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<document id="B16F65E6ED6CB051094C40C6CD50C0DC" ID-DOI="10.1649/0010-065X-73.3.473" ID-ISSN="1938-4394" ID-Zenodo-Dep="5377336" ID-ZooBank="urn:lsid:zoobank.org:pub:234159CD-885B-4D4B-8ECF-8C2D1A930ADE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630282929292" checkinUser="valdenar" docAuthor="Suzumura, Alyssa Lee, Kobayashi, Norio &amp; Ôhara, Masahiro" docDate="2019" docId="03FD136EB94BA93FFED5EEA4FCDDFD90" docLanguage="en" docName="0010-065X-73.3.473.pdf" docOrigin="The Coleopterists Bulletin 73 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-73.3.473" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Cercyon Leach 1817" docType="treatment" docVersion="4" lastPageNumber="475" masterDocId="FFC46B16B948A93CFFC4ED74FFACFFF5" masterDocTitle="Revision of the Beach-Dwelling Species of Cercyon Leach (Coleoptera: Hydrophilidae) of the West Coast of North America" masterLastPageNumber="493" masterPageNumber="473" pageNumber="475" updateTime="1699213074183" updateUser="plazi">
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<mods:title id="71CF717A7E8B33968881F653B4EBB31D">Revision of the Beach-Dwelling Species of Cercyon Leach (Coleoptera: Hydrophilidae) of the West Coast of North America</mods:title>
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<mods:namePart id="1A66FDB5604DFBE2D8613AAEF0F43B04">Suzumura, Alyssa Lee</mods:namePart>
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<mods:namePart id="C4DC0D6EB8F49FBE3760337A1E9D58F8">Kobayashi, Norio</mods:namePart>
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<mods:namePart id="BD1A6A7E4DF3EAA18A145D3EC4A28788">Ôhara, Masahiro</mods:namePart>
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<mods:title id="8189346D14412056BFB16FD68B647E1D">The Coleopterists Bulletin</mods:title>
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<subSubSection id="C34EF1F3B94BA93FFED5EEA4FD83FC10" box="[273,559,975,997]" pageId="3" pageNumber="475" type="nomenclature">
<paragraph id="8BEBA278B94BA93FFED5EEA4FD83FC10" blockId="3.[273,559,975,997]" box="[273,559,975,997]" pageId="3" pageNumber="475">
<heading id="D0A31514B94BA93FFED5EEA4FD83FC10" box="[273,559,975,997]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="475" reason="2">
<emphasis id="B9207E6AB94BA93FFED5EEA4FD83FC10" bold="true" box="[273,559,975,997]" pageId="3" pageNumber="475">
Genus
<taxonomicName id="4C54D9FBB94BA93FFE9DEEBBFD83FC10" ID-CoL="8VRKH" authority="Leach, 1817" authorityName="Leach" authorityYear="1817" box="[345,559,975,997]" class="Insecta" family="Hydrophilidae" genus="Cercyon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="B9207E6AB94BA93FFE9DEEBBFE07FC11" bold="true" box="[345,427,975,996]" italics="true" pageId="3" pageNumber="475">Cercyon</emphasis>
<bibRefCitation id="EFC5DF89B94BA93FFE77EEA4FD83FC10" author="Leach, W. E." box="[435,559,976,997]" pageId="3" pageNumber="475" pagination="121 - 150" refId="ref13528" refString="Leach, W. E. 1817. Descriptions of new or interesting animals. The Zoological Miscellany 3 (151): 121 - 150." type="journal article" year="1817">Leach, 1817</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34EF1F3B94BA93FFF01EE8EFCDDFD90" pageId="3" pageNumber="475" type="diagnosis">
<paragraph id="8BEBA278B94BA93FFF01EE8EFCA2FEAE" blockId="3.[173,659,1018,1730]" lastBlockId="3.[707,1193,194,613]" pageId="3" pageNumber="475">
<emphasis id="B9207E6AB94BA93FFF01EE8EFE80FBFA" bold="true" box="[197,300,1018,1039]" pageId="3" pageNumber="475">Diagnosis.</emphasis>
Body variously oval, rather flat to strongly convex; pronotum and elytra forming a continuous curve in lateral view. Dorsal surface glabrous, with punctation and sometimes with microsculpture or indistinct pubescence; pronotum without longitudinal grooves or pit-like impressions except often with 2 lateroposterior depressions; color ranging from pale yellow to black. Ventral surface covered in dense, short pubescence except raised surfaces of meso- and metaventrites. Antennae 9-segmented, terminal 3 antennomeres forming compact club; bases visible dorsally, not hidden by lateral margin of head. Clypeus not separated from frons by a distinct suture, often truncate, extending anterior to eyes when viewed dorsally; mentum of variable shape, anterior margin subtruncate or emarginate, often impressed anteromedially. Maxillary palpi shorter than antennae, 2
<superScript id="7C210F30B94BA93FFF7DE8A7FF67FA14" attach="left" box="[185,203,1491,1505]" fontSize="6" pageId="3" pageNumber="475">nd</superScript>
palpomere swollen apically, length of apical segment equal to or slightly longer than penultimate; male maxilla bearing adhesion discs, though may be indistinct in smaller species. Elytra with distinct epipleura, at least in basal half. Middle portion of prosternum elevated to form a tectiform longitudinal keel, not differentiated laterally from antennal excavations. Median portion of mesoventrite elevated to form a median keel or plate; contacting metaventrite at a single point or separated from it by a narrow gap. Outer margin of profemora rounded; tarsal formula 5-5-5 with 1
<superScript id="7C210F30B94BA93FFB59ED85FB05FF0A" attach="none" box="[1181,1193,241,255]" fontSize="6" pageId="3" pageNumber="475">st</superScript>
tarsomere always longer than 2
<superScript id="7C210F30B94BA93FFC2EEC78FC50FEEF" attach="none" box="[1002,1020,268,282]" fontSize="6" pageId="3" pageNumber="475">nd</superScript>
. Male genitalia of trilobed
<typeStatus id="54EF1CDAB94BA93FFCD1EC58FCECFEB4" box="[789,832,300,321]" pageId="3" pageNumber="475">type</typeStatus>
, as in
<figureCitation id="136FBEFDB94BA93FFC42EC58FC6FFEB4" box="[902,963,300,321]" captionStart="Fig" captionStartId="11.[197,228,1593,1612]" captionTargetBox="[177,1187,196,1575]" captionTargetId="figure-0@11.[175,1190,194,1577]" captionTargetPageId="11" captionText="Fig. 7. Cercyon species, male genitalia (parameres and basal piece, median lobe, and ninth ventrite, respectively). AC) C. whuljensis (BC14MO-025), DF) C. dux (BC14MO-030), GI) C. fimbriatus (BC16MO-033), JL) C. luniger (WA18AS-008), MO) C. setulosus (WA16AS-006). Bp = basal piece; Co = corona; Ed = ejaculatory duct; Pm = parameres. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5377350" httpUri="https://zenodo.org/record/5377350/files/figure.png" pageId="3" pageNumber="475">Fig. 7</figureCitation>
. Female genitalia as in
<figureCitation id="136FBEFDB94BA93FFD07EC32FCA6FEAE" box="[707,778,326,347]" captionStart="Fig" captionStartId="17.[197,228,1111,1130]" captionTargetBox="[177,655,196,1093]" captionTargetId="figure-513@17.[175,657,194,1095]" captionTargetPageId="17" captionText="Fig. 13. Cercyon setulosus, female genitalia (WA17AS-004). b = bursa; g = gonocoxite; IX = ninth segment; ltg = laterotergite of segment VIII; s = spermatheca; sd = spermathecal duct; sg = spermathecal gland." figureDoi="http://doi.org/10.5281/zenodo.5377365" httpUri="https://zenodo.org/record/5377365/files/figure.png" pageId="3" pageNumber="475">Fig. 13</figureCitation>
.
</paragraph>
<paragraph id="8BEBA278B94BA93FFD1FEC15FCDDFD90" blockId="3.[707,1193,194,613]" pageId="3" pageNumber="475">
<emphasis id="B9207E6AB94BA93FFD1FEC15FC68FE83" bold="true" box="[731,964,353,374]" pageId="3" pageNumber="475">Notes on Identification.</emphasis>
Along with members of the
<taxonomicName id="4C54D9FBB94BA93FFD29EC08FCDEFE64" box="[749,882,380,401]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
,
<taxonomicName id="4C54D9FBB94BA93FFC47EC0FFC7FFE64" authorityName="Leach" authorityYear="1817" box="[899,979,379,401]" class="Insecta" family="Hydrophilidae" genus="Cercyon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="B9207E6AB94BA93FFC47EC0FFC7FFE64" box="[899,979,379,401]" italics="true" pageId="3" pageNumber="475">Cercyon</emphasis>
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is one of the more common beach-dwelling beetle taxa. As it is the only beach-dwelling hydrophilid genus in most regions, excluding
<collectingCountry id="F343E2E8B94BA93FFCBBECBFFC79FE15" box="[895,981,459,480]" name="Australia" pageId="3" pageNumber="475">Australia</collectingCountry>
and
<collectingCountry id="F343E2E8B94BA93FFBCEECBFFB23FE15" box="[1034,1167,459,480]" name="New Zealand" pageId="3" pageNumber="475">New Zealand</collectingCountry>
, it can be easily differentiated from other beach beetles by its evenly oval habitus, uninterrupted outline between pronotum and elytra, presence of a mesoventral tablet, maxillary adhesion discs, and clubbed antennae.
</paragraph>
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