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<document id="410871CCB3A88B3A5B8D21B013431F37" ID-CLB-Dataset="3885" ID-DOI="10.5852/ejt.2021.775.1555" ID-GBIF-Dataset="e0c68699-90c8-4b60-8ed4-e0bd6a7d066f" ID-ISSN="2118-9773" ID-Zenodo-Dep="5596674" ID-ZooBank="264FC47B-9D2B-47F2-9461-F3A6CA5EB58B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635170404667" checkinUser="felipe" docAuthor="Haase, Martin, Grego, Jozef, Erőss, Zoltán Péter, Farkas, Roland &amp; Fehér, Zoltán" docDate="2021" docId="03E11505C62BFFD5694C6331FE507297" docLanguage="en" docName="ejt-775_143-184.pdf" docOrigin="European Journal of Taxonomy 775" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hauffenia lozekiana Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér 2021, sp. nov." docType="treatment" docUuid="63FE4F3F-32A5-4F83-869F-C72B6E2D56AE" docUuidSource="ZooBank" docVersion="10" lastPageNumber="166" masterDocId="FFD86D7DC63EFFC26B216053FFE6761D" masterDocTitle="On the origin and diversification of the stygobiotic freshwater snail genus Hauffenia (Caenogastropoda: Hydrobiidae) with special focus on the northern species and the description of two new species" masterLastPageNumber="184" masterPageNumber="143" pageNumber="164" updateTime="1699464333268" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9F8546D5E6D052263FEECC62BD1169AD">On the origin and diversification of the stygobiotic freshwater snail genus Hauffenia (Caenogastropoda: Hydrobiidae) with special focus on the northern species and the description of two new species</mods:title>
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<mods:namePart id="444EC4E8DFB8C925F505D32DE0AE02E4">Haase, Martin</mods:namePart>
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<mods:affiliation id="0065456126BA992D5D8FE869C7FF64B9">Vogelwarte, Zoological Institute and Museum, University of Greifswald, Soldmannstrasse 23, 17489 Greifswald, Germany.</mods:affiliation>
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<mods:namePart id="5508C0F3D1490BB0A7F0A99795B417CB">Grego, Jozef</mods:namePart>
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<mods:affiliation id="CD7C2EA6582D1C85AA8B73EAFFA2F1D6">Horná Mičiná 219, 97401 Banská Bystrica, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="B3794895B59689D37F7BE057FFE4FF39" type="email">jozef.grego@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2E284BD5BDEE55AD082E824D471CC264">Erőss, Zoltán Péter</mods:namePart>
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<mods:affiliation id="C8DC2C70511928E53569F16571A4BA3A">Levendula utca 68 / 4, 2119 Pécel, Hungary.</mods:affiliation>
<mods:nameIdentifier id="CA1FCAD0DB89B8D77ACCF29DC30183B1" type="email">erosspeter@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="B6E2A8B4F485BFACD43FBB74905CC2CE">Farkas, Roland</mods:namePart>
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<mods:affiliation id="21703BA2B635C7886B8502FF8F375719">Bükk National Park Directorate, Sánc utca 6, 3304 Eger, Hungary.</mods:affiliation>
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<mods:namePart id="53AB9DB5B10DB827B70844DAFA83C104">Fehér, Zoltán</mods:namePart>
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<mods:affiliation id="DAAB35EF28201079FD22B2DEC3C74F70">Hungary, Álmos vezér útja 69 / A, 1141 Budapest, Hungary.</mods:affiliation>
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<mods:title id="95DE1E4D660B0F3459E6D9438AECD44C">European Journal of Taxonomy</mods:title>
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<mods:date id="C06A6E55A1A36FE3F534354461381F1E">2021</mods:date>
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<subSubSection id="C352F798C62BFFD7694C6331FC217560" box="[621,967,866,893]" pageId="21" pageNumber="164" type="nomenclature">
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<heading id="D0BF137FC62BFFD7694C6331FC217560" box="[621,967,866,893]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="164" reason="2">
<taxonomicName id="4C48DF90C62BFFD7694C6331FC837560" ID-CoL="8RX4M" authority="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér, 2021" authorityName="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér" authorityYear="2021" box="[621,869,866,893]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="164" phylum="Mollusca" rank="species" species="lozekiana" status="sp. nov.">
<emphasis id="B93C7801C62BFFD7694C6331FC837560" bold="true" box="[621,869,866,893]" italics="true" pageId="21" pageNumber="164">Hauffenia lozekiana</emphasis>
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<taxonomicNameLabel id="A20FC57AC62BFFD7684D6330FC217560" box="[876,967,867,893]" pageId="21" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BF7A413C62BFFD76A5A63FAFB5F75DE" blockId="21.[369,1218,866,964]" box="[379,1209,937,964]" pageId="21" pageNumber="164">
<figureCitation id="1373B896C62BFFD76A5A63FAFE2375DE" box="[379,453,937,963]" captionStart="Fig" captionStartId="8.[189,232,1986,2012]" captionTargetBox="[189,1398,285,1894]" captionTargetId="figure-14@8.[189,1398,285,1927]" captionTargetPageId="8" captionText="Fig. 2. Localities in Slovakia (A) and Hungary (B). Abbreviation: LT = type locality." figureDoi="http://doi.org/10.5281/zenodo.5596680" httpUri="https://zenodo.org/record/5596680/files/figure.png" pageId="21" pageNumber="164">Figs 2</figureCitation>
,
<figureCitation id="1373B896C62BFFD76AF363FAFE0775D9" box="[466,481,937,964]" captionStart="Fig" captionStartId="11.[191,234,1958,1984]" captionTargetBox="[189,1398,1104,1917]" captionTargetId="figure-350@11.[189,1398,1104,1917]" captionTargetPageId="11" captionText="Fig. 5. Type locality of H. lozekiana sp. nov. (71) and Dabarska Pećina, a locality of H. steffeki sp. nov. (72) (see Table 1)." figureDoi="http://doi.org/10.5281/zenodo.5596686" httpUri="https://zenodo.org/record/5596686/files/figure.png" pageId="21" pageNumber="164">5</figureCitation>
71ab, 12AC, 13AB, 14A, 15AC, 17AB, 18;
<tableCitation id="C6CA91A8C62BFFD76F1863FAFB5F75DE" box="[1081,1209,937,963]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="24.[186,256,548,576]" captionStartId-1="28.[189,259,265,291]" captionTargetPageId-0="24" captionTargetPageId-1="28" captionText-0="Table 3 (continued on next page). Diagnostic sites of types 1, 3, and 4 of COI for H. lozekiana sp. nov. and H. steffeki sp. nov. For reference alignment see Supp. file 2." captionText-1="Table 4. Morphometrics of the new species. Abbreviations: AH = aperture height; AW = aperture width; CV = coefficient of variation corrected for unequal sample sizes; max = maximum; min = minimum; SD = standard deviation; SH = shell height; SW = shell width; W = number of whorls (to the nearest eighth). Measurements in mm." httpUri-0="http://table.plazi.org/id/DF37F49BC626FFDA6B9B6277FE937479" httpUri-1="http://table.plazi.org/id/DF37F49BC622FFDE6B9C615AFDC07790" pageId="21" pageNumber="164" tableUuid-0="DF37F49BC626FFDA6B9B6277FE937479" tableUuid-1="DF37F49BC622FFDE6B9C615AFDC07790">Tables 34</tableCitation>
</paragraph>
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<heading id="D0BF137FC62BFFD76B9C63B9FED07219" bold="true" box="[189,310,1002,1028]" fontSize="11" level="3" pageId="21" pageNumber="164" reason="3">
<emphasis id="B93C7801C62BFFD76B9C63B9FED07219" bold="true" box="[189,310,1002,1028]" pageId="21" pageNumber="164">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF7A413C62BFFD76B9C644AFC7372F9" blockId="21.[189,1399,1049,1252]" pageId="21" pageNumber="164">
The combination of valvatiform shell with eventually detached peristome and an only initially sculptured embryonic shell, a thin, yellow operculum without peg-like structure in the nucleus and the massive, trapezoidal penis, whose left side is flattened, distinguishes
<taxonomicName id="4C48DF90C62BFFD76F396433FB4B7264" authority="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér, 2021" authorityName="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér" authorityYear="2021" box="[1048,1197,1119,1145]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="164" phylum="Mollusca" rank="species" species="lozekiana" status="sp. nov.">
<emphasis id="B93C7801C62BFFD76F396433FB4B7264" box="[1048,1197,1119,1145]" italics="true" pageId="21" pageNumber="164">H. lozekiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20FC57AC62BFFD76F946433FAF77267" box="[1205,1297,1120,1146]" pageId="21" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
from all other congeneric species. Only three diagnostic positions of the COI alignment were of
<typeStatus id="54F31AB1C62BFFD76FE464D0FB1E7280" box="[1221,1272,1155,1181]" pageId="21" pageNumber="164">type</typeStatus>
1, a single of
<typeStatus id="54F31AB1C62BFFD76BFF64F4FEF772DC" box="[222,273,1191,1217]" pageId="21" pageNumber="164">type</typeStatus>
3, and 107 of
<typeStatus id="54F31AB1C62BFFD76AE064F4FE1272DC" box="[449,500,1191,1217]" pageId="21" pageNumber="164">type</typeStatus>
4 (
<tableCitation id="C6CA91A8C62BFFD7693D64F5FD9172DD" box="[540,631,1190,1216]" captionStart="Table 3" captionStartId="24.[186,256,548,576]" captionTargetPageId="24" captionText="Table 3 (continued on next page). Diagnostic sites of types 1, 3, and 4 of COI for H. lozekiana sp. nov. and H. steffeki sp. nov. For reference alignment see Supp. file 2." httpUri="http://table.plazi.org/id/DF37F49BC626FFDA6B9B6277FE937479" pageId="21" pageNumber="164" tableUuid="DF37F49BC626FFDA6B9B6277FE937479">Table 3</tableCitation>
). Uncorrected genetic distances to congeners ranged from 0.077 (
<taxonomicName id="4C48DF90C62BFFD76BE66499FE8172FE" baseAuthorityName="Zimmermann" baseAuthorityYear="1930" box="[199,359,1225,1251]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="164" phylum="Mollusca" rank="species" species="kerschneri">
<emphasis id="B93C7801C62BFFD76BE66499FE8172FE" box="[199,359,1225,1251]" italics="true" pageId="21" pageNumber="164">H. kerschneri</emphasis>
</taxonomicName>
) to 0.133 (
<taxonomicName id="4C48DF90C62BFFD76AC86499FD8672FE" authority="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér, 2021" authorityName="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér" authorityYear="2021" box="[489,608,1225,1251]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="164" phylum="Mollusca" rank="species" species="steffeki" status="sp. nov.">
<emphasis id="B93C7801C62BFFD76AC86499FD8672FE" box="[489,608,1225,1251]" italics="true" pageId="21" pageNumber="164">H. steffeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20FC57AC62BFFD769466499FD2372F9" box="[615,709,1226,1252]" pageId="21" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
; see Supp. file 3).
</paragraph>
</subSubSection>
<subSubSection id="C352F798C62BFFD76B9C6558FE2073A0" pageId="21" pageNumber="164" type="etymology">
<paragraph id="8BF7A413C62BFFD76B9C6558FEA27338" blockId="21.[189,324,1291,1317]" box="[189,324,1291,1317]" pageId="21" pageNumber="164">
<heading id="D0BF137FC62BFFD76B9C6558FEA27338" bold="true" box="[189,324,1291,1317]" fontSize="11" level="3" pageId="21" pageNumber="164" reason="3">
<emphasis id="B93C7801C62BFFD76B9C6558FEA27338" bold="true" box="[189,324,1291,1317]" pageId="21" pageNumber="164">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF7A413C62BFFD76B9C656AFE2073A0" blockId="21.[189,1399,1337,1469]" pageId="21" pageNumber="164">
<taxonomicName id="4C48DF90C62BFFD76B9C656AFE48734E" authority="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér, 2021" authorityName="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér" authorityYear="2021" box="[189,430,1337,1363]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="164" phylum="Mollusca" rank="species" species="lozekiana" status="sp. nov.">
<emphasis id="B93C7801C62BFFD76B9C656AFE48734E" box="[189,430,1337,1363]" italics="true" pageId="21" pageNumber="164">Hauffenia lozekiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20FC57AC62BFFD76A9A6569FDFD7349" box="[443,539,1338,1364]" pageId="21" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
is dedicated to Vojen Ložek (
<date id="FFF682D3C62BFFD768B2656AFBC97349" box="[915,1071,1337,1364]" pageId="21" pageNumber="164" value="1925-07-26">26 Jul. 1925</date>
<date id="FFF682D3C62BFFD76F1E656AFB0C734E" box="[1087,1258,1337,1364]" pageId="21" pageNumber="164" value="2020-08-15">15 Aug. 2020</date>
), renowned and influential Czech malacologist and paleontologist, who contributed significantly to Quaternary malacostratigraphy, Czechoslovakian zoogeography of mollusks and was the first to discover the genus
<taxonomicName id="4C48DF90C62BFFD76B9C65F0FED473A0" authorityName="Pollonera" authorityYear="1898" box="[189,306,1443,1469]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="164" phylum="Mollusca" rank="genus">
<emphasis id="B93C7801C62BFFD76B9C65F0FED473A0" box="[189,306,1443,1469]" italics="true" pageId="21" pageNumber="164">Hauffenia</emphasis>
</taxonomicName>
in
<collectingCountry id="F35FE483C62BFFD76A7965F0FE2473A0" box="[344,450,1443,1469]" name="Slovakia" pageId="21" pageNumber="164">Slovakia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C352F798C62BFFD46B9C65B7FB97775B" lastPageId="22" lastPageNumber="165" pageId="21" pageNumber="164" type="materials_examined">
<paragraph id="8BF7A413C62BFFD76B9C65B7FE4C73E3" blockId="21.[189,426,1508,1534]" box="[189,426,1508,1534]" pageId="21" pageNumber="164">
<heading id="D0BF137FC62BFFD76B9C65B7FE4C73E3" bold="true" box="[189,426,1508,1534]" fontSize="11" level="3" pageId="21" pageNumber="164" reason="3">
<emphasis id="B93C7801C62BFFD76B9C65B7FE4C73E3" bold="true" box="[189,426,1508,1534]" pageId="21" pageNumber="164">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF7A413C62BFFD76BCB6640FEE8708A" blockId="21.[189,1399,1555,1687]" pageId="21" pageNumber="164">
<materialsCitation id="3B20AE4EC62BFFD76BCB6640FEE8708A" ID-GBIF-Occurrence="3392791313" collectingDate="2014-01-04" collectionCode="NHMW" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="NHMW 113638" specimenCount="1" typeStatus="holotype">
<emphasis id="B93C7801C62BFFD76BCB6640FEBA7030" bold="true" box="[234,348,1555,1581]" pageId="21" pageNumber="164">
<typeStatus id="54F31AB1C62BFFD76BCB6640FEBA7030" box="[234,348,1555,1581]" pageId="21" pageNumber="164" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="1373B896C62BFFD76A4D6640FE1C7030" box="[364,506,1555,1581]" captionStart="Fig" captionStartId="22.[189,232,1945,1971]" captionTargetBox="[189,1398,1264,1906]" captionTargetId="figure-285@22.[189,1398,1264,1906]" captionTargetPageId="22" captionText="Fig. 12. Holotypes. AC. H. lozekiana sp. nov. (NHMW 113638). DE. H. steffeki sp. nov. (NHMW 113640). Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5596702" httpUri="https://zenodo.org/record/5596702/files/figure.png" pageId="21" pageNumber="164">Fig. 12AC</figureCitation>
)
<collectingCountry id="F35FE483C62BFFD76B9C6665FEA9704D" box="[189,335,1590,1616]" name="Slovakia" pageId="21" pageNumber="164">SLOVAKIA</collectingCountry>
<location id="8E97F2C8C62BFFD76A5A6665FD8D704D" LSID="urn:lsid:plazi:treatment:03E11505C62BFFD5694C6331FE507297:8E97F2C8C62BFFD76A5A6665FD8D704D" box="[379,619,1590,1616]" country="Slovakia" county="Kalinovsky" latitude="48.70045" longLatPrecision="1" longitude="19.265867" municipality="Oravce" name="Bystrica Highlands" pageId="21" pageNumber="164">Bystrica Highlands</location>
,
<locationDeviation id="9A45BD31C62BFFD769A06665FC2F704D" box="[641,969,1590,1616]" pageId="21" pageNumber="164">vicinity of Poniky/Oravce</locationDeviation>
,
<collectingCounty id="6296DC9FC62BFFD768FE6665FB80704D" box="[991,1126,1590,1616]" pageId="21" pageNumber="164">Kalinovský</collectingCounty>
prameň (Kalinovec);
<geoCoordinate id="EE7CC2D4C62BFFD76B9C660BFEBA7069" box="[189,348,1624,1652]" degrees="48" direction="north" minutes="42.027" orientation="latitude" pageId="21" pageNumber="164" precision="1" value="48.70045">48°42.027 N</geoCoordinate>
,
<geoCoordinate id="EE7CC2D4C62BFFD76A46660BFDE47069" box="[359,514,1624,1652]" degrees="19" direction="east" minutes="15.952" orientation="longitude" pageId="21" pageNumber="164" precision="1" value="19.265867">19°15.952 E</geoCoordinate>
; alt.
<quantity id="4CB009F6C62BFFD76918660AFD637069" box="[569,645,1625,1652]" metricMagnitude="2" metricUnit="m" metricValue="4.82" pageId="21" pageNumber="164" unit="m" value="482.0">
<elevation id="00654320C62BFFD76918660AFD637069" box="[569,645,1625,1652]" metricMagnitude="2" metricUnit="m" metricValue="4.82" pageId="21" pageNumber="164" unit="m" value="482.0">482 m</elevation>
</quantity>
;
<date id="FFF682D3C62BFFD769B0660AFCFF706E" box="[657,793,1625,1652]" pageId="21" pageNumber="164" value="2014-01-04">
<collectingDate id="EFB27B3BC62BFFD769B0660AFCFF706E" box="[657,793,1625,1652]" pageId="21" pageNumber="164" value="2014-01-04">4 Jan. 2014</collectingDate>
</date>
;
<collectorName id="26BDC1C5C62BFFD768056609FC60706E" box="[804,902,1625,1652]" pageId="21" pageNumber="164">J. Grego</collectorName>
leg.; karst spring;
<specimenCode id="DBEE0C68C62BFFD76F7F6609FAF9706E" box="[1118,1311,1625,1652]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="164" type="Museum">NHMW 113638</specimenCode>
. (
<figureCitation id="1373B896C62BFFD76E13660AFA917069" box="[1330,1399,1625,1652]" captionStart="Fig" captionStartId="11.[191,234,1958,1984]" captionTargetBox="[189,1398,1104,1917]" captionTargetId="figure-350@11.[189,1398,1104,1917]" captionTargetPageId="11" captionText="Fig. 5. Type locality of H. lozekiana sp. nov. (71) and Dabarska Pećina, a locality of H. steffeki sp. nov. (72) (see Table 1)." figureDoi="http://doi.org/10.5281/zenodo.5596686" httpUri="https://zenodo.org/record/5596686/files/figure.png" pageId="21" pageNumber="164">Fig. 5</figureCitation>
71ab)
</materialsCitation>
</paragraph>
<paragraph id="8BF7A413C62BFFD46BCB66EDFB97775B" blockId="21.[189,1399,1726,1964]" lastBlockId="22.[189,1398,265,327]" lastPageId="22" lastPageNumber="165" pageId="21" pageNumber="164">
<materialsCitation id="3B20AE4EC62BFFD76BCB66EDFBE470E6" ID-GBIF-Occurrence="3392791304" collectingDate="2014-01-04" collectionCode="JG" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="SLOVAKIA" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="JG F1684" specimenCount="5" typeStatus="paratype">
<emphasis id="B93C7801C62BFFD76BCB66EDFE8170C5" bold="true" box="[234,359,1726,1752]" pageId="21" pageNumber="164">
<typeStatus id="54F31AB1C62BFFD76BCB66EDFE8170C5" box="[234,359,1726,1752]" pageId="21" pageNumber="164" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<figureCitation id="1373B896C62BFFD76A5966EDFDE370C5" box="[376,517,1726,1752]" captionStart="Fig" captionStartId="23.[189,232,1947,1973]" captionTargetBox="[189,1398,1659,1906]" captionTargetId="figure-353@23.[189,1398,1659,1906]" captionTargetPageId="23" captionText="Fig. 13. Paratypes from type localities (SEM micrographs). AB. H. lozekiana sp. nov. (NHMW 113639). CD. H. steffeki sp. nov. (NHMW 113641). Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5596704" httpUri="https://zenodo.org/record/5596704/files/figure.png" pageId="21" pageNumber="164">Fig. 13AB</figureCitation>
)
<collectingCountry id="F35FE483C62BFFD76B9C66B2FEA970E6" box="[189,335,1761,1787]" name="Slovakia" pageId="21" pageNumber="164">SLOVAKIA</collectingCountry>
<specimenCount id="9D4E6F9AC62BFFD76A4866B2FE2570E1" box="[361,451,1761,1788]" pageId="21" pageNumber="164">5 specs</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBEE0C68C62BFFD768AF66B2FBE470E6" box="[910,1026,1761,1787]" pageId="21" pageNumber="164">JG F1684</specimenCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC62BFFD76F3C66B2FDA67103" ID-GBIF-Occurrence="3392791323" collectingDate="2014-01-04" collectionCode="NHMW" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="NHMW 113639" specimenCount="4">
<specimenCount id="9D4E6F9AC62BFFD76F3C66B2FB9170E1" box="[1053,1143,1761,1788]" pageId="21" pageNumber="164">4 specs</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBEE0C68C62BFFD76AA06756FDA67103" box="[385,576,1796,1823]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="164" type="Museum">NHMW 113639</specimenCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC62BFFD7697D6757FB507102" ID-GBIF-Occurrence="3392791319" box="[604,1206,1796,1823]" collectingDate="2014-01-04" collectionCode="ZPE" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCount="2">
<specimenCount id="9D4E6F9AC62BFFD7697D6757FD507102" box="[604,694,1796,1823]" pageId="21" pageNumber="164">2 specs</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED593CD6C62BFFD76FA36756FB507102" box="[1154,1206,1797,1823]" pageId="21" pageNumber="164">ZPE</collectionCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC62BFFD76FF36757FC4E715F" ID-GBIF-Occurrence="3392791301" collectingDate="2014-01-25" collectionCode="HNHM" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="HNHM 105307" specimenCount="2">
<specimenCount id="9D4E6F9AC62BFFD76FF36757FACB7102" box="[1234,1325,1796,1823]" pageId="21" pageNumber="164">2 specs</specimenCount>
; same collection data as for holotype;
<date id="FFF682D3C62BFFD7691F6774FD3B715C" box="[574,733,1831,1858]" pageId="21" pageNumber="164" value="2014-01-25">
<collectingDate id="EFB27B3BC62BFFD7691F6774FD3B715C" box="[574,733,1831,1858]" pageId="21" pageNumber="164" value="2014-01-25">25 Jan. 2014</collectingDate>
</date>
;
<specimenCode id="DBEE0C68C62BFFD769CD677BFC4E715F" box="[748,936,1831,1858]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="21" pageNumber="164">HNHM 105307</specimenCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC62BFFD768E66774FE417179" ID-GBIF-Occurrence="3392791312" collectingDate="2014-01-25" collectionCode="JG" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="JG F1649" specimenCount="2">
<specimenCount id="9D4E6F9AC62BFFD768E66774FBC5715F" box="[967,1059,1831,1858]" pageId="21" pageNumber="164">2 specs</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBEE0C68C62BFFD76A126718FE417179" box="[307,423,1866,1893]" pageId="21" pageNumber="164">JG F1649</specimenCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC62BFFD76AE16718FB187178" ID-GBIF-Occurrence="3392791309" box="[448,1278,1866,1893]" collectingDate="2017-05-18" collectionCode="JG" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="JG F1650" specimenCount="2">
<specimenCount id="9D4E6F9AC62BFFD76AE16718FDFC7178" box="[448,538,1867,1893]" pageId="21" pageNumber="164">2 specs</specimenCount>
; same collection data as for holotype;
<date id="FFF682D3C62BFFD768FF6718FB997178" box="[990,1151,1867,1893]" pageId="21" pageNumber="164" value="2017-05-18">
<collectingDate id="EFB27B3BC62BFFD768FF6718FB997178" box="[990,1151,1867,1893]" pageId="21" pageNumber="164" value="2017-05-18">18 May 2017</collectingDate>
</date>
;
<specimenCode id="DBEE0C68C62BFFD76FAA6718FB187178" box="[1163,1278,1866,1893]" pageId="21" pageNumber="164">JG F1650</specimenCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC62BFFD76E396718FC797195" ID-GBIF-Occurrence="3392791310" collectingDate="2020-07-02" collectionCode="JG" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="JG F1886" specimenCount="4">
<specimenCount id="9D4E6F9AC62BFFD76E396718FA947178" box="[1304,1394,1867,1893]" pageId="21" pageNumber="164">4 specs</specimenCount>
; same collection data as for holotype;
<date id="FFF682D3C62BFFD769AA673DFCFE7195" box="[651,792,1902,1928]" pageId="21" pageNumber="164" value="2020-07-02">
<collectingDate id="EFB27B3BC62BFFD769AA673DFCFE7195" box="[651,792,1902,1928]" pageId="21" pageNumber="164" value="2020-07-02">2 Jul. 2020</collectingDate>
</date>
;
<specimenCode id="DBEE0C68C62BFFD76809673DFC797195" box="[808,927,1902,1928]" pageId="21" pageNumber="164">JG F1886</specimenCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC62BFFD768E1673DFE0E71B1" ID-GBIF-Occurrence="3392791320" collectingDate="2020-07-02" collectionCode="NMBE" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="NMBE 569388" specimenCount="2">
<specimenCount id="9D4E6F9AC62BFFD768E1673DFBFB7195" box="[960,1053,1902,1928]" pageId="21" pageNumber="164">2 specs</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBEE0C68C62BFFD76A1567C1FE0E71B1" box="[308,488,1937,1964]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="21" pageNumber="164">NMBE 569388</specimenCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC62BFFD7692367C2FB5D71B1" ID-GBIF-Occurrence="3392791321" box="[514,1211,1937,1964]" collectingDate="2020-07-02" collectionCode="SMF" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="SMF 363255" specimenCount="2">
<specimenCount id="9D4E6F9AC62BFFD7692367C2FDBD71B1" box="[514,603,1937,1964]" pageId="21" pageNumber="164">2 specs</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBEE0C68C62BFFD76F0167C2FB5D71B1" box="[1056,1211,1937,1964]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="164">SMF 363255</specimenCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC62BFFD46FF567C2FD1C773E" ID-GBIF-Occurrence="3392791326" collectingDate="2020-07-02" collectionCode="SMOPAJ" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" lastPageId="22" lastPageNumber="165" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="21" pageNumber="164" specimenCode="SMOPAJ 1" specimenCount="2">
<specimenCount id="9D4E6F9AC62BFFD76FF567C2FAC871B1" box="[1236,1326,1937,1964]" pageId="21" pageNumber="164">2 specs</specimenCount>
; same collection data as for holotype; SMOPAJ 1/2021
</materialsCitation>
<materialsCitation id="3B20AE4EC628FFD46834615AFEF3775B" ID-GBIF-Occurrence="3392791327" collectingDate="2020-07-02" collectionCode="ZMH" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="22" pageNumber="165" specimenCode="ZMH 140884" specimenCount="2">
<specimenCount id="9D4E6F9AC628FFD46834615AFC88773E" box="[789,878,265,291]" pageId="22" pageNumber="165">2 specs</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBEE0C68C628FFD46E14615AFEF3775B" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="22" pageNumber="165" type="Museum">ZMH 140884</specimenCode>
</materialsCitation>
<materialsCitation id="3B20AE4EC628FFD46A0F617FFB8B775B" ID-GBIF-Occurrence="3392791325" box="[302,1133,300,327]" collectingDate="2020-12-21" collectionCode="JG" collectorName="J. Grego" country="Slovakia" county="Kalinovsky" elevation="482" latitude="48.70045" location="Bystrica Highlands" longLatPrecision="1" longitude="19.265867" municipality="Oravce" pageId="22" pageNumber="165" specimenCode="JG F1887" specimenCount="3">
<specimenCount id="9D4E6F9AC628FFD46A0F617FFE61775A" box="[302,391,300,327]" pageId="22" pageNumber="165">3 specs</specimenCount>
; same collection data as for holotype;
<date id="FFF682D3C628FFD46868617FFC0C775B" box="[841,1002,300,327]" pageId="22" pageNumber="165" value="2020-12-21">
<collectingDate id="EFB27B3BC628FFD46868617FFC0C775B" box="[841,1002,300,327]" pageId="22" pageNumber="165" value="2020-12-21">21 Dec. 2020</collectingDate>
</date>
;
<specimenCode id="DBEE0C68C628FFD468D7617FFB8B775B" box="[1014,1133,300,326]" pageId="22" pageNumber="165">JG F1887</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C352F798C628FFD56B9C613EFD5D7497" lastPageId="23" lastPageNumber="166" pageId="22" pageNumber="165" type="description">
<paragraph id="8BF7A413C628FFD46B9C613EFEA8779A" blockId="22.[189,334,365,391]" box="[189,334,365,391]" pageId="22" pageNumber="165">
<heading id="D0BF137FC628FFD46B9C613EFEA8779A" bold="true" box="[189,334,365,391]" fontSize="11" level="3" pageId="22" pageNumber="165" reason="3">
<emphasis id="B93C7801C628FFD46B9C613EFEA8779A" bold="true" box="[189,334,365,391]" pageId="22" pageNumber="165">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF7A413C628FFD46B9C61CFFE1A747B" blockId="22.[189,1399,412,614]" pageId="22" pageNumber="165">
SHELL (
<figureCitation id="1373B896C628FFD46A3D61CFFE5077AB" box="[284,438,412,438]" captionStart="Fig" captionStartId="22.[189,232,1945,1971]" captionTargetBox="[189,1398,1264,1906]" captionTargetId="figure-285@22.[189,1398,1264,1906]" captionTargetPageId="22" captionText="Fig. 12. Holotypes. AC. H. lozekiana sp. nov. (NHMW 113638). DE. H. steffeki sp. nov. (NHMW 113640). Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5596702" httpUri="https://zenodo.org/record/5596702/files/figure.png" pageId="22" pageNumber="165">Figs 12AC</figureCitation>
,
<figureCitation id="1373B896C628FFD46AEB61CFFDC477AB" box="[458,546,412,438]" captionStart="Fig" captionStartId="23.[189,232,1947,1973]" captionTargetBox="[189,1398,1659,1906]" captionTargetId="figure-353@23.[189,1398,1659,1906]" captionTargetPageId="23" captionText="Fig. 13. Paratypes from type localities (SEM micrographs). AB. H. lozekiana sp. nov. (NHMW 113639). CD. H. steffeki sp. nov. (NHMW 113641). Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5596704" httpUri="https://zenodo.org/record/5596704/files/figure.png" pageId="22" pageNumber="165">13AB</figureCitation>
,
<figureCitation id="1373B896C628FFD4691761CFFD8A77AB" box="[566,620,412,438]" captionStart="Fig" captionStartId="26.[189,232,1945,1971]" captionTargetBox="[200,1388,1376,1884]" captionTargetId="figure-409@26.[189,1398,1366,1893]" captionTargetPageId="26" captionText="Fig. 14. Protoconchs (SEM micrographs). A. H. lozekiana sp. nov., paratype (NHMW 113639). B. H. steffeki sp. nov., paratype (NHMW 113641). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5596706" httpUri="https://zenodo.org/record/5596706/files/figure.png" pageId="22" pageNumber="165">14A</figureCitation>
;
<tableCitation id="C6CA91A8C628FFD469A161CFFD0477AB" box="[640,738,412,438]" captionStart="Table 4" captionStartId="28.[189,259,265,291]" captionTargetPageId="28" captionText="Table 4. Morphometrics of the new species. Abbreviations: AH = aperture height; AW = aperture width; CV = coefficient of variation corrected for unequal sample sizes; max = maximum; min = minimum; SD = standard deviation; SH = shell height; SW = shell width; W = number of whorls (to the nearest eighth). Measurements in mm." httpUri="http://table.plazi.org/id/DF37F49BC622FFDE6B9C615AFDC07790" pageId="22" pageNumber="165" tableUuid="DF37F49BC622FFDE6B9C615AFDC07790">Table 4</tableCitation>
). Valvatiform, maximum height measured
<quantity id="4CB009F6C628FFD46FDE61CFFA9477AA" box="[1279,1394,412,439]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="22" pageNumber="165" unit="mm" value="1.03">1.03 mm</quantity>
, maximum width
<quantity id="4CB009F6C628FFD46AA961ECFE1377C7" box="[392,501,447,474]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="22" pageNumber="165" unit="mm" value="1.58">1.58 mm</quantity>
, on average twice as wide as high, with up to 2.75 whorls, final part of last whorl may be detached, fairly variable in shape with a coefficient of variation of 13.31% for the height/ width ratio; embryonic shell with 1.11.2 whorls, first 0.75 whorls wrinkled, sculpture then fading to become smooth; aperture almost round only slightly higher than wide, prosocline, lip continuous, not thickened; umbilicus wide.
</paragraph>
<paragraph id="8BF7A413C628FFD46B9C62DDFE6B74D6" blockId="22.[189,1398,653,715]" pageId="22" pageNumber="165">OPERCULUM. Thin, chitinous, yellow, almost circular with central nucleus, the latter without peg or peglike modification.</paragraph>
<paragraph id="8BF7A413C628FFD46B9C62A0FEC37532" blockId="22.[189,1398,754,815]" pageId="22" pageNumber="165">EXTERNAL FEATURES. Epidermis without pigment, no eye spots visible, tentacles without particular ciliation.</paragraph>
<paragraph id="8BF7A413C628FFD46B9C6304FB53756C" blockId="22.[189,1205,855,881]" box="[189,1205,855,881]" pageId="22" pageNumber="165">MANTLE CAVITY. 9 gill filaments (N = 1), osphradium short, oval, behind middle of gill.</paragraph>
<paragraph id="8BF7A413C628FFD46B9C63CAFB7C75E4" blockId="22.[188,1400,920,1017]" pageId="22" pageNumber="165">
DIGESTIVE SYSTEM (
<figureCitation id="1373B896C628FFD46ABF63CBFDD575AE" box="[414,563,920,947]" captionStart="Fig" captionStartId="27.[189,232,1913,1939]" captionTargetBox="[205,1380,529,1859]" captionTargetId="figure-96@27.[190,1397,511,1877]" captionTargetPageId="27" captionText="Fig. 15. Radulae (SEM micrographs). AC. H. lozekiana sp. nov. DF. H. steffeki sp. nov. D. Type locality. EF. Spring Krivina (locality 74 from Table 1). Scale bars: A = 10 µm; BE = 2 µm; F = 3 µm." figureDoi="http://doi.org/10.5281/zenodo.5596708" httpUri="https://zenodo.org/record/5596708/files/figure.png" pageId="22" pageNumber="165">Figs 15AC</figureCitation>
,
<figureCitation id="1373B896C628FFD4696063CBFD8475AE" box="[577,610,920,947]" captionStart="Fig" captionStartId="29.[189,232,1973,1999]" captionTargetId="figure-195@29.[416,1172,935,1935]" captionTargetPageId="29" captionText="Fig. 16. Schematic representations of intestinal loop configurations." figureDoi="http://doi.org/10.5281/zenodo.5597874" httpUri="https://zenodo.org/record/5597874/files/figure.png" pageId="22" pageNumber="165">16</figureCitation>
). Radula with formula R 4-5 1 4-5/1 1, L 4-6 1 5-6. M1 23-29, M2 15-20 (N = 3); intestine forming a long loop in roof of mantle cavity, the loop is additionally coiled, its appearance seems to vary depending on the presence and number of fecal pellets.
</paragraph>
<paragraph id="8BF7A413C628FFD46B9C6472FDA77240" blockId="22.[188,1399,1056,1118]" pageId="22" pageNumber="165">FEMALE GENITALIA. Not observed; the only live collected female specimen was subadult, hence, the distal genitalia were not yet developed.</paragraph>
<caption id="DF37F49BC628FFD46B9C67CAFDFA71CA" ID-DOI="http://doi.org/10.5281/zenodo.5596702" ID-Zenodo-Dep="5596702" httpUri="https://zenodo.org/record/5596702/files/figure.png" pageId="22" pageNumber="165" startId="22.[189,232,1945,1971]" targetBox="[189,1398,1264,1906]" targetPageId="22">
<paragraph id="8BF7A413C628FFD46B9C67CAFDFA71CA" blockId="22.[189,1399,1945,2007]" pageId="22" pageNumber="165">
<emphasis id="B93C7801C628FFD46B9C67CAFEFA71AE" bold="true" box="[189,284,1945,1971]" pageId="22" pageNumber="165">Fig. 12.</emphasis>
Holotypes.
<emphasis id="B93C7801C628FFD46A9167CAFE0171AE" bold="true" box="[432,487,1945,1971]" pageId="22" pageNumber="165">AC</emphasis>
.
<taxonomicName id="4C48DF90C628FFD46AD967C9FD6B71AE" authority="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér, 2021" authorityName="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér" authorityYear="2021" box="[504,653,1945,1971]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="165" phylum="Mollusca" rank="species" species="lozekiana" status="sp. nov.">
<emphasis id="B93C7801C628FFD46AD967C9FD6B71AE" box="[504,653,1945,1971]" italics="true" pageId="22" pageNumber="165">H. lozekiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20FC57AC628FFD469B667C9FD1571A9" box="[663,755,1946,1972]" pageId="22" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
(NHMW 113638). DE.
<taxonomicName id="4C48DF90C628FFD46F0967C9FB4771AE" authority="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér, 2021" authorityName="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér" authorityYear="2021" box="[1064,1185,1945,1971]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="165" phylum="Mollusca" rank="species" species="steffeki" status="sp. nov.">
<emphasis id="B93C7801C628FFD46F0967C9FB4771AE" box="[1064,1185,1945,1971]" italics="true" pageId="22" pageNumber="165">H. steffeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20FC57AC628FFD46F8967C9FAE271A9" box="[1192,1284,1946,1972]" pageId="22" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
(NHMW 113640). Scale bar = 500 µm.
</paragraph>
</caption>
<paragraph id="8BF7A413C629FFD56B9C615AFEC57774" blockId="23.[189,1399,265,361]" pageId="23" pageNumber="166">
MALE GENITALIA (
<figureCitation id="1373B896C629FFD56AAD615AFDC5773E" box="[396,547,265,291]" captionStart="Fig" captionStartId="30.[188,231,1913,1939]" captionTargetBox="[205,1378,1074,1859]" captionTargetId="figure-303@30.[189,1397,1058,1875]" captionTargetPageId="30" captionText="Fig. 17. Penes (SEM micrographs). AB. H. lozekiana sp. nov. (different specimens). CE. H. steffeki sp. nov. (one specimen). Arrow points at penial stylet, which is withdrawn in A and B. Scale bars: AC = 50 µm; D = 10 µm; E = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.5596710" httpUri="https://zenodo.org/record/5596710/files/figure.png" pageId="23" pageNumber="166">Figs 17AB</figureCitation>
,
<figureCitation id="1373B896C629FFD56910615AFDB4773E" box="[561,594,265,291]" captionStart="Fig" captionStartId="31.[189,232,1982,2008]" captionTargetBox="[368,1219,1271,1944]" captionTargetId="figure-281@31.[368,1219,1271,1944]" captionTargetPageId="31" captionText="Fig. 18. Penis of H. lozekiana sp. nov. Frontal view showing stylet. Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5596712" httpUri="https://zenodo.org/record/5596712/files/figure.png" pageId="23" pageNumber="166">18</figureCitation>
). Not observed except for kidney-shaped prostate and penis; penis massive, trapezoidal, left side flattened, small, brown, probably chitinous stylet anterior right near penial opening.
</paragraph>
<paragraph id="8BF7A413C629FFD56B9C61C3FEFB77B7" blockId="23.[189,285,400,426]" box="[189,285,400,426]" pageId="23" pageNumber="166">
<heading id="D0BF137FC629FFD56B9C61C3FEFB77B7" bold="true" box="[189,285,400,426]" fontSize="11" level="3" pageId="23" pageNumber="166" reason="3">
<emphasis id="B93C7801C629FFD56B9C61C3FEFB77B7" bold="true" box="[189,285,400,426]" pageId="23" pageNumber="166">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BF7A413C629FFD56B9C61ECFD5D7497" blockId="23.[189,1399,447,650]" pageId="23" pageNumber="166">
The type locality is a small permanent karst spring emerging from the small karst massif of Drienok formed by Middle Triassic (AegeanLadinian) Wetterstein, Reifling and Gutenstein Limestones of the Silicikum Unit (Silica Nappe) at the contact zone to impermeable Early Triassic shales and sandstones of Szin and Bódvaszilas Members with a small discontinuous stripe of alluvial gravel at the bottom of the spring valley. The character of the
<typeStatus id="54F31AB1C629FFD5695F621EFD57747A" box="[638,689,589,615]" pageId="23" pageNumber="166">type</typeStatus>
locality indicates the same habitat preference in the phreatic rhizosphere as discussed for
<taxonomicName id="4C48DF90C629FFD5692F6223FD527494" authorityName="Eross &amp; Petro" authorityYear="2008" box="[526,692,623,649]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="166" phylum="Mollusca" rank="species" species="kissdalmae">
<emphasis id="B93C7801C629FFD5692F6223FD527494" box="[526,692,623,649]" italics="true" pageId="23" pageNumber="166">H. kissdalmae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C352F798C629FFD56B9C62E3FD4574E4" box="[189,675,688,762]" pageId="23" pageNumber="166" type="distribution">
<paragraph id="8BF7A413C629FFD56B9C62E3FEB074D7" blockId="23.[189,342,688,714]" box="[189,342,688,714]" pageId="23" pageNumber="166">
<heading id="D0BF137FC629FFD56B9C62E3FEB074D7" bold="true" box="[189,342,688,714]" fontSize="11" level="3" pageId="23" pageNumber="166" reason="3">
<emphasis id="B93C7801C629FFD56B9C62E3FEB074D7" bold="true" box="[189,342,688,714]" pageId="23" pageNumber="166">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF7A413C629FFD56B9C62B3FD4574E4" blockId="23.[189,675,735,762]" box="[189,675,735,762]" pageId="23" pageNumber="166">
Known only from the
<typeStatus id="54F31AB1C629FFD56AE362B3FE1374E7" box="[450,501,736,762]" pageId="23" pageNumber="166">type</typeStatus>
locality so far.
</paragraph>
</subSubSection>
<subSubSection id="C352F798C629FFD56B9C6373FCD775CE" pageId="23" pageNumber="166" type="conservation">
<paragraph id="8BF7A413C629FFD56B9C6373FE837527" blockId="23.[189,357,800,826]" box="[189,357,800,826]" pageId="23" pageNumber="166">
<heading id="D0BF137FC629FFD56B9C6373FE837527" bold="true" box="[189,357,800,826]" fontSize="11" level="3" pageId="23" pageNumber="166" reason="3">
<emphasis id="B93C7801C629FFD56B9C6373FE837527" bold="true" box="[189,357,800,826]" pageId="23" pageNumber="166">Conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8BF7A413C629FFD56B9C631CFCD775CE" blockId="23.[188,1399,847,979]" pageId="23" pageNumber="166">
There is only one known location and the area of occupancy is probably considerably smaller than
<quantity id="4CB009F6C629FFD56B9C6320FEE07591" box="[189,262,882,909]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="166" unit="km" value="20.0">20 km</quantity>
<superScript id="7C3D095BC629FFD56A266321FEF6759D" attach="right" box="[263,272,882,896]" fontSize="6" pageId="23" pageNumber="166">2</superScript>
. The combination of the restricted range and the threat imposed by the apparent trend of logging in the area around the
<typeStatus id="54F31AB1C629FFD56AE063C5FE1275AD" box="[449,500,918,944]" pageId="23" pageNumber="166">type</typeStatus>
locality qualify for a threat assessment of Vulnerable (VU, D2) according to the
<bibRefCitation id="EFD9D9E2C629FFD56BC963EAFCC875CE" author="IUCN Standards &amp; Petitions Committee" box="[232,814,953,979]" pageId="23" pageNumber="166" refId="ref17487" refString="IUCN Standards and Petitions Committee. 2019. Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from http: // www. iucnredlist. org / documents / RedListGuidelines. pdf [accessed 12 Jun. 2021]." type="book" year="2019">IUCN Standards and Petitions Committee (2019)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C352F798C629FFD56B9C63A9FE507297" pageId="23" pageNumber="166" type="discussion">
<paragraph id="8BF7A413C629FFD56B9C63A9FEC97209" blockId="23.[189,303,1018,1044]" box="[189,303,1018,1044]" pageId="23" pageNumber="166">
<heading id="D0BF137FC629FFD56B9C63A9FEC97209" bold="true" box="[189,303,1018,1044]" fontSize="11" level="3" pageId="23" pageNumber="166" reason="3">
<emphasis id="B93C7801C629FFD56B9C63A9FEC97209" bold="true" box="[189,303,1018,1044]" pageId="23" pageNumber="166">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BF7A413C629FFD56B9C647AFE507297" blockId="23.[186,1398,1065,1162]" pageId="23" pageNumber="166">
The genetic distances of
<taxonomicName id="4C48DF90C629FFD56AFA6479FD88725E" authority="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér, 2021" authorityName="Haase &amp; Grego &amp; Erőss &amp; Farkas &amp; Fehér" authorityYear="2021" box="[475,622,1065,1091]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="166" phylum="Mollusca" rank="species" species="lozekiana" status="sp. nov.">
<emphasis id="B93C7801C629FFD56AFA6479FD88725E" box="[475,622,1065,1091]" italics="true" pageId="23" pageNumber="166">H. lozekiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20FC57AC629FFD56954647AFD28725E" box="[629,718,1065,1091]" pageId="23" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
from the more widespread
<taxonomicName id="4C48DF90C629FFD56F2D6479FB57725E" authorityName="Eross &amp; Petro" authorityYear="2008" box="[1036,1201,1065,1091]" class="Gastropoda" family="Hydrobiidae" genus="Hauffenia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="166" phylum="Mollusca" rank="species" species="kissdalmae">
<emphasis id="B93C7801C629FFD56F2D6479FB57725E" box="[1036,1201,1065,1091]" italics="true" pageId="23" pageNumber="166">H. kissdalmae</emphasis>
</taxonomicName>
and the Austrian species suggest that it is a relict probably from the earlier colonization of the Hron River Valley ca 4.5 Ma ago (see Discussion).
</paragraph>
</subSubSection>
</treatment>
</document>