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<document id="20EBC242F04CE2364844D756BAFFF902" ID-CLB-Dataset="21955" ID-DOI="10.5852/ejt.2017.368" ID-GBIF-Dataset="7d707e31-1104-4ed3-8493-ef9302b98796" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838412" ID-ZooBank="E4AEAA16-A546-48B8-99BA-2E0152C8C6F0" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1590087673171" checkinUser="carolina" docAuthor="Zlatkov, Boyan &amp; Huemer, Peter" docDate="2017" docId="03E9996C14444E70B53F6D1A85E2D817" docLanguage="en" docName="ejt-2017-368.pdf.imf" docOrigin="European Journal of Taxonomy 368" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phtheochroa apenninana Zlatkov &amp; Huemer 2017, sp. nov." docType="treatment" docUuid="12684A15-E318-488F-9908-F8D8541C089C" docUuidSource="ZooBank" docVersion="11" lastPageNumber="14" masterDocId="FFD0E114144F4E7DB7706C11845EDC35" masterDocTitle="Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="12" updateTime="1737036481411" updateUser="carolina" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6609695B5C93DD2437C0E70887B9A7E6">Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)</mods:title>
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<mods:namePart id="BEEAB8F0932135CD7E5FBB61B6D8BB3C">Zlatkov, Boyan</mods:namePart>
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<mods:affiliation id="7B5660FD3BBD8A29A61EFE8A42C54F59">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. &amp; National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. &amp; urn: lsid: zoobank. org: author: 3538 D 3 F 5 - 6822 - 403 F-B 9 E 6 - E 8 A 95 E 884404 &amp; Corresponding author: bzlatkov @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="C9878B065B52B49291B4B54B38B030EF" type="email">bzlatkov@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5B18F649EE092919A1F67A91ACAAAD34">Huemer, Peter</mods:namePart>
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<mods:affiliation id="06A80FC05DD8A396F29FB5439D11A557">Tiroler Landesmuseen Betriebsgesellschaft m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria. &amp; Email: p. huemer @ tiroler-landesmuseen. at &amp; urn: lsid: zoobank. org: author: 4 E 6 F 001 E-DB 9 D- 460 C-AB 39 - 0 CE 7 CC 773 B 59</mods:affiliation>
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<mods:title id="56101126B5CA7808BEE7E97504C3EE73">European Journal of Taxonomy</mods:title>
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<mods:date id="42200A3878C79BD9D13E0CEB38CCA8E6">2017</mods:date>
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<mods:number id="F03751F35D5C158C633E323191B814D6">2017-11-21</mods:number>
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<treatment id="03E9996C14444E70B53F6D1A85E2D817" ID-DOI="http://doi.org/10.5281/zenodo.3852270" ID-GBIF-Taxon="164246401" ID-Zenodo-Dep="3852270" LSID="urn:lsid:zoobank.org:act:12684A15-E318-488F-9908-F8D8541C089C" httpUri="http://treatment.plazi.org/id/03E9996C14444E70B53F6D1A85E2D817" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" scope_class="Insecta" scope_family="Tortricidae" scope_order="Lepidoptera" scope_subFamily="Tortricinae" scope_superFamily="Tortricoidea" scope_tribe="Cochylini">
<subSubSection id="C35A7BF114444E76B53F6D1A87BBDD13" box="[591,997,267,294]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BFF287A14444E76B53F6D1A87BBDD13" blockId="11.[378,1209,267,364]" box="[591,997,267,294]" pageId="11" pageNumber="12">
<heading id="D0B79F1614444E76B53F6D1A87BBDD13" box="[591,997,267,294]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<taxonomicName id="4C4053F914444E76B53F6D1A87DDDD10" ID-CoL="B3ZCF" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[591,899,267,294]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F46814444E76B53F6D1A87DDDD10" bold="true" box="[591,899,267,294]" italics="true" pageId="11" pageNumber="12">Phtheochroa apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314444E76B4FB6D1D87BBDD13" box="[907,997,268,294]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114444E76B60A6D3F8078DD59" pageId="11" pageNumber="12" type="description">
<paragraph id="8BFF287A14444E76B60A6D3F80E7DD7C" blockId="11.[378,1209,267,364]" box="[378,1209,302,329]" pageId="11" pageNumber="12">
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urn:lsid:zoobank.org:act:
<uuid id="FFE612AF14444E76B5EB6D3F80E7DD7C" box="[667,1209,302,329]" pageId="11" pageNumber="12">12684A15-E318-488F-9908-F8D8541C089C</uuid>
</uri>
</paragraph>
<paragraph id="8BFF287A14444E76B57E6D438078DD59" blockId="11.[378,1209,267,364]" box="[526,1062,338,364]" pageId="11" pageNumber="12">
<figureCitation id="137B34FF14444E76B57E6D438630DD59" box="[526,622,338,364]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="11" pageNumber="12">Figs 1G</figureCitation>
,
<figureCitation id="137B34FF14444E76B50A6D4386FEDD59" box="[634,672,338,364]" captionStart="Fig" captionStartId="5.[189,232,1813,1839]" captionTargetBox="[230,1355,468,1774]" captionTargetId="figure@5.[189,1398,417,1778]" captionTargetPageId="5" captionText="Fig. 2. Male genitalia (without phalli) of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov. A. Bulgaria, Rila Mts. B. Republic of Macedonia, Korab Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838416" httpUri="https://zenodo.org/record/3838416/files/figure.png" pageId="11" pageNumber="12">2D</figureCitation>
,
<figureCitation id="137B34FF14444E76B5DD6D43868DDD59" box="[685,723,338,364]" captionStart="Fig" captionStartId="6.[188,231,1807,1833]" captionTargetBox="[191,1389,1348,1764]" captionTargetId="figure@6.[188,1397,1344,1765]" captionTargetPageId="6" captionText="Fig. 3. Transtilla of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov., Bulgaria, Rila Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, paratype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838418" httpUri="https://zenodo.org/record/3838418/files/figure.png" pageId="11" pageNumber="12">3D</figureCitation>
,
<figureCitation id="137B34FF14444E76B5AF6D4386B3DD59" box="[735,749,338,364]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="11" pageNumber="12">4</figureCitation>
F–G, 6C–D, 9, 10;
<tableCitation id="C6C21DC114444E76B4BF6D438078DD59" box="[975,1062,338,364]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="11" pageNumber="12" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114444E76B7CD6D8285AADEE7" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BFF287A14444E76B7CD6D828568DD98" blockId="11.[189,310,403,429]" box="[189,310,403,429]" pageId="11" pageNumber="12">
<heading id="D0B79F1614444E76B7CD6D828568DD98" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B934F46814444E76B7CD6D828568DD98" bold="true" box="[189,310,403,429]" pageId="11" pageNumber="12">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A14444E76B7CD6DD385AADEE7" blockId="11.[189,1399,450,722]" pageId="11" pageNumber="12">
Externally,
<taxonomicName id="4C4053F914444E76B6376DD285AFDDE9" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[327,497,451,476]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F46814444E76B6376DD285AFDDE9" box="[327,497,451,476]" italics="true" pageId="11" pageNumber="12">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314444E76B68B6DD38606DDE9" box="[507,600,450,476]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C4053F914444E76B5916DD287D2DDE9" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[737,908,450,476]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814444E76B5916DD287D2DDE9" box="[737,908,450,476]" italics="true" pageId="11" pageNumber="12">P. schawerdae</emphasis>
</taxonomicName>
but the male genitalia are different. The vesica in
<taxonomicName id="4C4053F914444E76B65A6DF7858FDDCA" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[298,465,486,511]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F46814444E76B65A6DF7858FDDCA" box="[298,465,486,511]" italics="true" pageId="11" pageNumber="12">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314444E76B6A76DF4866FDDCA" box="[471,561,485,511]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
has a larger right part than in
<taxonomicName id="4C4053F914444E76B4FA6DF7806FDDCA" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[906,1073,485,511]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814444E76B4FA6DF7806FDDCA" box="[906,1073,485,511]" italics="true" pageId="11" pageNumber="12">P. schawerdae</emphasis>
</taxonomicName>
; the angle between the axes of the phallus and right diverticulum is smaller (maximum 90°) than in
<taxonomicName id="4C4053F914444E76B3756E1880F3DE17" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[1029,1197,520,546]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814444E76B3756E1880F3DE17" box="[1029,1197,520,546]" italics="true" pageId="11" pageNumber="12">P. schawerdae</emphasis>
</taxonomicName>
(100–120°). The phallic process in
<taxonomicName id="4C4053F914444E76B6E66E3D8661DE70" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[406,575,556,581]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F46814444E76B6E66E3D8661DE70" box="[406,575,556,581]" italics="true" pageId="11" pageNumber="12">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314444E76B5376E3D86FDDE73" box="[583,675,556,582]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
is shorter, wider and not strongly curved; the opposite is the case in
<taxonomicName id="4C4053F914444E76B6656E5E85E3DE5D" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[277,445,590,616]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814444E76B6656E5E85E3DE5D" box="[277,445,590,616]" italics="true" pageId="11" pageNumber="12">P. schawerdae</emphasis>
</taxonomicName>
. Both the wing colouration and male genitalia distinguish
<taxonomicName id="4C4053F914444E76B31D6E5E814BDE5D" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[1133,1301,591,616]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F46814444E76B31D6E5E814BDE5D" box="[1133,1301,591,616]" italics="true" pageId="11" pageNumber="12">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314444E76B26C6E5E8129DE5C" box="[1308,1399,591,617]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
from
<taxonomicName id="4C4053F914444E76B78D6E6285DEDEB9" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[253,384,626,652]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="alpinana" status="sp. nov.">
<emphasis id="B934F46814444E76B78D6E6285DEDEB9" box="[253,384,626,652]" italics="true" pageId="11" pageNumber="12">P. alpinana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314444E76B6F76E6385BADEB9" box="[391,484,626,652]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C4053F914444E76B69E6E628629DEB9" authorityName="Guenee, 1845 Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[494,631,626,652]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F46814444E76B69E6E628629DEB9" box="[494,631,626,652]" italics="true" pageId="11" pageNumber="12">P. frigidana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4053F914444E76B5DF6E628704DEB9" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[687,858,626,652]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
<emphasis id="B934F46814444E76B5DF6E628704DEB9" box="[687,858,626,652]" italics="true" pageId="11" pageNumber="12">P. cantabriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314444E76B4116E6387E5DEB9" box="[865,955,626,652]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
The vesicae of
<taxonomicName id="4C4053F914444E76B3006E628149DEB9" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[1136,1303,627,652]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F46814444E76B3006E628149DEB9" box="[1136,1303,627,652]" italics="true" pageId="11" pageNumber="12">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314444E76B26D6E638129DEB9" box="[1309,1399,626,652]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C4053F914444E76B79F6E8785C4DE9A" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[239,410,661,687]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
<emphasis id="B934F46814444E76B79F6E8785C4DE9A" box="[239,410,661,687]" italics="true" pageId="11" pageNumber="12">P. cantabriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314444E76B6D26E8485A2DE9A" box="[418,508,661,687]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
are similar but the angle between the axes of phallus and right diverticulum in
<taxonomicName id="4C4053F914444E76B7AB6EA885D9DEE7" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[219,391,696,722]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cantabriana">
<emphasis id="B934F46814444E76B7AB6EA885D9DEE7" box="[219,391,696,722]" italics="true" pageId="11" pageNumber="12">P. cantabriana</emphasis>
</taxonomicName>
is larger.
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114444E76B7CD6EEB80D4DF77" box="[189,1162,762,835]" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8BFF287A14444E76B7CD6EEB851ADF21" blockId="11.[189,324,762,788]" box="[189,324,762,788]" pageId="11" pageNumber="12">
<heading id="D0B79F1614444E76B7CD6EEB851ADF21" bold="true" box="[189,324,762,788]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B934F46814444E76B7CD6EEB851ADF21" bold="true" box="[189,324,762,788]" pageId="11" pageNumber="12">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A14444E76B7CD6F3980D4DF77" blockId="11.[189,1162,808,835]" box="[189,1162,808,835]" pageId="11" pageNumber="12">The specific name is a feminine adjective derived from the name of the Apennines.</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114444E76B7CD6F788166D911" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BFF287A14444E76B7CD6F7885F4DFB6" blockId="11.[189,426,873,899]" box="[189,426,873,899]" pageId="11" pageNumber="12">
<heading id="D0B79F1614444E76B7CD6F7885F4DFB6" bold="true" box="[189,426,873,899]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B934F46814444E76B7CD6F7885F4DFB6" bold="true" box="[189,426,873,899]" pageId="11" pageNumber="12">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B28222714444E76B79A6F898089D87F" ID-GBIF-Occurrence="2621725301" collectingDate="2010-07-15" collectionCode="TLMF" collectorName="Huemer" country="Italy" elevation="1750" latitude="42.426666" location="Prov. L'Aquila NP Gran Sasso ex Miniera di Lignite" longLatPrecision="114" longitude="13.713333" municipality="Miniera di Lignite" pageId="11" pageNumber="12" specimenCode="TLMF 2010" specimenCount="1" specimenCount-male="1" stateProvince="Prov. L'Aquila NP Gran Sasso ex Miniera di Lignite" typeStatus="holotype">
<paragraph id="8BFF287A14444E76B79A6F898502DF87" blockId="11.[234,348,920,946]" box="[234,348,920,946]" pageId="11" pageNumber="12">
<emphasis id="B934F46814444E76B79A6F898502DF87" bold="true" box="[234,348,920,946]" pageId="11" pageNumber="12">
<typeStatus id="54FB96D814444E76B79A6F898502DF87" box="[234,348,920,946]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFF287A14444E76B7CD6FD68089D87F" blockId="11.[189,1398,966,1099]" pageId="11" pageNumber="12">
<collectingCountry id="F35768EA14444E76B7CD6FD6854CDFD4" box="[189,274,967,993]" name="Italy" pageId="11" pageNumber="12">ITALY</collectingCountry>
:
<specimenCount id="9D46E3F314444E76B66E6FD68568DFD4" box="[286,310,967,993]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
, pinned, with partially spread wings, slightly worn, four labels: // Italia,
<collectingRegion id="4984E69814444E76B3EA6FD6860FD831" pageId="11" pageNumber="12">Prov. L’Aquila NP Gran Sasso ex Miniera di Lignite</collectingRegion>
<geoCoordinate id="EE744EBD14444E76B52B6FF98682D831" box="[603,732,1000,1028]" degrees="13" direction="east" minutes="42,8" orientation="longitude" pageId="11" pageNumber="12" precision="92" value="13.713333">13°42,8′ E</geoCoordinate>
,
<geoCoordinate id="EE744EBD14444E76B59B6FF98733D831" box="[747,877,1000,1028]" degrees="42" direction="north" minutes="25,6" orientation="latitude" pageId="11" pageNumber="12" precision="92" value="42.426666">42°25,6′ N</geoCoordinate>
<quantity id="4CB8859F14444E76B4076FFB878BD831" box="[887,981,1002,1029]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="11" pageNumber="12" unit="m" value="1750.0">
<elevation id="006DCF4914444E76B4076FFB878BD831" box="[887,981,1002,1029]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="11" pageNumber="12" unit="m" value="1750.0">1750 m</elevation>
</quantity>
,
<collectingDate id="EFBAF75214444E76B4946FFB80DFD830" box="[996,1153,1002,1029]" pageId="11" pageNumber="12">
14-
<date id="FFFE0EBA14444E76B3776FFB80DFD830" box="[1031,1153,1002,1029]" pageId="11" pageNumber="12" value="2010-07-15">15.7.2010</date>
</collectingDate>
leg.
<collectorName id="26B54DAC14444E76B3B06FFB8140D831" box="[1216,1310,1002,1028]" pageId="11" pageNumber="12">Huemer</collectorName>
<specimenCode id="DBE6800114444E76B2586FFB84A7D812" pageId="11" pageNumber="12">TLMF 2010</specimenCode>
-020 // BC TLMF Lep 01572 [green label] // Gen. prep. ♂ 1/14-
<date id="FFFE0EBA14444E76B4AB681C8006D81D" box="[987,1112,1037,1064]" pageId="11" pageNumber="12" value="2010-07-15">15.7.2010</date>
[genitalia slide number] // HOLOTYPE
<taxonomicName id="4C4053F914444E76B60768218790D87F" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[375,974,1072,1099]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apenninana">
<emphasis id="B934F46814444E76B607682186FED87F" box="[375,672,1072,1098]" italics="true" pageId="11" pageNumber="12">Phtheochroa apenninana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14444E76B30668218095D87F" box="[1142,1227,1072,1098]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="11" pageNumber="12">TLMF</collectionCode>
).
</paragraph>
</materialsCitation>
<materialsCitation id="3B28222714444E76B79A68638166D911" collectingDate="2010-07-13" collectionCode="TLMF" collectorName="Huemer" country="Italy" elevation="2010" latitude="42.448334" location="Campo Imperatore" longLatPrecision="114" longitude="13.573334" pageId="11" pageNumber="12" specimenCode="TLMF 2010" specimenCount="1" specimenCount-male="1" stateProvince="Prov. L'Aquila NP Gran Sasso" typeStatus="paratype">
<paragraph id="8BFF287A14444E76B79A68638502D8B9" blockId="11.[234,348,1138,1164]" box="[234,348,1138,1164]" pageId="11" pageNumber="12">
<emphasis id="B934F46814444E76B79A68638502D8B9" bold="true" box="[234,348,1138,1164]" pageId="11" pageNumber="12">
<typeStatus id="54FB96D814444E76B79A68638502D8B9" box="[234,348,1138,1164]" pageId="11" pageNumber="12" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFF287A14444E76B7CD68B18166D911" blockId="11.[189,1398,1184,1316]" pageId="11" pageNumber="12">
<collectingCountry id="F35768EA14444E76B7CD68B1854CD88F" box="[189,274,1184,1210]" name="Italy" pageId="11" pageNumber="12">ITALY</collectingCountry>
:
<specimenCount id="9D46E3F314444E76B66D68B18515D88E" box="[285,331,1184,1211]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
, pinned, with partially spread wings, slightly worn, four labels: // Italia,
<collectingRegion id="4984E69814444E76B3ED68B0851BD8E8" pageId="11" pageNumber="12">Prov. L’Aquila NP Gran Sasso</collectingRegion>
,
<location id="8E9F7EA114444E76B62068D28671D8EB" LSID="urn:lsid:plazi:treatment:03E9996C14444E70B53F6D1A85E2D817:8E9F7EA114444E76B62068D28671D8EB" box="[336,559,1219,1246]" country="Italy" latitude="42.448334" longLatPrecision="114" longitude="13.573334" name="Campo Imperatore" pageId="11" pageNumber="12" stateProvince="Prov. L'Aquila NP Gran Sasso">Campo Imperatore</location>
,
<location id="8E9F7EA114444E76B54A68D586B2D8E8" LSID="urn:lsid:plazi:treatment:03E9996C14444E70B53F6D1A85E2D817:8E9F7EA114444E76B54A68D586B2D8E8" box="[570,748,1219,1246]" country="Italy" latitude="42.448334" longLatPrecision="114" longitude="13.573334" name="E Observatorio" pageId="11" pageNumber="12" stateProvince="Prov. L'Aquila NP Gran Sasso">E Observatorio</location>
<geoCoordinate id="EE744EBD14444E76B58368D3872FD8EB" box="[755,881,1218,1246]" degrees="13" direction="east" minutes="34,4" orientation="longitude" pageId="11" pageNumber="12" precision="92" value="13.573334">13°34,4′ E</geoCoordinate>
,
<geoCoordinate id="EE744EBD14444E76B40D68D387A2D8EB" box="[893,1020,1218,1246]" degrees="42" direction="north" minutes="26,9" orientation="latitude" pageId="11" pageNumber="12" precision="92" value="42.448334">42°26,9′ N</geoCoordinate>
<quantity id="4CB8859F14444E76B37368D28000D8EB" box="[1027,1118,1219,1246]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="11" pageNumber="12" unit="m" value="2010.0">
<elevation id="006DCF4914444E76B37368D28000D8EB" box="[1027,1118,1219,1246]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="11" pageNumber="12" unit="m" value="2010.0">2010 m</elevation>
</quantity>
,
<date id="FFFE0EBA14444E76B31A68D28081D8E8" box="[1130,1247,1219,1245]" pageId="11" pageNumber="12" value="2010-07-13">
<collectingDate id="EFBAF75214444E76B31A68D28081D8E8" box="[1130,1247,1219,1245]" pageId="11" pageNumber="12" value="2010-07-13">13.7.2010</collectingDate>
</date>
leg.
<collectorName id="26B54DAC14444E76B26868D58128D8EB" box="[1304,1398,1220,1246]" pageId="11" pageNumber="12">Huemer</collectorName>
<specimenCode id="DBE6800114444E76B7CD68F6850CD934" box="[189,338,1255,1281]" pageId="11" pageNumber="12">TLMF 2010</specimenCode>
-020 // BC TLMF Lep 03037 [green label] // Gen. prep. ♂ 1/
<date id="FFFE0EBA14444E76B33068F680E4D934" box="[1088,1210,1255,1281]" pageId="11" pageNumber="12" value="2010-07-13">13.7.2010</date>
[genitalia slide number] // PARATYPE
<taxonomicName id="4C4053F914444E76B6A8691B806ED911" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[472,1072,1290,1316]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apenninana">
<emphasis id="B934F46814444E76B6A8691B875FD911" box="[472,769,1290,1316]" italics="true" pageId="11" pageNumber="12">Phtheochroa apenninana</emphasis>
Zlatkov &amp; Huemer, 2017
</taxonomicName>
[red label] // (
<collectionCode id="ED51B0BF14444E76B3A7691B8172D911" box="[1239,1324,1290,1316]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="11" pageNumber="12">TLMF</collectionCode>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C35A7BF114444E70B7CD695A8146DF2A" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="description">
<paragraph id="8BFF287A14444E76B7CD695A8510D950" blockId="11.[189,334,1355,1381]" box="[189,334,1355,1381]" pageId="11" pageNumber="12">
<heading id="D0B79F1614444E76B7CD695A8510D950" bold="true" box="[189,334,1355,1381]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B934F46814444E76B7CD695A8510D950" bold="true" box="[189,334,1355,1381]" pageId="11" pageNumber="12">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A14444E76B79A696B85C1D9A1" blockId="11.[234,415,1402,1428]" box="[234,415,1402,1428]" pageId="11" pageNumber="12">
<emphasis id="B934F46814444E76B79A696B8574D9A1" bold="true" box="[234,298,1402,1428]" pageId="11" pageNumber="12">Male</emphasis>
(
<figureCitation id="137B34FF14444E76B64A696B85CAD9A1" box="[314,404,1402,1428]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="11" pageNumber="12">Fig. 1G</figureCitation>
)
</paragraph>
<paragraph id="8BFF287A14444E76B7CD69B88149D9D0" blockId="11.[189,1398,1448,1510]" pageId="11" pageNumber="12">HEAD. Frons devoid of scales (worn specimens), vertex yellow brown, labial palps creamy. Antennae with scapus and pedicellus brown dorsally and creamy ventrally, flagellum with creamy scales.</paragraph>
<paragraph id="8BFF287A14444E76B7CD6A1C87B1DA81" blockId="11.[189,1399,1549,1722]" pageId="11" pageNumber="12">
THORAX. Dorsally, including tegulae, with yellow and brown scales, ventrally whitish, fore- and middle legs brown, hindlegs white. Forewing length
<quantity id="4CB8859F14444E76B5A06A218703DA7E" box="[720,861,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="7.4" metricValueMin="7.2" pageId="11" pageNumber="12" unit="mm" value="7.3" valueMax="7.4" valueMin="7.2">7.2–7.4 mm</quantity>
(n =2), upperside deep yellow, with remnants of rust dorsal blotch, cilia concolorous with upperside, underside dark grey, with white scales in costal area. Hindwing upper- and underside dark grey, with white costal area and white longitudinal line between veins M
<subScript id="17C42A3F14444E76B6F56ABD85D0DA8F" attach="left" box="[389,398,1708,1722]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
and M
<subScript id="17C42A3F14444E76B6916ABD85B4DA8F" attach="left" box="[481,490,1708,1722]" fontSize="6" pageId="11" pageNumber="12">2</subScript>
starting from discal cell and ending in cilia.
</paragraph>
<paragraph id="8BFF287A14444E76B7CD6ACA8524DAC0" blockId="11.[189,378,1755,1782]" box="[189,378,1755,1782]" pageId="11" pageNumber="12">ABDOMEN. Grey.</paragraph>
<paragraph id="8BFF287A14444E70B7CD6B0C872ADE29" blockId="11.[189,1399,1820,2023]" lastBlockId="13.[189,1399,267,540]" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
GENITALIA (
<figureCitation id="137B34FF14444E76B6336B0D85FADB02" box="[323,420,1820,1847]" captionStart="Fig" captionStartId="5.[189,232,1813,1839]" captionTargetBox="[230,1355,468,1774]" captionTargetId="figure@5.[189,1398,417,1778]" captionTargetPageId="5" captionText="Fig. 2. Male genitalia (without phalli) of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov. A. Bulgaria, Rila Mts. B. Republic of Macedonia, Korab Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838416" httpUri="https://zenodo.org/record/3838416/files/figure.png" pageId="11" pageNumber="12">Fig. 2D</figureCitation>
). Valva with weak curvature, appears slender but this may be due to preparation. Rest of valva, uncus and socii very similar to those of
<taxonomicName id="4C4053F914444E76B44E6B5087B9DB6F" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[830,999,1856,1882]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814444E76B44E6B5087B9DB6F" box="[830,999,1856,1882]" italics="true" pageId="11" pageNumber="12">P. schawerdae</emphasis>
</taxonomicName>
. Transtilla (
<figureCitation id="137B34FF14444E76B3056B51808FDB6F" box="[1141,1233,1856,1882]" captionStart="Fig" captionStartId="6.[188,231,1807,1833]" captionTargetBox="[191,1389,1348,1764]" captionTargetId="figure@6.[188,1397,1344,1765]" captionTargetPageId="6" captionText="Fig. 3. Transtilla of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov., Bulgaria, Rila Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, paratype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838418" httpUri="https://zenodo.org/record/3838418/files/figure.png" pageId="11" pageNumber="12">Fig. 3D</figureCitation>
) large, round. Phallus (
<figureCitation id="137B34FF14444E76B6556B728535DB4B" box="[293,363,1891,1918]" captionStart="Fig" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[262,1281,269,1845]" captionTargetId="figure@12.[253,1334,265,1851]" captionTargetPageId="12" captionText="Fig. 6. Phalli with vesica everted of Phtheochroa spp. A–B. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – C–D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype.A, C: left. B, D: dorsal. Abbreviations: dd = dorsal diverticulum; vd = ventral diverticulum; other abbreviations as in Fig. 5. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838424" httpUri="https://zenodo.org/record/3838424/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
C–D) slightly curved ventrally, with wide and blunt medioventral process (
<figureCitation id="137B34FF14444E76B39A6B72816CDB48" box="[1258,1330,1891,1917]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
F–G), with apex curved to the right. Vesica in general resembles that of
<taxonomicName id="4C4053F914444E76B4BD6B968028DB95" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[973,1142,1926,1952]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814444E76B4BD6B968028DB95" box="[973,1142,1926,1952]" italics="true" pageId="11" pageNumber="12">P. schawerdae</emphasis>
</taxonomicName>
, with large main part protruded at right from sclerotized phallus and two large diverticula bearing long, robust cornuti. Small, conical undulating structure (diverticulum) emerges ventrally from main part. Gonopore located dorsally, surrounded by semicylindrical sclerotized plate with minute, lentiform tubercles. Main part medially constricted by circular furrow in which gonopore is sunk. Portion located distally of furrow very large, heart-shaped in dorsal view. Main part of vesica looks ovoid in lateral left view. Diverticula emerge from right portion in opposing directions, right one straight, emerging from ventral half of right part and directed laterally, left one curved and also pointed laterally. Length and orientation of diverticula relatively constant. Axes of right diverticulum and sclerotized phallus form an angle of maximum 90°. Cornuti aciculate, capitate, of equal length, left one slightly bent. Barely discernible acanthae present on left portion of main part of vesica and at end of diverticula.
</paragraph>
<caption id="DF3F78F214434E71B7CD6B778642DBDE" ID-DOI="http://doi.org/10.5281/zenodo.3838424" ID-Zenodo-Dep="3838424" httpUri="https://zenodo.org/record/3838424/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1894,1920]" targetBox="[262,1281,269,1845]" targetPageId="12">
<paragraph id="8BFF287A14434E71B7CD6B778642DBDE" blockId="12.[189,1399,1894,2027]" pageId="12" pageNumber="13">
<emphasis id="B934F46814434E71B7CD6B77854FDBB5" bold="true" box="[189,273,1894,1920]" pageId="12" pageNumber="13">Fig. 6.</emphasis>
Phalli with vesica everted of
<taxonomicName id="4C4053F914434E71B5E36B778777DBB5" authorityName="Stephens" authorityYear="1829" box="[659,809,1894,1920]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B934F46814434E71B5E36B778777DBB5" box="[659,809,1894,1920]" italics="true" pageId="12" pageNumber="13">Phtheochroa</emphasis>
</taxonomicName>
spp.
<emphasis id="B934F46814434E71B4046B7787F5DBB5" bold="true" box="[884,939,1894,1920]" pageId="12" pageNumber="13">A–B</emphasis>
.
<taxonomicName id="4C4053F914434E71B4B16B768012DBB5" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[961,1100,1894,1920]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="alpinana" status="sp. nov.">
<emphasis id="B934F46814434E71B4B16B768012DBB5" box="[961,1100,1894,1920]" italics="true" pageId="12" pageNumber="13">P. alpinana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314434E71B32A6B7680E3DBB4" box="[1114,1213,1895,1921]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, France, Alpes Maritimes, holotype. –
<emphasis id="B934F46814434E71B6B96B98865CDB96" bold="true" box="[457,514,1929,1955]" pageId="12" pageNumber="13">C–D</emphasis>
.
<taxonomicName id="4C4053F914434E71B57F6B9B86EBDB96" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[527,693,1930,1955]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F46814434E71B57F6B9B86EBDB96" box="[527,693,1930,1955]" italics="true" pageId="12" pageNumber="13">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207491314434E71B5CA6B9B874BDB91" box="[698,789,1930,1956]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, Italy, Gran Sasso National Park, holotype.A, C: left. B, D: dorsal. Abbreviations: dd = dorsal diverticulum; vd = ventral diverticulum; other abbreviations as in Fig. 5. Scale bar = 250 µm.
</paragraph>
</caption>
<paragraph id="8BFF287A14424E70B79A6E52851BDE68" blockId="13.[234,325,579,605]" box="[234,325,579,605]" pageId="13" pageNumber="14">
<emphasis id="B934F46814424E70B79A6E52851BDE68" bold="true" box="[234,325,579,605]" pageId="13" pageNumber="14">Female</emphasis>
</paragraph>
<paragraph id="8BFF287A14424E70B7CD6E638567DEB9" blockId="13.[189,313,626,652]" box="[189,313,626,652]" pageId="13" pageNumber="14">Unknown.</paragraph>
<paragraph id="8BFF287A14424E70B7CD6EA2861FDEF8" blockId="13.[189,577,691,717]" box="[189,577,691,717]" pageId="13" pageNumber="14">
<heading id="D0B79F1614424E70B7CD6EA28525DEF8" box="[189,379,691,717]" pageId="13" pageNumber="14">
<emphasis id="B934F46814424E70B7CD6EA28525DEF8" bold="true" box="[189,379,691,717]" pageId="13" pageNumber="14">Molecular data</emphasis>
</heading>
(
<tableCitation id="C6C21DC114424E70B6FB6EA285BADEF8" box="[395,484,691,717]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="13" pageNumber="14" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
,
<figureCitation id="137B34FF14424E70B6826EA28669DEF8" box="[498,567,691,717]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[190,1397,1281,1884]" captionTargetId="figure@18.[172,1412,1251,1902]" captionTargetPageId="18" captionText="Fig. 9. Maximum likelihood tree (built with MEGA6) of cytochrome c oxidase subunit I (COI) barcode fragments. Values at the nodes are bootstrap support values based on 500 replicates." figureDoi="http://doi.org/10.5281/zenodo.3838430" httpUri="https://zenodo.org/record/3838430/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation>
)
</paragraph>
<paragraph id="8BFF287A14424E70B7CD6EF38146DF2A" blockId="13.[189,1398,737,799]" pageId="13" pageNumber="14">
BIN URI: BOLD:ACA9597. The maximum intraspecific divergence of the barcode region is 0% (n= 2). The minimum distance to the nearest neighbour,
<taxonomicName id="4C4053F914424E70B58B6F1787FDDF2A" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[763,931,773,799]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F46814424E70B58B6F1787FDDF2A" box="[763,931,773,799]" italics="true" pageId="13" pageNumber="14">P. schawerdae</emphasis>
</taxonomicName>
from the Dinaric Mts is 3.69%.
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114424E70B7CD6F568741DF87" pageId="13" pageNumber="14" type="biology_ecology">
<paragraph id="8BFF287A14424E70B7CD6F56857EDF54" blockId="13.[189,288,839,865]" box="[189,288,839,865]" pageId="13" pageNumber="14">
<heading id="D0B79F1614424E70B7CD6F56857EDF54" bold="true" box="[189,288,839,865]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B934F46814424E70B7CD6F56857EDF54" bold="true" box="[189,288,839,865]" pageId="13" pageNumber="14">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF287A14424E70B7CD6F648741DF87" blockId="13.[189,1398,885,947]" pageId="13" pageNumber="14">
Preimaginal stages and larval host plant are unknown. The moths fly in July. Habitat: the
<typeStatus id="54FB96D814424E70B3AC6F648151DFBA" box="[1244,1295,885,911]" pageId="13" pageNumber="14">type</typeStatus>
material was collected in alpine grassland above the treeline.
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF114424E70B7CD6FC885E2D817" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BFF287A14424E70B7CD6FC8859ADFC1" blockId="13.[189,452,985,1012]" box="[189,452,985,1012]" pageId="13" pageNumber="14">
<heading id="D0B79F1614424E70B7CD6FC88508DFC6" box="[189,342,985,1011]" pageId="13" pageNumber="14">
<emphasis id="B934F46814424E70B7CD6FC88508DFC6" bold="true" box="[189,342,985,1011]" pageId="13" pageNumber="14">Distribution</emphasis>
</heading>
(
<figureCitation id="137B34FF14424E70B6166FC885E5DFC1" box="[358,443,985,1012]" captionStart="Fig" captionStartId="20.[189,232,1990,2016]" captionTargetBox="[189,1398,1296,1948]" captionTargetId="figure@20.[183,1404,1290,1954]" captionTargetPageId="20" captionText="Fig. 10. Distribution of the Phtheochroa frigidana s. lat. species complex based on examined material." figureDoi="http://doi.org/10.5281/zenodo.3838432" httpUri="https://zenodo.org/record/3838432/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
)
</paragraph>
<paragraph id="8BFF287A14424E70B7CD681985E2D817" blockId="13.[189,444,1032,1058]" box="[189,444,1032,1058]" pageId="13" pageNumber="14">Apennine Mountains.</paragraph>
</subSubSection>
</treatment>
</document>
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