253 lines
26 KiB
XML
253 lines
26 KiB
XML
<document id="41F73BD53CEB88F937285E0AFF31EB6A" ID-DOI="10.11646/zootaxa.3948.1.2" ID-GBIF-Dataset="9e109059-3d75-4e2c-997e-6f4217d201d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244003" ID-ZooBank="9A0330AD-EEF7-4A80-8D80-1B6116BD856F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461042933937" checkinUser="plazi" docAuthor="Filippini, Valentina, Galante, Eduardo & Micó, Estefanía" docDate="2015" docId="03E1981EB53BFFEFFF59FCAD00E4EC88" docLanguage="en" docName="zt03948p040.pdf" docOrigin="Zootaxa 3948 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Callistethus ruteloides Filippini, Galante" docType="treatment" docVersion="7" masterDocId="FFD8E066B530FFE2FFCEFFD3014CEC5D" masterDocTitle="Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)" updateTime="1698576078296" updateUser="plazi">
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<mods:mods id="3BCDA1DF6B7E5001E072C7E4994DCD67" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A6E617CAEB9D00146BC9C7A221989CD6">
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<mods:title id="123A1F6E4F4AD9FB36CE3679E5FB20E4">Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="3E59FA5F118CCAE7822D7C413749E408" type="personal">
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<mods:role id="6951302AEF40ADBA68C5CF6D911D497E">
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<mods:roleTerm id="FEE55B5E86CAEB218F36A81A41CD14D7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="871A59BC7AF75089D8B3371B6ECD4205">Filippini, Valentina</mods:namePart>
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</mods:name>
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<mods:name id="C2B6A6F209D4B92421477CA577DC0A0D" type="personal">
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<mods:role id="709C7380A5749865FD36C84BEC8D6922">
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<mods:roleTerm id="EA38DBA921D831B5242C224D6111BACB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A05B7C99787C4D427E515C8770500948">Galante, Eduardo</mods:namePart>
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</mods:name>
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<mods:name id="85328817F73C8626C0824F7522B87D78" type="personal">
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<mods:role id="9B4E616BED88E25324471615C24FF4DC">
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<mods:roleTerm id="34D1C061903340FA0F41F1A9DDEF2376">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E9435176D93D50566D25DC97DEB8CFB3">Micó, Estefanía</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="EBA7CCAF9E6728181686B9ED1C20A696">text</mods:typeOfResource>
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<mods:relatedItem id="35BF869AF68359E2DE027D4ACC2EDEC6" type="host">
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<mods:titleInfo id="F0A8917D4AB6457AC48D9D2D86C86151">
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<mods:title id="9E469EBA1E2EC2BB64F59FEE541D772C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CA0CF2D4AC03BF61DDDE99DB67980ACC">
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<mods:date id="7328E8C4F837432C60BF014EB9E56561">2015</mods:date>
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<mods:detail id="F52FB362D61E691113B353A5ADEAEF95" type="volume">
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<mods:number id="EFB04623B391405F2322A471CDD4E5C6">3948</mods:number>
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</mods:detail>
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<mods:detail id="9062907BADBE3511B789565A7682202E" type="issue">
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<mods:number id="AED2818B918492C002F5EB6CF8F9076A">1</mods:number>
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</mods:detail>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="5BB29BFCEE8CF1E4C0EF10EC52AE8E53">journal volume</mods:classification>
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<mods:identifier id="E14C3C2F51ADE35B52E9C8D41EC89219" type="DOI">10.11646/zootaxa.3948.1.2</mods:identifier>
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<mods:identifier id="B59F15131046FA591A80D998CBB2520E" type="GBIF-Dataset">9e109059-3d75-4e2c-997e-6f4217d201d0</mods:identifier>
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<mods:identifier id="8A37D25CCB88694CD1FF7ADC7194C8A7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="78C940E6F7F7C24D6B198C33DF8E8F83" type="Zenodo-Dep">244003</mods:identifier>
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<mods:identifier id="4711D874428C10226AB5811296D8A08D" type="ZooBank">9A0330AD-EEF7-4A80-8D80-1B6116BD856F</mods:identifier>
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</mods:mods>
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<treatment id="03E1981EB53BFFEFFF59FCAD00E4EC88" ID-DOI="http://doi.org/10.5281/zenodo.6122708" ID-GBIF-Taxon="119609988" ID-Zenodo-Dep="6122708" LSID="urn:lsid:plazi:treatment:03E1981EB53BFFEFFF59FCAD00E4EC88" httpUri="http://treatment.plazi.org/id/03E1981EB53BFFEFFF59FCAD00E4EC88" lastPageId="13" pageId="11">
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<subSubSection id="C3527A83B53BFFE9FF59FCAD0195EFE7" pageId="11" type="nomenclature">
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<paragraph id="8BF72908B53BFFE9FF59FCAD02D9EFC5" blockId="11.[151,917,894,954]" box="[151,917,894,920]" pageId="11">
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<heading id="D0BF9E64B53BFFE9FF59FCAD02D9EFC5" bold="true" box="[151,917,894,920]" fontSize="11" level="1" pageId="11" reason="1">
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<emphasis id="B93CF51AB53BFFE9FF59FCAD02D9EFC5" bold="true" box="[151,917,894,920]" pageId="11">
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<taxonomicName id="4C48528BB53BFFE9FF59FCAD03C7EFC5" authority="Filippini, Galante" authorityName="Filippini, Galante" box="[151,651,894,920]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="ruteloides">
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<emphasis id="B93CF51AB53BFFE9FF59FCAD00E9EFC5" bold="true" box="[151,421,894,920]" italics="true" pageId="11">Callistethus ruteloides</emphasis>
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<collectingCountry id="F35F6998B53BFFE9FE62FCAD0355EFC5" box="[428,537,894,920]" name="Philippines" pageId="11">Filippini</collectingCountry>
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, Galante
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</taxonomicName>
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, & Micó, new species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF72908B53BFFE9FF59FC720195EFE7" blockId="11.[151,917,894,954]" box="[151,217,929,954]" pageId="11">
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<figureCitation id="1373358DB53BFFE9FF59FC720195EFE7" box="[151,217,929,954]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[186,1400,705,1952]" captionTargetId="figure@4.[179,1408,697,1957]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Habitus. 1: Anomala aglaos (holotype). 2: A. estrella (holotype). 3: A. inbio (holotype). 4: A. pincelada (holotype). 5: Callistethus ruteloides (holotype). 6: C. yalizo (holotype). Scale = 5 mm." httpUri="https://zenodo.org/record/244004/files/figure.png" pageId="11">Fig. 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3527A83B53BFFEEFF59FC3A0234EDA8" lastPageId="12" pageId="11" type="materials_examined">
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<paragraph id="8BF72908B53BFFE9FF59FC3A0285E87B" blockId="11.[151,1437,1001,2037]" pageId="11">
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<emphasis id="B93CF51AB53BFFE9FF59FC3A0009E85F" bold="true" box="[151,325,1001,1026]" pageId="11">
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<typeStatus id="54F397AAB53BFFE9FF59FC3A019EE85F" box="[151,210,1001,1026]" pageId="11">Type</typeStatus>
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material.
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||
</emphasis>
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||
<typeStatus id="54F397AAB53BFFE9FE82FC3A00F5E85F" box="[332,441,1001,1026]" pageId="11" type="holotype">Holotype</typeStatus>
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: ♂ "Quebrada Segunda, P. N. Tapanti,
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<quantity id="4CB084EDB53BFFE9FCA5FC3A028EE85C" box="[875,962,1001,1026]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="11" unit="m" value="1250.0">1250 m</quantity>
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, Prov. Cartago,
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<collectingCountry id="F35F6998B53BFFE9FBB9FC3905B8E85F" box="[1143,1268,1001,1026]" name="Costa Rica" pageId="11">Costa Rica</collectingCountry>
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, G. Mora,
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<date id="FFF60FC8B53BFFE9FAA7FC390183E87B" pageId="11" value="1992-05">May 1992</date>
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L-N 194000_560000 / INBIOCRI000959028" deposited in
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<taxonomicName id="4C48528BB53BFFE9FCBDFBDD0288E87B" box="[883,964,1038,1062]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="inbio">
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<collectionCode id="ED59B1CDB53BFFE9FCBDFBDD0288E87B" box="[883,964,1038,1062]" pageId="11">INBIO</collectionCode>
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||
</taxonomicName>
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||
.
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</paragraph>
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<paragraph id="8BF72908B53BFFE9FF09FBE103D4E937" blockId="11.[151,1437,1001,2037]" pageId="11">
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<typeStatus id="54F397AAB53BFFE9FF09FBE10078E817" box="[199,308,1074,1098]" pageId="11" type="paratype">Paratypes</typeStatus>
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(6):
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<specimenCount id="9D4EE281B53BFFE9FEA5FBE100C2E817" box="[363,398,1074,1098]" pageId="11" type="generic">1♂</specimenCount>
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"Est. Biol. Las Alturas,
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<quantity id="4CB084EDB53BFFE9FD6FFBE203B5E814" box="[673,761,1073,1098]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" unit="m" value="1500.0">1500 m</quantity>
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, Coto Brus, Prov. Puntarenas,
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<collectingCountry id="F35F6998B53BFFE9FB9EFBE10582E817" box="[1104,1230,1073,1098]" name="Costa Rica" pageId="11">Costa Rica</collectingCountry>
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. M. Ramirez.
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<date id="FFF60FC8B53BFFE9FAA3FBE10183E833" pageId="11" value="1992-03">Mar 1992</date>
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L-S 322500, 591300 / INBIOCRI000987456"; 1♀ "Tapantí (
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<collectingCountry id="F35F6998B53BFFE9FC4CFB85054CE833" box="[898,1024,1109,1134]" name="Costa Rica" pageId="11">Costa Rica</collectingCountry>
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)
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<quantity id="4CB084EDB53BFFE9FBC1FB850529E830" box="[1039,1125,1110,1134]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="11" unit="m" value="1300.0">1300 m</quantity>
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<date id="FFF60FC8B53BFFE9FBA2FB860589E833" box="[1132,1221,1109,1134]" pageId="11" value="2006-09-29">29-9-06</date>
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T. luz 17:00-23:00 Leg.: Micó, García, Galante /
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<collectionCode id="ED59B1CDB53BFFE9FE3AFBAA030FE8CF" box="[500,579,1145,1170]" pageId="11">CEUA</collectionCode>
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00019649"; 1♀ "El Copal, Prov. Cartago,
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<collectingCountry id="F35F6998B53BFFE9FBE7FBA905E6E8CF" box="[1065,1194,1145,1170]" name="Costa Rica" pageId="11">Costa Rica</collectingCountry>
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.
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<date id="FFF60FC8B53BFFE9FB79FBAA047AE8CF" box="[1207,1334,1145,1170]" pageId="11" value="2007-06-17">17/06/2007</date>
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Leg. M. Moraga /
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<collectionCode id="ED59B1CDB53BFFE9FEC2FB4E0017E8EB" box="[268,347,1181,1206]" pageId="11">CEUA</collectionCode>
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00106211"; 1♀ "
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||
<collectingCountry id="F35F6998B53BFFE9FDD3FB4D0380E8EB" box="[541,716,1182,1206]" name="Costa Rica" pageId="11">COSTA RICA</collectingCountry>
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. Prov. Alajuela. Upala. P.N. Volcán Tenorio. Alb. Heliconias. Send. a Laguna Danta.
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<specimenCount id="9D4EE281B53BFFE9FE76FB110342E884" box="[440,526,1218,1242]" pageId="11" type="generic">
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||
<quantity id="4CB084EDB53BFFE9FE76FB110345E884" box="[440,521,1218,1242]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="11" unit="m" value="900.0">900 m</quantity>
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.
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||
</specimenCount>
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||
<date id="FFF60FC8B53BFFE9FDD5FB12038FE887" box="[539,707,1217,1242]" pageId="11" value="2007-06-16">16 JUN 2007</date>
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||
. J. D. Gutiérrez. Tp. Luz 2. L_N_299100_424000 #92208 /
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<collectionCode id="ED59B1CDB53BFFE9FF59FB3601AAE8A3" box="[151,230,1253,1278]" pageId="11">CEUA</collectionCode>
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00106212"; 1♀ "Q. Segunda, P.N. Tapantí, Prov. Carta,
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<collectingCountry id="F35F6998B53BFFE9FCA4FB350559E8A3" box="[874,1045,1254,1278]" name="Costa Rica" pageId="11">COSTA RICA</collectingCountry>
|
||
.
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||
<quantity id="4CB084EDB53BFFE9FBEFFB350537E8A0" box="[1057,1147,1254,1278]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="11" unit="m" value="1300.0">1300 m</quantity>
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.
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<date id="FFF60FC8B53BFFE9FB47FB3505BDE8A3" box="[1161,1265,1254,1278]" pageId="11" value="1993-09">Set 1993</date>
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. G. Mora, L N 194000_559800 #2326 / INBIOCRI001669692"; 1♀ "
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<collectingCountry id="F35F6998B53BFFE9FCE4FAD902E8E97F" box="[810,932,1290,1314]" name="Panama" pageId="11">PANAMA</collectingCountry>
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- CHIRIQUI Santa Clara env.
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<quantity id="4CB084EDB53BFFE9FAF3FADA04D7E97C" box="[1341,1435,1289,1314]" metricMagnitude="3" metricUnit="m" metricValue="1.5459999999999998" pageId="11" unit="m" value="1546.0">1546 m</quantity>
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||
<geoCoordinate id="EE7C4FCFB53BFFE9FF59FAFE007FE91B" box="[151,307,1325,1350]" direction="north" orientation="latitude" pageId="11" precision="1" value="8.861723">08°51'42,2"N</geoCoordinate>
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||
;
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||
<geoCoordinate id="EE7C4FCFB53BFFE9FEF0FAFE00A7E91B" box="[318,491,1325,1350]" direction="west" orientation="longitude" pageId="11" precision="1" value="-82.74347">
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||
082°44'36,5"
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||
<geoCoordinate id="EE7C4FCFB53BFFE9FE01FAFE00A7E91B" box="[463,491,1325,1350]" direction="west" orientation="longitude" pageId="11" precision="5555" value="-17.6">W</geoCoordinate>
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</geoCoordinate>
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17.6.-
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<date id="FFF60FC8B53BFFE9FDE1FAFE0330E91B" box="[559,636,1325,1350]" pageId="11" value="06-07-04">4.7.06</date>
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||
; V.
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<collectingCountry id="F35F6998B53BFFE9FD51FAFE0397E91B" box="[671,731,1325,1350]" name="Serbia" pageId="11">Malý</collectingCountry>
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||
lgt. P 7 / coll. Vladislv
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<collectingCountry id="F35F6998B53BFFE9FC2CFAFE0551E91B" box="[994,1053,1325,1350]" name="Serbia" pageId="11">Malý</collectingCountry>
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<collectingCountry id="F35F6998B53BFFE9FBEAFAFE0525E91B" box="[1060,1129,1325,1350]" name="Czech Republic" pageId="11">Česká</collectingCountry>
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Rep. Praha" Vladislv
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<collectingCountry id="F35F6998B53BFFE9FAAFFAFE04D0E91B" box="[1377,1436,1325,1350]" name="Serbia" pageId="11">Malý</collectingCountry>
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||
(
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||
<collectingCountry id="F35F6998B53BFFE9FF51FA8201A4E937" box="[159,232,1361,1386]" name="Czech Republic" pageId="11">Česká</collectingCountry>
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,
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<collectingCountry id="F35F6998B53BFFE9FF3DFA8200E5E937" box="[243,425,1361,1386]" name="Czech Republic" pageId="11">Czech Republic</collectingCountry>
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), personal collection.
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||
</paragraph>
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<paragraph id="8BF72908B53BFFE9FF09FAA60505E9D0" blockId="11.[151,1437,1001,2037]" box="[199,1097,1397,1422]" pageId="11">
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||
<emphasis id="B93CF51AB53BFFE9FF09FAA60344E9D3" bold="true" box="[199,520,1397,1422]" pageId="11">
|
||
<typeStatus id="54F397AAB53BFFE9FF09FAA60078E9D3" box="[199,308,1397,1422]" pageId="11" type="holotype">Holotype</typeStatus>
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||
male description.
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||
</emphasis>
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||
Body shape oval. Length
|
||
<quantity id="4CB084EDB53BFFE9FCE0FAA602D9E9D0" box="[814,917,1397,1422]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="11" unit="mm" value="17.5">17.5 mm</quantity>
|
||
. Width
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<quantity id="4CB084EDB53BFFE9FC25FAA50509E9D0" box="[1003,1093,1398,1422]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="11" unit="mm" value="9.8">9.8 mm</quantity>
|
||
.
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||
</paragraph>
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||
<paragraph id="8BF72908B53BFFE9FF09FA4A03BBE9A7" blockId="11.[151,1437,1001,2037]" pageId="11">Color. Head pale ochre with a sinuate brown macula on frons. Pronotum pale ochre with 2 lateral brown bands. Scutellum pale ochre with red margins. Elytra pale ochre with 2 transversal sinuate beige bands, 1 median and 1 apical. Pygidium and legs ochre. Venter orange-ochre.</paragraph>
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||
<paragraph id="8BF72908B53BFFE9FF09F9D6056AEAEC" blockId="11.[151,1437,1001,2037]" pageId="11">Head large, almost 2/3 of the width of pronotum. Clypeus rectangular, with anterior angles widely curved. Clypeus ratio width/length 2.1; surface densely punctate-reticulate, convex; anterior margin straight, nearly as long as frons. Frons moderately punctate, flat. Ocular canthum long, thin, with rounded apex. Interocular ratio (interocular width/width of eye): 3.0. Antenna: ratio funiculus/club 0.6. Mandibles with apex curved ventrally, maxillae with 2nd and 3rd tooth fused together. Labium and labrum quadrilobulate.</paragraph>
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<paragraph id="8BF72908B53BFFE9FF09F96E00CEEB41" blockId="11.[151,1437,1001,2037]" pageId="11">Pronotum trapezoidal. Width 1.9 times length. Lateral margins curved. Anterior angles quadrate with blunt vertex; basal angles obtuse, blunt. Basal margin narrow, sinuate, and obliterated in the middle. Surface with fine and dense punctures.</paragraph>
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<paragraph id="8BF72908B53BFFE9FF09F8FB0518EB1C" blockId="11.[151,1437,1001,2037]" box="[199,1108,1832,1857]" pageId="11">Scutellum. Shape subtriangular, surface sparsely punctate; ratio width/length 1.6.</paragraph>
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<paragraph id="8BF72908B53BFFE9FF09F89F05FEEB39" blockId="11.[151,1437,1001,2037]" box="[199,1202,1868,1893]" pageId="11">Elytra with striae defined by regular rows of punctures. Marginal membrane very narrow.</paragraph>
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<paragraph id="8BF72908B53BFFE9FF09F8A3007BEBF0" blockId="11.[151,1437,1001,2037]" pageId="11">Pygidium strigate. In lateral view slightly convex. Triangular in shape; with short setae on disc, long setae at apical margin.</paragraph>
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<paragraph id="8BF72908B53BFFE9FF09F86B0219EBA9" blockId="11.[151,1437,1001,2037]" pageId="11">
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Mesosternal process long, with tapering apex in ventral view; width at base: 1.0 mm (
|
||
<figureCitation id="1373358DB53BFFE9FAE1F86B04C3EB8C" box="[1327,1423,1976,2001]" captionStart="FIGURES 25 – 26" captionStartId="13.[151,264,1743,1766]" captionTargetBox="[263,1342,924,1701]" captionTargetId="figure@13.[227,1360,911,1721]" captionTargetPageId="13" captionText="FIGURES 25 – 26. Mesometasternal process in 25: Callistethus ruteloides (holotype). 26: C. yalizo (paratype, Río Grande de Orosí, Cartago, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/244008/files/figure.png" pageId="11">Fig. 25</figureCitation>
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). Mesometasternal suture at about half the length of the process.
|
||
</paragraph>
|
||
<paragraph id="8BF72908B53CFFEEFF09FF4402B7EC88" blockId="12.[151,1437,151,717]" pageId="12">Abdominal sternites with 1–3 rows of setae, denser at sides, and 4–6 rows of punctures per sternite. Terminal sternite strigate, with apical margin thick and sinuate; apical membrane wide.</paragraph>
|
||
<paragraph id="8BF72908B53CFFEEFF09FF330373EDD5" blockId="12.[151,1437,151,717]" pageId="12">Legs. Protibia bidentate (Fig. 11); apical tooth long and straight; basal tooth below internal apex of protibia triangular in shape, obtuse. Metatibia slender, slightly narrower subapically. Ratio length/width 4.1. First external carina well developed. Surface rugose. Protarsomere 1 equal in length to protarsomeres 2–4, tarsomeres stout. Protarsal claws: external claw curved, internal claw bifurcate, upper branch equal in length and 2/3 the width of the lower branch, inferior margin sinuate.</paragraph>
|
||
<paragraph id="8BF72908B53CFFEEFF09FE470234EDA8" blockId="12.[151,1437,151,717]" pageId="12">
|
||
Aedeagus (
|
||
<figureCitation id="1373358DB53CFFEEFE88FE4700D6EDF1" box="[326,410,404,429]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,264,1885,1908]" captionTargetBox="[153,1429,1020,1837]" captionTargetId="figure@6.[151,1436,1010,1864]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Shape of aedeagus, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 13: Anomala aglaos (paratype, Isla Bonita, Costa Rica). 14: A. estrella (holotype). 15: A. inbio (holotype). 16: A. pincelada (holotype). 17: Callistethus ruteloides (holotype). 18: C. yalizo (Cerro Chompipe, Heredia). Scale = 1 mm." httpUri="https://zenodo.org/record/244005/files/figure.png" pageId="12">Fig. 17</figureCitation>
|
||
): parameres with a narrow and acute apex in a lateral view, pointing downwards, flattened dorsoventrally; apical portion of ventral plate membranous. Endophallus (
|
||
<figureCitation id="1373358DB53CFFEEFC16FE640566ED8D" box="[984,1066,439,464]" captionStart="FIGURES 23 – 24" captionStartId="12.[151,264,1577,1600]" captionTargetBox="[158,1428,776,1549]" captionTargetId="figure@12.[151,1436,769,1557]" captionTargetPageId="12" captionText="FIGURES 23 – 24. Endophallus of 23: Callistethus ruteloides (holotype). 24: C. yalizo (holotype). Scale = 1 mm." httpUri="https://zenodo.org/record/244007/files/figure.png" pageId="12">Fig. 23</figureCitation>
|
||
) short, with patches of short and long setae in different positions. Ejaculatory duct opening frontal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3527A83B53CFFEEFF09FE2C0279EE91" pageId="12" type="description">
|
||
<paragraph id="8BF72908B53CFFEEFF09FE2C0461EE02" blockId="12.[151,1437,151,717]" pageId="12">
|
||
<emphasis id="B93CF51AB53CFFEEFF09FE2C0077EE45" bold="true" box="[199,315,511,536]" pageId="12">Variation.</emphasis>
|
||
Background color from pale ochre to orange; transversal bands on elytra more or less evident. Female similar to male, antennal club shorter than in male; protibia with longer and wider apical tooth (Fig. 11); internal protarsal claw narrower with curved inferior margin; terminal sternite with slightly sinuate apex.
|
||
</paragraph>
|
||
<paragraph id="8BF72908B53CFFEEFF09FDBF0279EE91" blockId="12.[151,1437,151,717]" pageId="12">
|
||
Body length
|
||
<quantity id="4CB084EDB53CFFEEFEAEFDBF0345EED9" box="[352,521,620,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" metricValueMax="2.01" metricValueMin="1.75" pageId="12" unit="mm" value="18.8" valueMax="20.1" valueMin="17.5">17.5–20.1 mm</quantity>
|
||
, body width
|
||
<quantity id="4CB084EDB53CFFEEFD69FDBF020EEED9" box="[679,834,620,644]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" metricValueMax="1.14" metricValueMin="0.98" pageId="12" unit="mm" value="10.6" valueMax="11.4" valueMin="9.8">9.8–11.4 mm</quantity>
|
||
. Clypeus w/l: 1.9–2.1. Interocular ratio (interocular width/width of eye): 3.2–4.2. Pronotum w/l: 1.8–1.9. Scutellum w/l: 1.5–1.6. Pygidium w/l: 1.5–1.6. Metatibia w/l: 3.8–4.1. Width of mesosternal process at base: 1.0–
|
||
<quantity id="4CB084EDB53CFFEEFD16FD67027CEE91" box="[728,816,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="12" unit="mm" value="1.2">1.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF377980B53CFFEEFF59F9FA0454EA1D" box="[151,1304,1577,1600]" httpUri="https://zenodo.org/record/244007/files/figure.png" pageId="12" targetBox="[158,1428,776,1549]" targetPageId="12">
|
||
<paragraph id="8BF72908B53CFFEEFF59F9FA0454EA1D" blockId="12.[151,1304,1577,1600]" box="[151,1304,1577,1600]" pageId="12">
|
||
<emphasis id="B93CF51AB53CFFEEFF59F9FA0001EA62" bold="true" box="[151,333,1577,1600]" pageId="12">FIGURES 23–24</emphasis>
|
||
. Endophallus of 23:
|
||
<taxonomicName id="4C48528BB53CFFEEFDE9F9F90247EA62" box="[551,779,1578,1599]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="ruteloides">
|
||
<emphasis id="B93CF51AB53CFFEEFDE9F9F90247EA62" box="[551,779,1578,1599]" italics="true" pageId="12">Callistethus ruteloides</emphasis>
|
||
</taxonomicName>
|
||
(holotype). 24:
|
||
<taxonomicName id="4C48528BB53CFFEEFC60F9F80544EA62" box="[942,1032,1578,1600]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="yalizo">
|
||
<emphasis id="B93CF51AB53CFFEEFC60F9F80544EA62" box="[942,1032,1578,1600]" italics="true" pageId="12">C. yalizo</emphasis>
|
||
</taxonomicName>
|
||
(holotype). Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3527A83B53CFFEEFF09F9BF054FEAF4" pageId="12" type="diagnosis">
|
||
<paragraph id="8BF72908B53CFFEEFF09F9BF054FEAF4" blockId="12.[151,1437,1644,2030]" pageId="12">
|
||
<emphasis id="B93CF51AB53CFFEEFF09F9BF000DEAD8" bold="true" box="[199,321,1644,1669]" pageId="12">Diagnosis.</emphasis>
|
||
Size large, color pale ochre with darker bands on pronotum and elytra, clypeus long, head large, pronotum short, mesosternal process long, parameres flattened dorsoventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3527A83B53CFFEEFF09F966009EEAAC" pageId="12" type="etymology">
|
||
<paragraph id="8BF72908B53CFFEEFF09F966009EEAAC" blockId="12.[151,1437,1644,2030]" pageId="12">
|
||
<emphasis id="B93CF51AB53CFFEEFF09F9660002EA93" bold="true" box="[199,334,1717,1742]" pageId="12">Etymology.</emphasis>
|
||
Latinized from the name of genus
|
||
<taxonomicName id="4C48528BB53CFFEEFD38F9660273EA90" box="[758,831,1717,1741]" class="Insecta" family="Rutelidae" genus="Rutela" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CF51AB53CFFEEFD38F9660273EA90" box="[758,831,1717,1741]" italics="true" pageId="12">Rutela</emphasis>
|
||
</taxonomicName>
|
||
and suffix -oides, for its general resemblance with species of the tribe
|
||
<taxonomicName id="4C48528BB53CFFEEFEBFF90B0083EAAC" box="[369,463,1752,1777]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="tribe" tribe="Rutelini">Rutelini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3527A83B53CFFEFFF09F92E00E4EC88" lastPageId="13" pageId="12" type="distribution">
|
||
<paragraph id="8BF72908B53CFFEEFF09F92E0068EB64" blockId="12.[151,1437,1644,2030]" pageId="12">
|
||
<emphasis id="B93CF51AB53CFFEEFF09F92E002CEB4B" bold="true" box="[199,352,1789,1814]" pageId="12">Distribution.</emphasis>
|
||
Guanacaste and Talamanca mountain ranges,
|
||
<collectingCountry id="F35F6998B53CFFEEFCBEF92E02A7EB4B" box="[880,1003,1789,1814]" name="Costa Rica" pageId="12">Costa Rica</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="F35F6998B53CFFEEFB42F92D05A9EB48" box="[1164,1253,1790,1813]" name="Panama" pageId="12">Panama</collectingCountry>
|
||
from
|
||
<quantity id="4CB084EDB53CFFEEFAE2F92E01E3EB65" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.6" metricValueMin="0.9" pageId="12" unit="m" value="1250.0" valueMax="1600.0" valueMin="900.0">900–1600 m</quantity>
|
||
, (
|
||
<figureCitation id="1373358DB53CFFEEFF0DF8F3005BEB64" box="[195,279,1824,1849]" captionStart="FIGURE 34 – 37" captionStartId="15.[151,250,1947,1969]" captionTargetBox="[158,1427,933,1891]" captionTargetId="figure@15.[151,1436,925,1898]" captionTargetPageId="15" captionText="FIGURE 34 – 37. Distribution of 34: Anomala aglaos and A. estrella. 35: A. inbio and A. pincelada. 36: Callistethus ruteloides. 37: C. yalizo." httpUri="https://zenodo.org/record/244010/files/figure.png" pageId="12">Fig. 36</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF72908B53CFFEFFF09F89600E4EC88" blockId="12.[151,1437,1644,2030]" lastBlockId="13.[151,1436,151,213]" lastPageId="13" pageId="12">
|
||
<emphasis id="B93CF51AB53CFFEEFF09F89601B3EB03" bold="true" box="[199,255,1861,1886]" pageId="12">Note</emphasis>
|
||
. This species is tentatively placed in the genus
|
||
<taxonomicName id="4C48528BB53CFFEEFCD5F89602EDEB00" box="[795,929,1861,1885]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CF51AB53CFFEEFCD5F89602EDEB00" box="[795,929,1861,1885]" italics="true" pageId="12">Callistethus</emphasis>
|
||
</taxonomicName>
|
||
, as it has some distinct features not found in other
|
||
<taxonomicName id="4C48528BB53CFFEEFF19F8BA0011EBDC" box="[215,349,1897,1921]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CF51AB53CFFEEFF19F8BA0011EBDC" box="[215,349,1897,1921]" italics="true" pageId="12">Callistethus</emphasis>
|
||
</taxonomicName>
|
||
species. It could be placed in a separate genus, but the lack of known similar species would make a proper description and diagnosis of the new genus difficult. Singular characters are: long clypeus (nearly as long as frons); head large (2/3 of pronotum width); mandibles with apex curved ventrally; maxillae with second and third tooth fused together; labrum and labium quadrilobulated; pronotum short; tarsomeres short and stout; mesometasternal suture high, nearly at half the length of the mesosternal process; aedeagus with parameres flattened dorsoventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |