE68B347374FC9CD76AFC358D8A" bold="true" box="[886,901,1657,1681]" pageId="6" pageNumber="157">F</emphasis>
— 
<taxonomicName id="4AF04D778B347374FC74D76AFF578DAE" authority="Sokolov and Kavanaugh. Legend" authorityName="Sokolov and Kavanaugh. Legend" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="BF84EAE68B347374FC74D76AFB398D8A" box="[926,1161,1657,1681]" italics="true" pageId="6" pageNumber="157">Geocharidius minimus</emphasis>
Sokolov and Kavanaugh. Legend
</taxonomicName>
: ca—cardo, bs—basistipes, bss—basal stipital setae, mss—medial stipital setae, pf—palpifer. Scale = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8D4F36F48B347375FF2DD7F0FC6C8D90" blockId="6.[151,1437,1763,2042]" lastBlockId="7.[151,1437,1505,2035]" lastPageId="7" lastPageNumber="158" pageId="6" pageNumber="157">
The members of the tribe 
<taxonomicName id="4AF04D778B347374FE07D7F0FDF48DE6" authorityName="Jeannel" authorityYear="1937" box="[493,580,1763,1789]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
are flightless species, and it is clear that different parts of their pterothorax may have undergone different degrees of reduction over the course of their evolution. For 
<taxonomicName id="4AF04D778B347374FB52D614FAA28C3A" authorityName="Jeannel" authorityYear="1937" box="[1208,1298,1799,1825]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, this is true in terms of the length of the metaventrite and the shape of the metendosternite. In our case, the members of 
<taxonomicName id="4AF04D778B347374FF7DD65CFEF78C72" authorityName="Sokolov" authorityYear="2023" box="[151,327,1871,1897]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
possess the shortest metaventrite among the species of 
<taxonomicName id="4AF04D778B347374FC46D65CFBB38C72" authorityName="Jeannel" authorityYear="1937" box="[940,1027,1871,1897]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
(cf. 
<figureCitation id="15CB2A718B347374FBDFD65CFB1E8C72" box="[1077,1198,1871,1897]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="6" pageNumber="157">Figs 4A–B</figureCitation>
with 
<figureCitation id="15CB2A718B347374FB06D65CFAED8C72" box="[1260,1373,1871,1897]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="6" pageNumber="157">Figs 4C–I</figureCitation>
, mtv) in which the configuration of metaventrite is documented. Given that many taxa among 
<taxonomicName id="4AF04D778B347374FB67D660FB548C96" authorityName="Jeannel" authorityYear="1937" box="[1165,1252,1907,1933]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
exhibit reduced elytra, which sometimes can reach only half the length of the abdomen, it is reasonable to expect that species with very short metaventrite may be discovered in the future among other anillines also. Additionally, the members of 
<taxonomicName id="4AF04D778B347374FF7DD6CCFEF78CE2" authorityName="Sokolov" authorityYear="2023" box="[151,327,2015,2041]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
possess reduced configuration of metendoventrite without distinct lateral arms. They share the lack of lateral arms with the species of several genera whose structures of metendosternites are documented, like 
<taxonomicName id="4AF04D778B357375FF7DD716FF4C8D04" authorityName="Casey" authorityYear="1918" box="[151,252,1541,1567]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B357375FF7DD716FF4C8D04" box="[151,252,1541,1567]" italics="true" pageId="7" pageNumber="158">Anillinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4AF04D778B357375FEE1D716FE378D04" box="[267,391,1541,1567]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B357375FEE1D716FE378D04" box="[267,391,1541,1567]" italics="true" pageId="7" pageNumber="158">Serranillus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4AF04D778B357375FE2DD716FD0C8D04" authority="Jeannel" authorityName="Jeannel" authorityYear="1963" box="[455,700,1541,1567]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B357375FE2DD716FDEF8D04" box="[455,607,1541,1567]" italics="true" pageId="7" pageNumber="158">Geocharidius</emphasis>
Jeannel
</taxonomicName>
(cf. 
<figureCitation id="15CB2A718B357375FD18D716FCDE8D3B" box="[754,878,1541,1568]" captionStart="FIGURE 5" captionStartId="9.[151,250,1796,1820]" captionTargetBox="[185,1401,181,1771]" captionTargetId="figure-15@9.[185,1401,181,1772]" captionTargetPageId="9" captionText="FIGURE 5. Digital illustrations of structural features of pterothorax of Anillini, ventral aspect: A–D—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Nesamblyops brouni Sokolov; D—Nesamblyops viator n.sp.; E—Anillinus langdoni Sokolov and Carlton; F—Anillinus Cherokee Sokolov and Carlton; G—Geocharidius longinoi Sokolov and Kavanaugh; H—Geocharidius disjunctus Sokolov and Kavanaugh; I—Zapotecanillus longinoi Sokolov; J—Zeanillus pellucidus Sokolov; K—Zeanillus pallidus (Broun); L—Zeanillus nunni Sokolov; M—Pelodiaetodes insularis Sokolov; N—Pelodiaetodes nunni Sokolov; O—Pelodiaetodes moorei Sokolov; P—Pelodiaetus sulcatipennis Jeannel.Legend: mtes—metendosternite. Scales intentionally omitted." figureDoi="http://doi.org/10.5281/zenodo.10184235" httpUri="https://zenodo.org/record/10184235/files/figure.png" pageId="7" pageNumber="158">Figs 5A–D</figureCitation>
with 
<figureCitation id="15CB2A718B357375FC5AD716FB9C8D3B" box="[944,1068,1541,1568]" captionStart="FIGURE 5" captionStartId="9.[151,250,1796,1820]" captionTargetBox="[185,1401,181,1771]" captionTargetId="figure-15@9.[185,1401,181,1772]" captionTargetPageId="9" captionText="FIGURE 5. Digital illustrations of structural features of pterothorax of Anillini, ventral aspect: A–D—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Nesamblyops brouni Sokolov; D—Nesamblyops viator n.sp.; E—Anillinus langdoni Sokolov and Carlton; F—Anillinus Cherokee Sokolov and Carlton; G—Geocharidius longinoi Sokolov and Kavanaugh; H—Geocharidius disjunctus Sokolov and Kavanaugh; I—Zapotecanillus longinoi Sokolov; J—Zeanillus pellucidus Sokolov; K—Zeanillus pallidus (Broun); L—Zeanillus nunni Sokolov; M—Pelodiaetodes insularis Sokolov; N—Pelodiaetodes nunni Sokolov; O—Pelodiaetodes moorei Sokolov; P—Pelodiaetus sulcatipennis Jeannel.Legend: mtes—metendosternite. Scales intentionally omitted." figureDoi="http://doi.org/10.5281/zenodo.10184235" httpUri="https://zenodo.org/record/10184235/files/figure.png" pageId="7" pageNumber="158">Figs 5E–H</figureCitation>
, mtes, and both with the species that have lateral arms as in 
<figureCitation id="15CB2A718B357375FE34D73AFDFD8D5F" box="[478,589,1577,1604]" captionStart="FIGURE 5" captionStartId="9.[151,250,1796,1820]" captionTargetBox="[185,1401,181,1771]" captionTargetId="figure-15@9.[185,1401,181,1772]" captionTargetPageId="9" captionText="FIGURE 5. Digital illustrations of structural features of pterothorax of Anillini, ventral aspect: A–D—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Nesamblyops brouni Sokolov; D—Nesamblyops viator n.sp.; E—Anillinus langdoni Sokolov and Carlton; F—Anillinus Cherokee Sokolov and Carlton; G—Geocharidius longinoi Sokolov and Kavanaugh; H—Geocharidius disjunctus Sokolov and Kavanaugh; I—Zapotecanillus longinoi Sokolov; J—Zeanillus pellucidus Sokolov; K—Zeanillus pallidus (Broun); L—Zeanillus nunni Sokolov; M—Pelodiaetodes insularis Sokolov; N—Pelodiaetodes nunni Sokolov; O—Pelodiaetodes moorei Sokolov; P—Pelodiaetus sulcatipennis Jeannel.Legend: mtes—metendosternite. Scales intentionally omitted." figureDoi="http://doi.org/10.5281/zenodo.10184235" httpUri="https://zenodo.org/record/10184235/files/figure.png" pageId="7" pageNumber="158">Figs 5I–P</figureCitation>
, mtes). Together with the fact that 
<taxonomicName id="4AF04D778B357375FC00D73AFBFF8D58" authorityName="Casey" authorityYear="1918" box="[1002,1103,1577,1603]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B357375FC00D73AFBFF8D58" box="[1002,1103,1577,1603]" italics="true" pageId="7" pageNumber="158">Anillinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4AF04D778B357375FB67D73AFA958D58" authorityName="Jeannel" authorityYear="1963" box="[1165,1317,1577,1603]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B357375FB67D73AFA958D58" box="[1165,1317,1577,1603]" italics="true" pageId="7" pageNumber="158">Geocharidius</emphasis>
</taxonomicName>
belong to different clades on the phylogenetic tree of 
<taxonomicName id="4AF04D778B357375FD6FD75EFBB38D7C" authority="(Maddison et al. 2019)" baseAuthorityName="Maddison" baseAuthorityYear="2019" box="[645,1027,1613,1639]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="subFamily" subFamily="Trechinae">
Trechinae (
<bibRefCitation id="E9614B058B357375FCEED75EFC4B8D7C" author="Maddison, D. R. &amp; Kanda, K. &amp; Boyd, O. F. &amp; Faille, A. &amp; Porch, N. &amp; Erwin, T. L. &amp; Roig-Junent, S." box="[772,1019,1613,1639]" pageId="7" pageNumber="158" pagination="151 - 176" refId="ref26793" refString="Maddison, D. R., Kanda, K., Boyd, O. F., Faille, A., Porch, N., Erwin, T. L. &amp; Roig-Junent, S. (2019) Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): Unexpected clades, isolated lineages, and morphological convergence. Molecular Phylogenetics and Evolution, 132, 151 - 176. https: // doi. org / 10.1016 / j. ympev. 2018.11.006." type="journal article" year="2019">
Maddison 
<emphasis id="BF84EAE68B357375FC94D75DFC078D7C" box="[894,951,1613,1639]" italics="true" pageId="7" pageNumber="158">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
, in both cases states of metaventrite and metendosternite in 
<taxonomicName id="4AF04D778B357375FE77D762FDFD8D90" authorityName="Sokolov" authorityYear="2023" box="[413,589,1649,1675]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
may be considered as homoplastic.
</paragraph>
<caption id="D98F667C8B357375FF2DD418FD6C8EA8" ID-DOI="http://doi.org/10.5281/zenodo.10184231" ID-Zenodo-Dep="10184231" httpUri="https://zenodo.org/record/10184231/files/figure.png" pageId="7" pageNumber="158" startId="7.[199,298,1291,1315]" targetBox="[151,1436,181,1267]" targetPageId="7" targetType="figure">
<paragraph id="8D4F36F48B357375FF2DD418FD6C8EA8" blockId="7.[151,1437,1291,1459]" pageId="7" pageNumber="158">
<emphasis id="BF84EAE68B357375FF2DD418FE8A8E38" bold="true" box="[199,314,1291,1315]" pageId="7" pageNumber="158">FIGURE 3</emphasis>
. SEM illustrations of structural features of pronota of 
<taxonomicName id="4AF04D778B357375FCA2D418FC2B8E38" authorityName="Jeannel" authorityYear="1937" box="[840,923,1291,1315]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
: 
<emphasis id="BF84EAE68B357375FC4BD418FC618E38" bold="true" box="[929,977,1291,1315]" pageId="7" pageNumber="158">A–B</emphasis>
—subtribe 
<taxonomicName id="4AF04D778B357375FBAAD418FB528E38" authorityName="Sokolov" authorityYear="2023" box="[1088,1250,1291,1315]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
; 
<emphasis id="BF84EAE68B357375FB03D418FB4B8E38" bold="true" box="[1257,1275,1291,1315]" pageId="7" pageNumber="158">A</emphasis>
— 
<taxonomicName id="4AF04D778B357375FAFFD418FEF98E5C" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B357375FAFFD418FF5F8E5C" italics="true" pageId="7" pageNumber="158">Nesamblyops oreobius</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B357375FEBED43CFED58E5C" bold="true" box="[340,357,1327,1351]" pageId="7" pageNumber="158">B</emphasis>
— 
<taxonomicName id="4AF04D778B357375FE94D43CFD7A8E5C" authority="(Sharp)" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[382,714,1327,1351]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B357375FE94D43CFDC48E5C" box="[382,628,1327,1351]" italics="true" pageId="7" pageNumber="158">Nesamblyops subcaecus</emphasis>
(Sharp)
</taxonomicName>
; 
<emphasis id="BF84EAE68B357375FD3FD43CFD578E5C" bold="true" box="[725,743,1327,1351]" pageId="7" pageNumber="158">C</emphasis>
— 
<taxonomicName id="4AF04D778B357375FCEBD43CFBC98E5C" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[769,1145,1327,1351]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="alleni">
<emphasis id="BF84EAE68B357375FCEBD43CFC2D8E5C" box="[769,925,1327,1351]" italics="true" pageId="7" pageNumber="158">Anillinus alleni</emphasis>
Sokolov and Carlton
</taxonomicName>
; 
<emphasis id="BF84EAE68B357375FB6ED43CFB268E5C" bold="true" box="[1156,1174,1327,1351]" pageId="7" pageNumber="158">D</emphasis>
— 
<taxonomicName id="4AF04D778B357375FB5AD43CFEDA8E70" authority="Sokolov and Reddell" authorityName="Sokolov and Reddell" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="forthoodensis">
<emphasis id="BF84EAE68B357375FB5AD43CFA2C8E5C" box="[1200,1436,1327,1351]" italics="true" pageId="7" pageNumber="158">Anillinus forthoodensis</emphasis>
Sokolov and Reddell
</taxonomicName>
; 
<emphasis id="BF84EAE68B357375FE98D440FE338E70" bold="true" box="[370,387,1363,1387]" pageId="7" pageNumber="158">E</emphasis>
— 
<taxonomicName id="4AF04D778B357375FE77D440FD278E70" authority="Barr" authorityName="Barr" authorityYear="1995" box="[413,663,1363,1387]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="BF84EAE68B357375FE77D440FDD38E70" box="[413,611,1363,1387]" italics="true" pageId="7" pageNumber="158">Serranillus jeanneli</emphasis>
Barr
</taxonomicName>
; 
<emphasis id="BF84EAE68B357375FD75D440FD1E8E70" bold="true" box="[671,686,1363,1387]" pageId="7" pageNumber="158">F</emphasis>
— 
<taxonomicName id="4AF04D778B357375FD22D440FB3E8E70" authority="Sokolov and Kavanaugh" authorityName="Sokolov and Kavanaugh" box="[712,1166,1363,1387]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="balini">
<emphasis id="BF84EAE68B357375FD22D440FC218E70" box="[712,913,1363,1387]" italics="true" pageId="7" pageNumber="158">Geocharidius balini</emphasis>
Sokolov and Kavanaugh
</taxonomicName>
; 
<emphasis id="BF84EAE68B357375FB7FD440FB198E70" bold="true" box="[1173,1193,1363,1387]" pageId="7" pageNumber="158">G</emphasis>
— 
<taxonomicName id="4AF04D778B357375FB28D440FF5A8E94" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1914" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="BF84EAE68B357375FB28D440FA2D8E70" box="[1218,1437,1363,1387]" italics="true" pageId="7" pageNumber="158">Zeanillus punctigerus</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B357375FF1ED464FEB88E94" bold="true" box="[244,264,1399,1423]" pageId="7" pageNumber="158">H</emphasis>
— 
<taxonomicName id="4AF04D778B357375FECBD464FDFE8E94" authority="Sokolov" authorityName="Sokolov" box="[289,590,1399,1423]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B357375FECBD464FE408E94" box="[289,496,1399,1423]" italics="true" pageId="7" pageNumber="158">Pelodiaetodes nunni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B357375FDBDD464FDD18E94" bold="true" box="[599,609,1399,1423]" pageId="7" pageNumber="158">I</emphasis>
— 
<taxonomicName id="4AF04D778B357375FD90D464FC568E94" authority="Sokolov. Legend" authorityName="Sokolov. Legend" box="[634,998,1399,1423]" class="Insecta" family="Carabidae" genus="Pelodiaetus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B357375FD90D464FC818E94" box="[634,817,1399,1423]" italics="true" pageId="7" pageNumber="158">Pelodiaetus nunni</emphasis>
Sokolov. Legend
</taxonomicName>
: ast—setae of anterior margin of pronotum, bst—setae of basal margin of pronotum. Scale = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8D4F36F48B357375FF2DD786FEF18CE8" blockId="7.[151,1437,1505,2035]" pageId="7" pageNumber="158">
<emphasis id="BF84EAE68B357375FF2DD786FE2C8DB4" box="[199,412,1685,1711]" italics="true" pageId="7" pageNumber="158">Elytral chaetotaxy.</emphasis>
The members of 
<taxonomicName id="4AF04D778B357375FD8ED786FCA48DB4" authorityName="Sokolov" authorityYear="2023" box="[612,788,1685,1711]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
have eight pores in the elytral umbilical series, while the vast majority of the species of 
<taxonomicName id="4AF04D778B357375FE03D7AAFDF08DC8" authorityName="Jeannel" authorityYear="1937" box="[489,576,1721,1747]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
have elytral umbilical series with nine pores (all species of subtribes 
<taxonomicName id="4AF04D778B357375FAAAD7AAFA2C8DC8" box="[1344,1436,1721,1747]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="subTribe" subTribe="Anillina">Anillina</taxonomicName>
and 
<taxonomicName id="4AF04D778B357375FF23D7CEFEED8DEC" box="[201,349,1757,1783]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="subTribe" subTribe="Scotodipnina">Scotodipnina</taxonomicName>
with fully developed elytra). Nevertheless, the eight-pore configuration in the umbilical series can be found in some members of the subtribe 
<taxonomicName id="4AF04D778B357375FD54D612FCEF8C00" box="[702,863,1793,1819]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="subTribe" subTribe="Typhlocharina">Typhlocharina</taxonomicName>
(e.g., cf. 
<figureCitation id="15CB2A718B357375FC38D612FBE18C00" box="[978,1105,1793,1819]" captionStart="FIGURE 6" captionStartId="10.[151,250,1867,1891]" captionTargetBox="[189,1398,181,1843]" captionTargetId="figure-32@10.[189,1398,181,1843]" captionTargetPageId="10" captionText="FIGURE 6. SEM illustrations of structural features of elytra of Anillini, left lateral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus carolinae Casey; D—Anillinus alleni Sokolov and Carlton; E—Serranillus donovani (Jeannel); F—Geocharidius jalapensis Sokolov and Kavanaugh; G—Zeanillus punctigerus (Broun); H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus nunni Sokolov. Legend: ed2—scutellar seta; ed6— discal seta, ed8—apical seta; eo1…eo9—setae 1–9 of umbilical series of pores. Scale = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184237" httpUri="https://zenodo.org/record/10184237/files/figure.png" pageId="7" pageNumber="158">Figs 6A–B</figureCitation>
, with the umbilical series of 
<taxonomicName id="4AF04D778B357375FF7DD636FE998C24" authorityName="Dieck" authorityYear="1869" box="[151,297,1829,1855]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B357375FF7DD636FE998C24" box="[151,297,1829,1855]" italics="true" pageId="7" pageNumber="158">Typhlocharis</emphasis>
</taxonomicName>
[now 
<taxonomicName id="4AF04D778B357375FE9AD636FE478C24" box="[368,503,1829,1855]" class="Insecta" family="Carabidae" genus="Lusotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B357375FE9AD636FE478C24" box="[368,503,1829,1855]" italics="true" pageId="7" pageNumber="158">Lusotyphlus</emphasis>
</taxonomicName>
] 
<emphasis id="BF84EAE68B357375FDE2D636FD398C24" box="[520,649,1829,1855]" italics="true" pageId="7" pageNumber="158">algarvensis</emphasis>
Coiffait on 
<figureCitation id="15CB2A718B357375FCFBD636FCE28C24" box="[785,850,1829,1855]" captionStart="FIGURE 1" captionStartId="5.[151,250,1388,1412]" captionTargetBox="[151,1436,179,1362]" captionTargetId="figure-15@5.[151,1436,179,1362]" captionTargetPageId="5" captionText="FIGURE 1. SEM illustrations of structural features of heads of Anillini: A–C—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Nesamblyops magnificus, n.sp.; D—Anillinus alleni Sokolov and Carlton; E—Anillinus acutipennis Sokolov and Reddell; F—Geocharidius jalapensis Sokolov and Kavanaugh; G—Zeanillus punctigerus (Broun); H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: cl—clypaeus, lse— lateral clypeal setula, mse—medial clypeal setula, om—ommatidium, trom—trace of disappeared ommatidium. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184226" httpUri="https://zenodo.org/record/10184226/files/figure.png" pageId="7" pageNumber="158">Fig. 1</figureCitation>
, p. 151, or with the umbilical series of 
<taxonomicName id="4AF04D778B357375FAE1D636FEEE8C78" authority="Dieck" authorityName="Dieck" authorityYear="1869" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="silvanoides">
<emphasis id="BF84EAE68B357375FAE1D636FEA58C78" italics="true" pageId="7" pageNumber="158">Typhlocharis silvanoides</emphasis>
Dieck
</taxonomicName>
on 
<figureCitation id="15CB2A718B357375FE6CD65AFE798C78" box="[390,457,1865,1891]" captionStart="FIGURE 2" captionStartId="6.[151,250,1585,1609]" captionTargetBox="[189,1397,181,1561]" captionTargetId="figure-32@6.[189,1397,181,1561]" captionTargetPageId="6" captionText="FIGURE 2. SEM illustrations of structural features of labial complex of Anillini: A–B—subtribe Nesamblyopina; A— Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Zeanillus punctigerus (Broun); D—Pelodiaetodes insularis Sokolov; E—Anillinus forthoodensis Sokolov and Reddell; F—Geocharidius minimus Sokolov and Kavanaugh. Legend: ca—cardo, bs—basistipes, bss—basal stipital setae, mss—medial stipital setae, pf—palpifer. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184228" httpUri="https://zenodo.org/record/10184228/files/figure.png" pageId="7" pageNumber="158">Fig. 2</figureCitation>
, p. 
<quantity id="4A089B118B357375FE05D65AFD858C78" box="[495,565,1865,1891]" metricMagnitude="0" metricUnit="m" metricValue="3.5814" pageId="7" pageNumber="158" unit="in" value="141.0">141 in</quantity>
<bibRefCitation id="E9614B058B357375FDD1D65AFC078C78" author="Zaballos, J. P. &amp; Perez-Gonzales, S." box="[571,951,1865,1891]" pageId="7" pageNumber="158" pagination="221 - 232" refId="ref27788" refString="Zaballos, J. P. &amp; Perez-Gonzales, S. (2010) Typhlocharis vicariantes del estrecho de Gibraltar. I: T. armata Coiffait, 1969 (Coleoptera, Caraboidea, Trechidae). Graellsia, 66, 221 - 232." type="journal article" year="2010">Zaballos &amp; Perez-Gonzales (2010</bibRefCitation>
, 
<bibRefCitation id="E9614B058B357375FC2BD65AFC4A8C78" author="Zaballos, J. P. &amp; Perez-Gonzales, S." box="[961,1018,1865,1891]" pageId="7" pageNumber="158" pagination="135 - 149" refId="ref27833" refString="Zaballos, J. P. &amp; Perez-Gonzales, S. (2011) Typhlocharis vicariantes del estrecho de Gibraltar. II: T. silvanoides Dieck, 1869 (Coleoptera, Caraboidea, Trechidae). Graellsia, 67, 135 - 149. https: // doi. org / 10.3989 / graellsia. 2011. v 67.041" type="journal article" year="2011">2011</bibRefCitation>
) respectively). It is supposed that this configuration of the umbilical series of pores in 
<taxonomicName id="4AF04D778B357375FD5CD67EFCE78C9C" box="[694,855,1901,1927]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="162" phylum="Arthropoda" rank="subTribe" subTribe="Typhlocharina">Typhlocharina</taxonomicName>
arose as a result of the adaptations to the endogean way of life (
<bibRefCitation id="E9614B058B357375FEF4D682FE1F8CB0" author="Jeanne, C." box="[286,431,1937,1963]" pageId="7" pageNumber="158" pagination="83 - 102" refId="ref26537" refString="Jeanne, C. (1973) Sur la classification des Bembidiides endoges de la region euro-mediterraneenne (Col. Carabidae, Bembidiinae, Anillini). Nouvelle Revue d'Entomologie, 3, 83 - 102." type="journal article" year="1973">Jeanne 1973</bibRefCitation>
). However, the majority of species of 
<taxonomicName id="4AF04D778B357375FCBDD682FBB78CB0" authorityName="Sokolov" authorityYear="2023" box="[855,1031,1937,1963]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
are litter-dwellers, so similar elytral chaetotaxy in these two subtribes presumably evolved independently, and it is likely, in this case, that the resemblance is homoplastic.
</paragraph>
<caption id="D98F667C8B3A737AFF7DD5C4FC268EB8" ID-DOI="http://doi.org/10.5281/zenodo.10184233" ID-Zenodo-Dep="10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="8" pageNumber="159" startId="8.[151,250,1239,1263]" targetBox="[151,1437,181,1215]" targetPageId="8" targetType="figure">
<paragraph id="8D4F36F48B3A737AFF7DD5C4FC268EB8" blockId="8.[151,1437,1239,1444]" pageId="8" pageNumber="159">
<emphasis id="BF84EAE68B3A737AFF7DD5C4FEBD8FF4" bold="true" box="[151,269,1239,1263]" pageId="8" pageNumber="159">FIGURE 4</emphasis>
. SEM illustrations of structural features of pterothorax of 
<taxonomicName id="4AF04D778B3A737AFCB5D5CBFC008FEB" authorityName="Jeannel" authorityYear="1937" box="[863,944,1240,1264]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, ventral aspect: 
<emphasis id="BF84EAE68B3A737AFBBFD5C4FB358FEB" bold="true" box="[1109,1157,1239,1264]" pageId="8" pageNumber="159">A–B</emphasis>
—subtribe 
<taxonomicName id="4AF04D778B3A737AFB1CD5CBFA288FEB" authorityName="Sokolov" authorityYear="2023" box="[1270,1432,1240,1264]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
; 
<emphasis id="BF84EAE68B3A737AFF7DD5E8FF198E08" bold="true" box="[151,169,1275,1299]" pageId="8" pageNumber="159">A</emphasis>
— 
<taxonomicName id="4AF04D778B3A737AFF29D5EFFDB78E0F" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[195,519,1276,1300]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B3A737AFF29D5EFFE1B8E0F" box="[195,427,1276,1300]" italics="true" pageId="8" pageNumber="159">Nesamblyops oreobius</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B3A737AFDFED5EFFD958E0F" bold="true" box="[532,549,1276,1300]" pageId="8" pageNumber="159">B</emphasis>
— 
<taxonomicName id="4AF04D778B3A737AFDD5D5EFFC3F8E0F" authority="(Sharp)" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[575,911,1276,1300]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B3A737AFDD5D5EFFC878E0F" box="[575,823,1276,1300]" italics="true" pageId="8" pageNumber="159">Nesamblyops subcaecus</emphasis>
(Sharp)
</taxonomicName>
; 
<emphasis id="BF84EAE68B3A737AFC76D5E8FC1E8E08" bold="true" box="[924,942,1275,1299]" pageId="8" pageNumber="159">C</emphasis>
— 
<taxonomicName id="4AF04D778B3A737AFC22D5EFFAD38E0F" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[968,1379,1276,1300]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="BF84EAE68B3A737AFC22D5EFFB308E0F" box="[968,1152,1276,1300]" italics="true" pageId="8" pageNumber="159">Anillinus lescheni</emphasis>
Sokolov and Carlton
</taxonomicName>
; 
<emphasis id="BF84EAE68B3A737AFA9BD5EFFA338E0F" bold="true" box="[1393,1411,1276,1300]" pageId="8" pageNumber="159">D</emphasis>
— 
<taxonomicName id="4AF04D778B3A737AFF7DD433FDDE8E23" authority="Sokolov and Reddell" authorityName="Sokolov and Reddell" box="[151,622,1312,1336]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="forthoodensis">
<emphasis id="BF84EAE68B3A737AFF7DD433FE368E23" box="[151,390,1312,1336]" italics="true" pageId="8" pageNumber="159">Anillinus forthoodensis</emphasis>
Sokolov and Reddell
</taxonomicName>
; 
<emphasis id="BF84EAE68B3A737AFD96D433FD3D8E23" bold="true" box="[636,653,1312,1336]" pageId="8" pageNumber="159">E</emphasis>
— 
<taxonomicName id="4AF04D778B3A737AFD4CD433FB1D8E23" authority="Sokolov and Kavanaugh" authorityName="Sokolov and Kavanaugh" box="[678,1197,1312,1336]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="jalapensis">
<emphasis id="BF84EAE68B3A737AFD4CD433FC108E23" box="[678,928,1312,1336]" italics="true" pageId="8" pageNumber="159">Geocharidius jalapensis</emphasis>
Sokolov and Kavanaugh
</taxonomicName>
; 
<emphasis id="BF84EAE68B3A737AFB50D433FB798E23" bold="true" box="[1210,1225,1312,1336]" pageId="8" pageNumber="159">F</emphasis>
— 
<taxonomicName id="4AF04D778B3A737AFB09D433FF5A8E47" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1884" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="BF84EAE68B3A737AFB09D433FA2D8E23" box="[1251,1437,1312,1336]" italics="true" pageId="8" pageNumber="159">Zeanillus pallidus</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B3A737AFF16D450FEA08E40" bold="true" box="[252,272,1347,1371]" pageId="8" pageNumber="159">G</emphasis>
— 
<taxonomicName id="4AF04D778B3A737AFEC3D457FD348E47" authority="Sokolov" authorityName="Sokolov" box="[297,644,1348,1372]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="BF84EAE68B3A737AFEC3D457FDAF8E47" box="[297,543,1348,1372]" italics="true" pageId="8" pageNumber="159">Pelodiaetodes insularis</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3A737AFD7FD457FD198E47" bold="true" box="[661,681,1348,1372]" pageId="8" pageNumber="159">H</emphasis>
— 
<taxonomicName id="4AF04D778B3A737AFD28D457FC4E8E47" authority="Sokolov" authorityName="Sokolov" box="[706,1022,1348,1372]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B3A737AFD28D457FC298E40" box="[706,921,1348,1372]" italics="true" pageId="8" pageNumber="159">Pelodiaetodes nunni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3A737AFBE5D457FBA98E47" bold="true" box="[1039,1049,1348,1372]" pageId="8" pageNumber="159">I</emphasis>
— 
<taxonomicName id="4AF04D778B3A737AFBD8D457FF578E9B" authority="Jeannel. Legend" authorityName="Jeannel. Legend" class="Insecta" family="Carabidae" genus="Pelodiaetus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="sulcatipennis">
<emphasis id="BF84EAE68B3A737AFBD8D457FA8D8E47" box="[1074,1341,1348,1372]" italics="true" pageId="8" pageNumber="159">Pelodiaetus sulcatipennis</emphasis>
Jeannel. Legend
</taxonomicName>
: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8D4F36F48B3A737AFF2DD4C1FCAD8DF3" blockId="8.[151,1437,1490,2020]" pageId="8" pageNumber="159">
<emphasis id="BF84EAE68B3A737AFF2DD4C1FE0A8EF7" box="[199,442,1490,1516]" italics="true" pageId="8" pageNumber="159">Structure of protibia.</emphasis>
The members of 
<taxonomicName id="4AF04D778B3A737AFD66D4C1FC8C8EF7" authorityName="Sokolov" authorityYear="2023" box="[652,828,1490,1516]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
exhibit an obliquely sloped apicolateral edge of the protibia. On the posterior surface of the protibia, there is a row of 4–6 setae (
<figureCitation id="15CB2A718B3A737AFBE3D4E5FB398D0B" box="[1033,1161,1526,1552]" captionStart="FIGURE 7" captionStartId="11.[151,250,1452,1476]" captionTargetBox="[168,1419,201,1429]" captionTargetId="figure-15@11.[168,1419,201,1429]" captionTargetPageId="11" captionText="FIGURE 7. SEM illustrations of structural features of protibia of Anillini, dorsal (E) and ventral (A–D, F–H) aspects: A– B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D—Geocharidius comayaguanus Sokolov and Kavanaugh; E—Geocharidius zullini Vigna Taglianti; F—Zeanillus brouni Sokolov; G—Pelodiaetodes nunni Sokolov; H—Pelodiaetus sulcatipennis Jeannel. Legend: as—articulosetae; psr—posterior setal row. Scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10184239" httpUri="https://zenodo.org/record/10184239/files/figure.png" pageId="8" pageNumber="159">Figs 7A–B</figureCitation>
, psr) that are positioned adjacent to the edge and do not protrude beyond the slope edge (plesiomorphic state). This row is equivalent to the “distal cluster” of Hlavac (p. 52, 
<figureCitation id="15CB2A718B3A737AFDE0D72DFDFC8D43" box="[522,588,1598,1624]" captionStart="FIGURE3" captionStartId="7.[199,298,1291,1315]" captionTargetBox="[151,1436,181,1267]" captionTargetId="figure-15@7.[151,1436,181,1267]" captionTargetPageId="7" captionText="FIGURE3. SEM illustrationsof structuralfeatures of pronota ofAnillini:A–B—subtribe Nesamblyopina;A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus alleni Sokolov and Carlton; D—Anillinus forthoodensis Sokolov andReddell;E—Serranillusjeanneli Barr; F—Geocharidius balini Sokolov and Kavanaugh; G—Zeanillus punctigerus (Broun); H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus nunni Sokolov. Legend: ast—setae of anterior margin of pronotum, bst—setae of basal margin of pronotum. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184231" httpUri="https://zenodo.org/record/10184231/files/figure.png" pageId="8" pageNumber="159">Fig. 3</figureCitation>
, DC, abbreviations on p. 65 (1971) and the erroneously cited “distal comb” in 
<bibRefCitation id="E9614B058B3A737AFF5ED771FE078D67" author="Maddison, D. R. &amp; Kanda, K. &amp; Boyd, O. F. &amp; Faille, A. &amp; Porch, N. &amp; Erwin, T. L. &amp; Roig-Junent, S." box="[180,439,1634,1660]" pageId="8" pageNumber="159" pagination="151 - 176" refId="ref26793" refString="Maddison, D. R., Kanda, K., Boyd, O. F., Faille, A., Porch, N., Erwin, T. L. &amp; Roig-Junent, S. (2019) Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): Unexpected clades, isolated lineages, and morphological convergence. Molecular Phylogenetics and Evolution, 132, 151 - 176. https: // doi. org / 10.1016 / j. ympev. 2018.11.006." type="journal article" year="2019">
Maddison 
<emphasis id="BF84EAE68B3A737AFEC1D770FED38D67" box="[299,355,1634,1660]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
(2019)
</bibRefCitation>
. All other anillines with a documented protibial structure have an apicolateral edge of the protibia with a notch that varies in the degree of concavity. This notch is associated with a row of 9–15 setae on the posterior surface, which protrudes beyond the edge of concavity, thus forming a kind of “comb” on the protibial apex (derived state) (cf. 
<figureCitation id="15CB2A718B3A737AFE42D7DDFD938DF3" box="[424,547,1742,1768]" captionStart="FIGURE 7" captionStartId="11.[151,250,1452,1476]" captionTargetBox="[168,1419,201,1429]" captionTargetId="figure-15@11.[168,1419,201,1429]" captionTargetPageId="11" captionText="FIGURE 7. SEM illustrations of structural features of protibia of Anillini, dorsal (E) and ventral (A–D, F–H) aspects: A– B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D—Geocharidius comayaguanus Sokolov and Kavanaugh; E—Geocharidius zullini Vigna Taglianti; F—Zeanillus brouni Sokolov; G—Pelodiaetodes nunni Sokolov; H—Pelodiaetus sulcatipennis Jeannel. Legend: as—articulosetae; psr—posterior setal row. Scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10184239" httpUri="https://zenodo.org/record/10184239/files/figure.png" pageId="8" pageNumber="159">Figs 7C–H</figureCitation>
with 
<figureCitation id="15CB2A718B3A737AFD88D7DDFD6F8DF3" box="[610,735,1742,1768]" captionStart="FIGURE 7" captionStartId="11.[151,250,1452,1476]" captionTargetBox="[168,1419,201,1429]" captionTargetId="figure-15@11.[168,1419,201,1429]" captionTargetPageId="11" captionText="FIGURE 7. SEM illustrations of structural features of protibia of Anillini, dorsal (E) and ventral (A–D, F–H) aspects: A– B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D—Geocharidius comayaguanus Sokolov and Kavanaugh; E—Geocharidius zullini Vigna Taglianti; F—Zeanillus brouni Sokolov; G—Pelodiaetodes nunni Sokolov; H—Pelodiaetus sulcatipennis Jeannel. Legend: as—articulosetae; psr—posterior setal row. Scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10184239" httpUri="https://zenodo.org/record/10184239/files/figure.png" pageId="8" pageNumber="159">Figs 7A–B</figureCitation>
, psr).
</paragraph>
<paragraph id="8D4F36F48B3A737AFF2DD7E1FEFF8CFF" blockId="8.[151,1437,1490,2020]" pageId="8" pageNumber="159">
<emphasis id="BF84EAE68B3A737AFF2DD7E1FEC58C17" box="[199,373,1778,1804]" italics="true" pageId="8" pageNumber="159">Male genitalia.</emphasis>
The symmetry/equality of the basal lobes of the median lobe of males is considered to be a synapomorphy of 
<taxonomicName id="4AF04D778B3A737AFE8DD605FE728C2B" authorityName="Jeannel" authorityYear="1937" box="[359,450,1814,1840]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
. However, within the tribe, many taxa actually demonstrate unequal basal lobes. This character state may characterize either whole genera or a particular species (e.g., cf. images of male median lobe of 
<taxonomicName id="4AF04D778B3A737AFF5DD64DFE838C63" box="[183,307,1886,1912]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B3A737AFF5DD64DFE838C63" box="[183,307,1886,1912]" italics="true" pageId="8" pageNumber="159">Serranillus</emphasis>
</taxonomicName>
in 
<bibRefCitation id="E9614B058B3A737AFEB3D64DFD368C63" author="Sokolov, I. M. &amp; Carlton, C. E." box="[345,646,1886,1912]" pageId="8" pageNumber="159" pagination="245 - 249" refId="ref27479" refString="Sokolov, I. M. &amp; Carlton, C. E. (2012) Species of Anillinus Casey (Coleoptera: Carabidae: Trechinae) described from Brazil and their relation to North American representatives of the genus. The Coleopterists Bulletin, 66, 245 - 249. https: // doi. org / 10.1649 / 072.066.0310" type="journal article" year="2012">Sokolov &amp; Carlton (2012)</bibRefCitation>
, of 
<taxonomicName id="4AF04D778B3A737AFD59D64DFCFB8C63" authorityName="Jeannel" authorityYear="1963" box="[691,843,1886,1912]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B3A737AFD59D64DFCFB8C63" box="[691,843,1886,1912]" italics="true" pageId="8" pageNumber="159">Geocharidius</emphasis>
</taxonomicName>
in 
<bibRefCitation id="E9614B058B3A737AFC9BD64DFB7A8C63" author="Sokolov, I. M. &amp; Kavanaugh, D. H." box="[881,1226,1886,1912]" pageId="8" pageNumber="159" pagination="61 - 118" refId="ref27608" refString="Sokolov, I. M. &amp; Kavanaugh, D. H. (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys, 443, 61 - 118. https: // doi. org / 10.3897 / zookeys. 443.7880" type="journal article" year="2014">Sokolov &amp; Kavanaugh (2014)</bibRefCitation>
, of 
<taxonomicName id="4AF04D778B3A737AFB1CD64DFF448C87" authority="Sokolov" authorityName="Sokolov" authorityYear="2013" class="Insecta" family="Carabidae" genus="Zapotecanillus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B3A737AFB1CD64DFA2C8C63" box="[1270,1436,1886,1912]" italics="true" pageId="8" pageNumber="159">Zapotecanillus</emphasis>
Sokolov
</taxonomicName>
in 
<bibRefCitation id="E9614B058B3A737AFEFFD691FE758C87" author="Sokolov, I. M." box="[277,453,1922,1948]" pageId="8" pageNumber="159" pagination="51 - 92" refId="ref27353" refString="Sokolov, I. M. (2013) A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistics analysis and some notes on the evolution of the genus. ZooKeys, 352, 51 - 92. https: // doi. org / 10.3897 / zookeys. 352.6052" type="journal article" year="2013">Sokolov (2013)</bibRefCitation>
, of 
<taxonomicName id="4AF04D778B3A737AFE06D691FC308C87" authority="Sokolov &amp; Watrous" authorityName="Sokolov &amp; Watrous" authorityYear="2008" box="[492,896,1922,1948]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="aleyae">
<emphasis id="BF84EAE68B3A737AFE06D691FD108C87" box="[492,672,1922,1948]" italics="true" pageId="8" pageNumber="159">Anillinus aleyae</emphasis>
Sokolov &amp; Watrous
</taxonomicName>
in 
<bibRefCitation id="E9614B058B3A737AFC4BD691FB7D8C87" author="Sokolov, I. M. &amp; Watrous, L. E." box="[929,1229,1922,1948]" pageId="8" pageNumber="159" pagination="537 - 543" refId="ref27675" refString="Sokolov, I. M. &amp; Watrous, L. E. (2008) A new species and the first record of the genus Anillinus (Carabidae: Trechinae: Bembidiini) from the Ozark region. The Coleopterists Bulletin, 62, 537 - 543. https: // doi. org / 10.1649 / 1114.1" type="journal article" year="2008">Sokolov &amp; Watrous (2008)</bibRefCitation>
, and of 
<taxonomicName id="4AF04D778B3A737AFAC8D691FEDA8CDB" authority="Sokolov &amp; Carlton" authorityName="Sokolov &amp; Carlton" authorityYear="2004" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="robisoni">
<emphasis id="BF84EAE68B3A737AFAC8D691FA2C8C87" box="[1314,1436,1922,1948]" italics="true" pageId="8" pageNumber="159">A. robisoni</emphasis>
Sokolov &amp; Carlton
</taxonomicName>
in 
<bibRefCitation id="E9614B058B3A737AFE66D6B5FD308CDB" author="Sokolov, I. M. &amp; Carlton, C. E. &amp; Cornell, J. F." box="[396,640,1958,1984]" pageId="8" pageNumber="159" pagination="185 - 233" refId="ref27540" refString="Sokolov, I. M., Carlton, C. E. &amp; Cornell, J. F. (2004) Review of Anillinus with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Trechinae: Bembidiini). The Coleopterists Bulletin, 58, 185 - 233. https: // doi. org / 10.1649 / 611" type="journal article" year="2004">
Sokolov 
<emphasis id="BF84EAE68B3A737AFE05D6B4FD968CDB" box="[495,550,1958,1984]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
(2004))
</bibRefCitation>
and likely evolved independently in different taxa of 
<taxonomicName id="4AF04D778B3A737AFB24D6B5FA988CDB" authorityName="Jeannel" authorityYear="1937" box="[1230,1320,1958,1984]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, including 
<taxonomicName id="4AF04D778B3A737AFF7DD6D9FEFB8CFF" authorityName="Sokolov" authorityYear="2023" box="[151,331,1994,2020]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
.
</paragraph>
<caption id="D98F667C8B3B737BFF7DD617FD018CEF" ID-DOI="http://doi.org/10.5281/zenodo.10184235" ID-Zenodo-Dep="10184235" httpUri="https://zenodo.org/record/10184235/files/figure.png" pageId="9" pageNumber="160" startId="9.[151,250,1796,1820]" targetBox="[185,1401,181,1771]" targetPageId="9" targetType="figure">
<paragraph id="8D4F36F48B3B737BFF7DD617FD018CEF" blockId="9.[151,1437,1796,2036]" pageId="9" pageNumber="160">
<emphasis id="BF84EAE68B3B737BFF7DD617FEBC8C07" bold="true" box="[151,268,1796,1820]" pageId="9" pageNumber="160">FIGURE 5</emphasis>
. Digital illustrations of structural features of pterothorax of 
<taxonomicName id="4AF04D778B3B737BFC89D617FC048C07" authorityName="Jeannel" authorityYear="1937" box="[867,948,1796,1820]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, ventral aspect: 
<emphasis id="BF84EAE68B3B737BFBBFD617FB368C07" bold="true" box="[1109,1158,1796,1820]" pageId="9" pageNumber="160">A–D</emphasis>
—subtribe 
<taxonomicName id="4AF04D778B3B737BFB1CD617FA288C07" authorityName="Sokolov" authorityYear="2023" box="[1270,1432,1796,1820]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFF7DD63BFF198C5B" bold="true" box="[151,169,1832,1856]" pageId="9" pageNumber="160">A</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFF29D63BFDB08C5B" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[195,512,1832,1856]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B3B737BFF29D63BFE188C5B" box="[195,424,1832,1856]" italics="true" pageId="9" pageNumber="160">Nesamblyops oreobius</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFDE0D63BFDAB8C5B" bold="true" box="[522,539,1832,1856]" pageId="9" pageNumber="160">B</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFDDED63BFCCE8C5B" authority="(Sharp)" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[564,894,1832,1856]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B3B737BFDDED63BFC998C5B" box="[564,809,1832,1856]" italics="true" pageId="9" pageNumber="160">Nesamblyops subcaecus</emphasis>
(Sharp)
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFC62D63BFC2A8C5B" bold="true" box="[904,922,1832,1856]" pageId="9" pageNumber="160">C</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFC59D63BFB508C5B" authority="Sokolov" authorityName="Sokolov, B" authorityYear="2023" box="[947,1248,1832,1856]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="BF84EAE68B3B737BFC59D63BFB338C5B" box="[947,1155,1832,1856]" italics="true" pageId="9" pageNumber="160">Nesamblyops brouni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFB03D63BFB4B8C5B" bold="true" box="[1257,1275,1832,1856]" pageId="9" pageNumber="160">D</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFAFFD63BFF648C7F" authority="Sokolov, 2023" authorityName="Sokolov" authorityYear="2023" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="viator" status="sp. nov.">
<emphasis id="BF84EAE68B3B737BFAFFD63BFF648C7F" italics="true" pageId="9" pageNumber="160">Nesamblyops viator</emphasis>
</taxonomicName>
<emphasis id="BF84EAE68B3B737BFF34D65FFEA18C7F" bold="true" box="[222,273,1868,1892]" pageId="9" pageNumber="160">
<taxonomicNameLabel id="A4B7579D8B3B737BFF34D65FFEA18C7F" box="[222,273,1868,1892]" pageId="9" pageNumber="160" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
; 
<emphasis id="BF84EAE68B3B737BFEC8D65FFE838C7F" bold="true" box="[290,307,1868,1892]" pageId="9" pageNumber="160">E</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFEA6D65FFD438C7F" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[332,755,1868,1892]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="langdoni">
<emphasis id="BF84EAE68B3B737BFEA6D65FFDBC8C7F" box="[332,524,1868,1892]" italics="true" pageId="9" pageNumber="160">Anillinus langdoni</emphasis>
Sokolov and Carlton
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFCEBD65FFCA08C7F" bold="true" box="[769,784,1868,1892]" pageId="9" pageNumber="160">F</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFCC0D65FFB678C7F" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[810,1239,1868,1892]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="cherokee">
<emphasis id="BF84EAE68B3B737BFCC0D65FFC408C7F" box="[810,1008,1868,1892]" italics="true" pageId="9" pageNumber="160">Anillinus Cherokee</emphasis>
Sokolov and Carlton
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFB0FD65FFB498C7F" bold="true" box="[1253,1273,1868,1892]" pageId="9" pageNumber="160">G</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFAF8D65FFE5E8C93" authority="Sokolov and Kavanaugh" authorityName="Sokolov and Kavanaugh" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="longinoi">
<emphasis id="BF84EAE68B3B737BFAF8D65FFF5B8C93" italics="true" pageId="9" pageNumber="160">Geocharidius longinoi</emphasis>
Sokolov and Kavanaugh
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFE1DD663FDBB8C93" bold="true" box="[503,523,1904,1928]" pageId="9" pageNumber="160">H</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFDCED663FBAD8C93" authority="Sokolov and Kavanaugh" authorityName="Sokolov and Kavanaugh" box="[548,1053,1904,1928]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="disjunctus">
<emphasis id="BF84EAE68B3B737BFDCED663FCAB8C93" box="[548,795,1904,1928]" italics="true" pageId="9" pageNumber="160">Geocharidius disjunctus</emphasis>
Sokolov and Kavanaugh
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFBCCD663FB808C93" bold="true" box="[1062,1072,1904,1928]" pageId="9" pageNumber="160">I</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFBA0D663FA298C93" authority="Sokolov" authorityName="Sokolov" authorityYear="2013" box="[1098,1433,1904,1928]" class="Insecta" family="Carabidae" genus="Zapotecanillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="longinoi">
<emphasis id="BF84EAE68B3B737BFBA0D663FA8A8C93" box="[1098,1338,1904,1928]" italics="true" pageId="9" pageNumber="160">Zapotecanillus longinoi</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFF7DD687FF148CB7" bold="true" box="[151,164,1940,1964]" pageId="9" pageNumber="160">J</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFF57D687FE5C8CB7" authority="Sokolov" authorityName="Sokolov" authorityYear="2016" box="[189,492,1940,1964]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="BF84EAE68B3B737BFF57D687FE3C8CB7" box="[189,396,1940,1964]" italics="true" pageId="9" pageNumber="160">Zeanillus pellucidus</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFE12D687FDBC8CB7" bold="true" box="[504,524,1940,1964]" pageId="9" pageNumber="160">K</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFDCFD687FC888CB7" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1884" box="[549,824,1940,1964]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="BF84EAE68B3B737BFDCFD687FD6D8CB7" box="[549,733,1940,1964]" italics="true" pageId="9" pageNumber="160">Zeanillus pallidus</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFCAED687FCE58CB7" bold="true" box="[836,853,1940,1964]" pageId="9" pageNumber="160">L</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFC85D687FBDF8CB7" authority="Sokolov" authorityName="Sokolov" authorityYear="2016" box="[879,1135,1940,1964]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B3B737BFC85D687FBBF8CB7" box="[879,1039,1940,1964]" italics="true" pageId="9" pageNumber="160">Zeanillus nunni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFB91D687FB238CB7" bold="true" box="[1147,1171,1940,1964]" pageId="9" pageNumber="160">M</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFB46D687FF5F8CCB" authority="Sokolov" authorityName="Sokolov" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="BF84EAE68B3B737BFB46D687FA2D8CB7" box="[1196,1437,1940,1964]" italics="true" pageId="9" pageNumber="160">Pelodiaetodes insularis</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFF1DD6ABFEB98CCB" bold="true" box="[247,265,1976,2000]" pageId="9" pageNumber="160">N</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFEC9D6ABFDFE8CCB" authority="Sokolov" authorityName="Sokolov" box="[291,590,1976,2000]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B3B737BFEC9D6ABFE418CCB" box="[291,497,1976,2000]" italics="true" pageId="9" pageNumber="160">Pelodiaetodes nunni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFDBCD6ABFDDA8CCB" bold="true" box="[598,618,1976,2000]" pageId="9" pageNumber="160">O</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFD69D6ABFC0B8CCB" authority="Sokolov" authorityName="Sokolov" authorityYear="2015" box="[643,955,1976,2000]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="BF84EAE68B3B737BFD69D6ABFCEE8CCB" box="[643,862,1976,2000]" italics="true" pageId="9" pageNumber="160">Pelodiaetodes moorei</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3B737BFC29D6ABFC628CCB" bold="true" box="[963,978,1976,2000]" pageId="9" pageNumber="160">P</emphasis>
— 
<taxonomicName id="4AF04D778B3B737BFC06D6ABFA298CCB" authority="Jeannel. Legend" authorityName="Jeannel. Legend" box="[1004,1433,1976,2000]" class="Insecta" family="Carabidae" genus="Pelodiaetus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="sulcatipennis">
<emphasis id="BF84EAE68B3B737BFC06D6ABFB5E8CCB" box="[1004,1262,1976,2000]" italics="true" pageId="9" pageNumber="160">Pelodiaetus sulcatipennis</emphasis>
Jeannel. Legend
</taxonomicName>
: mtes—metendosternite. Scales intentionally omitted.
</paragraph>
</caption>
<caption id="D98F667C8B387378FF7DD658FBDA8CE8" ID-DOI="http://doi.org/10.5281/zenodo.10184237" ID-Zenodo-Dep="10184237" httpUri="https://zenodo.org/record/10184237/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,250,1867,1891]" targetBox="[189,1398,181,1843]" targetPageId="10" targetType="figure">
<paragraph id="8D4F36F48B387378FF7DD658FBDA8CE8" blockId="10.[151,1437,1867,2036]" pageId="10" pageNumber="161">
<emphasis id="BF84EAE68B387378FF7DD658FEBF8C78" bold="true" box="[151,271,1867,1891]" pageId="10" pageNumber="161">FIGURE 6</emphasis>
. SEM illustrations of structural features of elytra of 
<taxonomicName id="4AF04D778B387378FCDED658FC358C78" authorityName="Jeannel" authorityYear="1937" box="[820,901,1867,1891]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, left lateral aspect: 
<emphasis id="BF84EAE68B387378FBB9D658FB338C7F" bold="true" box="[1107,1155,1867,1892]" pageId="10" pageNumber="161">A–B</emphasis>
—subtribe 
<taxonomicName id="4AF04D778B387378FB1CD658FA288C78" authorityName="Sokolov" authorityYear="2023" box="[1270,1432,1867,1891]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
; 
<emphasis id="BF84EAE68B387378FF7DD67CFF198C9C" bold="true" box="[151,169,1903,1927]" pageId="10" pageNumber="161">A</emphasis>
— 
<taxonomicName id="4AF04D778B387378FF29D67CFDB38C93" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[195,515,1903,1928]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B387378FF29D67CFE198C9C" box="[195,425,1903,1927]" italics="true" pageId="10" pageNumber="161">Nesamblyops oreobius</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B387378FDE5D663FD908C93" bold="true" box="[527,544,1904,1928]" pageId="10" pageNumber="161">B</emphasis>
— 
<taxonomicName id="4AF04D778B387378FDD3D67CFC368C9C" authority="(Sharp)" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[569,902,1903,1927]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B387378FDD3D67CFC808C9C" box="[569,816,1903,1927]" italics="true" pageId="10" pageNumber="161">Nesamblyops subcaecus</emphasis>
(Sharp)
</taxonomicName>
; 
<emphasis id="BF84EAE68B387378FC78D67CFC148C9C" bold="true" box="[914,932,1903,1927]" pageId="10" pageNumber="161">C</emphasis>
— 
<taxonomicName id="4AF04D778B387378FC57D67CFB798C93" authority="Casey" authorityName="Casey" authorityYear="1918" box="[957,1225,1903,1928]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="carolinae">
<emphasis id="BF84EAE68B387378FC57D67CFB308C9C" box="[957,1152,1903,1927]" italics="true" pageId="10" pageNumber="161">Anillinus carolinae</emphasis>
Casey
</taxonomicName>
; 
<emphasis id="BF84EAE68B387378FB3ED663FB568C93" bold="true" box="[1236,1254,1904,1928]" pageId="10" pageNumber="161">D</emphasis>
— 
<taxonomicName id="4AF04D778B387378FAEAD67CFEDC8CB0" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="alleni">
<emphasis id="BF84EAE68B387378FAEAD67CFA2D8C9C" box="[1280,1437,1903,1927]" italics="true" pageId="10" pageNumber="161">Anillinus alleni</emphasis>
Sokolov and Carlton
</taxonomicName>
; 
<emphasis id="BF84EAE68B387378FE9CD687FE378CB7" bold="true" box="[374,391,1940,1964]" pageId="10" pageNumber="161">E</emphasis>
— 
<taxonomicName id="4AF04D778B387378FE4AD680FD6B8CB0" authority="(Jeannel)" baseAuthorityName="Jeannel" baseAuthorityYear="1963" box="[416,731,1939,1963]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="donovani">
<emphasis id="BF84EAE68B387378FE4AD680FDC58CB0" box="[416,629,1939,1963]" italics="true" pageId="10" pageNumber="161">Serranillus donovani</emphasis>
(Jeannel)
</taxonomicName>
; 
<emphasis id="BF84EAE68B387378FD0FD687FD448CB7" bold="true" box="[741,756,1940,1964]" pageId="10" pageNumber="161">F</emphasis>
— 
<taxonomicName id="4AF04D778B387378FCE4D680FAB78CB0" authority="Sokolov and Kavanaugh" authorityName="Sokolov and Kavanaugh" box="[782,1287,1939,1963]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="jalapensis">
<emphasis id="BF84EAE68B387378FCE4D680FBB58CB0" box="[782,1029,1939,1963]" italics="true" pageId="10" pageNumber="161">Geocharidius jalapensis</emphasis>
Sokolov and Kavanaugh
</taxonomicName>
; 
<emphasis id="BF84EAE68B387378FAFBD680FA958CB0" bold="true" box="[1297,1317,1939,1963]" pageId="10" pageNumber="161">G</emphasis>
— 
<taxonomicName id="4AF04D778B387378FAD4D680FEDB8CCB" authority="(Broun)" baseAuthorityName="Broun" baseAuthorityYear="1914" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="punctigerus">
<emphasis id="BF84EAE68B387378FAD4D680FEBF8CD4" italics="true" pageId="10" pageNumber="161">Zeanillus punctigerus</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B387378FE92D6ABFE3C8CCB" bold="true" box="[376,396,1976,2000]" pageId="10" pageNumber="161">H</emphasis>
— 
<taxonomicName id="4AF04D778B387378FE4FD6A4FD698CD4" authority="Sokolov" authorityName="Sokolov" box="[421,729,1975,1999]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B387378FE4FD6A4FDC88CD4" box="[421,632,1975,1999]" italics="true" pageId="10" pageNumber="161">Pelodiaetodes nunni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B387378FD0FD6ABFD5F8CCB" bold="true" box="[741,751,1976,2000]" pageId="10" pageNumber="161">I</emphasis>
— 
<taxonomicName id="4AF04D778B387378FCE2D6A4FBCE8CD4" authority="Sokolov. Legend" authorityName="Sokolov. Legend" box="[776,1150,1975,1999]" class="Insecta" family="Carabidae" genus="Pelodiaetus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B387378FCE2D6A4FC738CD4" box="[776,963,1975,1999]" italics="true" pageId="10" pageNumber="161">Pelodiaetus nunni</emphasis>
Sokolov. Legend
</taxonomicName>
: ed2—scutellar seta; ed6— discal seta, ed8—apical seta; eo1…eo9—setae 1–9 of umbilical series of pores. Scale = 0.2 mm.
</paragraph>
</caption>
<caption id="D98F667C8B397379FF7DD4BFFD3A8D4F" ID-DOI="http://doi.org/10.5281/zenodo.10184239" ID-Zenodo-Dep="10184239" httpUri="https://zenodo.org/record/10184239/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,250,1452,1476]" targetBox="[168,1419,201,1429]" targetPageId="11" targetType="figure">
<paragraph id="8D4F36F48B397379FF7DD4BFFD3A8D4F" blockId="11.[151,1437,1452,1621]" pageId="11" pageNumber="162">
<emphasis id="BF84EAE68B397379FF7DD4BFFEA08EDE" bold="true" box="[151,272,1452,1477]" pageId="11" pageNumber="162">FIGURE 7</emphasis>
. SEM illustrations of structural features of protibia of 
<taxonomicName id="4AF04D778B397379FCA5D4BEFC108EDE" authorityName="Jeannel" authorityYear="1937" box="[847,928,1453,1477]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, dorsal (E) and ventral (
<emphasis id="BF84EAE68B397379FB48D4BFFB638EDE" bold="true" box="[1186,1235,1452,1477]" pageId="11" pageNumber="162">A–D</emphasis>
, 
<emphasis id="BF84EAE68B397379FB09D4BEFAA38EDE" bold="true" box="[1251,1299,1453,1477]" pageId="11" pageNumber="162">F–H</emphasis>
) aspects: 
<emphasis id="BF84EAE68B397379FA97D4BFFF188EF2" bold="true" pageId="11" pageNumber="162">A– B</emphasis>
—subtribe 
<taxonomicName id="4AF04D778B397379FEF0D4C2FE0C8EF2" authorityName="Sokolov" authorityYear="2023" box="[282,444,1489,1513]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
; 
<emphasis id="BF84EAE68B397379FE2CD4C3FE688EF3" bold="true" box="[454,472,1488,1512]" pageId="11" pageNumber="162">A</emphasis>
— 
<taxonomicName id="4AF04D778B397379FE18D4C2FC818EF2" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[498,817,1489,1513]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B397379FE18D4C2FD678EF2" box="[498,727,1489,1513]" italics="true" pageId="11" pageNumber="162">Nesamblyops oreobius</emphasis>
(Broun)
</taxonomicName>
; 
<emphasis id="BF84EAE68B397379FCD1D4C2FCFC8EF2" bold="true" box="[827,844,1489,1513]" pageId="11" pageNumber="162">B</emphasis>
— 
<taxonomicName id="4AF04D778B397379FC8CD4C2FB018EF2" authority="(Sharp)" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[870,1201,1489,1513]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B397379FC8CD4C2FBEB8EF2" box="[870,1115,1489,1513]" italics="true" pageId="11" pageNumber="162">Nesamblyops subcaecus</emphasis>
(Sharp)
</taxonomicName>
; 
<emphasis id="BF84EAE68B397379FB56D4C3FB7E8EF3" bold="true" box="[1212,1230,1488,1512]" pageId="11" pageNumber="162">C</emphasis>
— 
<taxonomicName id="4AF04D778B397379FB0DD4C2FEC28D16" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="BF84EAE68B397379FB0DD4C2FA2C8EF2" box="[1255,1436,1489,1513]" italics="true" pageId="11" pageNumber="162">Anillinus lescheni</emphasis>
Sokolov and Carlton
</taxonomicName>
; 
<emphasis id="BF84EAE68B397379FE6AD4E6FE228D16" bold="true" box="[384,402,1525,1549]" pageId="11" pageNumber="162">D</emphasis>
— 
<taxonomicName id="4AF04D778B397379FE41D4E6FC538D16" authority="Sokolov and Kavanaugh" authorityName="Sokolov and Kavanaugh" box="[427,995,1525,1549]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="comayaguanus">
<emphasis id="BF84EAE68B397379FE41D4E6FD678D17" box="[427,727,1525,1549]" italics="true" pageId="11" pageNumber="162">Geocharidius comayaguanus</emphasis>
Sokolov and Kavanaugh
</taxonomicName>
; 
<emphasis id="BF84EAE68B397379FC1AD4E6FBB18D16" bold="true" box="[1008,1025,1525,1549]" pageId="11" pageNumber="162">E</emphasis>
— 
<taxonomicName id="4AF04D778B397379FBF0D4E6FA288D16" authority="Vigna Taglianti" authorityName="Vigna Taglianti" authorityYear="1973" box="[1050,1432,1525,1549]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="zullini">
<emphasis id="BF84EAE68B397379FBF0D4E6FB5C8D16" box="[1050,1260,1525,1549]" italics="true" pageId="11" pageNumber="162">Geocharidius zullini</emphasis>
Vigna Taglianti
</taxonomicName>
; 
<emphasis id="BF84EAE68B397379FF7DD70AFF168D2A" bold="true" box="[151,166,1561,1585]" pageId="11" pageNumber="162">F</emphasis>
— 
<taxonomicName id="4AF04D778B397379FF2AD70AFE758D2A" authority="Sokolov" authorityName="Sokolov" authorityYear="2016" box="[192,453,1561,1585]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="BF84EAE68B397379FF2AD70AFED78D2A" box="[192,359,1561,1585]" italics="true" pageId="11" pageNumber="162">Zeanillus brouni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B397379FE24D70BFE528D2B" bold="true" box="[462,482,1560,1584]" pageId="11" pageNumber="162">G</emphasis>
— 
<taxonomicName id="4AF04D778B397379FE11D70AFC988D2A" authority="Sokolov" authorityName="Sokolov" box="[507,808,1561,1585]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B397379FE11D70AFD7A8D2B" box="[507,714,1561,1585]" italics="true" pageId="11" pageNumber="162">Pelodiaetodes nunni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B397379FCDBD70AFCF58D2A" bold="true" box="[817,837,1561,1585]" pageId="11" pageNumber="162">H</emphasis>
— 
<taxonomicName id="4AF04D778B397379FCB4D70AFABF8D2A" authority="Jeannel. Legend" authorityName="Jeannel. Legend" box="[862,1295,1561,1585]" class="Insecta" family="Carabidae" genus="Pelodiaetus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="sulcatipennis">
<emphasis id="BF84EAE68B397379FCB4D70AFBD28D2A" box="[862,1122,1561,1585]" italics="true" pageId="11" pageNumber="162">Pelodiaetus sulcatipennis</emphasis>
Jeannel. Legend
</taxonomicName>
: as—articulosetae; psr—posterior setal row. Scale = 0.05 mm.
</paragraph>
</caption>
<paragraph id="8D4F36F48B397379FF2DD790FB318C82" blockId="11.[151,1437,1667,1945]" pageId="11" pageNumber="162">
<emphasis id="BF84EAE68B397379FF2DD790FE0E8D86" box="[199,446,1667,1693]" italics="true" pageId="11" pageNumber="162">Setation of ovipositor.</emphasis>
The members of 
<taxonomicName id="4AF04D778B397379FD6ED790FC848D86" authorityName="Sokolov" authorityYear="2023" box="[644,820,1667,1693]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
have ovipositor with three ensiform setae, represented by one medial and two lateral setae (
<figureCitation id="15CB2A718B397379FDA4D7B4FD608DDA" box="[590,720,1703,1729]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="11" pageNumber="162">Figs 4A–B</figureCitation>
, les1–2, mes). This state of ovipositor is not known in other species of 
<taxonomicName id="4AF04D778B397379FEFAD7D8FEDB8DFE" authorityName="Jeannel" authorityYear="1937" box="[272,363,1739,1765]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
. The representatives of the subtribes 
<taxonomicName id="4AF04D778B397379FCFFD7D8FCC18DFE" box="[789,881,1739,1765]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="subTribe" subTribe="Anillina">Anillina</taxonomicName>
and 
<taxonomicName id="4AF04D778B397379FC41D7D8FBF28DFE" box="[939,1090,1739,1765]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="subTribe" subTribe="Scotodipnina">Scotodipnina</taxonomicName>
, in which female genitalia are documented, have only two ensiform setae on the ovipositor, of which one is in a lateral position and the other is in a medial position (cf. 
<figureCitation id="15CB2A718B397379FE67D600FDB28C36" box="[397,514,1811,1837]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[167,1420,200,1667]" captionTargetId="figure-32@12.[167,1420,200,1667]" captionTargetPageId="12" captionText="FIGURE 8. SEM illustrations of ovipositor sclerites of Anillini, lateral (A, C–F) and ventral (B) aspects: A–B—subtribe Nesamblyopina; A—Nesamblyops brouni Sokolov, B—Nesamblyops brouni Sokolov; C—Anillinus lescheni Sokolov and Carlton; D—Geocharidius longinoi Sokolov and Kavanaugh; E—Pelodiaetodes moorei Sokolov; F—Pelodiaetodes nunni Sokolov. Legend: gc1—gonocoxite 1; gc2—gonocoxite 2; les1-2—lateral ensiform setae 1 and 2; mes—medial ensiform seta; ns—nematiform setae. Scale = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.10184243" httpUri="https://zenodo.org/record/10184243/files/figure.png" pageId="11" pageNumber="162">Figs 8C–F</figureCitation>
, les1, mes, with 
<figureCitation id="15CB2A718B397379FD57D600FC8A8C36" box="[701,826,1811,1837]" captionStart="FIGURE 4" captionStartId="8.[151,250,1239,1263]" captionTargetBox="[151,1437,181,1215]" captionTargetId="figure-32@8.[151,1437,181,1215]" captionTargetPageId="8" captionText="FIGURE 4. SEM illustrations of structural features of pterothorax of Anillini, ventral aspect: A–B—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Anillinus lescheni Sokolov and Carlton; D— Anillinus forthoodensis Sokolov and Reddell; E—Geocharidius jalapensis Sokolov and Kavanaugh; F—Zeanillus pallidus (Broun); G—Pelodiaetodes insularis Sokolov; H—Pelodiaetodes nunni Sokolov; I—Pelodiaetus sulcatipennis Jeannel. Legend: mes—mesanepisternum, mscx—mesocoxa, msts—mesothoracic anapleural suture, msv—mesoventrite, mtcx— metacoxal, mtp—metaventral process, mtv—metaventrite. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184233" httpUri="https://zenodo.org/record/10184233/files/figure.png" pageId="11" pageNumber="162">Figs 4A–B</figureCitation>
, les1–2, mes). Members of the subtribe 
<taxonomicName id="4AF04D778B397379FB11D600FA2C8C36" box="[1275,1436,1811,1837]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="subTribe" subTribe="Typhlocharina">Typhlocharina</taxonomicName>
demonstrate variations in the number of ensiform setae from 0 to 1. These cases of variation occur in parallel to the changes in shape of gonocoxite 2, which varies in the representatives of 
<taxonomicName id="4AF04D778B397379FC29D648FBD48C6E" box="[963,1124,1883,1909]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="subTribe" subTribe="Typhlocharina">Typhlocharina</taxonomicName>
from the typical unguiform to a unique tubular shape (
<bibRefCitation id="E9614B058B397379FE55D66CFC888C82" author="Perez-Gonzales, S. &amp; Zaballos, J. P." box="[447,824,1919,1945]" pageId="11" pageNumber="162" pagination="46 - 62" refId="ref27132" refString="Perez-Gonzales, S. &amp; Zaballos, J. P. (2012) Re-description of two species of Typhlocharis (Coleoptera: Carabidae: Anillini) and revision of the models of female genitalia within the genus. Zootaxa, 3279 (1), 46 - 62. https: // doi. org / 10.11646 / zootaxa. 3279.1.2" type="journal article" year="2012">Perez-Gonzales &amp; Zaballos 2012</bibRefCitation>
; 
<bibRefCitation id="E9614B058B397379FCA9D66CFBC58C82" author="Perez-Gonzales, S. &amp; Andujar, C. &amp; Zaballos, J. P." box="[835,1141,1919,1945]" pageId="11" pageNumber="162" pagination="359 - 383" refId="ref27060" refString="Perez-Gonzales, S., Andujar, C. &amp; Zaballos, J. P. (2018) Hidden biodiversity: total evidence phylogenetics and evolution of morphological traits in a highly diverse lineage of endogean ground beetles, Typhlocharis Dieck, 1869 (Carabidae, Trechinae, Anillini). Cladistics, 34, 359 - 383. https: // doi. org / 10.1111 / cla. 12208" type="journal article" year="2018">
Perez-Gonzales 
<emphasis id="BF84EAE68B397379FC13D693FB828C82" box="[1017,1074,1919,1945]" italics="true" pageId="11" pageNumber="162">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
<caption id="D98F667C8B3E737EFF7DD788FD868C59" ID-DOI="http://doi.org/10.5281/zenodo.10184243" ID-Zenodo-Dep="10184243" httpUri="https://zenodo.org/record/10184243/files/figure.png" pageId="12" pageNumber="163" startId="12.[151,250,1691,1715]" targetBox="[167,1420,200,1667]" targetPageId="12" targetType="figure">
<paragraph id="8D4F36F48B3E737EFF7DD788FD868C59" blockId="12.[151,1437,1691,1859]" pageId="12" pageNumber="163">
<emphasis id="BF84EAE68B3E737EFF7DD788FEA18DA8" bold="true" box="[151,273,1691,1715]" pageId="12" pageNumber="163">FIGURE 8</emphasis>
. SEM illustrations of ovipositor sclerites of 
<taxonomicName id="4AF04D778B3E737EFD00D788FC8B8DA8" authorityName="Jeannel" authorityYear="1937" box="[746,827,1691,1715]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, lateral (
<emphasis id="BF84EAE68B3E737EFC70D788FC1C8DA8" bold="true" box="[922,940,1691,1715]" pageId="12" pageNumber="163">A</emphasis>
, 
<emphasis id="BF84EAE68B3E737EFC57D788FC5B8DA8" bold="true" box="[957,1003,1691,1715]" pageId="12" pageNumber="163">C–F</emphasis>
) and ventral (
<emphasis id="BF84EAE68B3E737EFB6ED788FB258DA8" bold="true" box="[1156,1173,1691,1715]" pageId="12" pageNumber="163">B</emphasis>
) aspects: 
<emphasis id="BF84EAE68B3E737EFAEBD788FA818DA8" bold="true" box="[1281,1329,1691,1715]" pageId="12" pageNumber="163">A–B</emphasis>
—subtribe 
<taxonomicName id="4AF04D778B3E737EFF7DD7ACFE898DCC" authorityName="Sokolov" authorityYear="2023" box="[151,313,1727,1751]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
; 
<emphasis id="BF84EAE68B3E737EFEA2D7ACFEEA8DCC" bold="true" box="[328,346,1727,1751]" pageId="12" pageNumber="163">A</emphasis>
— 
<taxonomicName id="4AF04D778B3E737EFE9ED7ACFD798DCC" authority="Sokolov, B" authorityName="Sokolov, B" authorityYear="2023" box="[372,713,1727,1751]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="BF84EAE68B3E737EFE9ED7ACFDF98DCC" box="[372,585,1727,1751]" italics="true" pageId="12" pageNumber="163">Nesamblyops brouni</emphasis>
Sokolov, 
<emphasis id="BF84EAE68B3E737EFD52D7ACFD798DCC" bold="true" box="[696,713,1727,1751]" pageId="12" pageNumber="163">B</emphasis>
</taxonomicName>
— 
<taxonomicName id="4AF04D778B3E737EFD08D7ACFBAA8DCC" authority="Sokolov" authorityName="Sokolov, B" authorityYear="2023" box="[738,1050,1727,1751]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="BF84EAE68B3E737EFD08D7ACFC078DCC" box="[738,951,1727,1751]" italics="true" pageId="12" pageNumber="163">Nesamblyops brouni</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3E737EFBC2D7ACFB8A8DCC" bold="true" box="[1064,1082,1727,1751]" pageId="12" pageNumber="163">C</emphasis>
— 
<taxonomicName id="4AF04D778B3E737EFBBED7ACFF568DE0" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="BF84EAE68B3E737EFBBED7ACFABD8DCC" box="[1108,1293,1727,1751]" italics="true" pageId="12" pageNumber="163">Anillinus lescheni</emphasis>
Sokolov and Carlton
</taxonomicName>
; 
<emphasis id="BF84EAE68B3E737EFF1ED7F0FEB68DE0" bold="true" box="[244,262,1763,1787]" pageId="12" pageNumber="163">D</emphasis>
— 
<taxonomicName id="4AF04D778B3E737EFECAD7F0FCA78DE0" authority="Sokolov and Kavanaugh" authorityName="Sokolov and Kavanaugh" box="[288,791,1763,1787]" class="Insecta" family="Carabidae" genus="Geocharidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="longinoi">
<emphasis id="BF84EAE68B3E737EFECAD7F0FDB88DE0" box="[288,520,1763,1787]" italics="true" pageId="12" pageNumber="163">Geocharidius longinoi</emphasis>
Sokolov and Kavanaugh
</taxonomicName>
; 
<emphasis id="BF84EAE68B3E737EFCCFD7F0FC868DE0" bold="true" box="[805,822,1763,1787]" pageId="12" pageNumber="163">E</emphasis>
— 
<taxonomicName id="4AF04D778B3E737EFCA5D7F0FB228DE0" authority="Sokolov" authorityName="Sokolov" authorityYear="2015" box="[847,1170,1763,1787]" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="BF84EAE68B3E737EFCA5D7F0FB808DE0" box="[847,1072,1763,1787]" italics="true" pageId="12" pageNumber="163">Pelodiaetodes moorei</emphasis>
Sokolov
</taxonomicName>
; 
<emphasis id="BF84EAE68B3E737EFB4AD7F0FB1F8DE0" bold="true" box="[1184,1199,1763,1787]" pageId="12" pageNumber="163">F</emphasis>
— 
<taxonomicName id="4AF04D778B3E737EFB23D7F0FEF78C04" authority="Sokolov. Legend" authorityName="Sokolov. Legend" class="Insecta" family="Carabidae" genus="Pelodiaetodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="BF84EAE68B3E737EFB23D7F0FA2D8DE0" box="[1225,1437,1763,1787]" italics="true" pageId="12" pageNumber="163">Pelodiaetodes nunni</emphasis>
Sokolov. Legend
</taxonomicName>
: gc1—gonocoxite 1; gc2—gonocoxite 2; les1-2—lateral ensiform setae 1 and 2; mes—medial ensiform seta; ns—nematiform setae. Scale = 0.02 mm.
</paragraph>
</caption>
<paragraph id="8D4F36F48B3E737FFF2DD662FE528AEE" blockId="12.[151,1436,1905,2040]" lastBlockId="13.[151,1437,151,717]" lastPageId="13" lastPageNumber="164" pageId="12" pageNumber="163">
<emphasis id="BF84EAE68B3E737EFF2DD662FD968C90" bold="true" box="[199,550,1905,1931]" pageId="12" pageNumber="163">
Relationships outside 
<taxonomicName id="4AF04D778B3E737EFE26D662FD968C90" authorityName="Jeannel" authorityYear="1937" box="[460,550,1905,1931]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
</emphasis>
. As described above, a certain number of characters indicative for the subtribe have not yet been documented for the species of the tribe 
<taxonomicName id="4AF04D778B3E737EFCDAD686FC3B8CB4" authorityName="Jeannel" authorityYear="1937" box="[816,907,1941,1967]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
. To determine how widely such characters are distributed among the representatives of 
<taxonomicName id="4AF04D778B3E737EFD9AD6AAFD528CC8" box="[624,738,1977,2003]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="subFamily" subFamily="Trechinae">Trechinae</taxonomicName>
, especially within taxa from the sister clades to the 
<taxonomicName id="4AF04D778B3E737EFAAFD6AAFA2C8CC8" authorityName="Jeannel" authorityYear="1937" box="[1349,1436,1977,2003]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
clade on the phylogenetic tree of 
<taxonomicName id="4AF04D778B3E737EFDFFD6CEFC258CEC" authority="(Maddison et al. 2019)" baseAuthorityName="Maddison" baseAuthorityYear="2019" box="[533,917,2013,2039]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="subFamily" subFamily="Trechinae">
Trechinae (
<bibRefCitation id="E9614B058B3E737EFD7ED6CEFC3C8CEC" author="Maddison, D. R. &amp; Kanda, K. &amp; Boyd, O. F. &amp; Faille, A. &amp; Porch, N. &amp; Erwin, T. L. &amp; Roig-Junent, S." box="[660,908,2013,2039]" pageId="12" pageNumber="163" pagination="151 - 176" refId="ref26793" refString="Maddison, D. R., Kanda, K., Boyd, O. F., Faille, A., Porch, N., Erwin, T. L. &amp; Roig-Junent, S. (2019) Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): Unexpected clades, isolated lineages, and morphological convergence. Molecular Phylogenetics and Evolution, 132, 151 - 176. https: // doi. org / 10.1016 / j. ympev. 2018.11.006." type="journal article" year="2019">
Maddison 
<emphasis id="BF84EAE68B3E737EFCE4D6CDFCF88CEC" box="[782,840,2013,2039]" italics="true" pageId="12" pageNumber="163">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
, members of the genera of Sinozolini, 
<taxonomicName id="4AF04D778B3E737EFABBD6CEFA288CEC" box="[1361,1432,2013,2039]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="tribe" tribe="Zolini">Zolini</taxonomicName>
, 
<taxonomicName id="4AF04D778B3F737FFF7DD184FEA58BAA" box="[151,277,151,177]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="tribe" tribe="Bembidiini">Bembidiini</taxonomicName>
and of both subtribes of Tachyini (i.e., Tachyina and Xystosomina) were examined. Results of this investigation are presented in 
<tableCitation id="C072034F8B3F737FFE1BD1A8FDF88BCE" box="[497,584,187,213]" captionStart="TABLE 1" captionStartId="3.[150,238,494,520]" captionTargetId="graphics-35@3.[150,2038,522,1596]" captionTargetPageId="3" captionText="TABLE 1. Variation of character states among the representatives of the Trechinae (character states of Nesamblyopina are marked with blue color)." httpUri="http://table.plazi.org/id/D98F667C8B317371FF7CD0FDF90E8913" pageId="13" pageNumber="164" tableUuid="D98F667C8B317371FF7CD0FDF90E8913">Table 1</tableCitation>
. The results show that character states indicative for the new subtribe are widely distributed among the examined non-anilline genera and can thus be interpreted as plesiomorphic. Especially amazing is the concordance in character states between 
<taxonomicName id="4AF04D778B3F737FFCFFD010FC758A06" authorityName="Sokolov" authorityYear="2023" box="[789,965,259,285]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
and the species of the genus 
<taxonomicName id="4AF04D778B3F737FFAF2D010FDA48A5A" authority="Schaum (Xystosomina, Tachyini)" class="Insecta" family="Carabidae" genus="Xystosomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B3F737FFAF2D010FA2C8A07" box="[1304,1436,259,284]" italics="true" pageId="13" pageNumber="164">Xystosomus</emphasis>
Schaum (Xystosomina, Tachyini)
</taxonomicName>
. The members of 
<taxonomicName id="4AF04D778B3F737FFD0BD034FC218A5A" authorityName="Sokolov" authorityYear="2023" box="[737,913,295,321]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
share the states of seven from nine characters shown in 
<tableCitation id="C072034F8B3F737FFEE2D058FEEC8A7E" box="[264,348,331,357]" captionStart="TABLE 1" captionStartId="3.[150,238,494,520]" captionTargetId="graphics-35@3.[150,2038,522,1596]" captionTargetPageId="3" captionText="TABLE 1. Variation of character states among the representatives of the Trechinae (character states of Nesamblyopina are marked with blue color)." httpUri="http://table.plazi.org/id/D98F667C8B317371FF7CD0FDF90E8913" pageId="13" pageNumber="164" tableUuid="D98F667C8B317371FF7CD0FDF90E8913">Table 1</tableCitation>
with the species of the genus 
<taxonomicName id="4AF04D778B3F737FFD51D058FC8F8A7F" box="[699,831,331,356]" class="Insecta" family="Carabidae" genus="Xystosomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B3F737FFD51D058FC8F8A7F" box="[699,831,331,356]" italics="true" pageId="13" pageNumber="164">Xystosomus</emphasis>
</taxonomicName>
, while the other representatives of 
<taxonomicName id="4AF04D778B3F737FFB3ED058FA9B8A7E" authorityName="Jeannel" authorityYear="1937" box="[1236,1323,331,357]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
shown in 
<tableCitation id="C072034F8B3F737FFF7DD07CFF5A8A92" box="[151,234,367,393]" captionStart="TABLE 1" captionStartId="3.[150,238,494,520]" captionTargetId="graphics-35@3.[150,2038,522,1596]" captionTargetPageId="3" captionText="TABLE 1. Variation of character states among the representatives of the Trechinae (character states of Nesamblyopina are marked with blue color)." httpUri="http://table.plazi.org/id/D98F667C8B317371FF7CD0FDF90E8913" pageId="13" pageNumber="164" tableUuid="D98F667C8B317371FF7CD0FDF90E8913">Table 1</tableCitation>
share at most the state of only one character with the members of 
<taxonomicName id="4AF04D778B3F737FFC20D07CFBCE8A92" authorityName="Sokolov" authorityYear="2023" box="[970,1150,367,393]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
. Nevertheless, because of the paucity of detailed morphological descriptions for many genera of 
<taxonomicName id="4AF04D778B3F737FFC77D080FC448AB6" authorityName="Jeannel" authorityYear="1937" box="[925,1012,403,429]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
in particular and 
<taxonomicName id="4AF04D778B3F737FFB59D080FA928AB6" box="[1203,1314,403,429]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="subFamily" subFamily="Trechinae">Trechinae</taxonomicName>
in general, additional morphological investigations are necessary to elucidate the relationships between 
<taxonomicName id="4AF04D778B3F737FFB51D0A4FADB8ACA" authorityName="Sokolov" authorityYear="2023" box="[1211,1387,439,465]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
and other basal taxa of 
<taxonomicName id="4AF04D778B3F737FFE86D0C8FE6F8AEE" box="[364,479,475,501]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="subFamily" subFamily="Trechinae">Trechinae</taxonomicName>
.
</paragraph>
<paragraph id="8D4F36F48B3F737FFF2DD0ECFE2689B2" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="164">
By considering all comparative morphological data, it seems reasonable to state that the new subtribe is characterized by a combination of mostly plesiomorphic traits, pointing out that 
<taxonomicName id="4AF04D778B3F737FFB8DD330FAA78926" authorityName="Sokolov" authorityYear="2023" box="[1127,1303,547,573]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="163" phylum="Arthropoda" rank="subTribe" subTribe="Nesamblyopina">Nesamblyopina</taxonomicName>
constitutes presumably an ancient relic lineage within 
<taxonomicName id="4AF04D778B3F737FFD68D354FD6D897A" authorityName="Jeannel" authorityYear="1937" box="[642,733,583,609]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
. This assumption is in concordance with the basal position of the 
<taxonomicName id="4AF04D778B3F737FFF29D378FEE8899E" authorityName="Jeannel" authorityYear="1937" box="[195,344,619,645]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B3F737FFF29D378FEE8899E" box="[195,344,619,645]" italics="true" pageId="13" pageNumber="164">Nesamblyops</emphasis>
</taxonomicName>
clade on the phylogenetic tree of 
<taxonomicName id="4AF04D778B3F737FFD0BD378FC88899E" authorityName="Jeannel" authorityYear="1937" box="[737,824,619,645]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
obtained in molecular analyses (
<bibRefCitation id="E9614B058B3F737FFB58D378FA28899E" author="Andujar, C. &amp; Faille, A. &amp; Perez-Gonzalez, S. &amp; Zaballos, J. P. &amp; Vogler, A. P. &amp; Ribera, I." box="[1202,1432,619,645]" pageId="13" pageNumber="164" pagination="235 - 246" refId="ref25995" refString="Andujar, C., Faille, A., Perez-Gonzalez, S., Zaballos, J. P., Vogler, A. P. &amp; Ribera, I. (2016) Gondwanian relicts and oceanic dispersal in a cosmopolitan radiation of euedaphic ground beetles. Molecular Phylogenetics and Evolution, 99, 235 - 246. https: // doi. org / 10.1016 / j. ympev. 2016.03.013" type="journal article" year="2016">
Andujar 
<emphasis id="BF84EAE68B3F737FFAF3D378FAE4899E" box="[1305,1364,619,645]" italics="true" pageId="13" pageNumber="164">et al.</emphasis>
2016
</bibRefCitation>
; 
<bibRefCitation id="E9614B058B3F737FFF7DD39CFE3A89B2" author="Maddison, D. R. &amp; Kanda, K. &amp; Boyd, O. F. &amp; Faille, A. &amp; Porch, N. &amp; Erwin, T. L. &amp; Roig-Junent, S." box="[151,394,655,681]" pageId="13" pageNumber="164" pagination="151 - 176" refId="ref26793" refString="Maddison, D. R., Kanda, K., Boyd, O. F., Faille, A., Porch, N., Erwin, T. L. &amp; Roig-Junent, S. (2019) Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): Unexpected clades, isolated lineages, and morphological convergence. Molecular Phylogenetics and Evolution, 132, 151 - 176. https: // doi. org / 10.1016 / j. ympev. 2018.11.006." type="journal article" year="2019">
Maddison 
<emphasis id="BF84EAE68B3F737FFEE4D39CFEF689B2" box="[270,326,655,681]" italics="true" pageId="13" pageNumber="164">et al.</emphasis>
2019
</bibRefCitation>
).
</paragraph>
<paragraph id="8D4F36F48B3F737FFF2DD3A0FD4689D6" blockId="13.[151,1437,151,717]" box="[199,758,691,717]" pageId="13" pageNumber="164">At this time, the new subtribe includes one genus.</paragraph>
</subSubSection>
</treatment>
</document>