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<document id="0DD9087C1D43006DAA2177B0FD1094E1" ID-DOI="10.11646/zootaxa.5375.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10184224" ID-ZooBank="F3D0E008-556C-4FAD-BF51-4F1A714325DA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700642529072" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2023" docId="055987E28B2E736FFF7DD56FFDF7889A" docLanguage="en" docName="zootaxa.5375.2.1.pdf" docOrigin="Zootaxa 5375 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.2.1/52321" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nesamblyops solitarius Sokolov 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="180" masterDocId="F960FF9A8B327372FFEAD113FFB08B1B" masterDocTitle="Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe" masterLastPageNumber="192" masterPageNumber="151" pageNumber="179" updateTime="1700643183691" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="066D0D186FF733664161B047CB266AB2">Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe</mods:title>
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<mods:namePart id="8F3EF446BC7ADA2EFD2BD2262FA620F0">Sokolov, Igor M.</mods:namePart>
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<mods:title id="0BDFEE3EFAF8E77A9F6BBCC4BF7B20BF">Zootaxa</mods:title>
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<mods:date id="05712EED66232168DB120D6D1EF6C44D">2023</mods:date>
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<mods:number id="DCEBD2E6F1EC88A25C97F2F12DA4958B">2023-11-22</mods:number>
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<mods:identifier id="7482974A88ADA64A04FB66EC7A8B1098" type="DOI">10.11646/zootaxa.5375.2.1</mods:identifier>
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<treatment id="055987E28B2E736FFF7DD56FFDF7889A" LSID="urn:lsid:plazi:treatment:055987E28B2E736FFF7DD56FFDF7889A" httpUri="http://treatment.plazi.org/id/055987E28B2E736FFF7DD56FFDF7889A" lastPageId="29" lastPageNumber="180" pageId="28" pageNumber="179">
<subSubSection id="C5EA657F8B2E736EFF7DD56FFDAB8F8D" box="[151,539,1148,1175]" pageId="28" pageNumber="179" type="nomenclature">
<paragraph id="8D4F36F48B2E736EFF7DD56FFDAB8F8D" blockId="28.[151,539,1148,1211]" box="[151,539,1148,1175]" pageId="28" pageNumber="179">
<heading id="D60781988B2E736EFF7DD56FFDAB8F8D" bold="true" box="[151,539,1148,1175]" fontSize="11" level="1" pageId="28" pageNumber="179" reason="1">
<emphasis id="BF84EAE68B2E736EFF7DD56FFDAB8F8D" bold="true" box="[151,539,1148,1175]" pageId="28" pageNumber="179">
<taxonomicName id="4AF04D778B2E736EFF7DD56FFE1E8F8C" authority="Sokolov, 2023" authorityName="Sokolov" authorityYear="2023" box="[151,430,1148,1175]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="solitarius" status="sp. nov.">
<emphasis id="BF84EAE68B2E736EFF7DD56FFE1E8F8C" bold="true" box="[151,430,1148,1175]" italics="true" pageId="28" pageNumber="179">Nesamblyops solitarius</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A4B7579D8B2E736EFE57D56FFDAB8F8D" box="[445,539,1148,1174]" pageId="28" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2E736EFF7DD5B3FE428FA1" box="[151,498,1184,1211]" pageId="28" pageNumber="179" type="description">
<paragraph id="8D4F36F48B2E736EFF7DD5B3FE428FA1" blockId="28.[151,539,1148,1211]" box="[151,498,1184,1211]" pageId="28" pageNumber="179">
<figureCitation id="15CB2A718B2E736EFF7DD5B3FE998FA1" box="[151,297,1184,1210]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="28" pageNumber="179">Figures. 10E</figureCitation>
,
<figureCitation id="15CB2A718B2E736EFEDED5B3FED58FA1" box="[308,357,1184,1210]" captionStart="FIGURE 13" captionStartId="28.[151,250,934,958]" captionTargetBox="[157,1430,181,910]" captionTargetId="figure-32@28.[157,1430,181,910]" captionTargetPageId="28" captionText="FIGURE 13. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. rotundicollis (Onekaka, Nelson, SO), B—N. solitarius (Italian Creek, Capleston, Buller, SO), C—N. subcaecus (Greymouth, Buller, SO), D—N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO), E—N. viator (Secretary Island, Grono Bay, Fiordland, SO), F—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184253" httpUri="https://zenodo.org/record/10184253/files/figure.png" pageId="28" pageNumber="179">13B</figureCitation>
,
<figureCitation id="15CB2A718B2E736EFE9BD5B3FE788FA1" box="[369,456,1184,1211]" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="28" pageNumber="179">15MO</figureCitation>
,
<figureCitation id="15CB2A718B2E736EFE3CD5B3FE428FA1" box="[470,498,1184,1210]" captionStart="FIGURE 20" captionStartId="37.[151,250,944,968]" captionTargetBox="[151,1436,181,920]" captionTargetId="figure-15@37.[151,1436,181,920]" captionTargetPageId="37" captionText="FIGURE 20. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Green circles—N. solitarius; black circles—N. montanus; pink circles— N. rotundicollis; red circles—N. subrufus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184269" httpUri="https://zenodo.org/record/10184269/files/figure.png" pageId="28" pageNumber="179">20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2E736EFF7DD5FBFE848E89" pageId="28" pageNumber="179" type="materials_examined">
<paragraph id="8D4F36F48B2E736EFF7DD5FBFE848E89" blockId="28.[151,1438,1256,1967]" pageId="28" pageNumber="179">
<emphasis id="BF84EAE68B2E736EFF7DD5FBFEF88E19" bold="true" box="[151,328,1256,1282]" pageId="28" pageNumber="179">Type material.</emphasis>
<materialsCitation id="3D983CA98B2E736EFEB8D5FBFA348E3C" collectingDate="1971-11" collectionCode="NZAC, BR, DSIR" collectorName="J. McBurney Litter &amp; Beech Forest Utilization Project &amp; No" country="New Zealand" location="Boatmans Ck" municipality="Beech Forest" pageId="28" pageNumber="179" specimenCount="1" typeStatus="holotype">
<emphasis id="BF84EAE68B2E736EFEB8D5FBFE578E19" bold="true" box="[338,487,1256,1282]" italics="true" pageId="28" pageNumber="179">
<typeStatus id="524B88568B2E736EFEB8D5FBFE578E19" box="[338,487,1256,1282]" pageId="28" pageNumber="179" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
, male, in
<collectionCode id="EBE1AE318B2E736EFD8AD5FBFD1F8E19" box="[608,687,1256,1282]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="28" pageNumber="179" type="Other institutional type">NZAC</collectionCode>
, labeled, dissected: \
<collectingCountry id="F5E776648B2E736EFC43D5FAFBC78E19" box="[937,1143,1256,1283]" name="New Zealand" pageId="28" pageNumber="179">NEW ZEALAND</collectingCountry>
<collectionCode id="EBE1AE318B2E736EFB6BD5FAFB168E18" box="[1153,1190,1257,1283]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="28" pageNumber="179">BR</collectionCode>
<location id="882F602F8B2E736EFB5BD5FAFAFC8E19" LSID="urn:lsid:plazi:treatment:055987E28B2E736FFF7DD56FFDF7889A:882F602F8B2E736EFB5BD5FAFAFC8E19" box="[1201,1356,1256,1283]" country="New Zealand" municipality="Beech Forest" name="Boatmans Ck" pageId="28" pageNumber="179">Boatmans Ck</location>
<locationDeviation id="9CFD2FD68B2E736EFABFD5FBFEE28E3D" location="Cronodun" pageId="28" pageNumber="179">
<quantity id="4A089B118B2E736EFABFD5FBFA2C8E18" box="[1365,1436,1256,1283]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="28" pageNumber="179" unit="km" value="4.5">4.5km</quantity>
SE of Cronodun
</locationDeviation>
<date id="F94E10348B2E736EFEB6D41EFE7D8E3D" box="[348,461,1292,1319]" pageId="28" pageNumber="179" value="1971-11">
<collectingDate id="E90AE9DC8B2E736EFEB6D41EFE7D8E3D" box="[348,461,1292,1319]" pageId="28" pageNumber="179" value="1971-11">Nov 1971</collectingDate>
</date>
<collectorName id="200553228B2E736EFE3DD41EFD188E3D" box="[471,680,1292,1319]" pageId="28" pageNumber="179">J.McBurney Litter</collectorName>
71/140 \
<collectorName id="200553228B2E736EFCF0D41FFB3A8E3D" box="[794,1162,1292,1319]" pageId="28" pageNumber="179">
<collectingMunicipality id="6D2BAC8E8B2E736EFCF0D41FFC008E3C" box="[794,944,1292,1319]" pageId="28" pageNumber="179">Beech Forest</collectingMunicipality>
Utilization Project
</collectorName>
\
<collectionCode id="EBE1AE318B2E736EFB4FD41FFB558E3D" box="[1189,1253,1292,1318]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="28" pageNumber="179">DSIR</collectionCode>
\
<collectorName id="200553228B2E736EFB1CD41EFAA88E3C" box="[1270,1304,1293,1319]" pageId="28" pageNumber="179">No</collectorName>
\ NZ PB
</materialsCitation>
\.
<materialsCitation id="3D983CA98B2E736EFF7DD423FEAE8E88" collectingDate="1972-04-21" collectionCode="BR, DSIR" collectorName="J. S. Dugdale &amp; Beech Forest Utilization Project &amp; No" country="New Zealand" location="Beech Forest" pageId="28" pageNumber="179" specimenCount="5" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="BF84EAE68B2E736EFF7DD423FE828E51" bold="true" box="[151,306,1328,1354]" italics="true" pageId="28" pageNumber="179">
<typeStatus id="524B88568B2E736EFF7DD423FE828E51" box="[151,306,1328,1354]" pageId="28" pageNumber="179" type="paratype">PARATYPES</typeStatus>
</emphasis>
(
<specimenCount id="9BF6FD7D8B2E736EFEA9D423FE628E51" box="[323,466,1328,1354]" count="2" pageId="28" pageNumber="179" type="generic">2 specimens</specimenCount>
, dissected
<specimenCount id="9BF6FD7D8B2E736EFDA7D423FD378E51" box="[589,647,1328,1354]" count="1" pageId="28" pageNumber="179" type="generic">1 ex.</specimenCount>
),
<specimenCount id="9BF6FD7D8B2E736EFD77D423FD598E51" box="[669,745,1328,1354]" count="1" pageId="28" pageNumber="179" type="male">1 male</specimenCount>
and
<specimenCount id="9BF6FD7D8B2E736EFCC9D423FC358E51" box="[803,901,1328,1354]" count="1" pageId="28" pageNumber="179" type="female">1 female</specimenCount>
labeled: \
<collectingCountry id="F5E776648B2E736EFC14D422FB798E51" box="[1022,1225,1328,1355]" name="New Zealand" pageId="28" pageNumber="179">NEW ZEALAND</collectingCountry>
<collectionCode id="EBE1AE318B2E736EFB38D422FB478E50" box="[1234,1271,1329,1355]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="28" pageNumber="179">BR</collectionCode>
<locationDeviation id="9CFD2FD68B2E736EFB15D423FE3A8E75" location="Capleston Italians Ck" pageId="28" pageNumber="179">
<quantity id="4A089B118B2E736EFB15D423FAF68E51" box="[1279,1350,1328,1354]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="28" pageNumber="179" unit="km" value="1.6">1.6km</quantity>
NW of Capleston Italians Ck
</locationDeviation>
<date id="F94E10348B2E736EFE78D447FDAF8E75" box="[402,543,1364,1390]" pageId="28" pageNumber="179" value="1972-04-21">
<collectingDate id="E90AE9DC8B2E736EFE78D447FDAF8E75" box="[402,543,1364,1390]" pageId="28" pageNumber="179" value="1972-04-21">21 Apr 1972</collectingDate>
</date>
\
<collectorName id="200553228B2E736EFDDDD447FD768E75" box="[567,710,1364,1390]" pageId="28" pageNumber="179">J.S. Dugdale</collectorName>
moss and litter 71/98 \
<collectorName id="200553228B2E736EFC39D447FA8D8E75" box="[979,1341,1364,1391]" pageId="28" pageNumber="179">
<location id="882F602F8B2E736EFC39D447FBD78E74" LSID="urn:lsid:plazi:treatment:055987E28B2E736FFF7DD56FFDF7889A:882F602F8B2E736EFC39D447FBD78E74" box="[979,1127,1364,1391]" country="New Zealand" name="Beech Forest" pageId="28" pageNumber="179">Beech Forest</location>
Utilization Project
</collectorName>
\
<collectionCode id="EBE1AE318B2E736EFABFD447FA258E75" box="[1365,1429,1364,1390]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="28" pageNumber="179">DSIR</collectionCode>
\
<collectorName id="200553228B2E736EFF7DD46AFF098E88" box="[151,185,1401,1427]" pageId="28" pageNumber="179">No</collectorName>
\ NZ PB
</materialsCitation>
\.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2E736EFF2DD48FFA928EC0" pageId="28" pageNumber="179" type="etymology">
<paragraph id="8D4F36F48B2E736EFF2DD48FFA928EC0" blockId="28.[151,1438,1256,1967]" pageId="28" pageNumber="179">
<emphasis id="BF84EAE68B2E736EFF2DD48FFEFE8EAD" bold="true" box="[199,334,1436,1462]" pageId="28" pageNumber="179">Etymology.</emphasis>
The specific epithet is a Latin adjective,
<taxonomicName id="4AF04D778B2E736EFCFBD48FFCCB8EAD" authorityName="Sokolov" authorityYear="2023" box="[785,891,1436,1462]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="BF84EAE68B2E736EFCFBD48FFCCB8EAD" box="[785,891,1436,1462]" italics="true" pageId="28" pageNumber="179">solitarius</emphasis>
</taxonomicName>
(meaning “living alone”), in the masculine form, and is alluding to the small number of specimens of this species, which are known to science at present.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2E736EFF2DD4F7FE488D71" pageId="28" pageNumber="179" type="materials_examined">
<paragraph id="8D4F36F48B2E736EFF2DD4F7FACB8EE4" blockId="28.[151,1438,1256,1967]" box="[199,1403,1508,1535]" pageId="28" pageNumber="179">
<materialsCitation id="3D983CA98B2E736EFF2DD4F7FAC78EE4" box="[199,1399,1508,1535]" country="New Zealand" county="South Island" location="Capleston area" municipality="Victoria Range" pageId="28" pageNumber="179" specimenCount="1" stateProvince="West Coast" typeStatus="holotype">
<emphasis id="BF84EAE68B2E736EFF2DD4F7FED68EE5" bold="true" box="[199,358,1508,1534]" pageId="28" pageNumber="179">
<typeStatus id="524B88568B2E736EFF2DD4F7FEB28EE5" box="[199,258,1508,1534]" pageId="28" pageNumber="179">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F5E776648B2E736EFE86D4F6FDB58EE5" box="[364,517,1508,1535]" name="New Zealand" pageId="28" pageNumber="179">New Zealand</collectingCountry>
,
<collectingCounty id="642E4E788B2E736EFDFAD4F7FD108EE5" box="[528,672,1508,1534]" pageId="28" pageNumber="179">South Island</collectingCounty>
,
<collectingRegion id="4F34F8168B2E736EFD41D4F6FC9C8EE5" box="[683,812,1508,1535]" country="New Zealand" name="West Coast" pageId="28" pageNumber="179">West Coast</collectingRegion>
, western foothills of
<collectingMunicipality id="6D2BAC8E8B2E736EFBFDD4F7FB728EE4" box="[1047,1218,1508,1535]" pageId="28" pageNumber="179">Victoria Range</collectingMunicipality>
,
<location id="882F602F8B2E736EFB24D4F7FAC78EE4" LSID="urn:lsid:plazi:treatment:055987E28B2E736FFF7DD56FFDF7889A:882F602F8B2E736EFB24D4F7FAC78EE4" box="[1230,1399,1508,1535]" country="New Zealand" county="South Island" municipality="Victoria Range" name="Capleston area" pageId="28" pageNumber="179" stateProvince="West Coast">Capleston area</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8D4F36F48B2E736EFF2DD71BFE488D71" blockId="28.[151,1438,1256,1967]" pageId="28" pageNumber="179">
<emphasis id="BF84EAE68B2E736EFF2DD71BFEED8D39" bold="true" box="[199,349,1544,1570]" pageId="28" pageNumber="179">Recognition.</emphasis>
Adults of this species (
<figureCitation id="15CB2A718B2E736EFD95D71BFD588D39" box="[639,744,1544,1570]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="28" pageNumber="179">Fig. 10E</figureCitation>
) are practically indistinguishable from the adults of many
<taxonomicName id="4AF04D778B2E736EFF7DD73FFE398D5D" box="[151,393,1580,1606]" pageId="28" pageNumber="179">
<emphasis id="BF84EAE68B2E736EFF7DD73FFE9C8D5D" box="[151,300,1580,1606]" italics="true" pageId="28" pageNumber="179">Nesamblyops</emphasis>
species
</taxonomicName>
based on external characters (e.g.,
<figureCitation id="15CB2A718B2E736EFCC1D73FFC0A8D5D" box="[811,954,1580,1606]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="28" pageNumber="179">Figs 10D, F</figureCitation>
) and are distinguished from them by the structures of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2E736FFF2DD767FDF7889A" lastPageId="29" lastPageNumber="180" pageId="28" pageNumber="179" type="description">
<paragraph id="8D4F36F48B2E736EFF2DD767FB208D95" blockId="28.[151,1438,1256,1967]" box="[199,1168,1652,1679]" pageId="28" pageNumber="179">
<emphasis id="BF84EAE68B2E736EFF2DD767FEE98D95" bold="true" box="[199,345,1652,1678]" pageId="28" pageNumber="179">Description.</emphasis>
Large for genus (SBL range
<quantity id="4A089B118B2E736EFD75D767FCF48D94" box="[671,836,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.08" metricValueMin="2.02" pageId="28" pageNumber="179" unit="mm" value="2.05" valueMax="2.08" valueMin="2.02">2.022.08 mm</quantity>
, mean 2.05±
<quantity id="4A089B118B2E736EFC39D767FBF88D94" box="[979,1096,1652,1679]" metricMagnitude="-5" metricUnit="m" metricValue="4.2" pageId="28" pageNumber="179" unit="mm" value="0.042">0.042 mm</quantity>
, n=2).
</paragraph>
<paragraph id="8D4F36F48B2E736EFF2DD78BFD708DE1" blockId="28.[151,1438,1256,1967]" pageId="28" pageNumber="179">
<emphasis id="BF84EAE68B2E736EFF2DD78BFE978DA9" box="[199,295,1688,1714]" italics="true" pageId="28" pageNumber="179">Habitus.</emphasis>
Body form (
<figureCitation id="15CB2A718B2E736EFE5ED78BFDA68DA9" box="[436,534,1688,1714]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="28" pageNumber="179">Fig. 10E</figureCitation>
) moderately convex, elongate ovoid, general proportions slightly wide (WE/SBL 0.38±0.003), head narrow relative to pronotum (WH/WPm 0.70±0.005), proportions of pronotum in comparison to elytra average for genus (WPm/WE 0.76±0.003).
</paragraph>
<paragraph id="8D4F36F48B2E736EFF2DD617FCFC8C04" blockId="28.[151,1438,1256,1967]" box="[199,844,1796,1823]" pageId="28" pageNumber="179">
<emphasis id="BF84EAE68B2E736EFF2DD617FEBD8C05" box="[199,269,1796,1822]" italics="true" pageId="28" pageNumber="179">Color.</emphasis>
Body color rufotestaceous, appendages testaceous.
</paragraph>
<paragraph id="8D4F36F48B2E736EFF2DD63BFAE98CB4" blockId="28.[151,1438,1256,1967]" pageId="28" pageNumber="179">
<emphasis id="BF84EAE68B2E736EFF2DD63BFEF18C59" box="[199,321,1832,1858]" italics="true" pageId="28" pageNumber="179">Prothorax.</emphasis>
Pronotum (
<figureCitation id="15CB2A718B2E736EFE26D63BFD848C59" box="[460,564,1832,1858]" captionStart="FIGURE 13" captionStartId="28.[151,250,934,958]" captionTargetBox="[157,1430,181,910]" captionTargetId="figure-32@28.[157,1430,181,910]" captionTargetPageId="28" captionText="FIGURE 13. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. rotundicollis (Onekaka, Nelson, SO), B—N. solitarius (Italian Creek, Capleston, Buller, SO), C—N. subcaecus (Greymouth, Buller, SO), D—N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO), E—N. viator (Secretary Island, Grono Bay, Fiordland, SO), F—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184253" httpUri="https://zenodo.org/record/10184253/files/figure.png" pageId="28" pageNumber="179">Fig. 13B</figureCitation>
) moderately long in comparison to elytra (LP/LE 0.42±0.006) and slightly transverse (WPm/LP 1.17±0.012), with lateral margins slightly and rectilinear constricted posteriorly (WPm/ WPp 1.20±0.029). Anterior angles indistinct, posterior angles obtuse (124125°), widely rounded. Width between posterior angles greater than between anterior angles (WPa/WPp 0.83±0.014). Basal margin slightly convex.
</paragraph>
<paragraph id="8D4F36F48B2F736FFF2DD184FCD98BE2" blockId="29.[151,1437,150,897]" pageId="29" pageNumber="180">
<emphasis id="BF84EAE68B2F736FFF2DD184FEA58BAA" box="[199,277,151,177]" italics="true" pageId="29" pageNumber="180">Elytra.</emphasis>
Ovoid, moderately depressed along suture, comparatively long (LE/SBL 0.59±0.012) and moderately narrow (WE/LE 0.64±0.018). Humeri completely rounded. Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third.
</paragraph>
<paragraph id="8D4F36F48B2F736FFF2DD010FE198926" blockId="29.[151,1437,150,897]" pageId="29" pageNumber="180">
<emphasis id="BF84EAE68B2F736FFF2DD010FEC68A06" box="[199,374,259,285]" italics="true" pageId="29" pageNumber="180">Male genitalia.</emphasis>
Median lobe of aedeagus (
<figureCitation id="15CB2A718B2F736FFD5DD010FCAF8A06" box="[695,799,258,285]" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="29" pageNumber="180">Fig. 15O</figureCitation>
) moderately arcuate and slightly twisted. Shaft almost subparallel in basal half, moderately tapering in apical half. Apex long and straight with slightly dilated symmetrical rounded tip. Apical orifice long, occupies almost one half of the shaft length. Ventral margin of median lobe almost straight. Walls of shaft with several poriferous canals scattered across shaft body. Dorsal copulatory sclerites atypical for genus. V-contour and rC-sclerite lacking. Only dorsal membraneous field with narrow sclerotization at its dorsal edge exists at the middle of shaft (
<figureCitation id="15CB2A718B2F736FFDE4D0A4FDC18ACA" box="[526,625,438,465]" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="29" pageNumber="180">Fig. 15O</figureCitation>
). In addition, internal sac has a scaly membraneous field at apical orifice. Left paramere (
<figureCitation id="15CB2A718B2F736FFEAED0C8FE1E8AEE" box="[324,430,474,501]" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="29" pageNumber="180">Fig. 15M</figureCitation>
) comparatively wide, with shortly attenuate apex, bearing two long setae. Right paramere (
<figureCitation id="15CB2A718B2F736FFF75D0ECFEB58902" box="[159,261,510,537]" captionStart="FIGURE 15" captionStartId="32.[151,250,1380,1404]" captionTargetBox="[157,1431,188,1353]" captionTargetId="figure-32@32.[151,1436,181,1356]" captionTargetPageId="32" captionText="FIGURE 15. Line drawings of male genitalia of New Zealand Nesamblyops species. N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. ovipennis (Mount Arthur, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. rotundicollis (Onekaka, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. solitarius (Italian Creek, Capleston, Buller, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. subacecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="29" pageNumber="180">Fig. 15N</figureCitation>
) narrow, of moderate length, bearing two long setae, which are shorter than the length of paramere. Ring sclerite not investigated.
</paragraph>
<paragraph id="8D4F36F48B2F736FFF2DD354FD29897A" blockId="29.[151,1437,150,897]" box="[199,665,583,609]" pageId="29" pageNumber="180">
<emphasis id="BF84EAE68B2F736FFF2DD354FE5D897A" box="[199,493,583,609]" italics="true" pageId="29" pageNumber="180">Female internal genitalia.</emphasis>
Not examined.
</paragraph>
<paragraph id="8D4F36F48B2F736FFF2DD379FF4189D6" blockId="29.[151,1437,150,897]" pageId="29" pageNumber="180">
<emphasis id="BF84EAE68B2F736FFF2DD379FDB5899F" bold="true" box="[199,517,618,644]" pageId="29" pageNumber="180">Geographical distribution.</emphasis>
The range of the species lies in the northern half of the
<collectingRegion id="4F34F8168B2F736FFB96D378FAFC899E" box="[1148,1356,619,645]" country="New Zealand" name="West Coast" pageId="29" pageNumber="180">West Coast region</collectingRegion>
, where the species was collected in two localities around Capleston at the foothills of the Victoria Range (
<figureCitation id="15CB2A718B2F736FFB17D39CFAE289B2" box="[1277,1362,655,681]" captionStart="FIGURE 20" captionStartId="37.[151,250,944,968]" captionTargetBox="[151,1436,181,920]" captionTargetId="figure-15@37.[151,1436,181,920]" captionTargetPageId="37" captionText="FIGURE 20. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Green circles—N. solitarius; black circles—N. montanus; pink circles— N. rotundicollis; red circles—N. subrufus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184269" httpUri="https://zenodo.org/record/10184269/files/figure.png" pageId="29" pageNumber="180">Fig. 20</figureCitation>
, green circles).
</paragraph>
<paragraph id="8D4F36F48B2F736FFF2DD3C5FC1689EA" blockId="29.[151,1437,150,897]" box="[199,934,726,753]" pageId="29" pageNumber="180">
<emphasis id="BF84EAE68B2F736FFF2DD3C5FE9A89EB" bold="true" box="[199,298,726,752]" pageId="29" pageNumber="180">Habitat.</emphasis>
Specimens were collected from moss and litter samples.
</paragraph>
<paragraph id="8D4F36F48B2F736FFF2DD3E9FDF7889A" blockId="29.[151,1437,150,897]" pageId="29" pageNumber="180">
<emphasis id="BF84EAE68B2F736FFF2DD3E9FEC0880F" bold="true" box="[199,368,762,788]" pageId="29" pageNumber="180">Relationships.</emphasis>
The structure of the male genitalia and parameres of
<taxonomicName id="4AF04D778B2F736FFC34D3E8FBDB880E" authorityName="Sokolov" authorityYear="2023" box="[990,1131,763,789]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="BF84EAE68B2F736FFC34D3E8FBDB880E" box="[990,1131,763,789]" italics="true" pageId="29" pageNumber="180">N. solitarius</emphasis>
</taxonomicName>
suggests its relatedness to
<taxonomicName id="4AF04D778B2F736FFF7DD20CFEF88822" authorityName="Sokolov" authorityYear="2023" box="[151,328,799,825]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="BF84EAE68B2F736FFF7DD20CFEF88822" box="[151,328,799,825]" italics="true" pageId="29" pageNumber="180">N. rotundicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="4AF04D778B2F736FFE68D20CFDB48822" authorityName="Sokolov" authorityYear="2023" box="[386,516,799,825]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="subrufus">
<emphasis id="BF84EAE68B2F736FFE68D20CFDB48822" box="[386,516,799,825]" italics="true" pageId="29" pageNumber="180">N. subrufus</emphasis>
</taxonomicName>
, described below. All three species share bisetose parameres and the lack of rC-sclerite in the internal sac of median. Also, all three species have presumably a synapomorphic character—a scaly membraneous field near apical orifice.
</paragraph>
</subSubSection>
</treatment>
</document>