239 lines
38 KiB
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239 lines
38 KiB
XML
<document id="08538E32D41317AD7663EAA77A7FF958" ID-DOI="10.11646/zootaxa.5450.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="11232500" ID-ZooBank="1B5D192F-1D5B-4460-9133-9AEAE9C920BF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716281048833" checkinUser="plazi" docAuthor="Johnson, Robert G." docDate="2024" docId="03FD8227FFEFE311FF78FA17FC07849C" docLanguage="en" docName="zootaxa.5450.1.1.pdf" docOrigin="Zootaxa 5450 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5450.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Harpes oudris Johnson 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="50" masterDocId="FFC4FA5FFFDFE320FFEFF87CFFB58771" masterDocTitle="Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco" masterLastPageNumber="185" masterPageNumber="1" pageNumber="49" updateTime="1716282690877" updateUser="ExternalLinkService">
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<mods:title id="D231B2FB8D601C040FFCC067CA9D0CB3">Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco</mods:title>
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<mods:namePart id="577527D1C09EAA9B829313D2EBE5812B">Johnson, Robert G.</mods:namePart>
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<mods:typeOfResource id="F8010BEE25B9186A68F41516896ADC34">text</mods:typeOfResource>
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<mods:title id="0CFD0C553212A4A9EFBD7A59B54C9A0E">Zootaxa</mods:title>
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<mods:part id="D36C19FFEC8E5C583EFC1025166C593C">
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<mods:date id="38A8D0963F1506E88F6D3EDDEBF13C19">2024</mods:date>
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<mods:number id="F6088669E254FC3EEBFBA9B25F24C446">2024-05-15</mods:number>
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<mods:number id="B14B89009D6A37346C0C22579E4AC01D">5450</mods:number>
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<mods:classification id="2C58A5DF64A507D2129B731FFB2ACDD8">journal article</mods:classification>
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<mods:identifier id="CFB7569F97EE1776F57D9E1885994D39" type="DOI">10.11646/zootaxa.5450.1.1</mods:identifier>
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<mods:identifier id="D94BBA4A8034DEF6D4565038C768F3D8" type="ISSN">1175-5334</mods:identifier>
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</mods:mods>
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<treatment id="03FD8227FFEFE311FF78FA17FC07849C" LSID="urn:lsid:plazi:treatment:03FD8227FFEFE311FF78FA17FC07849C" httpUri="http://treatment.plazi.org/id/03FD8227FFEFE311FF78FA17FC07849C" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
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<subSubSection id="C34E60BAFFEFE310FF78FA17FE3885F4" box="[151,397,618,645]" pageId="48" pageNumber="49" type="nomenclature">
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<paragraph id="8BEB3331FFEFE310FF78FA17FE3885F4" blockId="48.[151,397,618,681]" box="[151,397,618,645]" pageId="48" pageNumber="49">
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<heading id="D0A3845DFFEFE310FF78FA17FE3885F4" bold="true" box="[151,397,618,645]" fontSize="11" level="2" pageId="48" pageNumber="49" reason="4">
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<emphasis id="B920EF23FFEFE310FF78FA17FE3885F4" bold="true" box="[151,397,618,645]" pageId="48" pageNumber="49">
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<taxonomicName id="4C5448B2FFEFE310FF78FA17FEF185F4" authority="Johnson, 2024" authorityName="Johnson" authorityYear="2024" box="[151,324,618,645]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="oudris" status="sp. nov.">
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<emphasis id="B920EF23FFEFE310FF78FA17FEF185F4" bold="true" box="[151,324,618,645]" italics="true" pageId="48" pageNumber="49">Harpes oudris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2135258FFEFE310FEA5FA17FE3885F4" box="[330,397,619,645]" pageId="48" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFEFE310FF78FAF3FEA985D8" box="[151,284,655,681]" pageId="48" pageNumber="49" type="description">
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<paragraph id="8BEB3331FFEFE310FF78FAF3FEA985D8" blockId="48.[151,397,618,681]" box="[151,284,655,681]" pageId="48" pageNumber="49">
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<figureCitation id="136F2FB4FFEFE310FF78FAF3FEA985D8" box="[151,284,655,681]" captionStart="PLATE 2" captionStartId="108.[151,234,1756,1780]" captionTargetBox="[202,1386,181,1732]" captionTargetId="figure-20@108.[202,1386,181,1732]" captionTargetPageId="108" captionText="PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232532" httpUri="https://zenodo.org/record/11232532/files/figure.png" pageId="48" pageNumber="49">Plate 2E–M</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFEFE310FF78FAAAFA9884F0" pageId="48" pageNumber="49" type="diagnosis">
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<paragraph id="8BEB3331FFEFE310FF78FAAAFA9884F0" blockId="48.[151,1437,726,2013]" pageId="48" pageNumber="49">
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<emphasis id="B920EF23FFEFE310FF78FAAAFEA78581" bold="true" box="[151,274,726,752]" pageId="48" pageNumber="49">Diagnosis.</emphasis>
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Axial furrows are convex and deep, and preglabellar furrow well defined. Tubercles and faint caeca on genal area. Perforations on genal roll same as on brim. Brim is moderately steeply-sloping and brim width ratio is between 0.68 and 0.75. Dorsal surface of external rim overhangs marginal band and carries tubercles. Prolongations over 90% of cephalic length (sag.), extension of girder meeting internal rim about halfway down prolongation. Internal rim in lateral view strongly wave-like in profile and distal end of prolongation sharply upturned.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFEFE310FF28FBF7FCFF84D4" box="[199,842,907,933]" pageId="48" pageNumber="49" type="etymology">
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<paragraph id="8BEB3331FFEFE310FF28FBF7FCFF84D4" blockId="48.[151,1437,726,2013]" box="[199,842,907,933]" pageId="48" pageNumber="49">
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<emphasis id="B920EF23FFEFE310FF28FBF7FEFB84D4" bold="true" box="[199,334,907,933]" pageId="48" pageNumber="49">Etymology.</emphasis>
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Named after Jbel Ou-Driss, the
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<typeStatus id="54EF8D93FFEFE310FD56FBF7FD5C84D4" box="[697,745,907,933]" pageId="48" pageNumber="49">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFEFE310FF28FBD2FD8C83B4" pageId="48" pageNumber="49" type="materials_examined">
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<paragraph id="8BEB3331FFEFE310FF28FBD2FC6F8328" blockId="48.[151,1437,726,2013]" pageId="48" pageNumber="49">
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<emphasis id="B920EF23FFEFE310FF28FBD2FE4F84B8" bold="true" box="[199,506,942,969]" pageId="48" pageNumber="49">Material and occurrence.</emphasis>
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<materialsCitation id="3B3C396CFFEFE310FDECFBD3FC608328" collectionCode="NHMUK" collectorName="Holoptype & It & Section & El Otfal Formation & Eifelian & Jbel Ou-Driss" location="lower Emsian" pageId="48" pageNumber="49" specimenCount="2" typeStatus="paratype">
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<collectorName id="26A156E7FFEFE310FDECFBD3FDCB84B8" box="[515,638,943,969]" pageId="48" pageNumber="49">Holoptype</collectorName>
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:
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<collectionCode id="ED45ABF4FFEFE310FD65FBD3FD4184B8" box="[650,756,943,969]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="48" pageNumber="49" type="Museum">NHMUK</collectionCode>
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<collectorName id="26A156E7FFEFE310FD12FBD3FCBB84B8" box="[765,782,943,969]" pageId="48" pageNumber="49">It</collectorName>
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29241,
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<figureCitation id="136F2FB4FFEFE310FC81FBD3FC6384B8" box="[878,982,943,969]" captionStart="PLATE 2" captionStartId="108.[151,234,1756,1780]" captionTargetBox="[202,1386,181,1732]" captionTargetId="figure-20@108.[202,1386,181,1732]" captionTargetPageId="108" captionText="PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232532" httpUri="https://zenodo.org/record/11232532/files/figure.png" pageId="48" pageNumber="49">Pl. 2I–M</figureCitation>
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, from
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<collectorName id="26A156E7FFEFE310FBCBFBD3FBCD84B8" box="[1060,1144,943,969]" pageId="48" pageNumber="49">Section</collectorName>
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1, horizon 5, (
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<figureCitation id="136F2FB4FFEFE310FACAFBD3FAD984B8" box="[1317,1388,943,969]" captionStart="FIGURE 6" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[154,1432,185,1829]" captionTargetId="figure-20@12.[151,1435,181,1833]" captionTargetPageId="12" captionText="FIGURE 6. Section 1 Jbel Ou-Driss" figureDoi="http://doi.org/10.5281/zenodo.11232515" httpUri="https://zenodo.org/record/11232515/files/figure.png" pageId="48" pageNumber="49">Fig. 6</figureCitation>
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),
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<collectorName id="26A156E7FFEFE310FA6CFBD3FEFA849C" pageId="48" pageNumber="49">El Otfal Formation</collectorName>
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,
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<collectorName id="26A156E7FFEFE310FEB7FBAFFE01849C" box="[344,436,979,1005]" pageId="48" pageNumber="49">Eifelian</collectorName>
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,
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<collectorName id="26A156E7FFEFE310FE51FBAFFDED849C" box="[446,600,979,1005]" pageId="48" pageNumber="49">Jbel Ou-Driss</collectorName>
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(
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<figureCitation id="136F2FB4FFEFE310FD89FBAFFD08849C" box="[614,701,979,1005]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="48" pageNumber="49">Fig. 2B</figureCitation>
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, Map 4, site 22), dorsal exoskeleton with thorax partially exposed.
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<typeStatus id="54EF8D93FFEFE310FF78FB8BFF488360" box="[151,253,1015,1041]" pageId="48" pageNumber="49" type="paratype">Paratype</typeStatus>
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:
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<collectionCode id="ED45ABF4FFEFE310FEE6FB8BFEC68360" box="[265,371,1015,1041]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="48" pageNumber="49" type="Museum">NHMUK</collectionCode>
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It 29323,
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<figureCitation id="136F2FB4FFEFE310FE05FB8BFDDE8360" box="[490,619,1015,1041]" captionStart="PLATE 22" captionStartId="128.[151,234,1645,1669]" captionTargetBox="[252,1335,181,1622]" captionTargetId="figure-20@128.[252,1335,181,1622]" captionTargetPageId="128" captionText="PLATE 22. A–E Maghroharpes cf. 1 laatchanensis, Ihandar Formation, Pragian, Jbel el Otfal (Fig. 2B, Map 9, site 8), dorsal exoskeleton of partial cephalon, NHMUK It 29311. A, dorsal, B, anterior and C, lateral view. D and E, dorsal view of brim perforations and occipital node and tubercles, respectively. F–M Maghroharpes aff.1 minutipunctus, Boutiskaoine Formation? lower Emsian. F–J dorsal exoskeleton of partial cephalon, NHMUK It 29322, Jbel Ou-Driss (North) (Fig. 2B, Map 4, site 7). F, dorsal, G, anterior and H, lateral view. I, lateral and J, dorsal view of eye lobe. K–M dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29323, Boutiskaouine Formation, lower Emsian, Ahware (Fig. 2B, Map 9, site 13). K, L and M, dorsal view of cephalon, brim perforations and eye lobe, respectively. N–Q Maghroharpes aff. 2 minutipunctus, Ihandar Formation Jbel Ou-Driss (West) (Fig. 2B, Map 4, site 20), dorsal exoskeleton of partial cephalon, NHMUK It 29324. N, dorsal, O, lateral and P, anterior view. Q, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232598" httpUri="https://zenodo.org/record/11232598/files/figure.png" pageId="48" pageNumber="49">Pl. 22K–M</figureCitation>
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, from a
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<location id="8E8B65EAFFEFE310FD24FB8BFCD28360" LSID="urn:lsid:plazi:treatment:03FD8227FFEFE311FF78FA17FC07849C:8E8B65EAFFEFE310FD24FB8BFCD28360" box="[715,871,1015,1041]" name="lower Emsian" pageId="48" pageNumber="49">lower Emsian</location>
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horizon, Boutiskaouine Formation, Awhare, Iferd Nou Haouar (
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<figureCitation id="136F2FB4FFEFE310FEDAFC67FE258344" box="[309,400,1051,1077]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="48" pageNumber="49">Fig. 2B</figureCitation>
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, Map 9, site 13), dorsal exoskeleton of partial cephalon.
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<typeStatus id="54EF8D93FFEFE310FBC9FC67FB398344" box="[1062,1164,1051,1077]" pageId="48" pageNumber="49" type="paratype">Paratype</typeStatus>
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:
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<collectionCode id="ED45ABF4FFEFE310FB76FC67FAB68344" box="[1177,1283,1051,1077]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="48" pageNumber="49" type="Museum">NHMUK</collectionCode>
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It 29240,
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<figureCitation id="136F2FB4FFEFE310FA91FC67FF6E8328" captionStart="PLATE 2" captionStartId="108.[151,234,1756,1780]" captionTargetBox="[202,1386,181,1732]" captionTargetId="figure-20@108.[202,1386,181,1732]" captionTargetPageId="108" captionText="PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232532" httpUri="https://zenodo.org/record/11232532/files/figure.png" pageId="48" pageNumber="49">Pl. 2E–H</figureCitation>
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, from type horizon, site 21, dorsal exoskeleton with thorax enrolled
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BEB3331FFEFE310FF28FC1FFD8C83B4" blockId="48.[151,1437,726,2013]" pageId="48" pageNumber="49">
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Other material:
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<taxonomicName id="4C5448B2FFEFE310FE9CFC1FFE4A830C" authorityName="Johnson" authorityYear="2024" box="[371,511,1123,1149]" class="Trilobita" family="Harpetidae" genus="Harpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="oudris">
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<emphasis id="B920EF23FFEFE310FE9CFC1FFE3A830D" box="[371,399,1123,1148]" italics="true" pageId="48" pageNumber="49">H.</emphasis>
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||
cf.
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<emphasis id="B920EF23FFEFE310FE5BFC1FFE4A830C" box="[436,511,1123,1149]" italics="true" pageId="48" pageNumber="49">oudris</emphasis>
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</taxonomicName>
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<emphasis id="B920EF23FFEFE310FE10FC1FFDB6830C" box="[511,515,1123,1149]" italics="true" pageId="48" pageNumber="49">.</emphasis>
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||
<collectionCode id="ED45ABF4FFEFE310FDE8FC1FFDC4830C" box="[519,625,1123,1149]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="48" pageNumber="49" type="Museum">NHMUK</collectionCode>
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It 29242,
|
||
<figureCitation id="136F2FB4FFEFE310FD33FC1FFCF3830C" box="[732,838,1123,1149]" captionStart="PLATE 6" captionStartId="112.[151,234,1747,1771]" captionTargetBox="[213,1375,181,1723]" captionTargetId="figure-20@112.[213,1375,181,1723]" captionTargetPageId="112" captionText="PLATE 6. A–H Harpes cf. oudris, dorsal exoskeleton with ventral surfaces exposed on anterior half of cephalon, NHMUK It 29242; from El Otfal Formation, Eifelian, horizon 4, Section 1 (Fig. 6), Jbel Ou-Driss (Fig. 2B, Map 4, site 10). A, dorsal, B, anterior and C, lateral view. D, lateral view of glabella (uncoated) and muscle insertion scars, E, ventral view of anterior half of cephalon, F, ventral view of hypostome in place, G, ventral view of anterior of rim, H, dorsal view of brim perforations. I–N Harpes cf. lahceni, dorsal exoskeleton of partial cephalon only, NHMUK It 29235, from “Erbenochile Couche”, Timrhanrhart Formation, Emsian, 23.8 km SW of Foum Zguid (Fig. 2A, Map 2, site 6). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations and M and N, dorsal and lateral view of eye lobe respectively. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232549" httpUri="https://zenodo.org/record/11232549/files/figure.png" pageId="48" pageNumber="49">Pl. 6A–H</figureCitation>
|
||
, from horizon 4, Section 1 (
|
||
<figureCitation id="136F2FB4FFEFE310FB9EFC1FFB07830C" box="[1137,1202,1123,1149]" captionStart="FIGURE 6" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[154,1432,185,1829]" captionTargetId="figure-20@12.[151,1435,181,1833]" captionTargetPageId="12" captionText="FIGURE 6. Section 1 Jbel Ou-Driss" figureDoi="http://doi.org/10.5281/zenodo.11232515" httpUri="https://zenodo.org/record/11232515/files/figure.png" pageId="48" pageNumber="49">Fig. 6</figureCitation>
|
||
), El Otfal Formation, Jbel Ou-Driss (
|
||
<figureCitation id="136F2FB4FFEFE310FEAFFCFBFE2283D0" box="[320,407,1159,1185]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="48" pageNumber="49">Fig. 2B</figureCitation>
|
||
, Map 4, site 10), an articulated dorsal exoskeleton with thorax enrolled with ventral surfaces exposed on anterior half of cephalon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFFEFE311FF28FCB2FDDE86F8" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="description">
|
||
<paragraph id="8BEB3331FFEFE310FF28FCB2FCC68120" blockId="48.[151,1437,726,2013]" pageId="48" pageNumber="49">
|
||
<emphasis id="B920EF23FFEFE310FF28FCB2FEEC8399" bold="true" box="[199,345,1230,1256]" pageId="48" pageNumber="49">Description.</emphasis>
|
||
Cephalon ovoid with widest (tr.) point at mid-point between eye lobes and alae. Genicranium narrow, widest at posterior border. Glabella fairly broad, width (tr.) 75% of length (sag.) of glabella anterior of S0, and with rounded crest and tubercles but no pitting. S1 rising out of axial furrow, sloping backwards and curving around top of muscle insertion area. No furrows at S2, S3 or S4 and L1 inflated. Axial furrows converge strongly anteriorly at nearly
|
||
<superScript id="7C219E79FFEFE310FE32FD23FDB4821D" attach="left" box="[477,513,1374,1401]" fontSize="6" pageId="48" pageNumber="49">200</superScript>
|
||
to axis and cross junction between alae and L1. Preglabellar furrow distinct and bowed upwards in anterior view. Occipital furrow deep, wide, wider than posterior border and straight in dorsal view. Occipital ring broad (sag.), broader than posterior border (exsag.) and same height in lateral view as glabella anterior to S0. Occipital node large, alae anterolaterally directed and large (tr.), over 35% of length (sag.) of glabella anterior to S0 and reaching around 70% of way across genal area. Alar furrow shallow, well-defined. Interalar furrow and ridge and alar depression present. Posterior border short (tr.), around 23% of width (tr.) of occipital ring. Preglabellar field narrow, length (sag.) around 17% of width (tr.).
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEFE310FF28FE27FAD8803C" blockId="48.[151,1437,726,2013]" pageId="48" pageNumber="49">
|
||
Genal area narrow, width (tr.) about 50% of length (exsag.) with faint caeca and pits and tubercles. Genal ridge present and eye lobes cone-shaped, strongly inflated and large (exsag.), just under 40% of width (tr.) of glabella at S1. Visual surface close to inner margin of fringe with eye lobes extending to edge of axial furrow. Eyes with two lenses, each lens rectangle shape with rounded corners and of approximately same size, and no eye ridge. Course of inner margin of fringe across anterior of genal area, convex. Boss barely inflated and not reaching down to girder. No girder kink and girder moderately narrow. Genal role sloping gently anteriorly at around
|
||
<superScript id="7C219E79FFEFE310FB40FF73FB66806D" attach="left" box="[1199,1235,1806,1833]" fontSize="6" pageId="48" pageNumber="49">470</superScript>
|
||
and more steeply laterally at around
|
||
<superScript id="7C219E79FFEFE310FE85FF4FFE3B8031" attach="none" box="[362,398,1842,1869]" fontSize="6" pageId="48" pageNumber="49">630</superScript>
|
||
. No caeca on genal roll. Perforations are approximately same diameter as those of brim.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEFE311FF28FF2BFB60866C" blockId="48.[151,1437,726,2013]" lastBlockId="49.[151,1437,151,1005]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">Brim slightly concave (sag.), straight laterally, width (sag.) 30% of cephalic length (sag.) and sloping moderately steeply at around 22%. Brim width ratio around 0.75. Standardised brim perforations becoming finer away from girder towards the centre, where they have a standardised diameter of around 145μm. Light caeca reaching across brim. External rim stout with fine tubercles on dorsal surface. Marginal band near vertical, overhung by external rim and with tubercles. Prolongations moderately long (exsag.) at around 87% of cephalic length (sag.) with extension of brim steepening quickly to near vertical. In dorsal view, internal and external rims curve adaxially and in lateral view, profile of internal rim strongly wave-like. Tubercles on dorsal surface of internal rim, large row of perforations below; and genal spine short. Extension of girder meets internal rim halfway down prolongation.</paragraph>
|
||
<paragraph id="8BEB3331FFEEE311FF3AF95BFB9F8614" blockId="49.[151,1437,151,1005]" pageId="49" pageNumber="50">Thorax with over 20 thoracic segments with no tubercles or pitting on axial rings. Pleural furrows broad (tr.), deep and u-shaped, and outer portion of pleurae straight with rounded distal ends.</paragraph>
|
||
<paragraph id="8BEB3331FFEEE311FF28F913FDDE86F8" blockId="49.[151,1437,151,1005]" box="[199,619,367,393]" pageId="49" pageNumber="50">Pygidium and hypostome not known.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFFEEE311FF28F9EFFC07849C" pageId="49" pageNumber="50" type="discussion">
|
||
<paragraph id="8BEB3331FFEEE311FF28F9EFFB808684" blockId="49.[151,1437,151,1005]" pageId="49" pageNumber="50">
|
||
<emphasis id="B920EF23FFEEE311FF28F9EFFE8E86DC" bold="true" box="[199,315,403,429]" pageId="49" pageNumber="50">Remarks.</emphasis>
|
||
<taxonomicName id="4C5448B2FFEEE311FEAAF9EFFE5C86DC" authorityName="Johnson" authorityYear="2024" box="[325,489,403,429]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oudris">
|
||
<emphasis id="B920EF23FFEEE311FEAAF9EFFE5C86DC" box="[325,489,403,429]" italics="true" pageId="49" pageNumber="50">Harpes oudris</emphasis>
|
||
</taxonomicName>
|
||
lacks three genus diagnostic characters shown in
|
||
<tableCitation id="C6D6068AFFEEE311FBC3F9EFFB2486DC" box="[1068,1169,402,429]" captionStart="TABLE 5" captionStartId="182.[151,239,503,529]" captionText="TABLE 5b. Proposed diagnostic characters for Devonian harpetid genera (continued)" pageId="49" pageNumber="50">Table 5a</tableCitation>
|
||
. Its brim width ratio is above the upper limit of 0.68, there are light caeca reaching across brim and the extension of the girder meets the internal rim halfway down the prolongation rather than one third of the way down.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEEE311FF28F983FBA38448" blockId="49.[151,1437,151,1005]" pageId="49" pageNumber="50">
|
||
<taxonomicName id="4C5448B2FFEEE311FF28F983FEDC8568" authorityName="Johnson" authorityYear="2024" box="[199,361,511,537]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oudris">
|
||
<emphasis id="B920EF23FFEEE311FF28F983FEDC8568" box="[199,361,511,537]" italics="true" pageId="49" pageNumber="50">Harpes oudris</emphasis>
|
||
</taxonomicName>
|
||
is similar to but differs from
|
||
<taxonomicName id="4C5448B2FFEEE311FD59F983FCEB8568" box="[694,862,511,537]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFEEE311FD59F983FCEB8568" box="[694,862,511,537]" italics="true" pageId="49" pageNumber="50">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
in having faint caeca on genal areas; axial furrows that are convex rather than straight (reflecting on the difference in glabella outline) and a distinct preglabellar furrow. Alae and eye lobes more inflated and posterior border (tr.) shorter, 25% of width (tr.) of occipital ring rather than between 30% and 34% in the case of
|
||
<taxonomicName id="4C5448B2FFEEE311FD96FA17FC9785F4" box="[633,802,619,645]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFEEE311FD96FA17FC9785F4" box="[633,802,619,645]" italics="true" pageId="49" pageNumber="50">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
. Glabella of
|
||
<taxonomicName id="4C5448B2FFEEE311FC58FA17FB9785F4" authorityName="Johnson" authorityYear="2024" box="[951,1058,619,645]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oudris">
|
||
<emphasis id="B920EF23FFEEE311FC58FA17FB9785F4" box="[951,1058,619,645]" italics="true" pageId="49" pageNumber="50">H. oudris</emphasis>
|
||
</taxonomicName>
|
||
not joined to anterior boss, genal roll steeper anteriorly and perforations on genal roll same size as on brim, not smaller. Brim of
|
||
<taxonomicName id="4C5448B2FFEEE311FB5FFAF3FAAC85D8" authorityName="Johnson" authorityYear="2024" box="[1200,1305,655,681]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oudris">
|
||
<emphasis id="B920EF23FFEEE311FB5FFAF3FAAC85D8" box="[1200,1305,655,681]" italics="true" pageId="49" pageNumber="50">H. oudris</emphasis>
|
||
</taxonomicName>
|
||
steeper and has light caeca reaching across it. Brim width ratio over 0.68, whereas that of
|
||
<taxonomicName id="4C5448B2FFEEE311FBEDFACFFB1E85BC" box="[1026,1195,691,717]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFEEE311FBEDFACFFB1E85BC" box="[1026,1195,691,717]" italics="true" pageId="49" pageNumber="50">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
is under 0.55. Dorsal surface of rim has fine tubercles and overhangs marginal band. Prolongations of
|
||
<taxonomicName id="4C5448B2FFEEE311FBF9FAABFB358580" authorityName="Johnson" authorityYear="2024" box="[1046,1152,727,753]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oudris">
|
||
<emphasis id="B920EF23FFEEE311FBF9FAABFB358580" box="[1046,1152,727,753]" italics="true" pageId="49" pageNumber="50">H. oudris</emphasis>
|
||
</taxonomicName>
|
||
are shorter and extension of girder meets internal rim halfway down prolongation, not a third of the way down.
|
||
<taxonomicName id="4C5448B2FFEEE311FB88FA87FABF8464" authorityName="Johnson" authorityYear="2024" box="[1127,1290,763,789]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oudris">
|
||
<emphasis id="B920EF23FFEEE311FB88FA87FABF8464" box="[1127,1290,763,789]" italics="true" pageId="49" pageNumber="50">Harpes oudris</emphasis>
|
||
</taxonomicName>
|
||
has 20 to 24 thoracic segments, not the 27 of
|
||
<emphasis id="B920EF23FFEEE311FDEBFB63FD068448" box="[516,691,799,825]" italics="true" pageId="49" pageNumber="50">
|
||
<taxonomicName id="4C5448B2FFEEE311FDEBFB63FD1A8448" box="[516,687,799,825]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="boudibensis">H. boudibensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and pleural furrows are deeper.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEEE311FF28FB3FFB6884F0" blockId="49.[151,1437,151,1005]" pageId="49" pageNumber="50">
|
||
<taxonomicName id="4C5448B2FFEEE311FF28FB3FFEE0842C" authorityName="Johnson" authorityYear="2024" box="[199,341,835,861]" class="Trilobita" family="Harpetidae" genus="Harpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oudris">
|
||
<emphasis id="B920EF23FFEEE311FF28FB3FFF56842D" box="[199,227,835,860]" italics="true" pageId="49" pageNumber="50">H.</emphasis>
|
||
cf.
|
||
<emphasis id="B920EF23FFEEE311FEE2FB3FFEE0842C" box="[269,341,835,861]" italics="true" pageId="49" pageNumber="50">oudris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136F2FB4FFEEE311FE8BFB3FFE23842C" box="[356,406,835,861]" captionStart="PLATE 6" captionStartId="112.[151,234,1747,1771]" captionTargetBox="[213,1375,181,1723]" captionTargetId="figure-20@112.[213,1375,181,1723]" captionTargetPageId="112" captionText="PLATE 6. A–H Harpes cf. oudris, dorsal exoskeleton with ventral surfaces exposed on anterior half of cephalon, NHMUK It 29242; from El Otfal Formation, Eifelian, horizon 4, Section 1 (Fig. 6), Jbel Ou-Driss (Fig. 2B, Map 4, site 10). A, dorsal, B, anterior and C, lateral view. D, lateral view of glabella (uncoated) and muscle insertion scars, E, ventral view of anterior half of cephalon, F, ventral view of hypostome in place, G, ventral view of anterior of rim, H, dorsal view of brim perforations. I–N Harpes cf. lahceni, dorsal exoskeleton of partial cephalon only, NHMUK It 29235, from “Erbenochile Couche”, Timrhanrhart Formation, Emsian, 23.8 km SW of Foum Zguid (Fig. 2A, Map 2, site 6). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations and M and N, dorsal and lateral view of eye lobe respectively. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232549" httpUri="https://zenodo.org/record/11232549/files/figure.png" pageId="49" pageNumber="50">Pl. 6</figureCitation>
|
||
A-H) differs from
|
||
<taxonomicName id="4C5448B2FFEEE311FD89FB3FFD65842C" authorityName="Johnson" authorityYear="2024" box="[614,720,835,861]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oudris">
|
||
<emphasis id="B920EF23FFEEE311FD89FB3FFD65842C" box="[614,720,835,861]" italics="true" pageId="49" pageNumber="50">H. oudris</emphasis>
|
||
</taxonomicName>
|
||
in having a gently convex brim, a brim width ratio which is just over 10% higher at 0.77, and an internal rim that, in lateral view, is straight rather than wave-like.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFEEE311FF28FBF7FC07849C" blockId="49.[151,1437,151,1005]" pageId="49" pageNumber="50">
|
||
The
|
||
<taxonomicName id="4C5448B2FFEEE311FF17FBF7FE3684D4" authorityName="Johnson" authorityYear="2024" box="[248,387,907,933]" class="Trilobita" family="Harpetidae" genus="Harpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oudris">
|
||
<emphasis id="B920EF23FFEEE311FF17FBF7FEA184D5" box="[248,276,907,932]" italics="true" pageId="49" pageNumber="50">H.</emphasis>
|
||
cf.
|
||
<emphasis id="B920EF23FFEEE311FED4FBF7FE3684D4" box="[315,387,907,933]" italics="true" pageId="49" pageNumber="50">oudris</emphasis>
|
||
</taxonomicName>
|
||
material contains an exposed hypostome (see
|
||
<figureCitation id="136F2FB4FFEEE311FC6FFBF7FC5784D4" box="[896,994,907,933]" captionStart="PLATE 6" captionStartId="112.[151,234,1747,1771]" captionTargetBox="[213,1375,181,1723]" captionTargetId="figure-20@112.[213,1375,181,1723]" captionTargetPageId="112" captionText="PLATE 6. A–H Harpes cf. oudris, dorsal exoskeleton with ventral surfaces exposed on anterior half of cephalon, NHMUK It 29242; from El Otfal Formation, Eifelian, horizon 4, Section 1 (Fig. 6), Jbel Ou-Driss (Fig. 2B, Map 4, site 10). A, dorsal, B, anterior and C, lateral view. D, lateral view of glabella (uncoated) and muscle insertion scars, E, ventral view of anterior half of cephalon, F, ventral view of hypostome in place, G, ventral view of anterior of rim, H, dorsal view of brim perforations. I–N Harpes cf. lahceni, dorsal exoskeleton of partial cephalon only, NHMUK It 29235, from “Erbenochile Couche”, Timrhanrhart Formation, Emsian, 23.8 km SW of Foum Zguid (Fig. 2A, Map 2, site 6). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations and M and N, dorsal and lateral view of eye lobe respectively. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232549" httpUri="https://zenodo.org/record/11232549/files/figure.png" pageId="49" pageNumber="50">Pl. 6 F, E</figureCitation>
|
||
) that differs from that of
|
||
<taxonomicName id="4C5448B2FFEEE311FB19FBF7FA2984D4" box="[1270,1436,907,933]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFEEE311FB19FBF7FA2984D4" box="[1270,1436,907,933]" italics="true" pageId="49" pageNumber="50">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="136F2FB4FFEEE311FF22FBD3FE8484B8" box="[205,305,943,969]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="49" pageNumber="50">Pl. 1H, I</figureCitation>
|
||
) in having a well-defined middle furrow, with raised maculae on the posterior edge of the furrow. These do not appear to be present on the hypostome of
|
||
<taxonomicName id="4C5448B2FFEEE311FCECFBAFFC1E849C" box="[771,939,979,1005]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFEEE311FCECFBAFFC1E849C" box="[771,939,979,1005]" italics="true" pageId="49" pageNumber="50">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |