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<document id="9E2FFDE5B9B3FC7E64D1CDC1CB32A814" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFE9501BFF23FC1EFCFB47BA" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus anosyensis Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="26" updateTime="1726053143081" updateUser="ExternalLinkService">
<mods:mods id="4E95AF603A4C5E9382214DA1E6E6FEDE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D443C93101AB76BE1FEE5DB6BE88D6DA">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="0646CE4DB5AE46F8E1580E9559C39D6C">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="7262DD9D4EFD66C49028B77A22D8D337">Zootaxa</mods:title>
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<mods:date id="6B74F04C1ABE106CAA8CBE454591D50F">2024</mods:date>
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<mods:classification id="EB807FA316D34A0D02343575881766EC">journal article</mods:classification>
<mods:identifier id="271C13044359F5319B698577C8707B83" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
<mods:identifier id="0D325CE29FD41F9C959A9D258EDD919B" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03E7FF50FFE9501BFF23FC1EFCFB47BA" ID-DOI="http://doi.org/10.5281/zenodo.13746027" ID-Zenodo-Dep="13746027" LSID="urn:lsid:plazi:treatment:03E7FF50FFE9501BFF23FC1EFCFB47BA" httpUri="http://treatment.plazi.org/id/03E7FF50FFE9501BFF23FC1EFCFB47BA" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="C3541DCDFFE95019FF23FC1EFD20479E" box="[151,741,978,1005]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BF14E46FFE95019FF23FC1EFD20479E" blockId="25.[151,741,978,1041]" box="[151,741,978,1005]" pageId="25" pageNumber="26">
<heading id="D0B9F92AFFE95019FF23FC1EFD20479E" bold="true" box="[151,741,978,1005]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="4">
<emphasis id="B93A9254FFE95019FF23FC1EFD20479E" bold="true" box="[151,741,978,1005]" pageId="25" pageNumber="26">
<taxonomicName id="4C4E35C5FFE95019FF23FC1EFDA9479F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,620,978,1005]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="anosyensis" status="comb. nov.">
<emphasis id="B93A9254FFE95019FF23FC1EFDC0479E" bold="true" box="[151,517,978,1005]" italics="true" pageId="25" pageNumber="26">Anhoraeomorphus anosyensis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFE95019FDCEFC1FFD20479E" box="[634,741,979,1005]" pageId="25" pageNumber="26" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE95019FF23FC3BFEE24062" box="[151,295,1015,1041]" pageId="25" pageNumber="26" type="description">
<paragraph id="8BF14E46FFE95019FF23FC3BFEE24062" blockId="25.[151,741,978,1041]" box="[151,295,1015,1041]" pageId="25" pageNumber="26">
(
<figureCitation id="137552C3FFE95019FF2BFC3BFEDA4062" box="[159,287,1015,1041]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Figs 91–98</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE95019FF23FBF7FDBD4020" box="[151,632,1083,1107]" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="8BF14E46FFE95019FF23FBF7FDBD4020" blockId="25.[151,632,1083,1107]" box="[151,632,1083,1107]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB5E6968FFE95019FF23FBF7FDBD4020" box="[151,632,1083,1107]" pageId="25" pageNumber="26">
<treatmentCitation id="0AEF6857FFE95019FF23FBF7FDB14020" author="Franz, H." box="[151,628,1083,1107]" page="151" pageId="25" pageNumber="26" year="1986">
<taxonomicName id="4C4E35C5FFE95019FF23FBF7FDB14020" ID-CoL="3MNJ8" authority="Franz, 1986 b: 151" authorityName="Franz" authorityPageNumber="151" authorityYear="1986" box="[151,628,1083,1107]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFE95019FF23FBF7FE774020" box="[151,434,1083,1107]" italics="true" pageId="25" pageNumber="26">Horaeomorphus anosyensis</emphasis>
<bibRefCitation id="EFDF33B7FFE95019FE0CFBF0FDB14020" author="Franz, H." box="[440,628,1083,1107]" pageId="25" pageNumber="26" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 151</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE95019FF23FB4DFDBF4193" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BF14E46FFE95019FF23FB4DFEA84090" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
<emphasis id="B93A9254FFE95019FF23FB4DFE6240E8" bold="true" box="[151,423,1153,1179]" pageId="25" pageNumber="26">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFE95019FE04FB4DFEAC4090" collectionCode="NHMW" collectorName="Foret N Ft. Dauphin" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B93A9254FFE95019FE04FB4DFD0840E8" bold="true" box="[432,717,1153,1179]" pageId="25" pageNumber="26">
<typeStatus id="54F5F0E4FFE95019FE04FB4DFDD840E8" box="[432,541,1153,1179]" pageId="25" pageNumber="26" type="paratype">Paratype</typeStatus>
(
<collectingCountry id="F3590ED6FFE95019FD9AFB4DFD0040E8" box="[558,709,1153,1179]" name="Madagascar" pageId="25" pageNumber="26">Madagascar</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D4885CFFFE95019FD69FB4EFD3140EF" box="[733,756,1154,1180]" count="1" pageId="25" pageNumber="26" type="male">♂</specimenCount>
, two labels: “Madagascar (Süd) /
<collectorName id="26BB2B90FFE95019FBC8FB4DFAA040E8" box="[1148,1381,1153,1180]" pageId="25" pageNumber="26">Forét N Ft. Dauphin</collectorName>
/ lg. Alluaud 1900” [white, printed], “
<taxonomicName id="4C4E35C5FFE95019FDBDFB69FD0540CC" box="[521,704,1189,1215]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ anosiensis {sic!} m. / det.
<determinerName id="7E2DA282FFE95019FC40FB69FB9740CC" box="[1012,1106,1189,1215]" pageId="25" pageNumber="26">H.Franz</determinerName>
” [yellowish, handwritten and printed] (
<collectionCode id="ED5FD683FFE95019FF4BFB05FEA74090" box="[255,354,1225,1251]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFE95019FF73FB21FDBF4193" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
<emphasis id="B93A9254FFE95019FF73FB21FE644174" bold="true" box="[199,417,1261,1287]" pageId="25" pageNumber="26">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFE95019FE40FB21FD804174" box="[500,581,1261,1287]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 91</figureCitation>
) slender; head (
<figureCitation id="137552C3FFE95019FD4CFB21FCBD4174" box="[760,888,1261,1287]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Figs 92–93</figureCitation>
) round, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFE95019FEA0FADDFEA34158" box="[276,358,1297,1323]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 91</figureCitation>
) bell-shaped, broadest in anterior third, with one pair of large antebasal pits; antennomeres 3–6 (
<figureCitation id="137552C3FFE95019FF2BFAF9FF36413C" box="[159,243,1333,1359]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 94</figureCitation>
) each elongate; all femora similarly slender; metatibiae in male modified, strongly broadening to distal third, then narrowing to apex, with tuft of setae on outer surface at widest point; aedeagus (
<figureCitation id="137552C3FFE95019FB14FA95FAE44107" box="[1184,1313,1369,1396]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Figs 95–98</figureCitation>
) in ventral view broadest near base, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites narrowly separated, each strongly elongate and with subtriangular apex, each paramere with subapical group of five short setae distant from apex.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE9501BFF73FA25FD9C464E" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
<paragraph id="8BF14E46FFE95019FF73FA25FEE9425B" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
<emphasis id="B93A9254FFE95019FF73FA25FEB04270" bold="true" box="[199,373,1513,1539]" pageId="25" pageNumber="26">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFE95019FDA8FA25FDAE4270" box="[540,619,1513,1539]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 91</figureCitation>
) moderately dark brown, tarsi and palps distinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFE95019FF75F9C1FEED425B" box="[193,296,1549,1576]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="25" pageNumber="26" unit="mm" value="2.15">2.15 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFE95019FF73F9FDFCE24350" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
Head (
<figureCitation id="137552C3FFE95019FEA0F9FDFEAC4238" box="[276,361,1585,1611]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 92</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFE95019FC0DF9FDFBE7423F" box="[953,1058,1585,1612]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="25" pageNumber="26" unit="mm" value="0.33">0.33 mm</quantity>
, HW
<quantity id="4CB6E3A3FFE95019FBDCF9FDFB17423F" box="[1128,1234,1585,1612]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="mm" value="0.4">0.40 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFE95019FF6AF999FEF5421C" box="[222,304,1621,1647]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 93</figureCitation>
) subequal to eye; vertex and frons confluent, together weakly convex and weakly transverse; vertex posteriorly broadly rounded; each eye moderately large and weakly convex, oval, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFE95019FC97F90DFCB042A8" box="[803,885,1729,1755]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="25" pageNumber="26">Fig. 94</figureCitation>
) slender but short, as long as half BL, AnL
<quantity id="4CB6E3A3FFE95019FADFF90DFF004373" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="25" pageNumber="26" unit="mm" value="1.05">1.05 mm</quantity>
, antennomeres 1–6 each weakly elongate, 7–10 each about as long as broad, 11 much shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFE95019FF73F8E2FAAA43C0" blockId="25.[151,1437,1153,1972]" pageId="25" pageNumber="26">
Pronotum bell-shaped, broadest in anterior third; PL
<quantity id="4CB6E3A3FFE95019FCA4F8E1FCB0433B" box="[784,885,1837,1864]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="25" pageNumber="26" unit="mm" value="0.58">0.58 mm</quantity>
, PW
<quantity id="4CB6E3A3FFE95019FC04F8E1FBD3433B" box="[944,1046,1837,1864]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="0.5">0.50 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, weakly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large round pits, and with short but distinct sublateral carinae. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<caption id="DF311ECEFFEA501AFF23F95DFEF54282" ID-DOI="http://doi.org/10.5281/zenodo.13745955" ID-Zenodo-Dep="13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,264,1681,1705]" targetBox="[151,1418,181,1636]" targetPageId="26" targetType="figure">
<paragraph id="8BF14E46FFEA501AFF23F95DFEF54282" blockId="26.[151,1436,1681,1777]" pageId="26" pageNumber="27">
<emphasis id="B93A9254FFEA501AFF23F95DFE9642DA" bold="true" box="[151,339,1681,1705]" pageId="26" pageNumber="27">FIGURES 91–98.</emphasis>
<taxonomicName id="4C4E35C5FFEA501AFEECF95DFD1F42DA" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[344,730,1681,1705]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFEA501AFEECF95DFD4D42DA" box="[344,648,1681,1705]" italics="true" pageId="26" pageNumber="27">Anhoraeomorphus anosyensis</emphasis>
(Franz)
</taxonomicName>
, paratype. Habitus in dorsal view (
<emphasis id="B93A9254FFEA501AFB82F95DFB8A42DA" bold="true" box="[1078,1103,1681,1705]" pageId="26" pageNumber="27">91</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFEA501AFAC7F95DFA4942DA" bold="true" box="[1395,1420,1681,1705]" pageId="26" pageNumber="27">92</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFEA501AFCE3F979FCB542BE" bold="true" box="[855,880,1717,1741]" pageId="26" pageNumber="27">93</emphasis>
); antenna (
<emphasis id="B93A9254FFEA501AFC57F979FC3942BE" bold="true" box="[995,1020,1717,1741]" pageId="26" pageNumber="27">94</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFEA501AFB50F979FAE642BE" bold="true" box="[1252,1315,1717,1741]" pageId="26" pageNumber="27">95, 97</emphasis>
) and lateral (
<emphasis id="B93A9254FFEA501AFF14F915FF1A4282" bold="true" box="[160,223,1753,1777]" pageId="26" pageNumber="27">96, 98</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFEA501AFF73F88FFEAE43BA" blockId="26.[151,1436,1859,1993]" pageId="26" pageNumber="27">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFEA501AFC39F88FFC31432D" box="[909,1012,1859,1886]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="26" pageNumber="27" unit="mm" value="1.25">1.25 mm</quantity>
, EW
<quantity id="4CB6E3A3FFEA501AFB87F88FFB5F432D" box="[1075,1178,1859,1886]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="26" pageNumber="27" unit="mm" value="0.85">0.85 mm</quantity>
, EI 1.47; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures small and shallow, but more distinct than those on pronotum, those in anterior half separated by spaces subequal to diameters of punctures; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FF5BFF3E456E" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">Legs moderately long, slender, unmodified except large and concave patch of adhesive setae on apical mesal region of protibia and slightly recurved and strongly broadening metatibiae, which are broadest near distal third, where outer margin bears tuft of long setae, and from this point tibia is narrowing distally and its outer margin is concave.</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FEEBFE67466A" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
Aedeagus (
<figureCitation id="137552C3FFEB501BFEF6FEEBFE054532" box="[322,448,294,321]" captionStart="FIGURES 91–98" captionStartId="26.[151,264,1681,1705]" captionTargetBox="[151,1418,181,1636]" captionTargetId="figure-28@26.[151,1435,181,1657]" captionTargetPageId="26" captionText="FIGURES 91–98. Anhoraeomorphus anosyensis (Franz), paratype. Habitus in dorsal view (91); head in anterodorsal view (92); head and prothorax in lateral view (only bristles on head are shown) (93); antenna (94); aedeagus in ventral (95, 97) and lateral (96, 98) views." figureDoi="http://doi.org/10.5281/zenodo.13745955" httpUri="https://zenodo.org/record/13745955/files/figure.png" pageId="27" pageNumber="28">Figs 95–98</figureCitation>
) slender,AeL
<quantity id="4CB6E3A3FFEB501BFDEFFEEBFD054532" box="[603,704,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="mm" value="0.4">0.40 mm</quantity>
; in ventral view broadest near base, drop-shaped, distally gradually narrowing towards broadly subtriangular apex with rounded tip; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with asymmetrical and conspicuously long distolateral sclerites and with strongly asymmetrical assemblage of median proximal sclerotized structures, distolateral sclerites narrowly separated, each with subtriangular apex; parameres broad and their apices slightly projecting beyond aedeagal apex, each paramere with twisted short distal region appearing in ventral view as an apical ‘loop’, and with subapical group of five short setae distant from apex.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FDEFFD9C464E" blockId="27.[151,1437,151,969]" box="[199,601,547,573]" pageId="27" pageNumber="28">Female. Not studied (see Remarks).</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF73FD8AFCD84612" box="[199,797,582,609]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BF14E46FFEB501BFF73FD8AFCD84612" blockId="27.[151,1437,151,969]" box="[199,797,582,609]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF73FD8AFEA44613" bold="true" box="[199,353,582,608]" pageId="27" pageNumber="28">Distribution.</emphasis>
Southern
<collectingCountry id="F3590ED6FFEB501BFE60FD8BFD9E4612" box="[468,603,583,609]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
(Anosy Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEB501BFF73FDA7FCFB47BA" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BF14E46FFEB501BFF73FDA7FAED474A" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
<emphasis id="B93A9254FFEB501BFF73FDA7FEFE46F6" bold="true" box="[199,315,619,645]" pageId="27" pageNumber="28">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFEB501BFEF7FDA7FD5146F7" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[323,660,619,645]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFEB501BFEF7FDA7FD5146F7" box="[323,660,619,645]" italics="true" pageId="27" pageNumber="28">Anhoraeomorphus anosyensis</emphasis>
</taxonomicName>
is highly similar to
<taxonomicName id="4C4E35C5FFEB501BFCCEFDA7FBEE46F7" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[890,1067,619,645]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFEB501BFCCEFDA7FBEE46F7" box="[890,1067,619,645]" italics="true" pageId="27" pageNumber="28">A. roussettensis</emphasis>
</taxonomicName>
. These are the only species with males having metatibiae strongly broadening distally, but in subapical regions rapidly narrowing, so that the outer subapical margin is concave, and there is a tuft of setae on the outer angulate broadening near distal third. These two species share also some structures of the aedeagus,
<emphasis id="B93A9254FFEB501BFCA1FD1BFCF84683" box="[789,829,727,752]" italics="true" pageId="27" pageNumber="28">e.g.</emphasis>
, a pair of very long and dark distolateral endophallic sclerites. However,
<taxonomicName id="4C4E35C5FFEB501BFEC1FD37FDCB4767" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[373,526,763,789]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFEB501BFEC1FD37FDCB4767" box="[373,526,763,789]" italics="true" pageId="27" pageNumber="28">A. anosyensis</emphasis>
</taxonomicName>
has distinctly slenderer antennae with antennomeres 3–6 each elongate, whereas in
<taxonomicName id="4C4E35C5FFEB501BFF00FCD3FEA6474B" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[180,355,799,825]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFEB501BFF00FCD3FEA6474B" box="[180,355,799,825]" italics="true" pageId="27" pageNumber="28">A. roussettensis</emphasis>
</taxonomicName>
antennomeres 3–6 are as long as broad, and the antennae are distinctly more massive.
</paragraph>
<paragraph id="8BF14E46FFEB501BFF73FC8FFCFB47BA" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
<materialsCitation id="3B26441BFFEB501BFF73FC8FFCFF47BA" collectionCode="NHMW" pageId="27" pageNumber="28" specimenCount="3" typeStatus="holotype">
The
<typeStatus id="54F5F0E4FFEB501BFF4DFC8FFE9E472E" box="[249,347,835,861]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
male and a
<typeStatus id="54F5F0E4FFEB501BFE69FC8FFDF8472E" box="[477,573,835,861]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
female of this species are deposited in Muséum National d'Histoire Naturelle, Paris and were not available for my study. The examined
<typeStatus id="54F5F0E4FFEB501BFC92FCABFC4347F2" box="[806,902,871,897]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
male (
<collectionCode id="ED5FD683FFEB501BFC60FCABFBF647F2" box="[980,1075,871,897]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="28" type="Museum">NHMW</collectionCode>
) has the aedeagus matching the illustration in the original description (including the uniquely shaped parameral apices). Fort-Dauphin indicated in label data is a French name for the Malagasy city Tolagnaro
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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