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<document id="0C2F4E5E7E21095089D945AABE061D5A" ID-CLB-Dataset="2408" ID-DOI="10.11646/zootaxa.5093.2.4" ID-GBIF-Dataset="a29a1307-a6a4-4198-8653-e79c13f4d008" ID-ISSN="1175-5326" ID-Zenodo-Dep="5905225" ID-ZooBank="FA40C88B-9784-4DC0-9109-90936BAFE808" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643182534301" checkinUser="plazi" docAuthor="Padate, Vinay P., Cubelio, Sherine Sonia &amp; Takeda, Masatsune" docDate="2022" docId="03FF87B3FFB34B5BFF54ABA28607C7AB" docLanguage="en" docName="zootaxa.5093.2.4.pdf" docOrigin="Zootaxa 5093 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guyanacaris keralam Padate &amp; Cubelio &amp; Takeda 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="209" masterDocId="FFC6FFCBFFB64B55FFC3AB35853EC67A" masterDocTitle="Two axiidean ghost shrimps (Crustacea: Decapoda) from India, Guyanacaris keralam sp. nov. (Axiidae) and Paragourretia galathea (K. Sakai, 2017) (Ctenochelidae)" masterLastPageNumber="217" masterPageNumber="195" pageNumber="200" updateTime="1699304328208" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="26C873C2B8C7DBBF413A359CB35F59CF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8BF4944A38FEE65D11970E006BEEC491">
<mods:title id="82F779C53C6440E8EE84E9DCA1CD70AD">Two axiidean ghost shrimps (Crustacea: Decapoda) from India, Guyanacaris keralam sp. nov. (Axiidae) and Paragourretia galathea (K. Sakai, 2017) (Ctenochelidae)</mods:title>
</mods:titleInfo>
<mods:name id="FE51D9955F5722D1C936BC94FAE6BEB1" type="personal">
<mods:role id="F86EE948BE857D12782EE307043180DA">
<mods:roleTerm id="7D09DBC9D27D7B1F5C1356C362954FDC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6EBDAD6E75A669DA074EB53F1CF353E6">Padate, Vinay P.</mods:namePart>
<mods:nameIdentifier id="C69456927D25B6A68060D2601EFAEC91" type="ORCID">0000-0002-2244-8338</mods:nameIdentifier>
<mods:affiliation id="418015820D7A141A10FF7ABC98BA71CF">Centre for Marine Living Resources &amp; Ecology, Ministry of Earth Sciences, Government of India, Atal Bhavan, LNG Terminus Road, Puthuvype, Kochi, 682508, India. &amp; vinaypadate @ gmail. com; https: // orcid. org / 0000 - 0002 - 2244 - 8338</mods:affiliation>
<mods:nameIdentifier id="2E1801E45BFD54E9F6675F57AB35BF31" type="email">vinaypadate@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="530884EADE7638F9B181E1D5E64F59F4" type="personal">
<mods:role id="B502A2F202763BC127B99D5E78D4C9B2">
<mods:roleTerm id="59293C66662A6E733FB415A2978E2AC2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF428CFC34DD8B7A2738EAC68C98187F">Cubelio, Sherine Sonia</mods:namePart>
<mods:affiliation id="352539AAB953BC56F9F74E2C1455CD18">Centre for Marine Living Resources &amp; Ecology, Ministry of Earth Sciences, Government of India, Atal Bhavan, LNG Terminus Road, Puthuvype, Kochi, 682508, India.</mods:affiliation>
</mods:name>
<mods:name id="13312EF070B2FDE91310395129D1BB99" type="personal">
<mods:role id="67C86A47E8B5AAAEED73153A2872F857">
<mods:roleTerm id="16148ADD7C4475781989162B352A7AE2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F298AEE991BE0AAD78BC8CA735A2A42E">Takeda, Masatsune</mods:namePart>
<mods:nameIdentifier id="7647B43E5691A120A1C6D1E938097A9F" type="ORCID">0000-0002-0028-1397</mods:nameIdentifier>
<mods:affiliation id="37356BEE13E4FEFC0AE543581287BD14">Department of Zoology, National Museum of Nature and Science, Tokyo. 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki, 305 - 0005, Japan. takeda @ kahaku. go. jp; https: // orcid. org / 0000 - 0002 - 0028 - 1397</mods:affiliation>
<mods:nameIdentifier id="340BB218206FDA5C5960F5AA2D6B4530" type="email">takeda@kahaku.go.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A5DAA44623CF088DFFED65BADFAB3077">text</mods:typeOfResource>
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<mods:titleInfo id="4BFEDBB5B53F1CD5BB9656E05B81A5A6">
<mods:title id="F614385EFF05615B4966366EEB614982">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="94203D5E74C10A83639BFF277E6F7860">
<mods:date id="A48CFD14868E536F106F194C0EE05AC7">2022</mods:date>
<mods:detail id="C7AFC3A44A1DB1A6DA545D7E67D9CA2C" type="pubDate">
<mods:number id="FD12DDDA966CBD4D118A45A0070C8DBE">2022-01-26</mods:number>
</mods:detail>
<mods:detail id="241C5E659C8CD447DAAE2833E52DBF4D" type="volume">
<mods:number id="33E077060C08D876B82893674AD95116">5093</mods:number>
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<mods:number id="7F5BAD89C814159C86A27C628FD51920">2</mods:number>
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<mods:start id="936477D524829C3C821198BC95A5F9F4">195</mods:start>
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<mods:classification id="C33EF2992BCF2C2D8B0ED0C703004EC7">journal article</mods:classification>
<mods:identifier id="72EE8AADBE877FD0411B823506E1F258" type="CLB-Dataset">2408</mods:identifier>
<mods:identifier id="BB245E556A8BC629A459B0C6BD332373" type="DOI">10.11646/zootaxa.5093.2.4</mods:identifier>
<mods:identifier id="B1B4D3B942D37D59AC1D7A8F5B55837B" type="GBIF-Dataset">a29a1307-a6a4-4198-8653-e79c13f4d008</mods:identifier>
<mods:identifier id="0281E2BC7422C045685B8BD26322553A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D16308B90A5F23445265555B10C2D22E" type="Zenodo-Dep">5905225</mods:identifier>
<mods:identifier id="B7C634EF73EF2B5E2E915ACCD4D89FB4" type="ZooBank">FA40C88B-9784-4DC0-9109-90936BAFE808</mods:identifier>
</mods:mods>
<treatment id="03FF87B3FFB34B5BFF54ABA28607C7AB" ID-DOI="http://doi.org/10.5281/zenodo.5906911" ID-GBIF-Taxon="192626888" ID-Zenodo-Dep="5906911" LSID="urn:lsid:plazi:treatment:03FF87B3FFB34B5BFF54ABA28607C7AB" httpUri="http://treatment.plazi.org/id/03FF87B3FFB34B5BFF54ABA28607C7AB" lastPageId="14" lastPageNumber="209" pageId="5" pageNumber="200">
<subSubSection id="C34C652EFFB34B50FF54ABA2873CC6CB" box="[151,514,151,178]" pageId="5" pageNumber="200" type="nomenclature">
<paragraph id="8BE936A5FFB34B50FF54ABA2873CC6CB" blockId="5.[151,960,151,250]" box="[151,514,151,178]" pageId="5" pageNumber="200">
<heading id="D0A181C9FFB34B50FF54ABA2873CC6CB" bold="true" box="[151,514,151,178]" fontSize="11" level="1" pageId="5" pageNumber="200" reason="1">
<emphasis id="B922EAB7FFB34B50FF54ABA2873CC6CB" bold="true" box="[151,514,151,178]" pageId="5" pageNumber="200">
<taxonomicName id="4C564D26FFB34B50FF54ABA284A2C6C8" ID-CoL="17f2f101-3deb-4b1d-ab7f-3a1094b63c59" authority="Padate &amp; Cubelio &amp; Takeda, 2022" authorityName="Padate &amp; Cubelio &amp; Takeda" authorityYear="2022" box="[151,412,151,178]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="keralam" status="sp. nov.">
<emphasis id="B922EAB7FFB34B50FF54ABA284A2C6C8" bold="true" box="[151,412,151,178]" italics="true" pageId="5" pageNumber="200">Guyanacaris keralam</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCFFB34B50FE67ABA2873CC6CB" box="[420,514,151,177]" pageId="5" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C652EFFB34B50FF54AB8E86FEC680" pageId="5" pageNumber="200" type="description">
<paragraph id="8BE936A5FFB34B50FF54AB8E8435C6AC" blockId="5.[151,960,151,250]" box="[151,267,187,214]" pageId="5" pageNumber="200">
(
<figureCitation id="136D2A20FFB34B50FF5CAB8E843DC6AC" box="[159,259,187,214]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="6.[151,250,1822,1846]" captionStartId-1="8.[151,250,1776,1800]" captionStartId-2="9.[151,250,1831,1855]" captionStartId-3="14.[151,250,1962,1986]" captionTargetBox-0="[163,1424,632,1786]" captionTargetBox-1="[164,1432,192,1744]" captionTargetBox-2="[157,1424,201,1786]" captionTargetBox-3="[157,1430,1254,1938]" captionTargetId-0="figure-247@6.[151,1436,624,1799]" captionTargetId-1="figure-21@8.[151,1436,181,1751]" captionTargetId-2="figure-18@9.[150,1438,181,1807]" captionTargetId-3="figure-292@14.[157,1430,1254,1938]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="14" captionText-0="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." captionText-1="FIGURE 3. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 1, lateral view; B, Right pereopod 1, mesial view; C, Left pereopod 1, lateral view; D, Left pereopod 1, mesial view. Scale bars: A–D = 10 mm." captionText-2="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." captionText-3="FIGURE 5. Paragourretia galathea (Sakai, 2017), male, CL 9.0 mm, TL 29.0 mm (CMLRE/IO/SS/AXI/00003). Photographs. A, Dorsal habitus; B, Lateral habitus. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5905229" figureDoi-1="http://doi.org/10.5281/zenodo.5905231" figureDoi-2="http://doi.org/10.5281/zenodo.5905233" figureDoi-3="http://doi.org/10.5281/zenodo.5905235" httpUri-0="https://zenodo.org/record/5905229/files/figure.png" httpUri-1="https://zenodo.org/record/5905231/files/figure.png" httpUri-2="https://zenodo.org/record/5905233/files/figure.png" httpUri-3="https://zenodo.org/record/5905235/files/figure.png" pageId="5" pageNumber="200">Figs 2–5</figureCitation>
)
</paragraph>
<paragraph id="8BE936A5FFB34B50FF54ABEA86FEC680" blockId="5.[151,960,151,250]" box="[151,960,223,250]" pageId="5" pageNumber="200">
<uri id="FFC73AA7FFB34B50FF54ABEA86FEC680" box="[151,960,223,250]" pageId="5" pageNumber="200">
urn:lsid:zoobank.org:act:
<uuid id="FFF00C70FFB34B50FE64ABEA86FEC680" box="[423,960,223,250]" pageId="5" pageNumber="200">6FE055EC-2D4E-46C7-80ED-4C30F540F940</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34C652EFFB34B50FF54AA158760C7FB" pageId="5" pageNumber="200" type="materials_examined">
<paragraph id="8BE936A5FFB34B50FF54AA158760C7FB" blockId="5.[151,1437,288,2040]" pageId="5" pageNumber="200">
<emphasis id="B922EAB7FFB34B50FF54AA158447C740" bold="true" box="[151,377,288,314]" pageId="5" pageNumber="200">Material examined</emphasis>
.
<materialsCitation id="3B3E3CF8FFB34B50FE44AA158760C7FB" ID-GBIF-Occurrence="3457616301" collectingDate="2013-08-26" collectionCode="CMLRE, IO" collectorName="Expo &amp; Dr. Rajool Shanis C. P." country="India" county="Kasaragod" elevation="321" latitude="12.1" location="Male" longLatPrecision="776" longitude="74.32" municipality="Arabian Sea" pageId="5" pageNumber="200" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<typeStatus id="54ED8807FFB34B50FE44AA1584CDC740" box="[391,499,288,314]" pageId="5" pageNumber="200" type="holotype">Holotype</typeStatus>
.
<location id="8E89607EFFB34B50FE3EAA158709C740" LSID="urn:lsid:plazi:treatment:03FF87B3FFB34B5BFF54ABA28607C7AB:8E89607EFFB34B50FE3EAA158709C740" box="[509,567,288,314]" country="India" county="Kasaragod" latitude="12.1" longLatPrecision="776" longitude="74.32" municipality="Arabian Sea" name="Male" pageId="5" pageNumber="200" stateProvince="Kerala">Male</location>
(
<collectionCode id="ED47AE60FFB34B50FD86AA148797C741" box="[581,681,289,315]" pageId="5" pageNumber="200">CMLRE</collectionCode>
/
<collectionCode id="ED47AE60FFB34B50FD72AA1487F0C741" box="[689,718,289,315]" country="Portugal" name="Instituto de Oceanografia da Universidade de Lisboa" pageId="5" pageNumber="200">IO</collectionCode>
/SS/AXI/00001, CL 50.0 mm, TL 119.0 mm),
<collectingMunicipality id="6B8DACDFFFB34B50FB11AA158060C741" box="[1234,1374,288,315]" pageId="5" pageNumber="200">Arabian Sea</collectingMunicipality>
, west off
<collectingCounty id="62884E29FFB34B50FF7DAA718407C724" box="[190,313,324,350]" pageId="5" pageNumber="200">Kasaragod</collectingCounty>
,
<collectingRegion id="4992F847FFB34B50FE87AA7184AFC724" box="[324,401,324,350]" country="India" name="Kerala" pageId="5" pageNumber="200">Kerala</collectingRegion>
,
<collectingCountry id="F3417635FFB34B50FE5EAA7184E7C724" box="[413,473,324,350]" name="India" pageId="5" pageNumber="200">India</collectingCountry>
, FORVSS station 31810,
<geoCoordinate id="EE625062FFB34B50FD3AAA718664C724" box="[761,858,324,350]" degrees="12.10" direction="north" orientation="latitude" pageId="5" pageNumber="200" precision="555" value="12.1">12.10°N</geoCoordinate>
,
<geoCoordinate id="EE625062FFB34B50FCA5AA7186FAC724" box="[870,964,324,350]" degrees="74.32" direction="east" orientation="longitude" pageId="5" pageNumber="200" precision="555" value="74.32">74.32°E</geoCoordinate>
,
<elevation id="007BD196FFB34B50FC0CAA7681A8C724" box="[975,1174,323,350]" metricMagnitude="2" metricUnit="m" metricValue="3.21" metricValueMax="3.26" metricValueMin="3.16" pageId="5" pageNumber="200" unit="m" value="321.0" valueMax="326.0" valueMin="316.0">
<quantity id="4CAE9B40FFB34B50FC0CAA768170C724" box="[975,1102,323,350]" metricMagnitude="2" metricUnit="m" metricValue="3.21" metricValueMax="3.26" metricValueMin="3.16" pageId="5" pageNumber="200" unit="m" value="321.0" valueMax="326.0" valueMin="316.0">316–326 m</quantity>
depth
</elevation>
,
<date id="FFE81065FFB34B50FB62AA768068C724" box="[1185,1366,323,350]" pageId="5" pageNumber="200" value="2013-08-26">
<collectingDate id="EFACE98DFFB34B50FB62AA768068C724" box="[1185,1366,323,350]" pageId="5" pageNumber="200" value="2013-08-26">26 August 2013</collectingDate>
</date>
,
<collectorName id="26A35373FFB34B50FAA2AA7180A2C724" box="[1377,1436,324,350]" pageId="5" pageNumber="200">Expo</collectorName>
model trawl, coll.
<collectorName id="26A35373FFB34B50FEA7AA528760C7FB" box="[356,606,359,385]" pageId="5" pageNumber="200">Dr. Rajool Shanis C.P.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34C652EFFB34B50FF04AABF874AC42E" pageId="5" pageNumber="200" type="diagnosis">
<paragraph id="8BE936A5FFB34B50FF04AABF874AC42E" blockId="5.[151,1437,288,2040]" pageId="5" pageNumber="200">
<emphasis id="B922EAB7FFB34B50FF04AABF847CC7DE" bold="true" box="[199,322,394,420]" pageId="5" pageNumber="200">Diagnosis.</emphasis>
Rostrum with 4 or 5 lateral spines. Lateral gastric carina with 5 spines; submedian gastric carina with 6 spines; median gastric carina with 3 anterior spines, 1 tubercle and 4 posterior spines. Eyestalk short of midlength of rostrum. Antennular peduncle not reaching to distal margin of antennal article 4. Antennal article 4 longer than article 2, scaphocerite overreaching midlength of article 4. Chelipeds subequal, major P1 dactylus with 6 erect dorsal spines, minor P1 dactylus with 5 erect dorsal spines; fingers at least 3 times palm length. P3–4 propodi and dactyli with corneous movable spines.
</paragraph>
</subSubSection>
<subSubSection id="C34C652EFFB34B52FF04A968860CC5DF" lastPageId="7" lastPageNumber="202" pageId="5" pageNumber="200" type="description">
<paragraph id="8BE936A5FFB34B50FF04A96880A2C5CE" blockId="5.[151,1437,288,2040]" pageId="5" pageNumber="200">
<emphasis id="B922EAB7FFB34B50FF04A96884DDC40D" bold="true" box="[199,483,605,631]" pageId="5" pageNumber="200">
Description of
<typeStatus id="54ED8807FFB34B50FEB6A96884E0C40D" box="[373,478,605,631]" pageId="5" pageNumber="200" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace minutely granular, sparsely setose (
<figureCitation id="136D2A20FFB34B50FC1DA9688164C40D" box="[990,1114,605,631]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="5" pageNumber="200">Figs 1A, C</figureCitation>
,
<figureCitation id="136D2A20FFB34B50FBA7A9688197C402" box="[1124,1193,605,632]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">2A, B</figureCitation>
). Rostrum acute, 0.19 times post-rostral carapace length, 0.45 times length of front-to-cervical groove, acute, short of base of distalmost antennular article, and midlength of penultimate antennal article (article4), bearing 4lateral spines on the right margin, 5 lateral spines on the left margin anterior to supraocular spine, weakly continuous with lateral gastric carinae (
<figureCitation id="136D2A20FFB34B50FAA9A9F285E2C57E" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="5" pageNumber="200">Figs 1A, C</figureCitation>
,
<figureCitation id="136D2A20FFB34B50FF2BA9DF846FC57F" box="[232,337,746,773]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">2A, B, D</figureCitation>
). Supraocular spine slightly shorter than rostral spines, anterolateral margin with 1 antennal spine. Lateral gastric carina bearing 5 spines; submedian gastric carina bearing 6 spines; median gastric carina bearing 3 anterior spines, 1 tubercle and 4 posterior spines; cervical groove distinct, extending anteriorly up to hepatic region, unarmed; post-cervical and branchiostegal regions unarmed, low median longitudinal post-cervical carina present (
<figureCitation id="136D2A20FFB34B50FF5CA8428420C5EB" box="[159,286,887,913]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="5" pageNumber="200">Figs 1A, C</figureCitation>
,
<figureCitation id="136D2A20FFB34B50FEE9A84284ADC5EB" box="[298,403,887,913]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">2A, B, D</figureCitation>
). Thoracic sternite 7 (P4) deeply divided in midline over posterior two-thirds, bearing sharp oblique lateral ridge; sternite 8 (P5) with setose semicircular flap on anterior face at base of P5 (
<figureCitation id="136D2A20FFB34B50FB3DA8AF8062C5CE" box="[1278,1372,922,948]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="5" pageNumber="200">Figs 1B</figureCitation>
,
<figureCitation id="136D2A20FFB34B50FAA8A8AF80AEC5CE" box="[1387,1424,922,948]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">2C</figureCitation>
).
</paragraph>
<paragraph id="8BE936A5FFB34B50FF04A8888407C21E" blockId="5.[151,1437,288,2040]" pageId="5" pageNumber="200">
Pleonal 1 pleuron twice as deep as middorsal length, bearing 1 ventral spine; pleuron 2 broad, lateral length 1.6 times dorsal length, gently convex anteroventrally, bearing 13 marginal spines ventrally; pleura 3–5 ventrally subquadrate, posterior margins almost straight, pleura 3–4 bearing 6 marginal spines each, pleuron 5 bearing 5 marginal spines; pleuron 6 bluntly triangular, bearing 1 ventral spine (
<figureCitation id="136D2A20FFB34B50FC5AAF12812AC23B" box="[921,1044,1063,1089]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="5" pageNumber="200">Figs 1A, C</figureCitation>
,
<figureCitation id="136D2A20FFB34B50FBDCAF12817BC23B" box="[1055,1093,1063,1089]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">2A</figureCitation>
); pleura 1–5 with lateral crease (
<figureCitation id="136D2A20FFB34B50FF5CAF7F85C4C21E" box="[159,250,1098,1124]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="5" pageNumber="200">Figs 1C</figureCitation>
,
<figureCitation id="136D2A20FFB34B50FEC5AF7F8412C21E" box="[262,300,1098,1124]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">2A</figureCitation>
).
</paragraph>
<paragraph id="8BE936A5FFB34B50FF04AF58800BC342" blockId="5.[151,1437,288,2040]" pageId="5" pageNumber="200">
Ocular peduncle 0.46 times rostral length; cornea lightly pigmented (
<figureCitation id="136D2A20FFB34B50FC29AF588152C2F2" box="[1002,1132,1133,1160]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">Fig. 2B, D</figureCitation>
). Antennular peduncle not reaching to distal margin of penultimate antennal article (article 4) (
<figureCitation id="136D2A20FFB34B50FC57AFA58170C2D1" box="[916,1102,1168,1195]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">Fig. 2B, D, E, F</figureCitation>
). Antennal article 1 bearing 1 spine on distoventral margin, ventral margin with 1 row of blunt spinules; article 2 distal spine (stylocerite) slender, directed anteriorly, short of midlength of antennal article 4; scaphocerite gently curved, overreaching midlength of article 4, bearing mesial spine at base; article 3 with broad mesiodistal spine on ventral margin; article 4 length 1.40 times length of article 2 (excluding distal spine); article 5 0.46 times length of article 4 (
<figureCitation id="136D2A20FFB34B50FBB2AE288016C342" box="[1137,1320,1309,1336]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">Fig. 2B, D, E, F</figureCitation>
).
</paragraph>
<paragraph id="8BE936A5FFB34B50FF04AE75867BC304" blockId="5.[151,1437,288,2040]" pageId="5" pageNumber="200">Pleurobranch gills absent; well-developed epipods and podobranch gills on maxillipeds 2–3 and pereopods 1–4; 2 arthrobranch gills each on maxilliped 3 and pereopods 1–4.</paragraph>
<paragraph id="8BE936A5FFB34B50FF04AEB28775C071" blockId="5.[151,1437,288,2040]" pageId="5" pageNumber="200">
Maxilliped 3 coxa bearing 1 spine; basis bearing 1 spine; ischium ventral margin bearing 3 spines, crista dentata with 17–21 teeth (
<figureCitation id="136D2A20FFB34B50FEA5AE9F84FCC3BE" box="[358,450,1450,1476]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">Fig. 2G</figureCitation>
); right merus ventral margin bearing 4 spines (
<figureCitation id="136D2A20FFB34B50FC14AE9F810FC3BE" box="[983,1073,1450,1476]" captionStart="FIGURE 3" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[164,1432,192,1744]" captionTargetId="figure-21@8.[151,1436,181,1751]" captionTargetPageId="8" captionText="FIGURE 3. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 1, lateral view; B, Right pereopod 1, mesial view; C, Left pereopod 1, lateral view; D, Left pereopod 1, mesial view. Scale bars: A–D = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905231" httpUri="https://zenodo.org/record/5905231/files/figure.png" pageId="5" pageNumber="200">Fig. 3G</figureCitation>
), left merus with 3 spines; right carpus ventral margin bearing 1 subdistal spine, left carpus spine damaged; exopod with well-developed flagellum, reaching distal end of merus (
<figureCitation id="136D2A20FFB34B50FE27AEC58700C071" box="[484,574,1520,1547]" captionStart="FIGURE 2" captionStartId="6.[151,250,1822,1846]" captionTargetBox="[163,1424,632,1786]" captionTargetId="figure-247@6.[151,1436,624,1799]" captionTargetPageId="6" captionText="FIGURE 2. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India.A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="5" pageNumber="200">Fig. 2H</figureCitation>
).
</paragraph>
<paragraph id="8BE936A5FFB34B50FF04AD2186FDC18D" blockId="5.[151,1437,288,2040]" pageId="5" pageNumber="200">
Chelipeds (PI) subequal in length, propodus of right P1 slightly more swollen, ornamentation slightly dissimilar. Major (right) P1 coxa bearing 1 curved spine proximally, 2 spines distally on ventral margin, lateral margin bearing 1 spine and 3 spinules; basis ventral surface bearing 1 spine, terminal spine on lateral margin behind ischial articulation; ischium ventral margin bearing 2 rows of 3–5 spines (
<figureCitation id="136D2A20FFB34B50FD3FAD488646C0E2" box="[764,888,1661,1688]" captionStart="FIGURE 3" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[164,1432,192,1744]" captionTargetId="figure-21@8.[151,1436,181,1751]" captionTargetPageId="8" captionText="FIGURE 3. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 1, lateral view; B, Right pereopod 1, mesial view; C, Left pereopod 1, lateral view; D, Left pereopod 1, mesial view. Scale bars: A–D = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905231" httpUri="https://zenodo.org/record/5905231/files/figure.png" pageId="5" pageNumber="200">Fig. 3A, B</figureCitation>
); merus dorsal margin bearing 5 spines, anterior 2 forming 1 pair with spinule at carpal articulation, outer ventral margin bearing 10 small spines and 8 spinules, inner ventral margin bearing 5 large spines, distoventral lobule with large hook-like spine, lateral surface sparsely spinose distally, mesial surface smooth (
<figureCitation id="136D2A20FFB34B50FD98ADD287EBC17B" box="[603,725,1767,1793]" captionStart="FIGURE 3" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[164,1432,192,1744]" captionTargetId="figure-21@8.[151,1436,181,1751]" captionTargetPageId="8" captionText="FIGURE 3. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 1, lateral view; B, Right pereopod 1, mesial view; C, Left pereopod 1, lateral view; D, Left pereopod 1, mesial view. Scale bars: A–D = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905231" httpUri="https://zenodo.org/record/5905231/files/figure.png" pageId="5" pageNumber="200">Fig. 3A, B</figureCitation>
); carpus dorsal surface bearing 3 spines, lateral surface bearing 12 spinules including 2 on distal margin, mesial surface bearing 1 pair of blunt spinules distally, ventral margin bearing 3 spinules distally (
<figureCitation id="136D2A20FFB34B50FE15AC18876AC132" box="[470,596,1837,1864]" captionStart="FIGURE 3" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[164,1432,192,1744]" captionTargetId="figure-21@8.[151,1436,181,1751]" captionTargetPageId="8" captionText="FIGURE 3. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 1, lateral view; B, Right pereopod 1, mesial view; C, Left pereopod 1, lateral view; D, Left pereopod 1, mesial view. Scale bars: A–D = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905231" httpUri="https://zenodo.org/record/5905231/files/figure.png" pageId="5" pageNumber="200">Fig. 3A, B</figureCitation>
); propodus dorsal margin bearing 2 irregular rows of 3–5 spines, ventral margin bearing 13 spines in lateral row, mesial surface inflated (
<figureCitation id="136D2A20FFB34B50FC9EAC65868CC111" box="[861,946,1872,1899]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="5" pageNumber="200">Fig. 1B</figureCitation>
), bearing 13 spines (anterior largest), pollex (fixed finger) inner surface bearing 5 spines dorsally, lateral surface sparsely spinose, 2 irregular rows of 3 spines each proximally; pollex 3.00 times as long as upper palm, cutting edge straight with 30 bluntly triangular teeth (
<figureCitation id="136D2A20FFB34B50FAACACA285E3C1AF" captionStart="FIGURE 3" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[164,1432,192,1744]" captionTargetId="figure-21@8.[151,1436,181,1751]" captionTargetPageId="8" captionText="FIGURE 3. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 1, lateral view; B, Right pereopod 1, mesial view; C, Left pereopod 1, lateral view; D, Left pereopod 1, mesial view. Scale bars: A–D = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905231" httpUri="https://zenodo.org/record/5905231/files/figure.png" pageId="5" pageNumber="200">Fig. 3A, B</figureCitation>
); dactylus dorsal margin with 6 spines, lateral and mesial surfaces unarmed, cutting edge bearing 30 bluntly triangular teeth (
<figureCitation id="136D2A20FFB34B50FE92ACE884F4C182" box="[337,458,2013,2040]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="5" pageNumber="200">Fig. 4A, B</figureCitation>
); both fingers heavily setose (
<figureCitation id="136D2A20FFB34B50FCD8ACE88688C18D" box="[795,950,2013,2040]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="5" pageNumber="200">Fig. 1A, B, C</figureCitation>
).
</paragraph>
<paragraph id="8BE936A5FFB04B53FF04ABA286FDC422" blockId="6.[151,1437,151,600]" pageId="6" pageNumber="201">
Minor (left) P1 coxa bearing 1 curved spine proximally, 2 spines distally on ventral margin, lateral margin bearing 1 spine and 3 spinules; basis ventral surface bearing 1 spine, terminal spine on lateral margin behind ischial articulation; ischium ventral margin bearing 2 rows of 3–5 spines; merus dorsal margin bearing 5 spines, 1 spinule at carpal articulation, outer ventral margin bearing 9 small spines and 8 spinules, inner ventral margin bearing 4 large spines, distoventral lobule with large hook-like spine, lateral surface sparsely spinose distally, mesial surface smooth; carpus dorsal surface bearing 2 spines, lateral surface bearing 11 spinules including 2 on distal margin, mesial surface bearing 1 pair of blunt spinules distally, ventral margin bearing 2 spinules distally; propodus dorsal margin bearing 2 irregular rows of 3–4 spines, ventral margin bearing 14 spines in lateral row, mesial surface gently inflated (
<figureCitation id="136D2A20FFB04B53FF3FAA85846DC7B0" box="[252,339,432,458]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="6" pageNumber="201">Fig. 1B</figureCitation>
), bearing 10 spines (anterior largest), pollex (fixed finger) inner surface bearing 3 spines dorsally, lateral surface sparsely spinose, 2 oblique rows of spines proximally, 2 spines in upper row, 5 spines in lower row; pollex 3.20 times as long as upper palm, cutting edge straight with approximately 29 bluntly triangular teeth (
<figureCitation id="136D2A20FFB04B53FAACAAC285E3C44E" captionStart="FIGURE 3" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[164,1432,192,1744]" captionTargetId="figure-21@8.[151,1436,181,1751]" captionTargetPageId="8" captionText="FIGURE 3. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 1, lateral view; B, Right pereopod 1, mesial view; C, Left pereopod 1, lateral view; D, Left pereopod 1, mesial view. Scale bars: A–D = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905231" httpUri="https://zenodo.org/record/5905231/files/figure.png" pageId="6" pageNumber="201">Fig. 3C, D</figureCitation>
); dactylus dorsal margin with 5 spines, lateral and mesial surfaces unarmed, cutting edge bearing 33 bluntly triangular teeth (
<figureCitation id="136D2A20FFB04B53FE92A90B84F5C422" box="[337,459,574,600]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="6" pageNumber="201">Fig. 4C, D</figureCitation>
); both fingers heavily setose (
<figureCitation id="136D2A20FFB04B53FCD8A90B8688C422" box="[795,950,574,600]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="6" pageNumber="201">Fig. 1A, B, C</figureCitation>
).
</paragraph>
<caption id="DF29662DFFB04B53FF54AC2B8169C1BC" ID-DOI="http://doi.org/10.5281/zenodo.5905229" ID-Zenodo-Dep="5905229" httpUri="https://zenodo.org/record/5905229/files/figure.png" pageId="6" pageNumber="201" startId="6.[151,250,1822,1846]" targetBox="[163,1424,632,1786]" targetPageId="6">
<paragraph id="8BE936A5FFB04B53FF54AC2B8169C1BC" blockId="6.[151,1436,1822,1990]" pageId="6" pageNumber="201">
<emphasis id="B922EAB7FFB04B53FF54AC2B842DC14C" bold="true" box="[151,275,1822,1846]" pageId="6" pageNumber="201">FIGURE 2.</emphasis>
<taxonomicName id="4C564D26FFB04B53FEDAAC2A84CBC14C" authority="Padate &amp; Cubelio &amp; Takeda, 2022" authorityName="Padate &amp; Cubelio &amp; Takeda" authorityYear="2022" box="[281,501,1822,1846]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="201" phylum="Arthropoda" rank="species" species="keralam" status="sp. nov.">
<emphasis id="B922EAB7FFB04B53FEDAAC2A84CBC14C" box="[281,501,1822,1846]" italics="true" pageId="6" pageNumber="201">Guyanacaris keralam</emphasis>
</taxonomicName>
<emphasis id="B922EAB7FFB04B53FE3FAC2B8772C14C" bold="true" box="[508,588,1822,1846]" pageId="6" pageNumber="201">
<taxonomicNameLabel id="A21157CCFFB04B53FE3FAC2B8772C14C" box="[508,588,1822,1846]" pageId="6" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Carapace, pleon and tail fan, lateral view; B, Carapace, left eye, left antennule and left antenna, dorsal view; C, Thoracic sternites 7 and 8; D, Rostrum, left eye, left antennule and left antenna, dorsal view; E, Left antennule and left antenna, lateral view; F, Left antennule, left antenna and epistome, ventral view; G, Right maxilliped 3 (denuded), inner view; H, Left maxilliped 3, outer view. Scale bars: A–B = 10 mm; C–F = 2 mm; G–H = 5 mm.
</paragraph>
</caption>
<paragraph id="8BE936A5FFB14B52FF04ABA28180C78F" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="202">
P2 ischium bearing 1 large subdistal and 4 proximal spinules ventrally; merus bearing 1 large subdistal and 2 smaller proximal spines ventrally; carpus 0.78 times as long as chela; propodus upper margin 0.77 length of dactylus (
<figureCitation id="136D2A20FFB14B52FF63ABEA85C4C683" box="[160,250,223,249]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4A</figureCitation>
), fingers bearing pectinate teeth on cutting margins (
<figureCitation id="136D2A20FFB14B52FC8EABEA86F6C683" box="[845,968,223,249]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4A, B</figureCitation>
). P3 ischium bearing 5 proximal spinules ventrally; merus bearing 1 distoventral spine; propodus 1.22 times as long as dactylus, lateral surface bearing transverse sets of corneous spines (
<figureCitation id="136D2A20FFB14B52FDF0AA1287AEC73B" box="[563,656,295,321]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4C</figureCitation>
); dactylus curved, lateral surface bearing 2 rows of 5–10 corneous spines, distal tip corneous, tapering to a claw (
<figureCitation id="136D2A20FFB14B52FD5AAA7E862DC71F" box="[665,787,331,357]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4C, D</figureCitation>
). P4 ischium bearing 5 proximal spinules ventrally; merus bearing 1 distoventral spine; propodus 3.04 times as long as dactylus, lateral surface bearing transverse sets of corneous spines (
<figureCitation id="136D2A20FFB14B52FE9AAAA6848CC7D7" box="[345,434,403,429]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4E</figureCitation>
); dactylus curved, lateral surface bearing 2 rows of 6–11 corneous movable spines, distal tip corneous, tapering to a claw (
<figureCitation id="136D2A20FFB14B52FE3CAA82874FC7AB" box="[511,625,439,465]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4E, F</figureCitation>
). P5 propodus 2.55 times as long as dactylus, subchelate, with short fixed finger (
<figureCitation id="136D2A20FFB14B52FF2AAAEE847AC78F" box="[233,324,475,501]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4G</figureCitation>
); dactylus unarmed, heavily setose proximally, distal tip corneous (
<figureCitation id="136D2A20FFB14B52FBF6AAEE818FC78F" box="[1077,1201,475,501]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4G, H</figureCitation>
).
</paragraph>
<paragraph id="8BE936A5FFB14B52FF04AACA8189C41B" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="202">
Pleopod 1 present, bearing 2 long setae distally (
<figureCitation id="136D2A20FFB14B52FD2FAACA8605C463" box="[748,827,511,537]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4I</figureCitation>
). Pleopod 2 endopod bearing appendix masculina and appendix interna attached at proximal one-thirds portion, appendix interna 0.33 times length of endopod, subequal to appendix masculina (
<figureCitation id="136D2A20FFB14B52FE61A97284CDC41B" box="[418,499,583,609]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4J</figureCitation>
). Pleopods 3–5 appendix interna 0.36 times length of endopod.
</paragraph>
<paragraph id="8BE936A5FFB14B52FF04A95E8439C527" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="202">
Telson 1.31 times as long as broad, lateral margin bearing 3 spines in the middle third portion, 1 spine in the posterior third portion, posterior margin broadly convex, densely setose with posteromedian spine, posterolateral angle unarmed; dorsal surface with 1 pair of submedian subcircular depressions anteriorly, 2 spines in each oblique row (
<figureCitation id="136D2A20FFB14B52FF17A9E2840FC48B" box="[212,305,727,753]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4K</figureCitation>
). Uropodal endopod 1.06 times as long as wide, with 3 lateral spines, longitudinal ridge bearing 5 spines. Uropodal exopod 1.34 times as long as wide, with 3 lateral spines, dorsal surface bearing 2 longitudinal ribs, inner rib unarmed, outer rib with 2 spines, posterolateral angle with 1 fixed spine; transverse suture with 9 spines (
<figureCitation id="136D2A20FFB14B52FF63A87685C4C527" box="[160,250,835,861]" captionStart="FIGURE 4" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[157,1424,201,1786]" captionTargetId="figure-18@9.[150,1438,181,1807]" captionTargetPageId="9" captionText="FIGURE 4. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="7" pageNumber="202">Fig. 4K</figureCitation>
).
</paragraph>
<paragraph id="8BE936A5FFB14B52FF04A852860CC5DF" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="202">
Preserved colouration (in formalin for 7 years and recently transferred to 70% ethanol) is brown with dark yellow setae; PI chelae and carpi light grey (
<figureCitation id="136D2A20FFB14B52FD4AA8BE861BC5DF" box="[649,805,907,933]" captionStart="FIGURE 1" captionStartId="4.[151,250,1805,1829]" captionTargetBox="[152,1436,181,1780]" captionTargetId="figure-21@4.[152,1436,181,1780]" captionTargetPageId="4" captionText="FIGURE 1. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. (A) Dorsal habitus; (B) Ventral habitus; (C) Lateral habitus. Scale bars: A–B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905227" httpUri="https://zenodo.org/record/5905227/files/figure.png" pageId="7" pageNumber="202">Fig. 1A, B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C652EFFB14B52FF04A89A865DC597" pageId="7" pageNumber="202" type="etymology">
<paragraph id="8BE936A5FFB14B52FF04A89A865DC597" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="202">
<emphasis id="B922EAB7FFB14B52FF04A89A8470C5B3" bold="true" box="[199,334,943,969]" pageId="7" pageNumber="202">Etymology.</emphasis>
The species name is derived from the vernacular Malayalam word “Keralam” denoting the southwestern Indian state of
<collectingRegion id="4992F847FFB14B52FE1AA8E68719C597" box="[473,551,979,1005]" country="India" name="Kerala" pageId="7" pageNumber="202">Kerala</collectingRegion>
; used as noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34C652EFFB14B5BFF04A8C285CFC7D7" lastPageId="14" lastPageNumber="209" pageId="7" pageNumber="202" type="discussion">
<paragraph id="8BE936A5FFB14B52FF04A8C281A6C293" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="202">
<emphasis id="B922EAB7FFB14B52FF04A8C28405C26B" bold="true" box="[199,315,1015,1041]" pageId="7" pageNumber="202">Remarks.</emphasis>
<bibRefCitation id="EFC74B54FFB14B52FE86A8C2876AC26B" author="Poore, G. C. B. &amp; Collins, DJ" box="[325,596,1015,1041]" pageId="7" pageNumber="202" pagination="221 - 287" refId="ref13390" refString="Poore, G. C. B. &amp; Collins, DJ. (2009) Australian Axiidae (Crustacea: Decapoda: Axiidea. Memoirs of Museum Victoria, 66 (2), 221 - 287. https: // doi. org / 10.24199 / j. mmv. 2009.66.20" type="journal article" year="2009">Poore &amp; Collins (2009)</bibRefCitation>
commented that
<taxonomicName id="4C564D26FFB14B52FCDDA8C285ECC24F" authority="Boesch &amp; Smalley, 1972" authorityName="Boesch &amp; Smalley" authorityYear="1972" class="Malacostraca" family="Axiidae" genus="Calocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutimana" subGenus="Calastacus">
<emphasis id="B922EAB7FFB14B52FCDDA8C28188C26B" box="[798,1206,1015,1041]" italics="true" pageId="7" pageNumber="202">Calocaris (Calastacus) hirsutimana</emphasis>
<bibRefCitation id="EFC74B54FFB14B52FB03A8C285ECC24F" author="Boesch, D. F. &amp; Smalley, A. E." pageId="7" pageNumber="202" pagination="45 - 52" refId="ref11230" refString="Boesch, D. F. &amp; Smalley, A. E. (1972) A new axiid (Decapoda, Thalassinidea) from the Northern Gulf of Mexico and tropical Atlantic. Bulletin of Marine Science, 22 (1), 45 - 52." type="journal article" year="1972">Boesch &amp; Smalley, 1972</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C564D26FFB14B52FF1DAF2E87E9C24F" authority="Squires, 1979" authorityName="Squires" authorityYear="1979" box="[222,727,1051,1077]" class="Malacostraca" family="Axiidae" genus="Axiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="caespitosa" subGenus="Axiopsis">
<emphasis id="B922EAB7FFB14B52FF1DAF2E870AC24E" box="[222,564,1051,1077]" italics="true" pageId="7" pageNumber="202">Axiopsis (Axiopsis) caespitosa</emphasis>
<bibRefCitation id="EFC74B54FFB14B52FDFFAF2E87E9C24F" author="Squires, H. J." box="[572,727,1051,1077]" pageId="7" pageNumber="202" pagination="1584 - 1591" refId="ref14776" refString="Squires, H. J. (1979) Axiopsis caespitosa (Thalassinidea, Axiidae), a new species from the Pacific coast of Colombia. Canadian Journal of Zoology, 57, 1584 - 1591. https: // doi. org / 10.1139 / z 79 - 207" type="journal article" year="1979">Squires, 1979</bibRefCitation>
</taxonomicName>
referred to
<taxonomicName id="4C564D26FFB14B52FC9FAF2E86D8C24F" authorityName="Sakai &amp; de Saint Laurent" authorityYear="1989" box="[860,998,1051,1077]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FC9FAF2E86D8C24F" box="[860,998,1051,1077]" italics="true" pageId="7" pageNumber="202">Acanthaxius</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC74B54FFB14B52FBD1AF2E80A9C24F" author="Sakai, K. &amp; Saint Laurent, M. de" box="[1042,1431,1051,1077]" pageId="7" pageNumber="202" pagination="1 - 104" refId="ref14618" refString="Sakai, K. &amp; Saint Laurent, M. de (1989) A check list of Axiidae (Decapoda, Crustacea, Thalassinidea, Anomura), with remarks and in addition descriptions of one new subfamily, eleven new genera and two new species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 3, 1 - 104." type="journal article" year="1989">Sakai and de Saint Laurent (1989)</bibRefCitation>
, and
<taxonomicName id="4C564D26FFB14B52FF0BAF0A874FC223" authority="Sakai, 1994" authorityName="K. Sakai" authorityYear="1994" box="[200,625,1087,1113]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="polychaetes">
<emphasis id="B922EAB7FFB14B52FF0BAF0A84E1C223" box="[200,479,1087,1113]" italics="true" pageId="7" pageNumber="202">Acanthaxius polychaetes</emphasis>
<bibRefCitation id="EFC74B54FFB14B52FE2BAF0A874FC223" author="Sakai, K." box="[488,625,1087,1113]" pageId="7" pageNumber="202" pagination="175 - 202" refId="ref14144" refString="Sakai, K. (1994) Eleven species of Australian Axiidae (Crustacea: Decapoda: Thalassinidea) with descriptions of one new genus and five new species. The Beagle, Occasional Papers of the Northern Territory Museum of Arts and Sciences, 11, 175 - 202." type="journal article" year="1994">Sakai, 1994</bibRefCitation>
</taxonomicName>
, differed from the typical
<taxonomicName id="4C564D26FFB14B52FC5FAF0A8118C223" authorityName="Sakai &amp; de Saint Laurent" authorityYear="1989" box="[924,1062,1087,1113]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FC5FAF0A8118C223" box="[924,1062,1087,1113]" italics="true" pageId="7" pageNumber="202">Acanthaxius</emphasis>
</taxonomicName>
in the presence of male pleopod 1, absence of prominent supraocular spine, comparatively more compact propodus on the major cheliped with few lateral spines, and presence of prominent lateral spine on the telson; additionally,
<taxonomicName id="4C564D26FFB14B52FBBBAFB28471C2BF" authority="(Rathbun, 1902)" baseAuthorityName="Rathbun" baseAuthorityYear="1902" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="spinulicauda">
<emphasis id="B922EAB7FFB14B52FBBBAFB280A3C2DB" box="[1144,1437,1159,1185]" italics="true" pageId="7" pageNumber="202">Acanthaxius spinulicauda</emphasis>
(
<bibRefCitation id="EFC74B54FFB14B52FF5CAF9E8478C2BF" author="Rathbun, M. J." box="[159,326,1195,1221]" pageId="7" pageNumber="202" pagination="885 - 905" refId="ref13652" refString="Rathbun, M. J. (1902) Descriptions of new decapod crustaceans from the west coast of North America. Proceedings of the United States National Museum, 24 (1272): 885 - 905. https: // doi. org / 10.5479 / si. 00963801.1272.885" type="journal article" year="1902">Rathbun, 1902</bibRefCitation>
)
</taxonomicName>
reportedly differed from the
<typeStatus id="54ED8807FFB14B52FD57AF9E87FAC2BF" box="[660,708,1195,1221]" pageId="7" pageNumber="202">type</typeStatus>
species
<taxonomicName id="4C564D26FFB14B52FCE0AF9E815CC2BF" authority="K. Sakai, 1987" authorityName="K. Sakai" authorityYear="1987" box="[803,1122,1195,1221]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="pilocheira">
<emphasis id="B922EAB7FFB14B52FCE0AF9E868BC2BF" box="[803,949,1195,1221]" italics="true" pageId="7" pageNumber="202">A. pilocheira</emphasis>
K.
<bibRefCitation id="EFC74B54FFB14B52FC1DAF9E815CC2BF" author="Sakai, K." box="[990,1122,1195,1221]" pageId="7" pageNumber="202" pagination="296 - 308" refId="ref14052" refString="Sakai, K. (1987) Two new Thalassinidea (Crustacea: Decapoda) from Japan, with the biogeographical distribution of the Japanese Thalassinidea. Bulletin of Marine Science, 41 (2), 296 - 308." type="journal article" year="1987">
Sakai,
<quantity id="4CAE9B40FFB14B52FBE9AF9E815CC2BF" box="[1066,1122,1195,1221]" metricMagnitude="1" metricUnit="m" metricValue="5.04698" pageId="7" pageNumber="202" unit="in" value="1987.0">1987</quantity>
</bibRefCitation>
</taxonomicName>
in possessing a less spinose carapace. They further suggested the creation of new genera to accommodate these species.
</paragraph>
<paragraph id="8BE936A5FFB14B52FF04AFC687CDC057" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="202">
<bibRefCitation id="EFC74B54FFB14B52FF04AFC68469C377" author="Sakai, K." box="[199,343,1267,1293]" pageId="7" pageNumber="202" pagination="1 - 616" refId="ref14380" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
retained
<taxonomicName id="4C564D26FFB14B52FE03AFC68751C376" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[448,623,1267,1293]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="spinosissima">
<emphasis id="B922EAB7FFB14B52FE03AFC68751C376" box="[448,623,1267,1293]" italics="true" pageId="7" pageNumber="202">A. spinosissima</emphasis>
</taxonomicName>
in
<taxonomicName id="4C564D26FFB14B52FD57AFC68620C377" authorityName="Sakai &amp; de Saint Laurent" authorityYear="1989" box="[660,798,1267,1293]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FD57AFC68620C377" box="[660,798,1267,1293]" italics="true" pageId="7" pageNumber="202">Acanthaxius</emphasis>
</taxonomicName>
due to the absence of male pleopod 1. He erected a new genus,
<emphasis id="B922EAB7FFB14B52FF26AE228442C34B" box="[229,380,1303,1329]" italics="true" pageId="7" pageNumber="202">
<taxonomicName id="4C564D26FFB14B52FF26AE228446C34B" authorityName="K. Sakai" authorityYear="2011" box="[229,376,1303,1329]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">Guyanacaris</taxonomicName>
,
</emphasis>
for
<taxonomicName id="4C564D26FFB14B52FE6BAE228770C34B" authorityName="Boesch &amp; Smalley" authorityYear="1972" box="[424,590,1303,1329]" class="Malacostraca" family="Axiidae" genus="Calocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutimana">
<emphasis id="B922EAB7FFB14B52FE6BAE228770C34B" box="[424,590,1303,1329]" italics="true" pageId="7" pageNumber="202">C. hirsutimana</emphasis>
</taxonomicName>
(as
<typeStatus id="54ED8807FFB14B52FDB9AE228794C34B" box="[634,682,1303,1329]" pageId="7" pageNumber="202">type</typeStatus>
species) and
<emphasis id="B922EAB7FFB14B52FCFEAE2286E9C34A" box="[829,983,1303,1329]" italics="true" pageId="7" pageNumber="202">
<taxonomicName id="4C564D26FFB14B52FCFEAE2286EDC34A" authorityName="Squires" authorityYear="1979" box="[829,979,1303,1329]" class="Malacostraca" family="Axiidae" genus="Axiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="caespitosa">A. caespitosa</taxonomicName>
,
</emphasis>
owing to an elongate, forwardly directed antennal scaphocerite and the presence of uniramous, bisegmented male pleopod 1. He erected another genus,
<taxonomicName id="4C564D26FFB14B52FF54AE6A84F8C303" authority="K. Sakai, 2011" authorityName="K. Sakai" authorityYear="2011" box="[151,454,1375,1401]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FF54AE6A842BC302" box="[151,277,1375,1400]" italics="true" pageId="7" pageNumber="202">Bruceaxius</emphasis>
K.
<bibRefCitation id="EFC74B54FFB14B52FEFCAE6A84F8C303" author="Sakai, K." box="[319,454,1375,1401]" pageId="7" pageNumber="202" pagination="1 - 616" refId="ref14380" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
, for
<taxonomicName id="4C564D26FFB14B52FE39AE6A87A3C303" authorityName="K. Sakai" authorityYear="1994" box="[506,669,1375,1401]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="polychaetes">
<emphasis id="B922EAB7FFB14B52FE39AE6A87A3C303" box="[506,669,1375,1401]" italics="true" pageId="7" pageNumber="202">A. polychaetes</emphasis>
</taxonomicName>
(as
<typeStatus id="54ED8807FFB14B52FD0EAE6A87C3C303" box="[717,765,1375,1401]" pageId="7" pageNumber="202">type</typeStatus>
species), owing to the presence of uniramous, bisegmented male pleopod 1, with a digitiform distal segment, carapace with anterolateral spine and postcervical carina, P1 chela with unarmed dorsal margin and thickly setose lateral surfaces, fingers of larger cheliped shorter than palm, those of smaller cheliped distinctly longer than palm, and pleonal 1–5 tergites and pleura separated by longitudinal carina. Moreover, he transferred
<taxonomicName id="4C564D26FFB14B52FDCBAEDA8786C073" baseAuthorityName="Rathbun" baseAuthorityYear="1902" box="[520,696,1519,1545]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="spinulicauda">
<emphasis id="B922EAB7FFB14B52FDCBAEDA8786C073" box="[520,696,1519,1545]" italics="true" pageId="7" pageNumber="202">A. spinulicauda</emphasis>
</taxonomicName>
to another genus
<taxonomicName id="4C564D26FFB14B52FC43AEDA81EAC073" authority="K. Sakai, 2011" authorityName="K. Sakai" authorityYear="2011" box="[896,1236,1519,1545]" class="Malacostraca" family="Axiidae" genus="Leonardsaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FC43AEDA811AC073" box="[896,1060,1519,1545]" italics="true" pageId="7" pageNumber="202">Leonardsaxius</emphasis>
K.
<bibRefCitation id="EFC74B54FFB14B52FB8DAEDA81EAC073" author="Sakai, K." box="[1102,1236,1519,1545]" pageId="7" pageNumber="202" pagination="1 - 616" refId="ref14380" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
, owing to smooth submedian and lateral gastric carinae on the carapace.
</paragraph>
<paragraph id="8BE936A5FFB14B52FF04AD028134C1A7" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="202">
<bibRefCitation id="EFC74B54FFB14B52FF04AD0284A3C02B" author="Dworschak, P. C." box="[199,413,1591,1617]" pageId="7" pageNumber="202" pagination="37 - 55" refId="ref11485" refString="Dworschak, P. C. (2013) Axiidea and Gebiidea (Crustacea: Decapoda) of Costa Rica. Annalen des Naturhistorischen Museums in Wien, Serie B, 115, 37 - 55." type="journal article" year="2013">Dworschak (2013)</bibRefCitation>
and
<bibRefCitation id="EFC74B54FFB14B52FE1DAD028608C02B" author="Dworschak, P. C. &amp; Poore, G. C. B." box="[478,822,1591,1617]" pageId="7" pageNumber="202" pagination="61 - 76" refId="ref11684" refString="Dworschak, P. C. &amp; Poore, G. C. B. (2018) More cautionary tales: family, generic and species synonymies of recently published taxa of ghost and mud shrimps (Decapoda: Axiidea and Gebiidea). Zootaxa, 4394 (1), 61 - 76. https: // doi. org / 10.11646 / zootaxa. 4394.1.3" type="journal article" year="2018">Dworschak and Poore (2018)</bibRefCitation>
recognized
<taxonomicName id="4C564D26FFB14B52FC0AAD028164C02B" authorityName="K. Sakai" authorityYear="2011" box="[969,1114,1591,1617]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FC0AAD028164C02B" box="[969,1114,1591,1617]" italics="true" pageId="7" pageNumber="202">Guyanacaris</emphasis>
</taxonomicName>
for
<taxonomicName id="4C564D26FFB14B52FB51AD02807EC02B" authorityName="Boesch &amp; Smalley" authorityYear="1972" box="[1170,1344,1591,1617]" class="Malacostraca" family="Axiidae" genus="Calocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutimana">
<emphasis id="B922EAB7FFB14B52FB51AD02807EC02B" box="[1170,1344,1591,1617]" italics="true" pageId="7" pageNumber="202">G. hirsutimana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C564D26FFB14B52FA42AD028433C00E" authorityName="Squires" authorityYear="1979" class="Malacostraca" family="Axiidae" genus="Axiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="caespitosa">
<emphasis id="B922EAB7FFB14B52FA42AD028433C00E" italics="true" pageId="7" pageNumber="202">G. caespitosa</emphasis>
</taxonomicName>
. Moreover,
<bibRefCitation id="EFC74B54FFB14B52FE5EAD6E87CFC00F" author="Dworschak, P. C. &amp; Poore, G. C. B." box="[413,753,1627,1653]" pageId="7" pageNumber="202" pagination="61 - 76" refId="ref11684" refString="Dworschak, P. C. &amp; Poore, G. C. B. (2018) More cautionary tales: family, generic and species synonymies of recently published taxa of ghost and mud shrimps (Decapoda: Axiidea and Gebiidea). Zootaxa, 4394 (1), 61 - 76. https: // doi. org / 10.11646 / zootaxa. 4394.1.3" type="journal article" year="2018">Dworschak and Poore (2018)</bibRefCitation>
considered
<bibRefCitation id="EFC74B54FFB14B52FCBDAD6E811DC00F" author="Sakai, K." box="[894,1059,1627,1653]" pageId="7" pageNumber="202" pagination="1 - 616" refId="ref14380" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai’s (2011)</bibRefCitation>
definition of
<taxonomicName id="4C564D26FFB14B52FB04AD6E8066C00F" authorityName="K. Sakai" authorityYear="2011" box="[1223,1368,1627,1653]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FB04AD6E8066C00F" box="[1223,1368,1627,1653]" italics="true" pageId="7" pageNumber="202">Guyanacaris</emphasis>
</taxonomicName>
to be weak.
<bibRefCitation id="EFC74B54FFB14B52FF21AD4A844AC0E3" author="Sakai, K." box="[226,372,1662,1689]" pageId="7" pageNumber="202" pagination="867 - 880" refId="ref14424" refString="Sakai, K. (2015) Two new species, Colemanaxius andamanensis sp. nov. and Bruceaxius thailandensis sp. nov. from the Andaman Sea, Thailand (infraorder Axiidea de Saint Laurent, 1979). Crustaceana, 88 (7 - 8), 867 - 880. http: // dx. doi. org / 10.1163 / 15685403 - 00003440" type="journal article" year="2015">Sakai (2015)</bibRefCitation>
described a second species of
<emphasis id="B922EAB7FFB14B52FD16AD4A8127C0E3" box="[725,1049,1663,1689]" italics="true" pageId="7" pageNumber="202">
<taxonomicName id="4C564D26FFB14B52FD16AD4A8668C0E2" authorityName="K. Sakai" authorityYear="2011" box="[725,854,1663,1688]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">Bruceaxius</taxonomicName>
,
<taxonomicName id="4C564D26FFB14B52FCA0AD4A8127C0E3" authorityName="K. Sakai" authorityYear="2015" box="[867,1049,1663,1689]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="thailandensis">B. thailandensis</taxonomicName>
</emphasis>
.
<bibRefCitation id="EFC74B54FFB14B52FBEAAD4A8180C0E3" author="Poore, G. C. B." box="[1065,1214,1663,1689]" pageId="7" pageNumber="202" pagination="259 - 367" refId="ref13306" refString="Poore, G. C. B. (2020) Axiid and micheleid lobsters from Indo-West Pacific deep-sea environments (Crustacea: Decapoda: Axiidea: Axiidae, Micheleidae). In: Corbari, L., Ahyong, S. T. &amp; Chan, T. - Y. (Eds), Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos 31. Memoires du Museum national d'Histoire naturelle, Paris, 213, 259 - 367." type="journal article" year="2020">Poore (2020)</bibRefCitation>
commented that
<taxonomicName id="4C564D26FFB14B52FA47AD4A84F2C0C7" authority="K. Sakai, 1994" authorityName="K. Sakai" authorityYear="1994" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="polychaetes">
<emphasis id="B922EAB7FFB14B52FA47AD4A8425C0C7" italics="true" pageId="7" pageNumber="202">A. polychaetes</emphasis>
K.
<bibRefCitation id="EFC74B54FFB14B52FE84AD9684F2C0C7" author="Sakai, K." box="[327,460,1699,1725]" pageId="7" pageNumber="202" pagination="175 - 202" refId="ref14144" refString="Sakai, K. (1994) Eleven species of Australian Axiidae (Crustacea: Decapoda: Thalassinidea) with descriptions of one new genus and five new species. The Beagle, Occasional Papers of the Northern Territory Museum of Arts and Sciences, 11, 175 - 202." type="journal article" year="1994">Sakai, 1994</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C564D26FFB14B52FDC6AD96864BC0C4" authority="(Rathbun, 1906)" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[517,885,1699,1726]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="spinosissima">
<emphasis id="B922EAB7FFB14B52FDC6AD96878AC0C6" box="[517,692,1699,1725]" italics="true" pageId="7" pageNumber="202">A. spinosissima</emphasis>
(
<bibRefCitation id="EFC74B54FFB14B52FD07AD968653C0C4" author="Rathbun, M. J." box="[708,877,1699,1726]" pageId="7" pageNumber="202" pagination="827 - 930" refId="ref13703" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. I - XIV." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</taxonomicName>
probably belonged to
<taxonomicName id="4C564D26FFB14B52FBB6AD968038C0C7" authorityName="K. Sakai" authorityYear="2011" box="[1141,1286,1699,1725]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FBB6AD968038C0C7" box="[1141,1286,1699,1725]" italics="true" pageId="7" pageNumber="202">Guyanacaris</emphasis>
</taxonomicName>
owing to the presence of multi-spinate lateral gastric carinae converging anteriorly on small supraocular spines, and a mesial spine at the base of scaphocerite. If
<emphasis id="B922EAB7FFB14B52FDE8ADDE87EBC17F" box="[555,725,1771,1797]" italics="true" pageId="7" pageNumber="202">
<taxonomicName id="4C564D26FFB14B52FDE8ADDE87EFC17F" authorityName="K. Sakai" authorityYear="1994" box="[555,721,1771,1797]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="polychaetes">A. polychaetes</taxonomicName>
,
</emphasis>
<typeStatus id="54ED8807FFB14B52FD1EADDE8633C17F" box="[733,781,1771,1797]" pageId="7" pageNumber="202">type</typeStatus>
species of
<emphasis id="B922EAB7FFB14B52FC48ADDE812EC17E" box="[907,1040,1771,1796]" italics="true" pageId="7" pageNumber="202">
<taxonomicName id="4C564D26FFB14B52FC48ADDE8132C17E" authorityName="K. Sakai" authorityYear="2011" box="[907,1036,1771,1796]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">Bruceaxius</taxonomicName>
,
</emphasis>
is considered to be a
<emphasis id="B922EAB7FFB14B52FAC6ADDE80A2C17F" box="[1285,1436,1771,1797]" italics="true" pageId="7" pageNumber="202">
<taxonomicName id="4C564D26FFB14B52FAC6ADDE80A6C17F" authorityName="K. Sakai" authorityYear="2011" box="[1285,1432,1771,1797]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">Guyanacaris</taxonomicName>
,
</emphasis>
then
<taxonomicName id="4C564D26FFB14B52FF13AC3A8470C152" authorityName="K. Sakai" authorityYear="2011" box="[208,334,1807,1832]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FF13AC3A8470C152" box="[208,334,1807,1832]" italics="true" pageId="7" pageNumber="202">Bruceaxius</emphasis>
</taxonomicName>
should be considered to be a synonym of
<taxonomicName id="4C564D26FFB14B52FCFBAC3A86F7C153" authorityName="K. Sakai" authorityYear="2011" box="[824,969,1807,1833]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FCFBAC3A86F7C153" box="[824,969,1807,1833]" italics="true" pageId="7" pageNumber="202">Guyanacaris</emphasis>
</taxonomicName>
. With regards to
<taxonomicName id="4C564D26FFB14B52FB51AC3A8076C153" authorityName="K. Sakai" authorityYear="2015" box="[1170,1352,1807,1833]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B922EAB7FFB14B52FB51AC3A8076C153" box="[1170,1352,1807,1833]" italics="true" pageId="7" pageNumber="202">B. thailandensis</emphasis>
</taxonomicName>
, it was noted that the line illustration of the antennal peduncle (
<bibRefCitation id="EFC74B54FFB14B52FCC3AC0686BCC137" author="Sakai, K." box="[768,898,1842,1869]" pageId="7" pageNumber="202" pagination="867 - 880" refId="ref14424" refString="Sakai, K. (2015) Two new species, Colemanaxius andamanensis sp. nov. and Bruceaxius thailandensis sp. nov. from the Andaman Sea, Thailand (infraorder Axiidea de Saint Laurent, 1979). Crustaceana, 88 (7 - 8), 867 - 880. http: // dx. doi. org / 10.1163 / 15685403 - 00003440" type="journal article" year="2015">Sakai 2015</bibRefCitation>
:
<figureCitation id="136D2A20FFB14B52FC4FAC0686DCC137" box="[908,994,1843,1869]" captionStart="FIGURE 3" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[164,1432,192,1744]" captionTargetId="figure-21@8.[151,1436,181,1751]" captionTargetPageId="8" captionText="FIGURE 3. Guyanacaris keralam sp. nov., holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 1, lateral view; B, Right pereopod 1, mesial view; C, Left pereopod 1, lateral view; D, Left pereopod 1, mesial view. Scale bars: A–D = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5905231" httpUri="https://zenodo.org/record/5905231/files/figure.png" pageId="7" pageNumber="202">Fig. 3A</figureCitation>
) did not clearly indicate the presence of a mesial spine at the base of scaphocerite. In view of this, it is herein considered that
<taxonomicName id="4C564D26FFB14B52FB9AAC628030C10B" authorityName="K. Sakai" authorityYear="2015" box="[1113,1294,1879,1905]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B922EAB7FFB14B52FB9AAC628030C10B" box="[1113,1294,1879,1905]" italics="true" pageId="7" pageNumber="202">B. thailandensis</emphasis>
</taxonomicName>
differs from the typical
<taxonomicName id="4C564D26FFB14B52FED0AC4E849AC1EF" authorityName="K. Sakai" authorityYear="2011" box="[275,420,1915,1941]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FED0AC4E849AC1EF" box="[275,420,1915,1941]" italics="true" pageId="7" pageNumber="202">Guyanacaris</emphasis>
</taxonomicName>
in the absence of a mesial spine at the base of scaphocerite, major P1 carpus, propodus and dactylus with tuberculate upper margins, male pleopod 2 with tri-segmented appendix masculina (
<tableCitation id="C6D4031EFFB14B52FB3DACAA8066C1C3" box="[1278,1368,1951,1977]" captionStart="TABLE 2" captionStartId="10.[151,239,466,492]" captionTargetBox="[165,2009,506,1696]" captionTargetPageId="10" captionText="TABLE 2. Morphological comparison among the species of Guyanacaris K. Sakai, 2011 and Bruceaxius K. Sakai, 2011." httpUri="http://table.plazi.org/id/DF29662DFFBC4B5FFF54AAE7809BC796" pageId="7" pageNumber="202" tableUuid="DF29662DFFBC4B5FFF54AAE7809BC796">Table 2</tableCitation>
). It is therefore suggested that this species may be tentatively retained in
<taxonomicName id="4C564D26FFB14B52FC46ACF6813DC1A6" authorityName="K. Sakai" authorityYear="2011" box="[901,1027,1987,2012]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB14B52FC46ACF6813DC1A6" box="[901,1027,1987,2012]" italics="true" pageId="7" pageNumber="202">Bruceaxius</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF29662DFFBE4B5DFF54ADC58608C12A" ID-DOI="http://doi.org/10.5281/zenodo.5905231" ID-Zenodo-Dep="5905231" httpUri="https://zenodo.org/record/5905231/files/figure.png" pageId="8" pageNumber="203" startId="8.[151,250,1776,1800]" targetBox="[164,1432,192,1744]" targetPageId="8">
<paragraph id="8BE936A5FFBE4B5DFF54ADC58608C12A" blockId="8.[151,1436,1776,1872]" pageId="8" pageNumber="203">
<emphasis id="B922EAB7FFBE4B5DFF54ADC5842DC172" bold="true" box="[151,275,1776,1800]" pageId="8" pageNumber="203">FIGURE 3.</emphasis>
<taxonomicName id="4C564D26FFBE4B5DFEDAADC484CBC172" authority="Padate &amp; Cubelio &amp; Takeda, 2022" authorityName="Padate &amp; Cubelio &amp; Takeda" authorityYear="2022" box="[281,501,1776,1800]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="203" phylum="Arthropoda" rank="species" species="keralam" status="sp. nov.">
<emphasis id="B922EAB7FFBE4B5DFEDAADC484CBC172" box="[281,501,1776,1800]" italics="true" pageId="8" pageNumber="203">Guyanacaris keralam</emphasis>
</taxonomicName>
<emphasis id="B922EAB7FFBE4B5DFE3FADC58772C172" bold="true" box="[508,588,1776,1800]" pageId="8" pageNumber="203">
<taxonomicNameLabel id="A21157CCFFBE4B5DFE3FADC58772C172" box="[508,588,1776,1800]" pageId="8" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 1, lateral view; B, Right pereopod 1, mesial view; C, Left pereopod 1, lateral view; D, Left pereopod 1, mesial view. Scale bars: A–D = 10 mm.
</paragraph>
</caption>
<caption id="DF29662DFFBF4B5CFF54AC128172C188" ID-DOI="http://doi.org/10.5281/zenodo.5905233" ID-Zenodo-Dep="5905233" httpUri="https://zenodo.org/record/5905233/files/figure.png" pageId="9" pageNumber="204" startId="9.[151,250,1831,1855]" targetBox="[157,1424,201,1786]" targetPageId="9">
<paragraph id="8BE936A5FFBF4B5CFF54AC128172C188" blockId="9.[151,1436,1831,2035]" pageId="9" pageNumber="204">
<emphasis id="B922EAB7FFBF4B5CFF54AC12842DC145" bold="true" box="[151,275,1831,1855]" pageId="9" pageNumber="204">FIGURE 4.</emphasis>
<taxonomicName id="4C564D26FFBF4B5CFEDAAC1284CBC145" authority="Padate &amp; Cubelio &amp; Takeda, 2022" authorityName="Padate &amp; Cubelio &amp; Takeda" authorityYear="2022" box="[281,501,1831,1855]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="204" phylum="Arthropoda" rank="species" species="keralam" status="sp. nov.">
<emphasis id="B922EAB7FFBF4B5CFEDAAC1284CBC145" box="[281,501,1831,1855]" italics="true" pageId="9" pageNumber="204">Guyanacaris keralam</emphasis>
</taxonomicName>
<emphasis id="B922EAB7FFBF4B5CFE3FAC128772C145" bold="true" box="[508,588,1831,1855]" pageId="9" pageNumber="204">
<taxonomicNameLabel id="A21157CCFFBF4B5CFE3FAC128772C145" box="[508,588,1831,1855]" pageId="9" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, CL 50.6 mm, TL 119.0 mm (CMLRE/IO/SS/AXI/00001) from the Arabian Sea off Kerala, India. A, Right pereopod 2, lateral view; B, Right Pereopod 2 chela, lateral view; C, Right pereopod 3, lateral view; D, Right pereopod 3 dactylus, sub-dorsal view; E, Right pereopod 4, lateral view; F, Right pereopod 4 dactylus, lateral view; G, Right pereopod 5, lateral view; H, Right pereopod 5 dactylus, lateral view; I, male pleopods 1, posterior view; J, Left pleopod 2 exopod, endopod, appendix interna and appendix masculine, anterior view; K, pleomere 6, telson and right uropods, dorsal view. Scale bars: K = 10 mm; A–C, E, G = 5 mm; F, H–J = 2 mm; D = 1 mm.
</paragraph>
</caption>
<caption id="DF29662DFFBC4B5FFF54AAE7809BC796" ID-Table-UUID="DF29662DFFBC4B5FFF54AAE7809BC796" box="[151,1445,466,492]" httpUri="http://table.plazi.org/id/DF29662DFFBC4B5FFF54AAE7809BC796" pageId="10" pageNumber="205" startId="10.[151,239,466,492]" targetBox="[165,2009,506,1696]" targetIsTable="true" targetPageId="10">
<paragraph id="8BE936A5FFBC4B5FFF54AAE7809BC796" blockId="10.[151,1445,466,492]" box="[151,1445,466,492]" pageId="10" pageNumber="205">
<emphasis id="B922EAB7FFBC4B5FFF54AAE78434C796" bold="true" box="[151,266,466,492]" pageId="10" pageNumber="205">TABLE 2.</emphasis>
Morphological comparison among the species of
<taxonomicName id="4C564D26FFBC4B5FFCE3AAE78170C796" authority="K. Sakai, 2011" authorityName="K. Sakai" authorityYear="2011" box="[800,1102,466,492]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFBC4B5FFCE3AAE78694C796" box="[800,938,466,492]" italics="true" pageId="10" pageNumber="205">Guyanacaris</emphasis>
K.
<bibRefCitation id="EFC74B54FFBC4B5FFC12AAE78170C796" author="Sakai, K." box="[977,1102,466,492]" pageId="10" pageNumber="205" pagination="1 - 616" refId="ref14380" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C564D26FFBC4B5FFB41AAE7809CC796" authority="K. Sakai, 2011" authorityName="K. Sakai" authorityYear="2011" box="[1154,1442,466,492]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFBC4B5FFB41AAE781C4C796" box="[1154,1274,466,492]" italics="true" pageId="10" pageNumber="205">Bruceaxius</emphasis>
K.
<bibRefCitation id="EFC74B54FFBC4B5FFAE2AAE7809CC796" author="Sakai, K." box="[1313,1442,466,492]" pageId="10" pageNumber="205" pagination="1 - 616" refId="ref14380" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE936A5FFBC4B5FFF66AACF82A0C0DA" pageId="10" pageNumber="205">
<table id="F956C405FFBCB4AAFF66AACF82E7C0DA" box="[165,2009,506,1696]" colsContinueIn="11.[151,1996,661,1529]" gridcols="4" gridrows="21" pageId="10" pageNumber="205">
<tr id="356634E7FFBCB4AAFF66AACF82E7C469" box="[165,2009,506,531]" gridrow="0" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AACF876BC469" box="[165,597,506,531]" gridcol="0" gridrow="0" pageId="10" pageNumber="205">
<emphasis id="B922EAB7FFBC4B5FFF66AACF844BC469" bold="true" box="[165,373,506,531]" pageId="10" pageNumber="205">Characters/Species</emphasis>
</th>
<th id="76B75D9BFFBCB4AAFD47AACF8106C469" box="[644,1080,506,531]" gridcol="1" gridrow="0" pageId="10" pageNumber="205">
<taxonomicName id="4C564D26FFBC4B5FFD47AACF868BC469" authority="(Squires, 1979)" baseAuthorityName="Squires" baseAuthorityYear="1979" box="[644,949,506,531]" class="Malacostraca" family="Axiidae" genus="Axiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="caespitosa">
<emphasis id="B922EAB7FFBC4B5FFD47AACF868BC469" bold="true" box="[644,949,506,531]" pageId="10" pageNumber="205">
<emphasis id="B922EAB7FFBC4B5FFD47AACF8630C469" bold="true" box="[644,782,506,531]" italics="true" pageId="10" pageNumber="205">G. caespitosa</emphasis>
(
<bibRefCitation id="EFC74B54FFBC4B5FFCD8AACF8690C469" author="Squires, H. J." box="[795,942,506,531]" pageId="10" pageNumber="205" pagination="1584 - 1591" refId="ref14776" refString="Squires, H. J. (1979) Axiopsis caespitosa (Thalassinidea, Axiidae), a new species from the Pacific coast of Colombia. Canadian Journal of Zoology, 57, 1584 - 1591. https: // doi. org / 10.1139 / z 79 - 207" type="journal article" year="1979">Squires, 1979</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="76B75D9BFFBCB4AAFB9BAACF832EC469" box="[1112,1552,506,531]" gridcol="2" gridrow="0" pageId="10" pageNumber="205">
<taxonomicName id="4C564D26FFBC4B5FFB9BAACF832EC469" authority="(Boesch &amp; Smalley, 1972)" baseAuthorityName="Boesch &amp; Smalley" baseAuthorityYear="1972" box="[1112,1552,506,531]" class="Malacostraca" family="Axiidae" genus="Calocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="hirsutimana">
<emphasis id="B922EAB7FFBC4B5FFB9BAACF832EC469" bold="true" box="[1112,1552,506,531]" pageId="10" pageNumber="205">
<emphasis id="B922EAB7FFBC4B5FFB9BAACF81C9C469" bold="true" box="[1112,1271,506,531]" italics="true" pageId="10" pageNumber="205">G. hirsutimana</emphasis>
(
<bibRefCitation id="EFC74B54FFBC4B5FFAC5AACF8337C469" author="Boesch, D. F. &amp; Smalley, A. E." box="[1286,1545,506,531]" pageId="10" pageNumber="205" pagination="45 - 52" refId="ref11230" refString="Boesch, D. F. &amp; Smalley, A. E. (1972) A new axiid (Decapoda, Thalassinidea) from the Northern Gulf of Mexico and tropical Atlantic. Bulletin of Marine Science, 22 (1), 45 - 52." type="journal article" year="1972">Boesch &amp; Smalley, 1972</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="76B75D9BFFBCB4AAF9EEAACF82E7C469" box="[1581,2009,506,531]" gridcol="3" gridrow="0" pageId="10" pageNumber="205">
<emphasis id="B922EAB7FFBC4B5FF9EEAACF83C6C469" bold="true" box="[1581,1784,506,531]" pageId="10" pageNumber="205">
<taxonomicName id="4C564D26FFBC4B5FF9EEAACF839CC469" authority="Padate &amp; Cubelio &amp; Takeda, 2022" authorityName="Padate &amp; Cubelio &amp; Takeda" authorityYear="2022" box="[1581,1698,506,531]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="205" phylum="Arthropoda" rank="species" species="keralam" status="sp. nov.">
<emphasis id="B922EAB7FFBC4B5FF9EEAACF839CC469" bold="true" box="[1581,1698,506,531]" italics="true" pageId="10" pageNumber="205">G. keralam</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCFFBC4B5FF96BAACF83C6C469" box="[1704,1784,506,531]" pageId="10" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="356634E7FFBCB4AAFF66A91782E7C425" box="[165,2009,546,607]" gridrow="1" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66A917876BC425" box="[165,597,546,607]" gridcol="0" gridrow="1" pageId="10" pageNumber="205">Rostrum</th>
<td id="76B75D9BFFBCB4AAFD47A9178106C425" box="[644,1080,546,607]" gridcol="1" gridrow="1" pageId="10" pageNumber="205">overreaching base of penultimate anten- nular article</td>
<td id="76B75D9BFFBCB4AAFB9BA917832EC425" box="[1112,1552,546,607]" gridcol="2" gridrow="1" pageId="10" pageNumber="205">not reaching base of penultimate anten- nular article</td>
<td id="76B75D9BFFBCB4AAF9EEA91782E7C425" box="[1581,2009,546,607]" gridcol="3" gridrow="1" pageId="10" pageNumber="205">not reaching base of distalmost antennular article</td>
</tr>
<tr id="356634E7FFBCB4AAFF66A95A82E7C4F2" box="[165,2009,623,648]" gridrow="2" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66A95A876BC4F2" box="[165,597,623,648]" gridcol="0" gridrow="2" pageId="10" pageNumber="205">Rostral lateral armature</th>
<td id="76B75D9BFFBCB4AAFD47A95A8106C4F2" box="[644,1080,623,648]" gridcol="1" gridrow="2" pageId="10" pageNumber="205">3 pairs of spines or 3–5 spinules</td>
<td id="76B75D9BFFBCB4AAFB9BA95A832EC4F2" box="[1112,1552,623,648]" gridcol="2" gridrow="2" pageId="10" pageNumber="205">5 pairs of spines</td>
<td id="76B75D9BFFBCB4AAF9EEA95A82E7C4F2" box="[1581,2009,623,648]" gridcol="3" gridrow="2" pageId="10" pageNumber="205">4–5 spines</td>
</tr>
<tr id="356634E7FFBCB4AAFF66A9A282E7C4AE" box="[165,2009,663,724]" gridrow="3" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66A9A2876BC4AE" box="[165,597,663,724]" gridcol="0" gridrow="3" pageId="10" pageNumber="205">Carapace (excluding gastric carinae) orna- mentation</th>
<td id="76B75D9BFFBCB4AAFD47A9A28106C4AE" box="[644,1080,663,724]" gridcol="1" gridrow="3" pageId="10" pageNumber="205">1 antennal spine</td>
<td id="76B75D9BFFBCB4AAFB9BA9A2832EC4AE" box="[1112,1552,663,724]" gridcol="2" gridrow="3" pageId="10" pageNumber="205">1 antennal spine</td>
<td id="76B75D9BFFBCB4AAF9EEA9A282E7C4AE" box="[1581,2009,663,724]" gridcol="3" gridrow="3" pageId="10" pageNumber="205">1 antennal spine</td>
</tr>
<tr id="356634E7FFBCB4AAFF66A9D682E7C486" box="[165,2009,739,764]" gridrow="4" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66A9D6876BC486" box="[165,597,739,764]" gridcol="0" gridrow="4" pageId="10" pageNumber="205">Carapace, lateral gastric carina</th>
<td id="76B75D9BFFBCB4AAFD47A9D68106C486" box="[644,1080,739,764]" gridcol="1" gridrow="4" pageId="10" pageNumber="205">3–4 spines or 6–8 spines</td>
<td id="76B75D9BFFBCB4AAFB9BA9D6832EC486" box="[1112,1552,739,764]" gridcol="2" gridrow="4" pageId="10" pageNumber="205">6–7 spines</td>
<td id="76B75D9BFFBCB4AAF9EEA9D682E7C486" box="[1581,2009,739,764]" gridcol="3" gridrow="4" pageId="10" pageNumber="205">5 spines</td>
</tr>
<tr id="356634E7FFBCB4AAFF66A83E82E7C532" box="[165,2009,779,840]" gridrow="5" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66A83E876BC532" box="[165,597,779,840]" gridcol="0" gridrow="5" pageId="10" pageNumber="205">Carapace, submedian gastric carina arma- ture</th>
<td id="76B75D9BFFBCB4AAFD47A83E8106C532" box="[644,1080,779,840]" gridcol="1" gridrow="5" pageId="10" pageNumber="205">4–5 spines or 5–7 spines</td>
<td id="76B75D9BFFBCB4AAFB9BA83E832EC532" box="[1112,1552,779,840]" gridcol="2" gridrow="5" pageId="10" pageNumber="205">5 spines</td>
<td id="76B75D9BFFBCB4AAF9EEA83E82E7C532" box="[1581,2009,779,840]" gridcol="3" gridrow="5" pageId="10" pageNumber="205">6 spines</td>
</tr>
<tr id="356634E7FFBCB4AAFF66A86D82E7C5EF" box="[165,2009,856,917]" gridrow="6" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66A86D876BC5EF" box="[165,597,856,917]" gridcol="0" gridrow="6" pageId="10" pageNumber="205">Carapace, median gastric carinae armature</th>
<td id="76B75D9BFFBCB4AAFD47A86D8106C5EF" box="[644,1080,856,917]" gridcol="1" gridrow="6" pageId="10" pageNumber="205">3 anterior spines followed by 1 tubercle, or 13 spines</td>
<td id="76B75D9BFFBCB4AAFB9BA86D832EC5EF" box="[1112,1552,856,917]" gridcol="2" gridrow="6" pageId="10" pageNumber="205">8 spines</td>
<td id="76B75D9BFFBCB4AAF9EEA86D82E7C5EF" box="[1581,2009,856,917]" gridcol="3" gridrow="6" pageId="10" pageNumber="205">3 anterior spines followed by 1 tubercle and 4 posterior spines</td>
</tr>
<tr id="356634E7FFBCB4AAFF66A89182E7C5C7" box="[165,2009,932,957]" gridrow="7" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66A891876BC5C7" box="[165,597,932,957]" gridcol="0" gridrow="7" pageId="10" pageNumber="205">Carapace, cervical groove armature</th>
<td id="76B75D9BFFBCB4AAFD47A8918106C5C7" box="[644,1080,932,957]" gridcol="1" gridrow="7" pageId="10" pageNumber="205">3 spines</td>
<td id="76B75D9BFFBCB4AAFB9BA891832EC5C7" box="[1112,1552,932,957]" gridcol="2" gridrow="7" pageId="10" pageNumber="205">smooth</td>
<td id="76B75D9BFFBCB4AAF9EEA89182E7C5C7" box="[1581,2009,932,957]" gridcol="3" gridrow="7" pageId="10" pageNumber="205">smooth</td>
</tr>
<tr id="356634E7FFBCB4AAFF66A8F982E7C59F" box="[165,2009,972,997]" gridrow="8" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66A8F9876BC59F" box="[165,597,972,997]" gridcol="0" gridrow="8" pageId="10" pageNumber="205">Pleuron 1</th>
<td id="76B75D9BFFBCB4AAFD47A8F98106C59F" box="[644,1080,972,997]" gridcol="1" gridrow="8" pageId="10" pageNumber="205">acute</td>
<td id="76B75D9BFFBCB4AAFB9BA8F9832EC59F" box="[1112,1552,972,997]" gridcol="2" gridrow="8" pageId="10" pageNumber="205">acute</td>
<td id="76B75D9BFFBCB4AAF9EEA8F982E7C59F" box="[1581,2009,972,997]" gridcol="3" gridrow="8" pageId="10" pageNumber="205">acute</td>
</tr>
<tr id="356634E7FFBCB4AAFF66A8C182E7C24B" box="[165,2009,1012,1073]" gridrow="9" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66A8C1876BC24B" box="[165,597,1012,1073]" gridcol="0" gridrow="9" pageId="10" pageNumber="205">Pleura 1–5, marginal armature</th>
<td id="76B75D9BFFBCB4AAFD47A8C18106C24B" box="[644,1080,1012,1073]" gridcol="1" gridrow="9" pageId="10" pageNumber="205">with several spines</td>
<td id="76B75D9BFFBCB4AAFB9BA8C1832EC24B" box="[1112,1552,1012,1073]" gridcol="2" gridrow="9" pageId="10" pageNumber="205">with several spines</td>
<td id="76B75D9BFFBCB4AAF9EEA8C182E7C24B" box="[1581,2009,1012,1073]" gridcol="3" gridrow="9" pageId="10" pageNumber="205">with 1, 13, 6, 6, 5 and 1 spines, respec- tively</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AF7482E7C220" box="[165,2009,1089,1114]" gridrow="10" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AF74876BC220" box="[165,597,1089,1114]" gridcol="0" gridrow="10" pageId="10" pageNumber="205">Ocular peduncle</th>
<td id="76B75D9BFFBCB4AAFD47AF748106C220" box="[644,1080,1089,1114]" gridcol="1" gridrow="10" pageId="10" pageNumber="205">overreaching distal tip of rostrum</td>
<td id="76B75D9BFFBCB4AAFB9BAF74832EC220" box="[1112,1552,1089,1114]" gridcol="2" gridrow="10" pageId="10" pageNumber="205">overreaching distal tip of rostrum</td>
<td id="76B75D9BFFBCB4AAF9EEAF7482E7C220" box="[1581,2009,1089,1114]" gridcol="3" gridrow="10" pageId="10" pageNumber="205">not reaching midlength of rostrum</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AF5C82E7C2DC" box="[165,2009,1129,1190]" gridrow="11" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AF5C876BC2DC" box="[165,597,1129,1190]" gridcol="0" gridrow="11" pageId="10" pageNumber="205">Antennular peduncle</th>
<td id="76B75D9BFFBCB4AAFD47AF5C8106C2DC" box="[644,1080,1129,1190]" gridcol="1" gridrow="11" pageId="10" pageNumber="205">reaching base of penultimate antennal article</td>
<td id="76B75D9BFFBCB4AAFB9BAF5C832EC2DC" box="[1112,1552,1129,1190]" gridcol="2" gridrow="11" pageId="10" pageNumber="205">not reaching midlength of penultimate antennal article</td>
<td id="76B75D9BFFBCB4AAF9EEAF5C82E7C2DC" box="[1581,2009,1129,1190]" gridcol="3" gridrow="11" pageId="10" pageNumber="205">not reaching distal margin of penultimate antennal article</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AF8082E7C2B4" box="[165,2009,1205,1230]" gridrow="12" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AF80876BC2B4" box="[165,597,1205,1230]" gridcol="0" gridrow="12" pageId="10" pageNumber="205">Antennal article 2, distal spine orientation</th>
<td id="76B75D9BFFBCB4AAFD47AF808106C2B4" box="[644,1080,1205,1230]" gridcol="1" gridrow="12" pageId="10" pageNumber="205">anterior</td>
<td id="76B75D9BFFBCB4AAFB9BAF80832EC2B4" box="[1112,1552,1205,1230]" gridcol="2" gridrow="12" pageId="10" pageNumber="205">anterior</td>
<td id="76B75D9BFFBCB4AAF9EEAF8082E7C2B4" box="[1581,2009,1205,1230]" gridcol="3" gridrow="12" pageId="10" pageNumber="205">anterior</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AFE882E7C360" box="[165,2009,1245,1306]" gridrow="13" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AFE8876BC360" box="[165,597,1245,1306]" gridcol="0" gridrow="13" pageId="10" pageNumber="205">Scaphocerite</th>
<td id="76B75D9BFFBCB4AAFD47AFE88106C360" box="[644,1080,1245,1306]" gridcol="1" gridrow="13" pageId="10" pageNumber="205">not reaching distal margin of penultimate antennal article</td>
<td id="76B75D9BFFBCB4AAFB9BAFE8832EC360" box="[1112,1552,1245,1306]" gridcol="2" gridrow="13" pageId="10" pageNumber="205">not reaching distal margin of penultimate antennal article</td>
<td id="76B75D9BFFBCB4AAF9EEAFE882E7C360" box="[1581,2009,1245,1306]" gridcol="3" gridrow="13" pageId="10" pageNumber="205">not reaching distal margin of penultimate antennal article</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AE1F82E7C339" box="[165,2009,1322,1347]" gridrow="14" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AE1F876BC339" box="[165,597,1322,1347]" gridcol="0" gridrow="14" pageId="10" pageNumber="205">Scaphocerite, mesial spine at base</th>
<td id="76B75D9BFFBCB4AAFD47AE1F8106C339" box="[644,1080,1322,1347]" gridcol="1" gridrow="14" pageId="10" pageNumber="205">present</td>
<td id="76B75D9BFFBCB4AAFB9BAE1F832EC339" box="[1112,1552,1322,1347]" gridcol="2" gridrow="14" pageId="10" pageNumber="205">present</td>
<td id="76B75D9BFFBCB4AAF9EEAE1F82E7C339" box="[1581,2009,1322,1347]" gridcol="3" gridrow="14" pageId="10" pageNumber="205">present</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AE6782E7C3F5" box="[165,2009,1362,1423]" gridrow="15" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AE67876BC3F5" box="[165,597,1362,1423]" gridcol="0" gridrow="15" pageId="10" pageNumber="205">Maxilliped 3, ventral margin armature</th>
<td id="76B75D9BFFBCB4AAFD47AE678106C3F5" box="[644,1080,1362,1423]" gridcol="1" gridrow="15" pageId="10" pageNumber="205">not known</td>
<td id="76B75D9BFFBCB4AAFB9BAE67832EC3F5" box="[1112,1552,1362,1423]" gridcol="2" gridrow="15" pageId="10" pageNumber="205">ischium with 2 spines, merus with 3 spines, carpus with 1 subdistal spine</td>
<td id="76B75D9BFFBCB4AAF9EEAE6782E7C3F5" box="[1581,2009,1362,1423]" gridcol="3" gridrow="15" pageId="10" pageNumber="205">ischium with 3 spines, merus with 3–4 spines, carpus with 1 subdistal spine</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AEAB82E7C3CD" box="[165,2009,1438,1463]" gridrow="16" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AEAB876BC3CD" box="[165,597,1438,1463]" gridcol="0" gridrow="16" pageId="10" pageNumber="205">Relative size of chelipeds</th>
<td id="76B75D9BFFBCB4AAFD47AEAB8106C3CD" box="[644,1080,1438,1463]" gridcol="1" gridrow="16" pageId="10" pageNumber="205">unequal</td>
<td id="76B75D9BFFBCB4AAFB9BAEAB832EC3CD" box="[1112,1552,1438,1463]" gridcol="2" gridrow="16" pageId="10" pageNumber="205">unequal</td>
<td id="76B75D9BFFBCB4AAF9EEAEAB82E7C3CD" box="[1581,2009,1438,1463]" gridcol="3" gridrow="16" pageId="10" pageNumber="205">Subequal</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AEF382E7C05D" box="[165,2009,1478,1575]" gridrow="17" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AEF3876BC05D" box="[165,597,1478,1575]" gridcol="0" gridrow="17" pageId="10" pageNumber="205">Length of pollex relative to upper palm</th>
<td id="76B75D9BFFBCB4AAFD47AEF38106C05D" box="[644,1080,1478,1575]" gridcol="1" gridrow="17" pageId="10" pageNumber="205">pollex of major P1 1.4 times upper palm length, that of minor P1 3.1 times upper palm length</td>
<td id="76B75D9BFFBCB4AAFB9BAEF3832EC05D" box="[1112,1552,1478,1575]" gridcol="2" gridrow="17" pageId="10" pageNumber="205">pollex of major P1 1.5 times upper palm length, that of minor P1 1.66 times upper palm length</td>
<td id="76B75D9BFFBCB4AAF9EEAEF382E7C05D" box="[1581,2009,1478,1575]" gridcol="3" gridrow="17" pageId="10" pageNumber="205">pollex of major P1 3.0 times upper palm length, that of minor P1 3.2 times upper palm length</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AD0282E7C02A" box="[165,2009,1591,1616]" gridrow="18" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AD02876BC02A" box="[165,597,1591,1616]" gridcol="0" gridrow="18" pageId="10" pageNumber="205">P1 carpus, dorsal surface armature</th>
<td id="76B75D9BFFBCB4AAFD47AD028106C02A" box="[644,1080,1591,1616]" gridcol="1" gridrow="18" pageId="10" pageNumber="205">5–8 spines or numerous denticles</td>
<td id="76B75D9BFFBCB4AAFB9BAD02832EC02A" box="[1112,1552,1591,1616]" gridcol="2" gridrow="18" pageId="10" pageNumber="205">2–3 spines</td>
<td id="76B75D9BFFBCB4AAF9EEAD0282E7C02A" box="[1581,2009,1591,1616]" gridcol="3" gridrow="18" pageId="10" pageNumber="205">2–3 spines</td>
</tr>
<tr id="356634E7FFBCB4AAFF66AD6A82E7C002" box="[165,2009,1631,1656]" gridrow="19" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66AD6A876BC002" box="[165,597,1631,1656]" gridcol="0" gridrow="19" pageId="10" pageNumber="205">P1 propodus, dorsal margin armature</th>
<td id="76B75D9BFFBCB4AAFD47AD6A8106C002" box="[644,1080,1631,1656]" gridcol="1" gridrow="19" pageId="10" pageNumber="205">2–3 pairs of spines or numerous denticles</td>
<td id="76B75D9BFFBCB4AAFB9BAD6A832EC002" box="[1112,1552,1631,1656]" gridcol="2" gridrow="19" pageId="10" pageNumber="205">2–3 spines</td>
<td id="76B75D9BFFBCB4AAF9EEAD6A82E7C002" box="[1581,2009,1631,1656]" gridcol="3" gridrow="19" pageId="10" pageNumber="205">2 rows of 3–5 spines</td>
</tr>
<tr id="356634E7FFBCB4AAFF66ADB282E7C0DA" box="[165,2009,1671,1696]" gridrow="20" pageId="10" pageNumber="205">
<th id="76B75D9BFFBCB4AAFF66ADB2876BC0DA" box="[165,597,1671,1696]" gridcol="0" gridrow="20" pageId="10" pageNumber="205">P1 dactylus, dorsal margin armature</th>
<td id="76B75D9BFFBCB4AAFD47ADB28106C0DA" box="[644,1080,1671,1696]" gridcol="1" gridrow="20" pageId="10" pageNumber="205">5 spines on minor P1</td>
<td id="76B75D9BFFBCB4AAFB9BADB2832EC0DA" box="[1112,1552,1671,1696]" gridcol="2" gridrow="20" pageId="10" pageNumber="205">2 spines each on both P1</td>
<td id="76B75D9BFFBCB4AAF9EEADB282E7C0DA" box="[1581,2009,1671,1696]" gridcol="3" gridrow="20" pageId="10" pageNumber="205">6 spines on major P1, 5 on minor P1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE936A5FFBC4B5FF96DAD8582DAC0B3" blockId="10.[1710,2020,1712,1737]" box="[1710,2020,1712,1737]" pageId="10" pageNumber="205">......continued on the next page</paragraph>
<caption id="DF29662DFFBD4B5EFF54A95884B0C4FC" box="[151,398,621,646]" isContinuationCaption="true" pageId="11" pageNumber="206" startId="11.[151,235,621,646]" targetBox="[151,1996,661,1529]" targetIsTable="true" targetPageId="11">
<paragraph id="8BE936A5FFBD4B5EFF54A95884B0C4FC" blockId="11.[151,398,621,646]" box="[151,398,621,646]" pageId="11" pageNumber="206">TABLE 2. (Continued)</paragraph>
</caption>
<paragraph id="8BE936A5FFBD4B5EFF54A9A0827EC3AF" pageId="11" pageNumber="206">
<table id="F956C405FFBDB4AAFF54A9A082F2C383" box="[151,1996,661,1529]" colsContinueFrom="10.[165,2009,506,1696]" gridcols="4" gridrows="13" pageId="11" pageNumber="206">
<tr id="356634E7FFBDB4AAFF54A9A082F2C4D4" box="[151,1996,661,686]" gridrow="0" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54A9A08770C4D4" box="[151,590,661,686]" gridcol="0" gridrow="0" pageId="11" pageNumber="206">
<emphasis id="B922EAB7FFBD4B5EFF54A9A08458C4D4" bold="true" box="[151,358,661,686]" pageId="11" pageNumber="206">Characters/Species</emphasis>
</th>
<th id="76B75D9BFFBDB4AAFDB6A9A08121C4D4" box="[629,1055,661,686]" gridcol="1" gridrow="0" pageId="11" pageNumber="206">
<taxonomicName id="4C564D26FFBD4B5EFDB6A9A08699C4D4" authority="(Squires, 1979)" baseAuthorityName="Squires" baseAuthorityYear="1979" box="[629,935,661,686]" class="Malacostraca" family="Axiidae" genus="Axiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="caespitosa">
<emphasis id="B922EAB7FFBD4B5EFDB6A9A08699C4D4" bold="true" box="[629,935,661,686]" pageId="11" pageNumber="206">
<emphasis id="B922EAB7FFBD4B5EFDB6A9A087C1C4D4" bold="true" box="[629,767,661,686]" italics="true" pageId="11" pageNumber="206">G. caespitosa</emphasis>
(
<bibRefCitation id="EFC74B54FFBD4B5EFCCFA9A0869EC4D4" author="Squires, H. J." box="[780,928,661,686]" pageId="11" pageNumber="206" pagination="1584 - 1591" refId="ref14776" refString="Squires, H. J. (1979) Axiopsis caespitosa (Thalassinidea, Axiidae), a new species from the Pacific coast of Colombia. Canadian Journal of Zoology, 57, 1584 - 1591. https: // doi. org / 10.1139 / z 79 - 207" type="journal article" year="1979">Squires, 1979</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="76B75D9BFFBDB4AAFB89A9A0833FC4D4" box="[1098,1537,661,686]" gridcol="2" gridrow="0" pageId="11" pageNumber="206">
<taxonomicName id="4C564D26FFBD4B5EFB89A9A0833FC4D4" authority="(Boesch &amp; Smalley, 1972)" baseAuthorityName="Boesch &amp; Smalley" baseAuthorityYear="1972" box="[1098,1537,661,686]" class="Malacostraca" family="Axiidae" genus="Calocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="hirsutimana">
<emphasis id="B922EAB7FFBD4B5EFB89A9A0833FC4D4" bold="true" box="[1098,1537,661,686]" pageId="11" pageNumber="206">
<emphasis id="B922EAB7FFBD4B5EFB89A9A081D7C4D4" bold="true" box="[1098,1257,661,686]" italics="true" pageId="11" pageNumber="206">G. hirsutimana</emphasis>
(
<bibRefCitation id="EFC74B54FFBD4B5EFB3BA9A080C4C4D4" author="Boesch, D. F. &amp; Smalley, A. E." box="[1272,1530,661,686]" pageId="11" pageNumber="206" pagination="45 - 52" refId="ref11230" refString="Boesch, D. F. &amp; Smalley, A. E. (1972) A new axiid (Decapoda, Thalassinidea) from the Northern Gulf of Mexico and tropical Atlantic. Bulletin of Marine Science, 22 (1), 45 - 52." type="journal article" year="1972">Boesch &amp; Smalley, 1972</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="76B75D9BFFBDB4AAF9DDA9A082F2C4D4" box="[1566,1996,661,686]" gridcol="3" gridrow="0" pageId="11" pageNumber="206">
<emphasis id="B922EAB7FFBD4B5EF9DDA9A083D4C4D4" bold="true" box="[1566,1770,661,686]" pageId="11" pageNumber="206">
<taxonomicName id="4C564D26FFBD4B5EF9DDA9A083ADC4D4" authority="Padate &amp; Cubelio &amp; Takeda, 2022" authorityName="Padate &amp; Cubelio &amp; Takeda" authorityYear="2022" box="[1566,1683,661,686]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="keralam" status="sp. nov.">
<emphasis id="B922EAB7FFBD4B5EF9DDA9A083ADC4D4" bold="true" box="[1566,1683,661,686]" italics="true" pageId="11" pageNumber="206">G. keralam</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCFFBD4B5EF95AA9A083D4C4D4" box="[1689,1770,661,686]" pageId="11" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="356634E7FFBDB4AAFF54A98B82F2C4AD" box="[151,1996,702,727]" gridrow="1" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54A98B8770C4AD" box="[151,590,702,727]" gridcol="0" gridrow="1" pageId="11" pageNumber="206">P2 merus, ventral margin armature</th>
<td id="76B75D9BFFBDB4AAFDB6A98B8121C4AD" box="[629,1055,702,727]" gridcol="1" gridrow="1" pageId="11" pageNumber="206">3 or 4 spines</td>
<td id="76B75D9BFFBDB4AAFB89A98B833FC4AD" box="[1098,1537,702,727]" gridcol="2" gridrow="1" pageId="11" pageNumber="206">4 spines</td>
<td id="76B75D9BFFBDB4AAF9DDA98B82F2C4AD" box="[1566,1996,702,727]" gridcol="3" gridrow="1" pageId="11" pageNumber="206">2 proximal and 1 distal spine</td>
</tr>
<tr id="356634E7FFBDB4AAFF54A9D382F2C485" box="[151,1996,742,767]" gridrow="2" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54A9D38770C485" box="[151,590,742,767]" gridcol="0" gridrow="2" pageId="11" pageNumber="206">P2 carpus, ventral margin armature</th>
<td id="76B75D9BFFBDB4AAFDB6A9D38121C485" box="[629,1055,742,767]" gridcol="1" gridrow="2" pageId="11" pageNumber="206">absent</td>
<td id="76B75D9BFFBDB4AAFB89A9D3833FC485" box="[1098,1537,742,767]" gridcol="2" gridrow="2" pageId="11" pageNumber="206">absent</td>
<td id="76B75D9BFFBDB4AAF9DDA9D382F2C485" box="[1566,1996,742,767]" gridcol="3" gridrow="2" pageId="11" pageNumber="206">absent</td>
</tr>
<tr id="356634E7FFBDB4AAFF54A83B82F2C531" box="[151,1996,782,843]" gridrow="3" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54A83B8770C531" box="[151,590,782,843]" gridcol="0" gridrow="3" pageId="11" pageNumber="206">P3–4 propodus and dactylus armature</th>
<td id="76B75D9BFFBDB4AAFDB6A83B8121C531" box="[629,1055,782,843]" gridcol="1" gridrow="3" pageId="11" pageNumber="206">absent</td>
<td id="76B75D9BFFBDB4AAFB89A83B833FC531" box="[1098,1537,782,843]" gridcol="2" gridrow="3" pageId="11" pageNumber="206">absent</td>
<td id="76B75D9BFFBDB4AAF9DDA83B82F2C531" box="[1566,1996,782,843]" gridcol="3" gridrow="3" pageId="11" pageNumber="206">corneous movable spines on P3–4 propodi and dactyli</td>
</tr>
<tr id="356634E7FFBDB4AAFF54A86F82F2C509" box="[151,1996,858,883]" gridrow="4" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54A86F8770C509" box="[151,590,858,883]" gridcol="0" gridrow="4" pageId="11" pageNumber="206">P5 propodus and dactylus armature</th>
<td id="76B75D9BFFBDB4AAFDB6A86F8121C509" box="[629,1055,858,883]" gridcol="1" gridrow="4" pageId="11" pageNumber="206">corneous denticles on P5 propodus only</td>
<td id="76B75D9BFFBDB4AAFB89A86F833FC509" box="[1098,1537,858,883]" gridcol="2" gridrow="4" pageId="11" pageNumber="206">absent</td>
<td id="76B75D9BFFBDB4AAF9DDA86F82F2C509" box="[1566,1996,858,883]" gridcol="3" gridrow="4" pageId="11" pageNumber="206">absent</td>
</tr>
<tr id="356634E7FFBDB4AAFF54A8B782F2C5C5" box="[151,1996,898,959]" gridrow="5" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54A8B78770C5C5" box="[151,590,898,959]" gridcol="0" gridrow="5" pageId="11" pageNumber="206">Telson, lateral margin armature</th>
<td id="76B75D9BFFBDB4AAFDB6A8B78121C5C5" box="[629,1055,898,959]" gridcol="1" gridrow="5" pageId="11" pageNumber="206">2 spines</td>
<td id="76B75D9BFFBDB4AAFB89A8B7833FC5C5" box="[1098,1537,898,959]" gridcol="2" gridrow="5" pageId="11" pageNumber="206">2 proximal and 1 distal spine</td>
<td id="76B75D9BFFBDB4AAF9DDA8B782F2C5C5" box="[1566,1996,898,959]" gridcol="3" gridrow="5" pageId="11" pageNumber="206">3 spines along middle third portion and 1 distal spine</td>
</tr>
<tr id="356634E7FFBDB4AAFF54A8FA82F2C592" box="[151,1996,975,1000]" gridrow="6" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54A8FA8770C592" box="[151,590,975,1000]" gridcol="0" gridrow="6" pageId="11" pageNumber="206">Telson, posterolateral margin armature</th>
<td id="76B75D9BFFBDB4AAFDB6A8FA8121C592" box="[629,1055,975,1000]" gridcol="1" gridrow="6" pageId="11" pageNumber="206">absent</td>
<td id="76B75D9BFFBDB4AAFB89A8FA833FC592" box="[1098,1537,975,1000]" gridcol="2" gridrow="6" pageId="11" pageNumber="206">absent</td>
<td id="76B75D9BFFBDB4AAF9DDA8FA82F2C592" box="[1566,1996,975,1000]" gridcol="3" gridrow="6" pageId="11" pageNumber="206">absent</td>
</tr>
<tr id="356634E7FFBDB4AAFF54A8C282F2C26A" box="[151,1996,1015,1040]" gridrow="7" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54A8C28770C26A" box="[151,590,1015,1040]" gridcol="0" gridrow="7" pageId="11" pageNumber="206">Uropodal endopod, lateral margin armature</th>
<td id="76B75D9BFFBDB4AAFDB6A8C28121C26A" box="[629,1055,1015,1040]" gridcol="1" gridrow="7" pageId="11" pageNumber="206">2 spines</td>
<td id="76B75D9BFFBDB4AAFB89A8C2833FC26A" box="[1098,1537,1015,1040]" gridcol="2" gridrow="7" pageId="11" pageNumber="206">2 spines</td>
<td id="76B75D9BFFBDB4AAF9DDA8C282F2C26A" box="[1566,1996,1015,1040]" gridcol="3" gridrow="7" pageId="11" pageNumber="206">3 spines</td>
</tr>
<tr id="356634E7FFBDB4AAFF54AF2A82F2C226" box="[151,1996,1055,1116]" gridrow="8" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54AF2A8770C226" box="[151,590,1055,1116]" gridcol="0" gridrow="8" pageId="11" pageNumber="206">Uropodal exopod, lateral margin armature</th>
<td id="76B75D9BFFBDB4AAFDB6AF2A8121C226" box="[629,1055,1055,1116]" gridcol="1" gridrow="8" pageId="11" pageNumber="206">5 spines (including 2 movable spines at edge of transverse suture)</td>
<td id="76B75D9BFFBDB4AAFB89AF2A833FC226" box="[1098,1537,1055,1116]" gridcol="2" gridrow="8" pageId="11" pageNumber="206">2 spines</td>
<td id="76B75D9BFFBDB4AAF9DDAF2A82F2C226" box="[1566,1996,1055,1116]" gridcol="3" gridrow="8" pageId="11" pageNumber="206">4 spines</td>
</tr>
<tr id="356634E7FFBDB4AAFF54AF5E82F2C2D2" box="[151,1996,1131,1192]" gridrow="9" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54AF5E8770C2D2" box="[151,590,1131,1192]" gridcol="0" gridrow="9" pageId="11" pageNumber="206">Nature of appendix masculina on male pleopod 2</th>
<td id="76B75D9BFFBDB4AAFDB6AF5E8121C2D2" box="[629,1055,1131,1192]" gridcol="1" gridrow="9" pageId="11" pageNumber="206">uni-segmented, distinctly shorter than ap- pendix interna</td>
<td id="76B75D9BFFBDB4AAFB89AF5E833FC2D2" box="[1098,1537,1131,1192]" gridcol="2" gridrow="9" pageId="11" pageNumber="206">uni-segmented, distinctly shorter than ap- pendix interna</td>
<td id="76B75D9BFFBDB4AAF9DDAF5E82F2C2D2" box="[1566,1996,1131,1192]" gridcol="3" gridrow="9" pageId="11" pageNumber="206">uni-segmented, subequal to appendix interna</td>
</tr>
<tr id="356634E7FFBDB4AAFF54AF8282F2C31B" box="[151,1996,1207,1377]" gridrow="10" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54AF828770C31B" box="[151,590,1207,1377]" gridcol="0" gridrow="10" pageId="11" pageNumber="206">
<emphasis id="B922EAB7FFBD4B5EFF54AF82848EC2AA" bold="true" box="[151,432,1207,1232]" pageId="11" pageNumber="206">Geographical distribution</emphasis>
</th>
<td id="76B75D9BFFBDB4AAFDB6AF828121C31B" box="[629,1055,1207,1377]" gridcol="1" gridrow="10" pageId="11" pageNumber="206">
Gulf of California, Pacific coasts of Costa Rica, Mexico, Nicaragua, Colombia and Peru (
<bibRefCitation id="EFC74B54FFBD4B5EFD72AE358605C363" author="Squires, H. J." box="[689,827,1280,1305]" pageId="11" pageNumber="206" pagination="1584 - 1591" refId="ref14776" refString="Squires, H. J. (1979) Axiopsis caespitosa (Thalassinidea, Axiidae), a new species from the Pacific coast of Colombia. Canadian Journal of Zoology, 57, 1584 - 1591. https: // doi. org / 10.1139 / z 79 - 207" type="journal article" year="1979">Squires 1979</bibRefCitation>
;
<bibRefCitation id="EFC74B54FFBD4B5EFC86AE3586D5C363" author="Hendrickx, M." box="[837,1003,1280,1305]" pageId="11" pageNumber="206" pagination="355 - 358" refId="ref11841" refString="Hendrickx, M. (1987) The species of Axiidae (Crustacea: Thalassinidea) from the Pacific coast of Mexico, with a key for their identification. Revista de Biologia Tropical, 32 (2), 355 - 358" type="journal article" year="1987">Hendrickx 1987</bibRefCitation>
,
<bibRefCitation id="EFC74B54FFBD4B5EFDB6AE118795C347" author="Hendrickx, M." box="[629,683,1316,1341]" pageId="11" pageNumber="206" pagination="1277 - 1278" refId="ref11884" refString="Hendrickx, M. (2004) Additional records of Acanthaxius caespitosus (Squires, 1979) (Decapoda, Thalassinoidea, Axiidae) from the Eastern Tropical Pacific, Crustaceana, 77 (10), 1277 - 1278. https: // doi. org / 10.1163 / 1568540043166065" type="journal article" year="2004">2004</bibRefCitation>
;
<bibRefCitation id="EFC74B54FFBD4B5EFD76AE11865BC347" author="Dworschak, P. C." box="[693,869,1316,1341]" pageId="11" pageNumber="206" pagination="37 - 55" refId="ref11485" refString="Dworschak, P. C. (2013) Axiidea and Gebiidea (Crustacea: Decapoda) of Costa Rica. Annalen des Naturhistorischen Museums in Wien, Serie B, 115, 37 - 55." type="journal article" year="2013">Dworschak 2013</bibRefCitation>
;
<bibRefCitation id="EFC74B54FFBD4B5EFCACAE1187D5C31B" author="Dworschak, P. C. &amp; Poore, G. C. B." pageId="11" pageNumber="206" pagination="61 - 76" refId="ref11684" refString="Dworschak, P. C. &amp; Poore, G. C. B. (2018) More cautionary tales: family, generic and species synonymies of recently published taxa of ghost and mud shrimps (Decapoda: Axiidea and Gebiidea). Zootaxa, 4394 (1), 61 - 76. https: // doi. org / 10.11646 / zootaxa. 4394.1.3" type="journal article" year="2018">Dworschak &amp; Poore 2018</bibRefCitation>
)
</td>
<td id="76B75D9BFFBDB4AAFB89AF82833FC31B" box="[1098,1537,1207,1377]" gridcol="2" gridrow="10" pageId="11" pageNumber="206">
Gulf of Mexico, tropical Atlantic off Guyana, Costa Rica, South Atlantic Bight off Georgia, South Carolina (
<bibRefCitation id="EFC74B54FFBD4B5EFAB2AE3581E0C347" author="Boesch, D. F. &amp; Smalley, A. E." pageId="11" pageNumber="206" pagination="45 - 52" refId="ref11230" refString="Boesch, D. F. &amp; Smalley, A. E. (1972) A new axiid (Decapoda, Thalassinidea) from the Northern Gulf of Mexico and tropical Atlantic. Bulletin of Marine Science, 22 (1), 45 - 52." type="journal article" year="1972">Boesch &amp; Smalley, 1972</bibRefCitation>
;
<bibRefCitation id="EFC74B54FFBD4B5EFB2BAE1180A7C347" author="Heard, R. W. &amp; King, R. A. &amp; Knott, D. M. &amp; Thoma, B. P. &amp; Thornton-DeVictor, S." box="[1256,1433,1316,1341]" pageId="11" pageNumber="206" refId="ref11752" refString="Heard, R. W., King, R. A., Knott, D. M., Thoma, B. P. &amp; Thornton-DeVictor, S. (2007) A guide to the Thalassinidea (Crustacea: Malacostraca: Decapoda) of the South Atlantic bight. NOAA Professional Paper NMFS 8, Seattle, Washington, 30 pp. [https: // spo. nmfs. noaa. gov / sites / default / files / pp 8. pdf]" type="book" year="2007">
Heard
<emphasis id="B922EAB7FFBD4B5EFAEFAE118067C347" box="[1324,1369,1316,1341]" italics="true" pageId="11" pageNumber="206">et al</emphasis>
. 2007
</bibRefCitation>
)
</td>
<td id="76B75D9BFFBDB4AAF9DDAF8282F2C31B" box="[1566,1996,1207,1377]" gridcol="3" gridrow="10" pageId="11" pageNumber="206">Southeastern Arabian Sea (present study)</td>
</tr>
<tr id="356634E7FFBDB4AAFF54AE4582F2C3F3" box="[151,1996,1392,1417]" gridrow="11" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54AE458770C3F3" box="[151,590,1392,1417]" gridcol="0" gridrow="11" pageId="11" pageNumber="206">
<emphasis id="B922EAB7FFBD4B5EFF54AE45845CC3F3" bold="true" box="[151,354,1392,1417]" pageId="11" pageNumber="206">Bathymetric range</emphasis>
</th>
<td id="76B75D9BFFBDB4AAFDB6AE458121C3F3" box="[629,1055,1392,1417]" gridcol="1" gridrow="11" pageId="11" pageNumber="206">72–200 m</td>
<td id="76B75D9BFFBDB4AAFB89AE45833FC3F3" box="[1098,1537,1392,1417]" gridcol="2" gridrow="11" pageId="11" pageNumber="206">62–232 m</td>
<td id="76B75D9BFFBDB4AAF9DDAE4582F2C3F3" box="[1566,1996,1392,1417]" gridcol="3" gridrow="11" pageId="11" pageNumber="206">326 m</td>
</tr>
<tr id="356634E7FFBDB4AAFF54AEAD82F2C383" box="[151,1996,1432,1529]" gridrow="12" pageId="11" pageNumber="206">
<th id="76B75D9BFFBDB4AAFF54AEAD8770C383" box="[151,590,1432,1529]" gridcol="0" gridrow="12" pageId="11" pageNumber="206">
<emphasis id="B922EAB7FFBD4B5EFF54AEAD85F6C3AF" bold="true" pageId="11" pageNumber="206">References (for morphological charac- ters)</emphasis>
</th>
<td id="76B75D9BFFBDB4AAFDB6AEAD8121C383" box="[629,1055,1432,1529]" gridcol="1" gridrow="12" pageId="11" pageNumber="206">
<bibRefCitation id="EFC74B54FFBD4B5EFDB6AEAD8638C3CB" author="Squires, H. J." box="[629,774,1432,1457]" pageId="11" pageNumber="206" pagination="1584 - 1591" refId="ref14776" refString="Squires, H. J. (1979) Axiopsis caespitosa (Thalassinidea, Axiidae), a new species from the Pacific coast of Colombia. Canadian Journal of Zoology, 57, 1584 - 1591. https: // doi. org / 10.1139 / z 79 - 207" type="journal article" year="1979">Squires (1979</bibRefCitation>
: figs 2, 3),
<bibRefCitation id="EFC74B54FFBD4B5EFCBBAEAD813FC3CB" author="Sakai, K." box="[888,1025,1432,1457]" pageId="11" pageNumber="206" pagination="503 - 510" refId="ref14491" refString="Sakai, K. (2017 a) One new species of a new genus, Neoaxius gen. nov., in a new family, Neoaxiidae fam. nov., from the Gulf of Nicoya, Costa Rica (Decapoda, Axioidea). Crustaceana, 90 (4), 503 - 510. https: // doi. org / 10.1163 / 15685403 - 00003650" type="journal article" year="2017">Sakai (2017a</bibRefCitation>
: figs 1A–B, 2A–B, 3C),
<bibRefCitation id="EFC74B54FFBD4B5EFCA4AE8987C8C383" author="Dworschak, P. C. &amp; Poore, G. C. B." pageId="11" pageNumber="206" pagination="61 - 76" refId="ref11684" refString="Dworschak, P. C. &amp; Poore, G. C. B. (2018) More cautionary tales: family, generic and species synonymies of recently published taxa of ghost and mud shrimps (Decapoda: Axiidea and Gebiidea). Zootaxa, 4394 (1), 61 - 76. https: // doi. org / 10.11646 / zootaxa. 4394.1.3" type="journal article" year="2018">Dworschak &amp; Poore (2018</bibRefCitation>
: fig. 1C)
</td>
<td id="76B75D9BFFBDB4AAFB89AEAD833FC383" box="[1098,1537,1432,1529]" gridcol="2" gridrow="12" pageId="11" pageNumber="206">
<bibRefCitation id="EFC74B54FFBD4B5EFB89AEAD8073C3CB" author="Boesch, D. F. &amp; Smalley, A. E." box="[1098,1357,1432,1457]" pageId="11" pageNumber="206" pagination="45 - 52" refId="ref11230" refString="Boesch, D. F. &amp; Smalley, A. E. (1972) A new axiid (Decapoda, Thalassinidea) from the Northern Gulf of Mexico and tropical Atlantic. Bulletin of Marine Science, 22 (1), 45 - 52." type="journal article" year="1972">Boesch &amp; Smalley (1972</bibRefCitation>
: figs 1–8)
</td>
<td id="76B75D9BFFBDB4AAF9DDAEAD82F2C383" box="[1566,1996,1432,1529]" gridcol="3" gridrow="12" pageId="11" pageNumber="206">Present study (Figs 1A–C, 2A–B, D–E, G–H, 3A–D, 4A, C–H, J–K)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE936A5FFBD4B5EF95CAD3C82E8C058" blockId="11.[1695,2006,1545,1570]" box="[1695,2006,1545,1570]" pageId="11" pageNumber="206">......continued on the next page</paragraph>
<caption id="DF29662DFFBA4B59FF54AAF984B0C79F" box="[151,398,460,485]" isContinuationCaption="true" pageId="12" pageNumber="207" startId="12.[151,235,460,485]" targetBox="[151,2006,500,1686]" targetIsTable="true" targetPageId="12">
<paragraph id="8BE936A5FFBA4B59FF54AAF984B0C79F" blockId="12.[151,398,460,485]" box="[151,398,460,485]" pageId="12" pageNumber="207">TABLE 2. (Continued)</paragraph>
</caption>
<paragraph id="8BE936A5FFBA4B59FF54AAC18349C0EC" pageId="12" pageNumber="207">
<table id="F956C405FFBAB4AAFF54AAC182E8C0EC" box="[151,2006,500,1686]" colsContinueIn="13.[151,1972,698,1571]" gridcols="4" gridrows="20" pageId="12" pageNumber="207">
<tr id="356634E7FFBAB4AAFF54AAC182E8C477" box="[151,2006,500,525]" gridrow="0" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AAC18760C477" box="[151,606,500,525]" gridcol="0" gridrow="0" pageId="12" pageNumber="207">
<emphasis id="B922EAB7FFBA4B59FF54AAC18458C477" bold="true" box="[151,358,500,525]" pageId="12" pageNumber="207">Characters/Species</emphasis>
</th>
<th id="76B75D9BFFBAB4AAFD48AAC1816AC477" box="[651,1108,500,525]" gridcol="1" gridrow="0" pageId="12" pageNumber="207">
<taxonomicName id="4C564D26FFBA4B59FD48AAC186E8C477" authority="(K. Sakai, 1994)" baseAuthorityName="K. Sakai" baseAuthorityYear="1994" box="[651,982,500,525]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="207" phylum="Arthropoda" rank="species" species="polychaetes">
<emphasis id="B922EAB7FFBA4B59FD48AAC186E8C477" bold="true" box="[651,982,500,525]" pageId="12" pageNumber="207">
<emphasis id="B922EAB7FFBA4B59FD48AAC1861DC477" bold="true" box="[651,803,500,525]" italics="true" pageId="12" pageNumber="207">G. polychaetes</emphasis>
(K.
<bibRefCitation id="EFC74B54FFBA4B59FC91AAC186F1C477" author="Sakai, K." box="[850,975,500,525]" pageId="12" pageNumber="207" pagination="175 - 202" refId="ref14144" refString="Sakai, K. (1994) Eleven species of Australian Axiidae (Crustacea: Decapoda: Thalassinidea) with descriptions of one new genus and five new species. The Beagle, Occasional Papers of the Northern Territory Museum of Arts and Sciences, 11, 175 - 202." type="journal article" year="1994">Sakai, 1994</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="76B75D9BFFBAB4AAFBB7AAC180C4C477" box="[1140,1530,500,525]" gridcol="2" gridrow="0" pageId="12" pageNumber="207">
<taxonomicName id="4C564D26FFBA4B59FBB7AAC180F0C477" authority="(Rathbun, 1906)" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[1140,1486,500,525]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="207" phylum="Arthropoda" rank="species" species="spinosissima">
<emphasis id="B922EAB7FFBA4B59FBB7AAC180F0C477" bold="true" box="[1140,1486,500,525]" pageId="12" pageNumber="207">
<emphasis id="B922EAB7FFBA4B59FBB7AAC18029C477" bold="true" box="[1140,1303,500,525]" italics="true" pageId="12" pageNumber="207">G. spinosissima</emphasis>
(
<bibRefCitation id="EFC74B54FFBA4B59FAE6AAC180F9C477" author="Rathbun, M. J." box="[1317,1479,500,525]" pageId="12" pageNumber="207" pagination="827 - 930" refId="ref13703" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. I - XIV." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="76B75D9BFFBAB4AAF9D9AAC182E8C477" box="[1562,2006,500,525]" gridcol="3" gridrow="0" pageId="12" pageNumber="207">
<emphasis id="B922EAB7FFBA4B59F9D9AAC18258C477" bold="true" box="[1562,1894,500,525]" pageId="12" pageNumber="207">
<emphasis id="B922EAB7FFBA4B59F9D9AAC183FDC477" bold="true" box="[1562,1731,500,525]" italics="true" pageId="12" pageNumber="207">B. thailandensis</emphasis>
K.
<bibRefCitation id="EFC74B54FFBA4B59F929AAC18258C477" author="Sakai, K." box="[1770,1894,500,525]" pageId="12" pageNumber="207" pagination="867 - 880" refId="ref14424" refString="Sakai, K. (2015) Two new species, Colemanaxius andamanensis sp. nov. and Bruceaxius thailandensis sp. nov. from the Andaman Sea, Thailand (infraorder Axiidea de Saint Laurent, 1979). Crustaceana, 88 (7 - 8), 867 - 880. http: // dx. doi. org / 10.1163 / 15685403 - 00003440" type="journal article" year="2015">Sakai, 2015</bibRefCitation>
</emphasis>
</th>
</tr>
<tr id="356634E7FFBAB4AAFF54A92882E8C420" box="[151,2006,541,602]" gridrow="1" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54A9288760C420" box="[151,606,541,602]" gridcol="0" gridrow="1" pageId="12" pageNumber="207">Rostrum</th>
<td id="76B75D9BFFBAB4AAFD48A928816AC420" box="[651,1108,541,602]" gridcol="1" gridrow="1" pageId="12" pageNumber="207">not reaching base of distalmost antennular article</td>
<td id="76B75D9BFFBAB4AAFBB7A92880C4C420" box="[1140,1530,541,602]" gridcol="2" gridrow="1" pageId="12" pageNumber="207">reaching base of distalmost antennular article</td>
<td id="76B75D9BFFBAB4AAF9D9A92882E8C420" box="[1562,2006,541,602]" gridcol="3" gridrow="1" pageId="12" pageNumber="207">not reaching base of penultimate antennular article</td>
</tr>
<tr id="356634E7FFBAB4AAFF54A95C82E8C4F8" box="[151,2006,617,642]" gridrow="2" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54A95C8760C4F8" box="[151,606,617,642]" gridcol="0" gridrow="2" pageId="12" pageNumber="207">Rostral lateral armature</th>
<td id="76B75D9BFFBAB4AAFD48A95C816AC4F8" box="[651,1108,617,642]" gridcol="1" gridrow="2" pageId="12" pageNumber="207">6–7 spines</td>
<td id="76B75D9BFFBAB4AAFBB7A95C80C4C4F8" box="[1140,1530,617,642]" gridcol="2" gridrow="2" pageId="12" pageNumber="207">3–4 pairs of spines</td>
<td id="76B75D9BFFBAB4AAF9D9A95C82E8C4F8" box="[1562,2006,617,642]" gridcol="3" gridrow="2" pageId="12" pageNumber="207">5–8 spines</td>
</tr>
<tr id="356634E7FFBAB4AAFF54A9A482E8C4B4" box="[151,2006,657,718]" gridrow="3" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54A9A48760C4B4" box="[151,606,657,718]" gridcol="0" gridrow="3" pageId="12" pageNumber="207">Carapace (excluding gastric carinae) orna- mentation</th>
<td id="76B75D9BFFBAB4AAFD48A9A4816AC4B4" box="[651,1108,657,718]" gridcol="1" gridrow="3" pageId="12" pageNumber="207">1 antennal spine</td>
<td id="76B75D9BFFBAB4AAFBB7A9A480C4C4B4" box="[1140,1530,657,718]" gridcol="2" gridrow="3" pageId="12" pageNumber="207">3 hepatic spines, 2 anterolateral spines (including antennal spine)</td>
<td id="76B75D9BFFBAB4AAF9D9A9A482E8C4B4" box="[1562,2006,657,718]" gridcol="3" gridrow="3" pageId="12" pageNumber="207">1 antennal spine</td>
</tr>
<tr id="356634E7FFBAB4AAFF54A9E882E8C48C" box="[151,2006,733,758]" gridrow="4" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54A9E88760C48C" box="[151,606,733,758]" gridcol="0" gridrow="4" pageId="12" pageNumber="207">Carapace, lateral gastric carina</th>
<td id="76B75D9BFFBAB4AAFD48A9E8816AC48C" box="[651,1108,733,758]" gridcol="1" gridrow="4" pageId="12" pageNumber="207">5–6 spines</td>
<td id="76B75D9BFFBAB4AAFBB7A9E880C4C48C" box="[1140,1530,733,758]" gridcol="2" gridrow="4" pageId="12" pageNumber="207">7 spines</td>
<td id="76B75D9BFFBAB4AAF9D9A9E882E8C48C" box="[1562,2006,733,758]" gridcol="3" gridrow="4" pageId="12" pageNumber="207">7 spines</td>
</tr>
<tr id="356634E7FFBAB4AAFF54A83382E8C565" box="[151,2006,774,799]" gridrow="5" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54A8338760C565" box="[151,606,774,799]" gridcol="0" gridrow="5" pageId="12" pageNumber="207">Carapace, submedian gastric carina armature</th>
<td id="76B75D9BFFBAB4AAFD48A833816AC565" box="[651,1108,774,799]" gridcol="1" gridrow="5" pageId="12" pageNumber="207">5–6 spines</td>
<td id="76B75D9BFFBAB4AAFBB7A83380C4C565" box="[1140,1530,774,799]" gridcol="2" gridrow="5" pageId="12" pageNumber="207">8–9 spines</td>
<td id="76B75D9BFFBAB4AAF9D9A83382E8C565" box="[1562,2006,774,799]" gridcol="3" gridrow="5" pageId="12" pageNumber="207">6–7 spines</td>
</tr>
<tr id="356634E7FFBAB4AAFF54A81B82E8C511" box="[151,2006,814,875]" gridrow="6" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54A81B8760C511" box="[151,606,814,875]" gridcol="0" gridrow="6" pageId="12" pageNumber="207">Carapace, median gastric carinae armature</th>
<td id="76B75D9BFFBAB4AAFD48A81B816AC511" box="[651,1108,814,875]" gridcol="1" gridrow="6" pageId="12" pageNumber="207">4 anterior spines followed by 1 tubercle and 6 posterior spines</td>
<td id="76B75D9BFFBAB4AAFBB7A81B80C4C511" box="[1140,1530,814,875]" gridcol="2" gridrow="6" pageId="12" pageNumber="207">5 anterior spines followed by 1 tuber- cle and 7 posterior spines</td>
<td id="76B75D9BFFBAB4AAF9D9A81B82E8C511" box="[1562,2006,814,875]" gridcol="3" gridrow="6" pageId="12" pageNumber="207">17 spines and 1 median tubercle</td>
</tr>
<tr id="356634E7FFBAB4AAFF54A84F82E8C5E9" box="[151,2006,890,915]" gridrow="7" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54A84F8760C5E9" box="[151,606,890,915]" gridcol="0" gridrow="7" pageId="12" pageNumber="207">Carapace, cervical groove armature</th>
<td id="76B75D9BFFBAB4AAFD48A84F816AC5E9" box="[651,1108,890,915]" gridcol="1" gridrow="7" pageId="12" pageNumber="207">smooth</td>
<td id="76B75D9BFFBAB4AAFBB7A84F80C4C5E9" box="[1140,1530,890,915]" gridcol="2" gridrow="7" pageId="12" pageNumber="207">11 spines</td>
<td id="76B75D9BFFBAB4AAF9D9A84F82E8C5E9" box="[1562,2006,890,915]" gridcol="3" gridrow="7" pageId="12" pageNumber="207">groove finely denticulate</td>
</tr>
<tr id="356634E7FFBAB4AAFF54A89782E8C5C1" box="[151,2006,930,955]" gridrow="8" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54A8978760C5C1" box="[151,606,930,955]" gridcol="0" gridrow="8" pageId="12" pageNumber="207">Pleuron 1</th>
<td id="76B75D9BFFBAB4AAFD48A897816AC5C1" box="[651,1108,930,955]" gridcol="1" gridrow="8" pageId="12" pageNumber="207">acute</td>
<td id="76B75D9BFFBAB4AAFBB7A89780C4C5C1" box="[1140,1530,930,955]" gridcol="2" gridrow="8" pageId="12" pageNumber="207">acute</td>
<td id="76B75D9BFFBAB4AAF9D9A89782E8C5C1" box="[1562,2006,930,955]" gridcol="3" gridrow="8" pageId="12" pageNumber="207">less acute</td>
</tr>
<tr id="356634E7FFBAB4AAFF54A8FE82E8C272" box="[151,2006,971,1032]" gridrow="9" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54A8FE8760C272" box="[151,606,971,1032]" gridcol="0" gridrow="9" pageId="12" pageNumber="207">Pleura 1–5, marginal armature</th>
<td id="76B75D9BFFBAB4AAFD48A8FE816AC272" box="[651,1108,971,1032]" gridcol="1" gridrow="9" pageId="12" pageNumber="207">pleura 1, 3–6 with 1 spine each</td>
<td id="76B75D9BFFBAB4AAFBB7A8FE80C4C272" box="[1140,1530,971,1032]" gridcol="2" gridrow="9" pageId="12" pageNumber="207">pleuron 1 with several spines, pleura 2–3 with 2–3 spines</td>
<td id="76B75D9BFFBAB4AAF9D9A8FE82E8C272" box="[1562,2006,971,1032]" gridcol="3" gridrow="9" pageId="12" pageNumber="207">with 0, 9, 5, 4, 4 and 1 spines, respectively</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AF2282E8C24A" box="[151,2006,1047,1072]" gridrow="10" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AF228760C24A" box="[151,606,1047,1072]" gridcol="0" gridrow="10" pageId="12" pageNumber="207">Ocular peduncle</th>
<td id="76B75D9BFFBAB4AAFD48AF22816AC24A" box="[651,1108,1047,1072]" gridcol="1" gridrow="10" pageId="12" pageNumber="207">reaching midlength of rostrum</td>
<td id="76B75D9BFFBAB4AAFBB7AF2280C4C24A" box="[1140,1530,1047,1072]" gridcol="2" gridrow="10" pageId="12" pageNumber="207">reaching distal tip of rostrum</td>
<td id="76B75D9BFFBAB4AAF9D9AF2282E8C24A" box="[1562,2006,1047,1072]" gridcol="3" gridrow="10" pageId="12" pageNumber="207">not reaching distal tip of rostrum</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AF0A82E8C206" box="[151,2006,1087,1148]" gridrow="11" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AF0A8760C206" box="[151,606,1087,1148]" gridcol="0" gridrow="11" pageId="12" pageNumber="207">Antennular peduncle</th>
<td id="76B75D9BFFBAB4AAFD48AF0A816AC206" box="[651,1108,1087,1148]" gridcol="1" gridrow="11" pageId="12" pageNumber="207">not reaching distal margin of penultimate antennal article</td>
<td id="76B75D9BFFBAB4AAFBB7AF0A80C4C206" box="[1140,1530,1087,1148]" gridcol="2" gridrow="11" pageId="12" pageNumber="207">reaching distal margin of penultimate antennal article</td>
<td id="76B75D9BFFBAB4AAF9D9AF0A82E8C206" box="[1562,2006,1087,1148]" gridcol="3" gridrow="11" pageId="12" pageNumber="207">reaching distal margin of penultimate antennal article</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AFBE82E8C2DE" box="[151,2006,1163,1188]" gridrow="12" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AFBE8760C2DE" box="[151,606,1163,1188]" gridcol="0" gridrow="12" pageId="12" pageNumber="207">Antennal article 2, distal spine orientation</th>
<td id="76B75D9BFFBAB4AAFD48AFBE816AC2DE" box="[651,1108,1163,1188]" gridcol="1" gridrow="12" pageId="12" pageNumber="207">anterior</td>
<td id="76B75D9BFFBAB4AAFBB7AFBE80C4C2DE" box="[1140,1530,1163,1188]" gridcol="2" gridrow="12" pageId="12" pageNumber="207">anterior</td>
<td id="76B75D9BFFBAB4AAF9D9AFBE82E8C2DE" box="[1562,2006,1163,1188]" gridcol="3" gridrow="12" pageId="12" pageNumber="207">anterior</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AF8182E8C28B" box="[151,2006,1204,1265]" gridrow="13" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AF818760C28B" box="[151,606,1204,1265]" gridcol="0" gridrow="13" pageId="12" pageNumber="207">Scaphocerite</th>
<td id="76B75D9BFFBAB4AAFD48AF81816AC28B" box="[651,1108,1204,1265]" gridcol="1" gridrow="13" pageId="12" pageNumber="207">not reaching midlength of penultimate anten- nal article</td>
<td id="76B75D9BFFBAB4AAFBB7AF8180C4C28B" box="[1140,1530,1204,1265]" gridcol="2" gridrow="13" pageId="12" pageNumber="207">not reaching distal margin of penulti- mate antennal article</td>
<td id="76B75D9BFFBAB4AAF9D9AF8182E8C28B" box="[1562,2006,1204,1265]" gridcol="3" gridrow="13" pageId="12" pageNumber="207">not reaching distal margin of penultimate antennal article</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AE3582E8C363" box="[151,2006,1280,1305]" gridrow="14" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AE358760C363" box="[151,606,1280,1305]" gridcol="0" gridrow="14" pageId="12" pageNumber="207">Scaphocerite, mesial spine at base</th>
<td id="76B75D9BFFBAB4AAFD48AE35816AC363" box="[651,1108,1280,1305]" gridcol="1" gridrow="14" pageId="12" pageNumber="207">present</td>
<td id="76B75D9BFFBAB4AAFBB7AE3580C4C363" box="[1140,1530,1280,1305]" gridcol="2" gridrow="14" pageId="12" pageNumber="207">present</td>
<td id="76B75D9BFFBAB4AAF9D9AE3582E8C363" box="[1562,2006,1280,1305]" gridcol="3" gridrow="14" pageId="12" pageNumber="207">?absent</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AE1D82E8C31F" box="[151,2006,1320,1381]" gridrow="15" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AE1D8760C31F" box="[151,606,1320,1381]" gridcol="0" gridrow="15" pageId="12" pageNumber="207">Maxilliped 3, ventral margin armature</th>
<td id="76B75D9BFFBAB4AAFD48AE1D816AC31F" box="[651,1108,1320,1381]" gridcol="1" gridrow="15" pageId="12" pageNumber="207">ischium with 3 spines, merus with 3 spines, carpus with 1 subdistal spine</td>
<td id="76B75D9BFFBAB4AAFBB7AE1D80C4C31F" box="[1140,1530,1320,1381]" gridcol="2" gridrow="15" pageId="12" pageNumber="207">ischium with 3 spines, merus with 5 spines, carpus with 1 subdistal spine</td>
<td id="76B75D9BFFBAB4AAF9D9AE1D82E8C31F" box="[1562,2006,1320,1381]" gridcol="3" gridrow="15" pageId="12" pageNumber="207">ischium with 3 spines, merus with 6 spines, carpus unarmed</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AE4182E8C3F7" box="[151,2006,1396,1421]" gridrow="16" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AE418760C3F7" box="[151,606,1396,1421]" gridcol="0" gridrow="16" pageId="12" pageNumber="207">Relative size of chelipeds</th>
<td id="76B75D9BFFBAB4AAFD48AE41816AC3F7" box="[651,1108,1396,1421]" gridcol="1" gridrow="16" pageId="12" pageNumber="207">unequal</td>
<td id="76B75D9BFFBAB4AAFBB7AE4180C4C3F7" box="[1140,1530,1396,1421]" gridcol="2" gridrow="16" pageId="12" pageNumber="207">only 1 cheliped known</td>
<td id="76B75D9BFFBAB4AAF9D9AE4182E8C3F7" box="[1562,2006,1396,1421]" gridcol="3" gridrow="16" pageId="12" pageNumber="207">unequal</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AEA882E8C384" box="[151,2006,1437,1534]" gridrow="17" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AEA88760C384" box="[151,606,1437,1534]" gridcol="0" gridrow="17" pageId="12" pageNumber="207">Length of pollex relative to upper palm</th>
<td id="76B75D9BFFBAB4AAFD48AEA8816AC384" box="[651,1108,1437,1534]" gridcol="1" gridrow="17" pageId="12" pageNumber="207">pollex of major P1 0.7 times upper palm length, that of minor P1 subequal to upper palm length</td>
<td id="76B75D9BFFBAB4AAFBB7AEA880C4C384" box="[1140,1530,1437,1534]" gridcol="2" gridrow="17" pageId="12" pageNumber="207">pollex 1.1–1.25 times upper palm length</td>
<td id="76B75D9BFFBAB4AAF9D9AEA882E8C384" box="[1562,2006,1437,1534]" gridcol="3" gridrow="17" pageId="12" pageNumber="207">pollex of major P1 0.8 times upper palm length, that of minor P1 1.5 times upper palm length</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AD3882E8C030" box="[151,2006,1549,1610]" gridrow="18" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AD388760C030" box="[151,606,1549,1610]" gridcol="0" gridrow="18" pageId="12" pageNumber="207">P1 carpus, dorsal surface armature</th>
<td id="76B75D9BFFBAB4AAFD48AD38816AC030" box="[651,1108,1549,1610]" gridcol="1" gridrow="18" pageId="12" pageNumber="207">0–1 spine</td>
<td id="76B75D9BFFBAB4AAFBB7AD3880C4C030" box="[1140,1530,1549,1610]" gridcol="2" gridrow="18" pageId="12" pageNumber="207">2 spines</td>
<td id="76B75D9BFFBAB4AAF9D9AD3882E8C030" box="[1562,2006,1549,1610]" gridcol="3" gridrow="18" pageId="12" pageNumber="207">tuberculate on major P1, unarmed on minor P1</td>
</tr>
<tr id="356634E7FFBAB4AAFF54AD6C82E8C0EC" box="[151,2006,1625,1686]" gridrow="19" pageId="12" pageNumber="207">
<th id="76B75D9BFFBAB4AAFF54AD6C8760C0EC" box="[151,606,1625,1686]" gridcol="0" gridrow="19" pageId="12" pageNumber="207">P1 propodus, dorsal margin armature</th>
<td id="76B75D9BFFBAB4AAFD48AD6C816AC0EC" box="[651,1108,1625,1686]" gridcol="1" gridrow="19" pageId="12" pageNumber="207">0–3 spines</td>
<td id="76B75D9BFFBAB4AAFBB7AD6C80C4C0EC" box="[1140,1530,1625,1686]" gridcol="2" gridrow="19" pageId="12" pageNumber="207">7 spines</td>
<td id="76B75D9BFFBAB4AAF9D9AD6C82E8C0EC" box="[1562,2006,1625,1686]" gridcol="3" gridrow="19" pageId="12" pageNumber="207">tuberculate on major P1, denticulate on minor P1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE936A5FFBA4B59F960AD9382E4C0C5" blockId="12.[1699,2010,1702,1727]" box="[1699,2010,1702,1727]" pageId="12" pageNumber="207">......continued on the next page</paragraph>
<caption id="DF29662DFFBB4B58FF54A9A784B0C4D1" box="[151,398,658,683]" isContinuationCaption="true" pageId="13" pageNumber="208" startId="13.[151,235,658,683]" targetBox="[151,1972,698,1571]" targetIsTable="true" targetPageId="13">
<paragraph id="8BE936A5FFBB4B58FF54A9A784B0C4D1" blockId="13.[151,398,658,683]" box="[151,398,658,683]" pageId="13" pageNumber="208">TABLE 2. (Continued)</paragraph>
</caption>
<paragraph id="8BE936A5FFBB4B58FF54A98F82A6C385" pageId="13" pageNumber="208">
<table id="F956C405FFBBB4AAFF54A98F828AC059" box="[151,1972,698,1571]" colsContinueFrom="12.[151,2006,500,1686]" gridcols="4" gridrows="14" pageId="13" pageNumber="208">
<tr id="356634E7FFBBB4AAFF54A98F828AC4A9" box="[151,1972,698,723]" gridrow="0" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54A98F8761C4A9" box="[151,607,698,723]" gridcol="0" gridrow="0" pageId="13" pageNumber="208">
<emphasis id="B922EAB7FFBB4B58FF54A98F8458C4A9" bold="true" box="[151,358,698,723]" pageId="13" pageNumber="208">Characters/Species</emphasis>
</th>
<th id="76B75D9BFFBBB4AAFD48A98F816BC4A9" box="[651,1109,698,723]" gridcol="1" gridrow="0" pageId="13" pageNumber="208">
<taxonomicName id="4C564D26FFBB4B58FD48A98F86E8C4A9" authority="(K. Sakai, 1994)" baseAuthorityName="K. Sakai" baseAuthorityYear="1994" box="[651,982,698,723]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="208" phylum="Arthropoda" rank="species" species="polychaetes">
<emphasis id="B922EAB7FFBB4B58FD48A98F86E8C4A9" bold="true" box="[651,982,698,723]" pageId="13" pageNumber="208">
<emphasis id="B922EAB7FFBB4B58FD48A98F861DC4A9" bold="true" box="[651,803,698,723]" italics="true" pageId="13" pageNumber="208">G. polychaetes</emphasis>
(K.
<bibRefCitation id="EFC74B54FFBB4B58FC91A98F86F1C4A9" author="Sakai, K." box="[850,975,698,723]" pageId="13" pageNumber="208" pagination="175 - 202" refId="ref14144" refString="Sakai, K. (1994) Eleven species of Australian Axiidae (Crustacea: Decapoda: Thalassinidea) with descriptions of one new genus and five new species. The Beagle, Occasional Papers of the Northern Territory Museum of Arts and Sciences, 11, 175 - 202." type="journal article" year="1994">Sakai, 1994</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="76B75D9BFFBBB4AAFBB7A98F80C5C4A9" box="[1140,1531,698,723]" gridcol="2" gridrow="0" pageId="13" pageNumber="208">
<taxonomicName id="4C564D26FFBB4B58FBB7A98F80F0C4A9" authority="(Rathbun, 1906)" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[1140,1486,698,723]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="208" phylum="Arthropoda" rank="species" species="spinosissima">
<emphasis id="B922EAB7FFBB4B58FBB7A98F80F0C4A9" bold="true" box="[1140,1486,698,723]" pageId="13" pageNumber="208">
<emphasis id="B922EAB7FFBB4B58FBB7A98F8029C4A9" bold="true" box="[1140,1303,698,723]" italics="true" pageId="13" pageNumber="208">G. spinosissima</emphasis>
(
<bibRefCitation id="EFC74B54FFBB4B58FAE6A98F80F9C4A9" author="Rathbun, M. J." box="[1317,1479,698,723]" pageId="13" pageNumber="208" pagination="827 - 930" refId="ref13703" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. I - XIV." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="76B75D9BFFBBB4AAF9D9A98F828AC4A9" box="[1562,1972,698,723]" gridcol="3" gridrow="0" pageId="13" pageNumber="208">
<taxonomicName id="4C564D26FFBB4B58F9D9A98F8258C4A9" authority="K. Sakai, 2015" authorityName="K. Sakai" authorityYear="2015" box="[1562,1894,698,723]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="208" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B922EAB7FFBB4B58F9D9A98F8258C4A9" bold="true" box="[1562,1894,698,723]" pageId="13" pageNumber="208">
<emphasis id="B922EAB7FFBB4B58F9D9A98F83FDC4A9" bold="true" box="[1562,1731,698,723]" italics="true" pageId="13" pageNumber="208">B. thailandensis</emphasis>
K.
<bibRefCitation id="EFC74B54FFBB4B58F929A98F8258C4A9" author="Sakai, K." box="[1770,1894,698,723]" pageId="13" pageNumber="208" pagination="867 - 880" refId="ref14424" refString="Sakai, K. (2015) Two new species, Colemanaxius andamanensis sp. nov. and Bruceaxius thailandensis sp. nov. from the Andaman Sea, Thailand (infraorder Axiidea de Saint Laurent, 1979). Crustaceana, 88 (7 - 8), 867 - 880. http: // dx. doi. org / 10.1163 / 15685403 - 00003440" type="journal article" year="2015">Sakai, 2015</bibRefCitation>
</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="356634E7FFBBB4AAFF54A9D6828AC55A" box="[151,1972,739,800]" gridrow="1" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54A9D68761C55A" box="[151,607,739,800]" gridcol="0" gridrow="1" pageId="13" pageNumber="208">P1 dactylus, dorsal margin armature</th>
<td id="76B75D9BFFBBB4AAFD48A9D6816BC55A" box="[651,1109,739,800]" gridcol="1" gridrow="1" pageId="13" pageNumber="208">absent</td>
<td id="76B75D9BFFBBB4AAFBB7A9D680C5C55A" box="[1140,1531,739,800]" gridcol="2" gridrow="1" pageId="13" pageNumber="208">absent</td>
<td id="76B75D9BFFBBB4AAF9D9A9D6828AC55A" box="[1562,1972,739,800]" gridcol="3" gridrow="1" pageId="13" pageNumber="208">tuberculate on major P1, denticulate on minor P1</td>
</tr>
<tr id="356634E7FFBBB4AAFF54A81A828AC532" box="[151,1972,815,840]" gridrow="2" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54A81A8761C532" box="[151,607,815,840]" gridcol="0" gridrow="2" pageId="13" pageNumber="208">P2 merus, ventral margin armature</th>
<td id="76B75D9BFFBBB4AAFD48A81A816BC532" box="[651,1109,815,840]" gridcol="1" gridrow="2" pageId="13" pageNumber="208">4 spines</td>
<td id="76B75D9BFFBBB4AAFBB7A81A80C5C532" box="[1140,1531,815,840]" gridcol="2" gridrow="2" pageId="13" pageNumber="208">4 spines</td>
<td id="76B75D9BFFBBB4AAF9D9A81A828AC532" box="[1562,1972,815,840]" gridcol="3" gridrow="2" pageId="13" pageNumber="208">not known</td>
</tr>
<tr id="356634E7FFBBB4AAFF54A86D828AC50B" box="[151,1972,856,881]" gridrow="3" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54A86D8761C50B" box="[151,607,856,881]" gridcol="0" gridrow="3" pageId="13" pageNumber="208">P2 carpus, ventral margin armature</th>
<td id="76B75D9BFFBBB4AAFD48A86D816BC50B" box="[651,1109,856,881]" gridcol="1" gridrow="3" pageId="13" pageNumber="208">absent</td>
<td id="76B75D9BFFBBB4AAFBB7A86D80C5C50B" box="[1140,1531,856,881]" gridcol="2" gridrow="3" pageId="13" pageNumber="208">1 spine</td>
<td id="76B75D9BFFBBB4AAF9D9A86D828AC50B" box="[1562,1972,856,881]" gridcol="3" gridrow="3" pageId="13" pageNumber="208">not known</td>
</tr>
<tr id="356634E7FFBBB4AAFF54A8B5828AC5C7" box="[151,1972,896,957]" gridrow="4" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54A8B58761C5C7" box="[151,607,896,957]" gridcol="0" gridrow="4" pageId="13" pageNumber="208">P3–4 propodus and dactylus armature</th>
<td id="76B75D9BFFBBB4AAFD48A8B5816BC5C7" box="[651,1109,896,957]" gridcol="1" gridrow="4" pageId="13" pageNumber="208">corneous movable spines on P3–4 propodi only</td>
<td id="76B75D9BFFBBB4AAFBB7A8B580C5C5C7" box="[1140,1531,896,957]" gridcol="2" gridrow="4" pageId="13" pageNumber="208">absent</td>
<td id="76B75D9BFFBBB4AAF9D9A8B5828AC5C7" box="[1562,1972,896,957]" gridcol="3" gridrow="4" pageId="13" pageNumber="208">not known</td>
</tr>
<tr id="356634E7FFBBB4AAFF54A8F9828AC273" box="[151,1972,972,1033]" gridrow="5" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54A8F98761C273" box="[151,607,972,1033]" gridcol="0" gridrow="5" pageId="13" pageNumber="208">P5 propodus and dactylus armature</th>
<td id="76B75D9BFFBBB4AAFD48A8F9816BC273" box="[651,1109,972,1033]" gridcol="1" gridrow="5" pageId="13" pageNumber="208">corneous denticles on P5 propodus and dactylus</td>
<td id="76B75D9BFFBBB4AAFBB7A8F980C5C273" box="[1140,1531,972,1033]" gridcol="2" gridrow="5" pageId="13" pageNumber="208">absent</td>
<td id="76B75D9BFFBBB4AAF9D9A8F9828AC273" box="[1562,1972,972,1033]" gridcol="3" gridrow="5" pageId="13" pageNumber="208">not known</td>
</tr>
<tr id="356634E7FFBBB4AAFF54AF2D828AC22F" box="[151,1972,1048,1109]" gridrow="6" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54AF2D8761C22F" box="[151,607,1048,1109]" gridcol="0" gridrow="6" pageId="13" pageNumber="208">Telson, lateral margin armature</th>
<td id="76B75D9BFFBBB4AAFD48AF2D816BC22F" box="[651,1109,1048,1109]" gridcol="1" gridrow="6" pageId="13" pageNumber="208">proximal spines variable (2 on right side, 4 on left side), and 1 distal spine</td>
<td id="76B75D9BFFBBB4AAFBB7AF2D80C5C22F" box="[1140,1531,1048,1109]" gridcol="2" gridrow="6" pageId="13" pageNumber="208">1 proximal spine followed by 1 spine at midlength</td>
<td id="76B75D9BFFBBB4AAF9D9AF2D828AC22F" box="[1562,1972,1048,1109]" gridcol="3" gridrow="6" pageId="13" pageNumber="208">1 proximal and 5 distal spines</td>
</tr>
<tr id="356634E7FFBBB4AAFF54AF51828AC207" box="[151,1972,1124,1149]" gridrow="7" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54AF518761C207" box="[151,607,1124,1149]" gridcol="0" gridrow="7" pageId="13" pageNumber="208">Telson, posterolateral margin armature</th>
<td id="76B75D9BFFBBB4AAFD48AF51816BC207" box="[651,1109,1124,1149]" gridcol="1" gridrow="7" pageId="13" pageNumber="208">absent</td>
<td id="76B75D9BFFBBB4AAFBB7AF5180C5C207" box="[1140,1531,1124,1149]" gridcol="2" gridrow="7" pageId="13" pageNumber="208">3 pairs of spines</td>
<td id="76B75D9BFFBBB4AAF9D9AF51828AC207" box="[1562,1972,1124,1149]" gridcol="3" gridrow="7" pageId="13" pageNumber="208">1 pair of spines</td>
</tr>
<tr id="356634E7FFBBB4AAFF54AFB8828AC2DC" box="[151,1972,1165,1190]" gridrow="8" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54AFB88761C2DC" box="[151,607,1165,1190]" gridcol="0" gridrow="8" pageId="13" pageNumber="208">Uropodal endopod, lateral margin armature</th>
<td id="76B75D9BFFBBB4AAFD48AFB8816BC2DC" box="[651,1109,1165,1190]" gridcol="1" gridrow="8" pageId="13" pageNumber="208">2 spines</td>
<td id="76B75D9BFFBBB4AAFBB7AFB880C5C2DC" box="[1140,1531,1165,1190]" gridcol="2" gridrow="8" pageId="13" pageNumber="208">3 spines</td>
<td id="76B75D9BFFBBB4AAF9D9AFB8828AC2DC" box="[1562,1972,1165,1190]" gridcol="3" gridrow="8" pageId="13" pageNumber="208">1 spine</td>
</tr>
<tr id="356634E7FFBBB4AAFF54AF80828AC2B4" box="[151,1972,1205,1230]" gridrow="9" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54AF808761C2B4" box="[151,607,1205,1230]" gridcol="0" gridrow="9" pageId="13" pageNumber="208">Uropodal exopod, lateral margin armature</th>
<td id="76B75D9BFFBBB4AAFD48AF80816BC2B4" box="[651,1109,1205,1230]" gridcol="1" gridrow="9" pageId="13" pageNumber="208">5–6 spines</td>
<td id="76B75D9BFFBBB4AAFBB7AF8080C5C2B4" box="[1140,1531,1205,1230]" gridcol="2" gridrow="9" pageId="13" pageNumber="208">4 spines</td>
<td id="76B75D9BFFBBB4AAF9D9AF80828AC2B4" box="[1562,1972,1205,1230]" gridcol="3" gridrow="9" pageId="13" pageNumber="208">5 spines</td>
</tr>
<tr id="356634E7FFBBB4AAFF54AFE8828AC360" box="[151,1972,1245,1306]" gridrow="10" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54AFE88761C360" box="[151,607,1245,1306]" gridcol="0" gridrow="10" pageId="13" pageNumber="208">Nature of appendix masculina on male pleo- pod 2</th>
<td id="76B75D9BFFBBB4AAFD48AFE8816BC360" box="[651,1109,1245,1306]" gridcol="1" gridrow="10" pageId="13" pageNumber="208">uni-segmented, slightly longer than appendix interna</td>
<td id="76B75D9BFFBBB4AAFBB7AFE880C5C360" box="[1140,1531,1245,1306]" gridcol="2" gridrow="10" pageId="13" pageNumber="208">not known</td>
<td id="76B75D9BFFBBB4AAF9D9AFE8828AC360" box="[1562,1972,1245,1306]" gridcol="3" gridrow="10" pageId="13" pageNumber="208">tri-segmented, distinctly longer than appendix interna</td>
</tr>
<tr id="356634E7FFBBB4AAFF54AE1C828AC3D4" box="[151,1972,1321,1454]" gridrow="11" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54AE1C8761C3D4" box="[151,607,1321,1454]" gridcol="0" gridrow="11" pageId="13" pageNumber="208">
<emphasis id="B922EAB7FFBB4B58FF54AE1C848EC338" bold="true" box="[151,432,1321,1346]" pageId="13" pageNumber="208">Geographical distribution</emphasis>
</th>
<td id="76B75D9BFFBBB4AAFD48AE1C816BC3D4" box="[651,1109,1321,1454]" gridcol="1" gridrow="11" pageId="13" pageNumber="208">
Coral Sea off Queensland, Australia (
<bibRefCitation id="EFC74B54FFBB4B58FBC6AE1C8781C31C" author="Sakai, K." pageId="13" pageNumber="208" pagination="175 - 202" refId="ref14144" refString="Sakai, K. (1994) Eleven species of Australian Axiidae (Crustacea: Decapoda: Thalassinidea) with descriptions of one new genus and five new species. The Beagle, Occasional Papers of the Northern Territory Museum of Arts and Sciences, 11, 175 - 202." type="journal article" year="1994">Sakai 1994</bibRefCitation>
)
</td>
<td id="76B75D9BFFBBB4AAFBB7AE1C80C5C3D4" box="[1140,1531,1321,1454]" gridcol="2" gridrow="11" pageId="13" pageNumber="208">
Hawaii (
<bibRefCitation id="EFC74B54FFBB4B58FB08AE1C8060C338" author="Rathbun, M. J." box="[1227,1374,1321,1346]" pageId="13" pageNumber="208" pagination="827 - 930" refId="ref13703" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. I - XIV." type="journal article" year="1906">Rathbun 1906</bibRefCitation>
), North of Buton strait, Indonesia (de
<bibRefCitation id="EFC74B54FFBB4B58FA44AE788194C3F0" author="Man, J. G. de" pageId="13" pageNumber="208" pagination="1 - 127" refId="ref12630" refString="Man, J. G. de (1925) The Decapoda of the Siboga-Expedition. Part VI. The Axiidae collected by the Siboga-Expedition. Siboga Expeditie, 39 a 5, 1 - 127." type="journal article" year="1925">Man 1925</bibRefCitation>
), Nuku Hiva, Marquesas Islands, French Polynesia (
<bibRefCitation id="EFC74B54FFBB4B58FAF2AEA080F5C3D4" author="Ngoc-Ho, N." box="[1329,1483,1429,1454]" pageId="13" pageNumber="208" pagination="47 - 83" refId="ref13184" refString="Ngoc-Ho, N. (2005) Thalassinidea (Crustacea, Decapoda) from French Polynesia. Zoosystema, 27 (1), 47 - 83." type="journal article" year="2005">Ngoc-Ho 2005</bibRefCitation>
)
</td>
<td id="76B75D9BFFBBB4AAF9D9AE1C828AC3D4" box="[1562,1972,1321,1454]" gridcol="3" gridrow="11" pageId="13" pageNumber="208">
Andaman Sea, Straits of Malacca (
<bibRefCitation id="EFC74B54FFBB4B58F8B9AE1C8370C31C" author="Sakai, K." pageId="13" pageNumber="208" pagination="867 - 880" refId="ref14424" refString="Sakai, K. (2015) Two new species, Colemanaxius andamanensis sp. nov. and Bruceaxius thailandensis sp. nov. from the Andaman Sea, Thailand (infraorder Axiidea de Saint Laurent, 1979). Crustaceana, 88 (7 - 8), 867 - 880. http: // dx. doi. org / 10.1163 / 15685403 - 00003440" type="journal article" year="2015">Sakai 2015</bibRefCitation>
)
</td>
</tr>
<tr id="356634E7FFBBB4AAFF54AE88828AC3AD" box="[151,1972,1469,1495]" gridrow="12" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54AE888761C3AD" box="[151,607,1469,1495]" gridcol="0" gridrow="12" pageId="13" pageNumber="208">
<emphasis id="B922EAB7FFBB4B58FF54AE88845CC3AC" bold="true" box="[151,354,1469,1494]" pageId="13" pageNumber="208">Bathymetric range</emphasis>
</th>
<td id="76B75D9BFFBBB4AAFD48AE88816BC3AD" box="[651,1109,1469,1495]" gridcol="1" gridrow="12" pageId="13" pageNumber="208">260 m</td>
<td id="76B75D9BFFBBB4AAFBB7AE8880C5C3AD" box="[1140,1531,1469,1495]" gridcol="2" gridrow="12" pageId="13" pageNumber="208">41–43 m, 75–94 m, 90–155 m</td>
<td id="76B75D9BFFBBB4AAF9D9AE88828AC3AD" box="[1562,1972,1469,1495]" gridcol="3" gridrow="12" pageId="13" pageNumber="208">64–83 m</td>
</tr>
<tr id="356634E7FFBBB4AAFF54AED3828AC059" box="[151,1972,1510,1571]" gridrow="13" pageId="13" pageNumber="208">
<th id="76B75D9BFFBBB4AAFF54AED38761C059" box="[151,607,1510,1571]" gridcol="0" gridrow="13" pageId="13" pageNumber="208">
<emphasis id="B922EAB7FFBB4B58FF54AED38761C385" bold="true" box="[151,607,1510,1535]" pageId="13" pageNumber="208">References (for morphological characters)</emphasis>
</th>
<td id="76B75D9BFFBBB4AAFD48AED3816BC059" box="[651,1109,1510,1571]" gridcol="1" gridrow="13" pageId="13" pageNumber="208">
<bibRefCitation id="EFC74B54FFBB4B58FD48AED38636C385" author="Sakai, K." box="[651,776,1510,1535]" pageId="13" pageNumber="208" pagination="175 - 202" refId="ref14144" refString="Sakai, K. (1994) Eleven species of Australian Axiidae (Crustacea: Decapoda: Thalassinidea) with descriptions of one new genus and five new species. The Beagle, Occasional Papers of the Northern Territory Museum of Arts and Sciences, 11, 175 - 202." type="journal article" year="1994">Sakai (1994</bibRefCitation>
: figs 11A–F, 12B, F, 13A–F, H)
</td>
<td id="76B75D9BFFBBB4AAFBB7AED380C5C059" box="[1140,1531,1510,1571]" gridcol="2" gridrow="13" pageId="13" pageNumber="208">
<bibRefCitation id="EFC74B54FFBB4B58FBB7AED38031C385" author="Rathbun, M. J." box="[1140,1295,1510,1535]" pageId="13" pageNumber="208" pagination="827 - 930" refId="ref13703" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. I - XIV." type="journal article" year="1906">Rathbun (1906</bibRefCitation>
: fig. 50a–b),
<bibRefCitation id="EFC74B54FFBB4B58FA56AED3818DC059" author="Ngoc-Ho, N." pageId="13" pageNumber="208" pagination="47 - 83" refId="ref13184" refString="Ngoc-Ho, N. (2005) Thalassinidea (Crustacea, Decapoda) from French Polynesia. Zoosystema, 27 (1), 47 - 83." type="journal article" year="2005">Ngoc-Ho (2005</bibRefCitation>
: fig. 1A–G)
</td>
<td id="76B75D9BFFBBB4AAF9D9AED3828AC059" box="[1562,1972,1510,1571]" gridcol="3" gridrow="13" pageId="13" pageNumber="208">
<bibRefCitation id="EFC74B54FFBB4B58F9D9AED383A9C385" author="Sakai, K." box="[1562,1687,1510,1535]" pageId="13" pageNumber="208" pagination="867 - 880" refId="ref14424" refString="Sakai, K. (2015) Two new species, Colemanaxius andamanensis sp. nov. and Bruceaxius thailandensis sp. nov. from the Andaman Sea, Thailand (infraorder Axiidea de Saint Laurent, 1979). Crustaceana, 88 (7 - 8), 867 - 880. http: // dx. doi. org / 10.1163 / 15685403 - 00003440" type="journal article" year="2015">Sakai (2015</bibRefCitation>
: figs 3A–D, 4A, C–E, G)
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE936A5FFB84B5BFF04ABA285CFC7D7" blockId="14.[151,1437,150,465]" pageId="14" pageNumber="209">
In view of this,
<taxonomicName id="4C564D26FFB84B5BFE47ABA287F3C6CB" authority="K. Sakai, 2011" authorityName="K. Sakai" authorityYear="2011" box="[388,717,151,177]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB84B5BFE47ABA2872BC6CB" box="[388,533,151,177]" italics="true" pageId="14" pageNumber="209">Guyanacaris</emphasis>
K.
<bibRefCitation id="EFC74B54FFB84B5BFD87ABA287F3C6CB" author="Sakai, K." box="[580,717,151,177]" pageId="14" pageNumber="209" pagination="1 - 616" refId="ref14380" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
, is presently considered to represent 5 species—
<taxonomicName id="4C564D26FFB84B5BFAC2ABA28479C6AF" authority="(Squires, 1979)" baseAuthorityName="Squires" baseAuthorityYear="1979" class="Malacostraca" family="Axiidae" genus="Axiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="caespitosa">
<emphasis id="B922EAB7FFB84B5BFAC2ABA280A2C6CA" box="[1281,1436,151,177]" italics="true" pageId="14" pageNumber="209">G. caespitosa</emphasis>
(
<bibRefCitation id="EFC74B54FFB84B5BFF5CAB8E847EC6AF" author="Squires, H. J." box="[159,320,187,213]" pageId="14" pageNumber="209" pagination="1584 - 1591" refId="ref14776" refString="Squires, H. J. (1979) Axiopsis caespitosa (Thalassinidea, Axiidae), a new species from the Pacific coast of Colombia. Canadian Journal of Zoology, 57, 1584 - 1591. https: // doi. org / 10.1139 / z 79 - 207" type="journal article" year="1979">Squires, 1979</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C564D26FFB84B5BFE97AB8E860BC6AF" authority="(Boesch &amp; Smalley, 1972)" baseAuthorityName="Boesch &amp; Smalley" baseAuthorityYear="1972" box="[340,821,187,213]" class="Malacostraca" family="Axiidae" genus="Calocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="hirsutimana">
<emphasis id="B922EAB7FFB84B5BFE97AB8E84C0C6AF" box="[340,510,187,213]" italics="true" pageId="14" pageNumber="209">G. hirsutimana</emphasis>
(
<bibRefCitation id="EFC74B54FFB84B5BFDCDAB8E8613C6AF" author="Boesch, D. F. &amp; Smalley, A. E." box="[526,813,187,213]" pageId="14" pageNumber="209" pagination="45 - 52" refId="ref11230" refString="Boesch, D. F. &amp; Smalley, A. E. (1972) A new axiid (Decapoda, Thalassinidea) from the Northern Gulf of Mexico and tropical Atlantic. Bulletin of Marine Science, 22 (1), 45 - 52." type="journal article" year="1972">Boesch &amp; Smalley, 1972</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C564D26FFB84B5BFC82AB8E86FEC6AF" authority="Padate &amp; Cubelio &amp; Takeda, 2022" authorityName="Padate &amp; Cubelio &amp; Takeda" authorityYear="2022" box="[833,960,187,213]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="keralam" status="sp. nov.">
<emphasis id="B922EAB7FFB84B5BFC82AB8E86FEC6AF" box="[833,960,187,213]" italics="true" pageId="14" pageNumber="209">G. keralam</emphasis>
</taxonomicName>
<emphasis id="B922EAB7FFB84B5BFC0BAB8E811DC6AF" bold="true" box="[968,1059,187,213]" pageId="14" pageNumber="209">
<taxonomicNameLabel id="A21157CCFFB84B5BFC0BAB8E811DC6AF" box="[968,1059,187,213]" pageId="14" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C564D26FFB84B5BFBF1AB8E80A2C6AF" authority="(K. Sakai, 1994)" baseAuthorityName="K. Sakai" baseAuthorityYear="1994" box="[1074,1436,187,213]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="polychaetes">
<emphasis id="B922EAB7FFB84B5BFBF1AB8E81E7C6AF" box="[1074,1241,187,213]" italics="true" pageId="14" pageNumber="209">G. polychaetes</emphasis>
(K.
<bibRefCitation id="EFC74B54FFB84B5BFACDAB8E80AAC6AF" author="Sakai, K." box="[1294,1428,187,213]" pageId="14" pageNumber="209" pagination="175 - 202" refId="ref14144" refString="Sakai, K. (1994) Eleven species of Australian Axiidae (Crustacea: Decapoda: Thalassinidea) with descriptions of one new genus and five new species. The Beagle, Occasional Papers of the Northern Territory Museum of Arts and Sciences, 11, 175 - 202." type="journal article" year="1994">Sakai, 1994</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C564D26FFB84B5BFF06ABEA870DC680" authority="(Rathbun, 1906)" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[197,563,223,250]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="spinosissima">
<emphasis id="B922EAB7FFB84B5BFF06ABEA844BC682" box="[197,373,223,249]" italics="true" pageId="14" pageNumber="209">G. spinosissima</emphasis>
(
<bibRefCitation id="EFC74B54FFB84B5BFE41ABEA8715C680" author="Rathbun, M. J." box="[386,555,223,250]" pageId="14" pageNumber="209" pagination="827 - 930" refId="ref13703" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. I - XIV." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</taxonomicName>
, with known geographical distribution in the Atlantic and Indo-Pacific Oceans. This genus differs from
<taxonomicName id="4C564D26FFB84B5BFE63AA3686B0C767" authority="Sakai &amp; de Saint Laurent, 1989" authorityName="Sakai &amp; de Saint Laurent" authorityYear="1989" box="[416,910,259,285]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB84B5BFE63AA368714C767" box="[416,554,259,285]" italics="true" pageId="14" pageNumber="209">Acanthaxius</emphasis>
<bibRefCitation id="EFC74B54FFB84B5BFDF3AA3686B0C767" author="Sakai, K. &amp; Saint Laurent, M. de" box="[560,910,259,285]" pageId="14" pageNumber="209" pagination="1 - 104" refId="ref14618" refString="Sakai, K. &amp; Saint Laurent, M. de (1989) A check list of Axiidae (Decapoda, Crustacea, Thalassinidea, Anomura), with remarks and in addition descriptions of one new subfamily, eleven new genera and two new species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 3, 1 - 104." type="journal article" year="1989">Sakai &amp; de Saint Laurent, 1989</bibRefCitation>
</taxonomicName>
, in having short supraocular spine, the presence of post-cervical carina on the carapace, dentate margins of pleura 1–2, angular to almost straight posterior margins of pleura 3–5, scaphocerite with mesial spine at base, and the presence of male pleopod 1 (
<bibRefCitation id="EFC74B54FFB84B5BFB4AAA7E8037C71F" author="Sakai, K." box="[1161,1289,331,357]" pageId="14" pageNumber="209" pagination="1 - 616" refId="ref14380" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai 2011</bibRefCitation>
;
<bibRefCitation id="EFC74B54FFB84B5BFAD6AA7E80A7C71F" author="Poore, G. C. B." box="[1301,1433,331,357]" pageId="14" pageNumber="209" pagination="259 - 367" refId="ref13306" refString="Poore, G. C. B. (2020) Axiid and micheleid lobsters from Indo-West Pacific deep-sea environments (Crustacea: Decapoda: Axiidea: Axiidae, Micheleidae). In: Corbari, L., Ahyong, S. T. &amp; Chan, T. - Y. (Eds), Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos 31. Memoires du Museum national d'Histoire naturelle, Paris, 213, 259 - 367." type="journal article" year="2020">Poore 2020</bibRefCitation>
; present study). Morphological comparisons among the species of
<taxonomicName id="4C564D26FFB84B5BFC56AA5A8118C7F3" authorityName="K. Sakai" authorityYear="2011" box="[917,1062,367,393]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB84B5BFC56AA5A8118C7F3" box="[917,1062,367,393]" italics="true" pageId="14" pageNumber="209">Guyanacaris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C564D26FFB84B5BFBA0AA5A81DFC7F2" authorityName="K. Sakai" authorityYear="2011" box="[1123,1249,367,392]" class="Malacostraca" family="Axiidae" genus="Bruceaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7FFB84B5BFBA0AA5A81DFC7F2" box="[1123,1249,367,392]" italics="true" pageId="14" pageNumber="209">Bruceaxius</emphasis>
</taxonomicName>
are provided in
<tableCitation id="C6D4031EFFB84B5BFF54AAA685D3C7D7" box="[151,237,403,429]" captionStart="TABLE 2" captionStartId="10.[151,239,466,492]" captionTargetBox="[165,2009,506,1696]" captionTargetPageId="10" captionText="TABLE 2. Morphological comparison among the species of Guyanacaris K. Sakai, 2011 and Bruceaxius K. Sakai, 2011." httpUri="http://table.plazi.org/id/DF29662DFFBC4B5FFF54AAE7809BC796" pageId="14" pageNumber="209" tableUuid="DF29662DFFBC4B5FFF54AAE7809BC796">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C652EFFB84B5BFF04AA838607C7AB" box="[199,825,438,465]" pageId="14" pageNumber="209" type="distribution">
<paragraph id="8BE936A5FFB84B5BFF04AA838607C7AB" blockId="14.[151,1437,150,465]" box="[199,825,438,465]" pageId="14" pageNumber="209">
<emphasis id="B922EAB7FFB84B5BFF04AA83845FC7AA" bold="true" box="[199,353,438,464]" pageId="14" pageNumber="209">Distribution.</emphasis>
Southeastern Arabian Sea,
<collectingCountry id="F3417635FFB84B5BFD56AA8287F1C7AB" box="[661,719,439,465]" name="India" pageId="14" pageNumber="209">India</collectingCountry>
—
<specimenCount id="9D50FD2CFFB84B5BFD2FAA828607C7AB" box="[748,825,439,465]" pageId="14" pageNumber="209" type="generic">
<quantity id="4CAE9B40FFB84B5BFD2FAA82860BC7AB" box="[748,821,439,465]" metricMagnitude="2" metricUnit="m" metricValue="3.26" pageId="14" pageNumber="209" unit="m" value="326.0">326 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>
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