161 lines
19 KiB
XML
161 lines
19 KiB
XML
<document id="B332A8A5D50A8F6CC98E2DFAF338F261" ID-DOI="10.5252/z2011n1a5" ID-ISSN="1638-9387" ID-Zenodo-Dep="4546704" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613600601004" checkinUser="felipe" docAuthor="Montreuil, Olivier" docDate="2011" docId="03DC9E64DA543A643AEAFF738706E449" docLanguage="fr" docName="Zoosystema.33.1.101-122.pdf" docOrigin="Zoosystema 33 (1)" docSource="http://dx.doi.org/10.5252/z2011n1a5" docStyle="DocumentStyle:EB3C2C73672D2E883259A050BD27978B.6:Zoosystema.2011-2012.journal_article" docStyleId="EB3C2C73672D2E883259A050BD27978B" docStyleName="Zoosystema.2011-2012.journal_article" docStyleVersion="6" docTitle="Arachnodes bicolor Montreuil 2011, n. sp." docType="treatment" docVersion="3" lastPageNumber="108" masterDocId="FFE5E61CDA533A633A1FFFA48336E764" masterDocTitle="Monophylie du « groupe Epilissus » et refonte des genres Epilissus Reiche, 1841, Arachnodes Westwood, 1847 et Apterepilissus n. gen. de Madagascar (Insecta, Coleoptera, Scarabaeidae, Epilissini)" masterLastPageNumber="122" masterPageNumber="101" pageNumber="108" updateTime="1698872838645" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6EDA87D972FF35C3482C65B88CC9D826">Monophylie du « groupe Epilissus » et refonte des genres Epilissus Reiche, 1841, Arachnodes Westwood, 1847 et Apterepilissus n. gen. de Madagascar (Insecta, Coleoptera, Scarabaeidae, Epilissini)</mods:title>
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<mods:name id="B93EF411894F610DC42F2CAADD43E82B" type="personal">
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<mods:role id="341205E3EEF8A8D4A4E36921097292F9">
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<mods:roleTerm id="44DF953C953F12648D291830779AEB39">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="848940B66001BD1EBFD0C81BFD6421DB">Montreuil, Olivier</mods:namePart>
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<mods:affiliation id="4FFA7CCE4FA33893E90988B22577D055">Muséum national d’Histoire naturelle, UMR 7205, Département Systématique et Évolution, Entomologie, case postale 50, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) o. montr. @ mnhn. fr</mods:affiliation>
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<mods:nameIdentifier id="7DC311ECFB66DBDF3EA9139EDB16B961" type="email">montr.@mnhn.fr</mods:nameIdentifier>
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<mods:typeOfResource id="BD26740C1CCD7BDCFA5E41866C1C64FF">text</mods:typeOfResource>
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<mods:title id="7ED0F86B85E92C5894E924C4DB17F3E3">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="474571DF034384B3AFE023940EC15AA7">
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<mods:date id="03B292B7D10DFA8940EF276722ED0B6D">2011</mods:date>
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<mods:number id="E41C6CFB5200F5DE50B1A8C0DBF1CCE6">2011-03-31</mods:number>
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<mods:detail id="0CC86D94AA8F4D2CAD8D5D9DD8EAF4A8" type="volume">
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<mods:number id="D394F34463B7C0BBC0D09F63485E72CA">33</mods:number>
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<mods:number id="816BBC395EF3D3EA1A8BFA5AD4199176">1</mods:number>
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<mods:location id="38EB5D844373FE6730A86ED51936A4F0">
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<mods:url id="77AF2C24F8E795FEE6DC38009D5A65CE">http://dx.doi.org/10.5252/z2011n1a5</mods:url>
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</mods:location>
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<mods:classification id="F71774C3718B077504C9A44C67080A3F">journal article</mods:classification>
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<mods:identifier id="6E8CAB58C5FCFB7AD3BF82DC17D65BBB" type="DOI">10.5252/z2011n1a5</mods:identifier>
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<mods:identifier id="6FC08DBE5E2D573E5A089AD5C05C95B5" type="ISSN">1638-9387</mods:identifier>
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<mods:identifier id="D10CFFC26CE0F637D96E43911CDC24F3" type="Zenodo-Dep">4546704</mods:identifier>
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<treatment id="03DC9E64DA543A643AEAFF738706E449" ID-DOI="http://doi.org/10.5281/zenodo.4546718" ID-Zenodo-Dep="4546718" LSID="urn:lsid:plazi:treatment:03DC9E64DA543A643AEAFF738706E449" httpUri="http://treatment.plazi.org/id/03DC9E64DA543A643AEAFF738706E449" lastPageNumber="108" pageId="7" pageNumber="108">
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<subSubSection id="C36F7CF9DA543A643AEAFF738135E797" box="[245,515,215,243]" pageId="7" pageNumber="108" type="nomenclature">
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<paragraph id="8BCA2F72DA543A643AEAFF738135E797" blockId="7.[245,515,215,243]" box="[245,515,215,243]" pageId="7" pageNumber="108">
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<heading id="D082981EDA543A643AEAFF738135E797" box="[245,515,215,243]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="108" reason="2">
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<taxonomicName id="4C7554F1DA543A643AEAFF7382F7E796" ID-CoL="G3C6" authority="Montreuil, 2011" authorityName="Montreuil" authorityYear="2011" box="[245,449,215,242]" class="Insecta" family="Scarabaeidae" genus="Arachnodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
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<emphasis id="B901F360DA543A643AEAFF7382F7E796" bold="true" box="[245,449,215,242]" italics="true" pageId="7" pageNumber="108">Arachnodes bicolor</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2324E1BDA543A643BD7FF7C8135E797" box="[456,515,216,243]" pageId="7" pageNumber="108" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F7CF9DA543A643A6EFEB78241E6C8" pageId="7" pageNumber="108" type="materials_examined">
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<paragraph id="8BCA2F72DA543A643A6EFEB78103E604" blockId="7.[112,647,275,352]" pageId="7" pageNumber="108">
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<materialsCitation id="3B1D252FDA543A643A6EFEB78103E604" collectingDate="2006-04-08" collectionCode="MNHN" collectorName="Zombitse & Sakaraha & I. Hanski-group" elevation="774" pageId="7" pageNumber="108" specimenCode="EC1906" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54CE91D0DA543A643A6EFEB783EAE64D" box="[113,220,275,299]" pageId="7" pageNumber="108" type="holotype">HOLOTYPE</typeStatus>
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. —
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<taxonomicName id="4C7554F1DA543A643B13FEB782B5E64F" authority="Montreuil, 2011" authorityName="Montreuil" authorityYear="2011" box="[268,387,275,299]" genus="Madagascar" pageId="7" pageNumber="102" rank="genus" status="gen. nov.">
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<emphasis id="B901F360DA543A643B13FEB782B5E64F" bold="true" box="[268,387,275,299]" pageId="7" pageNumber="108">Madagascar</emphasis>
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</taxonomicName>
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.
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<collectorName id="26804AA4DA543A643B8DFEB782C5E64F" box="[402,499,275,299]" pageId="7" pageNumber="108">Zombitse</collectorName>
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,
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<collectorName id="26804AA4DA543A643BE0FEB7816CE64F" box="[511,602,275,299]" pageId="7" pageNumber="108">Sakaraha</collectorName>
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, dry dec. forest,
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<date id="FFCB09B2DA543A643AC0FE898272E621" box="[223,324,301,325]" pageId="7" pageNumber="108" value="2006-04-08">
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<collectingDate id="EF8FF05ADA543A643AC0FE898272E621" box="[223,324,301,325]" pageId="7" pageNumber="108" value="2006-04-08">8.IV.2006</collectingDate>
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</date>
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, alt.
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<quantity id="4C8D8297DA543A643B70FE89829DE621" box="[367,427,301,325]" metricMagnitude="2" metricUnit="m" metricValue="7.74" pageId="7" pageNumber="108" unit="m" value="774.0">
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<elevation id="0058C841DA543A643B70FE89829DE621" box="[367,427,301,325]" metricMagnitude="2" metricUnit="m" metricValue="7.74" pageId="7" pageNumber="108" unit="m" value="774.0">774m</elevation>
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</quantity>
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,
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<collectingMethod id="52345765DA543A643BACFE8981B5E621" box="[435,643,301,325]" pageId="7" pageNumber="108">fish-baited pitfall trap</collectingMethod>
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,
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<collectorName id="26804AA4DA543A643A6FFEEC823BE604" box="[112,269,328,352]" pageId="7" pageNumber="108">I. Hanski-group</collectorName>
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leg., 1 ơ (
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<collectionCode id="ED64B7B7DA543A643B66FEEC82FBE604" box="[377,461,328,352]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="108">MNHN</collectionCode>
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<specimenCode id="DBD38709DA543A643BCCFEEC811CE604" box="[467,554,328,352]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="108">EC1906</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFEDE8241E6C8" blockId="7.[113,646,378,428]" pageId="7" pageNumber="108">
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<typeStatus id="54CE91D0DA543A643A6EFEDE83EEE6F4" box="[113,216,378,402]" pageId="7" pageNumber="108" type="paratype">PARATYPES</typeStatus>
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. — mêmes données que l’holotype, 6 ơơ et
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<specimenCount id="9D73E4FBDA543A643A6EFE3183A2E6C8" box="[113,148,405,428]" pageId="7" pageNumber="108" type="female">♀♀</specimenCount>
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, (
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<collectionCode id="ED64B7B7DA543A643AB7FE3083CAE6C8" box="[168,252,404,428]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="108">MNHN</collectionCode>
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et
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<collectionCode id="ED64B7B7DA543A643B04FE30825AE6C8" box="[283,364,404,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="108" type="Museum">FMNH</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F7CF9DA543A643A6EFE628706E449" pageId="7" pageNumber="108" type="description">
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<paragraph id="8BCA2F72DA543A643A6EFE628215E69C" blockId="7.[113,646,454,504]" pageId="7" pageNumber="108">ÉTYMOLOGIE. — Cette espèce est nommée en référence à son bicolorisme.</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFD908234E529" blockId="7.[112,647,564,686]" box="[113,258,564,591]" pageId="7" pageNumber="108">DESCRIPTION</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFDF08110E5CA" blockId="7.[112,647,564,686]" pageId="7" pageNumber="108">
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Longueur
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<quantity id="4C8D8297DA543A643AFCFDF0825CE50B" box="[227,362,596,623]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="7" pageNumber="108" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5,5-6,5 mm</quantity>
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. Noir, élytres à léger reflet bleuté, avant-corps vert olive luisant.Tarses, pièces buccales brun-rouge, antennes jaunâtres.
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</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFD7783ADE589" blockId="7.[111,647,723,940]" box="[113,155,723,749]" pageId="7" pageNumber="108">
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<emphasis id="B901F360DA543A643A6EFD7783ADE589" box="[113,155,723,749]" italics="true" pageId="7" pageNumber="108">Tête</emphasis>
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</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFD56820AE4C8" blockId="7.[111,647,723,940]" pageId="7" pageNumber="108">Inerme, clypéus faiblement bidenté.Côtés faiblement anguleux au niveau de la suture clypéo-génale. Tégument très finement microréticulé, à ponctuation très fine, éparse, imperceptible sur les joues, le front et l’arrière du clypéus, à peine plus marquée sur l’avant du clypéus.</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFC7583EEE48E" blockId="7.[111,649,977,1225]" box="[113,216,977,1002]" pageId="7" pageNumber="108">
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<emphasis id="B901F360DA543A643A6EFC7583EEE48E" box="[113,216,977,1002]" italics="true" pageId="7" pageNumber="108">Pronotum</emphasis>
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</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFC54829EE3AD" blockId="7.[111,649,977,1225]" pageId="7" pageNumber="108">Marge basale indistinctement rebordée. Marges latérales et antérieure bien rebordées.Côtés plongeant avant les angles postérieurs, vus du dessus droit dans les deux tiers postérieurs puis convergents. Angles antérieurs droits.Tégument finement microréticulé à ponctuation éparse, fine mais bien visible, un peu plus marquée près de la base.</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFB498385E26C" blockId="7.[113,646,1261,1383]" box="[113,179,1261,1288]" pageId="7" pageNumber="108">
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<emphasis id="B901F360DA543A643A6EFB498385E26C" box="[113,179,1261,1288]" italics="true" pageId="7" pageNumber="108">Élytres</emphasis>
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</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFAA98388E203" blockId="7.[113,646,1261,1383]" pageId="7" pageNumber="108">Stries très fines, à ponctuation peu marquée. Interstries finement microréticulés, sans ponctuation visible.</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EFA288205E162" blockId="7.[113,646,1420,1542]" pageId="7" pageNumber="108">
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<emphasis id="B901F360DA543A643A6EFA2883FAE2C1" box="[113,204,1420,1446]" italics="true" pageId="7" pageNumber="108">8e tergite</emphasis>
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Tégument finement microréticulé, à ponctuation très fine, peu visible sauf au niveau d’une large dépression basale.
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</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EF98F839AE120" blockId="7.[111,648,1579,1700]" box="[113,172,1579,1604]" pageId="7" pageNumber="108">
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<emphasis id="B901F360DA543A643A6EF98F839AE120" box="[113,172,1579,1604]" italics="true" pageId="7" pageNumber="108">Pattes</emphasis>
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</paragraph>
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<paragraph id="8BCA2F72DA543A643A6EF9EE8745E616" blockId="7.[111,648,1579,1700]" lastBlockId="7.[674,1211,216,370]" pageId="7" pageNumber="108">Protibias tridentés sur l’arête externe, la dent médiane très nettement plus rapprochée de l’apicale. Face dorsale sans carène le long de la marge interne.Marge interne légèrement élargie avant l’apex. Angle apical interne des protibias prolongé en une forte saillie dirigée en avant et vers le bas. Éperon apical effilé, courbé vers l’intérieur et également dirigé vers le bas. Métatibias légèrement recourbés vers le bas.</paragraph>
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<paragraph id="8BCA2F72DA543A6438BAFE3281DAE6D5" blockId="7.[677,1210,406,496]" box="[677,748,406,433]" pageId="7" pageNumber="108">
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<emphasis id="B901F360DA543A6438BAFE3281DAE6D5" box="[677,748,406,433]" italics="true" pageId="7" pageNumber="108">Édéage</emphasis>
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</paragraph>
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<paragraph id="8BCA2F72DA543A6438BAFE128057E694" blockId="7.[677,1210,406,496]" pageId="7" pageNumber="108">
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Apex des paramères régulièrement abaissé en vue latérale (
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<figureCitation id="134E33F7DA543A64391FFE718063E694" box="[768,853,469,496]" captionStart="FIG" captionStartId="8.[113,124,1043,1060]" captionTargetBox="[278,1045,215,1000]" captionTargetId="figure-213@8.[277,1033,217,1002]" captionTargetPageId="8" captionText="FIG. 3. — Paramères des Arachnodes, vue dorsale et latérale gauche: A, A. bicolor n. sp.; B, A. luctuosus (Boucomont, 1937); C, A. seyrigi Lebis, 1953; D, A. viettei n. sp.; E, A. zombitsyensis n. sp.; F, A. dichrous Paulian; G, A. mahafalyensis Montreuil, 2006 (vue latérale gauche seule). Échelles: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4546712" httpUri="https://zenodo.org/record/4546712/files/figure.png" pageId="7" pageNumber="108">Fig. 3A</figureCitation>
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).
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</paragraph>
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<paragraph id="8BCA2F72DA543A6438BAFDB18011E54A" blockId="7.[677,1212,533,813]" box="[677,807,533,560]" pageId="7" pageNumber="108">REMARQUES</paragraph>
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<paragraph id="8BCA2F72DA543A6438BAFD908706E449" blockId="7.[677,1212,533,813]" pageId="7" pageNumber="108">
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Par son système de coloration, cette espèce se rapproche d’
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<taxonomicName id="4C7554F1DA543A64392CFDF08783E50B" authority="Paulian, 1976" authorityName="Paulian" authorityYear="1976" box="[819,1205,596,623]" class="Insecta" family="Scarabaeidae" genus="Arachnodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="dichrous">
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<emphasis id="B901F360DA543A64392CFDF0873BE50A" box="[819,1037,596,622]" italics="true" pageId="7" pageNumber="108">Arachnodes dichrous</emphasis>
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Paulian, 1976
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</taxonomicName>
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, mais elle s’en sépare facilement par la forme des paramères et par l’avant-corps d’aspect fortement luisant et non métallique comme chez
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<emphasis id="B901F360DA543A643E20FD17878CE5A9" box="[1087,1210,691,717]" italics="true" pageId="7" pageNumber="108">
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<taxonomicName id="4C7554F1DA543A643E20FD178781E5A9" authorityName="Paulian" authorityYear="1976" box="[1087,1207,691,717]" class="Insecta" family="Scarabaeidae" genus="Arachnodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="dichrous">A. dichrous</taxonomicName>
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.
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</emphasis>
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La forme de l’apex des paramères d’
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<taxonomicName id="4C7554F1DA543A643E20FD77878FE589" authorityName="Paulian" authorityYear="1976" box="[1087,1209,723,749]" class="Insecta" family="Scarabaeidae" genus="Arachnodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="dichrous">
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<emphasis id="B901F360DA543A643E20FD77878FE589" box="[1087,1209,723,749]" italics="true" pageId="7" pageNumber="108">A. dichrous</emphasis>
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</taxonomicName>
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(
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<figureCitation id="134E33F7DA543A6438B3FD5681CCE469" box="[684,762,754,781]" captionStart="FIG" captionStartId="8.[113,124,1043,1060]" captionTargetBox="[278,1045,215,1000]" captionTargetId="figure-213@8.[277,1033,217,1002]" captionTargetPageId="8" captionText="FIG. 3. — Paramères des Arachnodes, vue dorsale et latérale gauche: A, A. bicolor n. sp.; B, A. luctuosus (Boucomont, 1937); C, A. seyrigi Lebis, 1953; D, A. viettei n. sp.; E, A. zombitsyensis n. sp.; F, A. dichrous Paulian; G, A. mahafalyensis Montreuil, 2006 (vue latérale gauche seule). Échelles: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4546712" httpUri="https://zenodo.org/record/4546712/files/figure.png" pageId="7" pageNumber="108">Fig. 3F</figureCitation>
|
||
) se rapproche davantage de celle observée chez
|
||
<taxonomicName id="4C7554F1DA543A6438C4FCB68088E449" authority="Lebis, 1953" authorityName="Lebis" authorityYear="1953" box="[731,958,786,813]" class="Insecta" family="Scarabaeidae" genus="Arachnodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="seyrigi">
|
||
<emphasis id="B901F360DA543A6438C4FCB6800FE448" box="[731,825,786,812]" italics="true" pageId="7" pageNumber="108">A. seyrigi</emphasis>
|
||
<bibRefCitation id="EFE45283DA543A64395FFCB68088E449" author="LEBIS E." box="[832,958,786,813]" pageId="7" pageNumber="108" pagination="107 - 252" refId="ref13078" refString="LEBIS E. 1953. - Revision des Canthoninae de Madagascar. Memoires de l'Institut scientifique de Madagascar, Serie E. 3: 107 - 252." type="journal article" year="1953">Lebis, 1953</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134E33F7DA543A6439D2FCB68714E449" box="[973,1058,786,813]" captionStart="FIG" captionStartId="8.[113,124,1043,1060]" captionTargetBox="[278,1045,215,1000]" captionTargetId="figure-213@8.[277,1033,217,1002]" captionTargetPageId="8" captionText="FIG. 3. — Paramères des Arachnodes, vue dorsale et latérale gauche: A, A. bicolor n. sp.; B, A. luctuosus (Boucomont, 1937); C, A. seyrigi Lebis, 1953; D, A. viettei n. sp.; E, A. zombitsyensis n. sp.; F, A. dichrous Paulian; G, A. mahafalyensis Montreuil, 2006 (vue latérale gauche seule). Échelles: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4546712" httpUri="https://zenodo.org/record/4546712/files/figure.png" pageId="7" pageNumber="108">Fig. 3C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |