treatments-xml/data/03/FF/87/03FF87E6CA053B14DE89064AFBA7FD67.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.3789506" ID-GBIF-Dataset="de55da44-3fa9-4b4a-80da-41641abbde8e" ID-GBIF-Taxon="163812768" ID-ISSN="1313–2970" ID-Zenodo-Dep="3789506" ID-ZooBank="964540F2-C9F8-413B-B1C2-56A8A2761B82" LSID="urn:lsid:zoobank.org:act:A0FF698D-9560-4401-BDC9-13529E6460A8" approvalRequired="8" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="5" checkinTime="1588014885007" checkinUser="plazi" docAuthor="Broad, Gavin, Sääksjärvi, Ilari &amp; Palacio, Edgard" docDate="2010" docId="03FF87E6CA053B14DE89064AFBA7FD67" docLanguage="en" docName="ZK_article_2152.pdf" docOrigin="ZooKeys 35 (35)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@7ea42c82" docStyleName="zookeys.2008.journal_article" docTitle="Umanella giacometti Broad &amp; Saaksjarviq 2010, sp. n." docType="treatment" docUuid="A0FF698D-9560-4401-BDC9-13529E6460A8" docUuidSource="ZooBank" docVersion="6" lastPageId="13" lastPageNumber="90" masterDocId="FFC6FF9ECA003B19DE050406FF86FF93" masterDocTitle="A new species of Umanella Gauld (Hymenoptera, Ichneumonidae) from Ecuador" masterLastPageNumber="90" masterPageNumber="77" pageId="5" pageNumber="82" updateTime="1643563056878" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title>A new species of Umanella Gauld (Hymenoptera, Ichneumonidae) from Ecuador</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Broad, Gavin</mods:namePart>
<mods:affiliation>The Natural History Museum, London, United Kingdom</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sääksjärvi, Ilari</mods:namePart>
<mods:affiliation>University of Turku, Turku ,, Finland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Palacio, Edgard</mods:namePart>
<mods:affiliation>, Bogota, Colombia</mods:affiliation>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2010</mods:date>
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<mods:number>2010-02-02</mods:number>
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<mods:number>35</mods:number>
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<mods:number>35</mods:number>
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<mods:identifier type="DOI">10.3897/zookeys.35.326</mods:identifier>
<mods:identifier type="GBIF-Dataset">de55da44-3fa9-4b4a-80da-41641abbde8e</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789506" ID-GBIF-Taxon="163812768" ID-Zenodo-Dep="3789506" LSID="urn:lsid:zoobank.org:act:A0FF698D-9560-4401-BDC9-13529E6460A8" httpUri="http://treatment.plazi.org/id/03FF87E6CA053B14DE89064AFBA7FD67" lastPageId="13" lastPageNumber="90" pageId="5" pageNumber="82">
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<heading box="[140,706,588,615]" fontSize="11" level="2" pageId="5" pageNumber="82" reason="3">
<emphasis bold="true" box="[140,706,588,615]" pageId="5" pageNumber="82">
<taxonomicName authority="Broad &amp; Saaksjarvi" authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[140,629,588,615]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="giacometti" status="sp. n.">
<emphasis bold="true" box="[140,387,588,615]" italics="true" pageId="5" pageNumber="82">Umanella giacometti</emphasis>
Broad &amp; Sääksjärvi
</taxonomicName>
, 
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</emphasis>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act: 
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<subSubSection lastPageId="9" lastPageNumber="86" pageId="5" pageNumber="82" type="description">
<paragraph blockId="5.[140,1108,694,1672]" pageId="5" pageNumber="82">
<emphasis bold="true" box="[140,292,694,720]" pageId="5" pageNumber="82">
<emphasis box="[140,287,694,720]" inLineHeading="true" pageId="5" pageNumber="82" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[303,389,695,721]" italics="true" pageId="5" pageNumber="82">Female.</emphasis>
Whole insect: Fig. 1. 'Fore wing length 18.5 (
<typeStatus box="[952,1056,695,722]" pageId="5" pageNumber="82">holotype</typeStatus>
)– 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="82" unit="mm" value="20.0">20 mm</quantity>
, body length (from antennal insertion to apex of metasoma) 
<quantity box="[919,1048,730,756]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="5" pageNumber="82" unit="mm" value="32.5" valueMax="34.0" valueMin="31.0">31-34 mm</quantity>
, ovipositor length 41 (
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)– 
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, ratio of ovipositor to fore wing length 2.22– 2.25. Occipital carina complete, joining hypostomal carina behind base of mandible, dipped mediodorsally with depression on vertex and deep groove on back of head. Epomia present for short distance across trough in pronotum. Notauli deep and long, converging but not meeting posteriorly, petering out at about half the length of mesoscutum. Epicnemial carina absent (
<figureCitation box="[604,675,941,967]" captionStart="Figure 2" captionStartId="6.[140,212,1160,1182]" captionTargetBox="[140,1108,566,1140]" captionTargetId="figure@6.[140,1108,566,1141]" captionTargetPageId="6" captionText="Figure 2. U. giacometti, female holotype, detail of head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.3770855" httpUri="https://zenodo.org/record/3770855/files/figure.png" pageId="5" pageNumber="82">Fig. 2</figureCitation>
). Mesopleurum with posterior suture weakly impressed dorsally, strong ventrally, smooth except for some weak crenulae, and deeply impressed furrow connnecting suture to episternal scrobe. Mesepisternal sulcus complete, strong and non-crentulate. Posterior transverse carina of mesosternum absent. Propodeum lacking all carinae except pleural and submetapleural carinae, which are complete, and stubs of median longitudinal carinae, present at posterior end of propodeum. Faint, narrow groove present on dorsal, ventral half of propodeum. Propodeal spiracle about twice as long as medially wide. Whole body elongate and integument entirely smooth and shining except mandible sparsely punctate basally, puncto-striate medially. Upper tooth of mandible slightly shorter than lower tooth. Hind coxa about 1.2 as long as dorsal face of propodeum. First tergite of metasoma 4.4 × as long as apically wide, second tergite 2.8 × as long as apically wide. First tergite with swollen, apical area, posterior end impressed behind it. Sclerotized part of first sternite with low, raised bump just before level of spiracle; extends to half distance between spiracle and posterior end of tergite. Second tergite with deep, diagonal grooves cutting off anterior corners; narrow, drop-shaped median area defined, sides of raised area concave (
<figureCitation box="[481,574,1504,1530]" captionStart="Figure 3" captionStartId="7.[140,212,1447,1469]" captionTargetBox="[199,1049,165,1427]" captionTargetId="figure@7.[199,1050,162,1427]" captionTargetPageId="7" captionText="Figure 3. Second tergite of female metasoma of a U. caerulea from Costa Rica b U. caerulea from Colombia c U. giacometti; anterior to the right." figureDoi="http://doi.org/10.5281/zenodo.3770857" httpUri="https://zenodo.org/record/3770857/files/figure.png" pageId="5" pageNumber="82">Fig. 3C</figureCitation>
). Th ird and fourth tergites with deep, lateral grooves. Metasoma with strong setae laterally. All tarsal claws with acute basal lobe. Fore tibia with longitudinal patch of stronger, darker setae. Propodeum with coxal foramen narrowly separated from metasomal foramen by sclerotized bridge. Ovipositor with tip angled downwards. Lower valve of ovipositor apically slightly overlapping
</paragraph>
<paragraph blockId="6.[140,777,497,519]" box="[140,777,497,519]" pageId="6" pageNumber="83">
<emphasis bold="true" box="[140,238,497,519]" pageId="6" pageNumber="83">Figure |.</emphasis>
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[244,428,497,519]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[244,428,497,519]" italics="true" pageId="6" pageNumber="83">Umanella giacometti</emphasis>
</taxonomicName>
, female 
<typeStatus box="[507,591,497,519]" pageId="6" pageNumber="83">holotype</typeStatus>
; scale bar = 
<quantity box="[702,773,497,518]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="83" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770855" ID-Zenodo-Dep="3770855" box="[140,810,1160,1182]" httpUri="https://zenodo.org/record/3770855/files/figure.png" pageId="6" pageNumber="83" startId="6.[140,212,1160,1182]" targetBox="[140,1108,566,1140]" targetPageId="6">
<paragraph blockId="6.[140,810,1160,1182]" box="[140,810,1160,1182]" pageId="6" pageNumber="83">
<emphasis bold="true" box="[140,238,1160,1182]" pageId="6" pageNumber="83">Figure 2.</emphasis>
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[244,363,1161,1182]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[244,363,1161,1182]" italics="true" pageId="6" pageNumber="83">U. giacometti</emphasis>
</taxonomicName>
, female holotype, detail of head and mesosoma.
</paragraph>
</caption>
<paragraph blockId="6.[140,1108,1223,1672]" pageId="6" pageNumber="83">dorsal valve, with 13 visible teeth, regularly spaced and inclined. Dorsal valve with row of lateral denticles above teeth on each side.</paragraph>
<paragraph blockId="6.[140,1108,1223,1672]" lastBlockId="7.[140,1107,1539,1672]" lastPageId="7" lastPageNumber="84" pageId="6" pageNumber="83">
Colour: metallic blue, duller towards apex of metasoma. Antennae black, except for white, sub-apical annulus on three and a half flagellomeres (23rd to 26th) to five (
<typeStatus box="[148,252,1363,1389]" pageId="6" pageNumber="83">holotype</typeStatus>
, 23rd to 27th) flagellomeres. Maxillary palps dark brown, a little paler centrally, ventrally. Tegula metallic blue. Wing venation, including stigma, dark brown. Wing membrane slightly infuscate basally (
<typeStatus box="[642,745,1434,1461]" pageId="6" pageNumber="83">holotype</typeStatus>
) or distinctly brown, distinctly infuscate in apical quarter. Legs with coxae, trochanters and trochantelli shiny, metallic (dark) blue. Fore leg with apical edge of trochanter creamy white with white streak extending over basal half of anterior surface of fore femur, extreme apex of trochantellus and base of femur brown; fore tibia with basal 0.6 brown (slightly paler sub-basally); fore tarsus brown except apical half of third and fourth tarsomeres creamy, fifth tarsomere black. Mid tibia black with small, dull creamy mark sub-basally and abruptly white on apical quarter (black at extreme apex) (
<figureCitation box="[689,781,1539,1566]" captionStart="Figure 4" captionStartId="8.[140,212,623,645]" captionTargetBox="[140,1108,165,603]" captionTargetId="figure@8.[137,1111,165,603]" captionTargetPageId="8" captionText="Figure 4. Mid tibia of a female of U. caerulea from Costa Rica b female of U. caerulea from Colombia (BMNH) c female of U. caerulea from Colombia (IAVH) d female of U. giacometti." figureDoi="http://doi.org/10.5281/zenodo.3770859" httpUri="https://zenodo.org/record/3770859/files/figure.png" pageId="7" pageNumber="84">Fig. 4D</figureCitation>
); mid tarsus brown to black, except fourth and apical half of third tarsomere creamy. Hind tibia with white annulus sub-basally; hind tarsus black on first, basal half of second and apical half of fifth tarsomeres, remainder creamy.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770857" ID-Zenodo-Dep="3770857" httpUri="https://zenodo.org/record/3770857/files/figure.png" pageId="7" pageNumber="84" startId="7.[140,212,1447,1469]" targetBox="[199,1049,165,1427]" targetPageId="7">
<paragraph blockId="7.[140,1107,1447,1501]" pageId="7" pageNumber="84">
<emphasis bold="true" box="[140,239,1447,1469]" pageId="7" pageNumber="84">Figure 3.</emphasis>
Second tergite of female metasoma of 
<emphasis bold="true" box="[602,615,1447,1469]" pageId="7" pageNumber="84">a</emphasis>
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[621,723,1447,1469]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[621,723,1447,1469]" italics="true" pageId="7" pageNumber="84">U. caerulea</emphasis>
</taxonomicName>
from Costa Rica 
<emphasis bold="true" box="[890,904,1447,1469]" pageId="7" pageNumber="84">b</emphasis>
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[910,1013,1447,1469]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[910,1013,1447,1469]" italics="true" pageId="7" pageNumber="84">U. caerulea</emphasis>
</taxonomicName>
from Colombia 
<emphasis bold="true" box="[210,222,1479,1501]" pageId="7" pageNumber="84">c</emphasis>
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[228,348,1480,1501]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[228,348,1480,1501]" italics="true" pageId="7" pageNumber="84">U. giacometti</emphasis>
</taxonomicName>
; anterior to the right.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770859" ID-Zenodo-Dep="3770859" httpUri="https://zenodo.org/record/3770859/files/figure.png" pageId="8" pageNumber="85" startId="8.[140,212,623,645]" targetBox="[140,1108,165,603]" targetPageId="8">
<paragraph blockId="8.[140,1108,623,677]" pageId="8" pageNumber="85">
<emphasis bold="true" box="[140,239,623,645]" pageId="8" pageNumber="85">Figure 4.</emphasis>
Mid tibia of 
<emphasis bold="true" box="[364,377,623,645]" pageId="8" pageNumber="85">a</emphasis>
female of 
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[475,577,623,645]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[475,577,623,645]" italics="true" pageId="8" pageNumber="85">U. caerulea</emphasis>
</taxonomicName>
from Costa Rica 
<emphasis bold="true" box="[742,756,623,645]" pageId="8" pageNumber="85">b</emphasis>
female of 
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[854,956,623,645]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[854,956,623,645]" italics="true" pageId="8" pageNumber="85">U. caerulea</emphasis>
</taxonomicName>
from Colombia (BMNH) 
<emphasis bold="true" box="[236,248,655,677]" pageId="8" pageNumber="85">c</emphasis>
female of 
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[345,446,655,677]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[345,446,655,677]" italics="true" pageId="8" pageNumber="85">U. caerulea</emphasis>
</taxonomicName>
from Colombia (IAVH) 
<emphasis bold="true" box="[681,695,655,677]" pageId="8" pageNumber="85">d</emphasis>
female of 
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[792,912,655,677]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[792,912,655,677]" italics="true" pageId="8" pageNumber="85">U. giacometti</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="8.[140,1108,730,1672]" pageId="8" pageNumber="85">
<emphasis box="[185,352,730,756]" italics="true" pageId="8" pageNumber="85">
<typeStatus box="[185,277,731,756]" pageId="8" pageNumber="85">Paratype</typeStatus>
males.
</emphasis>
Whole insect: 
<figureCitation box="[521,589,730,756]" captionStart="Figure 8" captionStartId="11.[140,212,1654,1676]" captionTargetBox="[225,1022,1163,1634]" captionTargetId="figure@11.[225,1023,1163,1634]" captionTargetPageId="11" captionText="Figure 8. Paratype male of U. giacometti (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.3770867" httpUri="https://zenodo.org/record/3770867/files/figure.png" pageId="8" pageNumber="85">Figs 8</figureCitation>
, 
<figureCitation box="[603,620,730,756]" captionStart="Figure 9" captionStartId="12.[140,212,838,860]" captionTargetBox="[178,1069,165,816]" captionTargetId="figure@12.[178,1070,165,817]" captionTargetPageId="12" captionText="Figure 9. Paratype male of U. giacometti (ZMUT/USNM)." figureDoi="http://doi.org/10.5281/zenodo.3770869" httpUri="https://zenodo.org/record/3770869/files/figure.png" pageId="8" pageNumber="85">9</figureCitation>
. Fore wing length 
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, body length (from antennal insertion to apex of genitalia) 
<quantity box="[651,781,765,792]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="8" pageNumber="85" unit="mm" value="18.0" valueMax="21.0" valueMin="15.0">15–21 mm</quantity>
. Submetapleural carina complete to about two thirds length of propodeum then abruptly weaker (barely traceable on smaller specimen) for remainder. First metasomal tergite 3.7 to 4.0 × as long as maximum width (
<figureCitation box="[350,439,870,897]" captionStart="Figure 5" captionStartId="9.[140,212,1307,1329]" captionTargetBox="[190,1058,165,1287]" captionTargetId="figure@9.[190,1058,165,1287]" captionTargetPageId="9" captionText="Figure 5. First tergite of male metasoma of a U. caerulea (BMNH) b U. caerulea (IAVH) c U. giacometti; anterior to the right." figureDoi="http://doi.org/10.5281/zenodo.3770861" httpUri="https://zenodo.org/record/3770861/files/figure.png" pageId="8" pageNumber="85">Fig. 5B</figureCitation>
), second tergite 2.7 to 3.1 × as long as wide. Second and third tergites with strong diagonal, basal grooves cutting off raised, central section which is raised posteriorly (Fig. 10). First and second tergites with strong setae laterally but smooth dorsally. Sclerotized part of first tergite extending to 0.5 of distance between spiracle and hind edge. Third tergite regularly punctate and setose dorsally. Fourth tergite onwards setose dorsally with inconspicuous punctures.
</paragraph>
<paragraph blockId="8.[140,1108,730,1672]" lastBlockId="9.[140,1107,1434,1672]" lastPageId="9" lastPageNumber="86" pageId="8" pageNumber="85">Colour: head and mesosoma metallic blue. Antennae dark brown to black, white on 3–11 (22nd to 32nd) flagellomeres. Maxillary and labial palps cream coloured. Tegula white-transparent. Wing venation, including stigma, dark brown. Wing membrane clear basally, infuscate in apical quarter. Fore leg white on fore side, except apical tarsomere black or dark brown, other tarsomeres off white, basal half of first tarsomere pale brown to white. Hind side of fore leg basally white, but coxa with slight brown hint, femur with broad ventral streak on apical 0.7, light or dark brown fading into metallic blue; fore tibia brown on basal half, apex narrowly dark brown. Mid leg with coxa and trochanter white, trochantellus dark brown to black, fore side of femur dark brown to black on basal 0.15, remainder white; mid tibia almost totally white or dark brown on basal half, except for slightly paler subbasal patch, narrowly black at apex; mid tarsus dark brown to black on first and fifth tarsomeres, second and third dark brown fading to off white, fourth white. Hind leg with metallic blue coxa, shiny black or dark brown trochanter, trochantellus and femur; inner apex of hind trochantellus off-white, base of femur narrowly whitish to brown; hind tibia black or brown with narrow white annulus sub-basally, but uniformly black or brown dorsally; hind tarsus dark brown to black on first, basal 0.7 of second and apical 0.2–0.5 of fifth tarsomeres, remainder white. Metasoma with first to fifth or sixth tergites metallic blue, fading to dark brown or near black, apical tergites dark brown to shiny black; first sternite with sclerotized part metallic blue, sclerotized sections of other sternites shiny black to brown, membranous parts of sternites white.</paragraph>
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<paragraph blockId="9.[140,1107,1307,1361]" pageId="9" pageNumber="86">
<emphasis bold="true" box="[140,237,1307,1329]" pageId="9" pageNumber="86">Figure 5.</emphasis>
First tergite of male metasoma of 
<emphasis bold="true" box="[546,559,1307,1329]" pageId="9" pageNumber="86">a</emphasis>
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[563,664,1307,1328]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[563,664,1307,1328]" italics="true" pageId="9" pageNumber="86">
U. 
<taxonomicName box="[592,664,1307,1328]" pageId="9" pageNumber="86" rank="species" species="caerulea">caerulea</taxonomicName>
</emphasis>
</taxonomicName>
(BMNH) 
<emphasis bold="true" box="[764,778,1307,1329]" pageId="9" pageNumber="86">b</emphasis>
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[783,883,1307,1328]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[783,883,1307,1328]" italics="true" pageId="9" pageNumber="86">
U. 
<taxonomicName box="[811,883,1307,1328]" pageId="9" pageNumber="86" rank="species" species="caerulea">caerulea</taxonomicName>
</emphasis>
</taxonomicName>
(IAVH) 
<emphasis bold="true" box="[965,977,1307,1329]" pageId="9" pageNumber="86">c</emphasis>
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[982,1101,1307,1328]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[982,1101,1307,1328]" italics="true" pageId="9" pageNumber="86">
U. 
<taxonomicName box="[1011,1101,1307,1328]" pageId="9" pageNumber="86" rank="species" species="giacometti">giacometti</taxonomicName>
</emphasis>
</taxonomicName>
; anterior to the right.
</paragraph>
</caption>
<paragraph blockId="9.[140,1107,1434,1672]" pageId="9" pageNumber="86">
<emphasis bold="true" box="[185,304,1609,1635]" pageId="9" pageNumber="86">
<emphasis box="[185,298,1609,1635]" inLineHeading="true" pageId="9" pageNumber="86" reason="1">Variation</emphasis>
.
</emphasis>
The male in USNM/ZMUT is smaller than the two males in BMNH and is more extensively white on the antennal flagellomeres and mid tibia.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770863" ID-Zenodo-Dep="3770863" box="[140,511,955,977]" httpUri="https://zenodo.org/record/3770863/files/figure.png" pageId="10" pageNumber="87" startId="10.[140,212,955,977]" targetBox="[140,1108,165,935]" targetPageId="10">
<paragraph blockId="10.[140,511,955,977]" box="[140,511,955,977]" pageId="10" pageNumber="87">
<emphasis bold="true" box="[140,238,955,977]" pageId="10" pageNumber="87">Figure 6.</emphasis>
Paratype male of 
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[404,505,955,977]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[404,505,955,977]" italics="true" pageId="10" pageNumber="87">U. caerulea</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection pageId="10" pageNumber="87" type="biology_ecology">
<paragraph blockId="10.[140,1108,1046,1672]" pageId="10" pageNumber="87">
<emphasis bold="true" box="[185,280,1046,1072]" pageId="10" pageNumber="87">
<emphasis box="[185,275,1046,1072]" inLineHeading="true" pageId="10" pageNumber="87" reason="1">Biology</emphasis>
.
</emphasis>
No specimens of 
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[490,594,1047,1073]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis box="[490,594,1047,1073]" italics="true" pageId="10" pageNumber="87">Umanella</emphasis>
</taxonomicName>
have been reared but the 
<typeStatus box="[895,991,1047,1073]" pageId="10" pageNumber="87">holotype</typeStatus>
and three 
<typeStatus box="[140,243,1082,1108]" pageId="10" pageNumber="87">paratypes</typeStatus>
(BMNH) were collected whilst flying around a standing, dead tree trunk. The hosts may be large coleopteran larvae. Th e ovipositor shape is rather similar to that of 
<taxonomicName authorityName="Smith" authorityYear="1877" box="[140,280,1152,1178]" class="Insecta" family="Ichneumonidae" genus="Dolichomitus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis box="[140,280,1152,1178]" italics="true" pageId="10" pageNumber="87">Dolichomitus</emphasis>
</taxonomicName>
species, which are parasitoids of beetle larvae in dead wood. Three specimens have been found at fairly high (
<quantity box="[478,567,1188,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="87" unit="m" value="1100.0">1,100m</quantity>
) altitude whilst one male was collected by canopy fogging at fairly low (
<quantity box="[372,447,1223,1249]" metricMagnitude="2" metricUnit="m" metricValue="2.16" pageId="10" pageNumber="87" unit="m" value="216.0">216 m</quantity>
) altitude. In this respect the new species resembles 
<taxonomicName authorityName="Gauld" authorityYear="1991" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis italics="true" pageId="10" pageNumber="87">Umanella caerulea</emphasis>
</taxonomicName>
, which also inhabits tropical forests of various altitudes (
<bibRefCitation author="Gauld ID &amp; Ugalde Gomez JA &amp; Hanson P" box="[842,1046,1258,1284]" pageId="10" pageNumber="87" pagination="1 - 189" refId="ref4786" refString="Gauld ID, Ugalde Gomez JA, Hanson P (1998) Guia de los Pimplinae de Costa Rica. Revista de Biologia Tropical 46 Supplement 1: 1 - 189." type="journal article" year="1998">Gauld et al., 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[185,666,1293,1319]" pageId="10" pageNumber="87" type="distribution">
<paragraph blockId="10.[140,1108,1046,1672]" box="[185,666,1293,1319]" pageId="10" pageNumber="87">
<emphasis bold="true" box="[185,344,1293,1319]" pageId="10" pageNumber="87">
<emphasis box="[185,339,1293,1319]" inLineHeading="true" pageId="10" pageNumber="87" reason="1">Distribution</emphasis>
.
</emphasis>
Only known from 
<collectingCountry box="[566,663,1293,1319]" name="Ecuador" pageId="10" pageNumber="87">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="87" type="materials_examined">
<paragraph blockId="10.[140,1108,1046,1672]" pageId="10" pageNumber="87">
<emphasis bold="true" box="[185,422,1328,1354]" pageId="10" pageNumber="87">
<emphasis box="[185,286,1328,1354]" inLineHeading="true" pageId="10" pageNumber="87" reason="1">Material</emphasis>
examined.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2609008350" collectingDate="1981-05-30" collectionCode="BMNH" collectorName="M. Cooper'" country="Ecuador" county="Macas" elevation="1100" location="Santiago" municipality="Cord. de Cutucu" pageId="10" pageNumber="87" specimenCount="1" typeStatus="holotype">
<typeStatus box="[435,542,1328,1354]" pageId="10" pageNumber="87">Holotype</typeStatus>
female: ‘ 
<collectingCountry box="[654,753,1328,1354]" name="Ecuador" pageId="10" pageNumber="87">Ecuador</collectingCountry>
, Morona- 
<location LSID="urn:lsid:plazi:treatment:03FF87E6CA053B14DE89064AFBA7FD67:8E89602BCA0A3B13DD710136FC5EFAD9" box="[884,984,1328,1354]" country="Ecuador" county="Macas" municipality="Cord. de Cutucu" name="Santiago" pageId="10" pageNumber="87">Santiago</location>
, 
<collectingMunicipality pageId="10" pageNumber="87">Cord. de Cutucu</collectingMunicipality>
<quantity box="[236,294,1363,1389]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="87" unit="km" value="6.0">6km</quantity>
.e. of 
<collectingCounty box="[358,429,1364,1390]" pageId="10" pageNumber="87">Macas</collectingCounty>
<quantity box="[439,529,1364,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="87" unit="m" value="1100.0">
<elevation box="[439,529,1364,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="87" unit="m" value="1100.0">1,100m</elevation>
</quantity>
<date box="[539,658,1364,1390]" pageId="10" pageNumber="87" value="1981-05-30">
<collectingDate box="[539,658,1364,1390]" pageId="10" pageNumber="87" value="1981-05-30">30.V.1981</collectingDate>
</date>
<collectorName box="[668,804,1364,1390]" pageId="10" pageNumber="87">M. Cooper’</collectorName>
(
<collectionCode box="[823,920,1364,1390]" pageId="10" pageNumber="87">BMNH</collectionCode>
)
</materialsCitation>
. 
<typeStatus box="[942,1052,1364,1390]" pageId="10" pageNumber="87">Paratypes</typeStatus>
: one female, same data as 
<typeStatus box="[390,489,1399,1425]" pageId="10" pageNumber="87">holotype</typeStatus>
except 
<date box="[582,717,1399,1425]" pageId="10" pageNumber="87" value="1981-06-26">26.VI.1981</date>
, one male with same data except 
<date box="[140,244,1434,1460]" pageId="10" pageNumber="87" value="1981-05-04">4.V.1981</date>
(
<collectionCode box="[263,359,1434,1460]" pageId="10" pageNumber="87">BMNH</collectionCode>
), one male with same data except 
<date box="[768,903,1434,1460]" pageId="10" pageNumber="87" value="1981-06-26">26.VI.1981</date>
, one male, 
<collectingCountry name="Ecuador" pageId="10" pageNumber="87">Ecuador</collectingCountry>
, 
<collectingRegion box="[194,295,1469,1495]" country="Ecuador" name="Orellana" pageId="10" pageNumber="87">Orellana</collectingRegion>
, Reserva Etnica Waorani, Onkone Gare Camp, 
<quantity box="[852,954,1469,1496]" metricMagnitude="2" metricUnit="m" metricValue="2.163" pageId="10" pageNumber="87" unit="m" value="216.3">216.3 m</quantity>
, 
<date box="[968,1103,1469,1495]" pageId="10" pageNumber="87" value="1994-06-21">21.VI.1994</date>
, 
<geoCoordinate box="[140,293,1504,1531]" degrees="00" direction="south" minutes="39" orientation="latitude" pageId="10" pageNumber="87" precision="1" seconds="25.7" value="-0.6571389">00°39'25.7&quot;S</geoCoordinate>
, 
<geoCoordinate box="[304,481,1504,1530]" degrees="076" direction="west" minutes="27" orientation="longitude" pageId="10" pageNumber="87" precision="1" seconds="10.8" value="-76.452995">076°27'10.8&quot;W</geoCoordinate>
, fogging, terra firme forest, T.L.Erwin et al. leg. (held in trust for 
<collectingCountry box="[239,333,1539,1565]" name="Ecuador" pageId="10" pageNumber="87">Ecuador</collectingCountry>
at 
<collectionCode box="[369,468,1540,1566]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="87" type="Museum">NMNH</collectionCode>
, but currently on loan to 
<collectionCode box="[760,852,1540,1566]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="10" pageNumber="87">ZMUT</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="90" pageId="10" pageNumber="87" type="etymology">
<paragraph blockId="10.[140,1108,1046,1672]" pageId="10" pageNumber="87">
<emphasis bold="true" box="[185,320,1575,1601]" pageId="10" pageNumber="87">
<emphasis box="[185,315,1575,1601]" inLineHeading="true" pageId="10" pageNumber="87" reason="1">Etymology</emphasis>
.
</emphasis>
We are pleased to give the name suggested by Mrs Jean Halperin, who won a competition to name this beautiful species in celebration of the opening of the Natural History Museum’s Darwin Centre Two, where the 
<typeStatus box="[830,929,1645,1671]" pageId="10" pageNumber="87">holotype</typeStatus>
is housed. The
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770865" ID-Zenodo-Dep="3770865" box="[140,613,1107,1130]" httpUri="https://zenodo.org/record/3770865/files/figure.png" pageId="11" pageNumber="88" startId="11.[140,212,1107,1129]" targetBox="[226,1021,165,1088]" targetPageId="11">
<paragraph blockId="11.[140,613,1107,1130]" box="[140,613,1107,1130]" pageId="11" pageNumber="88">
<emphasis bold="true" box="[140,238,1107,1129]" pageId="11" pageNumber="88">Figure 7.</emphasis>
Male of 
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[321,423,1108,1129]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[321,423,1108,1129]" italics="true" pageId="11" pageNumber="88">U. caerulea</emphasis>
</taxonomicName>
, Colombia (IAVH).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770867" ID-Zenodo-Dep="3770867" box="[140,626,1654,1677]" httpUri="https://zenodo.org/record/3770867/files/figure.png" pageId="11" pageNumber="88" startId="11.[140,212,1654,1676]" targetBox="[225,1022,1163,1634]" targetPageId="11">
<paragraph blockId="11.[140,626,1654,1677]" box="[140,626,1654,1677]" pageId="11" pageNumber="88">
<emphasis bold="true" box="[140,238,1654,1676]" pageId="11" pageNumber="88">Figure 8.</emphasis>
Paratype male of 
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[404,523,1655,1676]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[404,523,1655,1676]" italics="true" pageId="11" pageNumber="88">U. giacometti</emphasis>
</taxonomicName>
(BMNH).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770869" ID-Zenodo-Dep="3770869" box="[140,701,838,860]" httpUri="https://zenodo.org/record/3770869/files/figure.png" pageId="12" pageNumber="89" startId="12.[140,212,838,860]" targetBox="[178,1069,165,816]" targetPageId="12">
<paragraph blockId="12.[140,701,838,860]" box="[140,701,838,860]" pageId="12" pageNumber="89">
<emphasis bold="true" box="[140,238,838,860]" pageId="12" pageNumber="89">Figure 9.</emphasis>
Paratype male of 
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[404,523,839,860]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="89" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[404,523,839,860]" italics="true" pageId="12" pageNumber="89">U. giacometti</emphasis>
</taxonomicName>
(ZMUT/USNM).
</paragraph>
</caption>
<paragraph blockId="12.[140,944,1654,1677]" box="[140,944,1654,1677]" pageId="12" pageNumber="89">
<emphasis bold="true" box="[140,251,1654,1676]" pageId="12" pageNumber="89">Figure |0.</emphasis>
<typeStatus box="[257,336,1655,1676]" pageId="12" pageNumber="89">Paratype</typeStatus>
male of 
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[417,536,1655,1676]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="89" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[417,536,1655,1676]" italics="true" pageId="12" pageNumber="89">U. giacometti</emphasis>
</taxonomicName>
(ZMUT/USNM), first and second tergites.
</paragraph>
<paragraph blockId="13.[140,1107,167,228]" pageId="13" pageNumber="90">name refers to the wasp’s resemblance to the slender, attenuated figures of the Swiss artist, Alberto Giacometti and is a noun in apposition.</paragraph>
<paragraph blockId="13.[140,393,322,349]" box="[140,393,322,349]" pageId="13" pageNumber="90">
<emphasis bold="true" box="[140,393,322,349]" pageId="13" pageNumber="90">Acknowledgments</emphasis>
</paragraph>
<paragraph blockId="13.[140,1108,378,756]" pageId="13" pageNumber="90">
We are grateful to Harry Taylor, of the Natural History Museum photo unit, for taking superb photographs of the 
<typeStatus box="[447,546,413,439]" pageId="13" pageNumber="90">holotype</typeStatus>
of 
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[585,733,414,439]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[585,733,414,439]" italics="true" pageId="13" pageNumber="90">U. giacometti</emphasis>
</taxonomicName>
. Dr Vladimir Blagoderov helped with other photography and image stacking. We are indebted to Martin Cooper for collecting four of the type series of 
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[546,695,484,509]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[546,695,484,509]" italics="true" pageId="13" pageNumber="90">U. giacometti</emphasis>
</taxonomicName>
, and so many other interesting 
<taxonomicName class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
, and donating them to the Natural History Museum, and to Dr. Terry L. Erwin for providing us with his interesting canopy fogging samples which yielded one male of the type series of 
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[422,568,590,615]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[422,568,590,615]" italics="true" pageId="13" pageNumber="90">U. giacometti</emphasis>
</taxonomicName>
. We are grateful to MSc Anu Veijalainen for her help in sorting the canopy fogging samples. 
<taxonomicName authorityName="Gauld" authorityYear="1991" box="[629,736,624,650]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis box="[629,736,624,650]" italics="true" pageId="13" pageNumber="90">Umanella</emphasis>
</taxonomicName>
specimens in IAVH were collect- ed thanks to the support of NSF grant DEB 0205982 to M.J. Sharkey and B.V. Bryan. Claudia Alejandra Medina from IAVH kindly provided access to specimens under her care. And thank you to Jean Halperin for suggesting the species name, 
<taxonomicName authorityName="Broad &amp; Saaksjarviq" authorityYear="2010" box="[940,1051,731,756]" class="Insecta" family="Ichneumonidae" genus="Umanella" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="giacometti">
<emphasis box="[940,1051,731,756]" italics="true" pageId="13" pageNumber="90">giacometti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>