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119 lines
17 KiB
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<document id="5E95528AC4F13DC26F5EC47C2E08F18B" ID-CLB-Dataset="6402" ID-DOI="10.5252/g2013n4a5" ID-GBIF-Dataset="fbfe438c-1eae-40cb-b56f-ff8167b283c2" ID-ISSN="1638-9395" ID-Zenodo-Dep="4538200" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613149818085" checkinUser="felipe" docAuthor="Boessenecker, Robert W." docDate="2013" docId="03E7DD69FF998617244A78A8FDCD3E5E" docLanguage="en" docName="Geodiversitas.35.4.815-940.pdf" docOrigin="Geodiversitas 35 (4)" docSource="http://www.bioone.org/doi/abs/10.5252/g2013n4a5" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Phocoenidae Gray 1825" docType="treatment" docVersion="9" lastPageNumber="889" masterDocId="FFDEA511FFD0865D27477C65FF9B3B38" masterDocTitle="A new marine vertebrate assemblage from the Late Neogene Purisima Formation in Central California, part II: Pinnipeds and Cetaceans" masterLastPageNumber="940" masterPageNumber="815" pageNumber="888" updateTime="1698929391291" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="CA9F159A8235869D5B2D3723DE34EB70">
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<mods:title id="6773A3895CA3236D175DCD1C014B0E75">A new marine vertebrate assemblage from the Late Neogene Purisima Formation in Central California, part II: Pinnipeds and Cetaceans</mods:title>
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<mods:name id="A3F9118F80DA2976493C4629D4B89969" type="personal">
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<mods:namePart id="4C770C7A3726D9A4078DE7C052386AA1">Boessenecker, Robert W.</mods:namePart>
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<mods:typeOfResource id="A3DCCCC1091531A47E159DC94A51227E">text</mods:typeOfResource>
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<mods:title id="F703D2A273A9DBFAEE6E3A28F985DB58">Geodiversitas</mods:title>
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<mods:part id="EC8868F24FCD16B03F84EFD2704802B3">
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<mods:date id="C42C238501F5362E99C8601D0160E8C3">2013</mods:date>
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<mods:detail id="C7B77279DA9D42B0D0E2D3B68D58818A" type="pubDate">
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<mods:number id="BC96E7D450075E51A02C5C0DD8DED63D">2013-12-31</mods:number>
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<mods:number id="4FE9946DD7C49EB0E7C737080D69B826">35</mods:number>
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<mods:identifier id="2710B9527C791094108BB8C32929263F" type="DOI">10.5252/g2013n4a5</mods:identifier>
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<treatment id="03E7DD69FF998617244A78A8FDCD3E5E" ID-DOI="http://doi.org/10.5281/zenodo.4818517" ID-GBIF-Taxon="182400961" ID-Zenodo-Dep="4818517" LSID="urn:lsid:plazi:treatment:03E7DD69FF998617244A78A8FDCD3E5E" httpUri="http://treatment.plazi.org/id/03E7DD69FF998617244A78A8FDCD3E5E" lastPageId="74" lastPageNumber="889" pageId="73" pageNumber="888">
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<subSubSection id="C3543FF4FF998614244A78A8FC013E6C" pageId="73" pageNumber="888" type="nomenclature">
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<paragraph id="8BF16C7FFF998614244A78A8FBC93FDF" blockId="73.[781,1106,1229,1256]" box="[781,1106,1229,1256]" pageId="73" pageNumber="888">
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<heading id="D0B9DB13FF998614244A78A8FBC93FDF" box="[781,1106,1229,1256]" centered="true" fontSize="11" level="2" pageId="73" pageNumber="888" reason="2">
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<taxonomicName id="4C4E17FCFF998614244A78A8FC0C3FDF" ID-CoL="6269L" authorityName="Gray" authorityYear="1825" box="[781,919,1229,1255]" class="Mammalia" family="Phocoenidae" kingdom="Animalia" order="Cetacea" pageId="73" pageNumber="888" phylum="Chordata" rank="family">Phocoenidae</taxonomicName>
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gen. et sp. indet.
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</heading>
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</paragraph>
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<paragraph id="8BF16C7FFF99861425E27962FC013E6C" blockId="73.[675,1209,1286,1364]" pageId="73" pageNumber="888">
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REFERRED MATERIAL. — UCMP 219176, an isolated left tympanic bulla collected by R. W. Boessenecker from UCMP locality
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<accessionNumber id="941DF19CFF99861424047959FC0D3E6C" box="[835,918,1340,1364]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V99847" pageId="73" pageNumber="888" type="EnaNcbi">V99847</accessionNumber>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3543FF4FF99861725E27909FDF23917" lastPageId="74" lastPageNumber="889" pageId="73" pageNumber="888" type="description">
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<paragraph id="8BF16C7FFF99861425E27909FD713EEB" blockId="73.[675,1210,1387,1491]" pageId="73" pageNumber="888">
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STRATIGRAPHIC OCCURRENCE. — Middle part of the San Gregorio section of the Purisima Formation, Early-Late Pliocene (3.35-2.5 Ma; Piacenzian-Gelasian equivalent;
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<figureCitation id="137570FAFF99861425E479DFFD463EEB" box="[675,733,1466,1491]" captionStart="FIG" captionStartId="6.[113,124,1221,1239]" captionTargetBox="[761,1205,215,1066]" captionTargetId="figure-149@6.[759,1205,592,972]" captionTargetPageId="6" captionText="FIG. 2. — Stratigraphy of the Purisima Formation: A, Stratigraphic column of the San Gregorio section of the Purisima Formation (in meters), showing stratigraphic position of UCMP vertebrate localities, dated ash beds, diatom floras, and species of Carcharodon; thickness measurements start over above missing section in uppermost part due lack of overlapping strata; epoch and stage boundaries marked as "?" are estimated; modified from Boessenecker (2011a); Pliocene-Pleistocene boundary after Hilgen et al. (2012); B, photograph of 3-4 m thick ash within Purisima Formation at mouth of San Gregorio Creek correlated with 2.5 Ma Ishi Tuff (Powell et al.2007); C, trace fossils in pervasively bioturbated sandstone,Purisima Formation,mouth of San Gregorio Creek; D, swaley cross-stratified sandstone, mouth of San Gregorio Creek. Abbreviations: Qt, Pleistocene terrace deposits; Tp, Purisima Formation; C, claystone; St, siltstone; Ss, sandstone. Scale bars: A, 2 m; B, C, 10 cm." figureDoi="http://doi.org/10.5281/zenodo.4538204" httpUri="https://zenodo.org/record/4538204/files/figure.png" pageId="73" pageNumber="888">Fig. 2</figureCitation>
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).
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</paragraph>
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<paragraph id="8BF16C7FFF99861425E2798EFCAE3D3B" blockId="73.[675,1210,1515,1699]" box="[677,821,1515,1541]" pageId="73" pageNumber="888">DESCRIPTION</paragraph>
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<paragraph id="8BF16C7FFF99861725E27A6EFDF23917" blockId="73.[675,1210,1515,1699]" lastBlockId="74.[111,649,216,559]" lastPageId="74" lastPageNumber="889" pageId="73" pageNumber="888">
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Ŋe tympanic bulla is small (
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<tableCitation id="C6CC59C4FF99861424A27A6FFBD23D1D" box="[997,1097,1546,1573]" captionStart="TABLE" captionStartId="58.[161,172,1180,1198]" captionTargetPageId="58" captionText="TABLE 12. — Measurements (in mm) of odontocete tympanic bullae. Measurements given to nearest tenth of a millimeter." httpUri="http://table.plazi.org/id/DF313CF7FFEA866727E678F9FB173F96" pageId="73" pageNumber="888" tableUuid="DF313CF7FFEA866727E678F9FB173F96">Table 12</tableCitation>
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), and rectangular in dorsal aspect (
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<figureCitation id="137570FAFF99861424F57A4FFBCD3D7D" box="[946,1110,1578,1605]" captionStart="FIG" captionStartId="72.[113,124,1231,1249]" captionTargetBox="[124,1187,216,1190]" captionTargetId="figure-147@72.[570,1028,569,928]" captionTargetPageId="72" captionText="FIG. 39. — Tympanic bullae of delphinoid odontocetes; A, UCMP 219176, Phocoenidae gen. et sp. indet. in dorsal view; B, UCMP 219174,Delphinidae gen.et sp.indet.in dorsal view;C, UCMP 219034, cf. Globicephalinae gen. et sp. indet. in dorsal view; D, UCMP 219176 in medial view; E, UCMP 219174 in medial view; F, UCMP 219034 in medial view; H, UCMP 219176 in ventral view; I, UCMP 219174 in ventral view; J, UCMP 219034 in ventral view; K, comparison of interpretive line drawings of extant delphinids; specimen numbers for extant specimens are as follows: Globicephala macrorhynchus Gray, 1846, CAS 16466; Grampus griseus Cuvier, 1812, CAS 13461; Orcinus orca Linnaeus, 1758, CAS 23814; Pseudorca crassidens Owen, 1846, AMNH 18965; Tursiops truncatus Gray, 1866, CAS 16183). Scale bars: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4538293" httpUri="https://zenodo.org/record/4538293/files/figure.png" pageId="73" pageNumber="888">Fig. 39A, D, H</figureCitation>
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). UCMP 219176 has a small, equidimensional and pachyostotic posterior processes, with slightly convex and nearly smooth posterior bullar facet with faint lon- gitudinal striations. Ŋe inner and outer posterior prominences are similar in transverse width, but the outer posterior prominence is positioned slightly further posterior to the inner posterior prominence (
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<figureCitation id="137570FAFF9A861727307D33FF7B3A49" box="[119,224,342,369]" captionStart="FIG" captionStartId="72.[113,124,1231,1249]" captionTargetBox="[124,1187,216,1190]" captionTargetId="figure-147@72.[570,1028,569,928]" captionTargetPageId="72" captionText="FIG. 39. — Tympanic bullae of delphinoid odontocetes; A, UCMP 219176, Phocoenidae gen. et sp. indet. in dorsal view; B, UCMP 219174,Delphinidae gen.et sp.indet.in dorsal view;C, UCMP 219034, cf. Globicephalinae gen. et sp. indet. in dorsal view; D, UCMP 219176 in medial view; E, UCMP 219174 in medial view; F, UCMP 219034 in medial view; H, UCMP 219176 in ventral view; I, UCMP 219174 in ventral view; J, UCMP 219034 in ventral view; K, comparison of interpretive line drawings of extant delphinids; specimen numbers for extant specimens are as follows: Globicephala macrorhynchus Gray, 1846, CAS 16466; Grampus griseus Cuvier, 1812, CAS 13461; Orcinus orca Linnaeus, 1758, CAS 23814; Pseudorca crassidens Owen, 1846, AMNH 18965; Tursiops truncatus Gray, 1866, CAS 16183). Scale bars: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4538293" httpUri="https://zenodo.org/record/4538293/files/figure.png" pageId="74" pageNumber="889">Fig. 39H</figureCitation>
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). Ŋe involucrum of UCMP 219176 is gracile and lacks a strongly inflated anterior portion. In dorsal aspect, the inner posterior prominence of UCMP 219174 forms a more acute angle at its apex than in 219176. Although the outer lip is damaged, a slight inflection at the margin of the broken portion suggests it had a lateral furrow.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3543FF4FF9A861727367E30FDCD3E5E" pageId="74" pageNumber="889" type="discussion">
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<paragraph id="8BF16C7FFF9A861727367E30FE3C3955" blockId="74.[112,648,597,1383]" box="[113,423,597,623]" pageId="74" pageNumber="889">REMARKS AND COMPARISONS</paragraph>
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<caption id="DF313CF7FF9A861725E27E36FC733991" ID-DOI="http://doi.org/10.5281/zenodo.4538295" ID-Zenodo-Dep="4538295" httpUri="https://zenodo.org/record/4538295/files/figure.png" pageId="74" pageNumber="889" startId="74.[677,688,595,613]" targetBox="[687,1204,216,553]" targetPageId="74">
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<paragraph id="8BF16C7FFF9A861725E27E36FC733991" blockId="74.[676,1211,595,681]" pageId="74" pageNumber="889">
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FIG. 40.— Partial tooth of cf.
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<taxonomicName id="4C4E17FCFF9A861724D77E36FB8D395D" authorityName="Gray" authorityYear="1866" box="[912,1046,595,613]" class="Mammalia" family="Delphinidae" kingdom="Animalia" order="Cetacea" pageId="74" pageNumber="889" phylum="Chordata" rank="subFamily" subFamily="Globicephalinae">Globicephalinae</taxonomicName>
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gen.et sp.indet.in:
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<emphasis id="B93AB06DFF9A861725E27E0FFD2A3944" bold="true" box="[677,689,618,636]" pageId="74" pageNumber="889">A</emphasis>
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, anterior or posterior view (UCMP 219162);
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<emphasis id="B93AB06DFF9A8617234F7E0FFB8E3944" bold="true" box="[1032,1045,618,636]" pageId="74" pageNumber="889">B</emphasis>
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, anterior or posterior view (UCMP 219162);
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<emphasis id="B93AB06DFF9A861724277EE4FCF539AB" bold="true" box="[864,878,641,659]" pageId="74" pageNumber="889">C</emphasis>
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, lingual view (UCMP 219162);
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<emphasis id="B93AB06DFF9A861723287EE4FBE639AB" bold="true" box="[1135,1149,641,659]" pageId="74" pageNumber="889">D</emphasis>
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, apical view (UCMP 219162). Scale bar: 1 cm.
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</paragraph>
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</caption>
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<paragraph id="8BF16C7FFF9A861727367E11FDCD3E5E" blockId="74.[112,648,597,1383]" pageId="74" pageNumber="889">
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UCMP 219176 exhibits one synapomorphy of
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<taxonomicName id="4C4E17FCFF9A861727347EF6FF773995" authorityName="Muizon" authorityYear="1988" box="[115,236,659,685]" infraOrder="Delphinida" pageId="74" pageNumber="872" rank="infraOrder">Delphinida</taxonomicName>
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, an anteriorly inflated involucrum (
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<bibRefCitation id="EFDF118EFF9A861725157EF6FF7339F5" author="MUIZON C. DE" pageId="74" pageNumber="889" pagination="159 - 227" refId="ref72620" refString="MUIZON C. DE 1988 a. - Les relations phylogenetiques des Delphinida (Cetacea, Mammalia). Annales de Paleontologie 74: 159 - 227." type="journal article" year="1988">Muizon 1988a</bibRefCitation>
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). Furthermore, this specimen exhibits two phocoenid synapomorphies: a short posterior process with a smooth articular surface (
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<bibRefCitation id="EFDF118EFF9A861725687E97FF353814" author="MUIZON C. DE" pageId="74" pageNumber="889" pagination="1 - 188" refId="ref72574" refString="MUIZON C. DE 1984. - Les vertebres fossiles de la Formation Pisco (Perou), Deuxieme partie: Les odontocetes (Cetacea, Mammalia) du Pliocene inferieur de Sud-Sacaco. Travaux de l'Institut francais d'Etudes Andines 50: 1 - 188." type="journal article" year="1984">Muizon 1984</bibRefCitation>
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), and a pachyostotic posterior process (
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<bibRefCitation id="EFDF118EFF9A861725177F77FF773874" author="MUIZON C. DE" pageId="74" pageNumber="889" pagination="159 - 227" refId="ref72620" refString="MUIZON C. DE 1988 a. - Les relations phylogenetiques des Delphinida (Cetacea, Mammalia). Annales de Paleontologie 74: 159 - 227." type="journal article" year="1988">Muizon 1988a</bibRefCitation>
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). Ŋis specimen differs from
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<taxonomicName id="4C4E17FCFF9A8617257D7F57FF513853" authorityName="Almejas Fm. Barnes" authorityYear="1984" class="Mammalia" family="Albireonidae" genus="Albireo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="74" pageNumber="889" phylum="Chordata" rank="species" species="whistleri">
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<emphasis id="B93AB06DFF9A8617257D7F57FF513853" italics="true" pageId="74" pageNumber="889">Albireo whistleri</emphasis>
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</taxonomicName>
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in its smaller size, lacking a transversely massive involucrum with a convex medial margin in dorsal view, and exhibiting an outer posterior prominence that extends further posteriorly than the inner posterior prominence. UCMP 219176 differs from
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<taxonomicName id="4C4E17FCFF9A861727B37F95FEE23F32" authorityName="Gray" authorityYear="1821" box="[244,377,1008,1034]" class="Mammalia" family="Delphinidae" kingdom="Animalia" order="Cetacea" pageId="74" pageNumber="889" phylum="Chordata" rank="family">Delphinidae</taxonomicName>
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gen. et sp. indet. (UCMP 219174) described above in exhibiting a ventral crest on the medial lobe that extends further anteriorly, and in having a larger posterior process with a smooth posterior bullar facet; in UCMP 219174, the facet is more strongly striated. Ŋis specimen is identical to extant phocoenids and an undescribed fossil phocoenid (SDNHM 65276) from the San Diego Formation (
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<bibRefCitation id="EFDF118EFF9A8617267A7888FD943E3F" author="RACICOT R. & DEMERE T. A. & ROWE T." box="[317,527,1261,1287]" pageId="74" pageNumber="889" pagination="132" refId="ref74632" refString="RACICOT R., DEMERE T. A. & ROWE T. 2007. - Morphology of a bizarre new fossil porpoise (Cetacea: Phocoenidae) from the Pliocene San Diego Formation of southern California. Journal of Vertebrate Paleontology 27 (3, supplement): 132 A." type="journal article" year="2007">
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Racicot
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<emphasis id="B93AB06DFF9A861726DF788BFE573E3F" box="[408,460,1261,1287]" italics="true" pageId="74" pageNumber="889">et al.</emphasis>
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2007
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</bibRefCitation>
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; this taxon is equivalent to
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<taxonomicName id="4C4E17FCFF9A8617266F7968FE2E3E1F" authorityName="Gray" authorityYear="1825" box="[296,437,1293,1319]" class="Mammalia" family="Phocoenidae" kingdom="Animalia" order="Cetacea" pageId="74" pageNumber="889" phylum="Chordata" rank="family">Phocoenidae</taxonomicName>
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unnamed genus 2 described below), and may be synonymous with any of the other phocoenids reported herein.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |