treatments-xml/data/05/59/87/055987E28B2F736DFF7DD2D6FB388F66.xml
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<document id="2EA4511C35963F8D0A529C2D84CDB229" ID-DOI="10.11646/zootaxa.5375.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10184224" ID-ZooBank="F3D0E008-556C-4FAD-BF51-4F1A714325DA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700642529072" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2023" docId="055987E28B2F736DFF7DD2D6FB388F66" docLanguage="en" docName="zootaxa.5375.2.1.pdf" docOrigin="Zootaxa 5375 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.2.1/52321" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nesamblyops subcaecus Greymouth, Buller, SO" docType="treatment" docVersion="2" lastPageNumber="182" masterDocId="F960FF9A8B327372FFEAD113FFB08B1B" masterDocTitle="Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe" masterLastPageNumber="192" masterPageNumber="151" pageNumber="180" updateTime="1700643183691" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="08F5418DA1C70D71CF3260017F9C7D61">Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe</mods:title>
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<paragraph id="8D4F36F48B2F736FFF7DD2D6FD4B88C4" blockId="29.[151,763,965,1028]" box="[151,763,965,992]" pageId="29" pageNumber="180">
<heading id="D60781988B2F736FFF7DD2D6FD4B88C4" bold="true" box="[151,763,965,992]" fontSize="11" level="1" pageId="29" pageNumber="180" reason="1">
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<taxonomicName id="4AF04D778B2F736FFF7DD2D6FD9188C4" ID-CoL="476SD" authority="(Sharp)" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[151,545,965,992]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2F736FFF7DD2D6FE0A88FB" bold="true" box="[151,442,965,992]" italics="true" pageId="29" pageNumber="180">Nesamblyops subcaecus</emphasis>
(Sharp)
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(dissected 5 exx.)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C5EA657F8B2F736FFF7DD2FAFDE58F18" box="[151,597,1001,1028]" pageId="29" pageNumber="180" type="description">
<paragraph id="8D4F36F48B2F736FFF7DD2FAFDE58F18" blockId="29.[151,763,965,1028]" box="[151,597,1001,1028]" pageId="29" pageNumber="180">
<figureCitation id="15CB2A718B2F736FFF7DD2FAFE958F18" box="[151,293,1001,1027]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="29" pageNumber="180">Figures. 10F</figureCitation>
,
<figureCitation id="15CB2A718B2F736FFEDBD2FAFED28F18" box="[305,354,1001,1027]" captionStart="FIGURE 13" captionStartId="28.[151,250,934,958]" captionTargetBox="[157,1430,181,910]" captionTargetId="figure-32@28.[157,1430,181,910]" captionTargetPageId="28" captionText="FIGURE 13. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. rotundicollis (Onekaka, Nelson, SO), B—N. solitarius (Italian Creek, Capleston, Buller, SO), C—N. subcaecus (Greymouth, Buller, SO), D—N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO), E—N. viator (Secretary Island, Grono Bay, Fiordland, SO), F—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184253" httpUri="https://zenodo.org/record/10184253/files/figure.png" pageId="29" pageNumber="180">13C</figureCitation>
,
<figureCitation id="15CB2A718B2F736FFE87D2FAFE0A8F1F" box="[365,442,1001,1028]" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="29" pageNumber="180">15PR</figureCitation>
,
<figureCitation id="15CB2A718B2F736FFE2DD2FAFE408F18" box="[455,496,1001,1027]" captionStart="FIGURE 17" captionStartId="34.[151,250,1191,1215]" captionTargetBox="[163,1421,187,1168]" captionTargetId="figure-32@34.[151,1436,181,1168]" captionTargetPageId="34" captionText="FIGURE 17. Line drawings of ring sclerite of New Zealand Nesamblyops species, male genitalia, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (Mount Domett, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO), G—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), H—N. ovipennis (Mount Arthur, Nelson, SO), I—N. rotundicollis (Onekaka, Nelson, SO), J—N. subcaecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO), K—N. viator (Secretary Island, Grono Bay, Fjordland, SO), L—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184261" httpUri="https://zenodo.org/record/10184261/files/figure.png" pageId="29" pageNumber="180">17J</figureCitation>
,
<figureCitation id="15CB2A718B2F736FFE16D2FAFD9E8F18" box="[508,558,1001,1027]" captionStart="FIGURE 18" captionStartId="35.[151,250,872,896]" captionTargetBox="[159,1434,181,829]" captionTargetId="figure-15@35.[151,1436,181,849]" captionTargetPageId="35" captionText="FIGURE 18. Line drawings of spermatheca of New Zealand Nesamblyops species. A—N. magnificus (Allison Reserve, Akatore, Dunedin, SO), B—N. rotundicollis (Parahamoi, Nelson, SO), C—N. subcaecus (Lake Kaniere Scenic Reserve, Mount Tuhua, Westland, SO). Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184265" httpUri="https://zenodo.org/record/10184265/files/figure.png" pageId="29" pageNumber="180">18C</figureCitation>
,
<figureCitation id="15CB2A718B2F736FFDD3D2FAFDE58F18" box="[569,597,1001,1027]" captionStart="FIGURE 21" captionStartId="37.[151,250,1860,1884]" captionTargetBox="[151,1437,1100,1836]" captionTargetId="figure-68@37.[151,1437,1100,1836]" captionTargetPageId="37" captionText="FIGURE 21. Distributional records for the Nesamblyops subcaecus and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Red star—the type locality of Cillenum subcaecum Sharp; black star—the type locality of Tachys coriaceus Broun. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184271" httpUri="https://zenodo.org/record/10184271/files/figure.png" pageId="29" pageNumber="180">21</figureCitation>
</paragraph>
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<emphasis id="BF84EAE68B2F736FFF7DD53DFF158F5D" bold="true" box="[151,165,1070,1094]" pageId="29" pageNumber="180">=</emphasis>
<treatmentCitation id="0C5110E58B2F736FFF4CD53DFD818F5D" author="Sharp, D." box="[166,561,1070,1094]" page="375" pageId="29" pageNumber="180" year="1886">
<taxonomicName id="4AF04D778B2F736FFF4CD53DFD818F5D" authority="Sharp, 1886: 375" authorityName="Sharp" authorityPageNumber="375" authorityYear="1886" box="[166,561,1070,1094]" class="Insecta" family="Carabidae" genus="Cillenum" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="subcaecum">
<emphasis id="BF84EAE68B2F736FFF4CD53DFECA8F5D" box="[166,378,1070,1094]" italics="true" pageId="29" pageNumber="180">Cillenum subcaecum</emphasis>
<bibRefCitation id="E9614B058B2F736FFE6AD53DFD818F5D" author="Sharp, D." box="[384,561,1070,1094]" pageId="29" pageNumber="180" pagination="351 - 454" refId="ref27315" refString="Sharp, D. (1886) On New Zealand Coleoptera, with descriptions of new genera and species. The Scientific Transactions of the Royal Dublin Society, 2 (3), 351 - 454." type="journal article" year="1886">Sharp, 1886: 375</bibRefCitation>
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.
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</paragraph>
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<emphasis id="BF84EAE68B2F736FFF7DD567FE318F95" bold="true" box="[151,385,1140,1166]" pageId="29" pageNumber="180">Material examined:</emphasis>
\
<collectingCountry id="F5E776648B2F736FFE78D566FDED8F95" box="[402,605,1140,1167]" name="New Zealand" pageId="29" pageNumber="180">NEW ZEALAND</collectingCountry>
,
<collectionCode id="EBE1AE318B2F736FFD8CD566FD3B8F94" box="[614,651,1141,1167]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="29" pageNumber="180">BR</collectionCode>
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King Domain
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<date id="F94E10348B2F736FFC1FD567FBCA8F95" box="[1013,1146,1140,1167]" pageId="29" pageNumber="180" value="1983-06-10">10 Jun 1983</date>
H.P. McColl \ Litter 15/83 \
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PB\ (
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); \ Kings Domain
<taxonomicName id="4AF04D778B2F736FFDA0D58BFC8A8FA9" authority="Westland" authorityName="Westland" box="[586,826,1176,1202]" domain="Greymouth" pageId="29" pageNumber="180" rank="domain">Greymouth Westland</taxonomicName>
\ Coll. E.Fairburn
<date id="F94E10348B2F736FFBF8D58BFBC98FA8" box="[1042,1145,1176,1203]" pageId="29" pageNumber="180" value="1945-01-08">8-1-1945</date>
\
<taxonomicName id="4AF04D778B2F736FFB7BD58BFAB98FA8" box="[1169,1289,1176,1203]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BF84EAE68B2F736FFB7BD58BFB508FA9" box="[1169,1248,1176,1202]" italics="true" pageId="29" pageNumber="180">Anillus</emphasis>
sp.
</taxonomicName>
\ Leafmould \ A.E. Brookes Collection \ (
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); \ Moa Basin \
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.Broun Collection \ A.E. Brookes Collection \
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\ Ns \
<collectingCountry id="F5E776648B2F736FFF7DD5F2FF0C8FE0" box="[151,188,1249,1275]" name="New Zealand" pageId="29" pageNumber="180">NZ</collectingCountry>
PB \ (
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); \ Arthurs Pass 2500 24.3.65 \ N.A. Walker \
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\ Ns \
<collectingCountry id="F5E776648B2F736FFC09D5F2FBB88FE0" box="[995,1032,1249,1275]" name="New Zealand" pageId="29" pageNumber="180">NZ</collectingCountry>
PB\ (
<specimenCount id="9BF6FD7D8B2F736FFBA1D5F3FB008FE1" box="[1099,1200,1248,1274]" count="1" pageId="29" pageNumber="180" type="female">1 female</specimenCount>
); \
<collectingCountry id="F5E776648B2F736FFB38D5F2FA2C8FE1" box="[1234,1436,1248,1275]" name="New Zealand" pageId="29" pageNumber="180">NEW ZEALAND</collectingCountry>
<collectionCode id="EBE1AE318B2F736FFF7DD417FF738E05" box="[151,195,1284,1310]" country="0" name="Museo de Cipolleti" pageId="29" pageNumber="180">MC</collectionCode>
Arthurs Pass Bealy Valley Margarets Tarn
<quantity id="4A089B118B2F736FFD59D417FD438E05" box="[691,755,1284,1310]" metricMagnitude="2" metricUnit="m" metricValue="8.83" pageId="29" pageNumber="180" unit="m" value="883.0">883m</quantity>
\
<date id="F94E10348B2F736FFCE6D417FC3E8E05" box="[780,910,1284,1310]" pageId="29" pageNumber="180" value="1982-02-08">8 Feb 1982</date>
J.S. Dugdale Litter 82/21 \
<collectingCountry id="F5E776648B2F736FFB3AD416FB458E04" box="[1232,1269,1285,1311]" name="New Zealand" pageId="29" pageNumber="180">NZ</collectingCountry>
PB \ (
<specimenCount id="9BF6FD7D8B2F736FFAA9D417FA2C8E05" box="[1347,1436,1284,1310]" count="4" pageId="29" pageNumber="180" type="male">4 males</specimenCount>
and females); \
<collectingCountry id="F5E776648B2F736FFEA7D43AFDAB8E59" box="[333,539,1320,1347]" name="New Zealand" pageId="29" pageNumber="180">NEW ZEALAND</collectingCountry>
<collectionCode id="EBE1AE318B2F736FFDCFD43BFDE18E59" box="[549,593,1320,1346]" country="0" name="Museo de Cipolleti" pageId="29" pageNumber="180">MC</collectionCode>
Arthurs Pass Dobson Nature Walk
<date id="F94E10348B2F736FFC05D43BFBC38E59" box="[1007,1139,1320,1346]" pageId="29" pageNumber="180" value="1982-02-08">8 Feb 1982</date>
\ C.F.Butcher sifted litter 82/26 \
<collectingCountry id="F5E776648B2F736FFF04D45EFEA38E7C" box="[238,275,1357,1383]" name="New Zealand" pageId="29" pageNumber="180">NZ</collectingCountry>
PB\ (
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); \ in moss Temple Basin Arthurs Pass 3600
<date id="F94E10348B2F736FFC4FD45FFB888E7D" box="[933,1080,1356,1383]" pageId="29" pageNumber="180" value="1966-11-12">12 Nov 1966</date>
B.M. May \ (
<specimenCount id="9BF6FD7D8B2F736FFB3FD45FFA8A8E7D" box="[1237,1338,1356,1382]" count="1" pageId="29" pageNumber="180" type="female">1 female</specimenCount>
); \
<collectingCountry id="F5E776648B2F736FFAB7D45EFEAB8E91" name="New Zealand" pageId="29" pageNumber="180">NEW ZEALAND</collectingCountry>
WD Mt Tuhua,
<quantity id="4A089B118B2F736FFE3BD463FDAF8E91" box="[465,543,1392,1418]" metricMagnitude="3" metricUnit="m" metricValue="1.067" pageId="29" pageNumber="180" unit="m" value="1067.0">1067m</quantity>
E.side of L. Kaniere
<date id="F94E10348B2F736FFCE1D463FC2C8E91" box="[779,924,1392,1419]" pageId="29" pageNumber="180" value="1984-11-20">20 Nov 1984</date>
\ C.F. Butcher Litter and mats 84/74 \
<collectingCountry id="F5E776648B2F736FFAADD462FADC8E90" box="[1351,1388,1393,1419]" name="New Zealand" pageId="29" pageNumber="180">NZ</collectingCountry>
PB\ (
<specimenCount id="9BF6FD7D8B2F736FFF4AD487FF478EB5" box="[160,247,1428,1454]" count="7" pageId="29" pageNumber="180" type="male">7 males</specimenCount>
and females).
</paragraph>
<paragraph id="8D4F36F48B2F736FFF2DD4ABFD8F8CF5" blockId="29.[151,1437,1140,2030]" pageId="29" pageNumber="180">
<emphasis id="BF84EAE68B2F736FFF2DD4ABFE018EC9" bold="true" box="[199,433,1464,1490]" pageId="29" pageNumber="180">Taxonomical notes.</emphasis>
My interpretation of
<taxonomicName id="4AF04D778B2F736FFD5ED4AAFCFC8EC9" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[692,844,1464,1490]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2F736FFD5ED4AAFCFC8EC9" box="[692,844,1464,1490]" italics="true" pageId="29" pageNumber="180">N. subcaecus</emphasis>
</taxonomicName>
is based on the specimen (see Material examined above) originated from the
<typeStatus id="524B88568B2F736FFE3CD4CEFDB68EEC" box="[470,518,1501,1527]" pageId="29" pageNumber="180">type</typeStatus>
locality of the species (“
<taxonomicName id="4AF04D778B2F736FFCC4D4CFFC018EED" box="[814,945,1500,1526]" domain="Greymouth" pageId="29" pageNumber="180" rank="domain">Greymouth</taxonomicName>
. Helms, ex. Reitter”, p. 375,
<bibRefCitation id="E9614B058B2F736FFAE2D4CFFA208EED" author="Sharp, D." box="[1288,1424,1500,1526]" pageId="29" pageNumber="180" pagination="351 - 454" refId="ref27315" refString="Sharp, D. (1886) On New Zealand Coleoptera, with descriptions of new genera and species. The Scientific Transactions of the Royal Dublin Society, 2 (3), 351 - 454." type="journal article" year="1886">Sharp 1886</bibRefCitation>
). Unfortunately, this specimen is a female, so, its association with a certain group of specimens is based exclusively on external morphological data. Another issue needs to be mentioned here is that the
<typeStatus id="524B88568B2F736FFB7BD736FB718D24" box="[1169,1217,1573,1599]" pageId="29" pageNumber="180">type</typeStatus>
locality of
<taxonomicName id="4AF04D778B2F736FFAA5D737FEE38D78" authority="Broun" authorityName="Broun" class="Insecta" family="Carabidae" genus="Tachys" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="BF84EAE68B2F736FFAA5D737FEB48D79" italics="true" pageId="29" pageNumber="180">Tachys coriaceus</emphasis>
Broun
</taxonomicName>
(synonym of
<taxonomicName id="4AF04D778B2F736FFE1DD75AFC988D79" authority="(Broun, 1893)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[503,808,1608,1634]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B2F736FFE1DD75AFDCB8D79" box="[503,635,1608,1634]" italics="true" pageId="29" pageNumber="180">N. oreobius</emphasis>
(
<bibRefCitation id="E9614B058B2F736FFD66D75BFCAF8D79" author="Broun, T." box="[652,799,1608,1634]" pageId="29" pageNumber="180" pagination="1395 - 1504" refId="ref26204" refString="Broun, T. (1893) New Zealand Coleoptera. Part VII. In: Manual of the New Zealand Coleoptera. Parts V., VI., VII. Government Printing Office, Wellington, pp. 1395 - 1504." type="book chapter" year="1893">Broun, 1893</bibRefCitation>
)
</taxonomicName>
, established by
<bibRefCitation id="E9614B058B2F736FFC0BD75AFBC58D79" author="Moore, B. P." box="[993,1141,1608,1635]" pageId="29" pageNumber="180" pagination="399 - 406" refId="ref26941" refString="Moore, B. P. (1980) A synopsis of the New Zealand Anillini (Coleoptera: Carabidae: Bembidiinae), with descriptions of new genera and species. New Zealand Journal of Zoology, 7, 399 - 406. https: // doi. org / 10.1080 / 03014223.1980.10423793" type="journal article" year="1980">Moore, 1980</bibRefCitation>
) located within the range of the species I consider to be named as
<taxonomicName id="4AF04D778B2F736FFD89D77EFD478D9D" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[611,759,1644,1670]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2F736FFD89D77EFD478D9D" box="[611,759,1644,1670]" italics="true" pageId="29" pageNumber="180">N. subcaecus</emphasis>
</taxonomicName>
. If the geographical label is correct, then the
<typeStatus id="524B88568B2F736FFB10D77EFA9A8D9C" box="[1274,1322,1645,1671]" pageId="29" pageNumber="180">type</typeStatus>
specimen of
<taxonomicName id="4AF04D778B2F736FFF5CD782FE8F8DB1" box="[182,319,1681,1706]" class="Insecta" family="Carabidae" genus="Tachys" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="BF84EAE68B2F736FFF5CD782FE8F8DB1" box="[182,319,1681,1706]" italics="true" pageId="29" pageNumber="180">T. coriaceus</emphasis>
</taxonomicName>
cannot be a synonym of
<taxonomicName id="4AF04D778B2F736FFDB1D782FD6D8DB1" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[603,733,1680,1706]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B2F736FFDB1D782FD6D8DB1" box="[603,733,1680,1706]" italics="true" pageId="29" pageNumber="180">N. oreobius</emphasis>
</taxonomicName>
, which range is limited to the central parts of the North Island (
<bibRefCitation id="E9614B058B2F736FFF75D7A7FEF18DD5" author="Sokolov, I. M." box="[159,321,1716,1742]" pageId="29" pageNumber="180" pagination="179 - 201" refId="ref27417" refString="Sokolov, I. M. (2023) Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa, 5230 (2), 179 - 201. https: // doi. org / 10.11646 / zootaxa. 5230.2.3" type="journal article" year="2023">Sokolov 2023</bibRefCitation>
). At the same time, it cannot be a synonym of
<taxonomicName id="4AF04D778B2F736FFCD0D7A6FC7C8DD5" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[826,972,1716,1742]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2F736FFCD0D7A6FC7C8DD5" box="[826,972,1716,1742]" italics="true" pageId="29" pageNumber="180">N. subcaecus</emphasis>
</taxonomicName>
also, because Thomas Broun writes in his description of
<taxonomicName id="4AF04D778B2F736FFEDCD7CAFE0E8DE9" box="[310,446,1753,1778]" class="Insecta" family="Carabidae" genus="Tachys" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="BF84EAE68B2F736FFEDCD7CAFE0E8DE9" box="[310,446,1753,1778]" italics="true" pageId="29" pageNumber="180">T. coriaceus</emphasis>
</taxonomicName>
that the specimen has “broader form, evidently more transverse thorax” than
<taxonomicName id="4AF04D778B2F736FFAFFD7CAFA258DE9" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[1301,1429,1752,1778]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B2F736FFAFFD7CAFA258DE9" box="[1301,1429,1752,1778]" italics="true" pageId="29" pageNumber="180">N. oreobius</emphasis>
</taxonomicName>
, described by him 15 years earlier. According to our data
<taxonomicName id="4AF04D778B2F736FFCFFD7EEFC188C0D" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[789,936,1788,1814]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2F736FFCFFD7EEFC188C0D" box="[789,936,1788,1814]" italics="true" pageId="29" pageNumber="180">N. subcaecus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4AF04D778B2F736FFB4ED7EEFA968C0D" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[1188,1318,1788,1814]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B2F736FFB4ED7EEFA968C0D" box="[1188,1318,1788,1814]" italics="true" pageId="29" pageNumber="180">N. oreobius</emphasis>
</taxonomicName>
by having narrower proportions (cf. pronotal proportions WPm/
<collectionCode id="EBE1AE318B2F736FFD00D632FCBA8C20" box="[746,778,1825,1851]" country="Russia" name="Laboratory of Palaeontology" pageId="29" pageNumber="180">LP</collectionCode>
1.22±0.025 vs 1.26±0.021, and elytral proportions WE/
<collectionCode id="EBE1AE318B2F736FFA90D632FA2C8C20" box="[1402,1436,1825,1851]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="29" pageNumber="180">LE</collectionCode>
0.65±0.020 vs 0.69±0.012, for
<taxonomicName id="4AF04D778B2F736FFE1CD656FD348C45" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[502,644,1860,1886]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2F736FFE1CD656FD348C45" box="[502,644,1860,1886]" italics="true" pageId="29" pageNumber="180">N subcaecus</emphasis>
</taxonomicName>
and
<taxonomicName id="4AF04D778B2F736FFD56D656FC8E8C45" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[700,830,1860,1886]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="BF84EAE68B2F736FFD56D656FC8E8C45" box="[700,830,1860,1886]" italics="true" pageId="29" pageNumber="180">N. oreobius</emphasis>
</taxonomicName>
subsequently). In addition, in the description the size of the
<typeStatus id="524B88568B2F736FFF0AD67AFEA08C98" box="[224,272,1897,1923]" pageId="29" pageNumber="180">type</typeStatus>
specimen of
<taxonomicName id="4AF04D778B2F736FFE42D67AFD828C99" box="[424,562,1897,1922]" class="Insecta" family="Carabidae" genus="Tachys" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="BF84EAE68B2F736FFE42D67AFD828C99" box="[424,562,1897,1922]" italics="true" pageId="29" pageNumber="180">T. coriaceus</emphasis>
</taxonomicName>
is mentioned as
<quantity id="4A089B118B2F736FFD1BD67BFCE58C98" box="[753,853,1896,1923]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="29" pageNumber="180" unit="mm" value="1.85">1.85 mm</quantity>
(“7/8 line”,
<bibRefCitation id="E9614B058B2F736FFC09D67AFBDC8C99" author="Broun, T." box="[995,1132,1896,1923]" pageId="29" pageNumber="180" pagination="405 - 422" refId="ref26248" refString="Broun, T. (1908) Descriptions of new species of New Zealand Coleoptera. Annals and Magazine of Natural History, Series 8, 2, 405 - 422. https: // doi. org / 10.1080 / 00222930808692504" type="journal article" year="1908">Broun 1908</bibRefCitation>
); this unequivocally points to affinity of the
<typeStatus id="524B88568B2F736FFEBFD69EFE358CBC" box="[341,389,1933,1959]" pageId="29" pageNumber="180">type</typeStatus>
to the group of large species of
<taxonomicName id="4AF04D778B2F736FFD05D69FFC348CBD" authorityName="Jeannel" authorityYear="1937" box="[751,900,1932,1958]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B2F736FFD05D69FFC348CBD" box="[751,900,1932,1958]" italics="true" pageId="29" pageNumber="180">Nesamblyops</emphasis>
</taxonomicName>
. In examined material none of the large species of
<taxonomicName id="4AF04D778B2F736FFF52D6A3FEFD8CD1" authorityName="Jeannel" authorityYear="1937" box="[184,333,1968,1994]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B2F736FFF52D6A3FEFD8CD1" box="[184,333,1968,1994]" italics="true" pageId="29" pageNumber="180">Nesamblyops</emphasis>
</taxonomicName>
has been collected in this area, hence at present the name
<taxonomicName id="4AF04D778B2F736FFC18D6A2FBCC8CD1" box="[1010,1148,1969,1994]" class="Insecta" family="Carabidae" genus="Tachys" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="BF84EAE68B2F736FFC18D6A2FBCC8CD1" box="[1010,1148,1969,1994]" italics="true" pageId="29" pageNumber="180">T. coriaceus</emphasis>
</taxonomicName>
cannot be attributed to a particular local species with certainty.
</paragraph>
<caption id="D98F667C8B2C736CFF7DD446FACF8D96" ID-DOI="http://doi.org/10.5281/zenodo.10184255" ID-Zenodo-Dep="10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="30" pageNumber="181" startId="30.[151,250,1365,1389]" targetBox="[157,1433,187,1336]" targetPageId="30" targetType="figure">
<paragraph id="8D4F36F48B2C736CFF7DD446FACF8D96" blockId="30.[151,1437,1365,1677]" pageId="30" pageNumber="181">
<emphasis id="BF84EAE68B2C736CFF7DD446FE918E76" bold="true" box="[151,289,1365,1389]" pageId="30" pageNumber="181">FIGURE 14.</emphasis>
Line drawings of male genitalia of New Zealand
<taxonomicName id="4AF04D778B2C736CFCC3D446FBB88E76" box="[809,1032,1365,1389]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BF84EAE68B2C736CFCC3D446FC008E76" box="[809,944,1365,1389]" italics="true" pageId="30" pageNumber="181">Nesamblyops</emphasis>
species.
</taxonomicName>
<taxonomicName id="4AF04D778B2C736CFBE5D445FB1B8E76" authorityName="Sokolov" authorityYear="2023" box="[1039,1195,1366,1389]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="canaanensis">
<emphasis id="BF84EAE68B2C736CFBE5D445FB1B8E76" box="[1039,1195,1366,1389]" italics="true" pageId="30" pageNumber="181">N. canaanensis</emphasis>
</taxonomicName>
(Canaan, Nelson, SO):
<emphasis id="BF84EAE68B2C736CFF7DD46AFF198E8A" bold="true" box="[151,169,1401,1425]" pageId="30" pageNumber="181">A</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B2C736CFDE7D46AFDAE8E8A" bold="true" box="[525,542,1401,1425]" pageId="30" pageNumber="181">B</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B2C736CFC74D46AFC008E8A" bold="true" box="[926,944,1401,1425]" pageId="30" pageNumber="181">C</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B2C736CFAF2D469FE778EAD" authority="(Fletchers Creek, Buller, SO)" baseAuthorityName="Fletchers Creek, Buller, SO" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="disjunctus">
<emphasis id="BF84EAE68B2C736CFAF2D469FA2C8E8A" box="[1304,1436,1401,1425]" italics="true" pageId="30" pageNumber="181">N. disjunctus</emphasis>
(Fletchers Creek, Buller, SO)
</taxonomicName>
:
<emphasis id="BF84EAE68B2C736CFE3ED48EFE568EAE" bold="true" box="[468,486,1437,1461]" pageId="30" pageNumber="181">D</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B2C736CFCBED48EFCD58EAE" bold="true" box="[852,869,1437,1461]" pageId="30" pageNumber="181">E</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B2C736CFB05D48EFB4E8EAE" bold="true" box="[1263,1278,1437,1461]" pageId="30" pageNumber="181">F</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B2C736CFE8DD4D1FE788EC2" authorityName="Sokolov" authorityYear="2023" box="[359,456,1473,1497]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="hobbit">
<emphasis id="BF84EAE68B2C736CFE8DD4D1FE788EC2" box="[359,456,1473,1497]" italics="true" pageId="30" pageNumber="181">N. hobbit</emphasis>
</taxonomicName>
(Mount Domett, Nelson, SO):
<emphasis id="BF84EAE68B2C736CFCF8D4D2FC968EC2" bold="true" box="[786,806,1473,1497]" pageId="30" pageNumber="181">G</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B2C736CFB73D4D2FB1D8EC2" bold="true" box="[1177,1197,1473,1497]" pageId="30" pageNumber="181">H</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B2C736CFEC4D4F6FE888EE6" bold="true" box="[302,312,1509,1533]" pageId="30" pageNumber="181">I</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B2C736CFD45D4F5FC988EE6" authorityName="Sokolov" authorityYear="2023" box="[687,808,1509,1533]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="karamea">
<emphasis id="BF84EAE68B2C736CFD45D4F5FC988EE6" box="[687,808,1509,1533]" italics="true" pageId="30" pageNumber="181">N. karamea</emphasis>
</taxonomicName>
(Karamea River Gorge, Nelson, SO):
<emphasis id="BF84EAE68B2C736CFB56D4F6FB798EE6" bold="true" box="[1212,1225,1509,1533]" pageId="30" pageNumber="181">J</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B2C736CFEC5D71AFEF38D3A" bold="true" box="[303,323,1545,1569]" pageId="30" pageNumber="181">K</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B2C736CFD3ED71AFD558D3A" bold="true" box="[724,741,1545,1569]" pageId="30" pageNumber="181">L</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B2C736CFBB5D719FB628D3A" authorityName="Sokolov" authorityYear="2023" box="[1119,1234,1545,1569]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="BF84EAE68B2C736CFBB5D719FB628D3A" box="[1119,1234,1545,1569]" italics="true" pageId="30" pageNumber="181">N. kuscheli</emphasis>
</taxonomicName>
(Flora Hut, Mount Arthur, Nelson, SO):
<emphasis id="BF84EAE68B2C736CFE87D73EFE358D5E" bold="true" box="[365,389,1581,1605]" pageId="30" pageNumber="181">M</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B2C736CFD0FD73EFD478D5E" bold="true" box="[741,759,1581,1605]" pageId="30" pageNumber="181">N</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B2C736CFB9ED73EFB388D5E" bold="true" box="[1140,1160,1581,1605]" pageId="30" pageNumber="181">O</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B2C736CFF09D741FCAC8D71" authority="(Allison Reserve, Akatore, Dunedin, SO)" baseAuthorityName="Allison Reserve, Akatore, Dunedin, SO" box="[227,796,1617,1642]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="magnificus">
<emphasis id="BF84EAE68B2C736CFF09D741FEC08D72" box="[227,368,1617,1641]" italics="true" pageId="30" pageNumber="181">N. magnificus</emphasis>
(Allison Reserve, Akatore, Dunedin, SO)
</taxonomicName>
:
<emphasis id="BF84EAE68B2C736CFCCDD742FC868D72" bold="true" box="[807,822,1617,1641]" pageId="30" pageNumber="181">P</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B2C736CFB74D742FB028D72" bold="true" box="[1182,1202,1617,1641]" pageId="30" pageNumber="181">Q</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B2C736CFEC3D766FE8B8D96" bold="true" box="[297,315,1653,1677]" pageId="30" pageNumber="181">R</emphasis>
—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8D4F36F48B2C736CFF2DD7A8FAE78DE2" blockId="30.[151,1436,1723,1966]" pageId="30" pageNumber="181">
The original description of
<taxonomicName id="4AF04D778B2C736CFDF9D7AFFD1D8DCE" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[531,685,1723,1749]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2C736CFDF9D7AFFD1D8DCE" box="[531,685,1723,1749]" italics="true" pageId="30" pageNumber="181">N. subcaecus</emphasis>
</taxonomicName>
contains little diagnostic information that would allow correct identification. Below, I redescribe the species to make comparison of
<taxonomicName id="4AF04D778B2C736CFC4ED7F3FB878DE2" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[932,1079,1759,1785]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2C736CFC4ED7F3FB878DE2" box="[932,1079,1759,1785]" italics="true" pageId="30" pageNumber="181">N. subcaecus</emphasis>
</taxonomicName>
with other species easier.
</paragraph>
<paragraph id="8D4F36F48B2C736CFF2DD617FC438C05" blockId="30.[151,1436,1723,1966]" box="[199,1011,1795,1822]" pageId="30" pageNumber="181">
<materialsCitation id="3D983CA98B2C736CFF2DD617FC5F8C05" box="[199,1007,1795,1822]" country="New Zealand" location="South Island" pageId="30" pageNumber="181" specimenCount="1" stateProvince="West Coast" typeStatus="holotype">
<emphasis id="BF84EAE68B2C736CFF2DD617FED68C06" bold="true" box="[199,358,1795,1822]" pageId="30" pageNumber="181">
<typeStatus id="524B88568B2C736CFF2DD617FEB28C05" box="[199,258,1796,1822]" pageId="30" pageNumber="181">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F5E776648B2C736CFE86D617FDB58C06" box="[364,517,1795,1822]" name="New Zealand" pageId="30" pageNumber="181">New Zealand</collectingCountry>
,
<location id="882F602F8B2C736CFDFAD610FD108C06" LSID="urn:lsid:plazi:treatment:055987E28B2F736DFF7DD2D6FB388F66:882F602F8B2C736CFDFAD610FD108C06" box="[528,672,1795,1821]" country="New Zealand" name="South Island" pageId="30" pageNumber="181" stateProvince="West Coast">South Island</location>
,
<collectingRegion id="4F34F8168B2C736CFD41D617FC9C8C05" box="[683,812,1796,1822]" country="New Zealand" name="West Coast" pageId="30" pageNumber="181">West Coast</collectingRegion>
,
<taxonomicName id="4AF04D778B2C736CFCDDD610FC078C06" box="[823,951,1795,1821]" domain="Greymouth" pageId="30" pageNumber="180" rank="domain">Greymouth</taxonomicName>
area
</materialsCitation>
.
</paragraph>
<paragraph id="8D4F36F48B2C736CFF2DD634FE488C92" blockId="30.[151,1436,1723,1966]" pageId="30" pageNumber="181">
<emphasis id="BF84EAE68B2C736CFF2DD634FEED8C5A" bold="true" box="[199,349,1831,1857]" pageId="30" pageNumber="181">Recognition.</emphasis>
Adults of this species (
<figureCitation id="15CB2A718B2C736CFD6AD634FD578C59" box="[640,743,1831,1858]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="30" pageNumber="181">Fig. 10F</figureCitation>
) are practically indistinguishable from the adults of many
<taxonomicName id="4AF04D778B2C736CFF7DD658FE398C7E" box="[151,393,1867,1893]" pageId="30" pageNumber="181">
<emphasis id="BF84EAE68B2C736CFF7DD658FE9C8C7E" box="[151,300,1867,1893]" italics="true" pageId="30" pageNumber="181">Nesamblyops</emphasis>
species
</taxonomicName>
based on external characters (e.g.,
<figureCitation id="15CB2A718B2C736CFCC6D658FC098C7D" box="[812,953,1867,1894]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="30" pageNumber="181">Figs 10DE</figureCitation>
) and are distinguished from them by the structures of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2C736DFF2DD680FB388F66" lastPageId="31" lastPageNumber="182" pageId="30" pageNumber="181" type="description">
<paragraph id="8D4F36F48B2C736CFF2DD680FB2E8CB5" blockId="30.[151,1436,1723,1966]" box="[199,1182,1939,1966]" pageId="30" pageNumber="181">
<emphasis id="BF84EAE68B2C736CFF2DD680FEE98CB6" bold="true" box="[199,345,1939,1965]" pageId="30" pageNumber="181">Description.</emphasis>
Small for genus (SBL range
<quantity id="4A089B118B2C736CFD4AD680FCF58CB5" box="[672,837,1939,1966]" metricMagnitude="-3" metricUnit="m" metricValue="1.605" metricValueMax="1.7" metricValueMin="1.51" pageId="30" pageNumber="181" unit="mm" value="1.605" valueMax="1.7" valueMin="1.51">1.511.70 mm</quantity>
, mean 1.62±
<quantity id="4A089B118B2C736CFC38D680FBF88CB5" box="[978,1096,1939,1966]" metricMagnitude="-5" metricUnit="m" metricValue="6.2" pageId="30" pageNumber="181" unit="mm" value="0.062">0.062 mm</quantity>
, n=14).
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD184FD708BE2" blockId="31.[151,1438,151,1149]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD184FE978BAA" box="[199,295,151,177]" italics="true" pageId="31" pageNumber="182">Habitus.</emphasis>
Body form (
<figureCitation id="15CB2A718B2D736DFE25D184FD9A8BAA" box="[463,554,151,177]" captionStart="FIGURE 2" captionStartId="6.[151,250,1585,1609]" captionTargetBox="[189,1397,181,1561]" captionTargetId="figure-32@6.[189,1397,181,1561]" captionTargetPageId="6" captionText="FIGURE 2. SEM illustrations of structural features of labial complex of Anillini: AB—subtribe Nesamblyopina; A— Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Zeanillus punctigerus (Broun); D—Pelodiaetodes insularis Sokolov; E—Anillinus forthoodensis Sokolov and Reddell; F—Geocharidius minimus Sokolov and Kavanaugh. Legend: ca—cardo, bs—basistipes, bss—basal stipital setae, mss—medial stipital setae, pf—palpifer. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184228" httpUri="https://zenodo.org/record/10184228/files/figure.png" pageId="31" pageNumber="182">Fig. 2F</figureCitation>
) moderately convex, elongate ovoid, general proportions wide (WE/SBL 0.39±0.009), head wide relative to pronotum (WH/WPm 0.76±0.023), proportions of pronotum in comparison to elytra average for genus (WPm/WE 0.73±0.015).
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD010FCFC8A06" blockId="31.[151,1438,151,1149]" box="[199,844,259,285]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD010FEBD8A06" box="[199,269,259,285]" italics="true" pageId="31" pageNumber="182">Color.</emphasis>
Body color rufotestaceous, appendages testaceous.
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD034FB198AB6" blockId="31.[151,1438,151,1149]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD034FEF18A5A" box="[199,321,295,321]" italics="true" pageId="31" pageNumber="182">Prothorax.</emphasis>
Pronotum (
<figureCitation id="15CB2A718B2D736DFE2CD034FD9A8A5A" box="[454,554,295,321]" captionStart="FIGURE 13" captionStartId="28.[151,250,934,958]" captionTargetBox="[157,1430,181,910]" captionTargetId="figure-32@28.[157,1430,181,910]" captionTargetPageId="28" captionText="FIGURE 13. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. rotundicollis (Onekaka, Nelson, SO), B—N. solitarius (Italian Creek, Capleston, Buller, SO), C—N. subcaecus (Greymouth, Buller, SO), D—N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO), E—N. viator (Secretary Island, Grono Bay, Fiordland, SO), F—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184253" httpUri="https://zenodo.org/record/10184253/files/figure.png" pageId="31" pageNumber="182">Fig. 13C</figureCitation>
) moderately long in comparison to elytra (LP/LE 0.39±0.010) and moderately transverse (WPm/LP 1.22±0.025), with lateral margins rectilinear constricted posteriorly (WPm/WPp 1.34±0.047). Anterior angles indistinct, posterior angles obtuse (123134°), widely rounded. Width between posterior angles equals width between anterior angles (WPa/WPp 1.00±0.040). Basal margin slightly convex.
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD0A4FCD98902" blockId="31.[151,1438,151,1149]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD0A4FEA58ACA" box="[199,277,439,465]" italics="true" pageId="31" pageNumber="182">Elytra.</emphasis>
Ovoid, moderately depressed along suture, comparatively long (LE/SBL 0.60±0.009) and moderately narrow (WE/LE 0.65±0.020). Humeri completely rounded. Lateral margins slightly divergent at basal half, subparallel at middle and evenly rounded to apex in apical third.
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD330FE798846" blockId="31.[151,1438,151,1149]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD330FEC78926" box="[199,375,547,573]" italics="true" pageId="31" pageNumber="182">Male genitalia.</emphasis>
Median lobe of aedeagus (
<figureCitation id="15CB2A718B2D736DFD52D330FCAF8926" box="[696,799,546,573]" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="31" pageNumber="182">Fig. 15R</figureCitation>
) moderately arcuate and slightly twisted. Shaft almost subparallel in basal half, moderately tapering in apical half. Apex moderately curved ventrally with tapering narrowly rounded tip. Apical orifice of moderate length occupies one third of shaft length. Ventral margin of median lobe straight in basal two thirds, slightly curved downward in apical third. Walls of shaft with several poriferous canals scattered in apical and basal parts of shaft. Dorsal copulatory sclerites atypical for genus. Only one weakly sclerotized field present at the middle of shaft in a form of short, wide, and curved stripe (
<figureCitation id="15CB2A718B2D736DFB48D3C4FAB789EA" box="[1186,1287,726,753]" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="31" pageNumber="182">Fig. 15R</figureCitation>
). V
<emphasis id="BF84EAE68B2D736DFADED3C4FA8D89EA" bold="true" box="[1332,1341,727,753]" pageId="31" pageNumber="182">-</emphasis>
and rC-sclerites lacking. Left paramere (
<figureCitation id="15CB2A718B2D736DFDE6D3E8FDDE880E" box="[524,622,762,789]" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="31" pageNumber="182">Fig. 15P</figureCitation>
) wide, apex not attenuate, bearing two long setae. Right paramere (
<figureCitation id="15CB2A718B2D736DFA85D3E8FF788822" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="31" pageNumber="182">Fig. 15Q</figureCitation>
) narrow, of moderate length, bearing three long setae, which are slightly shorter than the length of paramere. Ring sclerite as in
<figureCitation id="15CB2A718B2D736DFE82D250FE758846" box="[360,453,835,861]" captionStart="FIGURE 17" captionStartId="34.[151,250,1191,1215]" captionTargetBox="[163,1421,187,1168]" captionTargetId="figure-32@34.[151,1436,181,1168]" captionTargetPageId="34" captionText="FIGURE 17. Line drawings of ring sclerite of New Zealand Nesamblyops species, male genitalia, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (Mount Domett, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO), G—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), H—N. ovipennis (Mount Arthur, Nelson, SO), I—N. rotundicollis (Onekaka, Nelson, SO), J—N. subcaecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO), K—N. viator (Secretary Island, Grono Bay, Fjordland, SO), L—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184261" httpUri="https://zenodo.org/record/10184261/files/figure.png" pageId="31" pageNumber="182">Fig. 17J</figureCitation>
.
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD274FD29889A" blockId="31.[151,1438,151,1149]" box="[199,665,871,897]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD274FE5D889A" box="[199,493,871,897]" italics="true" pageId="31" pageNumber="182">Female internal genitalia.</emphasis>
Not examined.
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD299FCBF88F6" blockId="31.[151,1438,151,1149]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD299FDBA88BF" bold="true" box="[199,522,906,932]" pageId="31" pageNumber="182">Geographical distribution.</emphasis>
The range of the species occupies foothills and a part of the Southern Alps approximately from the catchment of Moa Stream and the Arthurs Pass area in the central
<collectingRegion id="4F34F8168B2D736DFB61D2BCFAB988D2" box="[1163,1289,943,969]" country="New Zealand" name="Canterbury" pageId="31" pageNumber="182">Canterbury</collectingRegion>
to the mouth of Grey River in the central
<collectingRegion id="4F34F8168B2D736DFE38D2C0FD2F88F6" box="[466,671,979,1005]" country="New Zealand" name="West Coast" pageId="31" pageNumber="182">West Coast region</collectingRegion>
(
<figureCitation id="15CB2A718B2D736DFD44D2C0FCB288F6" box="[686,770,979,1005]" captionStart="FIGURE 21" captionStartId="37.[151,250,1860,1884]" captionTargetBox="[151,1437,1100,1836]" captionTargetId="figure-68@37.[151,1437,1100,1836]" captionTargetPageId="37" captionText="FIGURE 21. Distributional records for the Nesamblyops subcaecus and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Red star—the type locality of Cillenum subcaecum Sharp; black star—the type locality of Tachys coriaceus Broun. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184271" httpUri="https://zenodo.org/record/10184271/files/figure.png" pageId="31" pageNumber="182">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD2E5FB458F0A" blockId="31.[151,1438,151,1149]" box="[199,1269,1014,1041]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD2E5FE9A8F0B" bold="true" box="[199,298,1014,1040]" pageId="31" pageNumber="182">Habitat.</emphasis>
Specimens were collected from moss, litter, leaf mold, litter and [plant] mats samples.
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD509FB388F66" blockId="31.[151,1438,151,1149]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD509FEC08F2F" bold="true" box="[199,368,1050,1076]" pageId="31" pageNumber="182">Relationships.</emphasis>
The structure of the male genitalia of
<taxonomicName id="4AF04D778B2D736DFCC8D508FC068F2E" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[802,950,1051,1077]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2D736DFCC8D508FC068F2E" box="[802,950,1051,1077]" italics="true" pageId="31" pageNumber="182">N. subcaecus</emphasis>
</taxonomicName>
suggests its relatedness to the species with reduced and simple armature of the internal sac. In having only one sclerotized copulatory sclerite
<taxonomicName id="4AF04D778B2D736DFAECD52CFA2C8F42" authorityName="Greymouth, Buller, SO" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[1286,1436,1087,1113]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="subcaecus">
<emphasis id="BF84EAE68B2D736DFAECD52CFA2C8F42" box="[1286,1436,1087,1113]" italics="true" pageId="31" pageNumber="182">N. subcaecus</emphasis>
</taxonomicName>
demonstrates affinity to and presumably forms one group with
<taxonomicName id="4AF04D778B2D736DFCB0D570FC0D8F67" authorityName="Sokolov" authorityYear="2023" box="[858,957,1123,1148]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="viator">
<emphasis id="BF84EAE68B2D736DFCB0D570FC0D8F67" box="[858,957,1123,1148]" italics="true" pageId="31" pageNumber="182">N. viator</emphasis>
</taxonomicName>
, described below.
</paragraph>
</subSubSection>
</treatment>
</document>