473 lines
58 KiB
XML
473 lines
58 KiB
XML
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<mods:title id="97798CF51ABF26267B22742FA8B69DA1">Revision of Hestiasulini Giglio-Tos, 1915 stat. rev. (Insecta: Mantodea: Hymenopodidae) of Borneo, with description of new taxa and comments on the taxonomy of the tribe</mods:title>
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<mods:namePart id="B583DAEC7C4822B40B7226C2AC89FA36">Schwarz, Christian J.</mods:namePart>
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<mods:namePart id="7D80E5A929BE395556EFEF74081CDB8F">Shcherbakov, Evgeny</mods:namePart>
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<mods:title id="14A7051C23ACD6E49AF67D47DFB0070C">Zootaxa</mods:title>
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<mods:date id="C9F6D31A528390494F1D2DA2B43B522D">2017</mods:date>
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<mods:number id="EB6D08A94AA278B99D66C48CD96251FB">4291</mods:number>
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<treatment id="03DC81241803A17AFF6BFB8EFE26671B" ID-DOI="http://doi.org/10.5281/zenodo.6050951" ID-GBIF-Taxon="132549693" ID-Zenodo-Dep="6050951" LSID="urn:lsid:plazi:treatment:03DC81241803A17AFF6BFB8EFE26671B" httpUri="http://treatment.plazi.org/id/03DC81241803A17AFF6BFB8EFE26671B" lastPageId="10" lastPageNumber="253" pageId="8" pageNumber="251">
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<subSubSection id="C36F63B91803A178FF6BFB8EFE2E671D" pageId="8" pageNumber="251" type="nomenclature">
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<paragraph id="8BCA30321803A178FF6BFB8EFD9E673C" blockId="8.[151,625,1087,1147]" box="[151,625,1087,1114]" pageId="8" pageNumber="251">
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<heading id="D082875E1803A178FF6BFB8EFD9E673C" bold="true" box="[151,625,1087,1114]" fontSize="11" level="1" pageId="8" pageNumber="251" reason="1">
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<taxonomicName id="4C754BB11803A178FF6BFB8EFD9E673C" ID-CoL="RTPT" authority="Giglio-Tos, 1915" authorityName="Giglio-Tos" authorityYear="1915" box="[151,625,1087,1114]" class="Insecta" family="Hymenopodidae" genus="Catestiasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="8" pageNumber="251" phylum="Arthropoda" rank="species" species="moultoni">
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<emphasis id="B901EC201803A178FF6BFB8EFD9E673C" bold="true" box="[151,625,1087,1114]" pageId="8" pageNumber="251">
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<emphasis id="B901EC201803A178FF6BFB8EFE70673F" bold="true" box="[151,415,1087,1113]" italics="true" pageId="8" pageNumber="251">Catestiasula moultoni</emphasis>
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<bibRefCitation id="EFE44DC31803A178FE5BFBF1FD9E673C" author="Giglio-Tos" box="[423,625,1088,1114]" pageId="8" pageNumber="251" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos, 1915</bibRefCitation>
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</emphasis>
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</paragraph>
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<paragraph id="8BCA30321803A178FF6BFBD3FE2E671D" blockId="8.[151,625,1087,1147]" box="[151,449,1122,1147]" pageId="8" pageNumber="251">
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(
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<figureCitation id="134E2CB71803A178FF63FBD3FF08671D" box="[159,231,1122,1147]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="8" pageNumber="251">Figs 8</figureCitation>
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,
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<figureCitation id="134E2CB71803A178FF0FFBD2FED3671D" box="[243,316,1123,1147]" captionStart="FIGURES 9 – 20" captionStartId="7.[151,264,1281,1304]" captionTargetBox="[206,1384,223,1244]" captionTargetId="figure@7.[182,1404,195,1257]" captionTargetPageId="7" captionText="FIGURES 9 – 20. Fore legs of Hestiasulini treated in this paper, anterior view. 9. A. phyllopus, male from the Malay Peninsula (ES). 10. A. phyllopus, male from Brunei, mirrored (CS). 11. A. phyllopus, female from Brunei (CS). 12. A. sarawaca, male from Brunei (CS). 13. P. borneana, holotype (SMNK). 14. H. brunneriana, male from Sri Lanka (CS). 15. H. brunneriana, female from Sri Lanka (CS). 16. A. sarawaca, female from Kalimantan; Photo C. J. Schwarz © RMNH (RMNH). 17. C. moultoni, male from Sarawak (CS). 18. C. moultoni, female from Brunei, mirrored (CS). 19. C. nitida, male from Myanmar (SMNK). 20. C. nitida, male from Sumatra (SMNK) (not to scale)." httpUri="https://zenodo.org/record/829535/files/figure.png" pageId="8" pageNumber="251">17–18</figureCitation>
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,
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<figureCitation id="134E2CB71803A178FEBBFBD2FE8A671D" box="[327,357,1123,1147]" captionStart="FIGURES 21 – 22" captionStartId="9.[151,264,1104,1127]" captionTargetBox="[180,1396,199,1063]" captionTargetId="figure@9.[159,1428,173,1083]" captionTargetPageId="9" captionText="FIGURES 21 – 22. Subgenital plate and genitalia of 21. H. brunneriana from India (ES) and 22. C. moultoni from Sabah (ES). Arrows indicate weakly sclerotized membrane at base of sclerotized left margin of ventral phallomere (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829537/files/figure.png" pageId="8" pageNumber="251">22</figureCitation>
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,
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<figureCitation id="134E2CB71803A178FE8DFBD3FE60671D" box="[369,399,1122,1147]" captionStart="FIGURES 51 – 52" captionStartId="23.[151,264,1634,1657]" captionTargetBox="[359,1219,469,1613]" captionTargetId="figure@23.[340,1247,469,1613]" captionTargetPageId="23" captionText="FIGURES 51 – 52. Life aspect of Catestiasula. 51. Male of C. moultoni, Mt. Trus Madi, Sabah, Malaysia; Photo © Alexey Yakovlev. 52. Male of unidentified Catestiasula specimen from Ulu Gombak NP, Selangor, Malaysia; Photo © Bernhard Jacobi." httpUri="https://zenodo.org/record/829551/files/figure.png" pageId="8" pageNumber="251">51</figureCitation>
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,
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<figureCitation id="134E2CB71803A178FE67FBD3FE57671D" box="[411,440,1122,1147]" captionStart="FIGURES 60 – 62" captionStartId="27.[151,264,1721,1744]" captionTargetBox="[176,1416,359,1677]" captionTargetId="figure@27.[151,1436,339,1700]" captionTargetPageId="27" captionText="FIGURES 60 – 62. Distribution maps of Bornean Hestiasulini. 60. C. moultoni (circles) and C. nitida (squares); open square indicates unprecise locality information. 61. A. phyllopus. 62. A. sarawaca (circles) and P. borneana (square); open circle indicates unprecise locality information." httpUri="https://zenodo.org/record/829557/files/figure.png" pageId="8" pageNumber="251">60</figureCitation>
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)
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</paragraph>
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<subSubSection id="C36F63B91803A178FF6BFB1BFB076798" pageId="8" pageNumber="251" type="reference_group">
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<paragraph id="8BCA30321803A178FF6BFB1BFDB267A6" blockId="8.[151,1256,1194,1278]" box="[151,605,1194,1216]" pageId="8" pageNumber="251">
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<treatmentCitationGroup id="AB65171C1803A178FF6BFB1BFDB267A6" box="[151,605,1194,1216]" pageId="8" pageNumber="251">
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<taxonomicName id="4C754BB11803A178FF6BFB1BFDB667A6" ID-CoL="RTPT" authority="Giglio-Tos 1915: 101" authorityName="Giglio-Tos" authorityPageNumber="101" authorityYear="1915" box="[151,601,1194,1216]" class="Insecta" family="Hymenopodidae" genus="Catestiasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="8" pageNumber="251" phylum="Arthropoda" rank="species" species="moultoni">
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<emphasis id="B901EC201803A178FF6BFB1BFE9967A6" box="[151,374,1194,1216]" italics="true" pageId="8" pageNumber="251">Catestiasula moultoni</emphasis>
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<treatmentCitation id="0AD416231803A178FE80FB1BFDB667A6" author="Giglio-Tos" box="[380,601,1194,1216]" page="101" pageId="8" pageNumber="251" year="1915">
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<bibRefCitation id="EFE44DC31803A178FE80FB1BFDC967A6" author="Giglio-Tos" box="[380,550,1194,1216]" pageId="8" pageNumber="251" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos 1915</bibRefCitation>
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||
: 101
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||
</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BCA30321803A178FF6BFB78FCD767B9" blockId="8.[151,1256,1194,1278]" box="[151,824,1225,1247]" pageId="8" pageNumber="251">
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<treatmentCitationGroup id="AB65171C1803A178FF6BFB78FCD767B9" box="[151,824,1225,1247]" pageId="8" pageNumber="251">
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<taxonomicName id="4C754BB11803A178FF6BFB78FDC367B9" ID-CoL="RTPT" authority="Giglio-Tos, 1915" authorityName="Giglio-Tos" authorityYear="1915" box="[151,556,1225,1247]" class="Insecta" family="Hymenopodidae" genus="Catestiasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="8" pageNumber="251" phylum="Arthropoda" rank="species" species="moultoni">
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<emphasis id="B901EC201803A178FF6BFB78FE9967B9" box="[151,374,1225,1247]" italics="true" pageId="8" pageNumber="251">Catestiasula moultoni</emphasis>
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<bibRefCitation id="EFE44DC31803A178FE80FB78FDC367B9" author="Giglio-Tos" box="[380,556,1225,1247]" pageId="8" pageNumber="251" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos, 1915</bibRefCitation>
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</taxonomicName>
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. In:
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<treatmentCitation id="0AD416231803A178FDA4FB78FCDB67B9" author="Giglio-Tos" box="[600,820,1225,1247]" page="547" pageId="8" pageNumber="251" year="1927">
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<bibRefCitation id="EFE44DC31803A178FDA4FB78FCED67B9" author="Giglio-Tos" box="[600,770,1225,1247]" pageId="8" pageNumber="251" refString="Giglio-Tos, E. (1927) Das Tierreich. 50. Lfg. Orthoptera Mantidae. Walter de Gruyter & Co., Berlin & Leipzig, XL + 707 pp." type="book" year="1927">Giglio-Tos 1927</bibRefCitation>
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: 547
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BCA30321803A178FF6BFB59FB076798" blockId="8.[151,1256,1194,1278]" box="[151,1256,1256,1278]" pageId="8" pageNumber="251">
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<treatmentCitationGroup id="AB65171C1803A178FF6BFB59FB076798" box="[151,1256,1256,1278]" pageId="8" pageNumber="251">
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<taxonomicName id="4C754BB11803A178FF6BFB59FDC56798" authority="Giglio-Tos, 1915" authorityName="Giglio-Tos" authorityYear="1915" box="[151,554,1256,1278]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="8" pageNumber="251" phylum="Arthropoda" rank="species" species="moultoni">
|
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<emphasis id="B901EC201803A178FF6BFB59FE8C6798" box="[151,355,1256,1278]" italics="true" pageId="8" pageNumber="251">Hestiasula moultoni</emphasis>
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(
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<bibRefCitation id="EFE44DC31803A178FE8CFB59FDCD6798" author="Giglio-Tos" box="[368,546,1256,1278]" pageId="8" pageNumber="251" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos, 1915</bibRefCitation>
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)
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</taxonomicName>
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. In:
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<treatmentCitation id="0AD416231803A178FDAAFB59FD006798" author="Beier" box="[598,751,1256,1278]" page="17" pageId="8" pageNumber="251" year="1934">
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<bibRefCitation id="EFE44DC31803A178FDAAFB59FD266798" author="Beier" box="[598,713,1256,1278]" pageId="8" pageNumber="251" refString="Beier, M. (1934) Mantodea. Fam. Mantidae. Subfam. Hymenopodinae. Genera Insectorum, Fascicule 196, 1 - 37, pls. 1 - 2 [Nachtrage 1937, 3 pp.]" type="book chapter" year="1934">Beier 1934</bibRefCitation>
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: 17
|
||
</treatmentCitation>
|
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;
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||
<treatmentCitation id="0AD416231803A178FD06FB59FC296798" author="Ehrmann" box="[762,966,1256,1278]" page="171" pageId="8" pageNumber="251" year="2002">
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<bibRefCitation id="EFE44DC31803A178FD06FB59FC7B6798" author="Ehrmann" box="[762,916,1256,1278]" pageId="8" pageNumber="251" refString="Ehrmann, R. (2002) Mantodea - Gottesanbeterinnen der Welt. Natur und Tier-Verlag, Munster, 519 pp." type="book" year="2002">Ehrmann 2002</bibRefCitation>
|
||
: 171
|
||
</treatmentCitation>
|
||
;
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||
<treatmentCitation id="0AD416231803A178FC2CFB59FB0B6798" author="Otte" box="[976,1252,1256,1278]" page="78" pageId="8" pageNumber="251" year="2005">
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<bibRefCitation id="EFE44DC31803A178FC2CFB59FB516798" author="Otte" box="[976,1214,1256,1278]" pageId="8" pageNumber="251" refString="Otte, D. & Spearman, L. (2005) Mantida Species File. Catalog of the Mantids of the World. Publication Number 1. Association of the Insect Diversity, Philadelphia, 489 pp." type="book" year="2005">Otte & Spearman 2005</bibRefCitation>
|
||
: 78
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</treatmentCitation>
|
||
.
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||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F63B91803A178FF6BFA9BFCC765E1" pageId="8" pageNumber="251" type="materials_examined">
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<paragraph id="8BCA30321803A178FF6BFA9BFCC765E1" blockId="8.[151,1437,1322,2031]" pageId="8" pageNumber="251">
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<materialsCitation id="3B1D3A6F1803A178FF6BFA9BFDF36624" ID-GBIF-Occurrence="1572326316" box="[151,540,1322,1347]" collectorName="Material." country="Malaysia" location="Malaysia" pageId="8" pageNumber="251" specimenCount="1" specimenCount-male="1">
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<collectorName id="268055E41803A178FF6BFA9BFEE96625" box="[151,262,1322,1347]" pageId="8" pageNumber="251">
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<emphasis id="B901EC201803A178FF6BFA9BFEE96625" bold="true" box="[151,262,1322,1347]" pageId="8" pageNumber="251">Material.</emphasis>
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</collectorName>
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<specimenCount id="9D73FBBB1803A178FEECFA9AFEAF6624" box="[272,320,1323,1347]" pageId="8" pageNumber="251" type="male">1 ♂</specimenCount>
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,
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||
<collectingCountry id="F36270A21803A178FEB3FA9BFE566625" box="[335,441,1322,1347]" name="Malaysia" pageId="8" pageNumber="251">Malaysia</collectingCountry>
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, Borneo
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</materialsCitation>
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||
, Sarawak, Lanjak Entimau Wildlife Sanctuary,
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||
<materialsCitation id="3B1D3A6F1803A178FBB9FA9BFE7066EC" ID-GBIF-Occurrence="1572326336" collectingDate="2008-06-15" collectingDateMax="2008-06-27" collectingDateMin="2008-06-15" collectorName="O. Konopik" country="Malaysia" location="Sarawak" pageId="8" pageNumber="251" specimenCount="1" specimenCount-female="1">
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<collectingCountry id="F36270A21803A178FBB9FA9BFB4A6625" box="[1093,1189,1322,1347]" name="Malaysia" pageId="8" pageNumber="251">Sarawak</collectingCountry>
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Forestry Department Research Station, at station lights, N 01˚38’43.58”–E 112˚16’41.98”,
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<collectingDate id="EF8FEF1A1803A178FC29FAFFFB676601" box="[981,1160,1358,1383]" pageId="8" pageNumber="251" value="2008-06-15" valueMax="2008-06-27" valueMin="2008-06-15">
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<date id="FFCB16F21803A178FC29FAFFFB676601" box="[981,1160,1358,1383]" pageId="8" pageNumber="251" value="2008-06-15" valueMax="2008-06-27" valueMin="2008-06-15">15.–27.VI.2008</date>
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</collectingDate>
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,
|
||
<collectorName id="268055E41803A178FB66FAFEFACA6601" box="[1178,1317,1358,1383]" pageId="8" pageNumber="251">O. Konopik</collectorName>
|
||
leg. (CS; pinned); 1 damaged
|
||
<specimenCount id="9D73FBBB1803A178FE76FAC2FE7066EC" box="[394,415,1395,1418]" pageId="8" pageNumber="251" type="female">♀</specimenCount>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B1D3A6F1803A178FE4DFAC3FD5266C8" ID-GBIF-Occurrence="1572326304" collectingDate="2013-02" collectorName="O. Konopik" country="Brunei" location="Kuala Belalong Field Studies Centre" municipality="Borneo" pageId="8" pageNumber="251" specimenCount="3" specimenCount-male="3" stateProvince="Temburong">
|
||
<collectingCountry id="F36270A21803A178FE4DFAC3FDEF66ED" box="[433,512,1394,1419]" name="Brunei" pageId="8" pageNumber="251">Brunei</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6BAEAA481803A178FDECFAC2FD8A66EC" box="[528,613,1395,1418]" pageId="8" pageNumber="251">Borneo</collectingMunicipality>
|
||
, Ulu
|
||
<collectingRegion id="49B1FED01803A178FD57FAC3FCC366ED" box="[683,812,1394,1419]" country="Brunei" name="Temburong" pageId="8" pageNumber="251">Temburong</collectingRegion>
|
||
National Park,
|
||
<location id="8EAA66E91803A178FC15FAC3FA7866ED" LSID="urn:lsid:plazi:treatment:03DC81241803A17AFF6BFB8EFE26671B:8EAA66E91803A178FC15FAC3FA7866ED" box="[1001,1431,1394,1419]" country="Brunei" municipality="Borneo" name="Kuala Belalong Field Studies Centre" pageId="8" pageNumber="251" stateProvince="Temburong">Kuala Belalong Field Studies Centre</location>
|
||
,
|
||
<collectingDate id="EF8FEF1A1803A178FF6BFA26FF0466C9" box="[151,235,1431,1455]" pageId="8" pageNumber="251" value="2013-02">
|
||
<date id="FFCB16F21803A178FF6BFA26FF0466C9" box="[151,235,1431,1455]" pageId="8" pageNumber="251" value="2013-02">II.2013</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="268055E41803A178FF00FA26FE6866C9" box="[252,391,1430,1455]" pageId="8" pageNumber="251">O. Konopik</collectorName>
|
||
leg. (CS; pinned);
|
||
<specimenCount id="9D73FBBB1803A178FD8AFA26FD5266C8" box="[630,701,1431,1455]" pageId="8" pageNumber="251" type="male">3 ♂♂</specimenCount>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B1D3A6F1803A178FD33FA27FEC96691" ID-GBIF-Occurrence="1572326309" collectingDate="2011-03-28" collectingDateMax="2011-04-10" collectingDateMin="2011-03-28" collectorName="P. Udovichenko & Borneo Jungle Girl" country="Malaysia" county="Borneo" elevation="1150" latitude="5.4430556" location="Trus Madi Forest Reserve" municipality="Keningau distr." pageId="8" pageNumber="251" specimenCount="1" specimenCount-male="1" stateProvince="Sabah">
|
||
<collectingCountry id="F36270A21803A178FD33FA27FCD666C9" box="[719,825,1430,1455]" name="Malaysia" pageId="8" pageNumber="251">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCounty id="62AB48BE1803A178FCB6FA26FC7066C8" box="[842,927,1431,1454]" pageId="8" pageNumber="251">Borneo</collectingCounty>
|
||
,
|
||
<collectingRegion id="49B1FED01803A178FC4CFA27FC1866C9" box="[944,1015,1430,1455]" country="Malaysia" name="Sabah" pageId="8" pageNumber="251">Sabah</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BAEAA481803A178FBF5FA27FB5266C9" box="[1033,1213,1430,1455]" pageId="8" pageNumber="251">Keningau distr.</collectingMunicipality>
|
||
,
|
||
<location id="8EAA66E91803A178FB32FA26FF1B66B4" LSID="urn:lsid:plazi:treatment:03DC81241803A17AFF6BFB8EFE26671B:8EAA66E91803A178FB32FA26FF1B66B4" country="Malaysia" county="Borneo" latitude="5.4430556" municipality="Keningau distr." name="Trus Madi Forest Reserve" pageId="8" pageNumber="251" stateProvince="Sabah">
|
||
<location id="8EAA66E91803A178FB32FA26FA7366C8" LSID="urn:lsid:plazi:treatment:03DC81241803A17AFF6BFB8EFE26671B:8EAA66E91803A178FB32FA26FA7366C8" box="[1230,1436,1430,1455]" country="Malaysia" county="Borneo" latitude="5.4430556" municipality="Keningau distr." name="Trus Madi Forest" pageId="8" pageNumber="251" stateProvince="Sabah">Trus Madi Forest</location>
|
||
Reserve
|
||
</location>
|
||
,
|
||
<elevation id="0058D7011803A178FF03FA0BFEBA66B4" box="[255,341,1466,1491]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="251" unit="m" value="1150.0">
|
||
<quantity id="4C8D9DD71803A178FF03FA0BFEBA66B4" box="[255,341,1466,1491]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="251" unit="m" value="1150.0">1150 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EF8FEF1A1803A178FE9CFA0AFDC466B5" box="[352,555,1467,1491]" pageId="8" pageNumber="251" value="2011-03-28" valueMax="2011-04-10" valueMin="2011-03-28">
|
||
<date id="FFCB16F21803A178FE9CFA0AFDC466B5" box="[352,555,1467,1491]" pageId="8" pageNumber="251" value="2011-03-28" valueMax="2011-04-10" valueMin="2011-03-28">28.III–10.IV.2011</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="268055E41803A178FDCAFA0AFD0866B5" box="[566,743,1466,1491]" pageId="8" pageNumber="251">P. Udovichenko</collectorName>
|
||
leg. (ES; pinned);
|
||
<specimenCount id="9D73FBBB1803A178FC46FA0AFC0966B4" box="[954,998,1467,1491]" pageId="8" pageNumber="251" type="male">1 ♂</specimenCount>
|
||
, same place,
|
||
<collectorName id="268055E41803A178FB80FA0AFABD66B5" box="[1148,1362,1466,1491]" pageId="8" pageNumber="251">Borneo Jungle Girl</collectorName>
|
||
camp,
|
||
<geoCoordinate id="EE4156F51803A178FF6BFA6EFEC96691" box="[151,294,1502,1527]" direction="north" orientation="latitude" pageId="8" pageNumber="251" precision="15" value="5.4430556">N 05°26’35”</geoCoordinate>
|
||
</materialsCitation>
|
||
–E 116°27’5”,
|
||
<quantity id="4C8D9DD71803A178FE30FA6FFDCD6690" box="[460,546,1502,1527]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="251" unit="m" value="1250.0">1250 m</quantity>
|
||
,
|
||
<date id="FFCB16F21803A178FDD1FA6EFD6D6691" box="[557,642,1503,1527]" pageId="8" pageNumber="251" value="2014-02">II.2014</date>
|
||
, A. Klimenko leg. (ES; pinned); 20 ♂♂, same place,
|
||
<date id="FFCB16F21803A178FB2FFA6EFA956691" box="[1235,1402,1503,1527]" pageId="8" pageNumber="251" value="2014-05-22" valueMax="2014-05-27" valueMin="2014-05-22">22.–27.V.2014</date>
|
||
, E. Shcherbakov leg. (ES; ethanol);
|
||
<materialsCitation id="3B1D3A6F1803A178FDD0F9B2FEFB6559" ID-GBIF-Occurrence="1572326331" country="Malaysia" county="Borneo" latitude="4.5824666" location="Pensiangan" municipality="Pensiangan distr." pageId="8" pageNumber="251" specimenCount="3" specimenCount-male="3" stateProvince="Sabah">
|
||
<specimenCount id="9D73FBBB1803A178FDD0F9B2FD98657C" box="[556,631,1539,1563]" pageId="8" pageNumber="251" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<collectingCountry id="F36270A21803A178FD71F9B3FD19657D" box="[653,758,1538,1563]" name="Malaysia" pageId="8" pageNumber="251">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCounty id="62AB48BE1803A178FCF7F9B2FC8F657C" box="[779,864,1539,1562]" pageId="8" pageNumber="251">Borneo</collectingCounty>
|
||
,
|
||
<collectingRegion id="49B1FED01803A178FC8AF9B3FC52657D" box="[886,957,1538,1563]" country="Malaysia" name="Sabah" pageId="8" pageNumber="251">Sabah</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BAEAA481803A178FC2EF9B3FB72657D" box="[978,1181,1538,1563]" pageId="8" pageNumber="251">Pensiangan distr.</collectingMunicipality>
|
||
,
|
||
<location id="8EAA66E91803A178FB4EF9B3FADE657D" LSID="urn:lsid:plazi:treatment:03DC81241803A17AFF6BFB8EFE26671B:8EAA66E91803A178FB4EF9B3FADE657D" box="[1202,1329,1538,1563]" country="Malaysia" county="Borneo" latitude="4.5824666" municipality="Pensiangan distr." name="Pensiangan" pageId="8" pageNumber="251" stateProvince="Sabah">Pensiangan</location>
|
||
env.,
|
||
<geoCoordinate id="EE4156F51803A178FA74F9B2FEFB6559" direction="north" orientation="latitude" pageId="8" pageNumber="251" precision="1" value="4.5824666">N 04°34.948’</geoCoordinate>
|
||
</materialsCitation>
|
||
–E 116°19.935’,
|
||
<quantity id="4C8D9DD71803A178FE2AF997FDCD6558" box="[470,546,1574,1599]" metricMagnitude="2" metricUnit="m" metricValue="5.16" pageId="8" pageNumber="251" unit="m" value="516.0">516 m</quantity>
|
||
,
|
||
<date id="FFCB16F21803A178FDCCF996FD056559" box="[560,746,1575,1599]" pageId="8" pageNumber="251" value="2014-05-27" valueMax="2014-06-02" valueMin="2014-05-27">27.V–2.VI.2014</date>
|
||
, E. Shcherbakov leg. (ES, ethanol);
|
||
<materialsCitation id="3B1D3A6F1803A178FB73F996FDEF6505" ID-GBIF-Occurrence="1572326297" country="Malaysia" latitude="45.95" location="Danum Valley" municipality="Borneo" pageId="8" pageNumber="251" specimenCount="1" specimenCount-male="1" stateProvince="Sabah">
|
||
<specimenCount id="9D73FBBB1803A178FB73F996FB526558" box="[1167,1213,1575,1599]" pageId="8" pageNumber="251" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingCountry id="F36270A21803A178FB30F997FADA6559" box="[1228,1333,1574,1599]" name="Malaysia" pageId="8" pageNumber="251">Malaysia</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6BAEAA481803A178FABFF996FA776558" box="[1347,1432,1575,1598]" pageId="8" pageNumber="251">Borneo</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49B1FED01803A178FF6BF9FBFF316505" box="[151,222,1610,1635]" country="Malaysia" name="Sabah" pageId="8" pageNumber="251">Sabah</collectingRegion>
|
||
,
|
||
<location id="8EAA66E91803A178FF16F9FAFE626505" LSID="urn:lsid:plazi:treatment:03DC81241803A17AFF6BFB8EFE26671B:8EAA66E91803A178FF16F9FAFE626505" box="[234,397,1610,1635]" country="Malaysia" latitude="45.95" municipality="Borneo" name="Danum Valley" pageId="8" pageNumber="251" stateProvince="Sabah">Danum Valley</location>
|
||
,
|
||
<geoCoordinate id="EE4156F51803A178FE64F9FAFDEF6505" box="[408,512,1610,1635]" direction="north" orientation="latitude" pageId="8" pageNumber="251" precision="925" value="45.95">N 45°57’</geoCoordinate>
|
||
</materialsCitation>
|
||
–E 117°47’,
|
||
<quantity id="4C8D9DD71803A178FD70F9FAFD396504" box="[652,726,1611,1635]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="251" unit="m" value="220.0">220 m</quantity>
|
||
, primary forest, canopy
|
||
<quantity id="4C8D9DD71803A178FC14F9FAFBCB6504" box="[1000,1060,1611,1635]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="251" unit="m" value="40.0">40 m</quantity>
|
||
, lux,
|
||
<date id="FFCB16F21803A178FB9EF9FAFB0C6505" box="[1122,1251,1611,1635]" pageId="8" pageNumber="251" value="2003-03-22">22.III.2003</date>
|
||
, M. Helmkampf leg. (genitalia preparation Schwarz No. 189) (CS; pinned).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F63B91803A178FF3BF923FAD765CD" box="[199,1336,1682,1707]" pageId="8" pageNumber="251" type="distribution">
|
||
<paragraph id="8BCA30321803A178FF3BF923FAD765CD" blockId="8.[151,1437,1322,2031]" box="[199,1336,1682,1707]" pageId="8" pageNumber="251">
|
||
<emphasis id="B901EC201803A178FF3BF923FE8F65CD" bold="true" box="[199,352,1682,1707]" pageId="8" pageNumber="251">Distribution.</emphasis>
|
||
N Borneo (
|
||
<bibRefCitation id="EFE44DC31803A178FE1FF923FD4F65CD" author="Giglio-Tos" box="[483,672,1682,1707]" pageId="8" pageNumber="251" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos 1915</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE44DC31803A178FD57F922FD0965CD" author="Giglio-Tos" box="[683,742,1683,1707]" pageId="8" pageNumber="251" refString="Giglio-Tos, E. (1927) Das Tierreich. 50. Lfg. Orthoptera Mantidae. Walter de Gruyter & Co., Berlin & Leipzig, XL + 707 pp." type="book" year="1927">1927</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE44DC31803A178FD0DF923FC1865CD" author="Helmkampf" box="[753,1015,1682,1707]" pageId="8" pageNumber="251" refString="Helmkampf, M., Schwarz, C. J. & Beck, J. (2007) A first look at the biodiversity of praying mantids (Insecta: Mantodea) in Sabah, Borneo. Sepilok Bulletin, 7, 1 - 13." type="journal article" year="2007">
|
||
Helmkampf
|
||
<emphasis id="B901EC201803A178FC81F925FC4165CD" box="[893,942,1682,1707]" italics="true" pageId="8" pageNumber="251">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE44DC31803A178FBFEF923FAC365CD" author="Schwarz" box="[1026,1324,1682,1707]" pageId="8" pageNumber="251" refString="Schwarz, C. J. & Konopik, O. (2014) An annotated checklist of the praying mantises (Mantodea) of Borneo, including the results of the 2008 scientific expedition to Lanjak Entimau Wildlife Sanctuary, Sarawak. Zootaxa, 3797 (1), 130 - 168. https: // doi. org / 10.11646 / zootaxa. 3797.1.12" type="journal article" year="2014">Schwarz & Konopik 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F63B91803A17AFF3BF907FCEC63D6" lastPageId="10" lastPageNumber="253" pageId="8" pageNumber="251" type="description">
|
||
<paragraph id="8BCA30321803A178FF3BF907FEB3645D" blockId="8.[151,1437,1322,2031]" pageId="8" pageNumber="251">
|
||
<emphasis id="B901EC201803A178FF3BF907FE9965A9" bold="true" box="[199,374,1718,1743]" pageId="8" pageNumber="251">Measurements</emphasis>
|
||
in mm.
|
||
<emphasis id="B901EC201803A178FE2FF907FDE265A9" box="[467,525,1718,1743]" italics="true" pageId="8" pageNumber="251">Male</emphasis>
|
||
: body length 16.3–18.6; head width 3.2–3.3; pronotum length 2.5–2.6; pronotum width 2.3–2.4; metazona 1.2; fore coxa length 4.3–4.4; fore femur length 5.4–5.6; fore femur width 2.9–3.0; fore tibia length 3.4–3.6; hind femur length 3.9–4.2; fore wing length 15.0–16.7; fore wing width 4.2–4.4; hind wing length 14.2–14.6.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321803A178FF3BF8F7FB0A64E5" blockId="8.[151,1437,1322,2031]" pageId="8" pageNumber="251">
|
||
<emphasis id="B901EC201803A178FF3BF8F7FEF46439" box="[199,283,1862,1887]" italics="true" pageId="8" pageNumber="251">Female</emphasis>
|
||
: fore femur length 6.8; fore femur width 3.7; meso femur length 4.1; meso tibia length 3.7; hind femur length 4.9; hind tibia length 5.3; fore wing length 18.2; fore wing width 4.8; costal field width 0.9.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321803A178FF3BF83EFD176489" blockId="8.[151,1437,1322,2031]" pageId="8" pageNumber="251">
|
||
Among a collection of mantodeans from
|
||
<collectingCountry id="F36270A21803A178FD6BF83FFD0C64C1" box="[663,739,1934,1959]" name="Brunei" pageId="8" pageNumber="251">Brunei</collectingCountry>
|
||
submitted to CS for study was a damaged female attributable to this species. Despite its bad condition, it is useful to give a description of the specimen in order to complete the generic and specific diagnosis (see also
|
||
<bibRefCitation id="EFE44DC31803A178FDABF866FD046489" author="Werner" box="[599,747,2007,2031]" pageId="8" pageNumber="251" refString="Werner, F. (1933) Funfter Beitrag zur Kenntnis der Mantodeen von Niederlandisch-Indien. Treubia, 14 (2), 255 - 272, pl. 8." type="journal article" year="1933">Werner 1933</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0A60BA1802A179FF6BFBE1FAB167E0" httpUri="https://zenodo.org/record/829537/files/figure.png" pageId="9" pageNumber="252" targetBox="[180,1396,199,1063]" targetPageId="9">
|
||
<paragraph id="8BCA30321802A179FF6BFBE1FAB167E0" blockId="9.[151,1436,1104,1158]" pageId="9" pageNumber="252">
|
||
<emphasis id="B901EC201802A179FF6BFBE1FEBB6700" bold="true" box="[151,340,1104,1127]" pageId="9" pageNumber="252">FIGURES 21–22.</emphasis>
|
||
Subgenital plate and genitalia of 21.
|
||
<taxonomicName id="4C754BB11802A179FD33FBE3FC816701" box="[719,878,1105,1127]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="9" pageNumber="252" phylum="Arthropoda" rank="species" species="brunneriana">
|
||
<emphasis id="B901EC201802A179FD33FBE3FC816701" box="[719,878,1105,1127]" italics="true" pageId="9" pageNumber="252">H. brunneriana</emphasis>
|
||
</taxonomicName>
|
||
from India (ES) and 22.
|
||
<taxonomicName id="4C754BB11802A179FB90FBE3FB0B6701" box="[1132,1252,1105,1127]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="9" pageNumber="252" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201802A179FB90FBE3FB0B6701" box="[1132,1252,1105,1127]" italics="true" pageId="9" pageNumber="252">C. moultoni</emphasis>
|
||
</taxonomicName>
|
||
from Sabah (ES). Arrows indicate weakly sclerotized membrane at base of sclerotized left margin of ventral phallomere (scale bar: 1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCA30321802A179FF3BFB03FF1E6789" blockId="9.[151,1437,1202,2019]" pageId="9" pageNumber="252">
|
||
<emphasis id="B901EC201802A179FF3BFB03FE3B67AD" bold="true" box="[199,468,1202,1227]" pageId="9" pageNumber="252">Description of female.</emphasis>
|
||
Specimen lacks head, pronotum and forelegs except one right femur, and the right tegmen.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321802A179FF3BFB4AFDFA6619" blockId="9.[151,1437,1202,2019]" pageId="9" pageNumber="252">
|
||
<emphasis id="B901EC201802A179FF3BFB4AFEA86675" box="[199,327,1275,1299]" italics="true" pageId="9" pageNumber="252">Fore femur</emphasis>
|
||
(
|
||
<figureCitation id="134E2CB71802A179FEA6FB4BFE416675" box="[346,430,1274,1299]" captionStart="FIGURES 9 – 20" captionStartId="7.[151,264,1281,1304]" captionTargetBox="[206,1384,223,1244]" captionTargetId="figure@7.[182,1404,195,1257]" captionTargetPageId="7" captionText="FIGURES 9 – 20. Fore legs of Hestiasulini treated in this paper, anterior view. 9. A. phyllopus, male from the Malay Peninsula (ES). 10. A. phyllopus, male from Brunei, mirrored (CS). 11. A. phyllopus, female from Brunei (CS). 12. A. sarawaca, male from Brunei (CS). 13. P. borneana, holotype (SMNK). 14. H. brunneriana, male from Sri Lanka (CS). 15. H. brunneriana, female from Sri Lanka (CS). 16. A. sarawaca, female from Kalimantan; Photo C. J. Schwarz © RMNH (RMNH). 17. C. moultoni, male from Sarawak (CS). 18. C. moultoni, female from Brunei, mirrored (CS). 19. C. nitida, male from Myanmar (SMNK). 20. C. nitida, male from Sumatra (SMNK) (not to scale)." httpUri="https://zenodo.org/record/829535/files/figure.png" pageId="9" pageNumber="252">Fig. 18</figureCitation>
|
||
) with 14 antero-ventral spines; spination formula iIiIiIiIiIiiiI. Posterior side with some tubercles on non-dilated part of femur. Postero-ventral margin very slightly serrulated between postero-ventral spines, with small pit between proximal first and second postero-ventral spines to accommodate distal-most postero-ventral spine of fore tibia.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321802A179FF3BFA3BFD3366A1" blockId="9.[151,1437,1202,2019]" pageId="9" pageNumber="252">
|
||
<emphasis id="B901EC201802A179FF3BFA3BFE7D66C5" box="[199,402,1418,1443]" italics="true" pageId="9" pageNumber="252">Mid and hind legs</emphasis>
|
||
with a ventral carina, hind femora additionally with a small proximal lobe. Hind metatarsus slightly shorter than remaining segments combined.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321802A179FF3BFA62FBDB6569" blockId="9.[151,1437,1202,2019]" pageId="9" pageNumber="252">
|
||
<emphasis id="B901EC201802A179FF3BFA62FEE5668C" box="[199,266,1491,1514]" italics="true" pageId="9" pageNumber="252">Wings</emphasis>
|
||
longer than abdomen, iridescent, slightly infumate, with opaque veins. Costal field of tegmen narrow (less than one third of tegmen width), with irregular cross-veins. Stigma indistinct.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321802A179FF3BF9ABFB856555" blockId="9.[151,1437,1202,2019]" box="[199,1130,1562,1587]" pageId="9" pageNumber="252">
|
||
<emphasis id="B901EC201802A179FF3BF9ABFEDE6555" box="[199,305,1562,1587]" italics="true" pageId="9" pageNumber="252">Abdomen</emphasis>
|
||
simple, fusiform, without lobes. Cerci short, pilose, last segment conical.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321802A179FF3BF98FFD67646D" blockId="9.[151,1437,1202,2019]" pageId="9" pageNumber="252">
|
||
<emphasis id="B901EC201802A179FF3BF98FFEA56531" box="[199,330,1598,1623]" italics="true" pageId="9" pageNumber="252">Coloration.</emphasis>
|
||
Main color dark brown. Distal part of fore coxa blackish, dorsal apical lobe light brown. Trochanter blackish. Posterior side of fore femur brown, dilated part with two irregular light markings, one at base, the second in distal third. Ventral part of fore femur black anteriorly, with a whitish patch traversing the clawgroove. Dilated part blackish ventrally, becoming somewhat lighter towards posterior margin, with two irregular yellowish markings at the same positions as seen on posterior side. Postero-ventral spines brown with dark apex, antero-ventral and discoidal spines blackish.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321802A179FF3BF8A7FCFE6435" blockId="9.[151,1437,1202,2019]" pageId="9" pageNumber="252">Mid and hind legs whitish, strongly mottled with dark, mottling pattern creating three indistinct annulations on femora and tibiae. Tarsomeres slightly darkened distally.</paragraph>
|
||
<paragraph id="8BCA30321802A179FF3BF8EFFAD56485" blockId="9.[151,1437,1202,2019]" pageId="9" pageNumber="252">Costal field, and area between subcostal and cubital vein in proximal part of tegmen opaque reddish-brown, remaining parts of tegmen perfectly hyaline, albeit slightly infumate, with reddish veins. Hindwings hyaline, slightly infumate, more so towards apex, area between costal and radial veins blackish opaque, discoidal area with blackish veins, anal area with blackish main and reddish cross-veins. All hyaline parts strongly iridescent.</paragraph>
|
||
<paragraph id="8BCA30321801A17AFF3BFF26FCEC63D6" blockId="10.[151,1437,151,1149]" box="[199,771,151,176]" pageId="10" pageNumber="253">Abdomen shiny black dorsally, brownish ventrally.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F63B91801A17AFF3BFF0DFE26671B" pageId="10" pageNumber="253" type="discussion">
|
||
<paragraph id="8BCA30321801A17AFF3BFF0DFEAF62CB" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="253">
|
||
<emphasis id="B901EC201801A17AFF3BFF0DFED463B3" bold="true" box="[199,315,188,213]" pageId="10" pageNumber="253">Remarks.</emphasis>
|
||
<bibRefCitation id="EFE44DC31801A17AFEBDFF0DFDE463B3" author="Giglio-Tos" box="[321,523,188,213]" pageId="10" pageNumber="253" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos (1915)</bibRefCitation>
|
||
created the genus
|
||
<taxonomicName id="4C754BB11801A17AFD24FF0DFC8B63B3" box="[728,868,188,213]" class="Insecta" family="Hymenopodidae" genus="Catestiasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EC201801A17AFD24FF0DFC8B63B3" box="[728,868,188,213]" italics="true" pageId="10" pageNumber="253">Catestiasula</emphasis>
|
||
</taxonomicName>
|
||
for two species:
|
||
<taxonomicName id="4C754BB11801A17AFBDDFF0DFB1663B3" box="[1057,1273,188,213]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="nitida">
|
||
<emphasis id="B901EC201801A17AFBDDFF0DFB1663B3" box="[1057,1273,188,213]" italics="true" pageId="10" pageNumber="253">Pachymantis nitida</emphasis>
|
||
</taxonomicName>
|
||
, designated as
|
||
<typeStatus id="54CE8E901801A17AFF6BFF51FF28639E" box="[151,199,224,248]" pageId="10" pageNumber="253">type</typeStatus>
|
||
species, and his newly described
|
||
<taxonomicName id="4C754BB11801A17AFDB8FF51FD27639E" box="[580,712,223,248]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201801A17AFDB8FF51FD27639E" box="[580,712,223,248]" italics="true" pageId="10" pageNumber="253">C. moultoni</emphasis>
|
||
</taxonomicName>
|
||
. In the original description and in his subsequent monograph on
|
||
<taxonomicName id="4C754BB11801A17AFF6BFEB5FE38627A" authority="Giglio-Tos 1927" authorityName="Giglio-Tos" authorityYear="1927" box="[151,471,260,285]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="order">
|
||
Mantodea (
|
||
<bibRefCitation id="EFE44DC31801A17AFEE9FEB5FE20627A" author="Giglio-Tos" box="[277,463,260,285]" pageId="10" pageNumber="253" refString="Giglio-Tos, E. (1927) Das Tierreich. 50. Lfg. Orthoptera Mantidae. Walter de Gruyter & Co., Berlin & Leipzig, XL + 707 pp." type="book" year="1927">Giglio-Tos 1927</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
he distinguished the two species based on the color of forewing main veins, which are allegedly blackish in
|
||
<taxonomicName id="4C754BB11801A17AFE7AFE99FE076226" box="[390,488,295,320]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="nitida">
|
||
<emphasis id="B901EC201801A17AFE7AFE99FE076226" box="[390,488,295,320]" italics="true" pageId="10" pageNumber="253">C. nitida</emphasis>
|
||
</taxonomicName>
|
||
and reddish in
|
||
<taxonomicName id="4C754BB11801A17AFD6AFE99FCF46226" box="[662,795,295,320]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201801A17AFD6AFE99FCF46226" box="[662,795,295,320]" italics="true" pageId="10" pageNumber="253">C. moultoni</emphasis>
|
||
</taxonomicName>
|
||
, and the color of the mid and hind tarsomeres, which are supposed to be simply brownish (“testaceous”) in
|
||
<taxonomicName id="4C754BB11801A17AFD35FEFDFCC26203" box="[713,813,332,357]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201801A17AFD35FEFDFCC26203" box="[713,813,332,357]" italics="true" pageId="10" pageNumber="253">moultoni</emphasis>
|
||
</taxonomicName>
|
||
but with black apex in
|
||
<taxonomicName id="4C754BB11801A17AFBCFFEFDFB9B6203" box="[1075,1140,332,357]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="nitida">
|
||
<emphasis id="B901EC201801A17AFBCFFEFDFB9B6203" box="[1075,1140,332,357]" italics="true" pageId="10" pageNumber="253">nitida</emphasis>
|
||
</taxonomicName>
|
||
. The wings of
|
||
<taxonomicName id="4C754BB11801A17AFAE1FEFDFAB06203" box="[1309,1375,332,357]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="nitida">
|
||
<emphasis id="B901EC201801A17AFAE1FEFDFAB06203" box="[1309,1375,332,357]" italics="true" pageId="10" pageNumber="253">nitida</emphasis>
|
||
</taxonomicName>
|
||
were characterized as being more infumate than in
|
||
<taxonomicName id="4C754BB11801A17AFD6DFEDEFD1A62EE" box="[657,757,367,392]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201801A17AFD6DFEDEFD1A62EE" box="[657,757,367,392]" italics="true" pageId="10" pageNumber="253">moultoni</emphasis>
|
||
</taxonomicName>
|
||
. At the same time he mentioned nothing about the coloration of fore femora.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321801A17AFF3BFE06FBE06292" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="253">
|
||
The types of
|
||
<taxonomicName id="4C754BB11801A17AFEA0FE09FE0E62B6" box="[348,481,439,464]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201801A17AFEA0FE09FE0E62B6" box="[348,481,439,464]" italics="true" pageId="10" pageNumber="253">C. moultoni</emphasis>
|
||
</taxonomicName>
|
||
, collected around Kuching, could not be located in the Sarawak Museum of Natural History, Kuching, where they had presumably been deposited (
|
||
<bibRefCitation id="EFE44DC31801A17AFCABFE6DFBEC6292" author="Ehrmann" box="[855,1027,476,501]" pageId="10" pageNumber="253" refString="Ehrmann, R. (2002) Mantodea - Gottesanbeterinnen der Welt. Natur und Tier-Verlag, Munster, 519 pp." type="book" year="2002">Ehrmann 2002</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCA30321801A17AFF3BFE4EFB9961E3" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="253">
|
||
The characters used by
|
||
<bibRefCitation id="EFE44DC31801A17AFE2EFE4EFD08617E" author="Giglio-Tos" box="[466,743,511,536]" pageId="10" pageNumber="253" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos (1915, 1 927)</bibRefCitation>
|
||
, i. e. the color of the veins and of the tarsi, are of little use in distinguishing between the species of this genus. The male specimens tentatively identified as
|
||
<taxonomicName id="4C754BB11801A17AFB42FD94FACE615B" box="[1214,1313,548,573]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="nitida">
|
||
<emphasis id="B901EC201801A17AFB42FD94FACE615B" box="[1214,1313,548,573]" italics="true" pageId="10" pageNumber="253">C. nitida</emphasis>
|
||
</taxonomicName>
|
||
have three maculations on the fore femur, which may vary in their extent but seem to be always present.
|
||
<taxonomicName id="4C754BB11801A17AFB24FDF9FAD26106" box="[1240,1341,583,608]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="nitida">
|
||
<emphasis id="B901EC201801A17AFB24FDF9FAD26106" box="[1240,1341,583,608]" italics="true" pageId="10" pageNumber="253">C. nitida</emphasis>
|
||
</taxonomicName>
|
||
is more robust and slightly larger than
|
||
<taxonomicName id="4C754BB11801A17AFE11FDDCFD9D61E3" box="[493,626,620,645]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201801A17AFE11FDDCFD9D61E3" box="[493,626,620,645]" italics="true" pageId="10" pageNumber="253">C. moultoni</emphasis>
|
||
</taxonomicName>
|
||
, with longer wings and more robust genitalia.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321801A17AFF3BFD3EFABE6051" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="253">
|
||
Both sexes of
|
||
<taxonomicName id="4C754BB11801A17AFE99FD21FE0661CE" box="[357,489,655,680]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201801A17AFE99FD21FE0661CE" box="[357,489,655,680]" italics="true" pageId="10" pageNumber="253">C. moultoni</emphasis>
|
||
</taxonomicName>
|
||
have forewing veins of various hues, mostly dark red to reddish-brown, and mid and hind tarsomeres with dark apices. Most of them have the antero-ventral side of the fore femur uniformly black, with only a small white spot in the claw groove (
|
||
<figureCitation id="134E2CB71801A17AFD37FD66FCF06196" box="[715,799,727,752]" captionStart="FIGURES 9 – 20" captionStartId="7.[151,264,1281,1304]" captionTargetBox="[206,1384,223,1244]" captionTargetId="figure@7.[182,1404,195,1257]" captionTargetPageId="7" captionText="FIGURES 9 – 20. Fore legs of Hestiasulini treated in this paper, anterior view. 9. A. phyllopus, male from the Malay Peninsula (ES). 10. A. phyllopus, male from Brunei, mirrored (CS). 11. A. phyllopus, female from Brunei (CS). 12. A. sarawaca, male from Brunei (CS). 13. P. borneana, holotype (SMNK). 14. H. brunneriana, male from Sri Lanka (CS). 15. H. brunneriana, female from Sri Lanka (CS). 16. A. sarawaca, female from Kalimantan; Photo C. J. Schwarz © RMNH (RMNH). 17. C. moultoni, male from Sarawak (CS). 18. C. moultoni, female from Brunei, mirrored (CS). 19. C. nitida, male from Myanmar (SMNK). 20. C. nitida, male from Sumatra (SMNK) (not to scale)." httpUri="https://zenodo.org/record/829535/files/figure.png" pageId="10" pageNumber="253">Fig. 17</figureCitation>
|
||
), but several specimens, including the female (
|
||
<figureCitation id="134E2CB71801A17AFAC6FD66FA616196" box="[1338,1422,727,752]" captionStart="FIGURES 9 – 20" captionStartId="7.[151,264,1281,1304]" captionTargetBox="[206,1384,223,1244]" captionTargetId="figure@7.[182,1404,195,1257]" captionTargetPageId="7" captionText="FIGURES 9 – 20. Fore legs of Hestiasulini treated in this paper, anterior view. 9. A. phyllopus, male from the Malay Peninsula (ES). 10. A. phyllopus, male from Brunei, mirrored (CS). 11. A. phyllopus, female from Brunei (CS). 12. A. sarawaca, male from Brunei (CS). 13. P. borneana, holotype (SMNK). 14. H. brunneriana, male from Sri Lanka (CS). 15. H. brunneriana, female from Sri Lanka (CS). 16. A. sarawaca, female from Kalimantan; Photo C. J. Schwarz © RMNH (RMNH). 17. C. moultoni, male from Sarawak (CS). 18. C. moultoni, female from Brunei, mirrored (CS). 19. C. nitida, male from Myanmar (SMNK). 20. C. nitida, male from Sumatra (SMNK) (not to scale)." httpUri="https://zenodo.org/record/829535/files/figure.png" pageId="10" pageNumber="253">Fig. 18</figureCitation>
|
||
), also have two very weakly expressed, more distal spots, of the same shape and position as in
|
||
<taxonomicName id="4C754BB11801A17AFB5EFD4CFAEC6073" box="[1186,1283,764,789]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="nitida">
|
||
<emphasis id="B901EC201801A17AFB5EFD4CFAEC6073" box="[1186,1283,764,789]" italics="true" pageId="10" pageNumber="253">C. nitida</emphasis>
|
||
</taxonomicName>
|
||
. Despite this, truly intermediate specimens do not seem to exist, pointing to the validity of both
|
||
<taxonomicName id="4C754BB11801A17AFBD1FC91FB7F605E" box="[1069,1168,799,824]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="nitida">
|
||
<emphasis id="B901EC201801A17AFBD1FC91FB7F605E" box="[1069,1168,799,824]" italics="true" pageId="10" pageNumber="253">C. nitida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C754BB11801A17AFB39FC91FAA6605E" box="[1221,1353,799,824]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201801A17AFB39FC91FAA6605E" box="[1221,1353,799,824]" italics="true" pageId="10" pageNumber="253">C. moultoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCA30321801A17AFF3BFCF5FE26671B" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="253">
|
||
The situation is further complicated by the existence of continental specimens without spots on the fore femur.
|
||
<taxonomicName id="4C754BB11801A17AFF6BFCD6FDF360E6" authority="Zhang, 1992" authorityName="Zhang" authorityYear="1992" box="[151,540,871,896]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="seminigra">
|
||
<emphasis id="B901EC201801A17AFF6BFCD6FE6A60E6" box="[151,389,871,896]" italics="true" pageId="10" pageNumber="253">Hestiasula seminigra</emphasis>
|
||
<bibRefCitation id="EFE44DC31801A17AFE70FCD6FDF360E6" author="Zhang" box="[396,540,871,896]" pageId="10" pageNumber="253" refString="Zhang, G. (1992) Two new species of the genus Hestiasula Saussure from China (Mantidae: Hymenopodinae). Acta Zootaxonomica Sinica, 17 (1), 67 - 70." type="journal article" year="1992">Zhang, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="F36270A21801A17AFD98FCD6FD4860E6" box="[612,679,871,896]" name="China" pageId="10" pageNumber="253">China</collectingCountry>
|
||
undoubtedly belongs to
|
||
<taxonomicName id="4C754BB11801A17AFC47FCD6FBA860E6" box="[955,1095,871,896]" class="Insecta" family="Hymenopodidae" genus="Catestiasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EC201801A17AFC47FCD6FBA860E6" box="[955,1095,871,896]" italics="true" pageId="10" pageNumber="253">Catestiasula</emphasis>
|
||
</taxonomicName>
|
||
, being very similar to both SE Asian species in all respects. Accordingly, we transfer it here to this genus as
|
||
<taxonomicName id="4C754BB11801A17AFC08FC3DFA7360C2" authority="Zhang, 1992" authorityName="Zhang" authorityYear="1992" box="[1012,1436,908,933]" class="Insecta" family="Hymenopodidae" genus="Catestiasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="seminigra" status="comb. nov.">
|
||
<emphasis id="B901EC201801A17AFC08FC3DFB1960C2" box="[1012,1270,908,933]" italics="true" pageId="10" pageNumber="253">Catestiasula seminigra</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE44DC31801A17AFAF8FC3DFA7B60C2" author="Zhang" box="[1284,1428,908,933]" pageId="10" pageNumber="253" refString="Zhang, G. (1992) Two new species of the genus Hestiasula Saussure from China (Mantidae: Hymenopodinae). Acta Zootaxonomica Sinica, 17 (1), 67 - 70." type="journal article" year="1992">Zhang, 1992</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A232515B1801A17AFF6BFC1EFF1360AE" box="[151,252,943,968]" pageId="10" pageNumber="253">
|
||
<emphasis id="B901EC201801A17AFF6BFC1EFF1360AE" bold="true" box="[151,252,943,968]" pageId="10" pageNumber="253">n. comb.</emphasis>
|
||
</taxonomicNameLabel>
|
||
. This species is particularly close to or even identical with
|
||
<taxonomicName id="4C754BB11801A17AFC68FC01FBF660AE" box="[916,1049,943,968]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="B901EC201801A17AFC68FC01FBF660AE" box="[916,1049,943,968]" italics="true" pageId="10" pageNumber="253">C. moultoni</emphasis>
|
||
</taxonomicName>
|
||
. Specimens probably representing the latter species are also known from
|
||
<collectingRegion id="49B1FED01801A17AFDA5FC65FD54608B" box="[601,699,980,1005]" country="Thailand" name="Sara Buri" pageId="10" pageNumber="253">Saraburi</collectingRegion>
|
||
, S
|
||
<collectingCountry id="F36270A21801A17AFD1EFC65FCA9608B" box="[738,838,980,1005]" name="Thailand" pageId="10" pageNumber="253">Thailand</collectingCountry>
|
||
(SMNK), and
|
||
<collectingRegion id="49B1FED01801A17AFC0AFC65FBB4608B" box="[1014,1115,980,1005]" country="Malaysia" name="Selangor" pageId="10" pageNumber="253">Selangor</collectingRegion>
|
||
, Malay Peninsula (
|
||
<figureCitation id="134E2CB71801A17AFAC5FC65FA7F608B" box="[1337,1424,980,1005]" captionStart="FIGURES 51 – 52" captionStartId="23.[151,264,1634,1657]" captionTargetBox="[359,1219,469,1613]" captionTargetId="figure@23.[340,1247,469,1613]" captionTargetPageId="23" captionText="FIGURES 51 – 52. Life aspect of Catestiasula. 51. Male of C. moultoni, Mt. Trus Madi, Sabah, Malaysia; Photo © Alexey Yakovlev. 52. Male of unidentified Catestiasula specimen from Ulu Gombak NP, Selangor, Malaysia; Photo © Bernhard Jacobi." httpUri="https://zenodo.org/record/829551/files/figure.png" pageId="10" pageNumber="253">Fig. 52</figureCitation>
|
||
). There is also an unidentified
|
||
<taxonomicName id="4C754BB11801A17AFE21FC46FD866776" box="[477,617,1015,1040]" class="Insecta" family="Hymenopodidae" genus="Catestiasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="10" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EC201801A17AFE21FC46FD866776" box="[477,617,1015,1040]" italics="true" pageId="10" pageNumber="253">Catestiasula</emphasis>
|
||
</taxonomicName>
|
||
specimen from
|
||
<collectingRegion id="49B1FED01801A17AFCE3FC46FC836776" box="[799,876,1015,1040]" country="India" name="Kerala" pageId="10" pageNumber="253">Kerala</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36270A21801A17AFC8BFC46FC5E6776" box="[887,945,1015,1040]" name="India" pageId="10" pageNumber="253">India</collectingCountry>
|
||
in the SMNK, extending the distribution of the genus to the Indian subcontinent. Its genitalia are depicted in
|
||
<figureCitation id="134E2CB71801A17AFC7AFBADFC346752" box="[902,987,1052,1077]" captionStart="FIGURES 43 – 50" captionStartId="22.[151,264,1570,1593]" captionTargetBox="[151,1436,467,1550]" captionTargetId="figure@22.[151,1436,467,1550]" captionTargetPageId="22" captionText="FIGURES 43 – 50. Genitalia of various Hestiasulini. 43. A. phyllopus from Sabah (CS). 44. A. basinigra from Myanmar (SMNK). 45. C. nitida from Myanmar (SMNK). 46. C. nitida from Sumatra (SMNK). 47. C. sp. from Kerala (SMNK). 48. Ephestiasula amoena from Nepal (ES). 49. E. pictipes from India (CS). 50. H. gyldenstolpei from Thailand (ES) (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829549/files/figure.png" pageId="10" pageNumber="253">Fig. 47</figureCitation>
|
||
. A revision of the whole complex with examination of additional material from various locations throughout the range is highly desirable to determine the true diversity of this genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |