treatments-xml/data/05/59/87/055987E28B2D7353FF7DD5D2FE2F8E52.xml
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<document id="365AA905D21288C9856130FC1CA15AC3" ID-DOI="10.11646/zootaxa.5375.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10184224" ID-ZooBank="F3D0E008-556C-4FAD-BF51-4F1A714325DA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700642529072" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2023" docId="055987E28B2D7353FF7DD5D2FE2F8E52" docLanguage="en" docName="zootaxa.5375.2.1.pdf" docOrigin="Zootaxa 5375 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.2.1/52321" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nesamblyops subrufus Sokolov 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="184" masterDocId="F960FF9A8B327372FFEAD113FFB08B1B" masterDocTitle="Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe" masterLastPageNumber="192" masterPageNumber="151" pageNumber="182" updateTime="1700643183691" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EBDB509A9B76235183909582711F74F8">Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe</mods:title>
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<mods:namePart id="480B7C675C48E6FD0CA67A5E3D0E325F">Sokolov, Igor M.</mods:namePart>
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<mods:date id="33559EB89BF910D7441867647C87AC0A">2023</mods:date>
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<mods:number id="3CF54D0BD62F660E0E0052E0EA73D217">2023-11-22</mods:number>
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<treatment id="055987E28B2D7353FF7DD5D2FE2F8E52" LSID="urn:lsid:plazi:treatment:055987E28B2D7353FF7DD5D2FE2F8E52" httpUri="http://treatment.plazi.org/id/055987E28B2D7353FF7DD5D2FE2F8E52" lastPageId="33" lastPageNumber="184" pageId="31" pageNumber="182">
<subSubSection id="C5EA657F8B2D736DFF7DD5D2FDA68FC0" box="[151,534,1217,1244]" pageId="31" pageNumber="182" type="nomenclature">
<paragraph id="8D4F36F48B2D736DFF7DD5D2FDA68FC0" blockId="31.[151,534,1217,1279]" box="[151,534,1217,1244]" pageId="31" pageNumber="182">
<heading id="D60781988B2D736DFF7DD5D2FDA68FC0" bold="true" box="[151,534,1217,1244]" fontSize="11" level="1" pageId="31" pageNumber="182" reason="1">
<emphasis id="BF84EAE68B2D736DFF7DD5D2FDA68FC0" bold="true" box="[151,534,1217,1244]" pageId="31" pageNumber="182">
<taxonomicName id="4AF04D778B2D736DFF7DD5D2FE1A8FC7" authority="Sokolov, 2023" authorityName="Sokolov" authorityYear="2023" box="[151,426,1217,1244]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="subrufus" status="sp. nov.">
<emphasis id="BF84EAE68B2D736DFF7DD5D2FE1A8FC7" bold="true" box="[151,426,1217,1244]" italics="true" pageId="31" pageNumber="182">Nesamblyops subrufus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A4B7579D8B2D736DFE52D5D2FDA68FC0" box="[440,534,1217,1243]" pageId="31" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2D736DFF7DD5F6FE5F8FE4" box="[151,495,1253,1279]" pageId="31" pageNumber="182" type="description">
<paragraph id="8D4F36F48B2D736DFF7DD5F6FE5F8FE4" blockId="31.[151,534,1217,1279]" box="[151,495,1253,1279]" pageId="31" pageNumber="182">
<figureCitation id="15CB2A718B2D736DFF7DD5F6FE9A8FE4" box="[151,298,1253,1279]" captionStart="FIGURE 11" captionStartId="24.[151,250,886,910]" captionTargetBox="[160,1434,198,862]" captionTargetId="figure-32@24.[151,1437,181,862]" captionTargetPageId="24" captionText="FIGURE 11. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO), B—N. viator (Secretary Island, Grono Bay, Fiordland, SO), C—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184249" httpUri="https://zenodo.org/record/10184249/files/figure.png" pageId="31" pageNumber="182">Figures. 11A</figureCitation>
,
<figureCitation id="15CB2A718B2D736DFEDCD5F6FED98FE4" box="[310,361,1253,1279]" captionStart="FIGURE 13" captionStartId="28.[151,250,934,958]" captionTargetBox="[157,1430,181,910]" captionTargetId="figure-32@28.[157,1430,181,910]" captionTargetPageId="28" captionText="FIGURE 13. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. rotundicollis (Onekaka, Nelson, SO), B—N. solitarius (Italian Creek, Capleston, Buller, SO), C—N. subcaecus (Greymouth, Buller, SO), D—N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO), E—N. viator (Secretary Island, Grono Bay, Fiordland, SO), F—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184253" httpUri="https://zenodo.org/record/10184253/files/figure.png" pageId="31" pageNumber="182">13D</figureCitation>
,
<figureCitation id="15CB2A718B2D736DFE9FD5F6FE788FE4" box="[373,456,1253,1279]" captionStart="FIGURE 16" captionStartId="33.[151,250,788,812]" captionTargetBox="[158,1431,188,760]" captionTargetId="figure-15@33.[152,1436,181,764]" captionTargetPageId="33" captionText="FIGURE 16. Line drawings of male genitalia of New Zealand Nesamblyops species. N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. viator (Secretary Island, Grono Bay, Fjordland, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. victoriae (Capleston, Victoria Range, Buller, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184259" httpUri="https://zenodo.org/record/10184259/files/figure.png" pageId="31" pageNumber="182">16AC</figureCitation>
,
<figureCitation id="15CB2A718B2D736DFE39D5F6FE5F8FE4" box="[467,495,1253,1279]" captionStart="FIGURE 20" captionStartId="37.[151,250,944,968]" captionTargetBox="[151,1436,181,920]" captionTargetId="figure-15@37.[151,1436,181,920]" captionTargetPageId="37" captionText="FIGURE 20. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Green circles—N. solitarius; black circles—N. montanus; pink circles— N. rotundicollis; red circles—N. subrufus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184269" httpUri="https://zenodo.org/record/10184269/files/figure.png" pageId="31" pageNumber="182">20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2D736DFF7DD43EFBC58E70" pageId="31" pageNumber="182" type="materials_examined">
<paragraph id="8D4F36F48B2D736DFF7DD43EFBC58E70" blockId="31.[151,1438,1325,2036]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF7DD43EFEF48E5C" bold="true" box="[151,324,1325,1351]" pageId="31" pageNumber="182">Type material.</emphasis>
<materialsCitation id="3D983CA98B2D736DFEA0D43EFEC88E77" collectionCode="NZAC, DSIR" collectorName="G. Kuschel" country="New Zealand" location="Ns" pageId="31" pageNumber="182" specimenCount="1" typeStatus="holotype">
<emphasis id="BF84EAE68B2D736DFEA0D43EFE6F8E5C" bold="true" box="[330,479,1325,1351]" italics="true" pageId="31" pageNumber="182">
<typeStatus id="524B88568B2D736DFEA0D43EFE6F8E5C" box="[330,479,1325,1351]" pageId="31" pageNumber="182" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
, male, in
<collectionCode id="EBE1AE318B2D736DFDA1D43EFD2A8E5C" box="[587,666,1325,1351]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="31" pageNumber="182" type="Other institutional type">NZAC</collectionCode>
, labeled, dissected: \ Dublin Terrace 25.11.61
<collectorName id="200553228B2D736DFB7DD43EFABE8E5C" box="[1175,1294,1325,1352]" pageId="31" pageNumber="182">G.Kuschel</collectorName>
\ Leafmould \
<collectionCode id="EBE1AE318B2D736DFF4CD442FF568E70" box="[166,230,1361,1387]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="31" pageNumber="182">DSIR</collectionCode>
\
<location id="882F602F8B2D736DFF1FD441FEA48E77" LSID="urn:lsid:plazi:treatment:055987E28B2D7353FF7DD5D2FE2F8E52:882F602F8B2D736DFF1FD441FEA48E77" box="[245,276,1362,1388]" country="New Zealand" name="Ns" pageId="31" pageNumber="182">Ns</location>
\
<collectingCountry id="F5E776648B2D736DFEC3D441FEFE8E77" box="[297,334,1362,1388]" name="New Zealand" pageId="31" pageNumber="182">NZ</collectingCountry>
PB
</materialsCitation>
\.
<emphasis id="BF84EAE68B2D736DFE7FD442FD808E70" bold="true" box="[405,560,1361,1387]" italics="true" pageId="31" pageNumber="182">
<typeStatus id="524B88568B2D736DFE7FD442FD808E70" box="[405,560,1361,1387]" pageId="31" pageNumber="182" type="paratype">PARATYPES</typeStatus>
</emphasis>
(
<specimenCount id="9BF6FD7D8B2D736DFDD5D442FD7E8E70" box="[575,718,1361,1387]" count="2" pageId="31" pageNumber="182" type="generic">2 specimens</specimenCount>
),
<specimenCount id="9BF6FD7D8B2D736DFD0BD442FCFD8E70" box="[737,845,1361,1387]" count="2" pageId="31" pageNumber="182" type="female">2 females</specimenCount>
labeled same as
<typeStatus id="524B88568B2D736DFBE1D442FBC18E70" box="[1035,1137,1361,1387]" pageId="31" pageNumber="182" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2D736DFF2DD466FDDA8EA8" pageId="31" pageNumber="182" type="etymology">
<paragraph id="8D4F36F48B2D736DFF2DD466FDDA8EA8" blockId="31.[151,1438,1325,2036]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD466FEFE8E94" bold="true" box="[199,334,1397,1423]" pageId="31" pageNumber="182">Etymology.</emphasis>
The specific epithet is a Latin adjective
<taxonomicName id="4AF04D778B2D736DFCE4D466FCDE8E94" authorityName="Sokolov" authorityYear="2023" box="[782,878,1397,1423]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="subrufus">
<emphasis id="BF84EAE68B2D736DFCE4D466FCDE8E94" box="[782,878,1397,1423]" italics="true" pageId="31" pageNumber="182">subrufus</emphasis>
</taxonomicName>
(meaning “somewhat red”) in the masculine form and refers to the color of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2D736DFF2DD4AEFE5D8D58" pageId="31" pageNumber="182" type="materials_examined">
<paragraph id="8D4F36F48B2D736DFF2DD4AEFB648EC3" blockId="31.[151,1438,1325,2036]" box="[199,1236,1469,1496]" pageId="31" pageNumber="182">
<materialsCitation id="3D983CA98B2D736DFF2DD4AEFB7F8EC3" box="[199,1231,1469,1496]" country="New Zealand" county="South Island" location="Dublin Terrace" municipality="Upper Buller Gorge" pageId="31" pageNumber="182" specimenCount="1" stateProvince="West Coast" typeStatus="holotype">
<emphasis id="BF84EAE68B2D736DFF2DD4AEFED68ECC" bold="true" box="[199,358,1469,1495]" pageId="31" pageNumber="182">
<typeStatus id="524B88568B2D736DFF2DD4AEFEB28ECC" box="[199,258,1469,1495]" pageId="31" pageNumber="182">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F5E776648B2D736DFE86D4ADFDB58ECC" box="[364,517,1469,1496]" name="New Zealand" pageId="31" pageNumber="182">New Zealand</collectingCountry>
,
<collectingCounty id="642E4E788B2D736DFDFAD4AEFD108ECC" box="[528,672,1469,1495]" pageId="31" pageNumber="182">South Island</collectingCounty>
,
<collectingRegion id="4F34F8168B2D736DFD41D4ADFC9C8ECC" box="[683,812,1469,1496]" country="New Zealand" name="West Coast" pageId="31" pageNumber="182">West Coast</collectingRegion>
,
<collectingMunicipality id="6D2BAC8E8B2D736DFCDDD4ADFBA98ECC" box="[823,1049,1469,1496]" pageId="31" pageNumber="182">Upper Buller Gorge</collectingMunicipality>
,
<location id="882F602F8B2D736DFBCED4AEFB7F8EC3" LSID="urn:lsid:plazi:treatment:055987E28B2D7353FF7DD5D2FE2F8E52:882F602F8B2D736DFBCED4AEFB7F8EC3" box="[1060,1231,1469,1496]" country="New Zealand" county="South Island" municipality="Upper Buller Gorge" name="Dublin Terrace" pageId="31" pageNumber="182" stateProvince="West Coast">Dublin Terrace</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD4F2FE5D8D58" blockId="31.[151,1438,1325,2036]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD4F2FEED8EE0" bold="true" box="[199,349,1505,1531]" pageId="31" pageNumber="182">Recognition.</emphasis>
Adults of this species (
<figureCitation id="15CB2A718B2D736DFD95D4F2FD598EE0" box="[639,745,1505,1531]" captionStart="FIGURE 11" captionStartId="24.[151,250,886,910]" captionTargetBox="[160,1434,198,862]" captionTargetId="figure-32@24.[151,1437,181,862]" captionTargetPageId="24" captionText="FIGURE 11. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO), B—N. viator (Secretary Island, Grono Bay, Fiordland, SO), C—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184249" httpUri="https://zenodo.org/record/10184249/files/figure.png" pageId="31" pageNumber="182">Fig. 11A</figureCitation>
) are practically indistinguishable from the adults of many
<taxonomicName id="4AF04D778B2D736DFF7DD716FE388D04" box="[151,392,1541,1567]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF7DD716FE9C8D04" box="[151,300,1541,1567]" italics="true" pageId="31" pageNumber="182">Nesamblyops</emphasis>
species
</taxonomicName>
based on external characters (e.g.,
<figureCitation id="15CB2A718B2D736DFCC3D716FC0C8D04" box="[809,956,1541,1567]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="31" pageNumber="182">Figs. 10DE</figureCitation>
) and are distinguished from them by the structure of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B2D7353FF2DD75EFE2F8E52" lastPageId="33" lastPageNumber="184" pageId="31" pageNumber="182" type="description">
<paragraph id="8D4F36F48B2D736DFF2DD75EFB208D7C" blockId="31.[151,1438,1325,2036]" box="[199,1168,1613,1640]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD75EFEE98D7C" bold="true" box="[199,345,1613,1639]" pageId="31" pageNumber="182">Description.</emphasis>
Large for genus (SBL range
<quantity id="4A089B118B2D736DFD75D75EFCF48D73" box="[671,836,1613,1640]" metricMagnitude="-3" metricUnit="m" metricValue="1.855" metricValueMax="1.91" metricValueMin="1.8" pageId="31" pageNumber="182" unit="mm" value="1.855" valueMax="1.91" valueMin="1.8">1.801.91 mm</quantity>
, mean 1.87±
<quantity id="4A089B118B2D736DFC39D75EFBF88D73" box="[979,1096,1613,1640]" metricMagnitude="-5" metricUnit="m" metricValue="6.1" pageId="31" pageNumber="182" unit="mm" value="0.061">0.061 mm</quantity>
, n=3).
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD762FCD68DCF" blockId="31.[151,1438,1325,2036]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD762FE978D90" box="[199,295,1649,1675]" italics="true" pageId="31" pageNumber="182">Habitus.</emphasis>
Body form (
<figureCitation id="15CB2A718B2D736DFE52D762FDAE8D90" box="[440,542,1649,1675]" captionStart="FIGURE 11" captionStartId="24.[151,250,886,910]" captionTargetBox="[160,1434,198,862]" captionTargetId="figure-32@24.[151,1437,181,862]" captionTargetPageId="24" captionText="FIGURE 11. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO), B—N. viator (Secretary Island, Grono Bay, Fiordland, SO), C—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184249" httpUri="https://zenodo.org/record/10184249/files/figure.png" pageId="31" pageNumber="182">Fig. 11A</figureCitation>
) markedly convex, elongate ovoid, general proportions slightly wide (WE/SBL 0.37±0.006), head moderately wide relative to pronotum (WH/WPm 0.75±0.025), proportions of pronotum in comparison to elytra average for genus (WPm/WE 0.74±0.005).
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD7CEFCFC8DE3" blockId="31.[151,1438,1325,2036]" box="[199,844,1757,1784]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD7CEFEBD8DEC" box="[199,269,1757,1783]" italics="true" pageId="31" pageNumber="182">Color.</emphasis>
Body color rufotestaceous, appendages testaceous.
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD612FB108C93" blockId="31.[151,1438,1325,2036]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD612FEF18C00" box="[199,321,1793,1819]" italics="true" pageId="31" pageNumber="182">Prothorax.</emphasis>
Pronotum (
<figureCitation id="15CB2A718B2D736DFE2FD612FD9B8C00" box="[453,555,1793,1819]" captionStart="FIGURE 13" captionStartId="28.[151,250,934,958]" captionTargetBox="[157,1430,181,910]" captionTargetId="figure-32@28.[157,1430,181,910]" captionTargetPageId="28" captionText="FIGURE 13. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. rotundicollis (Onekaka, Nelson, SO), B—N. solitarius (Italian Creek, Capleston, Buller, SO), C—N. subcaecus (Greymouth, Buller, SO), D—N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO), E—N. viator (Secretary Island, Grono Bay, Fiordland, SO), F—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184253" httpUri="https://zenodo.org/record/10184253/files/figure.png" pageId="31" pageNumber="182">Fig. 13D</figureCitation>
) moderately long in comparison to elytra (LP/LE 0.39±0.004) and moderately transverse (WPm/LP 1.18±0.018), with lateral margins rectilinear constricted posteriorly (WPm/WPp 1.21±0.029). Anterior angles indistinct, posterior angles obtuse (119124°), widely rounded. Width between posterior angles greater than between anterior angles (WPa/WPp 0.91±0.046). Basal margin slightly convex.
</paragraph>
<paragraph id="8D4F36F48B2D736DFF2DD682FCD98CE8" blockId="31.[151,1438,1325,2036]" pageId="31" pageNumber="182">
<emphasis id="BF84EAE68B2D736DFF2DD682FEA58CB0" box="[199,277,1937,1963]" italics="true" pageId="31" pageNumber="182">Elytra.</emphasis>
Ovoid, moderately depressed along suture, comparatively long (LE/SBL 0.59±0.002) and moderately narrow (WE/LE 0.63±0.008). Humeri completely rounded. Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third.
</paragraph>
<caption id="D98F667C8B127352FF7DD477FB738D87" ID-DOI="http://doi.org/10.5281/zenodo.10184257" ID-Zenodo-Dep="10184257" httpUri="https://zenodo.org/record/10184257/files/figure.png" pageId="32" pageNumber="183" startId="32.[151,250,1380,1404]" targetBox="[157,1431,188,1353]" targetPageId="32" targetType="figure">
<paragraph id="8D4F36F48B127352FF7DD477FB738D87" blockId="32.[151,1437,1380,1693]" pageId="32" pageNumber="183">
<emphasis id="BF84EAE68B127352FF7DD477FEAF8E67" bold="true" box="[151,287,1380,1404]" pageId="32" pageNumber="183">FIGURE 15.</emphasis>
Line drawings of male genitalia of New Zealand
<taxonomicName id="4AF04D778B127352FCE4D477FC5B8E67" box="[782,1003,1380,1404]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BF84EAE68B127352FCE4D477FC258E67" box="[782,917,1380,1404]" italics="true" pageId="32" pageNumber="183">Nesamblyops</emphasis>
species.
</taxonomicName>
<taxonomicName id="4AF04D778B127352FC05D476FBDE8E67" authorityName="Sokolov" authorityYear="2023" box="[1007,1134,1381,1404]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="BF84EAE68B127352FC05D476FBDE8E67" box="[1007,1134,1381,1404]" italics="true" pageId="32" pageNumber="183">N. montanus</emphasis>
</taxonomicName>
(Relax Shelter, Mount Robert Ridge, Nelson, SO):
<emphasis id="BF84EAE68B127352FE80D49BFECC8EBB" bold="true" box="[362,380,1416,1440]" pageId="32" pageNumber="183">A</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B127352FD0BD49AFD428EBA" bold="true" box="[737,754,1417,1441]" pageId="32" pageNumber="183">B</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B127352FB99D49BFB358EBB" bold="true" box="[1139,1157,1416,1440]" pageId="32" pageNumber="183">C</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B127352FF0FD4BEFCF48EDE" authority="(Kangaroo Creek, Mawhera Forest, Buller, SO)" baseAuthorityName="Kangaroo Creek, Mawhera Forest, Buller, SO" box="[229,836,1452,1477]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="BF84EAE68B127352FF0FD4BEFEFC8EDF" box="[229,332,1453,1476]" italics="true" pageId="32" pageNumber="183">N. moorei</emphasis>
(Kangaroo Creek, Mawhera Forest, Buller, SO)
</taxonomicName>
:
<emphasis id="BF84EAE68B127352FCBBD4BEFCD38EDE" bold="true" box="[849,867,1453,1477]" pageId="32" pageNumber="183">D</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B127352FB3FD4BEFB568EDE" bold="true" box="[1237,1254,1453,1477]" pageId="32" pageNumber="183">E</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B127352FEB1D4C2FEDA8EF2" bold="true" box="[347,362,1489,1513]" pageId="32" pageNumber="183">F</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B127352FD25D4C2FCFD8EF3" authorityName="Sokolov" authorityYear="2023" box="[719,845,1489,1512]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="ovipennis">
<emphasis id="BF84EAE68B127352FD25D4C2FCFD8EF3" box="[719,845,1489,1512]" italics="true" pageId="32" pageNumber="183">N. ovipennis</emphasis>
</taxonomicName>
(Mount Arthur, Nelson, SO):
<emphasis id="BF84EAE68B127352FB92D4C3FB3C8EF3" bold="true" box="[1144,1164,1488,1512]" pageId="32" pageNumber="183">G</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B127352FF0FD4E6FF498D16" bold="true" box="[229,249,1525,1549]" pageId="32" pageNumber="183">H</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B127352FD68D4E6FD3C8D16" bold="true" box="[642,652,1525,1549]" pageId="32" pageNumber="183">I</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B127352FC14D4E6FB2E8D17" authorityName="Sokolov" authorityYear="2023" box="[1022,1182,1524,1548]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="BF84EAE68B127352FC14D4E6FB2E8D17" box="[1022,1182,1524,1548]" italics="true" pageId="32" pageNumber="183">N. rotundicollis</emphasis>
</taxonomicName>
(Onekaka, Nelson, SO):
<emphasis id="BF84EAE68B127352FF7DD70AFF148D2A" bold="true" box="[151,164,1561,1585]" pageId="32" pageNumber="183">J</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B127352FDE1D70AFDAF8D2A" bold="true" box="[523,543,1561,1585]" pageId="32" pageNumber="183">K</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B127352FC48D70AFC038D2A" bold="true" box="[930,947,1561,1585]" pageId="32" pageNumber="183">L</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B127352FAF5D70AFDA48D4E" authority="(Italian Creek, Capleston, Buller, SO)" baseAuthorityName="Italian Creek, Capleston, Buller, SO" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="BF84EAE68B127352FAF5D70AFA2C8D2B" box="[1311,1436,1560,1584]" italics="true" pageId="32" pageNumber="183">N. solitarius</emphasis>
(Italian Creek, Capleston, Buller, SO)
</taxonomicName>
:
<emphasis id="BF84EAE68B127352FDF7D72EFD858D4E" bold="true" box="[541,565,1597,1621]" pageId="32" pageNumber="183">M</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B127352FC7CD72EFC188D4E" bold="true" box="[918,936,1597,1621]" pageId="32" pageNumber="183">N</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B127352FACFD72FFA898D4F" bold="true" box="[1317,1337,1596,1620]" pageId="32" pageNumber="183">O</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B127352FE7FD772FDAA8D63" baseAuthorityName="Sharp" baseAuthorityYear="1886" box="[405,538,1632,1656]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="subacecus">
<emphasis id="BF84EAE68B127352FE7FD772FDAA8D63" box="[405,538,1632,1656]" italics="true" pageId="32" pageNumber="183">N. subacecus</emphasis>
</taxonomicName>
(Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO):
<emphasis id="BF84EAE68B127352FB91D772FB3A8D62" bold="true" box="[1147,1162,1633,1657]" pageId="32" pageNumber="183">P</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B127352FF09D797FF478D87" bold="true" box="[227,247,1668,1692]" pageId="32" pageNumber="183">Q</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B127352FD93D796FD3B8D86" bold="true" box="[633,651,1669,1693]" pageId="32" pageNumber="183">R</emphasis>
—median lobe, right lateral aspect. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8D4F36F48B127352FF2DD7D8FA8C8CFA" blockId="32.[151,1437,1739,2017]" pageId="32" pageNumber="183">
<emphasis id="BF84EAE68B127352FF2DD7D8FEC58DFE" box="[199,373,1739,1765]" italics="true" pageId="32" pageNumber="183">Male genitalia.</emphasis>
Median lobe of aedeagus (
<figureCitation id="15CB2A718B127352FD47D7D8FCA38DFE" box="[685,787,1739,1765]" captionStart="FIGURE 16" captionStartId="33.[151,250,788,812]" captionTargetBox="[158,1431,188,760]" captionTargetId="figure-15@33.[152,1436,181,764]" captionTargetPageId="33" captionText="FIGURE 16. Line drawings of male genitalia of New Zealand Nesamblyops species. N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. viator (Secretary Island, Grono Bay, Fjordland, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. victoriae (Capleston, Victoria Range, Buller, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184259" httpUri="https://zenodo.org/record/10184259/files/figure.png" pageId="32" pageNumber="183">Fig. 16C</figureCitation>
) moderately arcuate and barely twisted. Shaft with sides diverging from basal orifice towards middle, then slightly tapering apically. Apex straight, short, enlarged, widely rounded. Apical orifice of moderate length occupies one third of the shaft length. Ventral margin of median lobe almost straight. Walls of shaft lacking poriferous canals. Dorsal copulatory sclerites moderately deviate from typical state. Dorsal field with V-contour small, but distinct, and medial rC-sclerite wide and weakly sclerotized (
<figureCitation id="15CB2A718B127352FA85D648FF7A8C82" captionStart="FIGURE 16" captionStartId="33.[151,250,788,812]" captionTargetBox="[158,1431,188,760]" captionTargetId="figure-15@33.[152,1436,181,764]" captionTargetPageId="33" captionText="FIGURE 16. Line drawings of male genitalia of New Zealand Nesamblyops species. N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. viator (Secretary Island, Grono Bay, Fjordland, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. victoriae (Capleston, Victoria Range, Buller, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184259" httpUri="https://zenodo.org/record/10184259/files/figure.png" pageId="32" pageNumber="183">Fig. 16C</figureCitation>
). Additionally, internal sac has a large scaly membraneous field near apical orifice. Left paramere (
<figureCitation id="15CB2A718B127352FAC7D66CFA248C82" box="[1325,1428,1919,1945]" captionStart="FIGURE 16" captionStartId="33.[151,250,788,812]" captionTargetBox="[158,1431,188,760]" captionTargetId="figure-15@33.[152,1436,181,764]" captionTargetPageId="33" captionText="FIGURE 16. Line drawings of male genitalia of New Zealand Nesamblyops species. N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. viator (Secretary Island, Grono Bay, Fjordland, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. victoriae (Capleston, Victoria Range, Buller, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184259" httpUri="https://zenodo.org/record/10184259/files/figure.png" pageId="32" pageNumber="183">Fig. 16A</figureCitation>
) comparatively narrow, with strongly attenuate apex, bearing two long setae. Right paramere (
<figureCitation id="15CB2A718B127352FB25D6B0FA858CA6" box="[1231,1333,1955,1981]" captionStart="FIGURE 16" captionStartId="33.[151,250,788,812]" captionTargetBox="[158,1431,188,760]" captionTargetId="figure-15@33.[152,1436,181,764]" captionTargetPageId="33" captionText="FIGURE 16. Line drawings of male genitalia of New Zealand Nesamblyops species. N. subrufus (Dublin Terrace, Upper Buller Gorge, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. viator (Secretary Island, Grono Bay, Fjordland, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. victoriae (Capleston, Victoria Range, Buller, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184259" httpUri="https://zenodo.org/record/10184259/files/figure.png" pageId="32" pageNumber="183">Fig. 16B</figureCitation>
) narrow, long, bearing two long setae, which are shorter than the length of paramere. Ring sclerite not investigated.
</paragraph>
<caption id="D98F667C8B137353FF7DD207FAB088A7" ID-DOI="http://doi.org/10.5281/zenodo.10184259" ID-Zenodo-Dep="10184259" httpUri="https://zenodo.org/record/10184259/files/figure.png" pageId="33" pageNumber="184" startId="33.[151,250,788,812]" targetBox="[158,1431,188,760]" targetPageId="33" targetType="figure">
<paragraph id="8D4F36F48B137353FF7DD207FAB088A7" blockId="33.[151,1436,788,957]" pageId="33" pageNumber="184">
<emphasis id="BF84EAE68B137353FF7DD207FEAF8837" bold="true" box="[151,287,788,812]" pageId="33" pageNumber="184">FIGURE 16.</emphasis>
Line drawings of male genitalia of New Zealand
<taxonomicName id="4AF04D778B137353FCFBD207FC5E8837" box="[785,1006,788,812]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BF84EAE68B137353FCFBD207FC288837" box="[785,920,788,812]" italics="true" pageId="33" pageNumber="184">Nesamblyops</emphasis>
species.
</taxonomicName>
<taxonomicName id="4AF04D778B137353FC19D206FBD78837" authorityName="Sokolov" authorityYear="2023" box="[1011,1127,788,812]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="subrufus">
<emphasis id="BF84EAE68B137353FC19D206FBD78837" box="[1011,1127,788,812]" italics="true" pageId="33" pageNumber="184">N. subrufus</emphasis>
</taxonomicName>
(Dublin Terrace, Upper Buller Gorge, Nelson, SO):
<emphasis id="BF84EAE68B137353FE86D22BFECE884B" bold="true" box="[364,382,824,848]" pageId="33" pageNumber="184">A</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B137353FD08D22AFD43884A" bold="true" box="[738,755,825,849]" pageId="33" pageNumber="184">B</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B137353FB99D22BFB35884B" bold="true" box="[1139,1157,824,848]" pageId="33" pageNumber="184">C</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B137353FF0ED24EFE8F886F" authorityName="Sokolov" authorityYear="2023" box="[228,319,861,884]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="viator">
<emphasis id="BF84EAE68B137353FF0ED24EFE8F886F" box="[228,319,861,884]" italics="true" pageId="33" pageNumber="184">N. viator</emphasis>
</taxonomicName>
(Secretary Island, Grono Bay, Fjordland, SO):
<emphasis id="BF84EAE68B137353FCCED24EFC86886E" bold="true" box="[804,822,861,885]" pageId="33" pageNumber="184">D</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B137353FB4AD24EFB01886E" bold="true" box="[1184,1201,861,885]" pageId="33" pageNumber="184">E</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B137353FEC2D292FE878882" bold="true" box="[296,311,897,921]" pageId="33" pageNumber="184">F</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="4AF04D778B137353FD4AD292FB018882" authority="(Capleston, Victoria Range, Buller, SO)" baseAuthorityName="Capleston, Victoria Range, Buller, SO" box="[672,1201,896,921]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="victoriae">
<emphasis id="BF84EAE68B137353FD4AD292FCA78883" box="[672,791,897,920]" italics="true" pageId="33" pageNumber="184">N. victoriae</emphasis>
(Capleston, Victoria Range, Buller, SO)
</taxonomicName>
:
<emphasis id="BF84EAE68B137353FB51D293FB7F8883" bold="true" box="[1211,1231,896,920]" pageId="33" pageNumber="184">G</emphasis>
—left paramere, left lateral aspect,
<emphasis id="BF84EAE68B137353FEC3D2B6FE8D88A6" bold="true" box="[297,317,933,957]" pageId="33" pageNumber="184">H</emphasis>
—right paramere, right lateral aspect,
<emphasis id="BF84EAE68B137353FD55D2B6FD7988A6" bold="true" box="[703,713,933,957]" pageId="33" pageNumber="184">I</emphasis>
—median lobe, right lateral aspect. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8D4F36F48B137353FF2DD2F8FD298F1E" blockId="33.[151,1436,1003,1353]" box="[199,665,1003,1029]" pageId="33" pageNumber="184">
<emphasis id="BF84EAE68B137353FF2DD2F8FE5D8F1E" box="[199,493,1003,1029]" italics="true" pageId="33" pageNumber="184">Female internal genitalia.</emphasis>
Not examined.
</paragraph>
<paragraph id="8D4F36F48B137353FF2DD51DFAB98F56" blockId="33.[151,1436,1003,1353]" pageId="33" pageNumber="184">
<emphasis id="BF84EAE68B137353FF2DD51DFDB78F33" bold="true" box="[199,519,1038,1064]" pageId="33" pageNumber="184">Geographical distribution.</emphasis>
This species is known from only one locality situated near Manuka Flat in the Upper Buller Gorge, upstream from Inangahua at the northern
<collectingRegion id="4F34F8168B137353FCBFD520FB928F56" box="[853,1058,1075,1101]" country="New Zealand" name="West Coast" pageId="33" pageNumber="184">West Coast region</collectingRegion>
(
<figureCitation id="15CB2A718B137353FBDBD520FB348F56" box="[1073,1156,1075,1101]" captionStart="FIGURE 20" captionStartId="37.[151,250,944,968]" captionTargetBox="[151,1436,181,920]" captionTargetId="figure-15@37.[151,1436,181,920]" captionTargetPageId="37" captionText="FIGURE 20. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Green circles—N. solitarius; black circles—N. montanus; pink circles— N. rotundicollis; red circles—N. subrufus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184269" httpUri="https://zenodo.org/record/10184269/files/figure.png" pageId="33" pageNumber="184">Fig. 20</figureCitation>
, red circle).
</paragraph>
<paragraph id="8D4F36F48B137353FF2DD545FBAC8F6A" blockId="33.[151,1436,1003,1353]" box="[199,1052,1110,1137]" pageId="33" pageNumber="184">
<emphasis id="BF84EAE68B137353FF2DD545FE9A8F6B" bold="true" box="[199,298,1110,1136]" pageId="33" pageNumber="184">Habitat.</emphasis>
All three known to date specimens were collected from leafmould.
</paragraph>
<paragraph id="8D4F36F48B137353FF2DD569FE2F8E52" blockId="33.[151,1436,1003,1353]" pageId="33" pageNumber="184">
<emphasis id="BF84EAE68B137353FF2DD569FEC08F8F" bold="true" box="[199,368,1146,1172]" pageId="33" pageNumber="184">Relationships.</emphasis>
The external morphology (body proportions, size) suggests relatedness of
<taxonomicName id="4AF04D778B137353FB31D568FAEF8F8E" authorityName="Sokolov" authorityYear="2023" box="[1243,1375,1147,1173]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="subrufus">
<emphasis id="BF84EAE68B137353FB31D568FAEF8F8E" box="[1243,1375,1147,1173]" italics="true" pageId="33" pageNumber="184">N. subrufus</emphasis>
</taxonomicName>
with
<taxonomicName id="4AF04D778B137353FF7DD58CFEF78FA2" authorityName="Sokolov" authorityYear="2023" box="[151,327,1183,1209]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="BF84EAE68B137353FF7DD58CFEF78FA2" box="[151,327,1183,1209]" italics="true" pageId="33" pageNumber="184">N. rotundicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="4AF04D778B137353FE6AD58CFDBC8FA2" box="[384,524,1183,1209]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="BF84EAE68B137353FE6AD58CFDBC8FA2" box="[384,524,1183,1209]" italics="true" pageId="33" pageNumber="184">N. solitarius</emphasis>
</taxonomicName>
. The structure of the parameres (two setae, attenuate apex of left paramere) and the male genitalia (presence of scaly field in apical part of median lobe) support this statement. However, the state of copulatory sclerites of
<taxonomicName id="4AF04D778B137353FE1CD5F4FDC88E1A" authorityName="Sokolov" authorityYear="2023" box="[502,632,1255,1281]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="subrufus">
<emphasis id="BF84EAE68B137353FE1CD5F4FDC88E1A" box="[502,632,1255,1281]" italics="true" pageId="33" pageNumber="184">N. subrufus</emphasis>
</taxonomicName>
are quite different from those of
<taxonomicName id="4AF04D778B137353FC1FD5F4FB158E1A" authorityName="Sokolov" authorityYear="2023" box="[1013,1189,1255,1281]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="BF84EAE68B137353FC1FD5F4FB158E1A" box="[1013,1189,1255,1281]" italics="true" pageId="33" pageNumber="184">N. rotundicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="4AF04D778B137353FB34D5F4FAD98E1A" box="[1246,1385,1255,1281]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="BF84EAE68B137353FB34D5F4FAD98E1A" box="[1246,1385,1255,1281]" italics="true" pageId="33" pageNumber="184">N. solitarius</emphasis>
</taxonomicName>
, but much closer to the copulatory sclerites of the other members of
<taxonomicName id="4AF04D778B137353FC68D418FBA78E3E" authorityName="Jeannel" authorityYear="1937" box="[898,1047,1291,1317]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="BF84EAE68B137353FC68D418FBA78E3E" box="[898,1047,1291,1317]" italics="true" pageId="33" pageNumber="184">Nesamblyops</emphasis>
</taxonomicName>
, thus making a link between two agglomerations of taxa.
</paragraph>
</subSubSection>
</treatment>
</document>