treatments-xml/data/03/FF/87/03FF87D6FFBB4467FF6BA94FFD77FB4A.xml
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<document id="3F4890CF3F7EF52641D93317C7EAF963" ID-CLB-Dataset="25263" ID-DOI="10.11646/zootaxa.4683.4.1" ID-GBIF-Dataset="48496621-de1a-4490-b866-1ae69d81ecbc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3479609" ID-ZooBank="982FE3FB-5610-44A1-AAE3-659B28F45307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570688900916" checkinUser="plazi" docAuthor="Lee, Chi-Feng &amp; Bezdĕk, Jan" docDate="2019" docId="03FF87D6FFBB4467FF6BA94FFD77FB4A" docLanguage="en" docName="zootaxa.4683.4.1.pdf" docOrigin="Zootaxa 4683 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Theopea yuae Lee &amp; Bezdĕk 2019, sp. nov." docType="treatment" docVersion="15" lastPageNumber="498" masterDocId="FFC6FFAEFF974448FFFCAF12FFAEFFA1" masterDocTitle="Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland and the Philippines: Redefinition of the genus" masterLastPageNumber="507" masterPageNumber="451" pageNumber="495" updateTime="1698758705207" updateUser="ExternalLinkService">
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<mods:title id="2F011516D672DC1549B8EF991F0D3E19">Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland and the Philippines: Redefinition of the genus</mods:title>
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<mods:namePart id="CD903169185AC279268AB81814550EA7">Lee, Chi-Feng</mods:namePart>
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<mods:namePart id="71B24F61529A3C368D494C1024E2FF0F">Bezdĕk, Jan</mods:namePart>
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<mods:title id="595721FFEE330AC18F475D84BED3353B">Zootaxa</mods:title>
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<treatment id="03FF87D6FFBB4467FF6BA94FFD77FB4A" ID-DOI="http://doi.org/10.5281/zenodo.4505281" ID-GBIF-Taxon="159690566" ID-Zenodo-Dep="4505281" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4467FF6BA94FFD77FB4A" httpUri="http://treatment.plazi.org/id/03FF87D6FFBB4467FF6BA94FFD77FB4A" lastPageId="47" lastPageNumber="498" pageId="44" pageNumber="495">
<subSubSection id="C34C654BFFBB4464FF6BA94FFE0BF9D9" box="[151,421,1629,1656]" pageId="44" pageNumber="495" type="nomenclature">
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<heading id="D0A181ACFFBB4464FF6BA94FFE0BF9D9" bold="true" box="[151,421,1629,1656]" fontSize="11" level="1" pageId="44" pageNumber="495" reason="1">
<emphasis id="B922EAD2FFBB4464FF6BA94FFE0BF9D9" bold="true" box="[151,421,1629,1656]" pageId="44" pageNumber="495">
<taxonomicName id="4C564D43FFBB4464FF6BA94FFEEEF9D9" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[151,320,1629,1656]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="yuae" status="sp. nov.">
<emphasis id="B922EAD2FFBB4464FF6BA94FFEEEF9D9" bold="true" box="[151,320,1629,1656]" italics="true" pageId="44" pageNumber="495">Theopea yuae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157A9FFBB4464FEBBA94CFE0BF9D9" box="[327,421,1630,1656]" pageId="44" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C34C654BFFBB4467FF6BA993FC9AFC06" lastPageId="47" lastPageNumber="498" pageId="44" pageNumber="495" type="description">
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(
<figureCitation id="136D2A45FFBB4464FF63A993FF5DF93A" box="[159,243,1665,1691]" captionStart="FIGURE 14" captionStartId="29.[151,250,1897,1922]" captionTargetBox="[238,1338,183,1873]" captionTargetId="figure@29.[232,1356,174,1878]" captionTargetPageId="29" captionText="FIGURE 14. Habitus of Theopea elegantula Baly, T. houjayi sp. nov., and T. yuae sp. nov. 14A. T. elegantula, male, dorsal view; 14B. Same, ventral view; 14C. T. houjayi sp. nov., male, dorsal view; 14D. Same, ventral view; 14E. T. houjayi sp. nov., female, dorsal view; 14F. T. yuae sp. nov., male, dorsal view; 14G. Same, ventral view; 14H. T. yuae sp. nov., male, dorsal view, color variation; 14I. T. yuae sp. nov., male, dorsal view, color variation." figureDoi="http://doi.org/10.5281/zenodo.3479637" httpUri="https://zenodo.org/record/3479637/files/figure.png" pageId="44" pageNumber="495">Figs 14</figureCitation>
F
<date id="FFE81000FFBB4464FEE8A993FE97F93A" box="[276,313,1665,1691]" pageId="44" pageNumber="495" value="2014-01">14I</date>
, 24)
</paragraph>
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<emphasis id="B922EAD2FFBB4464FF6BA9D8FE99F945" bold="true" box="[151,311,1737,1764]" pageId="44" pageNumber="495">Types (n= 8).</emphasis>
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<typeStatus id="54ED8862FFBB4464FEBCA9DBFE06F942" box="[320,424,1737,1763]" pageId="44" pageNumber="495" type="holotype">Holotype</typeStatus>
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(
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</emphasis>
. Sarawak.
<location id="8E89601BFFBB4464FCABA9DBFBB0F942" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4467FF6BA94FFD77FB4A:8E89601BFFBB4464FCABA9DBFBB0F942" box="[855,1054,1737,1764]" country="Malaysia" municipality="Matang Wildlife Centre" name="Kubah Nat. Park" pageId="44" pageNumber="495">Kubah Nat. Park</location>
.,
<collectingMunicipality id="6B8DACBAFFBB4464FBD2A9D8FA90F942" box="[1070,1342,1737,1764]" pageId="44" pageNumber="495">Matang Wildlife Centre</collectingMunicipality>
vic.,
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, leg.
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<materialsCitation id="3B3E3C9DFFBB4464FDCAA9FCFE6FF88A" ID-GBIF-Occurrence="2425473971" collectingDate="1996-06-24" collectingDateMax="1996-07-01" collectingDateMin="1996-06-24" collectionCode="SMNS" collectorName="J. Kodada" country="Malaysia" location="Batu Punggul Resort" pageId="44" pageNumber="495" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="54ED8862FFBB4464FDCAA9FCFD09F8A9" box="[566,679,1774,1800]" pageId="44" pageNumber="495" type="paratype">Paratypes</typeStatus>
.
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<collectingCountry id="F3417650FFBB4464FD4FA9FFFCE7F8A9" box="[691,841,1773,1800]" name="Malaysia" pageId="44" pageNumber="495">MALAYSIA</collectingCountry>
</emphasis>
.
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.
<specimenCount id="9D50FD49FFBB4464FC51A9FFFC7EF8A6" box="[941,976,1773,1799]" pageId="44" pageNumber="495" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FC1EA9FFFB9DF8A6" box="[994,1075,1773,1799]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="44" pageNumber="495">SMNS</collectionCode>
),
<location id="8E89601BFFBB4464FBB5A9FCFA97F8A9" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4467FF6BA94FFD77FB4A:8E89601BFFBB4464FBB5A9FCFA97F8A9" box="[1097,1337,1773,1800]" country="Malaysia" name="Batu Punggul Resort" pageId="44" pageNumber="495" stateProvince="Sabah">Batu Punggul Resort</location>
,
<date id="FFE81000FFBB4464FABBA9FFFEBAF88A" pageId="44" pageNumber="495" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">
<collectingDate id="EFACE9E8FFBB4464FABBA9FFFEBAF88A" pageId="44" pageNumber="495" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">24.VI. 1.VII.1996</collectingDate>
</date>
, leg.
<collectorName id="26A35316FFBB4464FEB2A800FE6FF88A" box="[334,449,1809,1836]" pageId="44" pageNumber="495">J. Kodada</collectorName>
</materialsCitation>
;
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<specimenCount id="9D50FD49FFBB4464FE37A803FE40F88A" box="[459,494,1809,1835]" pageId="44" pageNumber="495" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FE00A803FDE3F88A" box="[508,589,1809,1835]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="44" pageNumber="495">SMNS</collectionCode>
),
<collectorName id="26A35316FFBB4464FD9CA803FD71F88D" box="[608,735,1809,1836]" pageId="44" pageNumber="495">Bingkor, N</collectorName>
.
<location id="8E89601BFFBB4464FD15A803FCF7F88A" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4467FF6BA94FFD77FB4A:8E89601BFFBB4464FD15A803FCF7F88A" box="[745,857,1809,1835]" country="Malaysia" name="Keningau" pageId="44" pageNumber="495" stateProvince="Sabah">Keningau</location>
,
<date id="FFE81000FFBB4464FC9FA803FC4BF88A" box="[867,997,1809,1835]" pageId="44" pageNumber="495" value="1996-11-19">
<collectingDate id="EFACE9E8FFBB4464FC9FA803FC4BF88A" box="[867,997,1809,1835]" pageId="44" pageNumber="495" value="1996-11-19">19.XI.1996</collectingDate>
</date>
, leg.
<collectorName id="26A35316FFBB4464FBE3A800FB3BF88A" box="[1055,1173,1809,1836]" pageId="44" pageNumber="495">D. Grimm</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFFBB4464FB63A803FCDEF8EE" ID-GBIF-Occurrence="2425474126" collectingDate="2008-03-09" collectingDateMax="2008-03-12" collectingDateMin="2008-03-09" collectionCode="SMNS" collectorName="W. Schawaller" country="Malaysia" elevation="225" location="Gn. Gading" pageId="44" pageNumber="495" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry id="F3417650FFBB4464FB63A803FAAAF88A" box="[1183,1284,1809,1835]" name="Malaysia" pageId="44" pageNumber="495">Sarawak</collectingCountry>
:
<specimenCount id="9D50FD49FFBB4464FAF1A803FA9EF88A" box="[1293,1328,1809,1835]" pageId="44" pageNumber="495" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FAC2A803FA21F88A" box="[1342,1423,1809,1835]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="44" pageNumber="495">SMNS</collectionCode>
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<location id="8E89601BFFBB4464FF6BA827FEB7F8EE" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4467FF6BA94FFD77FB4A:8E89601BFFBB4464FF6BA827FEB7F8EE" box="[151,281,1845,1871]" country="Malaysia" name="Gn. Gading" pageId="44" pageNumber="495" stateProvince="Sabah">Gn. Gading</location>
N. P.,
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<date id="FFE81000FFBB4464FE0EA827FD20F8EE" box="[498,654,1845,1871]" pageId="44" pageNumber="495" value="2008-03-09" valueMax="2008-03-12" valueMin="2008-03-09">
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</date>
, leg.
<collectorName id="26A35316FFBB4464FD36A824FCDEF8EE" box="[714,880,1845,1872]" pageId="44" pageNumber="495">W. Schawaller</collectorName>
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;
<materialsCitation id="3B3E3C9DFFBB4464FC87A827FE90F8D2" ID-GBIF-Occurrence="2425474179" collectingDate="1999-05-12" collectionCode="NMPC" collectorName="P. Votruba" country="Malaysia" location="Miri" municipality="Lambir" pageId="44" pageNumber="495" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D50FD49FFBB4464FC87A827FC30F8EE" box="[891,926,1845,1871]" pageId="44" pageNumber="495" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FC51A827FBACF8EE" box="[941,1026,1845,1871]" country="Czech Republic" name="National Museum Prague" pageId="44" pageNumber="495">NMPC</collectionCode>
),
<collectingMunicipality id="6B8DACBAFFBB4464FBE9A827FBC9F8EE" box="[1045,1127,1845,1871]" pageId="44" pageNumber="495">Lambir</collectingMunicipality>
hills N.P.,
<location id="8E89601BFFBB4464FB1FA827FAB6F8EE" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4467FF6BA94FFD77FB4A:8E89601BFFBB4464FB1FA827FAB6F8EE" box="[1251,1304,1845,1871]" country="Malaysia" municipality="Lambir" name="Miri" pageId="44" pageNumber="495" stateProvince="Sabah">Miri</location>
,
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<collectingDate id="EFACE9E8FFBB4464FADFA827FA36F8EE" box="[1315,1432,1845,1871]" pageId="44" pageNumber="495" value="1999-05-12">12.V.1999</collectingDate>
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, leg.
<collectorName id="26A35316FFBB4464FF34A848FE90F8D2" box="[200,318,1881,1908]" pageId="44" pageNumber="495">P. Votruba</collectorName>
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;
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,
<specimenCount id="9D50FD49FFBB4464FE86A84BFE33F8D2" box="[378,413,1881,1907]" pageId="44" pageNumber="495" type="female">1♀</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FE50A84BFDA8F8D2" box="[428,518,1881,1907]" pageId="44" pageNumber="495">BMNH</collectionCode>
),
<location id="8E89601BFFBB4464FDE5A84BFDCBF8D2" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4467FF6BA94FFD77FB4A:8E89601BFFBB4464FDE5A84BFDCBF8D2" box="[537,613,1881,1907]" country="Malaysia" name="Lundu" pageId="44" pageNumber="495" stateProvince="Sabah">Lundu</location>
,
<date id="FFE81000FFBB4464FD8CA84BFD14F8D2" box="[624,698,1881,1907]" pageId="44" pageNumber="495" value="1914-01">
<collectingDate id="EFACE9E8FFBB4464FD8CA84BFD14F8D2" box="[624,698,1881,1907]" pageId="44" pageNumber="495" value="1914-01">I.1914</collectingDate>
</date>
, leg.
<collectorName id="26A35316FFBB4464FD0BA848FCE9F8D5" box="[759,839,1882,1908]" pageId="44" pageNumber="495">Bryant</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFFBB4464FCAEA84BFB45F8D2" ID-GBIF-Occurrence="2425473969" box="[850,1259,1881,1908]" collectingDate="1913-12" collectionCode="BMNH" country="Malaysia" location="Mt. Matang" pageId="44" pageNumber="495" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D50FD49FFBB4464FCAEA84BFCDBF8D2" box="[850,885,1881,1907]" pageId="44" pageNumber="495" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FC78A84BFC70F8D2" box="[900,990,1881,1907]" pageId="44" pageNumber="495">BMNH</collectionCode>
),
<location id="8E89601BFFBB4464FC0DA848FBD9F8D5" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4467FF6BA94FFD77FB4A:8E89601BFFBB4464FC0DA848FBD9F8D5" box="[1009,1143,1882,1908]" country="Malaysia" name="Mt. Matang" pageId="44" pageNumber="495" stateProvince="Sabah">Mt. Matang</location>
,
<date id="FFE81000FFBB4464FB7FA84BFB45F8D2" box="[1155,1259,1881,1907]" pageId="44" pageNumber="495" value="1913-12">
<collectingDate id="EFACE9E8FFBB4464FB7FA84BFB45F8D2" box="[1155,1259,1881,1907]" pageId="44" pageNumber="495" value="1913-12">XII.1913</collectingDate>
</date>
</materialsCitation>
, leg. G. E. Bry- ant;
<materialsCitation id="3B3E3C9DFFBB4464FF34A86FFDC1F836" ID-GBIF-Occurrence="2425474146" box="[200,623,1917,1944]" collectingDate="1914-01" collectionCode="BMNH" country="Malaysia" pageId="44" pageNumber="495" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D50FD49FFBB4464FF34A86FFF45F836" box="[200,235,1917,1943]" pageId="44" pageNumber="495" type="female">1♀</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FF06A86FFEFAF836" box="[250,340,1917,1943]" pageId="44" pageNumber="495">BMNH</collectionCode>
), same but with “
<date id="FFE81000FFBB4464FDEAA86FFDCAF836" box="[534,612,1917,1943]" pageId="44" pageNumber="495" value="1914-01">
<collectingDate id="EFACE9E8FFBB4464FDEAA86FFDCAF836" box="[534,612,1917,1943]" pageId="44" pageNumber="495" value="1914-01">I.1914</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFFBB4464FD86A86FFBE7F836" ID-GBIF-Occurrence="2425474198" box="[634,1097,1917,1944]" collectingDate="1914-01" collectingDateMax="1914-02-31" collectingDateMin="1914-01" collectionCode="BMNH" country="Malaysia" pageId="44" pageNumber="495" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D50FD49FFBB4464FD86A86FFD33F836" box="[634,669,1917,1943]" pageId="44" pageNumber="495" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FD50A86FFCA8F836" box="[684,774,1917,1943]" pageId="44" pageNumber="495">BMNH</collectionCode>
), same but with “
<date id="FFE81000FFBB4464FC34A86FFB90F836" box="[968,1086,1917,1943]" pageId="44" pageNumber="495" value="1914-01" valueMax="1914-02-31" valueMin="1914-01-00">
<collectingDate id="EFACE9E8FFBB4464FC34A86FFB90F836" box="[968,1086,1917,1943]" pageId="44" pageNumber="495" value="1914-01" valueMax="1914-02-31" valueMin="1914-01-00">I.II.1914</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFFBB4464FBA8A86FFD7DF81A" ID-GBIF-Occurrence="2425474203" collectingDate="1978-05" collectingDateMax="1978-08-31" collectingDateMin="1978-05" collectionCode="BMNH" collectorName="P. M. Hamond &amp; J. E. Marshall." country="Malaysia" location="Gn. Mulu" pageId="44" pageNumber="495" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D50FD49FFBB4464FBA8A86FFBD9F836" box="[1108,1143,1917,1943]" pageId="44" pageNumber="495" type="female">1♀</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FB7AA86FFB4EF836" box="[1158,1248,1917,1943]" pageId="44" pageNumber="495">BMNH</collectionCode>
),
<location id="8E89601BFFBB4464FB0FA86FFACEF836" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4467FF6BA94FFD77FB4A:8E89601BFFBB4464FB0FA86FFACEF836" box="[1267,1376,1917,1943]" country="Malaysia" name="Gn. Mulu" pageId="44" pageNumber="495" stateProvince="Sabah">Gn. Mulu</location>
N.P.,
<date id="FFE81000FFBB4464FF6BA8B3FE81F81A" box="[151,303,1953,1979]" pageId="44" pageNumber="495" value="1978-05" valueMax="1978-08-31" valueMin="1978-05-00">
<collectingDate id="EFACE9E8FFBB4464FF6BA8B3FE81F81A" box="[151,303,1953,1979]" pageId="44" pageNumber="495" value="1978-05" valueMax="1978-08-31" valueMin="1978-05-00">V.VIII.1978</collectingDate>
</date>
, leg.
<collectorName id="26A35316FFBB4464FE96A8B0FDA2F81A" box="[362,524,1953,1980]" pageId="44" pageNumber="495">P. M. Hamond</collectorName>
&amp;
<collectorName id="26A35316FFBB4464FDCCA8B0FD7DF81A" box="[560,723,1953,1980]" pageId="44" pageNumber="495">J. E. Marshall.</collectorName>
</materialsCitation>
</paragraph>
<caption id="DF296648FFBA4465FF6BA89EFE8FF84C" ID-DOI="http://doi.org/10.5281/zenodo.3479655" ID-Zenodo-Dep="3479655" httpUri="https://zenodo.org/record/3479655/files/figure.png" pageId="45" pageNumber="496" startId="45.[151,250,1932,1957]" targetBox="[163,1420,179,1908]" targetPageId="45">
<paragraph id="8BE936C0FFBA4465FF6BA89EFE8FF84C" blockId="45.[151,1437,1932,2029]" pageId="45" pageNumber="496">
<emphasis id="B922EAD2FFBA4465FF6BA89EFE8CF804" bold="true" box="[151,290,1932,1957]" pageId="45" pageNumber="496">FIGURE 23.</emphasis>
Diagnostic characters of
<taxonomicName id="4C564D43FFBA4465FDD2A89EFD11F805" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[558,703,1932,1956]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="496" phylum="Arthropoda" rank="species" species="tsoui" status="sp. nov.">
<emphasis id="B922EAD2FFBA4465FDD2A89EFD11F805" box="[558,703,1932,1956]" italics="true" pageId="45" pageNumber="496">Theopea tsoui</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFBA4465FD34A89FFCB3F804" bold="true" box="[712,797,1933,1957]" pageId="45" pageNumber="496">
<taxonomicNameLabel id="A21157A9FFBA4465FD34A89FFCB3F804" box="[712,797,1933,1957]" pageId="45" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
23A. Antenna, male; 23B. Antenna, female; 23C. Aedeagus, dorsal view; 23D. Aedeagus, lateral view; 23E. Aedeagus, ventral view; 23F. Abdominal ventrite VIII; 23G. Gonocoxae; 23H. Spermatheca.
</paragraph>
</caption>
<caption id="DF296648FFB94466FF6BA87EFE8FF86C" ID-DOI="http://doi.org/10.5281/zenodo.3479657" ID-Zenodo-Dep="3479657" httpUri="https://zenodo.org/record/3479657/files/figure.png" pageId="46" pageNumber="497" startId="46.[151,250,1900,1925]" targetBox="[166,1421,184,1873]" targetPageId="46">
<paragraph id="8BE936C0FFB94466FF6BA87EFE8FF86C" blockId="46.[151,1437,1900,1997]" pageId="46" pageNumber="497">
<emphasis id="B922EAD2FFB94466FF6BA87EFE8CF824" bold="true" box="[151,290,1900,1925]" pageId="46" pageNumber="497">FIGURE 24.</emphasis>
Diagnostic characters of
<taxonomicName id="4C564D43FFB94466FDD3A87EFD11F825" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[559,703,1900,1924]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="yuae" status="sp. nov.">
<emphasis id="B922EAD2FFB94466FDD3A87EFD11F825" box="[559,703,1900,1924]" italics="true" pageId="46" pageNumber="497">Theopea yuae</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB94466FD3BA87FFCB3F824" bold="true" box="[711,797,1901,1925]" pageId="46" pageNumber="497">
<taxonomicNameLabel id="A21157A9FFB94466FD3BA87FFCB3F824" box="[711,797,1901,1925]" pageId="46" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
24A. Antenna, male; 24B. Antenna, female; 24C. Aedeagus, dorsal view; 24D. Aedeagus, lateral view; 24E. Aedeagus, ventral view; 24F. Abdominal ventrite VIII; 24G. Gonocoxae; 24H. Spermatheca.
</paragraph>
</caption>
<paragraph id="8BE936C0FFB84467FF3BAF8AFC50FCFF" blockId="47.[151,1437,152,1259]" pageId="47" pageNumber="498">
<emphasis id="B922EAD2FFB84467FF3BAF8AFEF7FF12" bold="true" box="[199,345,152,179]" pageId="47" pageNumber="498">Description.</emphasis>
Length
<quantity id="4CAE9B25FFB84467FE46AF8BFDEBFF12" box="[442,581,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.2" metricValueMin="7.1" pageId="47" pageNumber="498" unit="mm" value="7.15" valueMax="7.2" valueMin="7.1">7.17.2 mm</quantity>
, width
<quantity id="4CAE9B25FFB84467FD67AF8BFC89FF12" box="[667,807,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="47" pageNumber="498" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.72.8 mm</quantity>
. General color (
<figureCitation id="136D2A45FFB84467FC1CAF8AFB2BFF12" box="[992,1157,152,179]" captionStart="FIGURE 14" captionStartId="29.[151,250,1897,1922]" captionTargetBox="[238,1338,183,1873]" captionTargetId="figure@29.[232,1356,174,1878]" captionTargetPageId="29" captionText="FIGURE 14. Habitus of Theopea elegantula Baly, T. houjayi sp. nov., and T. yuae sp. nov. 14A. T. elegantula, male, dorsal view; 14B. Same, ventral view; 14C. T. houjayi sp. nov., male, dorsal view; 14D. Same, ventral view; 14E. T. houjayi sp. nov., female, dorsal view; 14F. T. yuae sp. nov., male, dorsal view; 14G. Same, ventral view; 14H. T. yuae sp. nov., male, dorsal view, color variation; 14I. T. yuae sp. nov., male, dorsal view, color variation." figureDoi="http://doi.org/10.5281/zenodo.3479637" httpUri="https://zenodo.org/record/3479637/files/figure.png" pageId="47" pageNumber="498">Figs 14F, 14G</figureCitation>
) metallic blue; antennomeres IVIII black, IXXI white or pale yellow. Antennae filiform in male, antennomeres VII and VIII slightly swollen, outer margins of IX and X with processes at middles (
<figureCitation id="136D2A45FFB84467FCA1AFF2FC68FF5A" box="[861,966,224,251]" captionStart="FIGURE 24" captionStartId="46.[151,250,1900,1925]" captionTargetBox="[166,1421,184,1873]" captionTargetId="figure@46.[151,1436,174,1881]" captionTargetPageId="46" captionText="FIGURE 24. Diagnostic characters of Theopea yuae sp. nov. 24A. Antenna, male; 24B. Antenna, female; 24C. Aedeagus, dorsal view; 24D. Aedeagus, lateral view; 24E. Aedeagus, ventral view; 24F. Abdominal ventrite VIII; 24G. Gonocoxae; 24H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479657" httpUri="https://zenodo.org/record/3479657/files/figure.png" pageId="47" pageNumber="498">Fig. 24A</figureCitation>
), length ratios of antennomeres IXI 1.0: 0.3: 0.7: 0.9: 1.0: 0.9: 1.0: 1.0: 0.8: 0.7: 0.9, length to width ratios of antennomeres IXI 3.3: 1.4: 2.6: 3.3: 3.7: 3.3: 2.8: 3.3: 2.7: 2.8: 4.5; filiform in females (
<figureCitation id="136D2A45FFB84467FD03AE3AFCCBFEE2" box="[767,869,296,323]" captionStart="FIGURE 24" captionStartId="46.[151,250,1900,1925]" captionTargetBox="[166,1421,184,1873]" captionTargetId="figure@46.[151,1436,174,1881]" captionTargetPageId="46" captionText="FIGURE 24. Diagnostic characters of Theopea yuae sp. nov. 24A. Antenna, male; 24B. Antenna, female; 24C. Aedeagus, dorsal view; 24D. Aedeagus, lateral view; 24E. Aedeagus, ventral view; 24F. Abdominal ventrite VIII; 24G. Gonocoxae; 24H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479657" httpUri="https://zenodo.org/record/3479657/files/figure.png" pageId="47" pageNumber="498">Fig. 24B</figureCitation>
), length ratios of antennomeres IXI 1.0: 0.3: 0.7: 0.8: 0.9: 0.8: 0.9: 0.8: 0.8: 0.7: 0.9, length to width ratios of antennomeres IXI 3.3: 1.4: 2.9: 3.3: 3.5: 3.1: 3.3: 3.4: 3.4: 3.2: 4.2. Elytra elongate and parallel-sided, 1.8x longer than wide; disc with dense, coarse punctures, arranged into longitudinal rows, with one weak longitudinal ridge between two longitudinal rows of punctures, ridges near suture abbreviated. Tarsomeres I of front legs swollen in males; subparallel in females. Aedeagus (
<figureCitation id="136D2A45FFB84467FA96AEAAFF1DFE56" captionStart="FIGURE 24" captionStartId="46.[151,250,1900,1925]" captionTargetBox="[166,1421,184,1873]" captionTargetId="figure@46.[151,1436,174,1881]" captionTargetPageId="46" captionText="FIGURE 24. Diagnostic characters of Theopea yuae sp. nov. 24A. Antenna, male; 24B. Antenna, female; 24C. Aedeagus, dorsal view; 24D. Aedeagus, lateral view; 24E. Aedeagus, ventral view; 24F. Abdominal ventrite VIII; 24G. Gonocoxae; 24H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479657" httpUri="https://zenodo.org/record/3479657/files/figure.png" pageId="47" pageNumber="498">Figs 24</figureCitation>
C24E) extremely slender, 7.1x longer than wide; parallel-sided, slightly narrower at apical 1/6, apex truncate; tectum well sclerotized, basally broadened, broader than aedeagus; slightly curved in lateral view; ventral surface with deep notch from apex to base, narrowed at apical 1/6, abruptly broadened at apical 1/4, basal notch open; triangular sclerites present; internal sac with one broad, elongate sclerite, apically narrower, one extremely elongate sclerite located dorsally, with one pair of clusters of stout setae along sides, with one pair small stout setae located posteriorly. Gonocoxae (
<figureCitation id="136D2A45FFB84467FE52AD82FDBBFD0A" box="[430,533,656,683]" captionStart="FIGURE 24" captionStartId="46.[151,250,1900,1925]" captionTargetBox="[166,1421,184,1873]" captionTargetId="figure@46.[151,1436,174,1881]" captionTargetPageId="46" captionText="FIGURE 24. Diagnostic characters of Theopea yuae sp. nov. 24A. Antenna, male; 24B. Antenna, female; 24C. Aedeagus, dorsal view; 24D. Aedeagus, lateral view; 24E. Aedeagus, ventral view; 24F. Abdominal ventrite VIII; 24G. Gonocoxae; 24H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479657" httpUri="https://zenodo.org/record/3479657/files/figure.png" pageId="47" pageNumber="498">Fig. 24G</figureCitation>
) elongate, widest at apex, both gonocoxae combined together from basal 1/5 to apical 1/7; apices narrowly rounded, each gonocoxa with eight setae along outer margin from apex to apical 1/5; with one pair of short lateral processes at basal 2/5. Ventrite VIII (
<figureCitation id="136D2A45FFB84467FC83ADCAFC4FFD52" box="[895,993,728,755]" captionStart="FIGURE 24" captionStartId="46.[151,250,1900,1925]" captionTargetBox="[166,1421,184,1873]" captionTargetId="figure@46.[151,1436,174,1881]" captionTargetPageId="46" captionText="FIGURE 24. Diagnostic characters of Theopea yuae sp. nov. 24A. Antenna, male; 24B. Antenna, female; 24C. Aedeagus, dorsal view; 24D. Aedeagus, lateral view; 24E. Aedeagus, ventral view; 24F. Abdominal ventrite VIII; 24G. Gonocoxae; 24H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479657" httpUri="https://zenodo.org/record/3479657/files/figure.png" pageId="47" pageNumber="498">Fig. 24F</figureCitation>
) longitudinal and well sclerotized; disc with several long setae laterally and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (
<figureCitation id="136D2A45FFB84467FD46AC32FC8AFC9A" box="[698,804,800,827]" captionStart="FIGURE 24" captionStartId="46.[151,250,1900,1925]" captionTargetBox="[166,1421,184,1873]" captionTargetId="figure@46.[151,1436,174,1881]" captionTargetPageId="46" captionText="FIGURE 24. Diagnostic characters of Theopea yuae sp. nov. 24A. Antenna, male; 24B. Antenna, female; 24C. Aedeagus, dorsal view; 24D. Aedeagus, lateral view; 24E. Aedeagus, ventral view; 24F. Abdominal ventrite VIII; 24G. Gonocoxae; 24H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479657" httpUri="https://zenodo.org/record/3479657/files/figure.png" pageId="47" pageNumber="498">Fig. 24H</figureCitation>
) strongly swollen; pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, narrow and short.
</paragraph>
<paragraph id="8BE936C0FFB84467FF3BAC7AFC9AFC06" blockId="47.[151,1437,152,1259]" pageId="47" pageNumber="498">
<emphasis id="B922EAD2FFB84467FF3BAC7AFEE4FC22" bold="true" box="[199,330,872,899]" pageId="47" pageNumber="498">Variations.</emphasis>
Some specimens possess paler antennomeres V and VI and white antennomeres VIIXI (
<figureCitation id="136D2A45FFB84467FACEAC7AFA21FC22" box="[1330,1423,872,899]" captionStart="FIGURE 14" captionStartId="29.[151,250,1897,1922]" captionTargetBox="[238,1338,183,1873]" captionTargetId="figure@29.[232,1356,174,1878]" captionTargetPageId="29" captionText="FIGURE 14. Habitus of Theopea elegantula Baly, T. houjayi sp. nov., and T. yuae sp. nov. 14A. T. elegantula, male, dorsal view; 14B. Same, ventral view; 14C. T. houjayi sp. nov., male, dorsal view; 14D. Same, ventral view; 14E. T. houjayi sp. nov., female, dorsal view; 14F. T. yuae sp. nov., male, dorsal view; 14G. Same, ventral view; 14H. T. yuae sp. nov., male, dorsal view, color variation; 14I. T. yuae sp. nov., male, dorsal view, color variation." figureDoi="http://doi.org/10.5281/zenodo.3479637" httpUri="https://zenodo.org/record/3479637/files/figure.png" pageId="47" pageNumber="498">
Fig.
<date id="FFE81000FFB84467FA9BAC7BFA21FC22" box="[1383,1423,873,899]" pageId="47" pageNumber="498" value="2014-01">14I</date>
</figureCitation>
). A male from
<collectingRegion id="4992F822FFB84467FED0AC9EFEDEFC07" box="[300,368,908,934]" country="Malaysia" name="Sabah" pageId="47" pageNumber="498">Sabah</collectingRegion>
has black antennomeres VII (
<figureCitation id="136D2A45FFB84467FD43AC9EFC86FC06" box="[703,808,908,935]" captionStart="FIGURE 14" captionStartId="29.[151,250,1897,1922]" captionTargetBox="[238,1338,183,1873]" captionTargetId="figure@29.[232,1356,174,1878]" captionTargetPageId="29" captionText="FIGURE 14. Habitus of Theopea elegantula Baly, T. houjayi sp. nov., and T. yuae sp. nov. 14A. T. elegantula, male, dorsal view; 14B. Same, ventral view; 14C. T. houjayi sp. nov., male, dorsal view; 14D. Same, ventral view; 14E. T. houjayi sp. nov., female, dorsal view; 14F. T. yuae sp. nov., male, dorsal view; 14G. Same, ventral view; 14H. T. yuae sp. nov., male, dorsal view, color variation; 14I. T. yuae sp. nov., male, dorsal view, color variation." figureDoi="http://doi.org/10.5281/zenodo.3479637" httpUri="https://zenodo.org/record/3479637/files/figure.png" pageId="47" pageNumber="498">Fig. 14H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFB84467FF3BACA2FBC2FBDF" pageId="47" pageNumber="498" type="diagnosis">
<paragraph id="8BE936C0FFB84467FF3BACA2FBC2FBDF" blockId="47.[151,1437,152,1259]" pageId="47" pageNumber="498">
<emphasis id="B922EAD2FFB84467FF3BACA2FEECFC6A" bold="true" box="[199,322,944,971]" pageId="47" pageNumber="498">Diagnosis.</emphasis>
Specimens of
<taxonomicName id="4C564D43FFB84467FE15ACA3FD94FC6B" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[489,570,945,970]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="yuae" status="sp. nov.">
<emphasis id="B922EAD2FFB84467FE15ACA3FD94FC6B" box="[489,570,945,970]" italics="true" pageId="47" pageNumber="498">T. yuae</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB84467FDBEACA3FD32FC6A" bold="true" box="[578,668,945,971]" pageId="47" pageNumber="498">
<taxonomicNameLabel id="A21157A9FFB84467FDBEACA3FD32FC6A" box="[578,668,945,971]" pageId="47" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="4C564D43FFB84467FC5FACA3FBBDFC6B" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[931,1043,944,970]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="houjayi" status="sp. nov.">
<emphasis id="B922EAD2FFB84467FC5FACA3FBBDFC6B" box="[931,1043,944,970]" italics="true" pageId="47" pageNumber="498">T. houjayi</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB84467FBE7ACA3FBDBFC6A" bold="true" box="[1051,1141,945,971]" pageId="47" pageNumber="498">
<taxonomicNameLabel id="A21157A9FFB84467FBE7ACA3FBDBFC6A" box="[1051,1141,945,971]" pageId="47" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in possessing three apical white antennomeres (two apical white antennomeres in
<taxonomicName id="4C564D43FFB84467FD03ACC7FCE1FC4F" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[767,847,981,1006]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="tsoui" status="sp. nov.">
<emphasis id="B922EAD2FFB84467FD03ACC7FCE1FC4F" box="[767,847,981,1006]" italics="true" pageId="47" pageNumber="498">T. tsoui</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB84467FCAAACC7FC00FC4E" bold="true" box="[854,942,981,1007]" pageId="47" pageNumber="498">
<taxonomicNameLabel id="A21157A9FFB84467FCAAACC7FC00FC4E" box="[854,942,981,1007]" pageId="47" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C564D43FFB84467FC1EACC7FBD7FC4F" box="[994,1145,980,1006]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="kedenburgi">
<emphasis id="B922EAD2FFB84467FC1EACC7FBD7FC4F" box="[994,1145,980,1006]" italics="true" pageId="47" pageNumber="498">T. kedenburgi</emphasis>
</taxonomicName>
) and reduced longitudinal ridges near suture of elytra (distinct longitudinal ridges near suture of elytra in
<taxonomicName id="4C564D43FFB84467FBF0ACEBFB30FBB3" box="[1036,1182,1016,1042]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="elegantula">
<emphasis id="B922EAD2FFB84467FBF0ACEBFB30FBB3" box="[1036,1182,1016,1042]" italics="true" pageId="47" pageNumber="498">T. elegantula</emphasis>
</taxonomicName>
). These two cannot be separated from each other based on external morphology. However,
<taxonomicName id="4C564D43FFB84467FC64AB0FFC47FB97" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[920,1001,1053,1078]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="yuae" status="sp. nov.">
<emphasis id="B922EAD2FFB84467FC64AB0FFC47FB97" box="[920,1001,1053,1078]" italics="true" pageId="47" pageNumber="498">T. yuae</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB84467FC0DAB0FFBE5FB96" bold="true" box="[1009,1099,1053,1079]" pageId="47" pageNumber="498">
<taxonomicNameLabel id="A21157A9FFB84467FC0DAB0FFBE5FB96" box="[1009,1099,1053,1079]" pageId="47" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only found in
<collectingCountry id="F3417650FFB84467FAF0AB0EFAC2FB97" box="[1292,1388,1052,1078]" name="Malaysia" pageId="47" pageNumber="498">Sarawak</collectingCountry>
and males possess broad and rounded apices of the male aedeagi, in contrast to
<taxonomicName id="4C564D43FFB84467FC21AB53FBE2FBFB" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[989,1100,1088,1114]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="houjayi" status="sp. nov.">
<emphasis id="B922EAD2FFB84467FC21AB53FBE2FBFB" box="[989,1100,1088,1114]" italics="true" pageId="47" pageNumber="498">T. houjayi</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB84467FBAFAB53FB02FBFA" bold="true" box="[1107,1196,1089,1115]" pageId="47" pageNumber="498">
<taxonomicNameLabel id="A21157A9FFB84467FBAFAB53FB02FBFA" box="[1107,1196,1089,1115]" pageId="47" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which is confined to Kalimantan. Males of the latter possess slender and notched apices of the male aedeagi.
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFB84467FF3BAB9BFE21FB67" pageId="47" pageNumber="498" type="etymology">
<paragraph id="8BE936C0FFB84467FF3BAB9BFE21FB67" blockId="47.[151,1437,152,1259]" pageId="47" pageNumber="498">
<emphasis id="B922EAD2FFB84467FF3BAB9BFEE0FB02" bold="true" box="[199,334,1161,1187]" pageId="47" pageNumber="498">Etymology.</emphasis>
This new species is named after Mrs. Su-Fang Yu, a member of TCRT for her contribution to di- versity of leaf beetles.
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFB84467FF3BABC2FD77FB4A" box="[199,729,1232,1259]" pageId="47" pageNumber="498" type="distribution">
<paragraph id="8BE936C0FFB84467FF3BABC2FD77FB4A" blockId="47.[151,1437,152,1259]" box="[199,729,1232,1259]" pageId="47" pageNumber="498">
<emphasis id="B922EAD2FFB84467FF3BABC2FECFFB4A" bold="true" box="[199,353,1232,1259]" pageId="47" pageNumber="498">Distribution.</emphasis>
East
<collectingCountry id="F3417650FFB84467FE63ABC2FDA8FB4B" box="[415,518,1232,1258]" name="Malaysia" pageId="47" pageNumber="498">Malaysia</collectingCountry>
(
<collectingRegion id="4992F822FFB84467FDE8ABC2FDF3FB4B" box="[532,605,1232,1258]" country="Malaysia" name="Sabah" pageId="47" pageNumber="498">Sabah</collectingRegion>
,
<collectingCountry id="F3417650FFB84467FD94ABC2FD60FB4A" box="[616,718,1232,1259]" name="Malaysia" pageId="47" pageNumber="498">Sarawak</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>