treatments-xml/data/03/FF/ED/03FFED24342FCF5838F8FE05473EFDCA.xml

<document id="C74A73C78581304CA41C92C29806C211" ID-DOI="10.1515/aemnp-2017-0088" ID-ISSN="0374-1036" ID-Zenodo-Dep="5315619" ID-ZooBank="447405A9-868F-4D01-8C0B-E7EE7102AD08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630123064569" checkinUser="marcus" docAuthor="Guimarães, Bruno A. C. &amp; Ferreira-Jr, Nelson" docDate="2017" docId="03FFED24342FCF5838F8FE05473EFDCA" docLanguage="en" docName="ActaEntMusNatPra.2017.57.2.521-534.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57 (2)" docSource="http://dx.doi.org/10.1515/aemnp-2017-0088" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Canthysellus yawari Guimarães &amp; Ferreira-Jr 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="533" masterDocId="FFC6955C3426CF543943FF9E4358FF99" masterDocTitle="Four new species and new records of Canthysellus from Brazil (Coleoptera: Noteridae)" masterLastPageNumber="534" masterPageNumber="521" pageNumber="530" updateTime="1699161162332" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7AAE71EA56CA83ED45D4496426840B9B">Four new species and new records of Canthysellus from Brazil (Coleoptera: Noteridae)</mods:title>
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<mods:namePart id="B96BE1AD7F6EE171F732721AACBD06CD">Guimarães, Bruno A. C.</mods:namePart>
<mods:affiliation id="C5B2C74511B4681DDBC8769421A6C9BA">1,2) &amp;) &amp;) Laboratório de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro. Caixa Postal 68044, Rio de Janeiro, RJ, 21941 - 971, Brasil; e-mails: brunoaguilar 0044 @ gmail. com, nferrejr @ gmail. com &amp; 1,2) &amp;) Programa de Pós-graduação em Ciências Biológicas - Zoologia, Museu Nacional, Universidade Federal do Rio de Janeiro. Quinta da Boa Vista, São Cristóvão, Rio de Janeiro, RJ, 20940 - 040, Brasil</mods:affiliation>
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<mods:namePart id="BF2B7A78F43263990D254887A0CE309E">Ferreira-Jr, Nelson</mods:namePart>
<mods:affiliation id="193C1B6B6C7E09FF02071971233D154C">1,2) &amp;) &amp;) Laboratório de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro. Caixa Postal 68044, Rio de Janeiro, RJ, 21941 - 971, Brasil; e-mails: brunoaguilar 0044 @ gmail. com, nferrejr @ gmail. com</mods:affiliation>
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<taxonomicName id="4C5627B1342FCF5D38F8FE0541C2FE2A" authority="Guimarães &amp; Ferreira-Jr, 2017" authorityName="Guimarães &amp; Ferreira-Jr" authorityYear="2017" box="[443,666,411,435]" class="Insecta" family="Noteridae" genus="Canthysellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="yawari" status="sp. nov.">
<emphasis id="B9228020342FCF5D38F8FE0541C2FE2A" bold="true" box="[443,666,411,435]" italics="true" pageId="9" pageNumber="530">Canthysellus yawari</emphasis>
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(
<figureCitation id="136D40B7342FCF5D3B67FE244103FE54" box="[548,603,442,461]" captionStart="Figs 1–4" captionStartId="2.[151,188,982,1001]" captionTargetBox="[155,1114,233,950]" captionTargetId="figure-10@2.[151,1126,229,953]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Canthysellus species in lateral and dorsal view: 1 – C. kukrutkato sp. nov.; 2 – C. putkarot sp. nov.; 3 – C. omawe sp. nov.; 4 – C. yawari sp. nov. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315621" httpUri="https://zenodo.org/record/5315621/files/figure.png" pageId="9" pageNumber="530">Figs 4</figureCitation>
, 
<figureCitation id="136D40B7342FCF5D3B26FE2441D6FE54" box="[613,654,442,461]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="9" pageNumber="530">8a–e</figureCitation>
)
</paragraph>
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locality.
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<collectingCountry id="F3411CA2342FCF5D39ACFE7A427DFE6E" box="[239,293,484,503]" name="Brazil" pageId="9" pageNumber="530">Brazil</collectingCountry>
, Amazonas State, Barcelos Municipality, Aracá Sierra, 
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.
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<emphasis id="B9228020342FCF5D3932FE6043AFFD88" bold="true" box="[113,247,510,529]" pageId="9" pageNumber="530">Type material.</emphasis>
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<typeStatus id="54EDE290342FCF5D39BDFE604209FD89" box="[254,337,510,529]" pageId="9" pageNumber="530" type="holotype">HOLOTYPE</typeStatus>
: J, ‘Brazil, Amazonas State, Barcelos Municipality, Aracá Sierra, Point Bo2, puddles with foliage, 
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, 
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, Ferreira-Jr. N. leg.’ (
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. 
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: 
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: AMAZONAS:
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‘ 
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, 
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, 
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Miranda’s Site, 
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, puddle in the wood, trail to piaçaval, 
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, 
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</date>
, 
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leg.’ (
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/ 
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)
</materialsCitation>
; ‘ 
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, 
<collectingMunicipality id="6B8DC648342FCF5D3932FDF743A5FDE5" box="[113,253,617,636]" pageId="9" pageNumber="530">Amazonas State</collectingMunicipality>
, 
<collectorName id="26A339E4342FCF5D3847FDF742B2FDE5" box="[260,490,617,636]" pageId="9" pageNumber="530">Barcelos Municipality, Mr.</collectorName>
Miranda’s Site, 
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, puddle on the trail, 
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, 
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, 
<collectorName id="26A339E4342FCF5D383EFD1A42A0FD0E" box="[381,504,644,663]" pageId="9" pageNumber="530">Santos A.P.M.</collectorName>
&amp; 
<collectorName id="26A339E4342FCF5D3B50FD1A41CCFD0E" box="[531,660,644,663]" pageId="9" pageNumber="530">Nessimian J.L.</collectorName>
leg.’ (1 J / 
<collectionCode id="ED47C4F7342FCF5D3BBEFD1A4069FD0E" box="[765,817,644,663]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="530">INPA</collectionCode>
)
</materialsCitation>
; ‘ 
<materialsCitation id="3B3E566F342FCF5D3A00FD1A4159FD55" collectingDate="2009-08-23" collectionCode="DZRJ" collectorName="Point B" country="Brazil" county="Araca Sierra" latitude="0.86606115" location="Brazil" longLatPrecision="1" longitude="-63.46722" municipality="Barcelos Municipality" pageId="9" pageNumber="530" specimenCode="DZRJ 6064" specimenCount="5" specimenCount-female="5" typeStatus="paratype">
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, 
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, 
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, 
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, 
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02, 
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<geoCoordinate id="EE623AF5342FCF5D3B3CFD00405FFD28" box="[639,775,670,689]" degrees="63" direction="west" minutes="28" orientation="longitude" pageId="9" pageNumber="530" precision="1" seconds="01,99" value="-63.46722">63°28’01,99”W</geoCoordinate>
, puddles with foliage, 
<date id="FFE87AF2342FCF5D3A8FFD004765FD28" box="[972,1085,670,689]" pageId="9" pageNumber="530" value="2009-08-23">
<collectingDate id="EFAC831A342FCF5D3A8FFD004765FD28" box="[972,1085,670,689]" pageId="9" pageNumber="530" value="2009-08-23">23.VIII.2009</collectingDate>
</date>
, Ferreira-Jr. N. leg.’ (2 JJ 
<specimenCount id="9D5097BB342FCF5D3816FD2742DCFD52" box="[341,388,690,716]" pageId="9" pageNumber="530" type="female">5 ♀♀</specimenCount>
/ 
<specimenCode id="DBF0F449342FCF5D38D6FD2742A3FD55" box="[405,507,697,716]" collectionCode="DZRJ" pageId="9" pageNumber="530">DZRJ 6064</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C0FB9342FCF5E3932FD7B4207FAE6" lastPageId="10" lastPageNumber="531" pageId="9" pageNumber="530" type="diagnosis">
<paragraph id="8BE95C32342FCF5D3932FD7B4719FCE5" blockId="9.[113,1089,741,1564]" pageId="9" pageNumber="530">
<emphasis id="B9228020342FCF5D3932FD7B43BEFD65" bold="true" box="[113,230,741,764]" pageId="9" pageNumber="530">Diagnosis.</emphasis>
<taxonomicName id="4C5627B1342FCF5D39A8FD7B429CFD65" authority="Guimarães &amp; Ferreira-Jr, 2017" authorityName="Guimarães &amp; Ferreira-Jr" authorityYear="2017" box="[235,452,741,764]" class="Insecta" family="Noteridae" genus="Canthysellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="yawari" status="sp. nov.">
<emphasis id="B9228020342FCF5D39A8FD7B429CFD65" box="[235,452,741,764]" italics="true" pageId="9" pageNumber="530">Canthysellus yawari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2113D5B342FCF5D388AFD7B4143FD65" box="[457,539,741,764]" pageId="9" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from other species in the genus by the following characteristics: small size, 
<quantity id="4CAEF1D7342FCF5D3B6CFC9B419FFC85" box="[559,711,773,796]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.65" metricValueMin="2.45" pageId="9" pageNumber="530" unit="mm" value="2.55" valueMax="2.65" valueMin="2.45">2.45–2.65 mm</quantity>
; pronotum and elytra evenly covered with scattered punctures (
<figureCitation id="136D40B7342FCF5D38C5FCBB4291FCA5" box="[390,457,805,828]" captionStart="Figs 1–4" captionStartId="2.[151,188,982,1001]" captionTargetBox="[155,1114,233,950]" captionTargetId="figure-10@2.[151,1126,229,953]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Canthysellus species in lateral and dorsal view: 1 – C. kukrutkato sp. nov.; 2 – C. putkarot sp. nov.; 3 – C. omawe sp. nov.; 4 – C. yawari sp. nov. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315621" httpUri="https://zenodo.org/record/5315621/files/figure.png" pageId="9" pageNumber="530">Fig. 4</figureCitation>
); prosternal process glabrous; median lobe of aedeagus, in lateral aspect, long and slender with a small dorsal projection just before the apex (
<figureCitation id="136D40B7342FCF5D3A92FCDB476CFCC5" box="[977,1076,837,860]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="9" pageNumber="530">Figs 8a,c</figureCitation>
). 
<emphasis id="B9228020342FCF5D3932FCFB4221FCE5" bold="true" box="[113,377,869,892]" pageId="9" pageNumber="530">
Description of 
<typeStatus id="54EDE290342FCF5D3851FCFB422CFCE5" box="[274,372,869,892]" pageId="9" pageNumber="530" type="holotype">holotype</typeStatus>
.
</emphasis>
Body outline, in dorsal view oval (
<figureCitation id="136D40B7342FCF5D3B9EFCFB4043FCE5" box="[733,795,869,892]" captionStart="Figs 1–4" captionStartId="2.[151,188,982,1001]" captionTargetBox="[155,1114,233,950]" captionTargetId="figure-10@2.[151,1126,229,953]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Canthysellus species in lateral and dorsal view: 1 – C. kukrutkato sp. nov.; 2 – C. putkarot sp. nov.; 3 – C. omawe sp. nov.; 4 – C. yawari sp. nov. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315621" httpUri="https://zenodo.org/record/5315621/files/figure.png" pageId="9" pageNumber="530">Fig. 4</figureCitation>
), as in 
<taxonomicName id="4C5627B1342FCF5D3A22FCFB40B5FCE5" authority="Guimarães &amp; Ferreira-Jr, 2017" authorityName="Guimarães &amp; Ferreira-Jr" authorityYear="2017" box="[865,1005,869,892]" class="Insecta" family="Noteridae" genus="Canthysellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="kukrutkato" status="sp. nov.">
<emphasis id="B9228020342FCF5D3A22FCFB40B5FCE5" box="[865,1005,869,892]" italics="true" pageId="9" pageNumber="530">C. kukrutkato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2113D5B342FCF5D3AB1FCFB4719FCE5" box="[1010,1089,869,892]" pageId="9" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BE95C32342FCF5D39D3FC1B42E1FB85" blockId="9.[113,1089,741,1564]" pageId="9" pageNumber="530">
<emphasis id="B9228020342FCF5D39D3FC1B4396FC05" box="[144,206,901,924]" italics="true" pageId="9" pageNumber="530">Color</emphasis>
(
<figureCitation id="136D40B7342FCF5D3999FC1B4242FC05" box="[218,282,901,924]" captionStart="Figs 1–4" captionStartId="2.[151,188,982,1001]" captionTargetBox="[155,1114,233,950]" captionTargetId="figure-10@2.[151,1126,229,953]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Canthysellus species in lateral and dorsal view: 1 – C. kukrutkato sp. nov.; 2 – C. putkarot sp. nov.; 3 – C. omawe sp. nov.; 4 – C. yawari sp. nov. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315621" httpUri="https://zenodo.org/record/5315621/files/figure.png" pageId="9" pageNumber="530">Fig. 4</figureCitation>
). Head and pronotum light-yellowish brown. Elytra dark brown, each elytron with four pale yellowish-brown maculae: one laterally, a second on disc, both just anterior to midlength of elytron, and a third near suture, these three markings appearing as an arched and interrupted transverse band; finally, a fourth short transverse macula on distal third. Ventral surface light yellowish-brown.
</paragraph>
<paragraph id="8BE95C32342FCF5D39D3FBBB439CFBC5" blockId="9.[113,1089,741,1564]" pageId="9" pageNumber="530">
<emphasis id="B9228020342FCF5D39D3FBBB4389FBA5" box="[144,209,1061,1084]" italics="true" pageId="9" pageNumber="530">Head.</emphasis>
Surface smooth and shiny. Antennae and palpi similar to those in 
<taxonomicName id="4C5627B1342FCF5D3AEEFBBB4718FBA5" authority="Guimarães &amp; Ferreira-Jr, 2017" authorityName="Guimarães &amp; Ferreira-Jr" authorityYear="2017" box="[941,1088,1061,1084]" class="Insecta" family="Noteridae" genus="Canthysellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="kukrutkato" status="sp. nov.">
<emphasis id="B9228020342FCF5D3AEEFBBB4718FBA5" box="[941,1088,1061,1084]" italics="true" pageId="9" pageNumber="530">C. kukrutkato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2113D5B342FCF5D3932FBDB439CFBC5" box="[113,196,1093,1116]" pageId="9" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BE95C32342FCF5D39D3FBFB4089FA05" blockId="9.[113,1089,741,1564]" pageId="9" pageNumber="530">
<emphasis id="B9228020342FCF5D39D3FBFB43BCFBE5" box="[144,228,1125,1148]" italics="true" pageId="9" pageNumber="530">Thorax.</emphasis>
Pronotum shiny, evenly covered with scattered punctures and with transversal row of punctures bearing long and slender setae in anterior margin. Prosternum with dense, transverse tuft of five stiff setae at middle of discal surface; line of setae continuous medially, not extending beyond the lateral margins of prosternal process. Prosternal process glabrous on disc, with lateral submargins and apex sparsely covered by short and stout setae, setae distinctly spaced and evenly distributed (
<figureCitation id="136D40B7342FCF5D3B68FA9B4124FA85" box="[555,636,1285,1308]" captionStart="Figs 9–10" captionStartId="8.[151,188,900,919]" captionTargetBox="[202,1078,222,882]" captionTargetId="figure-259@8.[199,1081,219,886]" captionTargetPageId="8" captionText="Figs 9–10. Prosternum, meso- and metaventrum and noterid platform (metacoxae) of Canthysellus, depicting distribution of setae. 9 – C. buqueti (Laporte, 1835); 10 – C. omawe sp. nov. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5315627" httpUri="https://zenodo.org/record/5315627/files/figure.png" pageId="9" pageNumber="530">Fig. 10</figureCitation>
). Noterid platform completely covered by short and stout setae, distinctly spaced and evenly distributed. Elytron shiny, evenly covered with scattered punctures and with three longitudinal rows of punctures bearing long and slender setae (
<figureCitation id="136D40B7342FCF5D39F4FAFB43A2FAE5" box="[183,250,1381,1404]" captionStart="Figs 1–4" captionStartId="2.[151,188,982,1001]" captionTargetBox="[155,1114,233,950]" captionTargetId="figure-10@2.[151,1126,229,953]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Canthysellus species in lateral and dorsal view: 1 – C. kukrutkato sp. nov.; 2 – C. putkarot sp. nov.; 3 – C. omawe sp. nov.; 4 – C. yawari sp. nov. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315621" httpUri="https://zenodo.org/record/5315621/files/figure.png" pageId="9" pageNumber="530">Fig. 4</figureCitation>
). Metatibia with series of ten evenly spaced spiniform setae at posteroventral margin. Metatarsomere I with similar row of five setae on posteroventral margin.
</paragraph>
<paragraph id="8BE95C32342FCF5D39D3FA3B4194FA25" blockId="9.[113,1089,741,1564]" box="[144,716,1445,1468]" pageId="9" pageNumber="530">
<emphasis id="B9228020342FCF5D39D3FA3B43A4FA25" box="[144,252,1445,1468]" italics="true" pageId="9" pageNumber="530">Abdomen.</emphasis>
Ventral surface as in 
<taxonomicName id="4C5627B1342FCF5D38A2FA3B412AFA25" authority="Guimarães &amp; Ferreira-Jr, 2017" authorityName="Guimarães &amp; Ferreira-Jr" authorityYear="2017" box="[481,626,1445,1468]" class="Insecta" family="Noteridae" genus="Canthysellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="kukrutkato" status="sp. nov.">
<emphasis id="B9228020342FCF5D38A2FA3B412AFA25" box="[481,626,1445,1468]" italics="true" pageId="9" pageNumber="530">C. kukrutkato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2113D5B342FCF5D3B3AFA3B4194FA25" box="[633,716,1445,1468]" pageId="9" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BE95C32342FCF5E39D3FA5B4207FAE6" blockId="9.[113,1089,741,1564]" lastBlockId="10.[151,1127,1288,1567]" lastPageId="10" lastPageNumber="531" pageId="9" pageNumber="530">
<emphasis id="B9228020342FCF5D39D3FA5B43A7FA45" box="[144,255,1477,1500]" italics="true" pageId="9" pageNumber="530">Aedeagus.</emphasis>
Median lobe in lateral aspect long and slender, with ventral margin curved; small dorsal projection just before the apex and apex attenuate (
<figureCitation id="136D40B7342FCF5D3A4EFA7B4028FA65" box="[781,880,1509,1532]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="9" pageNumber="530">Figs 8a,c</figureCitation>
); left side with long carina arising basally and ceasing apically, constituting the major part of left side (
<figureCitation id="136D40B7342FCF5D3A89F99B476BF985" box="[970,1075,1541,1564]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="9" pageNumber="530">Figs 8a–c</figureCitation>
); in dorsal aspect, straight with apex slightly curved (
<figureCitation id="136D40B7342CCF5E3B99FA964074FA86" box="[730,812,1288,1311]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="10" pageNumber="531">Fig. 8b</figureCitation>
). Left lateral lobe, in lateral aspect, elongate and very slender with a dense tuft of setae at apex; setae distinctly extending beyond the dorsal margin (
<figureCitation id="136D40B7342CCF5E38F7FAD6415BFAC6" box="[436,515,1352,1375]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="10" pageNumber="531">Fig. 8d</figureCitation>
). Right lateral lobe, in lateral aspect, slender and rounded ventrally (
<figureCitation id="136D40B7342CCF5E3846FAF6420AFAE6" box="[261,338,1384,1407]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="10" pageNumber="531">Fig. 8e</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF290CBA342CCF5E39D4FB0241FFFB36" ID-DOI="http://doi.org/10.5281/zenodo.5315631" ID-Zenodo-Dep="5315631" box="[151,679,1180,1199]" httpUri="https://zenodo.org/record/5315631/files/figure.png" pageId="10" pageNumber="531" startId="10.[151,182,1180,1199]" targetBox="[217,1066,219,1151]" targetPageId="10">
<paragraph id="8BE95C32342CCF5E39D4FB0241FFFB36" blockId="10.[151,679,1180,1199]" box="[151,679,1180,1199]" pageId="10" pageNumber="531">
Fig. 11. Geographic distribution map of 
<taxonomicName id="4C5627B1342CCF5E38ACFB0241FFFB36" box="[495,679,1180,1199]" class="Insecta" family="Noteridae" genus="Canthysellus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9228020342CCF5E38ACFB024105FB36" box="[495,605,1180,1199]" italics="true" pageId="10" pageNumber="531">Canthysellus</emphasis>
species.
</taxonomicName>
</paragraph>
</caption>
<subSubSection id="C34C0FB9342CCF5F39D4FA16416EFECA" lastPageId="11" lastPageNumber="532" pageId="10" pageNumber="531" type="description">
<paragraph id="8BE95C32342CCF5E39D4FA1640BFF986" blockId="10.[151,1127,1288,1567]" pageId="10" pageNumber="531">
<emphasis id="B9228020342CCF5E39D4FA164252FA06" bold="true" box="[151,266,1416,1439]" pageId="10" pageNumber="531">Variation.</emphasis>
Almost no variation was observed in the prominence of the maculae. Some specimens had a darker coloration than that of the 
<typeStatus id="54EDE290342CCF5E3BC8FA3641E1FA26" box="[651,697,1448,1471]" pageId="10" pageNumber="531">type</typeStatus>
specimens. The number of stiff setae on the prosternal disc ranges from three to five in total. The setation on the lateral submargins of prosternal process is greatly reduced on many specimens. The number of setae on metatibia and metatarsomere I ranges from eight to twelve and three to four, respectively.
</paragraph>
<paragraph id="8BE95C32342DCF5F39D3FF42416EFECA" blockId="11.[113,1089,220,499]" pageId="11" pageNumber="532">
<emphasis id="B9228020342DCF5F39D3FF42426AFF6A" box="[144,306,220,243]" italics="true" pageId="11" pageNumber="532">Measurements.</emphasis>
<typeStatus id="54EDE290342DCF5F387BFF4242C7FF6A" box="[312,415,220,243]" pageId="11" pageNumber="532" type="holotype">Holotype</typeStatus>
: TL = 
<quantity id="4CAEF1D7342DCF5F38A4FF424113FF6A" box="[487,587,220,243]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="11" pageNumber="532" unit="mm" value="2.45">2.45 mm</quantity>
; MW = 
<quantity id="4CAEF1D7342DCF5F3BE7FF42405FFF6A" box="[676,775,220,243]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="532" unit="mm" value="1.3">1.30 mm</quantity>
; HW = 
<quantity id="4CAEF1D7342DCF5F3A1EFF424098FF6A" box="[861,960,220,243]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="532" unit="mm" value="0.7">0.70 mm</quantity>
; EW = 
<quantity id="4CAEF1D7342DCF5F3D51FF4243C6FE8A" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="532" unit="mm" value="0.45">0.45 mm</quantity>
; PntB = 
<quantity id="4CAEF1D7342DCF5F39B9FF624204FE8A" box="[250,348,252,275]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="532" unit="mm" value="0.05">0.05 mm</quantity>
; AntVII = 
<quantity id="4CAEF1D7342DCF5F388DFF624168FE8A" box="[462,560,252,275]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="532" unit="mm" value="0.05">0.05 mm</quantity>
. 
<typeStatus id="54EDE290342DCF5F3B79FF6241F1FE8A" box="[570,681,252,275]" pageId="11" pageNumber="532" type="paratype">Paratypes</typeStatus>
:TL = 
<quantity id="4CAEF1D7342DCF5F3BA9FF6240DDFE8A" box="[746,901,252,275]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.65" metricValueMin="2.45" pageId="11" pageNumber="532" unit="mm" value="2.55" valueMax="2.65" valueMin="2.45">2.45–2.65 mm</quantity>
(JJ = 
<quantity id="4CAEF1D7342DCF5F3A95FF6243C6FEAA" metricMagnitude="-3" metricUnit="m" metricValue="2.525" metricValueMax="2.6" metricValueMin="2.45" pageId="11" pageNumber="532" unit="mm" value="2.525" valueMax="2.6" valueMin="2.45">2.45–2.60 mm</quantity>
; 
<specimenCount id="9D5097BB342DCF5F39E9FE8B4388FEAD" box="[170,208,277,308]" pageId="11" pageNumber="532" type="female">♀♀</specimenCount>
= 
<quantity id="4CAEF1D7342DCF5F39AEFE8242D6FEAA" box="[237,398,284,307]" metricMagnitude="-3" metricUnit="m" metricValue="2.625" metricValueMax="2.65" metricValueMin="2.6" pageId="11" pageNumber="532" unit="mm" value="2.625" valueMax="2.65" valueMin="2.6">2.60–2.65 mm</quantity>
); MW = 
<quantity id="4CAEF1D7342DCF5F38ACFE8241D7FEAA" box="[495,655,284,307]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="11" pageNumber="532" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.30–1.50 mm</quantity>
; HW = 
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; EW = 
<quantity id="4CAEF1D7342DCF5F3A95FE8243C6FECA" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="11" pageNumber="532" unit="mm" value="0.475" valueMax="0.5" valueMin="0.45">0.45–0.50 mm</quantity>
; PntB = 
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; AntVII = 
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.
</paragraph>
</subSubSection>
<subSubSection id="C34C0FB9342DCF5F3932FEC241EDFE6A" pageId="11" pageNumber="532" type="etymology">
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<emphasis id="B9228020342DCF5F3932FEC243AAFEEA" bold="true" box="[113,242,348,371]" pageId="11" pageNumber="532">Etymology.</emphasis>
The specific name refers to the Yanomâmi’s forest spirit ‘Yawari’, in honor of the indigenous people present in the area of origin of the 
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. It is a noun in the nominative singular standing in apposition.
</paragraph>
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<emphasis id="B9228020342DCF5F3932FE2242D3FE4A" bold="true" box="[113,395,444,467]" pageId="11" pageNumber="532">Geographic distribution.</emphasis>
So far 
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<emphasis id="B9228020342DCF5F389AFE2241EBFE4A" box="[473,691,444,467]" italics="true" pageId="11" pageNumber="532">Canthysellus yawari</emphasis>
</taxonomicName>
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is known only from the Barcelos Municipality in the Brazilian state of 
<collectingRegion id="499292D0342DCF5F3B7EFE4241E9FE6A" box="[573,689,476,499]" country="Venezuela" name="Amazonas" pageId="11" pageNumber="532">Amazonas</collectingRegion>
.
</paragraph>
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<emphasis id="B9228020342DCF5F38DBFDBD4042FDA4" bold="true" box="[408,794,546,573]" pageId="11" pageNumber="532">
Key to species of 
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<emphasis id="B9228020342DCF5F3B3DFDBC4042FDA4" bold="true" box="[638,794,546,573]" italics="true" pageId="11" pageNumber="532">Canthysellus</emphasis>
</taxonomicName>
</emphasis>
(modified from 
<bibRefCitation id="EFC721C3342DCF5F3B75FDDB4046FDC5" author="BACA S. M. &amp; TOLEDO M." box="[566,798,581,604]" pageId="11" pageNumber="532" pagination="477 - 488" refId="ref7229" refString="BACA S. M. &amp; TOLEDO M. 2015: Canthysellus Baca and Toledo (Coleoptera: Noteridae: Noterini), a new genus of burrowing water beetle from South America. Coleopterists Bulletin 69: 477 - 488." type="journal article" year="2015">BACA &amp; TOLEDO 2015</bibRefCitation>
)
</paragraph>
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1 Pronotum and elytra smooth (
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); prosternal process completely covered by short and stout setae (
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). ................................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BE95C32342DCF5F3932FD2A4718FC92" blockId="11.[113,1089,628,1547]" pageId="11" pageNumber="532">
<keyLead id="30A7FF07342DCF5F3932FD2A4718FC92" pageId="11" pageNumber="532">
– Pronotum and elytra evenly covered with scattered punctures (
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); prosternal process glabrous on disc, with lateral sides and apex sparsely covered by short and stout setae, distinctly spaced and evenly distributed (
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). ................................................ 6
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2 Metatibia and metatarsomere I with inner margin densely setose (
<figureCitation id="136D40B7342DCF5F3A04FC8A40DEFCB2" box="[839,902,788,811]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="11" pageNumber="532">Fig. 7</figureCitation>
of BACA &amp; TOLE- DO 2015); setae slender, hair-like; aedeagus as in 
<figureCitation id="136D40B7342DCF5F3BE7FCAA4041FCD2" box="[676,793,820,843]" captionStart="Figs 9–10" captionStartId="8.[151,188,900,919]" captionTargetBox="[202,1078,222,882]" captionTargetId="figure-259@8.[199,1081,219,886]" captionTargetPageId="8" captionText="Figs 9–10. Prosternum, meso- and metaventrum and noterid platform (metacoxae) of Canthysellus, depicting distribution of setae. 9 – C. buqueti (Laporte, 1835); 10 – C. omawe sp. nov. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5315627" httpUri="https://zenodo.org/record/5315627/files/figure.png" pageId="11" pageNumber="532">Figs 10a–e</figureCitation>
of 
<bibRefCitation id="EFC721C3342DCF5F3A7CFCAA4765FCD2" author="BACA S. M. &amp; TOLEDO M." box="[831,1085,820,843]" pageId="11" pageNumber="532" pagination="477 - 488" refId="ref7229" refString="BACA S. M. &amp; TOLEDO M. 2015: Canthysellus Baca and Toledo (Coleoptera: Noteridae: Noterini), a new genus of burrowing water beetle from South America. Coleopterists Bulletin 69: 477 - 488." type="journal article" year="2015">BACA &amp; TOLEDO (2015)</bibRefCitation>
. ..................................................................................... 
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<emphasis id="B9228020342DCF5F3B8CFCCD400EFCF3" bold="true" box="[719,854,851,875]" italics="true" pageId="11" pageNumber="532">C. peruanus</emphasis>
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<keyLead id="30A7FF07342DCF5F3932FCEA4718FC32" pageId="11" pageNumber="532">
<emphasis id="B9228020342DCF5F3932FCEA4326FC12" bold="true" box="[113,126,884,907]" pageId="11" pageNumber="532">–</emphasis>
Metatibia and metatarsomere I with inner margin with single line of approximately 4–12 evenly spaced stiff setae (
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of 
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). ............................................ 3
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<keyLead id="30A7FF07342DCF5F3932FC2A4718FB92" pageId="11" pageNumber="532">
3 Prosternum with dense, transverse tuft of stiff setae at middle of discal surface, discontinuous medially, extending beyond the lateral margins of prosternal process (
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of 
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). .................................................................................................... 4
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<emphasis id="B9228020342DCF5F3932FB8A4326FBB2" bold="true" box="[113,126,1044,1067]" pageId="11" pageNumber="532">–</emphasis>
Prosternum with dense, transverse tuft of stiff setae at middle of discal surface, continuous medially, not extending beyond the lateral margins of prosternal process (
<figureCitation id="136D40B7342DCF5F3AE7FBAA40BBFBD2" box="[932,995,1076,1099]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="11" pageNumber="532">Fig. 5</figureCitation>
of 
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). .............................................................................................................. 5
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4 Larger size, 
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; base of head usually strongly infuscate, nearly black (
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of 
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); aedeagus as in 
<figureCitation id="136D40B7342DCF5F3B13FB0A41E0FB32" box="[592,696,1172,1195]" captionStart="Figs 9–10" captionStartId="8.[151,188,900,919]" captionTargetBox="[202,1078,222,882]" captionTargetId="figure-259@8.[199,1081,219,886]" captionTargetPageId="8" captionText="Figs 9–10. Prosternum, meso- and metaventrum and noterid platform (metacoxae) of Canthysellus, depicting distribution of setae. 9 – C. buqueti (Laporte, 1835); 10 – C. omawe sp. nov. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5315627" httpUri="https://zenodo.org/record/5315627/files/figure.png" pageId="11" pageNumber="532">Figs 9a–e</figureCitation>
of 
<bibRefCitation id="EFC721C3342DCF5F3B9EFB0A4090FB32" author="BACA S. M. &amp; TOLEDO M." box="[733,968,1172,1195]" pageId="11" pageNumber="532" pagination="477 - 488" refId="ref7229" refString="BACA S. M. &amp; TOLEDO M. 2015: Canthysellus Baca and Toledo (Coleoptera: Noteridae: Noterini), a new genus of burrowing water beetle from South America. Coleopterists Bulletin 69: 477 - 488." type="journal article" year="2015">BACA &amp; TOLEDO 2015</bibRefCitation>
, left lateral lobe very broad with setae extending just to or only slightly beyond the lobe margin (
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of 
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); median lobe with apex elongate in lateral aspect (
<figureCitation id="136D40B7342DCF5F3A9CFB4A4718FB72" box="[991,1088,1236,1259]" captionStart="Figs 9–10" captionStartId="8.[151,188,900,919]" captionTargetBox="[202,1078,222,882]" captionTargetId="figure-259@8.[199,1081,219,886]" captionTargetPageId="8" captionText="Figs 9–10. Prosternum, meso- and metaventrum and noterid platform (metacoxae) of Canthysellus, depicting distribution of setae. 9 – C. buqueti (Laporte, 1835); 10 – C. omawe sp. nov. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5315627" httpUri="https://zenodo.org/record/5315627/files/figure.png" pageId="11" pageNumber="532">Figs 9a,c</figureCitation>
of 
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). .......................................... 
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<emphasis id="B9228020342DCF5F3B8FFB6D400EFA92" bold="true" box="[716,854,1267,1291]" italics="true" pageId="11" pageNumber="532">C. sipaliwini</emphasis>
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– Smaller size, 
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; base of head usually only weakly infuscate (
<figureCitation id="136D40B7342DCF5F3AE7FA8A40BBFAB2" box="[932,995,1300,1323]" captionStart="Figs 1–4" captionStartId="2.[151,188,982,1001]" captionTargetBox="[155,1114,233,950]" captionTargetId="figure-10@2.[151,1126,229,953]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Canthysellus species in lateral and dorsal view: 1 – C. kukrutkato sp. nov.; 2 – C. putkarot sp. nov.; 3 – C. omawe sp. nov.; 4 – C. yawari sp. nov. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315621" httpUri="https://zenodo.org/record/5315621/files/figure.png" pageId="11" pageNumber="532">Fig. 1</figureCitation>
of 
<bibRefCitation id="EFC721C3342DCF5F3D44FA8A4264FAD2" author="BACA S. M. &amp; TOLEDO M." pageId="11" pageNumber="532" pagination="477 - 488" refId="ref7229" refString="BACA S. M. &amp; TOLEDO M. 2015: Canthysellus Baca and Toledo (Coleoptera: Noteridae: Noterini), a new genus of burrowing water beetle from South America. Coleopterists Bulletin 69: 477 - 488." type="journal article" year="2015">BACA &amp; TOLEDO 2015</bibRefCitation>
); aedeagus as in 
<figureCitation id="136D40B7342DCF5F38B1FAAA4101FAD2" box="[498,601,1332,1355]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="11" pageNumber="532">Figs 8a–e</figureCitation>
of 
<bibRefCitation id="EFC721C3342DCF5F3B3DFAAA4031FAD2" author="BACA S. M. &amp; TOLEDO M." box="[638,873,1332,1355]" pageId="11" pageNumber="532" pagination="477 - 488" refId="ref7229" refString="BACA S. M. &amp; TOLEDO M. 2015: Canthysellus Baca and Toledo (Coleoptera: Noteridae: Noterini), a new genus of burrowing water beetle from South America. Coleopterists Bulletin 69: 477 - 488." type="journal article" year="2015">BACA &amp; TOLEDO 2015</bibRefCitation>
, left lobe with setae extending far beyond lobe margin (
<figureCitation id="136D40B7342DCF5F3B4DFACA4104FAF2" box="[526,604,1364,1387]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="11" pageNumber="532">Fig. 8d</figureCitation>
of 
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); median lobe with apex pointed, but not elongate in lateral aspect (
<figureCitation id="136D40B7342DCF5F3BCDFAEA41A8FA12" box="[654,752,1396,1419]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="11" pageNumber="532">Figs 8a,c</figureCitation>
of 
<bibRefCitation id="EFC721C3342DCF5F3A50FAEA40A6FA12" author="BACA S. M. &amp; TOLEDO M." box="[787,1022,1396,1419]" pageId="11" pageNumber="532" pagination="477 - 488" refId="ref7229" refString="BACA S. M. &amp; TOLEDO M. 2015: Canthysellus Baca and Toledo (Coleoptera: Noteridae: Noterini), a new genus of burrowing water beetle from South America. Coleopterists Bulletin 69: 477 - 488." type="journal article" year="2015">BACA &amp; TOLEDO 2015</bibRefCitation>
). ....... ................................................................................................. 
<taxonomicName id="4C5627B1342DCF5F3A63FA0D4718FA32" authority="(Laporte, 1835)" baseAuthorityName="Laporte" baseAuthorityYear="1835" box="[800,1088,1427,1451]" class="Insecta" family="Noteridae" genus="Noterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="buqueti">
<emphasis id="B9228020342DCF5F3A63FA0D40CAFA32" bold="true" box="[800,914,1427,1451]" italics="true" pageId="11" pageNumber="532">C. buqueti</emphasis>
(Laporte, 1835)
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</keyStep>
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5 Large size, 
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; median lobe, in lateral aspect, elongate and thin with dorsobasal margin sharply narrowing before midlength of median lobe (
<figureCitation id="136D40B7342DCF5F3A78FA4A40C7FA72" box="[827,927,1492,1515]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="11" pageNumber="532">Figs 6a,c</figureCitation>
). ..................... ............................................................................................................ 
<emphasis id="B9228020342DCF5F3A2BFA6D4718F992" bold="true" box="[872,1088,1523,1547]" pageId="11" pageNumber="532">
<taxonomicName id="4C5627B1342DCF5F3A2BFA6D40BCF992" authority="Guimarães &amp; Ferreira-Jr, 2017" authorityName="Guimarães &amp; Ferreira-Jr" authorityYear="2017" box="[872,996,1523,1547]" class="Insecta" family="Noteridae" genus="Canthysellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="putkarot" status="sp. nov.">
<emphasis id="B9228020342DCF5F3A2BFA6D40BCF992" bold="true" box="[872,996,1523,1547]" italics="true" pageId="11" pageNumber="532">C. putkarot</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2113D5B342DCF5F3AA8FA6A4718F992" box="[1003,1088,1524,1547]" pageId="11" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE95C32342ACF5839D4FF42473EFEAA" blockId="12.[151,1127,220,595]" pageId="12" pageNumber="533">
<keyLead id="30A7FF07342ACF5839D4FF42473EFEAA" pageId="12" pageNumber="533">
<emphasis id="B9228020342ACF5839D4FF4243FCFF6A" bold="true" box="[151,164,220,243]" pageId="12" pageNumber="533">–</emphasis>
Medium to large size, 
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; median lobe, in lateral aspect, short and stout with dorsobasal margin oblique and distinctly projected, sharply narrowing at midlength of median lobe (
<figureCitation id="136D40B7342ACF583804FE8242F3FEAA" box="[327,427,284,307]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="12" pageNumber="533">Figs 5a,c</figureCitation>
). ................................................................ 
<emphasis id="B9228020342ACF583A32FE85473EFEAA" bold="true" box="[881,1126,283,307]" pageId="12" pageNumber="533">
<taxonomicName id="4C5627B1342ACF583A32FE854753FEAA" authority="Guimarães &amp; Ferreira-Jr, 2017" authorityName="Guimarães &amp; Ferreira-Jr" authorityYear="2017" box="[881,1035,283,307]" class="Insecta" family="Noteridae" genus="Canthysellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="533" phylum="Arthropoda" rank="species" species="kukrutkato" status="sp. nov.">
<emphasis id="B9228020342ACF583A32FE854753FEAA" bold="true" box="[881,1035,283,307]" italics="true" pageId="12" pageNumber="533">C. kukrutkato</emphasis>
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</emphasis>
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</keyStep>
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6 Larger, 
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; prosternum with dense, transverse tuft of stiff setae at middle of discal surface, discontinuous medially, extending beyond the lateral margins of prosternal process (
<figureCitation id="136D40B7342ACF583855FEE242DBFE0A" box="[278,387,380,403]" captionStart="Figs 1–4" captionStartId="2.[151,188,982,1001]" captionTargetBox="[155,1114,233,950]" captionTargetId="figure-10@2.[151,1126,229,953]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Canthysellus species in lateral and dorsal view: 1 – C. kukrutkato sp. nov.; 2 – C. putkarot sp. nov.; 3 – C. omawe sp. nov.; 4 – C. yawari sp. nov. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315621" httpUri="https://zenodo.org/record/5315621/files/figure.png" pageId="12" pageNumber="533">Figs 4a–b</figureCitation>
of 
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); median lobe, in lateral aspect, long and robust with a large dorsal projection just before apex; apex rounded and globose (
<figureCitation id="136D40B7342ACF583D74FE0243BCFE4A" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="12" pageNumber="533">Figs 7a,c</figureCitation>
). .................................................................................................... 
<emphasis id="B9228020342ACF583ADDFE25473EFE4A" bold="true" box="[926,1126,443,467]" pageId="12" pageNumber="533">
<taxonomicName id="4C5627B1342ACF583ADDFE254752FE4A" authority="Guimarães &amp; Ferreira-Jr, 2017" authorityName="Guimarães &amp; Ferreira-Jr" authorityYear="2017" box="[926,1034,443,467]" class="Insecta" family="Noteridae" genus="Canthysellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="533" phylum="Arthropoda" rank="species" species="omawe" status="sp. nov.">
<emphasis id="B9228020342ACF583ADDFE254752FE4A" bold="true" box="[926,1034,443,467]" italics="true" pageId="12" pageNumber="533">C. omawe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2113D5B342ACF583D52FE22473EFE4A" box="[1041,1126,444,467]" pageId="12" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE95C32342ACF5839D4FE42473EFDCA" blockId="12.[151,1127,220,595]" pageId="12" pageNumber="533">
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– Smaller, 
<quantity id="4CAEF1D7342ACF583857FE4242E8FE6A" box="[276,432,476,499]" metricMagnitude="-3" metricUnit="m" metricValue="2.525" metricValueMax="2.6" metricValueMin="2.45" pageId="12" pageNumber="533" unit="mm" value="2.525" valueMax="2.6" valueMin="2.45">2.45–2.60 mm</quantity>
; prosternum with dense, transverse tuft of stiff setae at middle of discal surface, continuous medially, not extending beyond the lateral margins of prosternal process (
<figureCitation id="136D40B7342ACF583850FD82420AFDAA" box="[275,338,540,563]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="12" pageNumber="533">Fig. 5</figureCitation>
of 
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); median lobe, in lateral aspect, long and slender with a small dorsal projection just before apex (
<figureCitation id="136D40B7342ACF583BE6FDA2405EFDCA" box="[677,774,572,595]" captionStart="Figs 5–8" captionStartId="4.[151,188,1505,1524]" captionTargetBox="[204,1072,228,1471]" captionTargetId="figure-10@4.[195,1082,219,1480]" captionTargetPageId="4" captionText="Figs 5–8.Aedeagi of Canthysellus: a – median lobe, left lateral aspect, b – same, dorsal aspect, c – same, right lateral aspect, d – left lateral lobe (paramere), e – right lateral lobe. 5 – C. kukrutkato sp. nov.; 6 – C. putkarot sp. nov.; 7 – C. omawe sp. nov.; 8 – C. yawari sp. nov. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5315625" httpUri="https://zenodo.org/record/5315625/files/figure.png" pageId="12" pageNumber="533">Figs 8a,c</figureCitation>
). .................... 
<emphasis id="B9228020342ACF583AE6FDA5473EFDCA" bold="true" box="[933,1126,571,595]" pageId="12" pageNumber="533">
<taxonomicName id="4C5627B1342ACF583AE6FDA54755FDCA" authority="Guimarães &amp; Ferreira-Jr, 2017" authorityName="Guimarães &amp; Ferreira-Jr" authorityYear="2017" box="[933,1037,571,595]" class="Insecta" family="Noteridae" genus="Canthysellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="533" phylum="Arthropoda" rank="species" species="yawari" status="sp. nov.">
<emphasis id="B9228020342ACF583AE6FDA54755FDCA" bold="true" box="[933,1037,571,595]" italics="true" pageId="12" pageNumber="533">C. yawari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2113D5B342ACF583D50FDA2473EFDCA" box="[1043,1126,572,595]" pageId="12" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
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