227 lines
34 KiB
XML
227 lines
34 KiB
XML
<document id="5470A2028BB99D2C57F01BF46778EAD6" ID-DOI="10.2478/aemnp-2018-0013" ID-GBIF-Dataset="7b94dd56-d3a7-4462-8da2-72ceb277835f" ID-ISSN="1804-6487" ID-Zenodo-Dep="3699163" ID-ZooBank="BF2CDC12-C555-4187-BFAB-274C9C674A5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583496890333" checkinUser="tatiana" docAuthor="Löbl, Ivan" docDate="2018" docId="03F94B1B217AFF896447E339005D85DC" docLanguage="en" docName="ActaEntMusNatPra.58.1.151-165.pdf.imf" docOrigin="Acta Entomologica Musei Nationalis Pragae 58 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Scaphisoma ancora Löbl 2018, sp. nov." docType="treatment" docVersion="13" lastPageNumber="153" masterDocId="FFC033632178FF8B656AE114055B8309" masterDocTitle="A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae)" masterLastPageNumber="165" masterPageNumber="151" pageNumber="153" updateTime="1698725294610" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
|
||
<mods:mods id="8A1AD51C5DBD9E5ABB54FB3F2BCD65BE" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="704A577AB9BEF8CACF1DF6E68FAAAF56">
|
||
<mods:title id="3AA9521BBBA3A365985EDB666C703BF2">A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="4C182A5D2DA0DB344EA735C16B5DBD43" type="personal">
|
||
<mods:role id="1AA30479B567963C2758AA40396171E0">
|
||
<mods:roleTerm id="25811E7D8B02BE497D58C21B0D564FC2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="ABBE1E210122D9E004894E8E49F8A125">Löbl, Ivan</mods:namePart>
|
||
<mods:affiliation id="3D81962BA3C89DC797880E1A16378342">Muséum d’histoire naturelle, Route de Malagnou 1, CH- 1208 Geneva, Switzerland</mods:affiliation>
|
||
<mods:nameIdentifier id="D50B3A329795FDD0B56E56F1BFBE56D0" type="email">ivan.lobl@bluewin.ch</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="DFEC71E4312AB7E6F4B927DB9CC19A42">text</mods:typeOfResource>
|
||
<mods:relatedItem id="84053F4471640081878C2707EB79D62C" type="host">
|
||
<mods:titleInfo id="7752B5603059EBF86CE5C48EDEE03C56">
|
||
<mods:title id="F4978F8846656C64117FECB6A1845A4A">Acta Entomologica Musei Nationalis Pragae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B262D306EFB24009F28474239894C87B">
|
||
<mods:date id="4512C7D0FBFDF86A4B37AE8D9A7B0F4E">2018</mods:date>
|
||
<mods:detail id="FFBBFEC9198F568CF6446EA81DA6139E" type="pubDate">
|
||
<mods:number id="3B218E7AAF9E950D726604559D463C83">2018-05-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="E533296349A3DD036976AC4993392C78" type="volume">
|
||
<mods:number id="5EA433408A470EDE302E4AC06F665830">58</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F79884F4F9F13CAF742C38EEE627FDA4" type="issue">
|
||
<mods:number id="CF8A20E8CA2BAF97B4F5633BC8A6FE37">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="77D7B40CD45316443410F4835B37E6FB" unit="page">
|
||
<mods:start id="7B9561DDAF5C86B2BF7750E8C719026C">151</mods:start>
|
||
<mods:end id="DAA4040EB59251A118EC30E4D628A734">165</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="991DB8AD3896427CC40443B9A06C17BF">journal article</mods:classification>
|
||
<mods:identifier id="BF915402EF1BCA8BB3E4DE66348B2ECB" type="DOI">10.2478/aemnp-2018-0013</mods:identifier>
|
||
<mods:identifier id="8BBD0D63DD13C2BEA8FEE2B38D8BC51F" type="GBIF-Dataset">7b94dd56-d3a7-4462-8da2-72ceb277835f</mods:identifier>
|
||
<mods:identifier id="1C9A88C59441E36D0E67380F59733528" type="ISSN">1804-6487</mods:identifier>
|
||
<mods:identifier id="97FC30CDB07ECDD2CD79E64C2DA0A123" type="Zenodo-Dep">3699163</mods:identifier>
|
||
<mods:identifier id="A321AF9BE014871ADDD82068C6FEA90B" type="ZooBank">BF2CDC12-C555-4187-BFAB-274C9C674A5D</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03F94B1B217AFF896447E339005D85DC" ID-DOI="http://doi.org/10.5281/zenodo.3706171" ID-GBIF-Taxon="162467443" ID-Zenodo-Dep="3706171" LSID="urn:lsid:plazi:treatment:03F94B1B217AFF896447E339005D85DC" httpUri="http://treatment.plazi.org/id/03F94B1B217AFF896447E339005D85DC" lastPageNumber="153" pageId="2" pageNumber="153">
|
||
<subSubSection id="C34AA986217AFF896447E3390739814D" box="[301,610,557,581]" pageId="2" pageNumber="153" type="nomenclature">
|
||
<paragraph id="8BEFFA0D217AFF896447E3390739814D" blockId="2.[301,610,557,606]" box="[301,610,557,581]" pageId="2" pageNumber="153">
|
||
<heading id="D0A74D61217AFF896447E3390739814D" box="[301,610,557,581]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="153" reason="3">
|
||
<emphasis id="B924261F217AFF896447E3390739814D" bold="true" box="[301,610,557,581]" pageId="2" pageNumber="153">
|
||
<taxonomicName id="4C50818E217AFF896447E339075D814C" ID-CoL="6Y5L7" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[301,518,557,581]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="ancora" status="sp. nov.">
|
||
<emphasis id="B924261F217AFF896447E339075D814C" bold="true" box="[301,518,557,581]" italics="true" pageId="2" pageNumber="153">Scaphisoma ancora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2179B64217AFF896767E3390739814D" box="[525,610,557,580]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA986217AFF89641CE35F07428157" box="[374,537,587,606]" pageId="2" pageNumber="153" type="description">
|
||
<paragraph id="8BEFFA0D217AFF89641CE35F07428157" blockId="2.[301,610,557,606]" box="[374,537,587,606]" pageId="2" pageNumber="153">
|
||
(
|
||
<figureCitation id="136BE688217AFF896416E35F04888157" box="[380,467,587,606]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="2" pageNumber="153">Figs 6–12</figureCitation>
|
||
,
|
||
<figureCitation id="136BE688217AFF8964B6E35F07498157" box="[476,530,587,606]" captionStart="Figs 50–53" captionStartId="12.[151,188,2027,2046]" captionTargetBox="[206,1382,457,2010]" captionTargetId="figure@12.[205,1382,456,2011]" captionTargetPageId="12" captionText="Figs 50–53. Habitus of Scaphisoma. 50, 51 – S. ancora sp. nov. (50 – male, 51 – female); 52 – S. sumpichi sp. nov., male; 53 – S. versicoloreum sp. nov., male." figureDoi="http://doi.org/10.5281/zenodo.3699171" httpUri="https://zenodo.org/record/3699171/files/figure.png" pageId="2" pageNumber="153">50, 51</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA986217AFF8965FDE361044981B4" pageId="2" pageNumber="153" type="materials_examined">
|
||
<paragraph id="8BEFFA0D217AFF8965FDE361044981B4" blockId="2.[151,760,629,808]" pageId="2" pageNumber="153">
|
||
<emphasis id="B924261F217AFF8965FDE36104488181" bold="true" box="[151,275,629,648]" pageId="2" pageNumber="153">
|
||
<typeStatus id="54EB44AF217AFF8965FDE361059F8181" box="[151,196,629,648]" pageId="2" pageNumber="153">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F347BA9D217AFF896470E36104348181" box="[282,367,629,648]" name="Indonesia" pageId="2" pageNumber="153">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="499434EF217AFF896410E361074B8181" box="[378,528,629,648]" country="Indonesia" name="Sulawesi Tengah" pageId="2" pageNumber="153">Central Sulawesi</collectingRegion>
|
||
, Palu District, Taman Nasional Lore Lindu [National Park], environs of Wuasa,
|
||
<geoCoordinate id="EE649CCA217AFF8967FBE39B07A881AB" box="[657,755,655,674]" degrees="1" direction="south" minutes="24,445" orientation="latitude" pageId="2" pageNumber="153" precision="1" value="-1.4074167">
|
||
1°24,445
|
||
<emphasis id="B924261F217AFF89678BE39B07BD81AB" box="[737,742,655,674]" italics="true" pageId="2" pageNumber="153">′</emphasis>
|
||
S
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE649CCA217AFF8965FDE3BE045581B4" box="[151,270,682,701]" degrees="120" direction="east" minutes="18,801" orientation="longitude" pageId="2" pageNumber="153" precision="1" value="120.31335">
|
||
120°18,801
|
||
<emphasis id="B924261F217AFF896590E3BE05A481B4" box="[250,255,682,701]" italics="true" pageId="2" pageNumber="153">′</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA986217AFF8965FDE3D104448021" pageId="2" pageNumber="153" type="materials_examined">
|
||
<paragraph id="8BEFFA0D217AFF8965FDE3D104448021" blockId="2.[151,760,629,808]" pageId="2" pageNumber="153">
|
||
<emphasis id="B924261F217AFF8965FDE3D1044581D1" bold="true" box="[151,286,709,728]" pageId="2" pageNumber="153">Type material.</emphasis>
|
||
<materialsCitation id="3B38F050217AFF89644EE3D105AF8004" ID-GBIF-Occurrence="3753676301" collectingDate="2013-08-15" collectionCode="NMPC" collectorName="V. Hula" latitude="-1.4074167" location="Lore Lindu N. P." longLatPrecision="1" longitude="20.31335" municipality="Palu dist." pageId="2" pageNumber="153" specimenCount="1" specimenCount-male="1" stateProvince="SULAWESI" typeStatus="holotype">
|
||
<typeStatus id="54EB44AF217AFF89644EE3D1042381DF" box="[292,376,709,728]" pageId="2" pageNumber="153" type="holotype">HOLOTYPE</typeStatus>
|
||
: (
|
||
<collectionCode id="ED4162C8217AFF8964C8E3D104B881D1" box="[418,483,709,728]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="153">NMPC</collectionCode>
|
||
),
|
||
<collectingRegion id="499434EF217AFF896499E3D1070481D1" box="[499,607,709,728]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="153">SULAWESI</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B8B6077217AFF896700E3D107E781D1" box="[618,700,709,728]" latitude="-1.4074167" longLatPrecision="1" longitude="20.31335" name="Palu dist." pageId="2" pageNumber="153" stateProvince="Sulawesi Utara">Palu dist.</collectingMunicipality>
|
||
/
|
||
<location id="8E8FACD6217AFF8967A5E3D105AF81FB" LSID="urn:lsid:plazi:treatment:03F94B1B217AFF896447E339005D85DC:8E8FACD6217AFF8967A5E3D105AF81FB" latitude="-1.4074167" longLatPrecision="1" longitude="20.31335" municipality="Palu dist." name="Lore Lindu N. P." pageId="2" pageNumber="153" stateProvince="SULAWESI">Lore Lindu N.P.</location>
|
||
,
|
||
<location id="8E8FACD6217AFF896590E3CB046881FB" LSID="urn:lsid:plazi:treatment:03F94B1B217AFF896447E339005D85DC:8E8FACD6217AFF896590E3CB046881FB" box="[250,307,735,754]" latitude="-1.4074167" longLatPrecision="1" longitude="20.31335" municipality="Palu dist." name="Wuasa" pageId="2" pageNumber="153" stateProvince="SULAWESI">Wuasa</location>
|
||
env., /
|
||
<geoCoordinate id="EE649CCA217AFF896407E3CB048B81FB" box="[365,464,735,754]" degrees="1" direction="south" minutes="24,445" orientation="latitude" pageId="2" pageNumber="153" precision="1" value="-1.4074167">1°24,445’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE649CCA217AFF8964BDE3CB071881FB" box="[471,579,735,754]" degrees="20" direction="east" minutes="18,801" orientation="longitude" pageId="2" pageNumber="153" precision="1" value="20.31335">20°18,801’E</geoCoordinate>
|
||
[sic!], /
|
||
<date id="FFEEDCCD217AFF8967E2E3CB07AE81FB" box="[648,757,735,754]" pageId="2" pageNumber="153" value="2013-08-15">
|
||
<collectingDate id="EFAA2525217AFF8967E2E3CB07AE81FB" box="[648,757,735,754]" pageId="2" pageNumber="153" value="2013-08-15">15.viii. 2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A59FDB217AFF8965FDE3EE05888004" box="[151,211,762,781]" pageId="2" pageNumber="153">V.Hula</collectorName>
|
||
lgt.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B38F050217AFF89659DE3EE04448021" collectingDate="2013-08-15" collectionCode="NMPC, MHNG" collectorName="V. Hula" latitude="-1.4074167" location="Lore Lindu N. P." longLatPrecision="1" longitude="20.31335" municipality="Palu dist." pageId="2" pageNumber="153" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="SULAWESI" typeStatus="paratype">
|
||
<typeStatus id="54EB44AF217AFF89659DE3EE04148002" box="[247,335,762,781]" pageId="2" pageNumber="153" type="paratype">PARATYPES</typeStatus>
|
||
: 5
|
||
<specimenCount id="9D563184217AFF8964E1E3EE04E38004" box="[395,440,758,781]" pageId="2" pageNumber="153" type="female">6 ♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED4162C8217AFF8964A9E3EE07588004" box="[451,515,762,781]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="153">NMPC</collectionCode>
|
||
,
|
||
<collectionCode id="ED4162C8217AFF896760E3EE07148004" box="[522,591,762,781]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="153" type="Museum">MHNG</collectionCode>
|
||
), with the same data as the holotype.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA986217AFF8965FDE25401A580DC" pageId="2" pageNumber="153" type="description">
|
||
<paragraph id="8BEFFA0D217AFF8965FDE254076E84FE" blockId="2.[151,760,832,2039]" pageId="2" pageNumber="153">
|
||
<emphasis id="B924261F217AFF8965FDE254047C805E" bold="true" box="[151,295,832,855]" pageId="2" pageNumber="153">Description.</emphasis>
|
||
Length
|
||
<quantity id="4CA857E8217AFF8964E0E254076A805E" box="[394,561,832,855]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" metricValueMax="1.8" metricValueMin="1.64" pageId="2" pageNumber="153" unit="mm" value="1.72" valueMax="1.8" valueMin="1.64">1.64–1.80 mm</quantity>
|
||
, width
|
||
<quantity id="4CA857E8217AFF8967E3E2540598807E" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="2" pageNumber="153" unit="mm" value="1.075" valueMax="1.1" valueMin="1.05">1.05–1.10 mm</quantity>
|
||
. Head, pronotum and elytra reddish-brown, abdomen and appendages lighter, ochreous. Antennae long, length/ width ratios of antennomeres as: III 10/9: IV 38/7: V 44/8: VI 39/7: VII 46/13: VIII 37/8: IX 48/12: X 44/12: XI 51/13. Pronotum and elytra not microsculptured. Pronotum weakly narrowed anteriad, with regularly rounded lateral margins, lateral margins carinae throughout visible in dorsal view; punctation on disc very fine, sparse, hardly visible at magnification 30×, consisting of sharply delimited punctures, puncture intervals much larger than puncture diameters, punctures along lateral carinae absent. Minute tip of scutellum exposed. Elytra fairly narrowed apically, lateral margins almost regularly curved, lateral margin carinae exposed throughout in dorsal view, shape of apices sexually dimorphic; sutural margin not raised, adsutural areas flat, narrow, each with single puncture row. Sutural striae converging apically, curved near pronotal lobe, not extended laterally along basal margins. Punctures along lateral margins absent. Elytral disc with punctures very fine and shallow, somewhat larger than those on pronotal disc, not clearly delimited, puncture intervals mostly about three to five times as large as puncture diameters. Hind wings completely reduced. Hypomera smooth. Mesepimera almost two times longer than interval to mesocoxae and about five times as long as wide. Metaventrite lacking microsculpture except on minute areas near metacoxae, entirely very finely and sparsely punctate, antecoxal puncture rows absent; margin of intercoxal process slightly concave; submesocoxal lines convex, very finely punctate; submesocoxal areas
|
||
<quantity id="4CA857E8217AFF8967A0E7F40599841E" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="153" unit="mm" value="0.04">0.04 mm</quantity>
|
||
, about as long as halves of shortest intervals between them and metacoxae. Metanepisterna flat, not narrowed anteriad, with straight suture. Abdominal ventrite I very finely and sparsely punctate, without obvious microsculpture; submetacoxal lines subparallel, distinctly punctate,
|
||
<quantity id="4CA857E8217AFF8967A0E694059984BE" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="153" unit="mm" value="0.03">0.03 mm</quantity>
|
||
, about as long as seventh of shortest intervals between them and apical margin of ventrite. Following ventrites with conspicuous punctulate microsculpture.
|
||
</paragraph>
|
||
<paragraph id="8BEFFA0D217AFF896630E1AA01B7801C" blockId="2.[828,1437,190,1749]" pageId="2" pageNumber="153">
|
||
<emphasis id="B924261F217AFF896630E1AA014383DC" bold="true" box="[858,1048,190,213]" italics="true" pageId="2" pageNumber="153">Male characters.</emphasis>
|
||
Protarsomeres I strongly widened, about as wide as apices of protibiae and almost twice as wide as tarsomeres III; protarsomeres II somewhat narrower than protarsomeres I (
|
||
<figureCitation id="136BE688217AFF8961E1E00A0190823C" box="[1163,1227,286,309]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="2" pageNumber="153">Fig. 8</figureCitation>
|
||
). Protibiae straight, mesotibiae straight near base, curved posterior basal fourth (
|
||
<figureCitation id="136BE688217AFF89662EE04A06DD827C" box="[836,902,350,373]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="2" pageNumber="153">Fig. 9</figureCitation>
|
||
), metatibiae slightly curved posterior basal third. Elytral apices truncate, perpendicular to sutural margin, inner apical angles in level with outer apical angles (
|
||
<figureCitation id="136BE688217AFF89601BE08A060182DC" captionStart="Figs 50–53" captionStartId="12.[151,188,2027,2046]" captionTargetBox="[206,1382,457,2010]" captionTargetId="figure@12.[205,1382,456,2011]" captionTargetPageId="12" captionText="Figs 50–53. Habitus of Scaphisoma. 50, 51 – S. ancora sp. nov. (50 – male, 51 – female); 52 – S. sumpichi sp. nov., male; 53 – S. versicoloreum sp. nov., male." figureDoi="http://doi.org/10.5281/zenodo.3699171" httpUri="https://zenodo.org/record/3699171/files/figure.png" pageId="2" pageNumber="153">Fig. 50</figureCitation>
|
||
). Median part of metaventrite with large, subtriangular impression. Abdominal ventrite VI with deeply emarginate apical margin (
|
||
<figureCitation id="136BE688217AFF896688E0EA016D811C" box="[994,1078,510,533]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="2" pageNumber="153">Fig. 10</figureCitation>
|
||
). Tergite IX with one macroseta on each plate, stalk widened basally, with lateral sections concavely impressed (
|
||
<figureCitation id="136BE688217AFF896143E32A012C815C" box="[1065,1143,574,597]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="2" pageNumber="153">Fig. 11</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="136BE688217AFF89606AE32A0004815C" box="[1280,1375,574,597]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="2" pageNumber="153">Figs 6, 7</figureCitation>
|
||
)
|
||
<quantity id="4CA857E8217AFF896004E32A063D817C" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="153" unit="mm" value="0.65">0.65 mm</quantity>
|
||
long, asymmetrical. Median lobe with small basal bulb bearing proximal anchor-like process, apical process asymmetrically curved, with small subapical tooth. Basal parts of parameres strongly expanded, apical parts lobate bearing very dense pores. Internal sac simple, tubular, membranous, lacking sclerotized pieces.
|
||
</paragraph>
|
||
<paragraph id="8BEFFA0D217AFF896630E20A01A580DC" blockId="2.[828,1437,190,1749]" pageId="2" pageNumber="153">
|
||
<emphasis id="B924261F217AFF896630E20A0164803C" bold="true" box="[858,1087,798,821]" italics="true" pageId="2" pageNumber="153">Female characters.</emphasis>
|
||
Elytra with prominent apical angles, apical margins oblique and concave, inner apical angles situated far posterior outer apical angles (
|
||
<figureCitation id="136BE688217AFF89602AE24A00CB807C" box="[1344,1424,862,885]" captionStart="Figs 50–53" captionStartId="12.[151,188,2027,2046]" captionTargetBox="[206,1382,457,2010]" captionTargetId="figure@12.[205,1382,456,2011]" captionTargetPageId="12" captionText="Figs 50–53. Habitus of Scaphisoma. 50, 51 – S. ancora sp. nov. (50 – male, 51 – female); 52 – S. sumpichi sp. nov., male; 53 – S. versicoloreum sp. nov., male." figureDoi="http://doi.org/10.5281/zenodo.3699171" httpUri="https://zenodo.org/record/3699171/files/figure.png" pageId="2" pageNumber="153">Fig. 51</figureCitation>
|
||
). Median part of metaventrite flattened. Pro- and mesotibiae straight, metatibiae hardly bent, tarsomeres not widened. Gonocoxite bent, evenly narrow (
|
||
<figureCitation id="136BE688217AFF8961C8E2AA01A980DC" box="[1186,1266,958,981]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="2" pageNumber="153">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA986217AFF896656E2CA0163855C" pageId="2" pageNumber="153" type="diagnosis">
|
||
<paragraph id="8BEFFA0D217AFF896656E2CA0163855C" blockId="2.[828,1437,190,1749]" pageId="2" pageNumber="153">
|
||
<emphasis id="B924261F217AFF896656E2CA016E80FC" bold="true" box="[828,1077,990,1013]" pageId="2" pageNumber="153">Differential diagnosis.</emphasis>
|
||
The sexually dimorphic elytra are unusual within the
|
||
<taxonomicName id="4C50818E217AFF89616CE2EA01F5871C" box="[1030,1198,1022,1045]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="tribe" tribe="Scaphisomatini">Scaphisomatini</taxonomicName>
|
||
. They are described in several species of the
|
||
<emphasis id="B924261F217AFF896142E50A0059873C" box="[1064,1282,1054,1077]" italics="true" pageId="2" pageNumber="153">
|
||
<taxonomicName id="4C50818E217AFF896142E50A01F2873C" authorityName="Leach" authorityYear="1815" box="[1064,1193,1054,1077]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="164" phylum="Arthropoda" rank="genus">Scaphisoma</taxonomicName>
|
||
tricolor
|
||
</emphasis>
|
||
species group (see
|
||
<bibRefCitation id="EFC187FC217AFF896605E52A010D875C" author="OGAWA R. & LOBL I." box="[879,1110,1086,1109]" firstAuthor="LOBL" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="2" pageNumber="153" pagination="155 - 169" part="63" refId="ref13146" refString="OGAWA R. & LOBL I. 2016: A review of the genus Xotidium Lobl, 1992 (Coleoptera, Staphylinidae, Scaphidiinae), with description of five new species. Deutsche Entomologische Zeitschrift 63: 155 - 169." title="A review of the genus Xotidium Lobl, 1992 (Coleoptera, Staphylinidae, Scaphidiinae), with description of five new species" type="journal article" year="2016">LÖBL & OGAWA 2016</bibRefCitation>
|
||
), which are quite unrelated to
|
||
<taxonomicName id="4C50818E217AFF896656E54B06F8877F" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[828,931,1119,1142]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="ancora" status="sp. nov.">
|
||
<emphasis id="B924261F217AFF896656E54B06F8877F" box="[828,931,1119,1142]" italics="true" pageId="2" pageNumber="153">S. ancora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2179B64217AFF8966C0E54A06A6877C" box="[938,1021,1118,1141]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Unlike in
|
||
<taxonomicName id="4C50818E217AFF896105E54B018D877F" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[1135,1238,1119,1142]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="ancora" status="sp. nov.">
|
||
<emphasis id="B924261F217AFF896105E54B018D877F" box="[1135,1238,1119,1142]" italics="true" pageId="2" pageNumber="153">S. ancora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2179B64217AFF8961B7E54A0068877C" box="[1245,1331,1118,1141]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, the inner apical elytral angles are usually denticulate in females of the
|
||
<emphasis id="B924261F217AFF89660CE58B068F87BC" box="[870,980,1182,1206]" italics="true" pageId="2" pageNumber="153">S. tricolor</emphasis>
|
||
group. Members of the
|
||
<emphasis id="B924261F217AFF89618FE58B000F87BC" box="[1253,1364,1182,1206]" italics="true" pageId="2" pageNumber="153">S. tricolor</emphasis>
|
||
group differ notably by the elytra with shortened sutural striae, the strigulate abdominal microsculpture, the aedeagi with large, weakly sclerotized basal bulbs lacking proximal processes, the apical processes of the median lobes consisting of asymmetrical dorsal and ventral branches, the parameres lobed and overlapping apically, and the complex internal sacs. An anchor-like aedeagal process is well developed in the Moluccan
|
||
<emphasis id="B924261F217AFF8966BFE48B010786BC" box="[981,1116,1438,1462]" italics="true" pageId="2" pageNumber="153">S. semibreve</emphasis>
|
||
Löbl, 2014. This species also shares reduced hind wings, likely correlated with a short metaventrite.
|
||
<emphasis id="B924261F217AFF8966A2E4CA01E986FC" box="[968,1202,1502,1525]" italics="true" pageId="2" pageNumber="153">
|
||
<taxonomicName id="4C50818E217AFF8966A2E4CA011D86FC" authorityName="Leach" authorityYear="1815" box="[968,1094,1502,1525]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="164" phylum="Arthropoda" rank="genus">Scaphisoma</taxonomicName>
|
||
semibreve
|
||
</emphasis>
|
||
differs notably from
|
||
<taxonomicName id="4C50818E217AFF8960E2E4CB06D3851F" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="ancora" status="sp. nov.">
|
||
<emphasis id="B924261F217AFF8960E2E4CB06D3851F" italics="true" pageId="2" pageNumber="153">S. ancora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2179B64217AFF8966E6E4EA0686851C" box="[908,989,1534,1557]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the internal sac of the aedeagus bearing rows of robust teeth-like sclerites, and by the abdomen lacking microsculpture.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA986217AFF896656E74A01F985BC" pageId="2" pageNumber="153" type="etymology">
|
||
<paragraph id="8BEFFA0D217AFF896656E74A01F985BC" blockId="2.[828,1437,190,1749]" pageId="2" pageNumber="153">
|
||
<emphasis id="B924261F217AFF896656E74A06E7857C" bold="true" box="[828,956,1630,1653]" pageId="2" pageNumber="153">Etymology.</emphasis>
|
||
The species epithet is a Latin noun in apposition,
|
||
<emphasis id="B924261F217AFF8966EFE76B068A859F" box="[901,977,1663,1686]" italics="true" pageId="2" pageNumber="153">ancora</emphasis>
|
||
, meaning ‘anchor’, and refers to the shape of the proximal aedeagal process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA986217AFF896656E7AA005D85DC" box="[828,1286,1726,1749]" pageId="2" pageNumber="153" type="distribution">
|
||
<paragraph id="8BEFFA0D217AFF896656E7AA005D85DC" blockId="2.[828,1437,190,1749]" box="[828,1286,1726,1749]" pageId="2" pageNumber="153">
|
||
<emphasis id="B924261F217AFF896656E7AA069585DC" bold="true" box="[828,974,1726,1749]" pageId="2" pageNumber="153">Distribution.</emphasis>
|
||
<collectingCountry id="F347BA9D217AFF8966BEE7AA016585DC" box="[980,1086,1726,1749]" name="Indonesia" pageId="2" pageNumber="153">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="499434EF217AFF896123E7AA005985DC" box="[1097,1282,1726,1749]" country="Indonesia" name="Sulawesi Tengah" pageId="2" pageNumber="153">Central Sulawesi</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |