treatments-xml/data/04/39/71/043971C107AC5C2B9BAEA85415594868.xml

<document ID-CLB-Dataset="21607" ID-DOI="http://dx.doi.org/10.3897/phytokeys.151.50538" ID-GBIF-Dataset="a62873a2-c4e1-4c6d-9baa-5c5fb94e7348" ID-PMC="PMC7329918" ID-Pensoft-Pub="1314-2003-151-107" ID-Pensoft-UUID="42D1EB2DF0545744B92BFCCCB395C45B" ID-PubMed="32655282" ModsDocID="1314-2003-151-107" checkinTime="1593036331120" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Cuta-Alarcon, Lia E., Bravo-Pedraza, William J. &amp; Soreng, Robert J." docDate="2020" docId="043971C107AC5C2B9BAEA85415594868" docLanguage="en" docName="PhytoKeys 151: 107-160" docOrigin="PhytoKeys 151" docSource="http://dx.doi.org/10.3897/phytokeys.151.50538" docTitle="Agrostis boyacensis Swallen &amp; García-Barr., Caldasia 2 (8): 303, fig. A. 1943" docType="treatment" docVersion="6" id="42D1EB2DF0545744B92BFCCCB395C45B" lastPageNumber="107" masterDocId="42D1EB2DF0545744B92BFCCCB395C45B" masterDocTitle="Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species" masterLastPageNumber="160" masterPageNumber="107" pageNumber="107" updateTime="1732688126400" updateUser="ExternalLinkService">
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<mods:title id="1B48DB3D0C53A9EDECEB890CD0AD08E4">Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species</mods:title>
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<mods:namePart id="B0D9A8F26A3E22CEC1778AD9CA1BF083">Sylvester, Steven P.</mods:namePart>
<mods:affiliation id="2EB33143020B1518909710CCE4E4E0DE">College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China &amp; Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
<mods:nameIdentifier id="87F268B709FF6049F7020D34DFF175FA" type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
<mods:nameIdentifier id="52626450D198366A742B8519B8A54448" type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="ABDE1ECB6A659218069B994BC76DB2FE">Cuta-Alarcon, Lia E.</mods:namePart>
<mods:affiliation id="1C0BE34FD0369687F3E64CD662EE52B8">Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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<mods:namePart id="04B86F5C2EB452D4F216D6D1BCB7D61E">Bravo-Pedraza, William J.</mods:namePart>
<mods:affiliation id="1DA156B7977550CF2075C18B916F6296">Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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<mods:namePart id="423327B83DBE4EECB869DABD65D66EDC">Soreng, Robert J.</mods:namePart>
<mods:affiliation id="7D7D7073D03FDB7708556562B4167165">Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
<mods:nameIdentifier id="4B698C6433823F4832D00704A23A0361" type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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<mods:title id="FE7E8E84B302084A1B11F36BA6875164">PhytoKeys</mods:title>
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<mods:date id="72A2514DC39FA98F8219314636B2CB75">2020</mods:date>
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<mods:number id="0D232D68F684C55B8D994D50B2558CCC">151</mods:number>
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<mods:start id="D5D82376C38E7B6ECF9EA85C16FE458B">107</mods:start>
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<mods:classification id="68EE1E2EDA109B36A329F6650976AE4D">journal article</mods:classification>
<mods:identifier id="59042F1D4BFDF07F14F88790B7442B48" type="DOI">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:identifier>
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<treatment id="043971C107AC5C2B9BAEA85415594868" ID-GBIF-Taxon="164730858" LSID="urn:lsid:plazi:treatment:043971C107AC5C2B9BAEA85415594868" httpUri="http://treatment.plazi.org/id/043971C107AC5C2B9BAEA85415594868" lastPageNumber="107" pageId="0" pageNumber="107" scope_family="Poaceae">
<subSubSection id="FE495A44937D37742F2AC6053EF3E2D5" pageId="0" pageNumber="107" type="nomenclature">
<paragraph id="A4EBBFF6DE772404A840B91B5617BF85" pageId="0" pageNumber="107">
<taxonomicName id="F622283690797963A1533258DD4936BD" ID-CoL="65TV3" LSID="043971C1-07AC-5C2B-9BAE-A85415594868" authority="Swallen &amp; Garcia-Barr., Caldasia 2 (8): 303, fig. A. 1943" authorityName="Swallen &amp; García-Barr., Caldasia 2 (8): 303, fig. A." authorityYear="1943" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis boyacensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
Agrostis boyacensis Swallen &amp; 
<normalizedToken id="43F67E1A51D1D0FE5BDFBB7ACDFE577D" originalValue="García-Barr">Garcia-Barr</normalizedToken>
., Caldasia 2 (8): 303, fig. A. 1943
</taxonomicName>
<figureCitation id="CD8DC10B04C584D7B1BD8DA338F22233" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Agrostis boyacensis, examples of specimens from paramo or superparamo of the Sierra Nevada del Cocuy A whole plant B close-up of inflorescence C spikelet at anthesis, lateral view D upper culm blades and ligular area E floret, ventral view F floret, dorsal view. Images A, C, D, F: Cleef 8504 (US 01247250) courtesy of United States National Herbarium (US); B, E: Cleef 5665 (US 2785695). Scale bar of E also for F." figureDoi="10.3897/phytokeys.151.50538.figure1" httpUri="https://binary.pensoft.net/fig/424839" pageId="0" pageNumber="107">Figs 1</figureCitation>
<figureCitation id="0DF4CAB1387859D8EAD6DFC1066EB377" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Agrostis boyacensis, example of a specimen encountered in paramos of Boyaca outside of the Sierra Nevada del Cocuy A whole plant B spikelet, glumes lateral view and floret ventral view showing anthers C spikelet, glumes lateral view with floret detached and raised above the glumes showing ventral surface. Images of Sylvester 3017 a (FMB)." figureDoi="10.3897/phytokeys.151.50538.figure2" httpUri="https://binary.pensoft.net/fig/424840" pageId="0" pageNumber="107">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="03131438F9BD1962316290C22E7DA5D9" pageId="0" pageNumber="107" type="materials_examined">
<paragraph id="6B409C6FCE6C2CA078AC2710230A5792" pageId="0" pageNumber="107">Type.</paragraph>
<paragraph id="59F6B213E8A807A744721279FD5F4F74" pageId="0" pageNumber="107">
Colombia. 
<normalizedToken id="7D0523B6C416ADDA58EEDC9039EDC07B" originalValue="Boyacá">Boyaca</normalizedToken>
: Nevado del Cocuy, alto valle de Las Lagunillas, [
<geoCoordinate id="509A8C479CD2880F364531363F268818" degrees="6.3906" direction="north" orientation="latitude" precision="5" value="6.3906">6.3906N</geoCoordinate>
, 
<geoCoordinate id="20B3A1E92082A153A0422C4DA14BFCD7" degrees="72.3542" direction="west" orientation="longitude" precision="5" value="-72.3542">72.3542W</geoCoordinate>
], 4000-4300 m alt., 12 Sep. 1938, J. Cuatrecasas &amp; H. 
<normalizedToken id="E485AA44B6CE44F9C484E01FDCF1DB1B" originalValue="García-Barriga">Garcia-Barriga</normalizedToken>
1459 (holotype: US (US00131729 [image!]); isotype: COL (COL000006092!), SI (SI000494 [image!] fragm. ex US)).
</paragraph>
</subSubSection>
<subSubSection id="83563DA9B275DAEB45FB857D13969AA6" pageId="0" pageNumber="107" type="description">
<paragraph id="2A870200615D43AFA40CF92B7D07483E" pageId="0" pageNumber="107">Description.</paragraph>
<paragraph id="9718C7933830AF5675DDAA3D58D8E727" pageId="0" pageNumber="107">
<emphasis id="2D4EB133F5B3C698502730BFFA7AFC0C" bold="true" pageId="0" pageNumber="107">Perennial herbs</emphasis>
, densely tufted, often stooling with perenniating many-branched culms, often with short lateral tending to ascending rhizomes. 
<emphasis id="CF797F06DA17940014AB3BDCAC351DC1" bold="true" pageId="0" pageNumber="107">Tillers</emphasis>
extravaginal, with cataphyllous shoots present. 
<emphasis id="4F655C8C5EF425645E5C44365AB238C4" bold="true" pageId="0" pageNumber="107">Culms</emphasis>
3-24 cm tall (to 37 cm tall in specimens from 
<normalizedToken id="F8F2BBD6B37C34F88A59D0B6952C2815" originalValue="Boyacán">Boyacan</normalizedToken>
<normalizedToken id="A637FC0BE9AD3F7037AE7FAA0DDD3B42" originalValue="páramos">paramos</normalizedToken>
outside the Sierra Nevada del Cocuy), erect to arched, firm, with 0(-2) nodes exerted at flowering, densely scabrous throughout or scabrous just below the nodes, or rarely smooth (specimens from 
<normalizedToken id="A61F55F0FC46B3F0493192BA28E7BB4A" originalValue="Boyacán">Boyacan</normalizedToken>
<normalizedToken id="2122DA6C844E175C0F7341F808825DA0" originalValue="páramos">paramos</normalizedToken>
outside the Sierra Nevada del Cocuy). 
<emphasis id="855A0E5F2F18C79288980BDB3C4FC519" bold="true" pageId="0" pageNumber="107">Leaves</emphasis>
basal and cauline, often mainly basal, glabrous, usually densely scabrous throughout, less often smooth or scaberulous (specimens from 
<normalizedToken id="BF139C7D802271E680A99BC63C1772BC" originalValue="Boyacán">Boyacan</normalizedToken>
<normalizedToken id="F78BB3902033FB23B29F53E631957799" originalValue="páramos">paramos</normalizedToken>
outside the Sierra Nevada del Cocuy); 
<emphasis id="2EB22D5D0B98BADB428F8A3755ACBEE5" bold="true" pageId="0" pageNumber="107">ligules</emphasis>
0.9-2 mm long, membranous or slightly scarious, truncate to triangular and obtuse, moderately to strongly decurrent with the sheath, abaxial surface smooth or scaberulous; 
<emphasis id="265D681AD5B19C78D468A17BAB412350" bold="true" pageId="0" pageNumber="107">blades</emphasis>
(2-)3-6(-12) cm long, 0.5-1.2(-1.5) mm in diameter as folded or rolled, convolute, involute, or strongly conduplicate, usually recurved or sometimes straight, firm to rigid, those of upper culm sometimes flat, firm, 2-3 mm wide, abaxial surface usually densely scabrous throughout, less often smooth to scaberulous (specimens from 
<normalizedToken id="DA1C6FBCA4A990F66425F9EC32ADEAE0" originalValue="Boyacán">Boyacan</normalizedToken>
<normalizedToken id="B6A7D6B64994833BF14BF2A3E3D05554" originalValue="páramos">paramos</normalizedToken>
outside the Sierra Nevada del Cocuy), adaxial surface and margins moderately to densely scabrous, apices blunt or slightly naviculate-acute. 
<emphasis id="C1E336D38770860AB7BC624E65FC8C9D" bold="true" pageId="0" pageNumber="107">Panicles</emphasis>
(2-)3-6(-8) cm long, c. 0.3-0.7(-0.9) cm wide, moderately to densely congested, sub-spikelike to spikelike, generally not interrupted, subincluded in the basal foliage to greatly exerted, lateral branches with spikelets usually present almost to the base, upper lateral branches short and held close to the central inflorescence axis, central axis and panicle branches moderately to densely scabrous, rarely smooth or scaberulous (specimens from 
<normalizedToken id="0659B3B29188C3874A3BB72570D60E57" originalValue="Boyacán">Boyacan</normalizedToken>
<normalizedToken id="B62DDF9398CF755F0888ABB451A9C6A4" originalValue="páramos">paramos</normalizedToken>
outside the Sierra Nevada del Cocuy); 
<emphasis id="1DBC09427F3711A8E965B1D22F22497D" bold="true" pageId="0" pageNumber="107">pedicels</emphasis>
0.8-2.5 mm long, usually shorter than their spikelets, not obviously dilated at their apex, moderately to densely scabrous, sometimes smooth or scaberulous (specimens from 
<normalizedToken id="7C03EA227D8330AA8B3B579AD7634C9E" originalValue="Boyacán">Boyacan</normalizedToken>
<normalizedToken id="2F9AF8245345055D1601E6AA80AC4EBF" originalValue="páramos">paramos</normalizedToken>
outside the Sierra Nevada del Cocuy). 
<emphasis id="EC03B8FF247D063566D2DD398E03F509" bold="true" pageId="0" pageNumber="107">Spikelets</emphasis>
1.8-2.4(-2.5) mm long; 
<emphasis id="E8E8F659967E6D009D520590203125F0" bold="true" pageId="0" pageNumber="107">glumes</emphasis>
usually unequal, sometimes subequal, the lower longer than the upper by up to 0.3 mm and usually wider than the upper, 1-veined, lower glume keel scabrous throughout to only in distal 1/3, upper glume scabrous only in the distal 1/3 or sometimes smooth throughout, apices acute; 
<emphasis id="705D78D2C7A636DC2E0555681B4D44C0" bold="true" pageId="0" pageNumber="107">floret</emphasis>
3/5-2/3(-3/4) the length of the glumes; 
<emphasis id="D0EBF7A722E864354D6AA181A5A2FA51" bold="true" pageId="0" pageNumber="107">calluses</emphasis>
glabrous or with 2 sparse tufts of very short hairs on the lateral sides; 
<emphasis id="DFA1520E9A5CD8D631B36CB6390EF0C2" bold="true" pageId="0" pageNumber="107">lemmas</emphasis>
1-1.5 mm long, glabrous, smooth, 5-veined, apex obtuse to slightly truncate, erose, muticous or exceptionally with a short straight awn inserted in the upper 
<normalizedToken id="39017D828B63748F4C34A4106A561A13" originalValue="½">1/2</normalizedToken>
of the lemma, to 0.5 mm long, not surpassing the glume apex, weak and falling easily (i.e. Sylvester 3071); 
<emphasis id="0190DA0EC8D0A157775F46B6720DD290" bold="true" pageId="0" pageNumber="107">paleas</emphasis>
absent or to 0.3 mm long, &lt;
<normalizedToken id="065075F63498207704AA8D940F7873CA" originalValue="¼">1/4</normalizedToken>
the length of the lemma; 
<emphasis id="574C682CA21A757D1FBF245262C98CAF" bold="true" pageId="0" pageNumber="107">rachilla</emphasis>
absent; 
<emphasis id="F74FB9410867AFE8D15D9F6FC08DD9AE" bold="true" pageId="0" pageNumber="107">anthers</emphasis>
0.5-0.8(-1) mm long.
</paragraph>
<caption id="7CF798801BE89F88AA11ADB2036EA94E" doi="10.3897/phytokeys.151.50538.figure1" httpUri="https://binary.pensoft.net/fig/424839" pageId="0" pageNumber="107" start="Figure 1" startId="F1">
<paragraph id="2C559B367ECFC8F14E7A03DF08140557" pageId="0" pageNumber="107">
<emphasis id="DA1E2594D1268070A00F84C5F28ADC9B" bold="true" pageId="0" pageNumber="107">Figure 1.</emphasis>
<taxonomicName id="CEA687CC09B779BA536936B53ED663A7" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis boyacensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
<emphasis id="B223CEB61637E93799B85096F597F12A" italics="true" pageId="0" pageNumber="107">Agrostis boyacensis</emphasis>
</taxonomicName>
, examples of specimens from 
<normalizedToken id="22BDE1D4F7C4EA105F85358EF291B647" originalValue="páramo">paramo</normalizedToken>
or 
<normalizedToken id="F96097E5414690553B6EE917B7644B3B" originalValue="superpáramo">superparamo</normalizedToken>
of the Sierra Nevada del Cocuy 
<emphasis id="BD81F74B0881571873EAC8AD20E9922C" bold="true" pageId="0" pageNumber="107">A</emphasis>
whole plant 
<emphasis id="DF2744B300DD475D972AAF8FB3554BC4" bold="true" pageId="0" pageNumber="107">B</emphasis>
close-up of inflorescence 
<emphasis id="63A5BBAC4E8CF3761A50FEEDD00D6D8F" bold="true" pageId="0" pageNumber="107">C</emphasis>
spikelet at anthesis, lateral view 
<emphasis id="8FCB302E895F57A5BED552E895C1D41D" bold="true" pageId="0" pageNumber="107">D</emphasis>
upper culm blades and ligular area 
<emphasis id="B376EB89A779C6C2F88C056F7FA4CB4B" bold="true" pageId="0" pageNumber="107">E</emphasis>
floret, ventral view 
<emphasis id="65CCDE49989277F8A74509A4D692138A" bold="true" pageId="0" pageNumber="107">F</emphasis>
floret, dorsal view. Images 
<emphasis id="9E0E9D678055EFCA7FD7B4B7BC6C88D5" bold="true" pageId="0" pageNumber="107">A, C, D, F</emphasis>
: Cleef 8504 (US01247250) courtesy of United States National Herbarium (US); 
<emphasis id="19A344D66F9AA3A2CC37C6EB8DAA4454" bold="true" pageId="0" pageNumber="107">B, E</emphasis>
: Cleef 5665 (US2785695). Scale bar of 
<emphasis id="4017B4EDD6BA245EAE8FD6E9136192A4" bold="true" pageId="0" pageNumber="107">E</emphasis>
also for 
<emphasis id="3E3A234F4D557196DF90B897464F823C" bold="true" pageId="0" pageNumber="107">F</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="619AA334B624D49AF460980331961E70" pageId="0" pageNumber="107" type="distribution">
<paragraph id="D5FA1C4A37C9C75D94EA482EDA3EA4FD" pageId="0" pageNumber="107">Distribution and ecology.</paragraph>
<paragraph id="C68F228FFDB1F8C7AF70B4C4EEB52536" pageId="0" pageNumber="107">
Colombia, Ecuador?. 
<normalizedToken id="8637F08198C9AA52A3EF773CABAA75AE" originalValue="Páramos">Paramos</normalizedToken>
and 
<normalizedToken id="38306944C3B6028D96E4054ABABF200D" originalValue="superpáramos">superparamos</normalizedToken>
, found in both zonal grass 
<normalizedToken id="0F69241EF0AFEF708B28136E4EE40B98" originalValue="páramo">paramo</normalizedToken>
and azonal high elevation moraine, 2850-4500 m alt. Within Colombia, the species seems to be found only in the Cordillera Oriental, with specimens only seen from Departamento 
<normalizedToken id="92D10F87D49B41BF7703343BB0C34FC8" originalValue="Boyacá">Boyaca</normalizedToken>
. 
<bibRefCitation id="8D1D44D336707529A2A07D0067D3011F" author="Luteyn, JL" journalOrPublisher="Memoirs of the New York Botanical Garden" pageId="0" pageNumber="107" pagination="1 - 278" refId="B20" refString="Luteyn, JL, 1999. Paramos, a checklist of plant diversity, geographical distribution, and botanical literature. Memoirs of the New York Botanical Garden 84: 1 - 278" title="Paramos, a checklist of plant diversity, geographical distribution, and botanical literature." volume="84" year="1999">Luteyn (1999)</bibRefCitation>
mentions the species to be found in Ecuador and the species is also mentioned in the key to some Ecuadorian taxa (
<bibRefCitation id="0937B2C74F3F56CF8B8F7D51331FBA22" author="Palacio, PC" journalOrPublisher="Nature" pageId="0" pageNumber="107" refId="B24" refString="Palacio, PC, Molina, AM, Rugolo de Agrasar, ZE, Vega, AS, in press. A new species of Agrostis (Poaceae, Pooideae, Poeae, Agrostidinae) from the Andean paramos of Ecuador. Novon." title="A new species of Agrostis (Poaceae, Pooideae, Poeae, Agrostidinae) from the Andean paramos of Ecuador. Novon." year="in press">Palacio et al. in press</bibRefCitation>
), but no specimens have been verified at US.
</paragraph>
</subSubSection>
<subSubSection id="724A8EC8FE6D27EA615F748774614C27" pageId="0" pageNumber="107" type="materials_examined">
<paragraph id="F732DACB48F64C444C6A008AEAB31B3B" pageId="0" pageNumber="107">Other specimens examined.</paragraph>
<paragraph id="4672B31C519F5E85B33DE5F48BA92110" pageId="0" pageNumber="107">
<emphasis id="AA6474D06A2C69FBCBC1F50D753898F5" bold="true" pageId="0" pageNumber="107">Colombia</emphasis>
. 
<emphasis id="BCEE811991A44F1A1B51E5F1ED78E225" bold="true" pageId="0" pageNumber="107">
<normalizedToken id="90E2992C544DC746B0F500E50C4B9189" originalValue="Boyacá">Boyaca</normalizedToken>
</emphasis>
: Munic. Chiscas, 
<normalizedToken id="BF2607550FC32D45E9E3832A601ABB61" originalValue="Páramo">Paramo</normalizedToken>
de Chacaritas, found on a rock escarpment 4 m tall, 
<geoCoordinate id="434C7174675D6E3E96898BDFA3ACBA2E" degrees="6.62227" direction="north" orientation="latitude" precision="1" value="6.62227">6.62227N</geoCoordinate>
, 
<geoCoordinate id="BC3436B954C2C879552A9C83E9C81D9F" degrees="72.39040" direction="west" orientation="longitude" precision="1" value="-72.3904">72.39040W</geoCoordinate>
, 4192 m alt., 4 Mar. 2018, S.P. Sylvester 3117 (K, UPTC, US); Munic. Chiscas, 
<normalizedToken id="34AFF8CBE6744349ABCBA54942F348DF" originalValue="Páramo">Paramo</normalizedToken>
de Chacaritas, close to the foot of the talus scree slope, 
<geoCoordinate id="B5117A45C0E9C9E681A67DFF33737160" degrees="6.61779" direction="north" orientation="latitude" precision="1" value="6.61779">6.61779N</geoCoordinate>
, 
<geoCoordinate id="34F877E35C4E3D070F94542B88679332" degrees="72.38899" direction="west" orientation="longitude" precision="1" value="-72.38899">72.38899W</geoCoordinate>
, 4354 m alt., 4 Mar. 2018, S.P. Sylvester 3129 (COL, FMB, K, US); Sierra Nevada del Cocuy, Laguna Grande, in open gravelly slopes, morranes and similar places at high altitude, [
<geoCoordinate id="EA2B4B5876FDE1C8DAB726A0C753A09D" degrees="6.5556" direction="north" orientation="latitude" precision="5" value="6.5556">6.5556N</geoCoordinate>
, 
<geoCoordinate id="9F86725F5C85E3055725870D356C4E75" degrees="72.3253" direction="west" orientation="longitude" precision="5" value="-72.3253">72.3253W</geoCoordinate>
], 4300-4500 m alt., 28 Dec. 1985, J.R.I. Wood 5247 (US3481052); Munic. Duitama, 
<normalizedToken id="6BC9161956493C773C79377D991568E9" originalValue="Páramo">Paramo</normalizedToken>
de la Rusia, Sector del 
<normalizedToken id="EF7EB25A692A92767FEBC1D3095D53CA" originalValue="Páramo">Paramo</normalizedToken>
de Agueros, en la 
<normalizedToken id="19B7F4FBBF2A9A827A59A8604084330F" originalValue="vía">via</normalizedToken>
que conduce a la vereda 
<normalizedToken id="4F11291625DAB2515E0D1F25D6FDE29C" originalValue="Avendaños">Avendanos</normalizedToken>
, 
<normalizedToken id="F5BCFA59173C27D4A5830A21FC29A37F" originalValue="páramo">paramo</normalizedToken>
dominado por 
<taxonomicName id="B58AADD8500E21FC9C74823CA974C7D0" class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="34F0CD00A25162F5C8BBF79250B696AD" italics="true" pageId="0" pageNumber="107">Chusquea</emphasis>
</taxonomicName>
, evidencia de pastoreo intenso, en alguna 
<normalizedToken id="FCE8BAA16D5AA3FF96842D66DFEB926F" originalValue="perturbación">perturbacion</normalizedToken>
humano / animal, 
<geoCoordinate id="D7842E716FCF66335C7A852C89B7B2B4" degrees="5.94611" direction="north" orientation="latitude" precision="1" value="5.94611">5.94611N</geoCoordinate>
, 
<geoCoordinate id="AECF73E8B42713E5FD488EB7559D5112" degrees="73.08481" direction="west" orientation="longitude" precision="1" value="-73.08481">73.08481W</geoCoordinate>
, 3768 m alt., 3 Oct. 2017, S.P. Sylvester 3017a (COL, K, SI, US); Munic. Arcabuco, Vereda el Carmen, 
<normalizedToken id="2AA746AA918DEE1EAF29003931DF5507" originalValue="Páramo">Paramo</normalizedToken>
del Valle, 
<normalizedToken id="8E3DC38B70DD526CE9F30D80F3AEFA16" originalValue="páramo">paramo</normalizedToken>
muy 
<normalizedToken id="654439DD8923D3E57A820B104A527A9F" originalValue="húmedo">humedo</normalizedToken>
dominado por el grupo 
<taxonomicName id="9A587EB76971B05FE57057501E611428" class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="EADEA69C2E3BE23667C5D7CBCD55EAEE" italics="true" pageId="0" pageNumber="107">Chusquea</emphasis>
</taxonomicName>
, pastoreo natural, 
<geoCoordinate id="98C75F75FB25D53A903998CC9A812BEE" degrees="5.75425" direction="north" orientation="latitude" precision="1" value="5.75425">5.75425N</geoCoordinate>
, 
<geoCoordinate id="C0869E13814EC75A8C84579FD257745F" degrees="73.38303" direction="west" orientation="longitude" precision="1" value="-73.38303">73.38303W</geoCoordinate>
, 3430 m alt., 15 Nov. 2017, S.P. Sylvester 3071 (K, US, SI). Munic. Chiscas, 
<normalizedToken id="7A3306C174C2968CA077B8A1F032A26C" originalValue="Páramo">Paramo</normalizedToken>
de Chacaritas, rocas expuestas en 
<normalizedToken id="830E362575229CA8729C2670686F683B" originalValue="vegetación">vegetacion</normalizedToken>
de 
<normalizedToken id="4F5D1CF99E4F06A810613AE219DFE6F4" originalValue="páramo">paramo</normalizedToken>
, 
<geoCoordinate id="D934CED85D9D937E9FC5587DAF90358A" degrees="6.61931" direction="north" orientation="latitude" precision="1" value="6.61931">6.61931N</geoCoordinate>
, 
<geoCoordinate id="F0BFA959CA42EF506F4D89685EC3009D" degrees="72.38898" direction="west" orientation="longitude" precision="1" value="-72.38898">72.38898W</geoCoordinate>
, 4287 m alt., 4 Mar. 2018, S.P. Sylvester 3124 (K, US, SI). Munic. Mongua, 
<normalizedToken id="A4B8F53E705682A47C352DA906683A72" originalValue="Páramo">Paramo</normalizedToken>
de 
<normalizedToken id="8178D110DD4C7DCA7D6C3632E9D87BC5" originalValue="Ocetá">Oceta</normalizedToken>
, valle de Laguna Negra, 
<normalizedToken id="1D66F525F51DF71A4C95DA752964F9FD" originalValue="vegetación">vegetacion</normalizedToken>
de frailejonal pajonal, con presencia de arbustos 
<normalizedToken id="686F8E59B27F883479812263235995DB" originalValue="pequeños">pequenos</normalizedToken>
, Se observan rastros de pastoreo, 
<geoCoordinate id="68469E81F7520684CB6B18D922EB2FFF" degrees="5.69525" direction="north" orientation="latitude" precision="1" value="5.69525">5.69525N</geoCoordinate>
, 
<geoCoordinate id="8DCDBD11CA32F3F1B7609F6CBB33D18F" degrees="72.79133" direction="west" orientation="longitude" precision="1" value="-72.79133">72.79133W</geoCoordinate>
, 3694 m alt., 29 Nov. 2017, L.E. 
<normalizedToken id="F7BAC140AEBA56386E6D73821CCED808" originalValue="Cuta-Alarcón">Cuta-Alarcon</normalizedToken>
353c (US); Planos del Toldadero, [
<geoCoordinate id="730EDE23A6ED3DA88CED34911F8DA91D" degrees="6.3669" direction="north" orientation="latitude" precision="5" value="6.3669">6.3669N</geoCoordinate>
, 
<geoCoordinate id="8569816722D67CB9AEA3666EC7A4055C" degrees="72.3342" direction="west" orientation="longitude" precision="5" value="-72.3342">72.3342W</geoCoordinate>
], 3950 m alt., 13 Sep. 1938, J. Cuatrecasas 1554 (US17730370); Sierra Nevada del Cocuy, alrededores de Salto de Correlitos, [
<geoCoordinate id="1B5F4C054A13F6A3B0840CA0DDD952FE" degrees="6.4444" direction="north" orientation="latitude" precision="5" value="6.4444">6.4444N</geoCoordinate>
, 
<geoCoordinate id="A416D03CF3F3F3AD9A829485766072CB" degrees="72.3175" direction="west" orientation="longitude" precision="5" value="-72.3175">72.3175W</geoCoordinate>
], 4400 m alt., 14 Apr. 1959, H.G. Barclay 7394 (US2434358); Sierra Nevada del Cocuy, Alto Ritacuva, [
<geoCoordinate id="65B91F457C3C0A9EECE6B52E20AFD8C9" degrees="6.5139" direction="north" orientation="latitude" precision="5" value="6.5139">6.5139N</geoCoordinate>
, 
<geoCoordinate id="BFD5AA0219BF6F935AD5703BEFC863D5" degrees="72.3531" direction="west" orientation="longitude" precision="5" value="-72.3531">72.3531W</geoCoordinate>
], 4450 m alt., between wet slopes which have dense 
<taxonomicName id="3DC2C215F786627698CF09A9B56A4EC8" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="A6B8EA3A161EC9EDFC4A413F2C62FDAF" italics="true" pageId="0" pageNumber="107">Espeletia</emphasis>
</taxonomicName>
, 24 Apr. 1959, H.G. Barclay 7453 (US2434361); Sierra Nevada del Cocuy, alrededores de Salto de Correlitos, rocky, southwest-facing slope, better vegetated between rocky ridges, east of Laguna San Paulino, [
<geoCoordinate id="B6F32088F4EC90E2C348E4296425AE4A" degrees="6.4444" direction="north" orientation="latitude" precision="5" value="6.4444">6.4444N</geoCoordinate>
, 
<geoCoordinate id="270E5389132EF252CD4EBBE0783C5183" degrees="72.3175" direction="west" orientation="longitude" precision="5" value="-72.3175">72.3175W</geoCoordinate>
], 4300 m alt., 14 Apr. 1959, H.G. Barclay 7381 (US2434355); Sierra Nevada del Cocuy, alto valle de Las Lagunillas, [
<geoCoordinate id="5AB19625264D4416B06DF68F6EE83761" degrees="6.3469" direction="north" orientation="latitude" precision="5" value="6.3469">6.3469N</geoCoordinate>
, 
<geoCoordinate id="ECCAC1BE511E28521E635A10E9319E28" degrees="72.3261" direction="west" orientation="longitude" precision="5" value="-72.3261">72.3261W</geoCoordinate>
], 4000-4300 m alt., 12 Sep. 1938, J. Cuatrecasas 1465 (US1772998); Sierra Nevada del Cocuy, Alto Valle Lagunillas, morrena seca pedregosa 100 m al SE de la Laguna Cuadrada, [
<geoCoordinate id="CB98C8BBFD35F712F7B52172373CE444" degrees="6.3619" direction="north" orientation="latitude" precision="5" value="6.3619">6.3619N</geoCoordinate>
, 
<geoCoordinate id="EB29B865EF71160070BE6069CE608AEA" degrees="72.3344" direction="west" orientation="longitude" precision="5" value="-72.3344">72.3344W</geoCoordinate>
], 4080 m alt., con 
<taxonomicName id="1FB53730707EF6D70EBC20EA8E35CE88" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis effusa" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="effusa">
<emphasis id="783A41620962CA2CAD9737413E807423" italics="true" pageId="0" pageNumber="107">Calamagrostis effusa</emphasis>
</taxonomicName>
[(Kunth) Steud. = 
<taxonomicName id="3FE07F84D9CD9BB91AE77716B7C39033" genus="Paramochloa" lsidName="Paramochloa effusa" pageId="0" pageNumber="107" rank="species" species="effusa">
<emphasis id="403CEA4DEA2E16547CF94FD7419EF69C" italics="true" pageId="0" pageNumber="107">Paramochloa effusa</emphasis>
</taxonomicName>
(Kunth) P.M. Peterson, Soreng, Romasch. &amp; 
<normalizedToken id="49A945E3D41121EA2FF2FF7D413AF2A7" originalValue="Barberá">Barbera</normalizedToken>
] y 
<taxonomicName id="68DA6C0461F284AC3F660BAC0812DAEB" authorityName="P.M.Peterson, R.J.Soreng, K.Romaschenko &amp; P.Barberá" authorityYear="2019" class="Jungermanniopsida" family="Gymnomitriaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Jungermanniales" pageId="0" pageNumber="107" phylum="Marchantiophyta" rank="family">Gymnomitriaceae</taxonomicName>
predominantes, asociadas con 
<taxonomicName id="DD4457B9CD9267A1BDED3FF76469C1D5" class="Liliopsida" family="Juncaceae" genus="Luzula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Luzula" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="6FECE70E0C5361691DCD7DE4ED54CA14" italics="true" pageId="0" pageNumber="107">Luzula</emphasis>
</taxonomicName>
, 
<taxonomicName id="CE67E676EAF9896BE32018F1FAC08FBF" authorityName="Cuatrec" authorityYear="1940" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia colombiana" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="colombiana">
<emphasis id="8D1789621A71A39535558744CBF272DF" italics="true" pageId="0" pageNumber="107">Espeletia colombiana</emphasis>
</taxonomicName>
[Cuatrec.] y musgos, 26 Nov. 1972, A.M. Cleef 5535 (US2785729); Sierra Nevada del Cocuy, Alto Valle Lagunillas, [
<geoCoordinate id="B64E2546E3882648913023A94CA21F8F" degrees="6.3906" direction="north" orientation="latitude" precision="5" value="6.3906">6.3906N</geoCoordinate>
, 
<geoCoordinate id="951C40BD18FB606084378D827820E5A5" degrees="72.3542" direction="west" orientation="longitude" precision="5" value="-72.3542">72.3542W</geoCoordinate>
], 3995 m alt., con 
<taxonomicName id="6A086F55248433BCACDD53FE9C9DD45A" authorityName="Cuatrec" authorityYear="1940" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia colombiana" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="colombiana">
<emphasis id="D080F735BB2E1DB217DC637E8130C249" italics="true" pageId="0" pageNumber="107">Espeletia colombiana</emphasis>
</taxonomicName>
predominante, asociada con 
<taxonomicName id="8A474B2F6522047E7D692DB927CAA1A2" authorityName="Benth" authorityYear="1881" class="Liliopsida" family="Poaceae" genus="Aciachne" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aciachne pulvinata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="pulvinata">
<emphasis id="A9FCF4B1DF9E502AEB5D6458C1345024" italics="true" pageId="0" pageNumber="107">Aciachne pulvinata</emphasis>
</taxonomicName>
[Benth.], 
<taxonomicName id="C44AF0144CEFB4014728FF8BF80D3FB2" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis breviculmis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="breviculmis">
<emphasis id="A5B0C63B751DBC9E0348A8F5CC71628F" italics="true" pageId="0" pageNumber="107">Agrostis breviculmis</emphasis>
</taxonomicName>
, 
<taxonomicName id="AA664F50293543EFEF653517965625FF" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis trichodes" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="trichodes">
<emphasis id="699886B0F20C516CAEB9FDB77928D7F1" italics="true" pageId="0" pageNumber="107">Agrostis trichodes</emphasis>
</taxonomicName>
[
<taxonomicName id="7515231D5428E8777149B795C73A9F9D" class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis trichodes" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="trichodes">
<emphasis id="ABE8C92B1EB81D469A7A133E3EB5F331" italics="true" pageId="0" pageNumber="107">Podagrostis trichodes</emphasis>
</taxonomicName>
] y 
<taxonomicName id="99489B8C108E184847513F583EB3A0DB" authorityName="Ruiz &amp; Pavon" authorityYear="1798" class="Magnoliopsida" family="Rosaceae" genus="Acaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acaena cylindristachya" order="Rosales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="cylindristachya">
<emphasis id="D91D7AACE6BC81E6863290AC7B0FC014" italics="true" pageId="0" pageNumber="107">Acaena cylindristachya</emphasis>
</taxonomicName>
[Ruiz &amp; Pav.], 
<normalizedToken id="099D1D0481099B927DBE7FBAE6526056" originalValue="gramínea">graminea</normalizedToken>
<normalizedToken id="848F6FB4F0D5745C44416F378C983C06" originalValue="común">comun</normalizedToken>
, 25 Sep. 1972, A.M. Cleef 5504 (US2785744); Sierra Nevada del Cocuy, Boqueron de Cusiri, 
<normalizedToken id="CFCC54FF279B642A63BC133AE36FBE39" originalValue="límite">limite</normalizedToken>
<normalizedToken id="3B630CF5137280152D6D6AA86DA38580" originalValue="superpáramo">superparamo</normalizedToken>
y 
<normalizedToken id="8A2B3B680D30CB139BBF8AFC41BB7160" originalValue="páramo">paramo</normalizedToken>
propiamente dicho, vertiente seco muy pedregoso, [
<geoCoordinate id="F032419F6601F2C003B70D3C37232B31" degrees="6.3431" direction="north" orientation="latitude" precision="5" value="6.3431">6.3431N</geoCoordinate>
, 
<geoCoordinate id="F06E5D2397651DAE454B570593E32C16" degrees="72.3128" direction="west" orientation="longitude" precision="5" value="-72.3128">72.3128W</geoCoordinate>
], 4320 m alt., con 
<taxonomicName id="6F83B24067F26121AE017F0533B658E6" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis effusa" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="effusa">
<emphasis id="EFB8C872DE5F218F75BE32625CE471DC" italics="true" pageId="0" pageNumber="107">Calamagrostis effusa</emphasis>
</taxonomicName>
[= 
<taxonomicName id="9F49BB73BF941F17BA8AE53CF5612993" genus="Paramochloa" lsidName="Paramochloa effusa" pageId="0" pageNumber="107" rank="species" species="effusa">
<emphasis id="ED35404DC863C3ECB6068FE273D8AE31" italics="true" pageId="0" pageNumber="107">Paramochloa effusa</emphasis>
</taxonomicName>
] y 
<taxonomicName id="8F6692FD69BD683D965D4D7316445092" authorityName="Cuatrec" authorityYear="1975" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia cleefii" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="cleefii">
<emphasis id="6BFE84312ADC4F6C434968C916481972" italics="true" pageId="0" pageNumber="107">Espeletia cleefii</emphasis>
</taxonomicName>
[Cuatrec.], 
<normalizedToken id="1150D0146676DF8636BC0934C78BDFE0" originalValue="gramínea">graminea</normalizedToken>
<normalizedToken id="51DE95153C98EDA1CA2BCC3C42C30F78" originalValue="común">comun</normalizedToken>
, 5 Mar. 1973, A.M. Cleef 8796 (US2785680); Sierra Nevada del Cocuy, Laguna Grande de la Sierra, [
<geoCoordinate id="DAE048741E6113A83407F17A7D3D4928" degrees="6.5556" direction="north" orientation="latitude" precision="5" value="6.5556">6.5556N</geoCoordinate>
, 
<geoCoordinate id="50FF1E9E4FE85B1D6E1A050A1BD5216A" degrees="72.3253" direction="west" orientation="longitude" precision="5" value="-72.3253">72.3253W</geoCoordinate>
], 4300-4500 m alt., 28 Dec. 1985, J.R.I. Wood 5247 (US3481052); Sierra Nevada del Cocuy, 
<normalizedToken id="D6C3CFD85A420FE68F0D0D9F7095401E" originalValue="Páramo">Paramo</normalizedToken>
<normalizedToken id="2221FBEC8BFB24362E6B5C897C90EFA1" originalValue="Cóncavo">Concavo</normalizedToken>
, Cuchilla Puentepiedra casi 2km al NE de la Laguna Pintada, vertiente pedregoso seco, [
<geoCoordinate id="ED7D91B3C2FC8B7571DE98D14176CD10" degrees="6.3723" direction="north" orientation="latitude" precision="5" value="6.3723">6.3723N</geoCoordinate>
, 
<geoCoordinate id="AE528166ADEFC4A1A1E78FA0AEA01314" degrees="72.3175" direction="west" orientation="longitude" precision="5" value="-72.3175">72.3175W</geoCoordinate>
], 4510 m alt., con 
<taxonomicName id="9A914AE0D73D4F887D62C6D8DEE78F1F" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis effusa" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="effusa">
<emphasis id="960164B12DAEECB5ACB2C3F098D9B08E" italics="true" pageId="0" pageNumber="107">Calamagrostis effusa</emphasis>
</taxonomicName>
[= 
<taxonomicName id="08514F50FAB1DAEC6CBAA2FB9DE5FD34" genus="Paramochloa" lsidName="Paramochloa effusa" pageId="0" pageNumber="107" rank="species" species="effusa">
<emphasis id="F9D3FEDF87144632D7382DDDFB69BCEE" italics="true" pageId="0" pageNumber="107">Paramochloa effusa</emphasis>
</taxonomicName>
], 
<taxonomicName id="7853E8A0BD933F59F045D9C18F5CCED2" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="7A63295C56573477B81638EA7CF8FF30" italics="true" pageId="0" pageNumber="107">Poa</emphasis>
</taxonomicName>
sp., asociadas con 
<taxonomicName id="507DC3DCFFACE5EB2BB55D136637CC11" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia lopezii" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="lopezii">
<emphasis id="F3269D84E1557887D699076311EB0321" italics="true" pageId="0" pageNumber="107">Espeletia lopezii</emphasis>
</taxonomicName>
fma. [Cuatrec.], y musgos, 30 Sep. 1972, A.M. Cleef 5665 (US2785695); Sierra Nevada del Cocuy, 
<normalizedToken id="3D096A6A6DE9ED240FAF2E611756369E" originalValue="Páramo">Paramo</normalizedToken>
<normalizedToken id="0CE67BC8637E3C99B8E6E78F87D2DCB4" originalValue="Cóncavo">Concavo</normalizedToken>
, morrena seca en el 
<normalizedToken id="EA303D561BE44DEAA3BB4E26260933C4" originalValue="límite">limite</normalizedToken>
<normalizedToken id="143A6ECBC05342EACDF90791CCAF2B64" originalValue="páramo">paramo</normalizedToken>
propiamente dicho y 
<normalizedToken id="86295779ABB5ECB778ED1980751656D9" originalValue="superpáramo">superparamo</normalizedToken>
, 3.5 km al NNW del Morro Pulpito del Diablo, [6.3995N, 72.3123W], 4325m alt., matorral de 
<taxonomicName id="A5794894A81C4524CA768B696A88690A" class="Magnoliopsida" family="Asteraceae" genus="Senecio" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Senecio vaccinioides" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="vaccinioides">
<emphasis id="F12EEBE9140FB9752C73806A71C9CAE6" italics="true" pageId="0" pageNumber="107">Senecio vaccinioides</emphasis>
</taxonomicName>
[Cuatrec.] con 
<taxonomicName id="BB156C9BBE9EE85D7BA82B1B614C395F" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alchemilla" order="Rosales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="53778C5E4260DD914A2BA0954006D4BD" italics="true" pageId="0" pageNumber="107">Alchemilla</emphasis>
</taxonomicName>
sp., 1 Mar. 1973, A.M. Cleef 8680 (US2785780); Sierra Nevada del Cocuy, 
<normalizedToken id="616B5E30552BA623FEF7659D5F33D12C" originalValue="Páramo">Paramo</normalizedToken>
<normalizedToken id="9A3509E4E96B0B11630EBEDBDDB10447" originalValue="Cóncavo">Concavo</normalizedToken>
, 
<normalizedToken id="D2F0EC87E907C52E422E866B1C685915" originalValue="superpáramo">superparamo</normalizedToken>
, morrena seca 3 km aprox. al Norte del morro Pulpito del Diablo. [
<geoCoordinate id="1AA85F1BE44D41EEF50AECD3796D9C72" degrees="6.3995" direction="north" orientation="latitude" precision="5" value="6.3995">6.3995N</geoCoordinate>
, 
<geoCoordinate id="B081B788F04F6AE8CBE8F33655716502" degrees="72.3123" direction="west" orientation="longitude" precision="5" value="-72.3123">72.3123W</geoCoordinate>
], 4375m alt., rastrojo de 
<taxonomicName id="643C3D5EBE4245F7697C637FC81A8045" authorityName="Cuatrecasas" authorityYear="1940" class="Magnoliopsida" family="Asteraceae" genus="Senecio" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Senecio guicanensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="guicanensis">
<emphasis id="7891FA901C2E5BEF27BA16FCF5A8A7F6" italics="true" pageId="0" pageNumber="107">Senecio guicanensis</emphasis>
</taxonomicName>
[Cuatrec.], 
<normalizedToken id="5B8CFCBE04067BC21ACFB593FDA3C027" originalValue="gramínea">graminea</normalizedToken>
<normalizedToken id="1A1C0E665573319772EDD8FC790ED34E" originalValue="común">comun</normalizedToken>
, 28 Feb. 1973, A.M. Cleef 8627 (US2785676); Sierra Nevada del Cocuy, 
<normalizedToken id="BA516D0613C20131413DC948BD390A26" originalValue="Páramo">Paramo</normalizedToken>
<normalizedToken id="E18D1696C52734C72293DBE1B2FE1C98" originalValue="Cóncavo">Concavo</normalizedToken>
, 
<normalizedToken id="4DC6B85D094B2CC950A6DE68684F37B8" originalValue="superpáramo">superparamo</normalizedToken>
3.5 km aprox. al NNW del morro Pulpito del Diablo, morrena seca muy pedegrosa, [
<geoCoordinate id="A7486381482DD24E3F41540CA77844F2" degrees="6.3995" direction="north" orientation="latitude" precision="5" value="6.3995">6.3995N</geoCoordinate>
, 
<geoCoordinate id="9C116153E0B620B6352178DCB92099DB" degrees="72.3122" direction="west" orientation="longitude" precision="5" value="-72.3122">72.3122W</geoCoordinate>
], 4315 m alt., con 
<taxonomicName id="2BF54C09131BBEE5A765B741D6F3A6FA" class="Liliopsida" family="Juncaceae" genus="Luzula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Luzula" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="4EE33B00A94DADEBD5F1331D8E0937F2" italics="true" pageId="0" pageNumber="107">Luzula</emphasis>
</taxonomicName>
y 
<taxonomicName id="85E45A197C3379DAC46A3FB26D5A3804" authority="var. prostrata" class="Magnoliopsida" family="Ericaceae" genus="Pernettya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pernettya prostrata subsp. var. var. prostrata" order="Ericales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="variety" species="prostrata" subSpecies="var." variety="prostrata">Pernettya prostrata var. prostrata</taxonomicName>
[(Cav.) DC.], 
<normalizedToken id="B5625AAA140064D703996E3C56231A8B" originalValue="gramínea">graminea</normalizedToken>
<normalizedToken id="F6C49BB806CA2C79E41024E66B8A89A6" originalValue="común">comun</normalizedToken>
, 26 Feb. 1973, A.M. Cleef 8504 (US01247250); Sierra Nevada del Cocuy, 
<normalizedToken id="A63B750E716E55FD77AAA178462AC9A6" originalValue="Páramo">Paramo</normalizedToken>
<normalizedToken id="517BCA0A63DF3346FE2CAE2806AAF9EC" originalValue="Cóncavo">Concavo</normalizedToken>
, 
<normalizedToken id="DFEF698DA634B58109B8A0ADBDC53A38" originalValue="superpáramo">superparamo</normalizedToken>
abrigo rocoso denominado Cueva de los Hombres, 3 km aprox. al N del morro Pulpito del Diablo, [
<geoCoordinate id="6AE1E2CD1B7B70527D202503B8DBDE5A" degrees="6.3995" direction="north" orientation="latitude" precision="5" value="6.3995">6.3995N</geoCoordinate>
, 
<geoCoordinate id="04B65307754382D730000EDDDF791930" degrees="72.3122" direction="west" orientation="longitude" precision="5" value="-72.3122">72.3122W</geoCoordinate>
], 4350 m alt., 28 Feb. 1973, A.M. Cleef 8608 (US2785679); Sierra Nevada del Cocuy, Quebrada Bocatoma, vertiente N del valle, 800 m al ENE [East North-East] de la Laguna Pintada, [6.4795N, 72.3167W], 4060 m alt., 
<taxonomicName id="F605B4061F2761B4BAD67438365FAD48" authorityName="J.B.Jack" authorityYear="1894" class="Insecta" family="Cecidomyiidae" genus="Stephaniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephaniella" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="14EA25579BA6A5786774E7ABC49CB5ED" italics="true" pageId="0" pageNumber="107">Stephaniella</emphasis>
</taxonomicName>
sp. predominante asociada con 
<taxonomicName id="CE82D8F40CAF53EB8BBA3C80B792D622" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis effusa" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="effusa">
<emphasis id="22D9D237478306D320350A6DC02E743C" italics="true" pageId="0" pageNumber="107">Calamagrostis effusa</emphasis>
</taxonomicName>
[= 
<taxonomicName id="4F3BEF37813FB92B639901ADD6DE6BBD" genus="Paramochloa" lsidName="Paramochloa effusa" pageId="0" pageNumber="107" rank="species" species="effusa">
<emphasis id="CB176FDED4C3EBD54C9FF690F9410224" italics="true" pageId="0" pageNumber="107">Paramochloa effusa</emphasis>
</taxonomicName>
], 
<taxonomicName id="1837F0F341C5D8CD8355405B4C9F6FE0" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alchemilla polylepis" order="Rosales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="polylepis">
<emphasis id="2D953E8BB09B24E74A0542E4A33AC3C2" italics="true" pageId="0" pageNumber="107">Alchemilla polylepis</emphasis>
</taxonomicName>
[Wedd.], 
<taxonomicName id="09237DD265E41183FA93106A2FB499DD" class="Magnoliopsida" family="Asteraceae" genus="Achyrocline" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achyrocline lehmanii" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="lehmanii">
<emphasis id="E09EF1769A63AAAA9928B4EFFA843A8E" italics="true" pageId="0" pageNumber="107">Achyrocline lehmanii</emphasis>
</taxonomicName>
[Hieron.] y 
<taxonomicName id="E94DD505C0D3CED81C8BC1BE94403290" class="Bryopsida" family="Polytrichaceae" genus="Polytrichum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Polytrichum" order="Polytrichales" pageId="0" pageNumber="107" phylum="Bryophyta" rank="genus">
<emphasis id="364113787296ADB54138F6AE37D9CCBB" italics="true" pageId="0" pageNumber="107">Polytrichum</emphasis>
</taxonomicName>
sp., 29 Sep. 1972, A.M. Cleef 5648 (US2785702).
</paragraph>
</subSubSection>
<subSubSection id="6B01BA5633F58352F4F77F1A3ACEE406" pageId="0" pageNumber="107" type="notes">
<paragraph id="0E2009737DF5B809ACC915A3A37D4451" pageId="0" pageNumber="107">Notes.</paragraph>
<paragraph id="E1B5FAE5A6C19FE9493C7288052C9CB9" pageId="0" pageNumber="107">
Specimens encountered from 
<normalizedToken id="4F6ABFE0DD7B69F038D19B2C850C7EEC" originalValue="páramos">paramos</normalizedToken>
not belonging to the Sierra Nevada del Cocuy, but within Departamento 
<normalizedToken id="AD1731489E9D5B54DD1982BEF49DD923" originalValue="Boyacá">Boyaca</normalizedToken>
, bear notable differences from those from the type locality and further study is needed to elucidate whether these are a distinct species or subspecies of 
<taxonomicName id="76FF271DC2823A7C5A24FFB3B70FE73C" lsidName="A. boyacensis" pageId="0" pageNumber="107" rank="species" species="boyacensis">
<emphasis id="1098234E590E020063CDBED17C087167" italics="true" pageId="0" pageNumber="107">A. boyacensis</emphasis>
</taxonomicName>
. These differences include leaf sheaths and blades, panicle branches, and pedicels being usually smooth or very lightly scaberulous, and plants being larger, usually&gt; 20 cm tall and up to 37 cm tall, and with the panicles largely exerted from the basal foliage (Fig. 
<figureCitation id="F759E310D1D8A22B0D066A067F0AAECF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Agrostis boyacensis, example of a specimen encountered in paramos of Boyaca outside of the Sierra Nevada del Cocuy A whole plant B spikelet, glumes lateral view and floret ventral view showing anthers C spikelet, glumes lateral view with floret detached and raised above the glumes showing ventral surface. Images of Sylvester 3017 a (FMB)." figureDoi="10.3897/phytokeys.151.50538.figure2" httpUri="https://binary.pensoft.net/fig/424840" pageId="0" pageNumber="107">2</figureCitation>
). These characteristics place it close to 
<taxonomicName id="28F4408ED86DF588BE47A7F2E7148C9F" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
<emphasis id="4484A614C6700F85970E815AA5718CC4" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
</taxonomicName>
, which is not known from Colombia (
<bibRefCitation id="CF69929E58F65BB7DD6D81D19172E76F" author="Giraldo-Canas, D" editor="Bernal, R" journalOrPublisher="2). Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota D. C." pageId="0" pageNumber="107" pagination="2127 - 2205" refId="B10" refString="Giraldo-Canas, D, Londono, X, Clark, LG, 2016. Poaceae. In: Bernal, R, Gradstein, SR, Celis, M, Eds., Catalogo de plantas y liquenes de Colombia (Vol. 2). Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota D.C.: 2127 - 2205" title="Poaceae." volumeTitle="Catalogo de plantas y liquenes de Colombia (Vol." year="2016">
<normalizedToken id="92AFF8F3B2A294A905F957DACD45C0F2" originalValue="Giraldo-Cañas">Giraldo-Canas</normalizedToken>
et al. 2016
</bibRefCitation>
) or 
<normalizedToken id="A050731F1A091F2D5AA81C2A5CE1A164" originalValue="páramos">paramos</normalizedToken>
in general (
<bibRefCitation id="BD7C7964209EEF1A3CC2D53148A3827A" author="Luteyn, JL" journalOrPublisher="Memoirs of the New York Botanical Garden" pageId="0" pageNumber="107" pagination="1 - 278" refId="B20" refString="Luteyn, JL, 1999. Paramos, a checklist of plant diversity, geographical distribution, and botanical literature. Memoirs of the New York Botanical Garden 84: 1 - 278" title="Paramos, a checklist of plant diversity, geographical distribution, and botanical literature." volume="84" year="1999">Luteyn 1999</bibRefCitation>
), although it is mentioned in the recently-published abbreviated key to some Ecuadorian species (
<bibRefCitation id="0F3840CC5FE41F802C450615787D1F24" author="Palacio, PC" journalOrPublisher="Nature" pageId="0" pageNumber="107" refId="B24" refString="Palacio, PC, Molina, AM, Rugolo de Agrasar, ZE, Vega, AS, in press. A new species of Agrostis (Poaceae, Pooideae, Poeae, Agrostidinae) from the Andean paramos of Ecuador. Novon." title="A new species of Agrostis (Poaceae, Pooideae, Poeae, Agrostidinae) from the Andean paramos of Ecuador. Novon." year="in press">Palacio et al. in press</bibRefCitation>
) (see below for how to differentiate these species).
</paragraph>
<paragraph id="70186A7AF746074388969B3E352B893C" pageId="0" pageNumber="107">
The habit of type specimens and some of the other specimens examined (e.g. Sylvester 3117 and 3129), with perenniating, many-branched culms, appears to be related to the habitat, with the Sylvester 3117 and 3129 specimens being found growing amongst moss. The leaf blades of these were also slightly laxer, albeit still involute or strongly conduplicate. In harsher conditions, such as open gravelly slopes, the tufts are more compact and leaf blades are rigid and strongly rolled and resemble a very large, densely-tufted 
<taxonomicName id="D2076570236E22B910BAEC81113E3938" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
<emphasis id="A2581D80CBF63ED6EBCC4D8A997F2A90" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
</taxonomicName>
with slightly laxer panicles.
</paragraph>
<caption id="E870670054FEBEA64AD86582F2C9BF92" doi="10.3897/phytokeys.151.50538.figure2" httpUri="https://binary.pensoft.net/fig/424840" pageId="0" pageNumber="107" start="Figure 2" startId="F2">
<paragraph id="BD60A50AEFE66C618B3E7073F0F0D0CE" pageId="0" pageNumber="107">
<emphasis id="3FA5AB17379D31D53FFECA465B212013" bold="true" pageId="0" pageNumber="107">Figure 2.</emphasis>
<taxonomicName id="2D806A3EF2AA1DC7CE1153B98B0F3805" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis boyacensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
<emphasis id="0CDD3219936BA6AD34AE07AF3CB67672" italics="true" pageId="0" pageNumber="107">Agrostis boyacensis</emphasis>
</taxonomicName>
, example of a specimen encountered in 
<normalizedToken id="6385ECAEAF0ADAEA6B41989558EA7F14" originalValue="páramos">paramos</normalizedToken>
of 
<normalizedToken id="DC476D91A0E37E47A4E3258FA980137E" originalValue="Boyacá">Boyaca</normalizedToken>
outside of the Sierra Nevada del Cocuy 
<emphasis id="6FA8D3797D08D5AF314672A91D481DD4" bold="true" pageId="0" pageNumber="107">A</emphasis>
whole plant 
<emphasis id="D54122FD577DA9A18F7A889A6F18321F" bold="true" pageId="0" pageNumber="107">B</emphasis>
spikelet, glumes lateral view and floret ventral view showing anthers 
<emphasis id="BE9A70F11B63E55456505ED75AF9FD45" bold="true" pageId="0" pageNumber="107">C</emphasis>
spikelet, glumes lateral view with floret detached and raised above the glumes showing ventral surface. Images of Sylvester 3017a (FMB).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="13247120013833445C3E00FABCE86E8F" pageId="0" pageNumber="107" type="similar species">
<paragraph id="BAF8D8D4764B1229AAC42867F4559602" pageId="0" pageNumber="107">Similar species.</paragraph>
<paragraph id="E643E93387D03FEFB44C402B353D0AE9" pageId="0" pageNumber="107">
<taxonomicName id="4FC85BB351EA32AFCDA693713F9F25FC" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis breviculmis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="breviculmis">
<emphasis id="89DD40384E5F1BEA5803ED4857EAA4AA" italics="true" pageId="0" pageNumber="107">Agrostis breviculmis</emphasis>
</taxonomicName>
and 
<taxonomicName id="6E703EB21DDDFF7704B69932695E119C" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
<emphasis id="72A504A33BAEEA6B4B6E14BAD9E5DEF7" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
</taxonomicName>
both have congested spikelike panicles and florets that lack prominent awns and well-developed paleas. 
<taxonomicName id="3EEA55C526F7BA6876E4B5503FB3EE67" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis breviculmis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="breviculmis">
<emphasis id="10861845FB245642D0EEBCE623DE371D" italics="true" pageId="0" pageNumber="107">Agrostis breviculmis</emphasis>
</taxonomicName>
bears close similarity in its convolute, often recurved, rigid to firm leaf blades that usually measure &lt;1 mm wide in diameter, and small spikelets with fairly well developed coarse scabers on the glume keels. 
<taxonomicName id="5379BA0C1D47ABDC8D93E85789851918" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis boyacensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
<emphasis id="6EAD03CC9D4EF8886CA880752E2C7BFF" italics="true" pageId="0" pageNumber="107">Agrostis boyacensis</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="C6EBE3CD907016797B42429934999286" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
<emphasis id="A9B55CE51A9A5B79EF78C80C25CA3B8E" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
</taxonomicName>
principally by the habit, with many extravaginally branched culms that form large dense tufts to 37 cm tall (vs. intravaginal innovations forming short tufts to 12(-15) cm tall in 
<taxonomicName id="73A8491E3A03EEE891C08377A41438FC" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
<emphasis id="DE7A21AC7B789CEC40E83252B49CD2E3" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
</taxonomicName>
), laxer panicles, 3-9 mm wide (vs. c. 0.5-2 mm wide in 
<taxonomicName id="1BC8998E20E4F890995F25A867FBB697" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
<emphasis id="E9E71115B977D5D20A1EC8FB92AC9356" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
</taxonomicName>
), and slightly larger spikelets, 1.8-2.4(-2.5) mm long (vs. usually 1.5-2.1 mm long in 
<taxonomicName id="022016CE15AFC752A9235889DDB9A49A" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
<emphasis id="79CA8052D96EC6B0C1CF82D7E0707C9B" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
</taxonomicName>
, noted to reach 2.5 mm long in Bolivia [
<bibRefCitation id="EEA62D42BE984911993292D4CA0D0BDD" author="Renvoize, SA" journalOrPublisher="Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico" pageId="0" pageNumber="107" refId="B29" refString="Renvoize, SA, 1998. Gramineas de Bolivia. Continental Printing, Belgium." title="Gramineas de Bolivia. Continental Printing, Belgium." year="1998">Renvoize 1998</bibRefCitation>
]).
</paragraph>
<paragraph id="74588A5E08132641FFF76660399C63CE" pageId="0" pageNumber="107">
<taxonomicName id="E489294DBB4E78374597F84D334DC24A" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis boyacensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
<emphasis id="E001673B36B4B5D83C8799FDEA3D82AB" italics="true" pageId="0" pageNumber="107">Agrostis boyacensis</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="F162C2A405752A003BD8AE6C59942B77" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
<emphasis id="6016053D6D0042A11A46538A4CACEB67" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
</taxonomicName>
principally by its robust convolute, involute or strongly conduplicate, usually recurved and rigid leaf blades (vs. laxer, weaker, flat or folded, usually filiform leaf blades in 
<taxonomicName id="57CEA6849A5F782254E08151C44BA938" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
<emphasis id="F99DA8CB9FFDFE35E16E77A26BF571FF" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
</taxonomicName>
). All specimens encountered from the Sierra Nevada del Cocuy can be easily differentiated by their panicle branches, pedicels, and leaf sheaths and blades that are moderately to densely scabrous (vs. panicle branches and pedicels smooth to lightly scaberulous, leaf sheaths smooth, blades smooth or scabrous in 
<taxonomicName id="35103A05DEE339098B161BB8B9B4E295" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
<emphasis id="3D5962CE35E9629EDD5D10CCEBB317D2" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
</taxonomicName>
). Specimens collected in other 
<normalizedToken id="591876A864A35E4EE0F30EF1E573205D" originalValue="páramos">paramos</normalizedToken>
from Departamento 
<normalizedToken id="B384571221F95EA79C194BE30DFF0FB1" originalValue="Boyacá">Boyaca</normalizedToken>
, outside of the Sierra Nevada del Cocuy, bear further similarities with 
<taxonomicName id="8CECFA695C865278468B5C727D1E5B92" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
<emphasis id="E953229162E4AF7306915A3A49322B8B" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
</taxonomicName>
, such as their culms and inflorescences being longer and greatly exerted from the basal foliage, and panicle branches, pedicels, and leaf blades that are usually smooth to lightly scaberulous (Fig. 
<figureCitation id="B82BA7B4658E46DA351EA65A00393BB8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Agrostis boyacensis, example of a specimen encountered in paramos of Boyaca outside of the Sierra Nevada del Cocuy A whole plant B spikelet, glumes lateral view and floret ventral view showing anthers C spikelet, glumes lateral view with floret detached and raised above the glumes showing ventral surface. Images of Sylvester 3017 a (FMB)." figureDoi="10.3897/phytokeys.151.50538.figure2" httpUri="https://binary.pensoft.net/fig/424840" pageId="0" pageNumber="107">2</figureCitation>
). Nevertheless, these can be differentiated from 
<taxonomicName id="FED90D5E29FE6C541ADB8463E635E94F" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
<emphasis id="F13D02463062FBC1ABA7C43789EB6103" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
</taxonomicName>
by a) their robust convolute, involute or strongly conduplicate, rigid and usually recurved leaf blades; b) the glumes usually being unequal, with the lower glume longer and usually wider than the upper; and c) the lower glume keel scabrous throughout to only in distal 1/3, the upper glume keel scabrous only in the distal 1/3 or sometimes smooth throughout.
</paragraph>
<paragraph id="5A826BB19301A5062ADC906B778CE96A" pageId="0" pageNumber="107">
<taxonomicName id="43A42CD2DFAD5CA8194535247F79A9D2" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">
<emphasis id="5492B76757BA88E1D0FF08CE6573A6B5" italics="true" pageId="0" pageNumber="107">Agrostis tolucensis</emphasis>
</taxonomicName>
has a congested spikelike panicle and florets with a minute palea and a lemma that can sometimes lack awns (see notes under 
<taxonomicName id="A5D1B6126EFB8EEE47C43EF43CB1F079" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
<emphasis id="F94409C21CECAC4186273E4D26E2BE09" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
</taxonomicName>
). This species usually has leaf blades filiform, flat, or folded, lax to firm, but can sometimes have basal leaf blades involute or convolute and firm to rigid. All specimens examined at US with involute or convolute and rigid leaf blades had lemmas with a well-developed dorsally inserted awn. Further distinction from 
<taxonomicName id="B22F93BDFF5A29C26332AC7F24F06E81" lsidName="A. boyacensis" pageId="0" pageNumber="107" rank="species" species="boyacensis">
<emphasis id="7CBAFDF00864FCDD259FFAC0FF558622" italics="true" pageId="0" pageNumber="107">A. boyacensis</emphasis>
</taxonomicName>
can be found in how the leaf blades are smooth or scabrous only on the margin and sometimes veins in 
<taxonomicName id="687310DD0B707E223F2FFB309867D72D" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
<emphasis id="DAA9893F85B07BF2EDA87193DEDD6D53" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
</taxonomicName>
, while scabrous throughout (margin, veins, and in-between veins) in 
<taxonomicName id="E3EA0896CA3188406E6B428C48C69B22" lsidName="A. boyacensis" pageId="0" pageNumber="107" rank="species" species="boyacensis">
<emphasis id="577D4E37A338EE03076A51BC970FAF81" italics="true" pageId="0" pageNumber="107">A. boyacensis</emphasis>
</taxonomicName>
from the Sierra Nevada del Cocuy, although 
<taxonomicName id="318837D1B03F123627D1B67A2EB73CBE" lsidName="A. boyacensis" pageId="0" pageNumber="107" rank="species" species="boyacensis">
<emphasis id="2E334B0291658858FFE8B0D0EA45213E" italics="true" pageId="0" pageNumber="107">A. boyacensis</emphasis>
</taxonomicName>
specimens from other Boyacan 
<normalizedToken id="0E92900E15058D893D880AAC6F4E2E4F" originalValue="páramos">paramos</normalizedToken>
have blades smooth to scaberulous.
</paragraph>
<paragraph id="8F2604D75AC0E379A1ED430563E08FE3" pageId="0" pageNumber="107">
The sometimes strongly conduplicate leaf blades of this species can also give it a resemblance to 
<taxonomicName id="87363E6617092FEF2CDD59B766F9BC52" lsidName="A. foliata" pageId="0" pageNumber="107" rank="species" species="foliata">
<emphasis id="96FFABE84A50BBC2ACBE16926865D976" italics="true" pageId="0" pageNumber="107">A. foliata</emphasis>
</taxonomicName>
, which has subcoriaceous to coriaceous leaf blades that can sometimes be folded, although these are usually&gt; 1.5 mm wide when opened out, and lemmas with well-developed awns inserted in the lower 1/2 of the lemma.
</paragraph>
</subSubSection>
</treatment>
</document>