1005 lines
121 KiB
XML
1005 lines
121 KiB
XML
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<mods:title id="438F349AB057BECB4220911817AB8C7D">Description of three new Boulenophrys species from eastern Guangdong, China, emphasizing the urgency of ecological conservation in this region (Anura, Megophryidae)</mods:title>
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<mods:namePart id="A283468680910AA720BAC0FFF32BC6FB">Wang, Jian</mods:namePart>
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<treatment id="03E94554FFF8A02E9BF6D7B8FC12C5DF" ID-DOI="http://doi.org/10.5281/zenodo.6309122" ID-GBIF-Taxon="193331550" ID-Zenodo-Dep="6309122" LSID="urn:lsid:plazi:treatment:03E94554FFF8A02E9BF6D7B8FC12C5DF" httpUri="http://treatment.plazi.org/id/03E94554FFF8A02E9BF6D7B8FC12C5DF" lastPageId="23" lastPageNumber="114" pageId="19" pageNumber="110">
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<taxonomicName id="4C408FC1FFF8A02A9BF6D7B8FE58C402" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[151,502,1773,1800]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="19" pageNumber="110" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
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<emphasis id="B9342850FFF8A02A9BF6D7B8FE58C402" bold="true" box="[151,502,1773,1800]" italics="true" pageId="19" pageNumber="110">Boulenophrys fengshunensis</emphasis>
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<taxonomicNameLabel id="A207952BFFF8A02A9A9FD7B8FDF5C40D" box="[510,603,1773,1799]" pageId="19" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
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Wang, Zeng, Lyu & Wang
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFFF442FFF8A02A9BF6D644FB98C426" blockId="19.[151,1078,1773,1907]" box="[151,1078,1809,1836]" pageId="19" pageNumber="110">Fengshun Horned Toad (in English) / Fēng Shùn Jiǎo Chán (丰NJDzdz in Chinese)</paragraph>
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</subSubSection>
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<emphasis id="B9342850FFF8A02A9BF6D660FE96C445" bold="true" box="[151,312,1845,1871]" pageId="19" pageNumber="110">Chresonymy:</emphasis>
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<taxonomicName id="4C408FC1FFF8A02A9A5ED660FE70C445" box="[319,478,1845,1871]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="19" pageNumber="110" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B9342850FFF8A02A9A5ED660FE13C445" box="[319,445,1845,1871]" italics="true" pageId="19" pageNumber="110">Megophrys</emphasis>
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sp
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</taxonomicName>
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14 —
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<bibRefCitation id="EFD189B3FFF8A02A9942D660FD66C445" author="Liu, Z. Y. & Chen, G. L. & Zhu, T. Q. & Zeng, Z. C. & Lyu, Z. T. & Wang, J. & Messenger, K. & Greenberg, A. J. & Guo, Z. X. & Yang, Z. H. & Shi, S. H. & Wang, Y. Y." box="[547,712,1845,1871]" pageId="19" pageNumber="110" pagination="723 - 731" refId="ref16294" refString="Liu, Z. Y., Chen, G. L., Zhu, T. Q., Zeng, Z. C., Lyu, Z. T., Wang, J., Messenger, K., Greenberg, A. J., Guo, Z. X., Yang, Z. H., Shi, S. H. & Wang, Y. Y. (2018) Prevalence of cryptic species in morphologically uniform taxa - Fast speciation and evolutionary radiation in Asian frogs. Molecular Phylogenetics and Evolution, 127, 723 - 731. https: // doi. org / 10.1016 / j. ympev. 2018.06.020" type="journal article" year="2018">
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Liu
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<emphasis id="B9342850FFF8A02A9930D663FD27C445" box="[593,649,1845,1871]" italics="true" pageId="19" pageNumber="110">et al.</emphasis>
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2018
|
||
</bibRefCitation>
|
||
</paragraph>
|
||
<paragraph id="8BFFF442FFF8A02A9BF6D60CFEB2C479" blockId="19.[151,1078,1773,1907]" box="[151,284,1881,1907]" pageId="19" pageNumber="110">
|
||
<figureCitation id="137BE8C7FFF8A02A9BF6D60CFEB2C479" box="[151,284,1881,1907]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="20.[151,250,1975,1999]" captionStartId-1="21.[151,250,2014,2038]" captionTargetBox-0="[170,1417,704,1951]" captionTargetBox-1="[170,1417,741,1991]" captionTargetId-0="figure-266@20.[170,1417,704,1951]" captionTargetId-1="figure-264@21.[170,1417,741,1991]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURE 7. The holotype (SYS a004744) of Boulenophrys fengshunensis sp. nov. in life." captionText-1="FIGURE 8. The paratype (SYS a005221) of Boulenophrys fengshunensis sp. nov. in life." figureDoi-0="http://doi.org/10.5281/zenodo.6036996" figureDoi-1="http://doi.org/10.5281/zenodo.6037003" httpUri-0="https://zenodo.org/record/6036996/files/figure.png" httpUri-1="https://zenodo.org/record/6037003/files/figure.png" pageId="19" pageNumber="110">Figures 7–8</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35AA7C9FFF8A02D9BF6D6F4FC18C3F2" lastPageId="20" lastPageNumber="111" pageId="19" pageNumber="110" type="materials_examined">
|
||
<paragraph id="8BFFF442FFF8A02A9BF6D6F4FA92C4D5" blockId="19.[151,1437,1953,2016]" pageId="19" pageNumber="110">
|
||
<materialsCitation id="3B28FE1FFFF8A02A9BF6D6F4FA92C4D5" ID-GBIF-Occurrence="3464452301" collectingDate="2016-05-13" collectorName="Jian Wang & Zhi-Tong Lyu" country="China" county="Meizhou" elevation="1500" latitude="24.17531" location="Mt. Tongguzhang" longLatPrecision="1" longitude="116.35073" municipality="Fengshun" pageId="19" pageNumber="110" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
|
||
<emphasis id="B9342850FFF8A02A9BF6D6F4FEA5C4B1" bold="true" box="[151,267,1953,1979]" pageId="19" pageNumber="110">
|
||
<typeStatus id="54FB4AE0FFF8A02A9BF6D6F4FEA8C4B1" box="[151,262,1953,1979]" pageId="19" pageNumber="110" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
SYS a004744, adult male, collected by
|
||
<collectorName id="26B59194FFF8A02A9995D6F4FCC0C4B6" box="[756,878,1953,1980]" pageId="19" pageNumber="110">Jian Wang</collectorName>
|
||
and
|
||
<collectorName id="26B59194FFF8A02A98CED6F4FBFEC4B6" box="[943,1104,1953,1980]" pageId="19" pageNumber="110">Zhi-Tong Lyu</collectorName>
|
||
on
|
||
<date id="FFFED282FFF8A02A9FE4D6F4FA8BC4B1" box="[1157,1317,1953,1980]" pageId="19" pageNumber="110" value="2016-05-13">
|
||
<collectingDate id="EFBA2B6AFFF8A02A9FE4D6F4FA8BC4B1" box="[1157,1317,1953,1980]" pageId="19" pageNumber="110" value="2016-05-13">13 May 2016</collectingDate>
|
||
</date>
|
||
from
|
||
<location id="8E9FA299FFF8A02A9E14D6F7FE81C4D5" LSID="urn:lsid:plazi:treatment:03E94554FFF8A02E9BF6D7B8FC12C5DF:8E9FA299FFF8A02A9E14D6F7FE81C4D5" country="China" county="Meizhou" latitude="24.17531" longLatPrecision="1" longitude="116.35073" municipality="Fengshun" name="Mt. Tongguzhang" pageId="19" pageNumber="110" stateProvince="Guangdong">Mt. Tongguzhang</location>
|
||
(
|
||
<geoCoordinate id="EE749285FFF8A02A9A5FD690FE47C4D5" box="[318,489,1989,2015]" degrees="24" direction="north" minutes="10" orientation="latitude" pageId="19" pageNumber="110" precision="1" seconds="31.12" value="24.17531">24°10'31.12"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE749285FFF8A02A9A95D690FD34C4D5" box="[500,666,1989,2015]" degrees="116" direction="east" minutes="21" orientation="longitude" pageId="19" pageNumber="110" precision="1" seconds="2.63" value="116.35073">116°21'2.63"E</geoCoordinate>
|
||
; ca.
|
||
<elevation id="006D1371FFF8A02A99ACD690FCF3C4EA" box="[717,861,1989,2016]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="110" unit="m" value="1500.0">
|
||
<quantity id="4CB859A7FFF8A02A99ACD690FC8CC4EA" box="[717,802,1989,2016]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="110" unit="m" value="1500.0">1500 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
),
|
||
<collectingMunicipality id="6B9B6E38FFF8A02A9815D690FC4AC4D5" box="[884,996,1989,2015]" pageId="19" pageNumber="110">Fengshun</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="629E8CCEFFF8A02A988ED690FBFBC4D5" box="[1007,1109,1989,2015]" pageId="19" pageNumber="110">Meizhou</collectingCounty>
|
||
,
|
||
<collectingRegion id="49843AA0FFF8A02A9F01D690FB48C4D5" box="[1120,1254,1989,2015]" country="China" name="Guangdong" pageId="19" pageNumber="110">Guangdong</collectingRegion>
|
||
,
|
||
<collectingCountry id="F357B4D2FFF8A02A9F90D690FA96C4D5" box="[1265,1336,1989,2015]" name="China" pageId="19" pageNumber="110">China</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BFFF442FFFFA02D9BA6D1C3FC18C3F2" blockId="20.[151,1437,150,681]" pageId="20" pageNumber="111">
|
||
<materialsCitation id="3B28FE1FFFFFA02D9BA6D1C3FC1CC3F2" ID-GBIF-Occurrence="3464452306" collectingDate="2016-05-11" collectingDateMax="2016-08-09" collectingDateMin="2016-05-11" collectionCode="CIB" collectorName="Jian Wang & Zhi-Tong Lyu" pageId="20" pageNumber="111" specimenCode="CIB118528" specimenCount="3" typeStatus="paratype">
|
||
<emphasis id="B9342850FFFFA02D9BA6D1C3FE3AC3BA" bold="true" box="[199,404,150,176]" pageId="20" pageNumber="111">
|
||
<typeStatus id="54FB4AE0FFFFA02D9BA6D1C3FE91C3BA" box="[199,319,150,176]" pageId="20" pageNumber="111" type="paratype">Paratypes</typeStatus>
|
||
(N=7).
|
||
</emphasis>
|
||
Adult males, SYS a004724/
|
||
<specimenCode id="DBE65C39FFFFA02D99BED1C3FCCEC3BB" box="[735,864,150,177]" collectionCode="CIB" country="China" httpUri="http://grbio.org/cool/vxkr-andy" name="Chengdu Institute of Biology" pageId="20" pageNumber="111">CIB118528</specimenCode>
|
||
, SYS a004725–4728, collected by
|
||
<collectorName id="26B59194FFFFA02D9F95D1C3FAC5C3BB" box="[1268,1387,150,177]" pageId="20" pageNumber="111">Jian Wang</collectorName>
|
||
and
|
||
<collectorName id="26B59194FFFFA02D9BF6D1EFFE9CC3DF" box="[151,306,186,213]" pageId="20" pageNumber="111">Zhi-Tong Lyu</collectorName>
|
||
on
|
||
<date id="FFFED282FFFFA02D9A3BD1EFFE42C3DE" box="[346,492,186,213]" pageId="20" pageNumber="111" value="2016-05-11">
|
||
<collectingDate id="EFBA2B6AFFFFA02D9A3BD1EFFE42C3DE" box="[346,492,186,213]" pageId="20" pageNumber="111" value="2016-05-11">11 May 2016</collectingDate>
|
||
</date>
|
||
from the same locality of the
|
||
<typeStatus id="54FB4AE0FFFFA02D9855D1EFFC34C3DE" box="[820,922,186,212]" pageId="20" pageNumber="111" type="holotype">holotype</typeStatus>
|
||
.
|
||
<collectorName id="26B59194FFFFA02D98C0D1EFFC4FC3DE" box="[929,993,186,212]" pageId="20" pageNumber="111">Adult</collectorName>
|
||
females, SYS a005220–5221, collected by
|
||
<collectorName id="26B59194FFFFA02D9BDBD18BFE81C3F3" box="[186,303,222,249]" pageId="20" pageNumber="111">Jian Wang</collectorName>
|
||
on
|
||
<date id="FFFED282FFFFA02D9A39D18BFE52C3F2" box="[344,508,222,248]" pageId="20" pageNumber="111" value="2016-08-09">
|
||
<collectingDate id="EFBA2B6AFFFFA02D9A39D18BFE52C3F2" box="[344,508,222,248]" pageId="20" pageNumber="111" value="2016-08-09">9 August 2016</collectingDate>
|
||
</date>
|
||
from the same locality of the
|
||
<typeStatus id="54FB4AE0FFFFA02D982DD18BFC1CC3F2" box="[844,946,222,248]" pageId="20" pageNumber="111" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35AA7C9FFFFA02D9BA6D057FF5EC24A" pageId="20" pageNumber="111" type="etymology">
|
||
<paragraph id="8BFFF442FFFFA02D9BA6D057FF5EC24A" blockId="20.[151,1437,150,681]" pageId="20" pageNumber="111">
|
||
<emphasis id="B9342850FFFFA02D9BA6D057FEE0C216" bold="true" box="[199,334,258,284]" pageId="20" pageNumber="111">Etymology.</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="4C408FC1FFFFA02D992AD057FD46C216" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[587,744,258,284]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="20" pageNumber="111" phylum="Chordata" rank="species" species="fengshunensis">
|
||
<emphasis id="B9342850FFFFA02D992AD057FD46C216" box="[587,744,258,284]" italics="true" pageId="20" pageNumber="111">fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
” refers to the
|
||
<typeStatus id="54FB4AE0FFFFA02D98EFD056FC10C217" box="[910,958,259,285]" pageId="20" pageNumber="111">type</typeStatus>
|
||
locality of the new species, the Fengshun County.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35AA7C9FFFFA02F9BA6D01FFE9CC057" lastPageId="22" lastPageNumber="113" pageId="20" pageNumber="111" type="diagnosis">
|
||
<paragraph id="8BFFF442FFFFA02D9BA6D01FFC04C1A2" blockId="20.[151,1437,150,681]" pageId="20" pageNumber="111">
|
||
<emphasis id="B9342850FFFFA02D9BA6D01FFEECC26E" bold="true" box="[199,322,330,356]" pageId="20" pageNumber="111">Diagnosis.</emphasis>
|
||
(1) Small body size, SVL
|
||
<quantity id="4CB859A7FFFFA02D991AD01FFC8EC26F" box="[635,800,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.6849999999999996" metricValueMax="3.94" metricValueMin="3.4299999999999997" pageId="20" pageNumber="111" unit="mm" value="36.849999999999994" valueMax="39.4" valueMin="34.3">34.3–39.4 mm</quantity>
|
||
(36.9 ± 1.7, N = 6) in adult males and SVL
|
||
<quantity id="4CB859A7FFFFA02D9E4DD01FFF6DC283" metricMagnitude="-2" metricUnit="m" metricValue="4.37" metricValueMax="4.49" metricValueMin="4.25" pageId="20" pageNumber="111" unit="mm" value="43.7" valueMax="44.9" valueMin="42.5">42.5–44.9 mm</quantity>
|
||
(N = 2) in adult females; (2) snout pointed in dorsal view; (3) tympanum moderate in size TD/ED 0.56–0.67, tubercles bearing spines on tips present on skin of temporal region including the tympanum; (4) vomerine ridge and vomerine teeth present; (5) margin of tongue rounded, not notched distally; (6) hindlimbs short, heels not meeting and tibio-tarsal articulation reaching forward to the region between tympanum and posterior corner of eye, TIB/SVL 0.37–0.43; (7) a subarticular tubercle present at the base of each fingers; (8) toes without lateral fringes and with rudiment of webbing; (9) distinct sparse enlarged tubercles on the surface of limbs, flanks, posterior part of belly and around the cloaca; (10) tips of the tubercles on ventral surface of thighs and around the cloaca bearing tiny spines; (11) single subgular vocal sac in males; (12) nuptial pads with well-developed villiform black nuptial spines on the dorsal surface of the first and second fingers in breeding males.
|
||
</paragraph>
|
||
<caption id="DF3FA4CAFFFFA02D9BF6D6E2FB89C4C5" ID-DOI="http://doi.org/10.5281/zenodo.6036996" ID-Zenodo-Dep="6036996" box="[151,1063,1975,2000]" httpUri="https://zenodo.org/record/6036996/files/figure.png" pageId="20" pageNumber="111" startId="20.[151,250,1975,1999]" targetBox="[170,1417,704,1951]" targetPageId="20">
|
||
<paragraph id="8BFFF442FFFFA02D9BF6D6E2FB89C4C5" blockId="20.[151,1063,1975,2000]" box="[151,1063,1975,2000]" pageId="20" pageNumber="111">
|
||
<emphasis id="B9342850FFFFA02D9BF6D6E2FEBDC4DA" bold="true" box="[151,275,1975,2000]" pageId="20" pageNumber="111">FIGURE 7.</emphasis>
|
||
The holotype (SYS a004744) of
|
||
<taxonomicName id="4C408FC1FFFFA02D9904D6E2FC28C4C5" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[613,902,1975,1999]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="20" pageNumber="111" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFFA02D9904D6E2FC28C4C5" box="[613,902,1975,1999]" italics="true" pageId="20" pageNumber="111">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFFA02D98EDD6EDFC73C4DA" bold="true" box="[908,989,1976,2000]" pageId="20" pageNumber="111">
|
||
<taxonomicNameLabel id="A207952BFFFFA02D98EDD6EDFC73C4DA" box="[908,989,1976,2000]" pageId="20" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFFF442FFFEA02C9BA6D1C3FDB3C3DF" blockId="21.[151,1437,150,718]" pageId="21" pageNumber="112">
|
||
<emphasis id="B9342850FFFEA02C9BA6D1C3FEC3C3BA" bold="true" box="[199,365,150,176]" pageId="21" pageNumber="112">Comparisons.</emphasis>
|
||
Comparative data of
|
||
<taxonomicName id="4C408FC1FFFEA02C993DD1C2FC35C3BB" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[604,923,151,177]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C993DD1C2FC35C3BB" box="[604,923,151,177]" italics="true" pageId="21" pageNumber="112">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C98C0D1C2FC54C3BB" bold="true" box="[929,1018,151,177]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C98C0D1C2FC54C3BB" box="[929,1018,151,177]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and the other recognized members of
|
||
<taxonomicName id="4C408FC1FFFEA02C9BF6D1EEFE9DC3DF" authorityName="Fei, Ye & Jiang" authorityYear="2016" box="[151,307,187,213]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9342850FFFEA02C9BF6D1EEFE9DC3DF" box="[151,307,187,213]" italics="true" pageId="21" pageNumber="112">Boulenophrys</emphasis>
|
||
</taxonomicName>
|
||
are listed in
|
||
<tableCitation id="C6C2C1F9FFFEA02C9AA2D1EEFDB6C3DF" box="[451,536,187,213]" captionStart="TABLE 3" captionStartId="7.[151,239,469,495]" captionTargetPageId="7" captionText="TABLE 3. Diagnostic characters separating all 64 species of Boulenophrys. “/” indicates that data are not available." httpUri="http://table.plazi.org/id/DF3FA4CAFFECA03E9BF6D080FAC5C2E5" pageId="21" pageNumber="112" tableUuid="DF3FA4CAFFECA03E9BF6D080FAC5C2E5">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFFF442FFFEA02C9BA6D18AFD9EC2DB" blockId="21.[151,1437,150,718]" pageId="21" pageNumber="112">
|
||
<taxonomicName id="4C408FC1FFFEA02C9BA6D18AFDA3C3F3" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[199,525,223,249]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C9BA6D18AFDA3C3F3" box="[199,525,223,249]" italics="true" pageId="21" pageNumber="112">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C997AD18AFDD5C3F3" bold="true" box="[539,635,223,249]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C997AD18AFDD5C3F3" box="[539,635,223,249]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="4C408FC1FFFEA02C98ABD18AFB57C3F2" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[970,1273,223,249]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="puningensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C98ABD18AFB57C3F2" box="[970,1273,223,249]" italics="true" pageId="21" pageNumber="112">Boulenophrys puningensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C9E66D18AFAC8C3F3" bold="true" box="[1287,1382,223,249]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C9E66D18AFAC8C3F3" box="[1287,1382,223,249]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C408FC1FFFEA02C9BF6D056FE3EC217" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[151,400,259,285]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="hungtai" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C9BF6D056FE3EC217" box="[151,400,259,285]" italics="true" pageId="21" pageNumber="112">Boulenophrys hungtai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C9AF7D056FE5EC217" bold="true" box="[406,496,259,285]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C9AF7D056FE5EC217" box="[406,496,259,285]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
having a tympanum with tubercles bearing spines on their tips (vs. tympanic region smooth without granules, tubercles or spines in
|
||
<taxonomicName id="4C408FC1FFFEA02C99D4D072FC71C24A" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[693,991,295,321]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="puningensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C99D4D072FC71C24A" box="[693,991,295,321]" italics="true" pageId="21" pageNumber="112">Boulenophrys puningensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C9886D072FBEFC24B" bold="true" box="[999,1089,295,321]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C9886D072FBEFC24B" box="[999,1089,295,321]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C408FC1FFFEA02C9F1BD072FADDC24B" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[1146,1395,295,321]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="hungtai" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C9F1BD072FADDC24B" box="[1146,1395,295,321]" italics="true" pageId="21" pageNumber="112">Boulenophrys hungtai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C9E1AD072FF66C26F" bold="true" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C9E1AD072FF66C26F" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
); the presence of both a vomerine ridge and vomerine teeth (vs. absence of vomerine ridge but with vomerine teeth present in
|
||
<taxonomicName id="4C408FC1FFFEA02C9A2AD03AFDEAC283" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[331,580,367,393]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="hungtai" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C9A2AD03AFDEAC283" box="[331,580,367,393]" italics="true" pageId="21" pageNumber="112">Boulenophrys hungtai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C992AD03AFD0BC283" bold="true" box="[587,677,367,393]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C992AD03AFD0BC283" box="[587,677,367,393]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
); tubercles on skin of the posterior part of belly without spines (vs. tubercles on skin of the posterior part of belly bearing spines on their tips in both
|
||
<taxonomicName id="4C408FC1FFFEA02C9F76D0C6FA90C2A6" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[1047,1342,403,429]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="puningensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C9F76D0C6FA90C2A6" box="[1047,1342,403,429]" italics="true" pageId="21" pageNumber="112">Boulenophrys puningensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C9E25D0C6FA32C2A7" bold="true" box="[1348,1436,403,429]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C9E25D0C6FA32C2A7" box="[1348,1436,403,429]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C408FC1FFFEA02C9BA6D0E2FE11C2DB" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[199,447,439,465]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="hungtai" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C9BA6D0E2FE11C2DB" box="[199,447,439,465]" italics="true" pageId="21" pageNumber="112">Boulenophrys hungtai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C9AA7D0E2FDB1C2DB" bold="true" box="[454,543,439,465]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C9AA7D0E2FDB1C2DB" box="[454,543,439,465]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFFF442FFFEA02C9BA6D08EFE7DC1C4" blockId="21.[151,1437,150,718]" pageId="21" pageNumber="112">
|
||
With a smaller body size, SVL
|
||
<quantity id="4CB859A7FFFEA02C9927D08EFD40C2FF" box="[582,750,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="3.6849999999999996" metricValueMax="3.94" metricValueMin="3.4299999999999997" pageId="21" pageNumber="112" unit="mm" value="36.849999999999994" valueMax="39.4" valueMin="34.3">34.3–39.4 mm</quantity>
|
||
in adult males and SVL
|
||
<quantity id="4CB859A7FFFEA02C9F42D08FFB62C2FF" box="[1059,1228,474,501]" metricMagnitude="-2" metricUnit="m" metricValue="4.37" metricValueMax="4.49" metricValueMin="4.25" pageId="21" pageNumber="112" unit="mm" value="43.7" valueMax="44.9" valueMin="42.5">42.5–44.9 mm</quantity>
|
||
in adult females,
|
||
<taxonomicName id="4C408FC1FFFEA02C9BF6D0AAFE79C113" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[151,471,511,537]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C9BF6D0AAFE79C113" box="[151,471,511,537]" italics="true" pageId="21" pageNumber="112">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C9ABFD0AAFD96C113" bold="true" box="[478,568,511,537]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C9ABFD0AAFD96C113" box="[478,568,511,537]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from the seven congeners whose SVL ≥
|
||
<quantity id="4CB859A7FFFEA02C9F34D0ABFB0AC113" box="[1109,1188,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="112" unit="mm" value="50.0">50 mm</quantity>
|
||
in males or SVL ≥
|
||
<quantity id="4CB859A7FFFEA02C9EE1D0AAFF6DC137" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="112" unit="mm" value="60.0">60 mm</quantity>
|
||
in females, including
|
||
<taxonomicName id="4C408FC1FFFEA02C9AA5D376FDDCC137" baseAuthorityName="Shen" baseAuthorityYear="1994" box="[452,626,547,573]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="caudoprocta">
|
||
<emphasis id="B9342850FFFEA02C9AA5D376FDDCC137" box="[452,626,547,573]" italics="true" pageId="21" pageNumber="112">B. caudoprocta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CB859A7FFFEA02C99E3D376FD47C137" box="[642,745,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="8.129999999999999" pageId="21" pageNumber="112" unit="mm" value="81.3">81.3 mm</quantity>
|
||
in a single male),
|
||
<taxonomicName id="4C408FC1FFFEA02C98A1D376FBD3C137" baseAuthorityName="Fei and Ye" baseAuthorityYear="1983" box="[960,1149,547,573]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="jingdongensis">
|
||
<emphasis id="B9342850FFFEA02C98A1D376FBD3C137" box="[960,1149,547,573]" italics="true" pageId="21" pageNumber="112">B. jingdongensis</emphasis>
|
||
</taxonomicName>
|
||
(53.0–
|
||
<quantity id="4CB859A7FFFEA02C9FAFD377FA9AC137" box="[1230,1332,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" pageId="21" pageNumber="112" unit="mm" value="56.5">56.5 mm</quantity>
|
||
in males and
|
||
<quantity id="4CB859A7FFFEA02C9BABD313FE9FC16B" box="[202,305,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" pageId="21" pageNumber="112" unit="mm" value="63.5">63.5 mm</quantity>
|
||
in a single female),
|
||
<taxonomicName id="4C408FC1FFFEA02C9945D312FD04C16B" baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang and Zhou" baseAuthorityYear="2017" box="[548,682,583,609]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="liboensis">
|
||
<emphasis id="B9342850FFFEA02C9945D312FD04C16B" box="[548,682,583,609]" italics="true" pageId="21" pageNumber="112">B. liboensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CB859A7FFFEA02C99DAD313FCCDC16B" box="[699,867,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="6.409999999999999" metricValueMax="6.77" metricValueMin="6.05" pageId="21" pageNumber="112" unit="mm" value="64.1" valueMax="67.7" valueMin="60.5">60.5–67.7 mm</quantity>
|
||
in males and
|
||
<quantity id="4CB859A7FFFEA02C9F6BD312FB34C16B" box="[1034,1178,583,609]" metricMagnitude="0" metricUnit="m" metricValue="1.66878" metricValueMax="1.7932399999999997" metricValueMin="1.54432" pageId="21" pageNumber="112" unit="in" value="65.69999999999999" valueMax="70.6" valueMin="60.8">60.8–70.6 in</quantity>
|
||
females),
|
||
<taxonomicName id="4C408FC1FFFEA02C9E74D312FA32C16B" baseAuthorityName="Lyu, Wang and Zhao" baseAuthorityYear="2020" box="[1301,1436,583,609]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="mirabilis">
|
||
<emphasis id="B9342850FFFEA02C9E74D312FA32C16B" box="[1301,1436,583,609]" italics="true" pageId="21" pageNumber="112">B. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CB859A7FFFEA02C9BFED33FFEE8C18F" box="[159,326,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="5.859999999999999" metricValueMax="6.14" metricValueMin="5.58" pageId="21" pageNumber="112" unit="mm" value="58.599999999999994" valueMax="61.4" valueMin="55.8">55.8–61.4 mm</quantity>
|
||
in males and
|
||
<quantity id="4CB859A7FFFEA02C9A8DD33FFD3FC18F" box="[492,657,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="7.165" metricValueMax="7.4799999999999995" metricValueMin="6.85" pageId="21" pageNumber="112" unit="mm" value="71.65" valueMax="74.8" valueMin="68.5">68.5–74.8 mm</quantity>
|
||
in females),
|
||
<taxonomicName id="4C408FC1FFFEA02C984AD33EFC60C18E" baseAuthorityName="Liu" baseAuthorityYear="1950" box="[811,974,619,644]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="omeimontis">
|
||
<emphasis id="B9342850FFFEA02C984AD33EFC60C18E" box="[811,974,619,644]" italics="true" pageId="21" pageNumber="112">B. omeimontis</emphasis>
|
||
</taxonomicName>
|
||
(56.0–
|
||
<quantity id="4CB859A7FFFEA02C9F41D33FFB29C18F" box="[1056,1159,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" pageId="21" pageNumber="112" unit="mm" value="59.5">59.5 mm</quantity>
|
||
in males and 68.0–
|
||
<quantity id="4CB859A7FFFEA02C9E0DD33FFF6DC1A3" metricMagnitude="-2" metricUnit="m" metricValue="7.25" pageId="21" pageNumber="112" unit="mm" value="72.5">72.5 mm</quantity>
|
||
in females),
|
||
<taxonomicName id="4C408FC1FFFEA02C9A35D3DAFDAFC1A3" baseAuthorityName="Jiang, Ye and Fei" baseAuthorityYear="2008" box="[340,513,655,681]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="sangzhiensis">
|
||
<emphasis id="B9342850FFFEA02C9A35D3DAFDAFC1A3" box="[340,513,655,681]" italics="true" pageId="21" pageNumber="112">B. sangzhiensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CB859A7FFFEA02C9971D3DBFDDBC1A3" box="[528,629,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="5.47" pageId="21" pageNumber="112" unit="mm" value="54.7">54.7 mm</quantity>
|
||
in a single male), and
|
||
<taxonomicName id="4C408FC1FFFEA02C9813D3DAFB95C1A3" baseAuthorityName="Tian and Sun" baseAuthorityYear="1995" box="[882,1083,655,681]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="shuichengensis">
|
||
<emphasis id="B9342850FFFEA02C9813D3DAFB95C1A3" box="[882,1083,655,681]" italics="true" pageId="21" pageNumber="112">B. shuichengensis</emphasis>
|
||
</taxonomicName>
|
||
(102.0–
|
||
<quantity id="4CB859A7FFFEA02C9FF9D3DAFAA7C1A3" box="[1176,1289,655,681]" metricMagnitude="-1" metricUnit="m" metricValue="1.183" pageId="21" pageNumber="112" unit="mm" value="118.3">118.3 mm</quantity>
|
||
in males and
|
||
<quantity id="4CB859A7FFFEA02C9BF6D3E7FEE9C1C7" box="[151,327,690,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.077" metricValueMax="1.156" metricValueMin="0.998" pageId="21" pageNumber="112" unit="mm" value="107.69999999999999" valueMax="115.6" valueMin="99.8">99.8–115.6 mm</quantity>
|
||
in females).
|
||
</paragraph>
|
||
<caption id="DF3FA4CAFFFEA02C9BF6D68BFB8BC4FC" ID-DOI="http://doi.org/10.5281/zenodo.6037003" ID-Zenodo-Dep="6037003" box="[151,1061,2014,2039]" httpUri="https://zenodo.org/record/6037003/files/figure.png" pageId="21" pageNumber="112" startId="21.[151,250,2014,2038]" targetBox="[170,1417,741,1991]" targetPageId="21">
|
||
<paragraph id="8BFFF442FFFEA02C9BF6D68BFB8BC4FC" blockId="21.[151,1061,2014,2039]" box="[151,1061,2014,2039]" pageId="21" pageNumber="112">
|
||
<emphasis id="B9342850FFFEA02C9BF6D68BFEBDC4FC" bold="true" box="[151,275,2014,2038]" pageId="21" pageNumber="112">FIGURE 8.</emphasis>
|
||
The paratype (SYS a005221) of
|
||
<taxonomicName id="4C408FC1FFFEA02C9902D68BFC2AC4FC" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[611,900,2014,2038]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="21" pageNumber="112" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFEA02C9902D68BFC2AC4FC" box="[611,900,2014,2038]" italics="true" pageId="21" pageNumber="112">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFEA02C98EAD68BFC75C4FC" bold="true" box="[907,987,2014,2038]" pageId="21" pageNumber="112">
|
||
<taxonomicNameLabel id="A207952BFFFEA02C98EAD68BFC75C4FC" box="[907,987,2014,2038]" pageId="21" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFFF442FFFDA02F9BA6D1C2FB8CC2A7" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="113">
|
||
In having relatively shorter shanks with heels that do not meet when the flexed hind limbs are held at right angles to the body axis,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A35D1EEFD3DC3DF" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[340,659,187,213]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFDA02F9A35D1EEFD3DC3DF" box="[340,659,187,213]" italics="true" pageId="22" pageNumber="113">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFDA02F99F8D1EEFD5CC3DF" bold="true" box="[665,754,187,213]" pageId="22" pageNumber="113">
|
||
<taxonomicNameLabel id="A207952BFFFDA02F99F8D1EEFD5CC3DF" box="[665,754,187,213]" pageId="22" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily distinguished from the following 35 congeners, i.e.
|
||
<taxonomicName id="4C408FC1FFFDA02F9BA1D18AFE8CC3F3" baseAuthorityName="Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy and Che" baseAuthorityYear="2019" box="[192,290,223,249]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="angka">
|
||
<emphasis id="B9342850FFFDA02F9BA1D18AFE8CC3F3" box="[192,290,223,249]" italics="true" pageId="22" pageNumber="113">B. angka</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A4ED18AFE7AC3F3" baseAuthorityName="Li, Lu, Liu and Wang" baseAuthorityYear="2020" box="[303,468,223,249]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="anlongensis">
|
||
<emphasis id="B9342850FFFDA02F9A4ED18AFE7AC3F3" box="[303,468,223,249]" italics="true" pageId="22" pageNumber="113">B. anlongensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A80D18AFD07C3F3" baseAuthorityName="Wu, Li, Liu, Wang and Wu" baseAuthorityYear="2020" box="[481,681,223,249]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="baishanzuensis">
|
||
<emphasis id="B9342850FFFDA02F9A80D18AFD07C3F3" box="[481,681,223,249]" italics="true" pageId="22" pageNumber="113">B. baishanzuensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F99D7D18AFCC6C3F3" baseAuthorityName="Ye, Fei and Xie" baseAuthorityYear="2007" box="[694,872,223,249]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="baolongensis">
|
||
<emphasis id="B9342850FFFDA02F99D7D18AFCC6C3F3" box="[694,872,223,249]" italics="true" pageId="22" pageNumber="113">B. baolongensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9814D18AFB9DC3F3" baseAuthorityName="Ye and Fei" baseAuthorityYear="1995" box="[885,1075,223,249]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="binchuanensis">
|
||
<emphasis id="B9342850FFFDA02F9814D18AFB9DC3F3" box="[885,1075,223,249]" italics="true" pageId="22" pageNumber="113">B. binchuanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9F20D18AFB48C3F3" baseAuthorityName="Jiang, Fei and Ye" baseAuthorityYear="2009" box="[1089,1254,223,249]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="binlingensis">
|
||
<emphasis id="B9342850FFFDA02F9F20D18AFB48C3F3" box="[1089,1254,223,249]" italics="true" pageId="22" pageNumber="113">B. binlingensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9F95D18AFAD9C3F3" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[1268,1399,223,249]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="boettgeri">
|
||
<emphasis id="B9342850FFFDA02F9F95D18AFAD9C3F3" box="[1268,1399,223,249]" italics="true" pageId="22" pageNumber="113">B. boettgeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9EE5D18AFE91C216" baseAuthorityName="Luo, Wang, Wang, Lu, Wang, Deng and Zhou" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="congjiangensis">
|
||
<emphasis id="B9342850FFFDA02F9EE5D18AFE91C216" italics="true" pageId="22" pageNumber="113">B. congjiangensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A2DD056FE08C217" baseAuthorityName="Wang and Liu" baseAuthorityYear="2014" box="[332,422,259,285]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="cheni">
|
||
<emphasis id="B9342850FFFDA02F9A2DD056FE08C217" box="[332,422,259,285]" italics="true" pageId="22" pageNumber="113">B. cheni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9AD3D056FDF7C217" baseAuthorityName="Xu, Li, Liu, Wei and Wang" baseAuthorityYear="2020" box="[434,601,259,285]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="chishuiensis">
|
||
<emphasis id="B9342850FFFDA02F9AD3D056FDF7C217" box="[434,601,259,285]" italics="true" pageId="22" pageNumber="113">B. chishuiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9904D056FCA8C217" baseAuthorityName="Lyu, Wang and Wang" baseAuthorityYear="2021" box="[613,774,259,285]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="daiyunensis">
|
||
<emphasis id="B9342850FFFDA02F9904D056FCA8C217" box="[613,774,259,285]" italics="true" pageId="22" pageNumber="113">B. daiyunensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9872D056FCDCC216" baseAuthorityName="Liu, Li, Wei, Xu, Cheng, Wang and Wu" baseAuthorityYear="2020" box="[787,882,259,284]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="jiangi">
|
||
<emphasis id="B9342850FFFDA02F9872D056FCDCC216" box="[787,882,259,284]" italics="true" pageId="22" pageNumber="113">B. jiangi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F981FD056FB96C216" baseAuthorityName="Wang" baseAuthorityYear="2012" box="[894,1080,259,284]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="jinggangensis">
|
||
<emphasis id="B9342850FFFDA02F981FD056FB96C216" box="[894,1080,259,284]" italics="true" pageId="22" pageNumber="113">B. jinggangensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9F25D056FB4CC217" box="[1092,1250,259,285]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="jiulianensis">
|
||
<emphasis id="B9342850FFFDA02F9F25D056FB4CC217" box="[1092,1250,259,285]" italics="true" pageId="22" pageNumber="113">B. jiulianensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9F8ED056FA3BC217" baseAuthorityName="Li, Xu, Liu, Jiang, Wei and Wang" baseAuthorityYear="2018" box="[1263,1429,259,285]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="leishanensis">
|
||
<emphasis id="B9342850FFFDA02F9F8ED056FA3BC217" box="[1263,1429,259,285]" italics="true" pageId="22" pageNumber="113">B. leishanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9BF6D072FF71C24B" baseAuthorityName="Wang and Yang" baseAuthorityYear="2014" box="[151,223,295,321]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="lini">
|
||
<emphasis id="B9342850FFFDA02F9BF6D072FF71C24B" box="[151,223,295,321]" italics="true" pageId="22" pageNumber="113">B. lini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9B91D072FEF8C24A" baseAuthorityName="Stejneger" baseAuthorityYear="1926" box="[240,342,295,320]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="minor">
|
||
<emphasis id="B9342850FFFDA02F9B91D072FEF8C24A" box="[240,342,295,320]" italics="true" pageId="22" pageNumber="113">B. minor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A06D072FD8AC24B" baseAuthorityName="Wang, Lyu and Wang" baseAuthorityYear="2019" box="[359,548,295,321]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="mufumontana">
|
||
<emphasis id="B9342850FFFDA02F9A06D072FD8AC24B" box="[359,548,295,321]" italics="true" pageId="22" pageNumber="113">B. mufumontana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9954D072FD48C24B" baseAuthorityName="Lyu, Wang, Liu and Wang" baseAuthorityYear="2019" box="[565,742,295,321]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="nanlingensis">
|
||
<emphasis id="B9342850FFFDA02F9954D072FD48C24B" box="[565,742,295,321]" italics="true" pageId="22" pageNumber="113">B. nanlingensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9996D072FC46C24B" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[759,1000,295,321]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="palpebralespinosa">
|
||
<emphasis id="B9342850FFFDA02F9996D072FC46C24B" box="[759,1000,295,321]" italics="true" pageId="22" pageNumber="113">B. palpebralespinosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9898D072FB06C24B" box="[1017,1192,295,321]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="qianbeiensis">
|
||
<emphasis id="B9342850FFFDA02F9898D072FB06C24B" box="[1017,1192,295,321]" italics="true" pageId="22" pageNumber="113">B. qianbeiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9FD8D072FADDC24A" baseAuthorityName="Lyu and Wang" baseAuthorityYear="2021" box="[1209,1395,295,320]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="sanmingensis">
|
||
<emphasis id="B9342850FFFDA02F9FD8D072FADDC24A" box="[1209,1395,295,320]" italics="true" pageId="22" pageNumber="113">B. sanmingensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9EE5D072FE8EC26F" baseAuthorityName="Lyu, Liu and Wang" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="shimentaina">
|
||
<emphasis id="B9342850FFFDA02F9EE5D072FE8EC26F" italics="true" pageId="22" pageNumber="113">B. shimentaina</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A56D01EFDBCC26F" baseAuthorityName="Wang, Deng, Liu, Wu and Liu" baseAuthorityYear="2019" box="[311,530,331,357]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="shunhuangensis">
|
||
<emphasis id="B9342850FFFDA02F9A56D01EFDBCC26F" box="[311,530,331,357]" italics="true" pageId="22" pageNumber="113">B. shunhuangensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9949D01EFD0CC26E" baseAuthorityName="Liu and Hu" baseAuthorityYear="1973" box="[552,674,331,356]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="spinata">
|
||
<emphasis id="B9342850FFFDA02F9949D01EFD0CC26E" box="[552,674,331,356]" italics="true" pageId="22" pageNumber="113">B. spinata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F99DBD01EFCC6C26F" baseAuthorityName="Wang and Lyu" baseAuthorityYear="2021" box="[698,872,331,357]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="tongboensis">
|
||
<emphasis id="B9342850FFFDA02F99DBD01EFCC6C26F" box="[698,872,331,357]" italics="true" pageId="22" pageNumber="113">B. tongboensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F981FD01EFBC9C26F" baseAuthorityName="Shen, Mo and Li" baseAuthorityYear="2010" box="[894,1127,331,357]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="tuberogranulatus">
|
||
<emphasis id="B9342850FFFDA02F981FD01EFBC9C26F" box="[894,1127,331,357]" italics="true" pageId="22" pageNumber="113">B. tuberogranulatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9F1FD01EFAC3C26F" baseAuthorityName="Ye and Fei" baseAuthorityYear="1995" box="[1150,1389,331,357]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="wuliangshanensis">
|
||
<emphasis id="B9342850FFFDA02F9F1FD01EFAC3C26F" box="[1150,1389,331,357]" italics="true" pageId="22" pageNumber="113">B. wuliangshanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9EE5D01EFE8BC283" baseAuthorityName="Ye and Fei" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="wushanensis">
|
||
<emphasis id="B9342850FFFDA02F9EE5D01EFE8BC283" italics="true" pageId="22" pageNumber="113">B. wushanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A51D03AFE42C282" baseAuthorityName="Lyu, Zeng and Wang" baseAuthorityYear="2020" box="[304,492,367,392]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="xiangnanensis">
|
||
<emphasis id="B9342850FFFDA02F9A51D03AFE42C282" box="[304,492,367,392]" italics="true" pageId="22" pageNumber="113">B. xiangnanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A96D03AFD3CC282" baseAuthorityName="Wang, Wu, Peng, Shi, Lu and Wu" baseAuthorityYear="2020" box="[503,658,367,392]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="xianjuensis">
|
||
<emphasis id="B9342850FFFDA02F9A96D03AFD3CC282" box="[503,658,367,392]" italics="true" pageId="22" pageNumber="113">B. xianjuensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F99FCD03AFCE0C283" authorityName="Qi, Mo, Lyu, Wang and Wang" authorityYear="2021" box="[669,846,367,393]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="yaoshanensis">
|
||
<emphasis id="B9342850FFFDA02F99FCD03AFCE0C283" box="[669,846,367,393]" italics="true" pageId="22" pageNumber="113">B. yaoshanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9838D03AFBB5C282" baseAuthorityName="Lyu, Zeng and Wang" baseAuthorityYear="2020" box="[857,1051,367,392]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="yangmingensis">
|
||
<emphasis id="B9342850FFFDA02F9838D03AFBB5C282" box="[857,1051,367,392]" italics="true" pageId="22" pageNumber="113">B.yangmingensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9F46D03AFB68C283" authorityName="Qi, Lyu, Wang and Wang" authorityYear="2021" box="[1063,1222,367,393]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="yingdeensis">
|
||
<emphasis id="B9342850FFFDA02F9F46D03AFB68C283" box="[1063,1222,367,393]" italics="true" pageId="22" pageNumber="113">B. yingdeensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C408FC1FFFDA02F9F97D03AFA38C283" authorityName="Qi, Wang, Lyu and Wang" authorityYear="2021" box="[1270,1430,367,393]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="yunkaiensis">
|
||
<emphasis id="B9342850FFFDA02F9F97D03AFA38C283" box="[1270,1430,367,393]" italics="true" pageId="22" pageNumber="113">B. yunkaiensis</emphasis>
|
||
</taxonomicName>
|
||
, all of which have relatively longer shanks with the heels meeting or overlapping.
|
||
</paragraph>
|
||
<paragraph id="8BFFF442FFFDA02F9BA6D0E2FE9CC057" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="113">
|
||
By the presence of vomerine teeth,
|
||
<taxonomicName id="4C408FC1FFFDA02F9933D0E2FC3FC2DB" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[594,913,439,465]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFDA02F9933D0E2FC3FC2DB" box="[594,913,439,465]" italics="true" pageId="22" pageNumber="113">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFDA02F98F9D0E2FC5FC2DB" bold="true" box="[920,1009,439,465]" pageId="22" pageNumber="113">
|
||
<taxonomicNameLabel id="A207952BFFFDA02F98F9D0E2FC5FC2DB" box="[920,1009,439,465]" pageId="22" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C408FC1FFFDA02F9FE2D0E2FB4EC2DA" baseAuthorityName="Wang, Li and Jin" baseAuthorityYear="2014" box="[1155,1248,439,464]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="acuta">
|
||
<emphasis id="B9342850FFFDA02F9FE2D0E2FB4EC2DA" box="[1155,1248,439,464]" italics="true" pageId="22" pageNumber="113">B. acuta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9F8FD0E2FA3BC2DB" baseAuthorityName="Inger and Romer" baseAuthorityYear="1961" box="[1262,1429,439,465]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="brachykolos">
|
||
<emphasis id="B9342850FFFDA02F9F8FD0E2FA3BC2DB" box="[1262,1429,439,465]" italics="true" pageId="22" pageNumber="113">B. brachykolos</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9BF6D08EFEE2C2FF" baseAuthorityName="Nguyen, Pham, Nguyen, Luong and Ziegler" baseAuthorityYear="2020" box="[151,332,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="caobangensis">
|
||
<emphasis id="B9342850FFFDA02F9BF6D08EFEE2C2FF" box="[151,332,475,501]" italics="true" pageId="22" pageNumber="113">B. caobangensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A38D08EFE1EC2FF" baseAuthorityName="Lyu, Zeng, Wang and Wang" baseAuthorityYear="2021" box="[345,432,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="daoji">
|
||
<emphasis id="B9342850FFFDA02F9A38D08EFE1EC2FF" box="[345,432,475,501]" italics="true" pageId="22" pageNumber="113">B. daoji</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9ADDD08EFD22C2FF" baseAuthorityName="Fei and Ye" baseAuthorityYear="2005" box="[444,652,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="huangshanensis">
|
||
<emphasis id="B9342850FFFDA02F9ADDD08EFD22C2FF" box="[444,652,475,501]" italics="true" pageId="22" pageNumber="113">B. huangshanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F99F9D08EFC94C2FF" baseAuthorityName="Pope" baseAuthorityYear="1929" box="[664,826,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="kuatunensis">
|
||
<emphasis id="B9342850FFFDA02F99F9D08EFC94C2FF" box="[664,826,475,501]" italics="true" pageId="22" pageNumber="113">B. kuatunensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9827D08EFC77C2FF" baseAuthorityName="Wang, Liu and Jiang" baseAuthorityYear="2017" box="[838,985,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="lishuiensis">
|
||
<emphasis id="B9342850FFFDA02F9827D08EFC77C2FF" box="[838,985,475,501]" italics="true" pageId="22" pageNumber="113">B. lishuiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9887D08EFBD1C2FF" baseAuthorityName="Shi, Li, Zhu, Jiang, Jiang and Wang" baseAuthorityYear="2021" box="[998,1151,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="lushuiensis">
|
||
<emphasis id="B9342850FFFDA02F9887D08EFBD1C2FF" box="[998,1151,475,501]" italics="true" pageId="22" pageNumber="113">B. lushuiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9FEDD08EFB44C2FF" baseAuthorityName="Wang, Li and Zhao" baseAuthorityYear="2014" box="[1164,1258,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="obesa">
|
||
<emphasis id="B9342850FFFDA02F9FEDD08EFB44C2FF" box="[1164,1258,475,501]" italics="true" pageId="22" pageNumber="113">B. obesa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9F97D08EFA38C2FF" baseAuthorityName="Messenger and Dahn" baseAuthorityYear="2019" box="[1270,1430,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="ombrophila">
|
||
<emphasis id="B9342850FFFDA02F9F97D08EFA38C2FF" box="[1270,1430,475,501]" italics="true" pageId="22" pageNumber="113">B. ombrophila</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C408FC1FFFDA02F9BAAD0AAFED1C112" baseAuthorityName="Wang, Lyu and Wang" baseAuthorityYear="2019" box="[203,383,511,536]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="wugongensis">
|
||
<emphasis id="B9342850FFFDA02F9BAAD0AAFED1C112" box="[203,383,511,536]" italics="true" pageId="22" pageNumber="113">B. wugongensis</emphasis>
|
||
</taxonomicName>
|
||
, all of which lack vomerine teeth. By having a rounded margin of the tongue that is not notched distally,
|
||
<taxonomicName id="4C408FC1FFFDA02F9A3CD376FD0EC137" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[349,672,547,573]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFDA02F9A3CD376FD0EC137" box="[349,672,547,573]" italics="true" pageId="22" pageNumber="113">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFDA02F99CBD376FCA9C137" bold="true" box="[682,775,547,573]" pageId="22" pageNumber="113">
|
||
<taxonomicNameLabel id="A207952BFFFDA02F99CBD376FCA9C137" box="[682,775,547,573]" pageId="22" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C408FC1FFFDA02F98C7D376FBDFC137" box="[934,1137,547,573]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="hoanglienensis">
|
||
<emphasis id="B9342850FFFDA02F98C7D376FBDFC137" box="[934,1137,547,573]" italics="true" pageId="22" pageNumber="113">B. hoanglienensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9FE3D376FAA9C137" baseAuthorityName="Wang, Liu, Lyu, Zeng and Wang" baseAuthorityYear="2017" box="[1154,1287,547,573]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="insularis">
|
||
<emphasis id="B9342850FFFDA02F9FE3D376FAA9C137" box="[1154,1287,547,573]" italics="true" pageId="22" pageNumber="113">B. insularis</emphasis>
|
||
</taxonomicName>
|
||
, which have a notched tongue. By the absence of lateral fringes on toes,
|
||
<taxonomicName id="4C408FC1FFFDA02F9827D312FB29C16B" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[838,1159,583,609]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFDA02F9827D312FB29C16B" box="[838,1159,583,609]" italics="true" pageId="22" pageNumber="113">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFDA02F9FF1D312FB45C16B" bold="true" box="[1168,1259,583,609]" pageId="22" pageNumber="113">
|
||
<taxonomicNameLabel id="A207952BFFFDA02F9FF1D312FB45C16B" box="[1168,1259,583,609]" pageId="22" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C408FC1FFFDA02F9EE5D312FEA4C18F" baseAuthorityName="Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong and Rowley" baseAuthorityYear="2017" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="rubrimera">
|
||
<emphasis id="B9342850FFFDA02F9EE5D312FEA4C18F" italics="true" pageId="22" pageNumber="113">B. rubrimera</emphasis>
|
||
</taxonomicName>
|
||
, which has narrow lateral fringes on toes. By the presence of rudimentary webbing on toes,
|
||
<taxonomicName id="4C408FC1FFFDA02F9E60D33EFE9AC1A3" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFDA02F9E60D33EFE9AC1A3" italics="true" pageId="22" pageNumber="113">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFDA02F9A5FD3DAFE37C1A3" bold="true" box="[318,409,655,681]" pageId="22" pageNumber="113">
|
||
<taxonomicNameLabel id="A207952BFFFDA02F9A5FD3DAFE37C1A3" box="[318,409,655,681]" pageId="22" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C408FC1FFFDA02F9955D3DAFD4AC1A3" baseAuthorityName="Rao and Yang" baseAuthorityYear="1997" box="[564,740,655,681]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="daweimontis">
|
||
<emphasis id="B9342850FFFDA02F9955D3DAFD4AC1A3" box="[564,740,655,681]" italics="true" pageId="22" pageNumber="113">B. daweimontis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F9992D3DAFC03C1A3" box="[755,941,655,681]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="fansipanensis">
|
||
<emphasis id="B9342850FFFDA02F9992D3DAFC03C1A3" box="[755,941,655,681]" italics="true" pageId="22" pageNumber="113">B. fansipanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C408FC1FFFDA02F98DFD3DAFB85C1A3" baseAuthorityName="Tapley, Cutajar, Nguyen, Portway, Mahony, Nguyen, Harding, Luong and Rowley" baseAuthorityYear="2021" box="[958,1067,655,681]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="frigida">
|
||
<emphasis id="B9342850FFFDA02F98DFD3DAFB85C1A3" box="[958,1067,655,681]" italics="true" pageId="22" pageNumber="113">B. frigida</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C408FC1FFFDA02F9F0CD3DAFAACC1A3" baseAuthorityName="Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong and Rowley" baseAuthorityYear="2017" box="[1133,1282,655,681]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="rubrimera">
|
||
<emphasis id="B9342850FFFDA02F9F0CD3DAFAACC1A3" box="[1133,1282,655,681]" italics="true" pageId="22" pageNumber="113">B. rubrimera</emphasis>
|
||
</taxonomicName>
|
||
, all of which lack webbing on toes.
|
||
<taxonomicName id="4C408FC1FFFDA02F9AFCD3E6FD7AC1C7" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[413,724,691,717]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="fengshunensi" status="sp. nov.">
|
||
<emphasis id="B9342850FFFDA02F9AFCD3E6FD7AC1C7" box="[413,724,691,717]" italics="true" pageId="22" pageNumber="113">Boulenophrys fengshunensi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFDA02F99BFD3E6FC94C1C7" bold="true" box="[734,826,691,717]" pageId="22" pageNumber="113">
|
||
<taxonomicNameLabel id="A207952BFFFDA02F99BFD3E6FC94C1C7" box="[734,826,691,717]" pageId="22" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
further differs from the remaining
|
||
<taxonomicName id="4C408FC1FFFDA02F9FB2D3E6FA32C1C7" baseAuthorityName="Wang and Wang" baseAuthorityYear="2019" box="[1235,1436,691,717]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="dongguanensis">
|
||
<emphasis id="B9342850FFFDA02F9FB2D3E6FA32C1C7" box="[1235,1436,691,717]" italics="true" pageId="22" pageNumber="113">B. dongguanensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C408FC1FFFDA02F9BA9D382FEDCC1FB" baseAuthorityName="Wang, Zeng and Wang" baseAuthorityYear="2019" box="[200,370,727,753]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="nankunensis">
|
||
<emphasis id="B9342850FFFDA02F9BA9D382FEDCC1FB" box="[200,370,727,753]" italics="true" pageId="22" pageNumber="113">B. nankunensis</emphasis>
|
||
</taxonomicName>
|
||
by having raised tubercles on the surface of posterior abdomen (vs. absence of such tubercles in both
|
||
<emphasis id="B9342850FFFDA02F9B91D3AEFDC3C01F" box="[240,621,763,789]" italics="true" pageId="22" pageNumber="113">
|
||
<taxonomicName id="4C408FC1FFFDA02F9B91D3AEFE17C01F" baseAuthorityName="Wang and Wang" baseAuthorityYear="2019" box="[240,441,763,789]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="dongguanensis">B. dongguanensis</taxonomicName>
|
||
<taxonomicName id="4C408FC1FFFDA02F9AA3D3AEFDC3C01F" baseAuthorityName="Wang, Zeng and Wang" baseAuthorityYear="2019" box="[450,621,763,789]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="nankunensis">B. nankunensis</taxonomicName>
|
||
</emphasis>
|
||
); the presence of tubercles bearing spines on their tips on the temporal region including the tympanum (vs. temporal region lacking tubercles or spines in both
|
||
<taxonomicName id="4C408FC1FFFDA02F9FE3D24AFAE5C033" baseAuthorityName="Wang and Wang" baseAuthorityYear="2019" box="[1154,1355,799,825]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="dongguanensis">
|
||
<emphasis id="B9342850FFFDA02F9FE3D24AFAE5C033" box="[1154,1355,799,825]" italics="true" pageId="22" pageNumber="113">B. dongguanensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C408FC1FFFDA02F9EE5D24AFE8FC057" baseAuthorityName="Wang, Zeng and Wang" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="22" pageNumber="113" phylum="Chordata" rank="species" species="nankunensis">
|
||
<emphasis id="B9342850FFFDA02F9EE5D24AFE8FC057" italics="true" pageId="22" pageNumber="113">B. nankunensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35AA7C9FFFDA02E9BA6D233FD8FC1C7" lastPageId="23" lastPageNumber="114" pageId="22" pageNumber="113" type="description">
|
||
<paragraph id="8BFFF442FFFDA02F9BA6D233FEE0C777" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="113">
|
||
<emphasis id="B9342850FFFDA02F9BA6D233FE47C08B" bold="true" box="[199,489,870,897]" pageId="22" pageNumber="113">
|
||
Description of
|
||
<typeStatus id="54FB4AE0FFFDA02F9A1DD232FE4BC08B" box="[380,485,871,897]" pageId="22" pageNumber="113" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male. Small body size, SVL
|
||
<quantity id="4CB859A7FFFDA02F98E9D232FC5FC08B" box="[904,1009,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="22" pageNumber="113" unit="mm" value="37.4">37.4 mm</quantity>
|
||
; head width slightly larger than head length, HWD/HDL 1.17; snout pointed in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eyes moderate in size, ED 0.37 of HDL, pupil vertical, near diamond-shaped; nostril oblique-ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance slightly larger than interorbital distance; tympanum moderate in size with an obvious margin, TD/ED 0.65; large ovoid choanae at base of maxilla; vomerine ridge and vomerine teeth present, maxillary teeth present; margin of tongue rounded, not notched distally; presence of single subgular vocal sac, and pair of slit-like openings at posterior of jaw.
|
||
</paragraph>
|
||
<paragraph id="8BFFF442FFFDA02F9BA6D5D2FD93C6CB" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="113">Forearm (i.e., radioulna) length 0.21 of SVL and hand 0.25 of SVL; hand without webbing, fingers without lateral fringes, relative finger length I = II <IV <III; tips of fingers slightly dilated, round; subarticular tubercles present and distinct; inner metacarpal tubercle larger and outer metacarpal tubercle smaller; a single nuptial pad bearing well-developed nuptial spines present on dorsal surface of first and second fingers, respectively. Hindlimbs short, tibio-tarsal articulation reaching forward to tympanum when hindlimb is stretched along side of body; heels not meeting when flexed hindlimbs are held at right angles to body axis; crus (i.e., tibiofibular) length 0.42 of SVL and foot length 0.59 of SVL; relative toe length I <II <V <III <IV; tips of toes slightly dilated, round; toes without lateral fringes and with rudimentary webbing; subarticular tubercle distinct; inner metatarsal tubercle long ovoid and lacking outer metatarsal tubercle.</paragraph>
|
||
<paragraph id="8BFFF442FFFDA02F9BA6D49EFD87C423" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="113">Dorsal skin rough, granular, with sparse tubercles; skin of dorsal body and limbs bearing tiny spines; sparse large conical tubercles on flanks; single horn-like prominent tubercle on edge of upper eyelid; obvious supratympanic fold curving posteroventrally from posterior corner of eye to level above insertion of arm; dense tubercles on skin of upper lip, upper eyelid, mandibular articulation, loreal, temporal region including tympanum and surface around cloaca; tip of tubercles on skin of temporal region excluding tympanum bearing tiny spine; single discontinuous “X” shaped ridge and few elongated tubercles arranged in longitudinal rows on two sides at mid-dorsum; sparse tubercles on dorsal shank and thigh; ventral skin of throat, chest and anterior part of belly smooth; ventral skin of thighs and posterior part of belly bearing dense tubercles; surface around cloaca with dense tubercles bearing tiny spines; small pectoral gland closer to axilla; single femoral gland positioned on posterior surface of thigh at midpoint between knee and cloaca..</paragraph>
|
||
<paragraph id="8BFFF442FFFDA02E9BA6D667FB9CC3F3" blockId="22.[151,1437,151,2013]" lastBlockId="23.[151,1437,151,717]" lastPageId="23" lastPageNumber="114" pageId="22" pageNumber="113">
|
||
<emphasis id="B9342850FFFDA02F9BA6D667FD9FC446" bold="true" box="[199,561,1842,1869]" pageId="22" pageNumber="113">
|
||
Coloration of
|
||
<typeStatus id="54FB4AE0FFFDA02F9A12D666FE74C447" box="[371,474,1843,1869]" pageId="22" pageNumber="113" type="holotype">holotype</typeStatus>
|
||
in life.
|
||
</emphasis>
|
||
Dorsal surface of body dark brown, with incomplete dark brown triangular marking between eyes. Two wide oblique black bands present on forearm. Dorsal surface of fingers and hindlimbs with dark grey transverse bands. Presence of vertical dark brown band below eye. Tip of tubercles on edge of upper eyelid greyish white. Supratympanic fold orangey brown. Ventral surface dark brown, black longitudinal band on surface of throat, posterior part of belly with irregular greyish white patches. Tubercles on ventral surface of posterior part of belly and thighs greyish white; spines on tips of tubercles on temporal region, ventral surface of thighs and around cloaca dark grey. Digits, inner and outer metacarpal tubercles, and inner metatarsal tubercle dark grey. Pectoral glands beige and femoral glands greyish white. Iris orangish brown.
|
||
</paragraph>
|
||
<paragraph id="8BFFF442FFFCA02E9BA6D057FC8DC283" blockId="23.[151,1437,151,717]" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BA6D057FD21C216" bold="true" box="[199,655,258,285]" pageId="23" pageNumber="114">
|
||
Coloration of
|
||
<typeStatus id="54FB4AE0FFFCA02E9A0AD056FE7CC217" box="[363,466,259,285]" pageId="23" pageNumber="114" type="holotype">holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
Dorsal surface of body dark brown. Triangular marking between eyes, vertical dark brown band below eye and transverse bands on dorsal forearms and hind limbs become indistinct. Supratympanic fold greyish white. All bands and patterns on ventral surface no longer apparent. Irregular greyish white patches on posterior part of belly become dark grey.
|
||
</paragraph>
|
||
<paragraph id="8BFFF442FFFCA02E9BA6D0C7FD8FC1C7" blockId="23.[151,1437,151,717]" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BA6D0C7FE91C2A6" bold="true" box="[199,319,402,428]" pageId="23" pageNumber="114">Variation.</emphasis>
|
||
Mensural data of the type series are listed in
|
||
<tableCitation id="C6C2C1F9FFFCA02E984AD0C6FCD1C2A7" box="[811,895,403,429]" captionStart="TABLE 6" captionStartId="23.[151,244,762,788]" captionTargetPageId="23" captionText="TABLE 6. Measurements (in mm) of the type series of Boulenophrys fengshunensis sp. nov., * indicates the holotype." httpUri="http://table.plazi.org/id/DF3FA4CAFFFCA02E9BF6D3AFFF4BC033" pageId="23" pageNumber="114" tableUuid="DF3FA4CAFFFCA02E9BF6D3AFFF4BC033">Table 6</tableCitation>
|
||
. Most of the
|
||
<typeStatus id="54FB4AE0FFFCA02E9F6DD0C6FBD9C2A7" box="[1036,1143,403,429]" pageId="23" pageNumber="114" type="paratype">paratypes</typeStatus>
|
||
are similar to the
|
||
<typeStatus id="54FB4AE0FFFCA02E9E5BD0C6FA32C2A7" box="[1338,1436,403,429]" pageId="23" pageNumber="114" type="holotype">holotype</typeStatus>
|
||
in morphology and color pattern, except for the following: dorsal surface and ventral surface dark brown, posterior part of belly with irregular greyish white patches, ventral surface of thighs dark grey, triangular marking between eyes is incomplete in the
|
||
<typeStatus id="54FB4AE0FFFCA02E9AD5D0AAFDB8C113" box="[436,534,511,537]" pageId="23" pageNumber="114" type="holotype">holotype</typeStatus>
|
||
(vs. dorsal surface beige and ventral surface light orange, posterior part of belly and ventral surface of thighs greyish white, triangular marking between eyes is complete in the female
|
||
<typeStatus id="54FB4AE0FFFCA02E9E62D376FACDC137" box="[1283,1379,547,573]" pageId="23" pageNumber="114" type="paratype">paratype</typeStatus>
|
||
SYS a005221 (
|
||
<figureCitation id="137BE8C7FFFCA02E9A66D312FEE4C16B" box="[263,330,583,609]" captionStart="FIGURE 8" captionStartId="21.[151,250,2014,2038]" captionTargetBox="[170,1417,741,1991]" captionTargetId="figure-264@21.[170,1417,741,1991]" captionTargetPageId="21" captionText="FIGURE 8. The paratype (SYS a005221) of Boulenophrys fengshunensis sp. nov. in life." figureDoi="http://doi.org/10.5281/zenodo.6037003" httpUri="https://zenodo.org/record/6037003/files/figure.png" pageId="23" pageNumber="114">Fig. 8</figureCitation>
|
||
); tibio-tarsal articulation reaching forward to middle of tympanum when hind limb stretched along body in the
|
||
<typeStatus id="54FB4AE0FFFCA02E9A7AD33EFED3C18F" box="[283,381,619,645]" pageId="23" pageNumber="114" type="holotype">holotype</typeStatus>
|
||
(vs. tibio-tarsal articulation reaching forward to posterior margin of tympanum in the
|
||
<typeStatus id="54FB4AE0FFFCA02E9E5DD33EFA32C18F" box="[1340,1436,619,645]" pageId="23" pageNumber="114" type="paratype">paratype</typeStatus>
|
||
SYS a004725; reaching forward to the region between tympanum and eye in the
|
||
<typeStatus id="54FB4AE0FFFCA02E9F44D3DAFB2BC1A3" box="[1061,1157,655,681]" pageId="23" pageNumber="114" type="paratype">paratype</typeStatus>
|
||
SYS a004726). Females are distinctly larger than the males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3FA4CAFFFCA02E9BF6D3AFFF4BC033" ID-Table-UUID="DF3FA4CAFFFCA02E9BF6D3AFFF4BC033" httpUri="http://table.plazi.org/id/DF3FA4CAFFFCA02E9BF6D3AFFF4BC033" pageId="23" pageNumber="114" startId="23.[151,244,762,788]" targetBox="[164,1406,841,1458]" targetIsTable="true" targetPageId="23">
|
||
<paragraph id="8BFFF442FFFCA02E9BF6D3AFFF4BC033" blockId="23.[151,1436,762,825]" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BF6D3AFFEBCC01E" bold="true" box="[151,274,762,788]" pageId="23" pageNumber="114">TABLE 6.</emphasis>
|
||
Measurements (in mm) of the type series of
|
||
<taxonomicName id="4C408FC1FFFCA02E9874D3AEFBF8C01F" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[789,1110,763,789]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="23" pageNumber="114" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFCA02E9874D3AEFBF8C01F" box="[789,1110,763,789]" italics="true" pageId="23" pageNumber="114">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207952BFFFCA02E9F3FD3AEFB14C01F" box="[1118,1210,763,789]" pageId="23" pageNumber="114" rank="species">
|
||
<emphasis id="B9342850FFFCA02E9F3FD3AEFB1AC01F" bold="true" box="[1118,1204,763,789]" pageId="23" pageNumber="114">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, * indicates the holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFFF442FFFCA02E9A44D21CFAC7C6BB" pageId="23" pageNumber="114">
|
||
<table id="F94006E2FFFC5FC69BC5D21CFAD0C6B8" box="[164,1406,841,1458]" gridcols="9" gridrows="15" pageId="23" pageNumber="114">
|
||
<tr id="3570F600FFFC5FC69BC5D21CFAD0C074" box="[164,1406,841,894]" gridrow="0" pageId="23" pageNumber="114" rowspan-0="1">
|
||
<th id="76A19F7CFFFC5FC69A6BD21CFEC0C074" box="[266,366,841,894]" gridcol="1" gridrow="0" pageId="23" pageNumber="114">SYS a004744*</th>
|
||
<th id="76A19F7CFFFC5FC69AC7D21CFE50C074" box="[422,510,841,894]" gridcol="2" gridrow="0" pageId="23" pageNumber="114">SYS a004724</th>
|
||
<th id="76A19F7CFFFC5FC6995DD21CFD3AC074" box="[572,660,841,894]" gridcol="3" gridrow="0" pageId="23" pageNumber="114">SYS a004725</th>
|
||
<th id="76A19F7CFFFC5FC699B0D21CFC87C074" box="[721,809,841,894]" gridcol="4" gridrow="0" pageId="23" pageNumber="114">SYS a004726</th>
|
||
<th id="76A19F7CFFFC5FC69806D21CFC10C074" box="[871,958,841,894]" gridcol="5" gridrow="0" pageId="23" pageNumber="114">SYS a004727</th>
|
||
<th id="76A19F7CFFFC5FC6989DD21CFBFAC074" box="[1020,1108,841,894]" gridcol="6" gridrow="0" pageId="23" pageNumber="114">SYS a004728</th>
|
||
<th id="76A19F7CFFFC5FC69FF3D21CFB47C074" box="[1170,1257,841,894]" gridcol="7" gridrow="0" pageId="23" pageNumber="114">SYS a005220</th>
|
||
<th id="76A19F7CFFFC5FC69E46D21CFAD0C074" box="[1319,1406,841,894]" gridcol="8" gridrow="0" pageId="23" pageNumber="114">SYS a005221</th>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D2DBFAD0C0AC" box="[164,1406,910,934]" gridrow="1" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D2DBFF4DC0AC" box="[164,227,910,934]" gridcol="0" gridrow="1" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BD0D2DBFF79C0AC" bold="true" box="[177,215,910,934]" pageId="23" pageNumber="114">Sex</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD2DBFEC0C0AC" box="[266,366,910,934]" gridcol="1" gridrow="1" pageId="23" pageNumber="114">Male</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D2DBFE50C0AC" box="[422,510,910,934]" gridcol="2" gridrow="1" pageId="23" pageNumber="114">Male</td>
|
||
<td id="76A19F7CFFFC5FC6995DD2DBFD3AC0AC" box="[572,660,910,934]" gridcol="3" gridrow="1" pageId="23" pageNumber="114">Male</td>
|
||
<td id="76A19F7CFFFC5FC699B0D2DBFC87C0AC" box="[721,809,910,934]" gridcol="4" gridrow="1" pageId="23" pageNumber="114">Male</td>
|
||
<td id="76A19F7CFFFC5FC69806D2DBFC10C0AC" box="[871,958,910,934]" gridcol="5" gridrow="1" pageId="23" pageNumber="114">Male</td>
|
||
<td id="76A19F7CFFFC5FC6989DD2DBFBFAC0AC" box="[1020,1108,910,934]" gridcol="6" gridrow="1" pageId="23" pageNumber="114">Male</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D2DBFB47C0AC" box="[1170,1257,910,934]" gridcol="7" gridrow="1" pageId="23" pageNumber="114">Female</td>
|
||
<td id="76A19F7CFFFC5FC69E46D2DBFAD0C0AC" box="[1319,1406,910,934]" gridcol="8" gridrow="1" pageId="23" pageNumber="114">Female</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D2E3FAD0C0C5" box="[164,1406,950,975]" gridrow="2" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D2E3FF4DC0C5" box="[164,227,950,975]" gridcol="0" gridrow="2" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BCAD2E3FF72C0C4" bold="true" box="[171,220,950,974]" pageId="23" pageNumber="114">SVL</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD2E3FEC0C0C5" box="[266,366,950,975]" gridcol="1" gridrow="2" pageId="23" pageNumber="114">37.4</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D2E3FE50C0C5" box="[422,510,950,975]" gridcol="2" gridrow="2" pageId="23" pageNumber="114">36.5</td>
|
||
<td id="76A19F7CFFFC5FC6995DD2E3FD3AC0C5" box="[572,660,950,975]" gridcol="3" gridrow="2" pageId="23" pageNumber="114">39.4</td>
|
||
<td id="76A19F7CFFFC5FC699B0D2E3FC87C0C5" box="[721,809,950,975]" gridcol="4" gridrow="2" pageId="23" pageNumber="114">34.3</td>
|
||
<td id="76A19F7CFFFC5FC69806D2E3FC10C0C5" box="[871,958,950,975]" gridcol="5" gridrow="2" pageId="23" pageNumber="114">37.5</td>
|
||
<td id="76A19F7CFFFC5FC6989DD2E3FBFAC0C5" box="[1020,1108,950,975]" gridcol="6" gridrow="2" pageId="23" pageNumber="114">36.4</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D2E3FB47C0C5" box="[1170,1257,950,975]" gridcol="7" gridrow="2" pageId="23" pageNumber="114">42.5</td>
|
||
<td id="76A19F7CFFFC5FC69E46D2E3FAD0C0C5" box="[1319,1406,950,975]" gridcol="8" gridrow="2" pageId="23" pageNumber="114">44.9</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D28AFAD0C0FD" box="[164,1406,991,1015]" gridrow="3" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D28AFF4DC0FD" box="[164,227,991,1015]" gridcol="0" gridrow="3" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BC9D28AFF71C0FD" bold="true" box="[168,223,991,1015]" pageId="23" pageNumber="114">HDL</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD28AFEC0C0FD" box="[266,366,991,1015]" gridcol="1" gridrow="3" pageId="23" pageNumber="114">12.4</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D28AFE50C0FD" box="[422,510,991,1015]" gridcol="2" gridrow="3" pageId="23" pageNumber="114">13.0</td>
|
||
<td id="76A19F7CFFFC5FC6995DD28AFD3AC0FD" box="[572,660,991,1015]" gridcol="3" gridrow="3" pageId="23" pageNumber="114">13.3</td>
|
||
<td id="76A19F7CFFFC5FC699B0D28AFC87C0FD" box="[721,809,991,1015]" gridcol="4" gridrow="3" pageId="23" pageNumber="114">12.3</td>
|
||
<td id="76A19F7CFFFC5FC69806D28AFC10C0FD" box="[871,958,991,1015]" gridcol="5" gridrow="3" pageId="23" pageNumber="114">12.2</td>
|
||
<td id="76A19F7CFFFC5FC6989DD28AFBFAC0FD" box="[1020,1108,991,1015]" gridcol="6" gridrow="3" pageId="23" pageNumber="114">12.3</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D28AFB47C0FD" box="[1170,1257,991,1015]" gridcol="7" gridrow="3" pageId="23" pageNumber="114">14.3</td>
|
||
<td id="76A19F7CFFFC5FC69E46D28AFAD0C0FD" box="[1319,1406,991,1015]" gridcol="8" gridrow="3" pageId="23" pageNumber="114">14.3</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D552FAD0C715" box="[164,1406,1031,1055]" gridrow="4" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D552FF4DC715" box="[164,227,1031,1055]" gridcol="0" gridrow="4" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BC5D552FF4DC715" bold="true" box="[164,227,1031,1055]" pageId="23" pageNumber="114">HDW</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD552FEC0C715" box="[266,366,1031,1055]" gridcol="1" gridrow="4" pageId="23" pageNumber="114">14.5</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D552FE50C715" box="[422,510,1031,1055]" gridcol="2" gridrow="4" pageId="23" pageNumber="114">13.7</td>
|
||
<td id="76A19F7CFFFC5FC6995DD552FD3AC715" box="[572,660,1031,1055]" gridcol="3" gridrow="4" pageId="23" pageNumber="114">14.1</td>
|
||
<td id="76A19F7CFFFC5FC699B0D552FC87C715" box="[721,809,1031,1055]" gridcol="4" gridrow="4" pageId="23" pageNumber="114">12.7</td>
|
||
<td id="76A19F7CFFFC5FC69806D552FC10C715" box="[871,958,1031,1055]" gridcol="5" gridrow="4" pageId="23" pageNumber="114">13.2</td>
|
||
<td id="76A19F7CFFFC5FC6989DD552FBFAC715" box="[1020,1108,1031,1055]" gridcol="6" gridrow="4" pageId="23" pageNumber="114">13.0</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D552FB47C715" box="[1170,1257,1031,1055]" gridcol="7" gridrow="4" pageId="23" pageNumber="114">15.2</td>
|
||
<td id="76A19F7CFFFC5FC69E46D552FAD0C715" box="[1319,1406,1031,1055]" gridcol="8" gridrow="4" pageId="23" pageNumber="114">15.3</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D57AFAD0C74D" box="[164,1406,1071,1095]" gridrow="5" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D57AFF4DC74D" box="[164,227,1071,1095]" gridcol="0" gridrow="5" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BCAD57AFF72C74D" bold="true" box="[171,220,1071,1095]" pageId="23" pageNumber="114">SNT</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD57AFEC0C74D" box="[266,366,1071,1095]" gridcol="1" gridrow="5" pageId="23" pageNumber="114">4.4</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D57AFE50C74D" box="[422,510,1071,1095]" gridcol="2" gridrow="5" pageId="23" pageNumber="114">4.5</td>
|
||
<td id="76A19F7CFFFC5FC6995DD57AFD3AC74D" box="[572,660,1071,1095]" gridcol="3" gridrow="5" pageId="23" pageNumber="114">5.1</td>
|
||
<td id="76A19F7CFFFC5FC699B0D57AFC87C74D" box="[721,809,1071,1095]" gridcol="4" gridrow="5" pageId="23" pageNumber="114">4.6</td>
|
||
<td id="76A19F7CFFFC5FC69806D57AFC10C74D" box="[871,958,1071,1095]" gridcol="5" gridrow="5" pageId="23" pageNumber="114">4.8</td>
|
||
<td id="76A19F7CFFFC5FC6989DD57AFBFAC74D" box="[1020,1108,1071,1095]" gridcol="6" gridrow="5" pageId="23" pageNumber="114">4.7</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D57AFB47C74D" box="[1170,1257,1071,1095]" gridcol="7" gridrow="5" pageId="23" pageNumber="114">5.0</td>
|
||
<td id="76A19F7CFFFC5FC69E46D57AFAD0C74D" box="[1319,1406,1071,1095]" gridcol="8" gridrow="5" pageId="23" pageNumber="114">4.9</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D502FAD0C77A" box="[164,1406,1111,1136]" gridrow="6" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D502FF4DC77A" box="[164,227,1111,1136]" gridcol="0" gridrow="6" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BCDD502FF74C765" bold="true" box="[172,218,1111,1135]" pageId="23" pageNumber="114">IND</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD502FEC0C77A" box="[266,366,1111,1136]" gridcol="1" gridrow="6" pageId="23" pageNumber="114">4.4</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D502FE50C77A" box="[422,510,1111,1136]" gridcol="2" gridrow="6" pageId="23" pageNumber="114">4.2</td>
|
||
<td id="76A19F7CFFFC5FC6995DD502FD3AC77A" box="[572,660,1111,1136]" gridcol="3" gridrow="6" pageId="23" pageNumber="114">4.3</td>
|
||
<td id="76A19F7CFFFC5FC699B0D502FC87C77A" box="[721,809,1111,1136]" gridcol="4" gridrow="6" pageId="23" pageNumber="114">3.9</td>
|
||
<td id="76A19F7CFFFC5FC69806D502FC10C77A" box="[871,958,1111,1136]" gridcol="5" gridrow="6" pageId="23" pageNumber="114">3.7</td>
|
||
<td id="76A19F7CFFFC5FC6989DD502FBFAC77A" box="[1020,1108,1111,1136]" gridcol="6" gridrow="6" pageId="23" pageNumber="114">4.2</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D502FB47C77A" box="[1170,1257,1111,1136]" gridcol="7" gridrow="6" pageId="23" pageNumber="114">4.3</td>
|
||
<td id="76A19F7CFFFC5FC69E46D502FAD0C77A" box="[1319,1406,1111,1136]" gridcol="8" gridrow="6" pageId="23" pageNumber="114">4.3</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D52AFAD0C792" box="[164,1406,1151,1176]" gridrow="7" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D52AFF4DC792" box="[164,227,1151,1176]" gridcol="0" gridrow="7" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BCDD52AFF72C79D" bold="true" box="[172,220,1151,1175]" pageId="23" pageNumber="114">IOD</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD52AFEC0C792" box="[266,366,1151,1176]" gridcol="1" gridrow="7" pageId="23" pageNumber="114">4.3</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D52AFE50C792" box="[422,510,1151,1176]" gridcol="2" gridrow="7" pageId="23" pageNumber="114">3.9</td>
|
||
<td id="76A19F7CFFFC5FC6995DD52AFD3AC792" box="[572,660,1151,1176]" gridcol="3" gridrow="7" pageId="23" pageNumber="114">4.0</td>
|
||
<td id="76A19F7CFFFC5FC699B0D52AFC87C792" box="[721,809,1151,1176]" gridcol="4" gridrow="7" pageId="23" pageNumber="114">3.6</td>
|
||
<td id="76A19F7CFFFC5FC69806D52AFC10C792" box="[871,958,1151,1176]" gridcol="5" gridrow="7" pageId="23" pageNumber="114">3.6</td>
|
||
<td id="76A19F7CFFFC5FC6989DD52AFBFAC792" box="[1020,1108,1151,1176]" gridcol="6" gridrow="7" pageId="23" pageNumber="114">3.9</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D52AFB47C792" box="[1170,1257,1151,1176]" gridcol="7" gridrow="7" pageId="23" pageNumber="114">4.1</td>
|
||
<td id="76A19F7CFFFC5FC69E46D52AFAD0C792" box="[1319,1406,1151,1176]" gridcol="8" gridrow="7" pageId="23" pageNumber="114">4.2</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D5FDFAD0C7CA" box="[164,1406,1192,1216]" gridrow="8" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D5FDFF4DC7CA" box="[164,227,1192,1216]" gridcol="0" gridrow="8" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BD3D5FDFF7BC7CA" bold="true" box="[178,213,1192,1216]" pageId="23" pageNumber="114">ED</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD5FDFEC0C7CA" box="[266,366,1192,1216]" gridcol="1" gridrow="8" pageId="23" pageNumber="114">4.6</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D5FDFE50C7CA" box="[422,510,1192,1216]" gridcol="2" gridrow="8" pageId="23" pageNumber="114">4.0</td>
|
||
<td id="76A19F7CFFFC5FC6995DD5FDFD3AC7CA" box="[572,660,1192,1216]" gridcol="3" gridrow="8" pageId="23" pageNumber="114">4.3</td>
|
||
<td id="76A19F7CFFFC5FC699B0D5FDFC87C7CA" box="[721,809,1192,1216]" gridcol="4" gridrow="8" pageId="23" pageNumber="114">3.7</td>
|
||
<td id="76A19F7CFFFC5FC69806D5FDFC10C7CA" box="[871,958,1192,1216]" gridcol="5" gridrow="8" pageId="23" pageNumber="114">4.4</td>
|
||
<td id="76A19F7CFFFC5FC6989DD5FDFBFAC7CA" box="[1020,1108,1192,1216]" gridcol="6" gridrow="8" pageId="23" pageNumber="114">4.1</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D5FDFB47C7CA" box="[1170,1257,1192,1216]" gridcol="7" gridrow="8" pageId="23" pageNumber="114">4.1</td>
|
||
<td id="76A19F7CFFFC5FC69E46D5FDFAD0C7CA" box="[1319,1406,1192,1216]" gridcol="8" gridrow="8" pageId="23" pageNumber="114">5.1</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D585FAD0C7E2" box="[164,1406,1232,1256]" gridrow="9" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D585FF4DC7E2" box="[164,227,1232,1256]" gridcol="0" gridrow="9" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BD3D585FF7BC7E2" bold="true" box="[178,213,1232,1256]" pageId="23" pageNumber="114">TD</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD585FEC0C7E2" box="[266,366,1232,1256]" gridcol="1" gridrow="9" pageId="23" pageNumber="114">3.0</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D585FE50C7E2" box="[422,510,1232,1256]" gridcol="2" gridrow="9" pageId="23" pageNumber="114">2.4</td>
|
||
<td id="76A19F7CFFFC5FC6995DD585FD3AC7E2" box="[572,660,1232,1256]" gridcol="3" gridrow="9" pageId="23" pageNumber="114">2.8</td>
|
||
<td id="76A19F7CFFFC5FC699B0D585FC87C7E2" box="[721,809,1232,1256]" gridcol="4" gridrow="9" pageId="23" pageNumber="114">2.5</td>
|
||
<td id="76A19F7CFFFC5FC69806D585FC10C7E2" box="[871,958,1232,1256]" gridcol="5" gridrow="9" pageId="23" pageNumber="114">2.6</td>
|
||
<td id="76A19F7CFFFC5FC6989DD585FBFAC7E2" box="[1020,1108,1232,1256]" gridcol="6" gridrow="9" pageId="23" pageNumber="114">2.4</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D585FB47C7E2" box="[1170,1257,1232,1256]" gridcol="7" gridrow="9" pageId="23" pageNumber="114">2.3</td>
|
||
<td id="76A19F7CFFFC5FC69E46D585FAD0C7E2" box="[1319,1406,1232,1256]" gridcol="8" gridrow="9" pageId="23" pageNumber="114">3.0</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D5ADFAD0C61B" box="[164,1406,1272,1297]" gridrow="10" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D5ADFF4DC61B" box="[164,227,1272,1297]" gridcol="0" gridrow="10" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BCBD5ADFF70C61A" bold="true" box="[170,222,1272,1296]" pageId="23" pageNumber="114">TED</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD5ADFEC0C61B" box="[266,366,1272,1297]" gridcol="1" gridrow="10" pageId="23" pageNumber="114">2.1</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D5ADFE50C61B" box="[422,510,1272,1297]" gridcol="2" gridrow="10" pageId="23" pageNumber="114">2.0</td>
|
||
<td id="76A19F7CFFFC5FC6995DD5ADFD3AC61B" box="[572,660,1272,1297]" gridcol="3" gridrow="10" pageId="23" pageNumber="114">2.1</td>
|
||
<td id="76A19F7CFFFC5FC699B0D5ADFC87C61B" box="[721,809,1272,1297]" gridcol="4" gridrow="10" pageId="23" pageNumber="114">2.2</td>
|
||
<td id="76A19F7CFFFC5FC69806D5ADFC10C61B" box="[871,958,1272,1297]" gridcol="5" gridrow="10" pageId="23" pageNumber="114">1.9</td>
|
||
<td id="76A19F7CFFFC5FC6989DD5ADFBFAC61B" box="[1020,1108,1272,1297]" gridcol="6" gridrow="10" pageId="23" pageNumber="114">2.0</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D5ADFB47C61B" box="[1170,1257,1272,1297]" gridcol="7" gridrow="10" pageId="23" pageNumber="114">2.2</td>
|
||
<td id="76A19F7CFFFC5FC69E46D5ADFAD0C61B" box="[1319,1406,1272,1297]" gridcol="8" gridrow="10" pageId="23" pageNumber="114">2.9</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D475FAD0C633" box="[164,1406,1312,1337]" gridrow="11" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D475FF4DC633" box="[164,227,1312,1337]" gridcol="0" gridrow="11" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BC9D474FF4EC633" bold="true" box="[168,224,1313,1337]" pageId="23" pageNumber="114">HND</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD475FEC0C633" box="[266,366,1312,1337]" gridcol="1" gridrow="11" pageId="23" pageNumber="114">9.4</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D475FE50C633" box="[422,510,1312,1337]" gridcol="2" gridrow="11" pageId="23" pageNumber="114">9.4</td>
|
||
<td id="76A19F7CFFFC5FC6995DD475FD3AC633" box="[572,660,1312,1337]" gridcol="3" gridrow="11" pageId="23" pageNumber="114">9.0</td>
|
||
<td id="76A19F7CFFFC5FC699B0D475FC87C633" box="[721,809,1312,1337]" gridcol="4" gridrow="11" pageId="23" pageNumber="114">9.5</td>
|
||
<td id="76A19F7CFFFC5FC69806D475FC10C633" box="[871,958,1312,1337]" gridcol="5" gridrow="11" pageId="23" pageNumber="114">9.1</td>
|
||
<td id="76A19F7CFFFC5FC6989DD475FBFAC633" box="[1020,1108,1312,1337]" gridcol="6" gridrow="11" pageId="23" pageNumber="114">9.2</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D475FB47C633" box="[1170,1257,1312,1337]" gridcol="7" gridrow="11" pageId="23" pageNumber="114">10.9</td>
|
||
<td id="76A19F7CFFFC5FC69E46D475FAD0C633" box="[1319,1406,1312,1337]" gridcol="8" gridrow="11" pageId="23" pageNumber="114">10.9</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D41DFAD0C66B" box="[164,1406,1352,1377]" gridrow="12" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D41DFF4DC66B" box="[164,227,1352,1377]" gridcol="0" gridrow="12" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BC9D41DFF71C66A" bold="true" box="[168,223,1352,1376]" pageId="23" pageNumber="114">RAD</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD41DFEC0C66B" box="[266,366,1352,1377]" gridcol="1" gridrow="12" pageId="23" pageNumber="114">7.9</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D41DFE50C66B" box="[422,510,1352,1377]" gridcol="2" gridrow="12" pageId="23" pageNumber="114">7.7</td>
|
||
<td id="76A19F7CFFFC5FC6995DD41DFD3AC66B" box="[572,660,1352,1377]" gridcol="3" gridrow="12" pageId="23" pageNumber="114">8.1</td>
|
||
<td id="76A19F7CFFFC5FC699B0D41DFC87C66B" box="[721,809,1352,1377]" gridcol="4" gridrow="12" pageId="23" pageNumber="114">8.1</td>
|
||
<td id="76A19F7CFFFC5FC69806D41DFC10C66B" box="[871,958,1352,1377]" gridcol="5" gridrow="12" pageId="23" pageNumber="114">8.2</td>
|
||
<td id="76A19F7CFFFC5FC6989DD41DFBFAC66B" box="[1020,1108,1352,1377]" gridcol="6" gridrow="12" pageId="23" pageNumber="114">8.2</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D41DFB47C66B" box="[1170,1257,1352,1377]" gridcol="7" gridrow="12" pageId="23" pageNumber="114">9.0</td>
|
||
<td id="76A19F7CFFFC5FC69E46D41DFAD0C66B" box="[1319,1406,1352,1377]" gridcol="8" gridrow="12" pageId="23" pageNumber="114">9.0</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D424FAD0C683" box="[164,1406,1393,1417]" gridrow="13" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D424FF4DC683" box="[164,227,1393,1417]" gridcol="0" gridrow="13" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BCFD424FF74C683" bold="true" box="[174,218,1393,1417]" pageId="23" pageNumber="114">TIB</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD424FEC0C683" box="[266,366,1393,1417]" gridcol="1" gridrow="13" pageId="23" pageNumber="114">15.66</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D424FE50C683" box="[422,510,1393,1417]" gridcol="2" gridrow="13" pageId="23" pageNumber="114">14.9</td>
|
||
<td id="76A19F7CFFFC5FC6995DD424FD3AC683" box="[572,660,1393,1417]" gridcol="3" gridrow="13" pageId="23" pageNumber="114">14.81</td>
|
||
<td id="76A19F7CFFFC5FC699B0D424FC87C683" box="[721,809,1393,1417]" gridcol="4" gridrow="13" pageId="23" pageNumber="114">14.73</td>
|
||
<td id="76A19F7CFFFC5FC69806D424FC10C683" box="[871,958,1393,1417]" gridcol="5" gridrow="13" pageId="23" pageNumber="114">15.24</td>
|
||
<td id="76A19F7CFFFC5FC6989DD424FBFAC683" box="[1020,1108,1393,1417]" gridcol="6" gridrow="13" pageId="23" pageNumber="114">15.6</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D424FB47C683" box="[1170,1257,1393,1417]" gridcol="7" gridrow="13" pageId="23" pageNumber="114">17.37</td>
|
||
<td id="76A19F7CFFFC5FC69E46D424FAD0C683" box="[1319,1406,1393,1417]" gridcol="8" gridrow="13" pageId="23" pageNumber="114">17.99</td>
|
||
</tr>
|
||
<tr id="3570F600FFFC5FC69BC5D4CCFAD0C6B8" box="[164,1406,1433,1458]" gridrow="14" pageId="23" pageNumber="114">
|
||
<th id="76A19F7CFFFC5FC69BC5D4CCFF4DC6B8" box="[164,227,1433,1458]" gridcol="0" gridrow="14" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BCAD4CCFF72C6BB" bold="true" box="[171,220,1433,1457]" pageId="23" pageNumber="114">FTL</emphasis>
|
||
</th>
|
||
<td id="76A19F7CFFFC5FC69A6BD4CCFEC0C6B8" box="[266,366,1433,1458]" gridcol="1" gridrow="14" pageId="23" pageNumber="114">22.2</td>
|
||
<td id="76A19F7CFFFC5FC69AC7D4CCFE50C6B8" box="[422,510,1433,1458]" gridcol="2" gridrow="14" pageId="23" pageNumber="114">21.4</td>
|
||
<td id="76A19F7CFFFC5FC6995DD4CCFD3AC6B8" box="[572,660,1433,1458]" gridcol="3" gridrow="14" pageId="23" pageNumber="114">21.3</td>
|
||
<td id="76A19F7CFFFC5FC699B0D4CCFC87C6B8" box="[721,809,1433,1458]" gridcol="4" gridrow="14" pageId="23" pageNumber="114">22.2</td>
|
||
<td id="76A19F7CFFFC5FC69806D4CCFC10C6B8" box="[871,958,1433,1458]" gridcol="5" gridrow="14" pageId="23" pageNumber="114">23.3</td>
|
||
<td id="76A19F7CFFFC5FC6989DD4CCFBFAC6B8" box="[1020,1108,1433,1458]" gridcol="6" gridrow="14" pageId="23" pageNumber="114">23.0</td>
|
||
<td id="76A19F7CFFFC5FC69FF3D4CCFB47C6B8" box="[1170,1257,1433,1458]" gridcol="7" gridrow="14" pageId="23" pageNumber="114">26.4</td>
|
||
<td id="76A19F7CFFFC5FC69E46D4CCFAD0C6B8" box="[1319,1406,1433,1458]" gridcol="8" gridrow="14" pageId="23" pageNumber="114">25.7</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C35AA7C9FFFCA02E9BA6D4B6FC12C5DF" pageId="23" pageNumber="114" type="distribution">
|
||
<paragraph id="8BFFF442FFFCA02E9BA6D4B6FC12C5DF" blockId="23.[151,1436,1507,1750]" pageId="23" pageNumber="114">
|
||
<emphasis id="B9342850FFFCA02E9BA6D4B6FDF6C6F7" bold="true" box="[199,600,1507,1533]" pageId="23" pageNumber="114">Distribution and natural history.</emphasis>
|
||
Currently,
|
||
<taxonomicName id="4C408FC1FFFCA02E99BCD4B6FB8EC6F7" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[733,1056,1507,1533]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="23" pageNumber="114" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFCA02E99BCD4B6FB8EC6F7" box="[733,1056,1507,1533]" italics="true" pageId="23" pageNumber="114">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFCA02E9F4BD4B6FB29C6F7" bold="true" box="[1066,1159,1507,1533]" pageId="23" pageNumber="114">
|
||
<taxonomicNameLabel id="A207952BFFFCA02E9F4BD4B6FB29C6F7" box="[1066,1159,1507,1533]" pageId="23" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is only known from its
|
||
<typeStatus id="54FB4AE0FFFCA02E9BF6D75DFF69C528" box="[151,199,1544,1570]" pageId="23" pageNumber="114">type</typeStatus>
|
||
locality, Mt. Tongguzhang of Fengshun. It inhabits flowing montane streams and the nearby forest floor and leaf litter at elevations between
|
||
<quantity id="4CB859A7FFFCA02E9962D77EFD3BC54C" box="[515,661,1579,1606]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="23" pageNumber="114" unit="m" value="1150.0" valueMax="1500.0" valueMin="800.0">800–1500 m</quantity>
|
||
.
|
||
<taxonomicName id="4C408FC1FFFCA02E99C0D77EFC4CC54F" authority="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang, 2022" authorityName="Wang & Zeng & Lyu & Qi & Liu & Chen & Lu & Xiao & Lin & Chen & Wang" authorityYear="2022" box="[673,994,1579,1605]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="23" pageNumber="114" phylum="Chordata" rank="species" species="fengshunensis" status="sp. nov.">
|
||
<emphasis id="B9342850FFFCA02E99C0D77EFC4CC54F" box="[673,994,1579,1605]" italics="true" pageId="23" pageNumber="114">Boulenophrys fengshunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9342850FFFCA02E988BD77EFBEBC54F" bold="true" box="[1002,1093,1579,1605]" pageId="23" pageNumber="114">
|
||
<taxonomicNameLabel id="A207952BFFFCA02E988BD77EFBEBC54F" box="[1002,1093,1579,1605]" pageId="23" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is found to be sympatric with
|
||
<taxonomicName id="4C408FC1FFFCA02E9BF6D71AFD99C563" authority="Chang, 1933" authorityName="Chang" authorityYear="1933" box="[151,567,1615,1641]" class="Amphibia" family="Salamandridae" genus="Pachytriton" kingdom="Animalia" order="Caudata" pageId="23" pageNumber="114" phylum="Chordata" rank="species" species="granulosus">
|
||
<emphasis id="B9342850FFFCA02E9BF6D71AFE31C563" box="[151,415,1615,1641]" italics="true" pageId="23" pageNumber="114">Pachytriton granulosus</emphasis>
|
||
<bibRefCitation id="EFD189B3FFFCA02E9AC6D71AFD99C563" author="Chang, M. L. Y." box="[423,567,1615,1641]" pageId="23" pageNumber="114" pagination="305 - 328" refId="ref14649" refString="Chang, M. L. Y. (1933) On the salamanders of Chekiang. Contributions from the Biological Laboratory of the Science Society of China. Zoological Series, 9, 305 - 328." type="journal article" year="1933">Chang, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C408FC1FFFCA02E990ED705FB44C563" authority="Yuan, Jiang, Ding, Zhang & Che, 2013" authorityName="Yuan, Jiang, Ding, Zhang & Che" authorityYear="2013" box="[623,1258,1615,1642]" class="Amphibia" family="Salamandridae" genus="Cynops" kingdom="Animalia" order="Caudata" pageId="23" pageNumber="114" phylum="Chordata" rank="species" species="glaucus">
|
||
<emphasis id="B9342850FFFCA02E990ED705FC8CC563" box="[623,802,1615,1642]" italics="true" pageId="23" pageNumber="114">Cynops glaucus</emphasis>
|
||
<bibRefCitation id="EFD189B3FFFCA02E9850D705FB44C563" author="Yuan, Z. Y. & Jiang, K. & Ding, L. & Zhang, L. & Che, J." box="[817,1258,1615,1642]" pageId="23" pageNumber="114" pagination="116 - 123" refId="ref19092" refString="Yuan, Z. Y., Jiang, K., Ding, L., Zhang, L. & Che, J. (2013) A new newt of the genus Cynops (Caudata: Salamandridae) from Guangdong, China. Asian Herpetological Research, 4 (2), 116 - 123. https: // doi. org / 10.3724 / SP. J. 1245.2013.00116" type="journal article" year="2013">Yuan, Jiang, Ding, Zhang & Che, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
. Advertisement calls of males were heard during April and June. Males were found calling under the leaf litters around the flowing seeps. Tadpoles could be found in this period. Female specimens collected during August had immature eggs; males were not heard calling during this period, but sub-adults were observed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |