283 lines
38 KiB
XML
283 lines
38 KiB
XML
<document id="A83C9FEBE8F03E8A8C14DBDEC0F746C4" ID="10.11646/zootaxa.4258.2.5SLASH10907" ID-CLB-Dataset="33155" ID-DOI="10.11646/zootaxa.4258.2.5" ID-GBIF-Dataset="ac8dfde6-c270-402e-835d-cd37633fdfca" ID-ISSN="1175-5326" ID-Zenodo-Dep="569442" ID-ZooBank="312CBC35-CBB7-438C-AF7D-F0B373A460BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493281446952" checkinUser="plazi" docAuthor="Sehnal, Richard" docDate="2017" docId="03FF87E2E11DFFAF18F7F926FF3AFEEE" docLanguage="en" docName="zootaxa.4258.2.5SLASH10907.pdf" docOrigin="Zootaxa 4258 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Entypophana njiapanda Sehnal, new species" docType="treatment" docVersion="7" lastPageNumber="169" masterDocId="FFC6FF9AE114FFA31860FFFFFFE5FF8A" masterDocTitle="Review of the central African leaf chafer genus Entypophana Moser, 1913 (Coleoptera: Scarabaeidae: Melolonthinae: Schizonychini)" masterLastPageNumber="171" masterPageNumber="157" pageNumber="166" updateTime="1698439120373" updateUser="ExternalLinkService">
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<mods:titleInfo id="00B9E71184C0262FB12E12B5AC87D71A">
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<mods:title id="60253568D7BCB93D654BB692471B98D4">Review of the central African leaf chafer genus Entypophana Moser, 1913 (Coleoptera: Scarabaeidae: Melolonthinae: Schizonychini)</mods:title>
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<mods:name id="F6FDF21FB630242D3C8D3CA9B351B372" type="personal">
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<mods:namePart id="21FB16FC3F61610B2ADE06F2C9C6D3F1">Sehnal, Richard</mods:namePart>
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<mods:title id="A6987BC9230E3AD2BCAC809C6D51C92A">Zootaxa</mods:title>
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<mods:part id="E227CCB99DCEA7AB3AC92D27C2E021E9">
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<mods:date id="D97CF5F5FBB3FF487C5E7E83A64CF9E6">2017</mods:date>
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<mods:number id="6816BFE9D7E631A49E48010AD35E146F">4258</mods:number>
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<mods:identifier id="FA1D355D82453D988DF40F475C9465B7" type="DOI">10.11646/zootaxa.4258.2.5</mods:identifier>
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<mods:identifier id="CD6C582CA7BEEA8DEFF95ED354660344" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FF87E2E11DFFAF18F7F926FF3AFEEE" ID-DOI="http://doi.org/10.5281/zenodo.6034962" ID-GBIF-Taxon="128810296" ID-Zenodo-Dep="6034962" LSID="urn:lsid:plazi:treatment:03FF87E2E11DFFAF18F7F926FF3AFEEE" httpUri="http://treatment.plazi.org/id/03FF87E2E11DFFAF18F7F926FF3AFEEE" lastPageId="12" lastPageNumber="169" pageId="9" pageNumber="166">
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<subSubSection id="C34C657FE11DFFAA18F7F926FE28F89F" pageId="9" pageNumber="166" type="nomenclature">
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<paragraph id="8BE936F4E11DFFAA18F7F926FD50F97E" blockId="9.[151,693,1753,1813]" box="[151,693,1753,1780]" pageId="9" pageNumber="166">
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<heading id="D0A18198E11DFFAA18F7F926FD50F97E" bold="true" box="[151,693,1753,1780]" fontSize="11" level="1" pageId="9" pageNumber="166" reason="1">
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<emphasis id="B922EAE6E11DFFAA18F7F926FD50F97E" bold="true" box="[151,693,1753,1780]" pageId="9" pageNumber="166">
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<taxonomicName id="4C564D77E11DFFAA18F7F926FDFCF97E" authority="Sehnal" authorityName="Sehnal" box="[151,537,1753,1780]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="166" phylum="Arthropoda" rank="species" species="njiapanda" status="sp. nov.">
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<emphasis id="B922EAE6E11DFFAA18F7F926FE5EF979" bold="true" box="[151,443,1753,1779]" italics="true" pageId="9" pageNumber="166">Entypophana njiapanda</emphasis>
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Sehnal
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="A211579DE11DFFAA1A46F925FD50F97E" box="[550,693,1754,1780]" pageId="9" pageNumber="166" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE936F4E11DFFAA18F7F903FE28F89F" blockId="9.[151,693,1753,1813]" box="[151,461,1788,1813]" pageId="9" pageNumber="166">
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<figureCitation id="136D2A71E11DFFAA18F7F903FEF3F89F" box="[151,278,1788,1813]" captionStart="FIGURES 6" captionStartId="10.[151,264,1055,1078]" captionTargetBox="[166,1431,203,1025]" captionTargetId="figure@10.[151,1435,193,1034]" captionTargetPageId="10" captionText="FIGURES 6 A – D. Entypophana njiapanda new species male, holotype; length 17.9 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569448/files/figure.png" pageId="9" pageNumber="166">Figs 6A–D</figureCitation>
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,
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<figureCitation id="136D2A71E11DFFAA1940F902FEDAF89E" box="[288,319,1789,1812]" captionStart="FIGURES 7" captionStartId="11.[151,264,1701,1724]" captionTargetBox="[185,1411,848,1668]" captionTargetId="figure@11.[177,1414,841,1674]" captionTargetPageId="11" captionText="FIGURES 7 A – F. Aedeagus in dorsal view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569449/files/figure.png" pageId="9" pageNumber="166">7F</figureCitation>
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,
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<figureCitation id="136D2A71E11DFFAA192AF902FE8DF89F" box="[330,360,1789,1813]" captionStart="FIGURES 8" captionStartId="12.[151,264,1326,1349]" captionTargetBox="[170,1427,419,1302]" captionTargetId="figure@12.[151,1437,411,1305]" captionTargetPageId="12" captionText="FIGURES 8 A – F. Aedeagus in lateral view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569450/files/figure.png" pageId="9" pageNumber="166">8F</figureCitation>
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,
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||
<figureCitation id="136D2A71E11DFFAA1913F902FE77F89F" box="[371,402,1789,1813]" captionStart="FIGURES 9" captionStartId="13.[151,264,925,948]" captionTargetBox="[159,1429,206,892]" captionTargetId="figure@13.[151,1436,195,904]" captionTargetPageId="13" captionText="FIGURES 9 A – F. Inner mesotarsal claw. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569451/files/figure.png" pageId="9" pageNumber="166">9F</figureCitation>
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,
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<specimenCount id="9D50FD7DE11DFFAA19FDF902FE28F89F" box="[413,461,1789,1813]" pageId="9" pageNumber="166" type="generic">
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<figureCitation id="136D2A71E11DFFAA19FDF902FE2CF89F" box="[413,457,1789,1813]" captionStart="FIGURES 10" captionStartId="13.[151,264,1927,1950]" captionTargetBox="[151,1435,1059,1906]" captionTargetId="figure@13.[151,1435,1059,1906]" captionTargetPageId="13" captionText="FIGURES 10 A – F. Map with currently known geographical distribution. A, Entypophana apicata Moser, 1913; B, Entypophana biapicata Moser, 1913; C, Entypophana hulstaerti Burgeon, 1946; D, Entypophana lujai Moser, 1917; E, Entypophana maynei Burgeon, 1946; F, Entypophana njiapanda new species." httpUri="https://zenodo.org/record/569452/files/figure.png" pageId="9" pageNumber="166">10F</figureCitation>
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.
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||
</specimenCount>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34C657FE11DFFA918F7F8BBFE0BFB19" lastPageId="10" lastPageNumber="167" pageId="9" pageNumber="166" type="materials_examined">
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<paragraph id="8BE936F4E11DFFAA18F7F8BBFCF8F8D7" blockId="9.[151,1436,1860,2029]" box="[151,797,1860,1885]" pageId="9" pageNumber="166">
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<materialsCitation id="3B3E3CA9E11DFFAA18F7F8BBFCF8F8D7" ID-GBIF-Occurrence="1499294118" box="[151,797,1860,1885]" country="Tanzania" location="Kilombero District" pageId="9" pageNumber="166" specimenCount="1" typeStatus="holotype">
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<emphasis id="B922EAE6E11DFFAA18F7F8BBFED0F8D7" bold="true" box="[151,309,1860,1885]" pageId="9" pageNumber="166">
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<typeStatus id="54ED8856E11DFFAA18F7F8BBFF37F8D7" box="[151,210,1860,1885]" pageId="9" pageNumber="166">Type</typeStatus>
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locality.
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||
</emphasis>
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||
<collectingCountry id="F3417664E11DFFAA195CF8BBFE41F8D7" box="[316,420,1860,1885]" name="Tanzania" pageId="9" pageNumber="166">Tanzania</collectingCountry>
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||
,
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||
<location id="8E89602FE11DFFAA19CFF8BBFD63F8D7" LSID="urn:lsid:plazi:treatment:03FF87E2E11DFFAF18F7F926FF3AFEEE:8E89602FE11DFFAA19CFF8BBFD63F8D7" box="[431,646,1860,1885]" country="Tanzania" name="Kilombero District" pageId="9" pageNumber="166">Kilombero District</location>
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||
, Mwanihana.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BE936F4E11DFFAA18A7F897FC13F867" blockId="9.[151,1436,1860,2029]" pageId="9" pageNumber="166">
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<emphasis id="B922EAE6E11DFFAA18A7F897FE92F80B" bold="true" box="[199,375,1896,1921]" pageId="9" pageNumber="166">Type material</emphasis>
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||
(2 specimens).
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||
<materialsCitation id="3B3E3CA9E11DFFAA1A24F897FDD4F867" ID-GBIF-Occurrence="1499294124" collectingDate="2010-11-27" collectingDateMax="2092" collectingDateMin="2010-11-27" collectionCode="BMNH" collectorName="Udzungwa Mts. N. P. & Smith, R & Takano, H & Schizonychini" country="Tanzania" county="Udzungwa Mts." determinerName="R. Sehnal" elevation="970" latitude="-7.791111" location="Njia Panda" longLatPrecision="21" longitude="36.82611" municipality="Mwanihana" pageId="9" pageNumber="166" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
|
||
<typeStatus id="54ED8856E11DFFAA1A24F897FD4AF80B" box="[580,687,1896,1921]" pageId="9" pageNumber="166" type="holotype">Holotype</typeStatus>
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||
,
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||
<specimenCount id="9D50FD7DE11DFFAA1AA4F896FD3EF80A" box="[708,731,1897,1920]" pageId="9" pageNumber="166" type="male">♂</specimenCount>
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||
, labelled: “
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||
<collectingCountry id="F3417664E11DFFAA1B0BF896FC1AF80B" box="[875,1023,1897,1921]" name="Tanzania" pageId="9" pageNumber="166">TANZANIA</collectingCountry>
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||
,
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||
<location id="8E89602FE11DFFAA1C73F897FB7DF80B" LSID="urn:lsid:plazi:treatment:03FF87E2E11DFFAF18F7F926FF3AFEEE:8E89602FE11DFFAA1C73F897FB7DF80B" box="[1043,1176,1896,1921]" country="Tanzania" county="Udzungwa Mts." latitude="-7.791111" longLatPrecision="21" longitude="36.82611" municipality="Mwanihana" name="Njia Panda" pageId="9" pageNumber="166">Njia Panda</location>
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||
<elevation id="007BD1C7E11DFFAA1CC8F896FB0DF80B" box="[1192,1256,1897,1921]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="9" pageNumber="166" unit="m" value="970.0">
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<quantity id="4CAE9B11E11DFFAA1CC8F896FB0DF80B" box="[1192,1256,1897,1921]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="9" pageNumber="166" unit="m" value="970.0">970m</quantity>
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||
</elevation>
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||
/
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||
<collectingMunicipality id="6B8DAC8EE11DFFAA1D71F897FA7DF80B" box="[1297,1432,1896,1921]" pageId="9" pageNumber="166">Mwanihana</collectingMunicipality>
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,
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||
<collectorName id="26A35322E11DFFAA18F7F873FE63F82E" box="[151,390,1932,1957]" pageId="9" pageNumber="166">
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||
<collectingCounty id="62884E78E11DFFAA18F7F873FEA8F82E" box="[151,333,1932,1957]" pageId="9" pageNumber="166">Udzungwa Mts.</collectingCounty>
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N.P.
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</collectorName>
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||
/
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||
<geoCoordinate id="EE625033E11DFFAA19C2F873FDC6F82F" box="[418,547,1932,1957]" direction="south" orientation="latitude" pageId="9" pageNumber="166" precision="15" value="-7.791111">S07°47′28″</geoCoordinate>
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<geoCoordinate id="EE625033E11DFFAA1A4CF873FD4BF82F" box="[556,686,1932,1957]" direction="east" orientation="longitude" pageId="9" pageNumber="166" precision="15" value="36.82611">E36°49′34″</geoCoordinate>
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||
/
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||
<collectingDate id="EFACE9DCE11DFFAA1AAAF873FC82F82F" box="[714,871,1932,1957]" pageId="9" pageNumber="166" value="2010-11-27" valueMax="2010-11-30" valueMin="2010-11-27">
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||
<date id="FFE81034E11DFFAA1AAAF873FC82F82F" box="[714,871,1932,1957]" pageId="9" pageNumber="166" value="2010-11-27" valueMax="2010-11-30" valueMin="2010-11-27">27-30.xi.2010</date>
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||
</collectingDate>
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||
<collectingMethod id="52174EE3E11DFFAA1B11F873FC09F82E" box="[881,1004,1932,1957]" pageId="9" pageNumber="166">Light Trap</collectingMethod>
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||
/ leg.
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||
<collectorName id="26A35322E11DFFAA1C5BF873FB46F82E" box="[1083,1187,1932,1957]" pageId="9" pageNumber="166">Smith, R</collectorName>
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||
&
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||
<collectorName id="26A35322E11DFFAA1CACF873FAA6F82E" box="[1228,1347,1932,1957]" pageId="9" pageNumber="166">Takano, H</collectorName>
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||
[white, printed] //
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||
<collectorName id="26A35322E11DFFAA196CF84FFE49F843" box="[268,428,1968,1993]" pageId="9" pageNumber="166">Schizonychini</collectorName>
|
||
/
|
||
<taxonomicName id="4C564D77E11DFFAA19A1F84FFDA8F842" box="[449,589,1968,1992]" class="Insecta" family="Melolonthidae" genus="Schizonycha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="166" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B922EAE6E11DFFAA19A1F84FFDA8F842" box="[449,589,1968,1992]" italics="true" pageId="9" pageNumber="166">Schizonycha</emphasis>
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||
</taxonomicName>
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||
<specimenCount id="9D50FD7DE11DFFAA1A34F84EFD8CF842" box="[596,617,1969,1992]" pageId="9" pageNumber="166" type="male">♂</specimenCount>
|
||
/ (
|
||
<taxonomicName id="4C564D77E11DFFAA1AE7F84FFCF9F842" box="[647,796,1968,1992]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="166" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B922EAE6E11DFFAA1AE7F84FFCF9F842" box="[647,796,1968,1992]" italics="true" pageId="9" pageNumber="166">Entypophana</emphasis>
|
||
</taxonomicName>
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||
) sp. /
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||
<determinerName id="7E35DA30E11DFFAA1B01F84EFC2BF843" box="[865,974,1968,1993]" pageId="9" pageNumber="166">R. Sehnal</determinerName>
|
||
det. 2012 [white, printed] //
|
||
<collectionCode id="ED47AE31E11DFFAA1D6EF84FFA8EF843" box="[1294,1387,1968,1993]" pageId="9" pageNumber="166">BMNH</collectionCode>
|
||
{E} /
|
||
<collectingDate id="EFACE9DCE11DFFAA18F7F82AFF11F867" box="[151,244,2005,2029]" pageId="9" pageNumber="166" value="2012" valueMax="2092">
|
||
<date id="FFE81034E11DFFAA18F7F82AFF11F867" box="[151,244,2005,2029]" pageId="9" pageNumber="166" value="2012" valueMax="2092">2012-92</date>
|
||
</collectingDate>
|
||
/ 1198576 [white, printed]”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3E3CA9E11DFFAA1A5BF82AFC13F867" ID-GBIF-Occurrence="1499294084" box="[571,1014,2004,2029]" collectingDate="2010-11-27" collectingDateMax="2092" collectingDateMin="2010-11-27" collectionCode="BMNH" collectorName="Udzungwa Mts. N. P. & Smith, R & Takano, H & Schizonychini" country="Tanzania" county="Udzungwa Mts." determinerName="R. Sehnal" elevation="970" latitude="-7.791111" location="Njia Panda" longLatPrecision="21" longitude="36.82611" municipality="Mwanihana" pageId="9" pageNumber="166" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<typeStatus id="54ED8856E11DFFAA1A5BF82AFD2CF867" box="[571,713,2005,2029]" pageId="9" pageNumber="166" type="paratype">PARATYPE</typeStatus>
|
||
,
|
||
<specimenCount id="9D50FD7DE11DFFAA1AB4F82AFD0EF866" box="[724,747,2005,2028]" pageId="9" pageNumber="166" type="female">♀</specimenCount>
|
||
: same data as holotype.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF29667CE11EFFA918F7FBE0FC64FBDE" httpUri="https://zenodo.org/record/569448/files/figure.png" pageId="10" pageNumber="167" targetBox="[166,1431,203,1025]" targetPageId="10">
|
||
<paragraph id="8BE936F4E11EFFA918F7FBE0FC64FBDE" blockId="10.[151,1436,1055,1108]" pageId="10" pageNumber="167">
|
||
<emphasis id="B922EAE6E11EFFA918F7FBE0FEB0FBBE" bold="true" box="[151,341,1055,1078]" pageId="10" pageNumber="167">FIGURES 6A–D.</emphasis>
|
||
<taxonomicName id="4C564D77E11EFFA9193DFBDFFDB1FBBF" box="[349,596,1056,1077]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="167" phylum="Arthropoda" rank="species" species="njiapanda" status="sp. nov.">
|
||
<emphasis id="B922EAE6E11EFFA9193DFBDFFDB1FBBF" box="[349,596,1056,1077]" italics="true" pageId="10" pageNumber="167">Entypophana njiapanda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A211579DE11EFFA91A3CFBDFFD3DFBBF" box="[604,728,1055,1077]" pageId="10" pageNumber="167" rank="species">new species</taxonomicNameLabel>
|
||
male, holotype; length 17.9 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE936F4E11EFFA918A7FB84FE0BFB19" blockId="10.[151,1437,1147,2036]" box="[199,494,1147,1172]" pageId="10" pageNumber="167">
|
||
<emphasis id="B922EAE6E11EFFA918A7FB84FE69FB1E" bold="true" box="[199,396,1147,1172]" pageId="10" pageNumber="167">Type depository.</emphasis>
|
||
BMNH.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C657FE11EFFA818A7FB60FC50F80C" lastPageId="11" lastPageNumber="168" pageId="10" pageNumber="167" type="description">
|
||
<paragraph id="8BE936F4E11EFFA918A7FB60FCC2FACD" blockId="10.[151,1437,1147,2036]" pageId="10" pageNumber="167">
|
||
<emphasis id="B922EAE6E11EFFA918A7FB60FDEDFB32" bold="true" box="[199,520,1183,1208]" pageId="10" pageNumber="167">
|
||
Description of the
|
||
<typeStatus id="54ED8856E11EFFA919C1FB60FDEDFB32" box="[417,520,1183,1208]" pageId="10" pageNumber="167" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="9D50FD7DE11EFFA91A78FB5FFDCAFB32" box="[536,559,1184,1208]" pageId="10" pageNumber="167" type="male">♂</specimenCount>
|
||
). Body length
|
||
<quantity id="4CAE9B11E11EFFA91AB3FB5FFCDDFB3D" box="[723,824,1184,1208]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="10" pageNumber="167" unit="mm" value="17.9">17.9 mm</quantity>
|
||
, elongate, almost parallel, strongly convex. Dorsal and ventral surface weakly shiny, head and pronotum dark brown, elytra brown, macrosetation pale (
|
||
<figureCitation id="136D2A71E11EFFA91C81FB3CFAD9FB56" box="[1249,1340,1219,1244]" captionStart="FIGURES 6" captionStartId="10.[151,264,1055,1078]" captionTargetBox="[166,1431,203,1025]" captionTargetId="figure@10.[151,1435,193,1034]" captionTargetPageId="10" captionText="FIGURES 6 A – D. Entypophana njiapanda new species male, holotype; length 17.9 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569448/files/figure.png" pageId="10" pageNumber="167">Fig. 6A</figureCitation>
|
||
). Dorsal surface of head, pronotum, scutellum, and elytra covered with white, short, recumbent, scale-like macrosetae; ventral surface of thorax and abdomen with yellow, recumbent, long macrosetae. Head appendages and legs covered with reddish-yellow, moderately long macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E11EFFA918A7FAACFBCFF972" blockId="10.[151,1437,1147,2036]" pageId="10" pageNumber="167">
|
||
Head with labrum transverse, deeply bilobed; lobes rounded, irregularly and weakly punctate; lobes covered with long, erect macrosetae. Head including clypeus smooth, glossy, fissured, densely coarsely punctate; each puncture with an erect macroseta shorter than puncture diameter. Frontoclypeal carina sinuate in the forward direction; surface behind carina densely but unevenly punctate except a narrow impunctate strip adjacent to frontoclypeal suture. Frontoclypeal suture undulate, medially arched toward occipital carina. Occipital carina always present, prominent, elevated, medially extended into a columnal impunctate summit, laterally diminishing (
|
||
<figureCitation id="136D2A71E11EFFA918FFF9D4FF13F9CE" box="[159,246,1579,1604]" captionStart="FIGURES 6" captionStartId="10.[151,264,1055,1078]" captionTargetBox="[166,1431,203,1025]" captionTargetId="figure@10.[151,1435,193,1034]" captionTargetPageId="10" captionText="FIGURES 6 A – D. Entypophana njiapanda new species male, holotype; length 17.9 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569448/files/figure.png" pageId="10" pageNumber="167">Fig. 6B</figureCitation>
|
||
). Space posterior of frontoclypeal suture with a large, triangular impunctate area. Occiput sparsely, evenly, moderately punctate. Canthus narrow, short, glabrous. Width of both eyes combined approximately equal to maximum width of frons between eyes, eye distinctly extending beyond canthus. Antenna with 10 antennomeres; club with three antennomeres, almost straight, short than antennal shaft (antennomeres 1–7 combined). Antennomeres 1–7 with sparse, long macrosetae; club sparsely, shortly macrosetaceous. Terminal maxillary palpomere expanded apically, slightly shorter than palpomeres 2 and 3 combined.
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E11EFFA918A7F8FBFB3BF87E" blockId="10.[151,1437,1147,2036]" pageId="10" pageNumber="167">
|
||
Pronotum transverse, widest behind midlength; with a large but shallow triangular, anteromedial depression as wide as head, which abruptly rises toward edge of crest. Front margin straight, with angles extended forward. Lateral margins crenulate and macrosetose. Base bordered, medially gently arched toward elytra, with smooth margin paralleled by a row of coarse punctures and hind angles broadly rounded. Crest delimiting anteromedial depression prominent, with a wide strip at edge and summit of crest impunctate. Depression densely covered by deep, asperate punctures with setae no longer than puncture diameters. Punctation of remaining surface also asperate, but sparser and punctures set in rows; setae yellow, scale-like, semirecumbent (
|
||
<figureCitation id="136D2A71E11EFFA91C18F824FB34F87E" box="[1144,1233,2011,2036]" captionStart="FIGURES 6" captionStartId="10.[151,264,1055,1078]" captionTargetBox="[166,1431,203,1025]" captionTargetId="figure@10.[151,1435,193,1034]" captionTargetPageId="10" captionText="FIGURES 6 A – D. Entypophana njiapanda new species male, holotype; length 17.9 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569448/files/figure.png" pageId="10" pageNumber="167">Fig. 6C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E11FFFA818A7FF68FB25FF5E" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="168">Scutellum large, almost equilaterally triangular, sides and apex rounded; sides with horseshoe-shaped punctures; each puncture bearing short, narrow, white, scale-like, almost recumbent macroseta.</paragraph>
|
||
<paragraph id="8BE936F4E11FFFA818A7FF20FDECFE0D" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="168">Elytra weakly convex, parallel-sided, rounded apically; apical angle approximately rectangular. Striae absent, except for feebly visible sutural stria. Humeral umbones present, weakly swollen. Surface not microsculptured, moderately shiny; punctures shallow, regularly spaced, separated by 2–3 diameters. Each puncture bearing shorter than their diameter, narrow, yellow scale-like, almost recumbent macroseta. Epipleuron distinct, complete, narrow, glabrous laterally. Macropterous.</paragraph>
|
||
<paragraph id="8BE936F4E11FFFA818A7FE6AFD7EFDB6" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="168">
|
||
Legs with femora narrow, shiny, irregularly punctate, macrosetaceous. Protibia narrow, distinctly tridentate; terminal spine inserted above medial dent. Protarsomeres 4–5 missing from both legs. Mesotibia and metatibia slightly expanded apicad, with one macrosetiferous, longitudinal carinae. Upper terminal spine of metatibia flattened, slightly curved, acute apically; 1/3 longer than lower, apically trunctate, chisel-shaped metatibial spine. Claws bifid, with ventrobasal teeth (
|
||
<figureCitation id="136D2A71E11FFFA81A4DFDDBFD68FDB6" box="[557,653,548,573]" captionStart="FIGURES 9" captionStartId="13.[151,264,925,948]" captionTargetBox="[159,1429,206,892]" captionTargetId="figure@13.[151,1436,195,904]" captionTargetPageId="13" captionText="FIGURES 9 A – F. Inner mesotarsal claw. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569451/files/figure.png" pageId="11" pageNumber="168">Figs. 9F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E11FFFA818A7FDB8FEEEFD2D" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="168">Ventral surface covered only by long, semierect, yellow macrosetae. Pygidium slightly transverse, convex, completely bordered; apically broadly rounded; irregularly covered by coarse, umbilicate, macrosetiferous punctures.</paragraph>
|
||
<paragraph id="8BE936F4E11FFFA818A7FD4BFD8CFC9E" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="168">
|
||
Male genitalia (
|
||
<figureCitation id="136D2A71E11FFFA819E1FD4BFE01FD46" box="[385,484,692,717]" captionStart="FIGURES 7" captionStartId="11.[151,264,1701,1724]" captionTargetBox="[185,1411,848,1668]" captionTargetId="figure@11.[177,1414,841,1674]" captionTargetPageId="11" captionText="FIGURES 7 A – F. Aedeagus in dorsal view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569449/files/figure.png" pageId="11" pageNumber="168">Figs. 7F</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A71E11FFFA81993FD4BFDF1FD46" box="[499,532,692,716]" captionStart="FIGURES 8" captionStartId="12.[151,264,1326,1349]" captionTargetBox="[170,1427,419,1302]" captionTargetId="figure@12.[151,1437,411,1305]" captionTargetPageId="12" captionText="FIGURES 8 A – F. Aedeagus in lateral view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569450/files/figure.png" pageId="11" pageNumber="168">8F</figureCitation>
|
||
). Parameres symmetrical, longer than phallobasis, regularly curved in lateral aspect, rounded with double hump apically in lateral aspect, on apex with double apex in dorsal aspect; covered by short, fine, semierect, yellow macrosetae.
|
||
</paragraph>
|
||
<caption id="DF29667CE11FFFA818F7F95AFADBF973" httpUri="https://zenodo.org/record/569449/files/figure.png" pageId="11" pageNumber="168" targetBox="[185,1411,848,1668]" targetPageId="11">
|
||
<paragraph id="8BE936F4E11FFFA818F7F95AFADBF973" blockId="11.[151,1436,1701,1785]" pageId="11" pageNumber="168">
|
||
<emphasis id="B922EAE6E11FFFA818F7F95AFEA9F931" bold="true" box="[151,332,1701,1724]" pageId="11" pageNumber="168">FIGURES 7A–F.</emphasis>
|
||
Aedeagus in dorsal view. A,
|
||
<taxonomicName id="4C564D77E11FFFA81A15F959FC33F931" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[629,982,1701,1723]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="apicata">
|
||
<emphasis id="B922EAE6E11FFFA81A15F959FCABF931" box="[629,846,1702,1723]" italics="true" pageId="11" pageNumber="168">Entypophana apicata</emphasis>
|
||
Moser, 1913
|
||
</taxonomicName>
|
||
, holotype; B,
|
||
<taxonomicName id="4C564D77E11FFFA81C03F959FF29F950" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="biapicata">
|
||
<emphasis id="B922EAE6E11FFFA81C03F959FAB5F931" box="[1123,1360,1702,1723]" italics="true" pageId="11" pageNumber="168">Entypophana biapicata</emphasis>
|
||
Moser, 1913
|
||
</taxonomicName>
|
||
, lectotype; C,
|
||
<taxonomicName id="4C564D77E11FFFA8190FF93BFCE8F950" authority="Burgeon, 1946" authorityName="Burgeon" authorityYear="1946" box="[367,781,1732,1754]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="hulstaerti">
|
||
<emphasis id="B922EAE6E11FFFA8190FF93BFD86F953" box="[367,611,1732,1753]" italics="true" pageId="11" pageNumber="168">Entypophana hulstaerti</emphasis>
|
||
Burgeon, 1946
|
||
</taxonomicName>
|
||
, lectotype; D,
|
||
<taxonomicName id="4C564D77E11FFFA81BD2F93BFAE3F950" authority="Moser, 1917" authorityName="Moser" authorityYear="1917" box="[946,1286,1732,1754]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="lujai">
|
||
<emphasis id="B922EAE6E11FFFA81BD2F93BFB96F953" box="[946,1139,1732,1753]" italics="true" pageId="11" pageNumber="168">Entypophana lujai</emphasis>
|
||
Moser, 1917
|
||
</taxonomicName>
|
||
, lectotype; E,
|
||
<taxonomicName id="4C564D77E11FFFA818F7F91CFDE9F973" authority="Burgeon, 1946" authorityName="Burgeon" authorityYear="1946" box="[151,524,1763,1785]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="maynei">
|
||
<emphasis id="B922EAE6E11FFFA818F7F91CFE88F973" box="[151,365,1763,1785]" italics="true" pageId="11" pageNumber="168">Entypophana maynei</emphasis>
|
||
Burgeon, 1946
|
||
</taxonomicName>
|
||
, lectotype; F,
|
||
<taxonomicName id="4C564D77E11FFFA81AF9F91CFC68F972" box="[665,909,1763,1784]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="njiapanda" status="sp. nov.">
|
||
<emphasis id="B922EAE6E11FFFA81AF9F91CFC68F972" box="[665,909,1763,1784]" italics="true" pageId="11" pageNumber="168">Entypophana njiapanda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A211579DE11FFFA81BF3F91BFBF5F973" box="[915,1040,1763,1785]" pageId="11" pageNumber="168" rank="species">new species</taxonomicNameLabel>
|
||
, holotype. Scale bar: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE936F4E11FFFA818A7F8DAFCBBF8E8" blockId="11.[151,1437,1829,2034]" pageId="11" pageNumber="168">
|
||
<emphasis id="B922EAE6E11FFFA818A7F8DAFE54F8B4" bold="true" box="[199,433,1829,1854]" pageId="11" pageNumber="168">Sexual dimorphism</emphasis>
|
||
. Female differs from male in the following characters: body length 20.0 mm, metatibia more strongly expanded apically; tarsomeres of all legs shorter.
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E11FFFA818A7F892FC50F80C" blockId="11.[151,1437,1829,2034]" box="[199,949,1901,1926]" pageId="11" pageNumber="168">
|
||
<emphasis id="B922EAE6E11FFFA818A7F892FE02F80C" bold="true" box="[199,487,1901,1926]" pageId="11" pageNumber="168">Geographic distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F3417664E11FFFA81995F892FDB9F80C" box="[501,604,1901,1926]" name="Tanzania" pageId="11" pageNumber="168">Tanzania</collectingCountry>
|
||
, Kilombero District (
|
||
<figureCitation id="136D2A71E11FFFA81B2CF892FC48F80C" box="[844,941,1901,1926]" captionStart="FIGURES 10" captionStartId="13.[151,264,1927,1950]" captionTargetBox="[151,1435,1059,1906]" captionTargetId="figure@13.[151,1435,1059,1906]" captionTargetPageId="13" captionText="FIGURES 10 A – F. Map with currently known geographical distribution. A, Entypophana apicata Moser, 1913; B, Entypophana biapicata Moser, 1913; C, Entypophana hulstaerti Burgeon, 1946; D, Entypophana lujai Moser, 1917; E, Entypophana maynei Burgeon, 1946; F, Entypophana njiapanda new species." httpUri="https://zenodo.org/record/569452/files/figure.png" pageId="11" pageNumber="168">Fig. 10F</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C657FE11FFFAF18A7F86EFED3FF72" lastPageId="12" lastPageNumber="169" pageId="11" pageNumber="168" type="diagnosis">
|
||
<paragraph id="8BE936F4E11FFFAF18A7F86EFED3FF72" blockId="11.[151,1437,1829,2034]" lastBlockId="12.[151,1436,151,356]" lastPageId="12" lastPageNumber="169" pageId="11" pageNumber="168">
|
||
<emphasis id="B922EAE6E11FFFA818A7F86EFE34F820" bold="true" box="[199,465,1937,1962]" pageId="11" pageNumber="168">Differential diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C564D77E11FFFA8198AF86DFCE1F820" box="[490,772,1938,1962]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="njiapanda">
|
||
<emphasis id="B922EAE6E11FFFA8198AF86DFCE1F820" box="[490,772,1938,1962]" italics="true" pageId="11" pageNumber="168">Entypophana njiapanda</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from all species of the genus
|
||
<taxonomicName id="4C564D77E11FFFA818F7F849FEC9F844" box="[151,300,1974,1998]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B922EAE6E11FFFA818F7F849FEC9F844" box="[151,300,1974,1998]" italics="true" pageId="11" pageNumber="168">Entypophana</emphasis>
|
||
</taxonomicName>
|
||
using the following diagnostic characters in males: frontoclypeal carina low, sinuate in the forward direction; surface behind frontoclypeal carina densely but unevenly punctate except for a narrow, impunctate strip adjacent to frontoclypeal suture (
|
||
<figureCitation id="136D2A71E118FFAF1A7FFF68FD99FF3A" box="[543,636,151,176]" captionStart="FIGURES 6" captionStartId="10.[151,264,1055,1078]" captionTargetBox="[166,1431,203,1025]" captionTargetId="figure@10.[151,1435,193,1034]" captionTargetPageId="10" captionText="FIGURES 6 A – D. Entypophana njiapanda new species male, holotype; length 17.9 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569448/files/figure.png" pageId="12" pageNumber="169">Fig. 6B</figureCitation>
|
||
); head including clypeus smooth, glossy, fissured, densely coarsely punctate; each head puncture with an erect macroseta shorter than puncture diameter (
|
||
<figureCitation id="136D2A71E118FFAF1C01FF43FB5CFF5F" box="[1121,1209,188,213]" captionStart="FIGURES 6" captionStartId="10.[151,264,1055,1078]" captionTargetBox="[166,1431,203,1025]" captionTargetId="figure@10.[151,1435,193,1034]" captionTargetPageId="10" captionText="FIGURES 6 A – D. Entypophana njiapanda new species male, holotype; length 17.9 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569448/files/figure.png" pageId="12" pageNumber="169">Fig. 6B</figureCitation>
|
||
); shape of aedeagus (
|
||
<figureCitation id="136D2A71E118FFAF18FFFF20FF18FF7D" box="[159,253,223,248]" captionStart="FIGURES 7" captionStartId="11.[151,264,1701,1724]" captionTargetBox="[185,1411,848,1668]" captionTargetId="figure@11.[177,1414,841,1674]" captionTargetPageId="11" captionText="FIGURES 7 A – F. Aedeagus in dorsal view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569449/files/figure.png" pageId="12" pageNumber="169">Figs. 7F</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A71E118FFAF1968FF1FFECDFF72" box="[264,296,224,248]" captionStart="FIGURES 8" captionStartId="12.[151,264,1326,1349]" captionTargetBox="[170,1427,419,1302]" captionTargetId="figure@12.[151,1437,411,1305]" captionTargetPageId="12" captionText="FIGURES 8 A – F. Aedeagus in lateral view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569450/files/figure.png" pageId="12" pageNumber="169">8F</figureCitation>
|
||
);
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C657FE118FFAF18A7FEFBFF3AFEEE" pageId="12" pageNumber="169" type="etymology">
|
||
<paragraph id="8BE936F4E118FFAF18A7FEFBFF3AFEEE" blockId="12.[151,1436,151,356]" pageId="12" pageNumber="169">
|
||
<emphasis id="B922EAE6E118FFAF18A7FEFBFEA3FE97" bold="true" box="[199,326,260,285]" pageId="12" pageNumber="169">Etymology</emphasis>
|
||
. Derived from name of
|
||
<typeStatus id="54ED8856E118FFAF1A35FEFAFD60FE97" box="[597,645,261,285]" pageId="12" pageNumber="169">type</typeStatus>
|
||
locality Njia Panda (
|
||
<collectingCountry id="F3417664E118FFAF1B11FEFBFC38FE97" box="[881,989,260,285]" name="Tanzania" pageId="12" pageNumber="169">Tanzania</collectingCountry>
|
||
); noun in apposition.
|
||
<emphasis id="B922EAE6E118FFAF18A7FED8FE17FECA" bold="true" box="[199,498,295,320]" pageId="12" pageNumber="169">Collecting circumstances</emphasis>
|
||
. Both
|
||
<typeStatus id="54ED8856E118FFAF1A20FED7FD95FECA" box="[576,624,296,320]" pageId="12" pageNumber="169">type</typeStatus>
|
||
specimens were attracted to light (H. Takano, personal communication 2013).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |