452 lines
69 KiB
XML
452 lines
69 KiB
XML
<document id="3C799371B0DB89DE86F643CD2864804B" ID-DOI="10.5281/zenodo.4504166" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504166" ID-ZooBank="924CD2C9-3007-4321-81E2-0237EE1359BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612465749776" checkinUser="carolina" docAuthor="Schubart, Christoph D. & Ng, Peter K. L." docDate="2014" docId="03EA5318167DE35DAAB3F6F1BFE62C15" docLanguage="en" docName="RafflesBZool.62.615-619.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Geosesarma raj Schubart & Ng 2014, new species" docType="treatment" docUuid="C89B65EA-D107-464B-9325-23A53624BE3B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="618" masterDocId="FFD32B60167CE35EA978F24CBC422B21" masterDocTitle="Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia" masterLastPageNumber="619" masterPageNumber="615" pageNumber="616" updateTime="1698866020970" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="690CDB220F31B0B6910D21F966797ECE">Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia</mods:title>
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<mods:namePart id="D9E14BC015CC8DA35A4924622349A8E8">Schubart, Christoph D.</mods:namePart>
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<mods:roleTerm id="06805A5864CBC99C152AA5C8F289C5AE">Author</mods:roleTerm>
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<mods:namePart id="210F84F0020DC13033E877B518DBFA43">Ng, Peter K. L.</mods:namePart>
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<mods:title id="11905C7C97D5F9F65974828079AE3425">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="DD33F4A1C24DF783CC368562C71E65B9">
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<mods:date id="59673239CBFEEA37009BE65A681FD895">2014</mods:date>
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<mods:number id="A6D0185B2FB20AA54EA3D9B0613A8850">2014-08-22</mods:number>
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<mods:number id="63AE9CF35B82381B24BA0A957768D7AC">62</mods:number>
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<subSubSection id="C359B185167DE35FAAB3F6F1B94C2FF4" box="[971,1294,1213,1237]" pageId="1" pageNumber="616" type="nomenclature">
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<paragraph id="8BFCE20E167DE35FAAB3F6F1B94C2FF4" blockId="1.[919,1345,1213,1269]" box="[971,1294,1213,1237]" pageId="1" pageNumber="616">
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<heading id="D0B45562167DE35FAAB3F6F1B94C2FF4" bold="true" box="[971,1294,1213,1237]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="616" reason="2">
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<emphasis id="B9373E1C167DE35FAAB3F6F1B94C2FF4" bold="true" box="[971,1294,1213,1237]" pageId="1" pageNumber="616">
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<taxonomicName id="4C43998D167DE35FAAB3F6F1B83B2FF4" ID-CoL="3FS9X" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[971,1145,1213,1237]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="raj" status="sp. nov.">
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<emphasis id="B9373E1C167DE35FAAB3F6F1B83B2FF4" bold="true" box="[971,1145,1213,1237]" italics="true" pageId="1" pageNumber="616">Geosesarma raj</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2048367167DE35FADF0F6F1B94C2FF4" box="[1160,1294,1213,1237]" pageId="1" pageNumber="616" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C359B185167DE35FAAEFF691B9032FD4" box="[919,1345,1245,1269]" pageId="1" pageNumber="616" type="description">
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<paragraph id="8BFCE20E167DE35FAAEFF691B9032FD4" blockId="1.[919,1345,1213,1269]" box="[919,1345,1245,1269]" pageId="1" pageNumber="616">
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(
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<figureCitation id="1378FE8B167DE35FAAE7F691B8622FD4" box="[927,1056,1245,1269]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="1" pageNumber="616">Figs. 1B, D</figureCitation>
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,
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<figureCitation id="1378FE8B167DE35FAD56F691B8302FD4" box="[1070,1138,1245,1269]" captionStart="Fig" captionStartId="1.[812,847,1977,1999]" captionTargetBox="[812,1451,1572,1954]" captionTargetId="figure-266@1.[812,1451,1572,1954]" captionTargetPageId="1" captionText="Fig. 2.A, C, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, frontal view of carapace; C, D, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4504172" httpUri="https://zenodo.org/record/4504172/files/figure.png" pageId="1" pageNumber="616">2B, D</figureCitation>
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,
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<figureCitation id="1378FE8B167DE35FAD07F691B89D2FD4" box="[1151,1247,1245,1269]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">3B, D, F</figureCitation>
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, J–M, 4)
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||
</paragraph>
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<subSubSection id="C359B185167DE35DAA55F751BDD62A77" lastPageId="3" lastPageNumber="618" pageId="1" pageNumber="616" type="materials_examined">
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<emphasis id="B9373E1C167DE35FAA55F751B8512E14" bold="true" box="[813,1043,1309,1333]" pageId="1" pageNumber="616">Material examined.</emphasis>
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<materialsCitation id="3B2BE853167DE35FAD65F751BF802EB4" collectingDate="2007-10-18" collectionCode="MZB" collectorName="G. Subaraj" country="Indonesia" location="northern Bintan Island" pageId="1" pageNumber="616" specimenCode="ZRC 2007.0676" specimenCount="1" stateProvince="Riau" typeStatus="holotype">
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<typeStatus id="54F85CAC167DE35FAD65F751B8C02E14" box="[1053,1154,1309,1333]" pageId="1" pageNumber="616" type="holotype">Holotype</typeStatus>
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– male (11.8 ×
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<quantity id="4CBB4FEB167DE35FAC47F751B9E62E14" box="[1343,1444,1309,1333]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="1" pageNumber="616" unit="mm" value="10.9">10.9 mm</quantity>
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) (
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<collectionCode id="ED527ACB167DE35FAA4CF771BF332E74" box="[820,881,1341,1365]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="616" type="Museum">MZB</collectionCode>
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Cru 3814, ex
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<specimenCode id="DBE54A75167DE35FAD59F771B89C2E74" box="[1057,1246,1341,1365]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="616">ZRC 2007.0676</specimenCode>
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),
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<collectingCountry id="F354A29E167DE35FAD8EF771B92A2E74" box="[1270,1384,1341,1365]" name="Indonesia" pageId="1" pageNumber="616">Indonesia</collectingCountry>
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:
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<collectingRegion id="49872CEC167DE35FAC0FF771B9E92E74" box="[1399,1451,1341,1365]" country="Indonesia" name="Riau" pageId="1" pageNumber="616">Riau</collectingRegion>
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Archipelago,
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<location id="8E9CB4D5167DE35FAABBF711B8822E54" LSID="urn:lsid:plazi:treatment:03EA5318167DE35DAAB3F6F1BFE62C15:8E9CB4D5167DE35FAABBF711B8822E54" box="[963,1216,1373,1397]" country="Indonesia" name="northern Bintan Island" pageId="1" pageNumber="616" stateProvince="Riau">northern Bintan Island</location>
|
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, coll.
|
||
<collectorName id="26B687D8167DE35FAC70F711B9C32E54" box="[1288,1409,1373,1397]" pageId="1" pageNumber="616">G. Subaraj</collectorName>
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,
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<date id="FFFDC4CE167DE35FACE8F711BF802EB4" pageId="1" pageNumber="616" value="2007-10-18">
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||
<collectingDate id="EFB93D26167DE35FACE8F711BF802EB4" pageId="1" pageNumber="616" value="2007-10-18">18 October 2007</collectingDate>
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</date>
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||
</materialsCitation>
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.
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||
<materialsCitation id="3B2BE853167DE35FAAB3F732B9572EF4" collectingDate="2007-10-21" collectionCode="ZRC" collectorName="G. Subaraj" country="Indonesia" location="Bintan Island" municipality="Archipelago" pageId="1" pageNumber="616" specimenCode="ZRC 2007.0677" specimenCount="1" specimenCount-female="1" stateProvince="Riau" typeStatus="paratype">
|
||
<typeStatus id="54F85CAC167DE35FAAB3F732B8712EB7" box="[971,1075,1406,1430]" pageId="1" pageNumber="616" type="paratype">Paratypes</typeStatus>
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||
–
|
||
<specimenCount id="9D452987167DE35FAD35F731B8EB2EB4" box="[1101,1193,1405,1429]" pageId="1" pageNumber="616" type="female">1 female</specimenCount>
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(11.8 ×
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<quantity id="4CBB4FEB167DE35FAC79F731B9232EB4" box="[1281,1377,1405,1429]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="1" pageNumber="616" unit="mm" value="10.7">10.7 mm</quantity>
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) (
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<specimenCode id="DBE54A75167DE35FAC0EF731BFE02E94" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="616">ZRC 2007.0677</specimenCode>
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),
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<collectingCountry id="F354A29E167DE35FAACDF7D1B8632E94" box="[949,1057,1437,1461]" name="Indonesia" pageId="1" pageNumber="616">Indonesia</collectingCountry>
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:
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<collectingRegion id="49872CEC167DE35FAD54F7D1B81C2E94" box="[1068,1118,1437,1461]" country="Indonesia" name="Riau" pageId="1" pageNumber="616">Riau</collectingRegion>
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<collectingMunicipality id="6B987874167DE35FAD1DF7D1B8AE2E94" box="[1125,1260,1437,1461]" pageId="1" pageNumber="616">Archipelago</collectingMunicipality>
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,
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<location id="8E9CB4D5167DE35FAD8FF7D1B9C82E94" LSID="urn:lsid:plazi:treatment:03EA5318167DE35DAAB3F6F1BFE62C15:8E9CB4D5167DE35FAD8FF7D1B9C82E94" box="[1271,1418,1437,1461]" country="Indonesia" municipality="Archipelago" name="Bintan Island" pageId="1" pageNumber="616" stateProvince="Riau">Bintan Island</location>
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, in tree hole, coll.
|
||
<collectorName id="26B687D8167DE35FAAADF7F1B80E2EF4" box="[981,1100,1469,1493]" pageId="1" pageNumber="616">G. Subaraj</collectorName>
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,
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<date id="FFFDC4CE167DE35FAD21F7F1B9572EF4" box="[1113,1301,1469,1493]" pageId="1" pageNumber="616" value="2007-10-21">
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<collectingDate id="EFB93D26167DE35FAD21F7F1B9572EF4" box="[1113,1301,1469,1493]" pageId="1" pageNumber="616" value="2007-10-21">21 October 2007</collectingDate>
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</date>
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</materialsCitation>
|
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;
|
||
<materialsCitation id="3B2BE853167DE35DAC5AF7F1BEAE2BD7" collectingDate="2007-06-23" collectionCode="ZRC" collectorName="G. Subaraj" country="Indonesia" county="Archipelago" lastPageId="3" lastPageNumber="618" location="Gunong" municipality="Bintan Island" pageId="1" pageNumber="616" specimenCode="ZRC 2007.0675" specimenCount="1" specimenCount-male="1" stateProvince="Riau" typeStatus="paratype">
|
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<specimenCount id="9D452987167DE35FAC5AF7F1B92E2EF4" box="[1314,1388,1469,1493]" pageId="1" pageNumber="616" type="male">1 male</specimenCount>
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(10.0 ×
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<quantity id="4CBB4FEB167DE35FAA3BF791BFD52ED4" box="[835,919,1501,1525]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="1" pageNumber="616" unit="mm" value="9.7">9.7 mm</quantity>
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) (
|
||
<specimenCode id="DBE54A75167DE35FAAD5F791B8222ED4" box="[941,1120,1501,1525]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="616">ZRC 2007.0675</specimenCode>
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),
|
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<collectingCountry id="F354A29E167DE35FAD0AF791B89C2ED4" box="[1138,1246,1501,1525]" name="Indonesia" pageId="1" pageNumber="616">Indonesia</collectingCountry>
|
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:
|
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<collectingRegion id="49872CEC167DE35FAD91F791B9592ED4" box="[1257,1307,1501,1525]" country="Indonesia" name="Riau" pageId="1" pageNumber="616">Riau</collectingRegion>
|
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<collectingCounty id="629D9A82167DE35FAC59F791B9EA2ED4" box="[1313,1448,1501,1525]" pageId="1" pageNumber="616">Archipelago</collectingCounty>
|
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,
|
||
<collectingMunicipality id="6B987874167FE35DA909F2F2BD522BF7" box="[113,272,190,214]" pageId="3" pageNumber="618">Bintan Island</collectingMunicipality>
|
||
:
|
||
<location id="8E9CB4D5167FE35DA867F2F2BD3A2BF7" LSID="urn:lsid:plazi:treatment:03EA5318167DE35DAAB3F6F1BFE62C15:8E9CB4D5167FE35DA867F2F2BD3A2BF7" box="[287,376,190,214]" country="Indonesia" county="Archipelago" municipality="Bintan Island" name="Gunong" pageId="3" pageNumber="618" stateProvince="Riau">Gunong</location>
|
||
(=
|
||
<location id="8E9CB4D5167FE35DA8D0F2F2BDB62BF7" LSID="urn:lsid:plazi:treatment:03EA5318167DE35DAAB3F6F1BFE62C15:8E9CB4D5167FE35DA8D0F2F2BDB62BF7" box="[424,500,190,214]" country="Indonesia" county="Archipelago" municipality="Bintan Island" name="Mount" pageId="3" pageNumber="618" stateProvince="Riau">Mount</location>
|
||
)
|
||
<location id="8E9CB4D5167FE35DAB7EF2F2BE162BF7" LSID="urn:lsid:plazi:treatment:03EA5318167DE35DAAB3F6F1BFE62C15:8E9CB4D5167FE35DAB7EF2F2BE162BF7" box="[518,596,190,214]" country="Indonesia" county="Archipelago" municipality="Bintan Island" name="Bintan" pageId="3" pageNumber="618" stateProvince="Riau">Bintan</location>
|
||
, above lower waterfalls (attacked by ants), coll.
|
||
<collectorName id="26B687D8167FE35DA8A6F292BE102BD7" box="[478,594,222,246]" pageId="3" pageNumber="618">G. Subaraj</collectorName>
|
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,
|
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<date id="FFFDC4CE167FE35DAB24F292BEAE2BD7" box="[604,748,222,246]" pageId="3" pageNumber="618" value="2007-06-23">
|
||
<collectingDate id="EFB93D26167FE35DAB24F292BEAE2BD7" box="[604,748,222,246]" pageId="3" pageNumber="618" value="2007-06-23">23 June 2007</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9D452987167FE35DA909F2B2BCF82A37" box="[113,186,254,278]" pageId="3" pageNumber="618" type="male">1 male</specimenCount>
|
||
(9.4 × 9.0 mm),
|
||
<materialsCitation id="3B2BE853167FE35DA80DF2B2BDD32A77" collectingDate="1996-02-19" collectionCode="ZRC" collectorName="Y. H. Lee" country="Indonesia" location="Gunung Bintan Besar" pageId="3" pageNumber="618" specimenCode="ZRC 1996.133, 135" specimenCount="2" specimenCount-female="2" stateProvince="Riau" typeStatus="paratype">
|
||
<specimenCount id="9D452987167FE35DA80DF2B2BD9F2A37" box="[373,477,254,278]" pageId="3" pageNumber="618" type="female">2 females</specimenCount>
|
||
(7.3 ×
|
||
<quantity id="4CBB4FEB167FE35DAB56F2B2BEC12A37" box="[558,643,254,278]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="618" unit="mm" value="6.8">6.8 mm</quantity>
|
||
, 6.4 ×
|
||
<quantity id="4CBB4FEB167FE35DABB7F2B2BCDE2A17" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="618" unit="mm" value="6.2">6.2 mm</quantity>
|
||
) (
|
||
<specimenCode id="DBE54A75167FE35DA9CDF352BD1E2A17" box="[181,348,286,310]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="618">ZRC 1996.133</specimenCode>
|
||
–
|
||
<specimenCode id="DBE54A75167FE35DA812F352BDD32A17" box="[362,401,286,310]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="618">135</specimenCode>
|
||
),
|
||
<location id="8E9CB4D5167FE35DA8DEF352BED62A17" LSID="urn:lsid:plazi:treatment:03EA5318167DE35DAAB3F6F1BFE62C15:8E9CB4D5167FE35DA8DEF352BED62A17" box="[422,660,286,310]" country="Indonesia" name="Gunung Bintan Besar" pageId="3" pageNumber="618" stateProvince="Riau">Gunung Bintan Besar</location>
|
||
, coll.
|
||
<collectorName id="26B687D8167FE35DABA2F353BCFD2A77" pageId="3" pageNumber="618">Y. H. Lee</collectorName>
|
||
,
|
||
<date id="FFFDC4CE167FE35DA9B3F372BDD32A77" box="[203,401,318,342]" pageId="3" pageNumber="618" value="1996-02-19">
|
||
<collectingDate id="EFB93D26167FE35DA9B3F372BDD32A77" box="[203,401,318,342]" pageId="3" pageNumber="618" value="1996-02-19">19 February 1996</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3CB286167DE35FA909F5F4BD5D2360" ID-DOI="http://doi.org/10.5281/zenodo.4504168" ID-Zenodo-Dep="4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="1" pageNumber="616" startId="1.[113,148,1976,1998]" targetBox="[113,752,1116,1953]" targetPageId="1">
|
||
<paragraph id="8BFCE20E167DE35FA909F5F4BD5D2360" blockId="1.[113,751,1976,2113]" pageId="1" pageNumber="616">
|
||
Fig. 1. A, C,
|
||
<taxonomicName id="4C43998D167DE35FA991F5F4BDE12CEC" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[233,419,1976,1997]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="bintan" status="sp. nov.">
|
||
<emphasis id="B9373E1C167DE35FA991F5F4BDE12CEC" box="[233,419,1976,1997]" italics="true" pageId="1" pageNumber="616">Geosesarma bintan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167DE35FA8C8F5F4BE672CEF" box="[432,549,1976,1998]" pageId="1" pageNumber="616" rank="species">new species</taxonomicNameLabel>
|
||
, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D,
|
||
<taxonomicName id="4C43998D167DE35FAB37F599BEA82CCB" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[591,746,2005,2026]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="raj" status="sp. nov.">
|
||
<emphasis id="B9373E1C167DE35FAB37F599BEA82CCB" box="[591,746,2005,2026]" italics="true" pageId="1" pageNumber="616">Geosesarma raj</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167DE35FA909F5BEBCA92329" box="[113,235,2034,2056]" pageId="1" pageNumber="616" rank="species">new species</taxonomicNameLabel>
|
||
, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3CB286167DE35FAA54F5F5B8212363" ID-DOI="http://doi.org/10.5281/zenodo.4504172" ID-Zenodo-Dep="4504172" httpUri="https://zenodo.org/record/4504172/files/figure.png" pageId="1" pageNumber="616" startId="1.[812,847,1977,1999]" targetBox="[812,1451,1572,1954]" targetPageId="1">
|
||
<paragraph id="8BFCE20E167DE35FAA54F5F5B8212363" blockId="1.[812,1453,1977,2114]" pageId="1" pageNumber="616">
|
||
Fig. 2. A, C,
|
||
<taxonomicName id="4C43998D167DE35FAADCF5F5B81D2CEF" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[932,1119,1977,1998]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="bintan" status="sp. nov.">
|
||
<emphasis id="B9373E1C167DE35FAADCF5F5B81D2CEF" box="[932,1119,1977,1998]" italics="true" pageId="1" pageNumber="616">Geosesarma bintan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167DE35FAD13F5F5B8A22CEE" box="[1131,1248,1977,1999]" pageId="1" pageNumber="616" rank="species">new species</taxonomicNameLabel>
|
||
, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D,
|
||
<taxonomicName id="4C43998D167DE35FAC73F59AB9E72CCA" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[1291,1445,2006,2027]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="raj" status="sp. nov.">
|
||
<emphasis id="B9373E1C167DE35FAC73F59AB9E72CCA" box="[1291,1445,2006,2027]" italics="true" pageId="1" pageNumber="616">Geosesarma raj</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167DE35FAA54F5BFBFE12329" box="[812,931,2034,2056]" pageId="1" pageNumber="616" rank="species">new species</taxonomicNameLabel>
|
||
, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, frontal view of carapace; C, D, outer view of right chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3CB286167EE35CA9F0F5F4B9062360" ID-DOI="http://doi.org/10.5281/zenodo.4504174" ID-Zenodo-Dep="4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="2" pageNumber="617" startId="2.[136,171,1976,1998]" targetBox="[183,1424,193,1952]" targetPageId="2">
|
||
<paragraph id="8BFCE20E167EE35CA9F0F5F4B9062360" blockId="2.[136,1476,1976,2113]" pageId="2" pageNumber="617">
|
||
Fig. 3. A, C, E, G–I,
|
||
<taxonomicName id="4C43998D167EE35CA818F5F5BE5C2CEC" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[352,542,1976,1998]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="617" phylum="Arthropoda" rank="species" species="bintan" status="sp. nov.">
|
||
<emphasis id="B9373E1C167EE35CA818F5F5BE5C2CEC" box="[352,542,1976,1998]" italics="true" pageId="2" pageNumber="617">Geosesarma bintan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167EE35CAB55F5F5BEE72CEF" box="[557,677,1976,1998]" pageId="2" pageNumber="617" rank="species">new species</taxonomicNameLabel>
|
||
, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M,
|
||
<taxonomicName id="4C43998D167EE35CA9F0F599BD662CCA" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[136,292,2005,2027]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="617" phylum="Arthropoda" rank="species" species="raj" status="sp. nov.">
|
||
<emphasis id="B9373E1C167EE35CA9F0F599BD662CCA" box="[136,292,2005,2027]" italics="true" pageId="2" pageNumber="617">Geosesarma raj</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167EE35CA849F59ABDEB2CCA" box="[305,425,2005,2027]" pageId="2" pageNumber="617" rank="species">new species</taxonomicNameLabel>
|
||
, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C359B185167FE35DA909F332BE2728D7" pageId="3" pageNumber="618" type="diagnosis">
|
||
<paragraph id="8BFCE20E167FE35DA909F332BD0328B7" blockId="3.[112,753,382,918]" pageId="3" pageNumber="618">
|
||
<emphasis id="B9373E1C167FE35DA909F332BCA52AB7" bold="true" box="[113,231,382,406]" pageId="3" pageNumber="618">Diagnosis.</emphasis>
|
||
Carapace almost squarish, not distinctly wider than long, lateral margins almost parallel (
|
||
<figureCitation id="1378FE8B167FE35DAB19F3D2BEA02A97" box="[609,738,414,438]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="3" pageNumber="618">Fig. 1B, D</figureCitation>
|
||
); dorsal surfaces evenly covered with numerous granules and striae; H-shaped median depression deep; postfrontal cristae prominent, margins sharp; exorbital tooth triangular, curving obliquely, with additional low tooth behind it (
|
||
<figureCitation id="1378FE8B167FE35DABFFF052BEA82917" box="[647,746,542,566]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="3" pageNumber="618">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="1378FE8B167FE35DA909F072BCD72977" box="[113,149,574,598]" captionStart="Fig" captionStartId="1.[812,847,1977,1999]" captionTargetBox="[812,1451,1572,1954]" captionTargetId="figure-266@1.[812,1451,1572,1954]" captionTargetPageId="1" captionText="Fig. 2.A, C, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, frontal view of carapace; C, D, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4504172" httpUri="https://zenodo.org/record/4504172/files/figure.png" pageId="3" pageNumber="618">2B</figureCitation>
|
||
). Exopod of third maxilliped relatively stout, without flagellum (
|
||
<figureCitation id="1378FE8B167FE35DA99BF012BD762957" box="[227,308,606,630]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3B</figureCitation>
|
||
). Male chelae with long fingers, numerous granules on palm (dorsally and ventrally) and proximal part of dactylus; dorsal margin of dactylus with 8 or 9 sharp granules on proximal two-thirds (
|
||
<figureCitation id="1378FE8B167FE35DA8AFF0F2BE6F29F7" box="[471,557,702,726]" captionStart="Fig" captionStartId="1.[812,847,1977,1999]" captionTargetBox="[812,1451,1572,1954]" captionTargetId="figure-266@1.[812,1451,1572,1954]" captionTargetPageId="1" captionText="Fig. 2.A, C, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, frontal view of carapace; C, D, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4504172" httpUri="https://zenodo.org/record/4504172/files/figure.png" pageId="3" pageNumber="618">Fig. 2D</figureCitation>
|
||
). Ambulatory legs with relatively slender meri, with sharp subdistal spine on dorsal margin, surface almost smooth; propodus elongate (
|
||
<figureCitation id="1378FE8B167FE35DA902F152BC9F2817" box="[122,221,798,822]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="3" pageNumber="618">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="1378FE8B167FE35DA995F152BD532817" box="[237,273,798,822]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">3D</figureCitation>
|
||
). Male abdomen relatively narrow; telson longer than broad, with rounded tip (
|
||
<figureCitation id="1378FE8B167FE35DA886F172BE122877" box="[510,592,830,854]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3F</figureCitation>
|
||
). G1 relatively stout, distal chitinous part elongate, tapering; subdistal part broad (
|
||
<figureCitation id="1378FE8B167FE35DA9B8F132BD4228B7" box="[192,256,894,918]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3</figureCitation>
|
||
J–M).
|
||
</paragraph>
|
||
<paragraph id="8BFCE20E167FE35DA909F1F2BE2728D7" blockId="3.[113,752,958,1015]" pageId="3" pageNumber="618">
|
||
<emphasis id="B9373E1C167FE35DA909F1F2BC8528F7" bold="true" box="[113,199,958,982]" pageId="3" pageNumber="618">Colour.</emphasis>
|
||
The carapace of this species is reddish brown in life, with the chelipeds bright orange (
|
||
<figureCitation id="1378FE8B167FE35DAB6CF192BE1A28D7" box="[532,600,990,1014]" captionStart="Fig" captionStartId="3.[113,148,2033,2055]" captionTargetBox="[112,752,1248,2008]" captionTargetId="figure-241@3.[112,752,1248,2008]" captionTargetPageId="3" captionText="Fig. 4. Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. Colour in life (Photograph by: T. M. Leong)." figureDoi="http://doi.org/10.5281/zenodo.4504176" httpUri="https://zenodo.org/record/4504176/files/figure.png" pageId="3" pageNumber="618">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359B185167FE35DA909F652BDF02F57" pageId="3" pageNumber="618" type="etymology">
|
||
<paragraph id="8BFCE20E167FE35DA909F652BDF02F57" blockId="3.[113,752,1054,1142]" pageId="3" pageNumber="618">
|
||
<emphasis id="B9373E1C167FE35DA909F652BCB02F17" bold="true" box="[113,242,1054,1078]" pageId="3" pageNumber="618">Etymology.</emphasis>
|
||
The species is named after G. Subaraj (“Raj” to all his friends), the collector. The specific epithet,
|
||
<taxonomicName id="4C43998D167FE35DABC8F673BE8D2F77" authorityName="Schubart & Ng" authorityYear="2014" box="[688,719,1087,1110]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="raj">
|
||
<emphasis id="B9373E1C167FE35DABC8F673BE8D2F77" box="[688,719,1087,1110]" italics="true" pageId="3" pageNumber="618">raj</emphasis>
|
||
</taxonomicName>
|
||
, is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359B185167FE35DAA55F2F1BFE62C15" pageId="3" pageNumber="618" type="discussion">
|
||
<paragraph id="8BFCE20E167FE35DAA55F2F1B87A2A94" blockId="3.[812,1452,189,437]" pageId="3" pageNumber="618">
|
||
<emphasis id="B9373E1C167FE35DAA55F2F1BFD92BF4" bold="true" box="[813,923,189,213]" pageId="3" pageNumber="618">Remarks.</emphasis>
|
||
The quadrate carapace, the absence of a flagellum on the exopod of the third maxilliped, and the stout G1, which has a prominent, chitinised, beak-like structure on the distal end with the subdistal part broad, ally
|
||
<taxonomicName id="4C43998D167FE35DAC5DF351BF0F2A75" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="raj" status="sp. nov.">
|
||
<emphasis id="B9373E1C167FE35DAC5DF351BF0F2A75" italics="true" pageId="3" pageNumber="618">Geosesarma raj</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167FE35DAA26F371BFA52A74" box="[862,999,317,341]" pageId="3" pageNumber="618" rank="species">new species</taxonomicNameLabel>
|
||
, with species like
|
||
<taxonomicName id="4C43998D167FE35DADC5F371BFDC2A55" authority="Ng & Lim, 1986" authorityName="Ng & Lim" authorityYear="1986" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="malayanum">
|
||
<emphasis id="B9373E1C167FE35DADC5F371B9232A74" box="[1213,1377,317,341]" italics="true" pageId="3" pageNumber="618">G. malayanum</emphasis>
|
||
Ng & Lim, 1986
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C43998D167FE35DAADFF311B8D62A55" authority="Ng, 1986" authorityName="Ng" authorityYear="1986" box="[935,1172,348,373]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="scandens">
|
||
<emphasis id="B9373E1C167FE35DAADFF311B86B2A54" box="[935,1065,349,373]" italics="true" pageId="3" pageNumber="618">G. scandens</emphasis>
|
||
<bibRefCitation id="EFD29FFF167FE35DAD57F311B8D62A55" author="Ng PKL" box="[1071,1172,348,373]" pageId="3" pageNumber="618" pagination="36 - 44" refId="ref3564" refString="Ng PKL (1986) Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea: Decapoda: Brachyura) from South East Asia. Journal of the Singapore National Academy of Science, 15: 36 - 44." type="journal article" year="1986">Ng, 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C43998D167FE35DADE6F311B9E52A55" authority="Ng, 1986" authorityName="Ng" authorityYear="1986" box="[1182,1447,348,373]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="tiomanicum">
|
||
<emphasis id="B9373E1C167FE35DADE6F311B97E2A55" box="[1182,1340,349,372]" italics="true" pageId="3" pageNumber="618">G. tiomanicum</emphasis>
|
||
<bibRefCitation id="EFD29FFF167FE35DAC3AF311B9E52A55" author="Ng PKL" box="[1346,1447,348,373]" pageId="3" pageNumber="618" pagination="36 - 44" refId="ref3564" refString="Ng PKL (1986) Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea: Decapoda: Brachyura) from South East Asia. Journal of the Singapore National Academy of Science, 15: 36 - 44." type="journal article" year="1986">Ng, 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
, from Peninsular
|
||
<collectingCountry id="F354A29E167FE35DAA9CF331B80A2AB4" box="[996,1096,381,405]" name="Malaysia" pageId="3" pageNumber="618">Malaysia</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C43998D167FE35DADFFF331B9282AB5" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[1159,1386,380,405]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B9373E1C167FE35DADFFF331B8B52AB4" box="[1159,1271,381,405]" italics="true" pageId="3" pageNumber="618">G. katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="F354A29E167FE35DAA55F3D1BFCE2A94" box="[813,908,413,437]" name="Malaysia" pageId="3" pageNumber="618">Sarawak</collectingCountry>
|
||
, East
|
||
<collectingCountry id="F354A29E167FE35DAAB6F3D1B8712A94" box="[974,1075,413,437]" name="Malaysia" pageId="3" pageNumber="618">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFCE20E167FE35DAA55F391BFB92E94" blockId="3.[811,1452,477,1461]" pageId="3" pageNumber="618">
|
||
<taxonomicName id="4C43998D167FE35DAA55F391BF9A2AD5" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[813,984,477,500]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="raj" status="sp. nov.">
|
||
<emphasis id="B9373E1C167FE35DAA55F391BF9A2AD5" box="[813,984,477,500]" italics="true" pageId="3" pageNumber="618">Geosesarma raj</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167FE35DAA9EF391B8282AD4" box="[998,1130,477,501]" pageId="3" pageNumber="618" rank="species">new species</taxonomicNameLabel>
|
||
, can be distinguished from
|
||
<taxonomicName id="4C43998D167FE35DACEAF391BFCD2934" authorityName="Ng" authorityYear="1986" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="scandens">
|
||
<emphasis id="B9373E1C167FE35DACEAF391BFCD2934" italics="true" pageId="3" pageNumber="618">G. scandens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54F85CAC167FE35DAAE3F3B1BF882934" box="[923,970,509,533]" pageId="3" pageNumber="618">type</typeStatus>
|
||
locality: Fraser’s Hill) by its relatively longer third maxilliped merus (
|
||
<figureCitation id="1378FE8B167FE35DAD40F051B8CF2914" box="[1080,1165,541,565]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3B</figureCitation>
|
||
) (shorter in
|
||
<taxonomicName id="4C43998D167FE35DAC66F051B9E62914" authorityName="Ng" authorityYear="1986" box="[1310,1444,541,565]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="scandens">
|
||
<emphasis id="B9373E1C167FE35DAC66F051B9E62914" box="[1310,1444,541,565]" italics="true" pageId="3" pageNumber="618">G. scandens</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167FE35DAA28F071BFFB2974" author="Ng PKL" box="[848,953,573,597]" pageId="3" pageNumber="618" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 62C) and the upward-directed (at a more gradual angle) chitinised distal part of the G1 (
|
||
<figureCitation id="1378FE8B167FE35DAC4AF011B9332954" box="[1330,1393,605,629]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3</figureCitation>
|
||
J–M) (chitinised distal part directed at a steeper angle in
|
||
<taxonomicName id="4C43998D167FE35DACE9F031BFCD2994" authorityName="Ng" authorityYear="1986" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="scandens">
|
||
<emphasis id="B9373E1C167FE35DACE9F031BFCD2994" italics="true" pageId="3" pageNumber="618">G. scandens</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167FE35DAAB9F0D1B86B2994" author="Ng PKL" box="[961,1065,669,693]" pageId="3" pageNumber="618" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 62D, E).
|
||
<taxonomicName id="4C43998D167FE35DADBCF0D1B9322995" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[1220,1392,669,692]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="raj" status="sp. nov.">
|
||
<emphasis id="B9373E1C167FE35DADBCF0D1B9322995" box="[1220,1392,669,692]" italics="true" pageId="3" pageNumber="618">Geosesarma raj</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167FE35DAC07F0D1BF3C29F4" pageId="3" pageNumber="618" rank="species">new species</taxonomicNameLabel>
|
||
, can be separated from
|
||
<taxonomicName id="4C43998D167FE35DAD0CF0F1B95229F4" authorityName="Ng & Lim" authorityYear="1986" box="[1140,1296,701,725]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="malayanum">
|
||
<emphasis id="B9373E1C167FE35DAD0CF0F1B95229F4" box="[1140,1296,701,725]" italics="true" pageId="3" pageNumber="618">G. malayanum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54F85CAC167FE35DAC67F0F1B90C29F4" box="[1311,1358,701,725]" pageId="3" pageNumber="618">type</typeStatus>
|
||
locality:
|
||
<collectingRegion id="49872CEC167FE35DAA54F091BF2B29D4" box="[812,873,733,757]" country="Malaysia" name="Johor" pageId="3" pageNumber="618">Johor</collectingRegion>
|
||
) by its more rugose dorsal surfaces of the carapace (
|
||
<figureCitation id="1378FE8B167FE35DAA4DF0B1BFE42834" box="[821,934,765,789]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="3" pageNumber="618">Fig. 1B, D</figureCitation>
|
||
) (relatively less rugose in
|
||
<taxonomicName id="4C43998D167FE35DADC6F0B1B9192834" authorityName="Ng & Lim" authorityYear="1986" box="[1214,1371,765,789]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="malayanum">
|
||
<emphasis id="B9373E1C167FE35DADC6F0B1B9192834" box="[1214,1371,765,789]" italics="true" pageId="3" pageNumber="618">G. malayanum</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167FE35DACF3F0B1BFFC2814" author="Ng PKL & Lim RP" pageId="3" pageNumber="618" pagination="393 - 402" refId="ref3829" refString="Ng PKL & Lim RP (1987) The taxonomy and biology of the nepenthiphilous freshwater sesarmine crab, Geosesarma malayanum Ng and Lim, 1986 (Crustacea: Decapoda: Brachyura: Grapsidae from Peninsular Malaysia). Malayan Nature Journal, 41: 393 - 402." type="journal article" year="1987">Ng & Lim, 1987</bibRefCitation>
|
||
: pl. 1;
|
||
<bibRefCitation id="EFD29FFF167FE35DAD74F151B8372814" author="Ng PKL" box="[1036,1141,797,821]" pageId="3" pageNumber="618" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 63A); relatively shorter ambulatory legs (
|
||
<figureCitation id="1378FE8B167FE35DAA90F171B8782874" box="[1000,1082,829,853]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="3" pageNumber="618">Fig. 1B</figureCitation>
|
||
) (longer in
|
||
<taxonomicName id="4C43998D167FE35DADC4F171B91B2874" authorityName="Ng & Lim" authorityYear="1986" box="[1212,1369,829,853]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="malayanum">
|
||
<emphasis id="B9373E1C167FE35DADC4F171B91B2874" box="[1212,1369,829,853]" italics="true" pageId="3" pageNumber="618">G. malayanum</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167FE35DACF3F171BFFC2854" author="Ng PKL & Lim RP" pageId="3" pageNumber="618" pagination="393 - 402" refId="ref3829" refString="Ng PKL & Lim RP (1987) The taxonomy and biology of the nepenthiphilous freshwater sesarmine crab, Geosesarma malayanum Ng and Lim, 1986 (Crustacea: Decapoda: Brachyura: Grapsidae from Peninsular Malaysia). Malayan Nature Journal, 41: 393 - 402." type="journal article" year="1987">Ng & Lim, 1987</bibRefCitation>
|
||
: pl. 1;
|
||
<bibRefCitation id="EFD29FFF167FE35DAD71F111B8332854" author="Ng PKL" box="[1033,1137,861,885]" pageId="3" pageNumber="618" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 63A); and the G1 which is slightly less stout, with the chitinised distal part relatively longer (
|
||
<figureCitation id="1378FE8B167FE35DAAFBF1D1BF862894" box="[899,964,925,949]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3</figureCitation>
|
||
J–M) (stouter with the chitinised distal part relatively short in
|
||
<taxonomicName id="4C43998D167FE35DAA8CF1F1B8D328F4" authorityName="Ng & Lim" authorityYear="1986" box="[1012,1169,957,981]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="malayanum">
|
||
<emphasis id="B9373E1C167FE35DAA8CF1F1B8D328F4" box="[1012,1169,957,981]" italics="true" pageId="3" pageNumber="618">G. malayanum</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167FE35DADBCF1F1B93F28F4" author="Ng PKL & Lim RP" box="[1220,1405,957,981]" pageId="3" pageNumber="618" pagination="393 - 402" refId="ref3829" refString="Ng PKL & Lim RP (1987) The taxonomy and biology of the nepenthiphilous freshwater sesarmine crab, Geosesarma malayanum Ng and Lim, 1986 (Crustacea: Decapoda: Brachyura: Grapsidae from Peninsular Malaysia). Malayan Nature Journal, 41: 393 - 402." type="journal article" year="1987">Ng & Lim, 1987</bibRefCitation>
|
||
: fig. 1;
|
||
<bibRefCitation id="EFD29FFF167FE35DAA31F191BFF028D4" author="Ng PKL" box="[841,946,989,1013]" pageId="3" pageNumber="618" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 63D–G). The new species differs from
|
||
<taxonomicName id="4C43998D167FE35DACE9F191BFE92F35" authorityName="Ng" authorityYear="1986" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="tiomanicum">
|
||
<emphasis id="B9373E1C167FE35DACE9F191BFE92F35" italics="true" pageId="3" pageNumber="618">G. tiomanicum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54F85CAC167FE35DAAC3F1B1BFA82F34" box="[955,1002,1021,1045]" pageId="3" pageNumber="618">type</typeStatus>
|
||
locality: Pulau Tioman) by its relatively longer third maxilliped merus (
|
||
<figureCitation id="1378FE8B167FE35DADFCF651B89B2F14" box="[1156,1241,1053,1077]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3B</figureCitation>
|
||
) (relatively shorter merus in
|
||
<taxonomicName id="4C43998D167FE35DAAE0F671B87F2F75" authorityName="Ng" authorityYear="1986" box="[920,1085,1085,1108]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="tiomanicum">
|
||
<emphasis id="B9373E1C167FE35DAAE0F671B87F2F75" box="[920,1085,1085,1108]" italics="true" pageId="3" pageNumber="618">G. tiomanicum</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167FE35DAD0DF671B8A32F74" author="Ng PKL" box="[1141,1249,1085,1109]" pageId="3" pageNumber="618" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 61C), the tip of the chitinised distal part of the G1 being subtruncate (
|
||
<figureCitation id="1378FE8B167FE35DAA4DF631BF312FB4" box="[821,883,1149,1173]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3</figureCitation>
|
||
J–M) (tip sharp; cf.
|
||
<bibRefCitation id="EFD29FFF167FE35DAD36F631B8F42FB4" author="Ng PKL" box="[1102,1206,1149,1173]" pageId="3" pageNumber="618" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 61D, E). From
|
||
<taxonomicName id="4C43998D167FE35DACE9F631BF382F94" authorityName="Ng" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B9373E1C167FE35DACE9F631BF382F94" italics="true" pageId="3" pageNumber="618">G. katibas</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54F85CAC167FE35DAAF3F6D1BFF92F94" box="[907,955,1181,1205]" pageId="3" pageNumber="618">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F354A29E167FE35DAD5FF6D1B8C52F94" box="[1063,1159,1181,1205]" name="Malaysia" pageId="3" pageNumber="618">Sarawak</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C43998D167FE35DADE5F6D1B8A22F95" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[1181,1248,1181,1204]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="raj" status="sp. nov.">
|
||
<emphasis id="B9373E1C167FE35DADE5F6D1B8A22F95" box="[1181,1248,1181,1204]" italics="true" pageId="3" pageNumber="618">G. raj</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167FE35DAD89F6D1B93A2F94" box="[1265,1400,1181,1205]" pageId="3" pageNumber="618" rank="species">new species</taxonomicNameLabel>
|
||
, can be distinguished by its relatively longer third maxilliped merus (
|
||
<figureCitation id="1378FE8B167FE35DAAF8F691BF942FD4" box="[896,982,1245,1269]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3B</figureCitation>
|
||
) (shorter in
|
||
<taxonomicName id="4C43998D167FE35DAD1EF691B89A2FD4" authorityName="Ng" authorityYear="1995" box="[1126,1240,1245,1269]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B9373E1C167FE35DAD1EF691B89A2FD4" box="[1126,1240,1245,1269]" italics="true" pageId="3" pageNumber="618">G. katibas</emphasis>
|
||
</taxonomicName>
|
||
; cf. Ng, 1995: fig. 11B); the dorsal margin of the dactylus of the male major chela having fewer (8 or 9) and less developed granules (
|
||
<figureCitation id="1378FE8B167FE35DAA4DF771BFCB2E74" box="[821,905,1341,1365]" captionStart="Fig" captionStartId="1.[812,847,1977,1999]" captionTargetBox="[812,1451,1572,1954]" captionTargetId="figure-266@1.[812,1451,1572,1954]" captionTargetPageId="1" captionText="Fig. 2.A, C, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, frontal view of carapace; C, D, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4504172" httpUri="https://zenodo.org/record/4504172/files/figure.png" pageId="3" pageNumber="618">Fig. 2B</figureCitation>
|
||
) (11 distinct granules; cf. Ng, 1995: fig. 11C, D); and the G1 being proportionately longer and less stout (
|
||
<figureCitation id="1378FE8B167FE35DACF8F711BF7A2EB4" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="3" pageNumber="618">Fig. 3</figureCitation>
|
||
J–M) (relatively shorter and stouter in
|
||
<taxonomicName id="4C43998D167FE35DAD9BF731B9122EB4" authorityName="Ng" authorityYear="1995" box="[1251,1360,1405,1429]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B9373E1C167FE35DAD9BF731B9122EB4" box="[1251,1360,1405,1429]" italics="true" pageId="3" pageNumber="618">G. katibas</emphasis>
|
||
</taxonomicName>
|
||
; cf. Ng, 1995: fig. 12A–E).
|
||
</paragraph>
|
||
<paragraph id="8BFCE20E167FE35DAA55F791BFE62C15" blockId="3.[811,1454,1501,1845]" pageId="3" pageNumber="618">
|
||
In many other parts of Peninsular
|
||
<collectingCountry id="F354A29E167FE35DADD8F791B9402ED4" box="[1184,1282,1501,1525]" name="Malaysia" pageId="3" pageNumber="618">Malaysia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F354A29E167FE35DAC41F791B9EA2ED4" box="[1337,1448,1501,1525]" name="Singapore" pageId="3" pageNumber="618">Singapore</collectingCountry>
|
||
, there is one lowland species living in swampy areas, often in burrows, and another that occurs in higher altitudes, under rocks in fast-flowing streams. In Johor and
|
||
<collectingCountry id="F354A29E167FE35DAC77F471B93C2D74" box="[1295,1406,1597,1621]" name="Singapore" pageId="3" pageNumber="618">Singapore</collectingCountry>
|
||
, for example,
|
||
<taxonomicName id="4C43998D167FE35DAAECF411B8542D55" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[916,1046,1629,1652]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="B9373E1C167FE35DAAECF411B8542D55" box="[916,1046,1629,1652]" italics="true" pageId="3" pageNumber="618">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
occurs in lowland freshwater swamps while
|
||
<taxonomicName id="4C43998D167FE35DAA09F431BFAA2DB5" authorityName="P.K.L.Ng" authorityYear="1986" box="[881,1000,1661,1684]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="nemesis">
|
||
<emphasis id="B9373E1C167FE35DAA09F431BFAA2DB5" box="[881,1000,1661,1684]" italics="true" pageId="3" pageNumber="618">G. nemesis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C43998D167FE35DAD59F431B8FD2DB4" authorityName="Ng & Lim" authorityYear="1986" box="[1057,1215,1661,1685]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="malayanum">
|
||
<emphasis id="B9373E1C167FE35DAD59F431B8FD2DB4" box="[1057,1215,1661,1685]" italics="true" pageId="3" pageNumber="618">G. malayanum</emphasis>
|
||
</taxonomicName>
|
||
prefer high-elevation areas with fast-flowing streams (see
|
||
<bibRefCitation id="EFD29FFF167FE35DADC4F4D1B9352D94" author="Ng PKL & Lim RP" box="[1212,1399,1693,1717]" pageId="3" pageNumber="618" pagination="393 - 402" refId="ref3829" refString="Ng PKL & Lim RP (1987) The taxonomy and biology of the nepenthiphilous freshwater sesarmine crab, Geosesarma malayanum Ng and Lim, 1986 (Crustacea: Decapoda: Brachyura: Grapsidae from Peninsular Malaysia). Malayan Nature Journal, 41: 393 - 402." type="journal article" year="1987">Ng & Lim, 1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD29FFF167FE35DACFCF4D1BF272DF4" author="Ng PKL" pageId="3" pageNumber="618" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD29FFF167FE35DAA0BF4F1BFEF2DF4" author="Ng PKL" box="[883,941,1725,1749]" pageId="3" pageNumber="618" pagination="189 - 204" refId="ref3658" refString="Ng PKL (1990) The freshwater crabs and prawns of Singapore. In: Chou LM & Ng PKL (eds.) Essays in Zoology. Department of Zoology, National University of Singapore. Pp. 189 - 204." type="book chapter" year="1990">1990</bibRefCitation>
|
||
). The same situation appears to be present in Pulau Bintan, with
|
||
<taxonomicName id="4C43998D167FE35DAD7AF491B82B2DD4" authorityName="Schubart & Ng" authorityYear="2014" box="[1026,1129,1757,1781]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="bintan">
|
||
<emphasis id="B9373E1C167FE35DAD7AF491B82B2DD4" box="[1026,1129,1757,1781]" italics="true" pageId="3" pageNumber="618">G. bintan</emphasis>
|
||
</taxonomicName>
|
||
as the lowland species while
|
||
<taxonomicName id="4C43998D167FE35DAA55F4B1BF2F2C35" authorityName="Schubart & Ng" authorityYear="2014" box="[813,877,1789,1812]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="618" phylum="Arthropoda" rank="species" species="raj">
|
||
<emphasis id="B9373E1C167FE35DAA55F4B1BF2F2C35" box="[813,877,1789,1812]" italics="true" pageId="3" pageNumber="618">G. raj</emphasis>
|
||
</taxonomicName>
|
||
occurs in more montane habitats, but not necessarily in streams.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |