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<document id="35B12D0E381CD91BB28B501C1DF53C98" ID-CLB-Dataset="45540" ID-DOI="10.11646/zootaxa.3814.3.2" ID-GBIF-Dataset="708ea530-4c8d-473b-8f29-1a54dea92f28" ID-ISSN="1175-5326" ID-Zenodo-Dep="231107" ID-ZooBank="5391C375-896D-4794-84A6-1846DF9E407D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460434473942" checkinUser="plazi" docAuthor="Kania, Iwona" docDate="2014" docId="03D787E6FFE6FFEAFF6BA56A3CB5BA94" docLanguage="en" docName="zt03814p352.pdf" docOrigin="Zootaxa 3814 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helius minutus Loew 1850" docType="treatment" docVersion="9" lastPageNumber="341" masterDocId="FFEEFF9EFFE1FFE2FFFCA26B3C50BD09" masterDocTitle="Subfamily Limoniinae Speiser, 1909 (Diptera, Limoniidae) from Baltic amber (Eocene): the genus Helius Lepeletier &amp; Serville, 1828" masterLastPageNumber="352" masterPageNumber="333" pageNumber="340" updateTime="1698331894480" updateUser="ExternalLinkService">
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<mods:title id="9DDF8720453ABCDE3117313CC7A161CD">Subfamily Limoniinae Speiser, 1909 (Diptera, Limoniidae) from Baltic amber (Eocene): the genus Helius Lepeletier &amp; Serville, 1828</mods:title>
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<treatment id="03D787E6FFE6FFEAFF6BA56A3CB5BA94" ID-DOI="http://doi.org/10.5281/zenodo.6138164" ID-GBIF-Taxon="119431365" ID-Zenodo-Dep="6138164" LSID="urn:lsid:plazi:treatment:03D787E6FFE6FFEAFF6BA56A3CB5BA94" httpUri="http://treatment.plazi.org/id/03D787E6FFE6FFEAFF6BA56A3CB5BA94" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340">
<subSubSection id="C364657BFFE6FFEAFF6BA56A3E95BF35" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340" type="nomenclature">
<paragraph id="8BC136F0FFE6FFE5FF6BA56A3DA8BA15" blockId="7.[151,504,1793,1853]" box="[151,504,1793,1820]" pageId="7" pageNumber="340">
<heading id="D089819CFFE6FFE5FF6BA56A3DA8BA15" bold="true" box="[151,504,1793,1820]" fontSize="11" level="1" pageId="7" pageNumber="340" reason="1">
<taxonomicName id="4C7E4D73FFE6FFE5FF6BA56A3DA8BA15" authority="Loew, 1850" authorityName="Loew" authorityYear="1850" box="[151,504,1793,1820]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE6FFE5FF6BA56A3DA8BA15" bold="true" box="[151,504,1793,1820]" pageId="7" pageNumber="340">
<emphasis id="B90AEAE2FFE6FFE5FF6BA56A3D01BA12" bold="true" box="[151,337,1793,1819]" italics="true" pageId="7" pageNumber="340">Helius minutus</emphasis>
(
<bibRefCitation id="EFEF4B01FFE6FFE5FE9DA5693DA0BA15" author="Loew" box="[353,496,1794,1820]" pageId="7" pageNumber="340" refString="Loew, H. (1850) Uber den Bernstein Und die Bernsteinfauna. Program der Keiserischen Realschule Meseritz, 44 pp." type="book" year="1850">Loew, 1850</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC136F0FFE6FFE5FF6BA54F3D74BA34" blockId="7.[151,504,1793,1853]" box="[151,292,1828,1853]" pageId="7" pageNumber="340">
(
<figureCitation id="13452A75FFE6FFE5FF63A54F3CB4BA34" box="[159,228,1828,1853]" captionStart="FIGURE 6" captionStartId="9.[151,250,1277,1299]" captionTargetBox="[329,1261,200,1245]" captionTargetId="figure@9.[322,1267,193,1252]" captionTargetPageId="9" captionText="FIGURE 6. Helius minutus Loew, 1850, No. MB. J. 225 (male); A, head (after Krzemiński 1993); B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231113/files/figure.png" pageId="7" pageNumber="340">Figs 6</figureCitation>
A–C)
</paragraph>
<paragraph id="8BC136F0FFE6FFE5FF6BA5073DB4BA8B" blockId="7.[151,1030,1900,2014]" box="[151,484,1900,1922]" pageId="7" pageNumber="340">
*1850
<taxonomicName id="4C7E4D73FFE6FFE5FF20A5073DF4BA8B" box="[220,420,1900,1922]" class="Insecta" family="Limoniidae" genus="Rhamphidia" higherTaxonomySource="CoL" kingdom="Plantae" order="Diptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B90AEAE2FFE6FFE5FF20A5073DF4BA8B" box="[220,420,1900,1922]" italics="true" pageId="7" pageNumber="340">Rhamphidia minuta</emphasis>
</taxonomicName>
, p. 37
</paragraph>
<paragraph id="8BC136F0FFE6FFE5FF6BA5E03E25BAA8" blockId="7.[151,1030,1900,2014]" box="[151,629,1931,1953]" pageId="7" pageNumber="340">
1856
<taxonomicName id="4C7E4D73FFE6FFE5FF2CA5E03D8CBAA8" authority="Loew" authorityName="Loew" box="[208,476,1931,1953]" class="Insecta" family="Limoniidae" genus="Rhamphidia" higherTaxonomySource="CoL" kingdom="Plantae" order="Diptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B90AEAE2FFE6FFE5FF2CA5E03DC8BAA8" box="[208,408,1931,1953]" italics="true" pageId="7" pageNumber="340">Rhamphidia minuta</emphasis>
Loew
</taxonomicName>
: Giebel, p. 242
</paragraph>
<paragraph id="8BC136F0FFE6FFE5FF6BA5C13ED6BAC9" blockId="7.[151,1030,1900,2014]" box="[151,646,1962,1984]" pageId="7" pageNumber="340">
1906
<taxonomicName id="4C7E4D73FFE6FFE5FF2CA5C13D8CBAC9" authority="Loew" authorityName="Loew" box="[208,476,1962,1984]" class="Insecta" family="Limoniidae" genus="Rhamphidia" higherTaxonomySource="CoL" kingdom="Plantae" order="Diptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B90AEAE2FFE6FFE5FF2CA5C13DC8BAC9" box="[208,408,1962,1984]" italics="true" pageId="7" pageNumber="340">Rhamphidia minuta</emphasis>
Loew
</taxonomicName>
: Meunier, p. 358
</paragraph>
<paragraph id="8BC136F0FFE6FFEAFF6BA5A33E3FBDC5" blockId="7.[151,1030,1900,2014]" lastBlockId="8.[151,968,152,573]" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340">
<date id="FFC01030FFE6FFE5FF6BA5A33C8BBAD7" box="[151,219,1992,2014]" pageId="7" pageNumber="340" value="1906-05">v.1906</date>
<taxonomicName id="4C7E4D73FFE6FFE5FF1DA5A33E33BAD7" authority="Loew (Meunier)" authorityName="Loew (Meunier" box="[225,611,1992,2014]" class="Insecta" family="Limoniidae" genus="Rhamphidia" higherTaxonomySource="CoL" kingdom="Plantae" order="Diptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B90AEAE2FFE6FFE5FF1DA5A33DE2BAD7" box="[225,434,1992,2014]" italics="true" pageId="7" pageNumber="340">Rhamphidia pulchra</emphasis>
Loew (Meunier)
</taxonomicName>
: Meunier, p. 358, p. 364, Pl. XII, fig.
<date id="FFC01030FFE6FFEAFC23A5A33C8BBDA7" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340" value="1906-05-03" valueMax="1906-05-04" valueMin="1906-05-03">3–4 v.1906</date>
<taxonomicName id="4C7E4D73FFE9FFEAFF1DA2F33DA4BDA7" authority="Meunier" authorityName="Meunier" box="[225,500,152,174]" class="Insecta" family="Limoniidae" genus="Antocha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="succinea">
<emphasis id="B90AEAE2FFE9FFEAFF1DA2F33DC5BDA7" box="[225,405,152,174]" italics="true" pageId="8" pageNumber="341">Antocha succinea</emphasis>
Meunier
</taxonomicName>
: Meunier, p. 358, 368, Pl. XII, fig. 8, 9 and 10 1931
<taxonomicName id="4C7E4D73FFE9FFEAFF2CA2DD3DECBDC5" authority="Loew" authorityName="Loew" box="[208,444,182,205]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFF2CA2DD3D38BDC4" box="[208,360,182,205]" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
(Loew)
</taxonomicName>
: Alexander, p. 42
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF6BA2BE3ECFBC21" blockId="8.[151,968,152,573]" pageId="8" pageNumber="341">
vp.1931
<taxonomicName id="4C7E4D73FFE9FFEAFF13A2BE3DA2BDE2" authority="Meunier" authorityName="Meunier" box="[239,498,213,235]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B90AEAE2FFE9FFEAFF13A2BE3DD5BDE2" box="[239,389,213,235]" italics="true" pageId="8" pageNumber="341">Helius pulcher</emphasis>
(Meunier)
</taxonomicName>
[sic!]: Alexander, p. 42 vp.1931
<taxonomicName id="4C7E4D73FFE9FFEAFF13A29F3E61BC03" authority="Meunier" authorityName="Meunier" box="[239,561,244,266]" class="Insecta" family="Limoniidae" genus="Rhamphidia" higherTaxonomySource="CoL" kingdom="Plantae" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B90AEAE2FFE9FFEAFF13A29F3D91BC03" box="[239,449,244,266]" italics="true" pageId="8" pageNumber="341">Rhamphidia pulchra</emphasis>
(Meunier)
</taxonomicName>
: Alexander, p. 42
<date id="FFC01030FFE9FFEAFF6BA3793C8BBC21" box="[151,219,274,296]" pageId="8" pageNumber="341" value="1931-05">v.1931</date>
<taxonomicName id="4C7E4D73FFE9FFEAFF1DA3793DA4BC21" authority="Meunier" authorityName="Meunier" box="[225,500,274,296]" class="Insecta" family="Limoniidae" genus="Antocha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="succinea">
<emphasis id="B90AEAE2FFE9FFEAFF1DA3793DC5BC21" box="[225,405,274,296]" italics="true" pageId="8" pageNumber="341">Antocha succinea</emphasis>
Meunier
</taxonomicName>
: Alexander, p.42
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF6BA35A3ED7BC4E" blockId="8.[151,968,152,573]" box="[151,647,305,327]" pageId="8" pageNumber="341">
1985
<taxonomicName id="4C7E4D73FFE9FFEAFF2CA35A3DECBC4E" authority="Loew" authorityName="Loew" box="[208,444,305,327]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFF2CA35A3D38BC4E" box="[208,360,305,327]" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
(Loew)
</taxonomicName>
: Krzemiński, p. 116
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF6BA33B3E65BC8D" blockId="8.[151,968,152,573]" pageId="8" pageNumber="341">
<date id="FFC01030FFE9FFEAFF6BA33B3C8BBC6F" box="[151,219,336,358]" pageId="8" pageNumber="341" value="1985-05">v.1985</date>
<taxonomicName id="4C7E4D73FFE9FFEAFF1DA33B3DA4BC6F" authority="Krzeminski" authorityName="Krzeminski" box="[225,500,336,358]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="abditus">
<emphasis id="B90AEAE2FFE9FFEAFF1DA33B3D25BC6F" box="[225,373,336,358]" italics="true" pageId="8" pageNumber="341">Helius abditus</emphasis>
Krzemiński
</taxonomicName>
: Krzemiński, p. 116, figs. 9, 10 1993
<taxonomicName id="4C7E4D73FFE9FFEAFF2CA3053DBCBC8D" authority="Krzeminski" authorityName="Krzeminski" box="[208,492,366,388]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B90AEAE2FFE9FFEAFF2CA3053D35BC8D" box="[208,357,366,388]" italics="true" pageId="8" pageNumber="341">Helius pulcher</emphasis>
: Krzemiński
</taxonomicName>
, p. 597
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF6BA3E63ED8BCAA" blockId="8.[151,968,152,573]" box="[151,648,397,419]" pageId="8" pageNumber="341">
1993
<taxonomicName id="4C7E4D73FFE9FFEAFF2CA3E63DECBCAA" authority="Loew" authorityName="Loew" box="[208,444,397,419]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFF2CA3E63D38BCAA" box="[208,360,397,419]" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
(Loew)
</taxonomicName>
: Krzemiński, p. 598
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF6BA3C73EFFBCE9" blockId="8.[151,968,152,573]" pageId="8" pageNumber="341">
<date id="FFC01030FFE9FFEAFF6BA3C73C8BBCCB" box="[151,219,428,450]" pageId="8" pageNumber="341" value="1993-05">v.1993</date>
<taxonomicName id="4C7E4D73FFE9FFEAFF1DA3C73D9CBCCB" authority="Loew" authorityName="Loew" box="[225,460,428,450]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFF1DA3C73D2ABCCB" box="[225,378,428,450]" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
(Loew)
</taxonomicName>
, p. 599,
<figureCitation id="13452A75FFE9FFEAFDDFA3C73E24BCCB" box="[547,628,428,450]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="7.[151,250,1282,1304]" captionStartId-1="9.[151,250,1277,1299]" captionStartId-2="10.[151,250,1974,1996]" captionTargetBox-0="[346,1240,198,1251]" captionTargetBox-1="[329,1261,200,1245]" captionTargetBox-2="[324,1276,1112,1932]" captionTargetId-0="figure@7.[346,1240,193,1260]" captionTargetId-1="figure@9.[322,1267,193,1252]" captionTargetId-2="figure@10.[319,1283,1097,1938]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 5. Helius hoffeinsorum sp. nov., No. 1462 (male), A, hypopygium (dorsal view); B, head; C, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); D, wing venation. Abbreviation of male hypopygium: gx—gonocoxite, ing—inner gonostylus, oug—outer gonostylus, aed—aedeagus, p—paramere." captionText-1="FIGURE 6. Helius minutus Loew, 1850, No. MB. J. 225 (male); A, head (after Krzemiński 1993); B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." captionText-2="FIGURE 7. Helius pulcher Loew, 1850, No. MB. J. 358 (female) A, head; B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri-0="https://zenodo.org/record/231112/files/figure.png" httpUri-1="https://zenodo.org/record/231113/files/figure.png" httpUri-2="https://zenodo.org/record/231114/files/figure.png" pageId="8" pageNumber="341">Fig. 5–7</figureCitation>
—
<typeStatus id="54C58852FFE9FFEAFD73A3C73EB1BCCB" box="[655,737,428,450]" pageId="8" pageNumber="341" type="neotype">neotype</typeStatus>
designation 1993
<taxonomicName id="4C7E4D73FFE9FFEAFF2CA3A13DB3BCE9" authority="Meunier" authorityName="Meunier" box="[208,483,458,480]" class="Insecta" family="Limoniidae" genus="Antocha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="succinea">
<emphasis id="B90AEAE2FFE9FFEAFF2CA3A13DD4BCE9" box="[208,388,458,480]" italics="true" pageId="8" pageNumber="341">Antocha succinea</emphasis>
Meunier
</taxonomicName>
: Krzemiński, p. 597
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF6BA3823E03BF17" blockId="8.[151,968,152,573]" pageId="8" pageNumber="341">
1993
<taxonomicName id="4C7E4D73FFE9FFEAFF2CA3823E51BCF6" authority="Krzeminski" authorityName="Krzeminski" box="[208,513,489,511]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="succineus">
<emphasis id="B90AEAE2FFE9FFEAFF2CA3823D2BBCF6" box="[208,379,489,511]" italics="true" pageId="8" pageNumber="341">Helius succineus</emphasis>
: Krzemiński
</taxonomicName>
, p. 597—nomen nudum 1994
<taxonomicName id="4C7E4D73FFE9FFEAFF2CA0633DFDBF17" authority="Loew" authorityName="Loew" box="[208,429,520,542]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFF2CA0633D38BF17" box="[208,360,520,542]" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
Loew
</taxonomicName>
: Evenhuis, p. 73
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF6BA04D3E95BF35" blockId="8.[151,968,152,573]" box="[151,709,550,573]" pageId="8" pageNumber="341">
1994
<taxonomicName id="4C7E4D73FFE9FFEAFF2CA04D3DB3BF35" authority="Meunier" authorityName="Meunier" box="[208,483,550,573]" class="Insecta" family="Limoniidae" genus="Antocha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="succinea">
<emphasis id="B90AEAE2FFE9FFEAFF2CA04D3DD4BF34" box="[208,388,550,573]" italics="true" pageId="8" pageNumber="341">Antocha succinea</emphasis>
Meunier
</taxonomicName>
: Evenhuis, p. 73 [sic!]
</paragraph>
</subSubSection>
<subSubSection id="C364657BFFE9FFEAFF6BA0023E1AB938" pageId="8" pageNumber="341" type="discussion">
<paragraph id="8BC136F0FFE9FFEAFF6BA0023E1AB938" blockId="8.[151,1437,617,1949]" pageId="8" pageNumber="341">
<emphasis id="B90AEAE2FFE9FFEAFF6BA0023D5BBF8B" bold="true" box="[151,267,617,642]" pageId="8" pageNumber="341">Remarks.</emphasis>
The species
<taxonomicName id="4C7E4D73FFE9FFEAFE58A0023F44BF8B" authority="Loew, 1850" authorityName="Loew" authorityYear="1850" box="[420,788,617,642]" class="Insecta" family="Limoniidae" genus="Rhamphidia" higherTaxonomySource="CoL" kingdom="Plantae" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B90AEAE2FFE9FFEAFE58A0023ED4BF88" box="[420,644,617,641]" italics="true" pageId="8" pageNumber="341">Rhamphidia minuta</emphasis>
<bibRefCitation id="EFEF4B01FFE9FFEAFD71A0013F44BF8B" author="Loew" box="[653,788,617,642]" pageId="8" pageNumber="341" refString="Loew, H. (1850) Uber den Bernstein Und die Bernsteinfauna. Program der Keiserischen Realschule Meseritz, 44 pp." type="book" year="1850">Loew, 1850</bibRefCitation>
</taxonomicName>
was not clearly characterized by Loew.
<bibRefCitation id="EFEF4B01FFE9FFEAFB15A00239CCBF8B" author="Meunier" box="[1257,1436,617,642]" pageId="8" pageNumber="341" refString="Meunier, F. (1906) Monographie des Tipulidae et des Dixidae de l'ambre de la Baltique. Annales des Science Naturelle Zoologie, 4, 349 - 401." type="journal article" year="1906">Meunier (1906)</bibRefCitation>
stated that he was not able to locate the specimen examined by Loew. The unavailability of Loew’s specimen was also noted by
<bibRefCitation id="EFEF4B01FFE9FFEAFEC2A0DA3E5BBFC3" author="Alexander" box="[318,523,689,714]" pageId="8" pageNumber="341" refString="Alexander, C. P. (1931) Crane-flies of the Baltic amber (Diptera). Berstein-Forschungen, 2, 1 - 135." type="journal article" year="1931">Alexander (1931)</bibRefCitation>
, who, however, placed it in the genus
<taxonomicName id="4C7E4D73FFE9FFEAFC28A0DA384DBFC0" box="[980,1053,689,713]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAE2FFE9FFEAFC28A0DA384DBFC0" box="[980,1053,689,713]" italics="true" pageId="8" pageNumber="341">Helius</emphasis>
</taxonomicName>
, resulting in combination
<taxonomicName id="4C7E4D73FFE9FFEAFAAFA0DA3DC8BFE7" authority="Loew, 1850" authorityName="Loew" authorityYear="1850" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFAAFA0DA3CA0BFE4" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
(
<bibRefCitation id="EFEF4B01FFE9FFEAFEFEA0BD3DC0BFE7" author="Loew" box="[258,400,725,750]" pageId="8" pageNumber="341" refString="Loew, H. (1850) Uber den Bernstein Und die Bernsteinfauna. Program der Keiserischen Realschule Meseritz, 44 pp." type="book" year="1850">Loew, 1850</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="EFEF4B01FFE9FFEAFE5BA0BE3E2FBFE7" author="Krzeminski" box="[423,639,725,750]" pageId="8" pageNumber="341" refString="Krzeminski, W. (1985) Limoniidae (Diptera Nematocera) from Baltic amber (in the collection of the Museum of the Earth in Warsaw). Part I. Subfamily Limoniinae. Prace Muzeum Ziemi, Warszawa, 37, 113 - 117." type="journal article" year="1985">Krzemiński (1985)</bibRefCitation>
stated that material of
<taxonomicName id="4C7E4D73FFE9FFEAFC6FA0BE38CDBFE7" authority="Loew" authorityName="Loew" box="[915,1181,725,750]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFC6FA0BE3810BFE4" box="[915,1088,725,749]" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
(Loew)
</taxonomicName>
is lost. Subsequently,
<bibRefCitation id="EFEF4B01FFE9FFEAFF6BA0923D3EBE1B" author="Krzeminski" box="[151,366,761,786]" pageId="8" pageNumber="341" refString="Krzeminski, W. (1993) Fossil Tipulomorpha (Diptera, Nematocera) from Baltic amber (Upper Eocene). - Revision of the genus Helius Lepeletier et Serville (Limoniidae). Acta Zoologica Cracoviensia, 35, 597 - 601." type="journal article" year="1993">Krzemiński (1993)</bibRefCitation>
designated
<typeStatus id="54C58852FFE9FFEAFE06A0913E03BE1B" box="[506,595,762,786]" pageId="8" pageNumber="341" type="neotype">neotype</typeStatus>
for this species, based on the specimen deposited in the Museum of the Earth, PAS, Warsaw. He also synonymized
<taxonomicName id="4C7E4D73FFE9FFEAFD72A1763858BE3F" authority="Krzeminski, 1985" authorityName="Krzeminski" authorityYear="1985" box="[654,1032,797,822]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="abditus">
<emphasis id="B90AEAE2FFE9FFEAFD72A1763F63BE3C" box="[654,819,797,821]" italics="true" pageId="8" pageNumber="341">Helius abditus</emphasis>
<bibRefCitation id="EFEF4B01FFE9FFEAFCC1A1763858BE3F" author="Krzeminski" box="[829,1032,797,822]" pageId="8" pageNumber="341" refString="Krzeminski, W. (1985) Limoniidae (Diptera Nematocera) from Baltic amber (in the collection of the Museum of the Earth in Warsaw). Part I. Subfamily Limoniinae. Prace Muzeum Ziemi, Warszawa, 37, 113 - 117." type="journal article" year="1985">Krzemiński, 1985</bibRefCitation>
</taxonomicName>
under
<taxonomicName id="4C7E4D73FFE9FFEAFBA7A1763936BE3F" authority="Loew" authorityName="Loew" box="[1115,1382,797,822]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFBA7A1763957BE3C" box="[1115,1287,797,821]" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
(Loew)
</taxonomicName>
, and proposed to synonymize
<taxonomicName id="4C7E4D73FFE9FFEAFE47A12A3F67BE53" authority="Meunier, 1906" authorityName="Meunier" authorityYear="1906" box="[443,823,833,858]" class="Insecta" family="Limoniidae" genus="Antocha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="succinea">
<emphasis id="B90AEAE2FFE9FFEAFE47A12A3ED5BE50" box="[443,645,833,857]" italics="true" pageId="8" pageNumber="341">Antocha succinea</emphasis>
<bibRefCitation id="EFEF4B01FFE9FFEAFD73A12A3F67BE53" author="Meunier" box="[655,823,833,858]" pageId="8" pageNumber="341" refString="Meunier, F. (1906) Monographie des Tipulidae et des Dixidae de l'ambre de la Baltique. Annales des Science Naturelle Zoologie, 4, 349 - 401." type="journal article" year="1906">Meunier, 1906</bibRefCitation>
</taxonomicName>
under
<taxonomicName id="4C7E4D73FFE9FFEAFC70A12A38C8BE53" authority="Loew" authorityName="Loew" box="[908,1176,833,858]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFC70A12A3868BE50" box="[908,1080,833,857]" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
(Loew)
</taxonomicName>
. It must be noted that specimen No.
<quantity id="4C869B15FFE9FFEAFEC6A10E3DC0BE77" box="[314,400,869,894]" metricMagnitude="2" metricUnit="m" metricValue="1.157478" pageId="8" pageNumber="341" unit="in" value="4557.0">4557 in</quantity>
<bibRefCitation id="EFEF4B01FFE9FFEAFE6BA10E3E19BE77" author="Meunier" box="[407,585,869,894]" pageId="8" pageNumber="341" refString="Meunier, F. (1906) Monographie des Tipulidae et des Dixidae de l'ambre de la Baltique. Annales des Science Naturelle Zoologie, 4, 349 - 401." type="journal article" year="1906">Meunier (1906)</bibRefCitation>
paper (specimen No. Z-4557, examined by Krzemiński, deposited recently in Palaeontological Institute in Göttingen—Klebs collection) is the
<typeStatus id="54C58852FFE9FFEAFC66A1E13F9ABEAB" box="[922,970,906,930]" pageId="8" pageNumber="341">type</typeStatus>
specimen of
<taxonomicName id="4C7E4D73FFE9FFEAFB9AA1E23C82BECF" authority="Meunier, 1906" authorityName="Meunier" authorityYear="1906" class="Insecta" family="Limoniidae" genus="Antocha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="succinea">
<emphasis id="B90AEAE2FFE9FFEAFB9AA1E2397EBEA8" box="[1126,1326,905,929]" italics="true" pageId="8" pageNumber="341">Antocha succinea</emphasis>
<bibRefCitation id="EFEF4B01FFE9FFEAFACBA1E23C82BECF" author="Meunier" pageId="8" pageNumber="341" refString="Meunier, F. (1906) Monographie des Tipulidae et des Dixidae de l'ambre de la Baltique. Annales des Science Naturelle Zoologie, 4, 349 - 401." type="journal article" year="1906">Meunier, 1906</bibRefCitation>
</taxonomicName>
, but
<bibRefCitation id="EFEF4B01FFE9FFEAFEF7A1C63DB0BECF" author="Krzeminski" box="[267,480,941,966]" pageId="8" pageNumber="341" refString="Krzeminski, W. (1993) Fossil Tipulomorpha (Diptera, Nematocera) from Baltic amber (Upper Eocene). - Revision of the genus Helius Lepeletier et Serville (Limoniidae). Acta Zoologica Cracoviensia, 35, 597 - 601." type="journal article" year="1993">Krzemiński (1993)</bibRefCitation>
did not indicate, that it is the
<typeStatus id="54C58852FFE9FFEAFCC7A1C53F3EBECF" box="[827,878,942,966]" pageId="8" pageNumber="341">type</typeStatus>
, and ascribed the specimen as material of
<taxonomicName id="4C7E4D73FFE9FFEAFAAFA1C63CA0BEE0" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFAAFA1C63CA0BEE0" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFEF4B01FFE9FFEAFEF9A1BA3DB0BEE3" author="Krzeminski" box="[261,480,977,1002]" pageId="8" pageNumber="341" refString="Krzeminski, W. (1993) Fossil Tipulomorpha (Diptera, Nematocera) from Baltic amber (Upper Eocene). - Revision of the genus Helius Lepeletier et Serville (Limoniidae). Acta Zoologica Cracoviensia, 35, 597 - 601." type="journal article" year="1993">Krzemiński (1993)</bibRefCitation>
also listed specimens: K-4265 (Klebs collection, No. 4265) and Z-253 (Klebs collection No. 253; both presently deposited in Göttingen), which were ascribed to
<taxonomicName id="4C7E4D73FFE9FFEAFBAEA19E38A9B904" box="[1106,1273,1013,1037]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B90AEAE2FFE9FFEAFBAEA19E38A9B904" box="[1106,1273,1013,1037]" italics="true" pageId="8" pageNumber="341">Helius pulcher</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFEF4B01FFE9FFEAFADBA19E3CB1B93B" author="Alexander" pageId="8" pageNumber="341" refString="Alexander, C. P. (1931) Crane-flies of the Baltic amber (Diptera). Berstein-Forschungen, 2, 1 - 135." type="journal article" year="1931">Alexander (1931)</bibRefCitation>
as belonging to
<taxonomicName id="4C7E4D73FFE9FFEAFE67A6723E14B938" box="[411,580,1049,1073]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE9FFEAFE67A6723E14B938" box="[411,580,1049,1073]" italics="true" pageId="8" pageNumber="341">Helius minutus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C364657BFFE9FFEAFF3BA6563EA5B803" pageId="8" pageNumber="341" type="materials_examined">
<paragraph id="8BC136F0FFE9FFEAFF3BA656393CB973" blockId="8.[151,1437,617,1949]" pageId="8" pageNumber="341">
<emphasis id="B90AEAE2FFE9FFEAFF3BA6563DE2B95F" bold="true" box="[199,434,1085,1110]" pageId="8" pageNumber="341">Material examined.</emphasis>
<typeStatus id="54C58852FFE9FFEAFE47A6553E4EB95F" box="[443,542,1086,1110]" pageId="8" pageNumber="341" type="neotype">Neotype</typeStatus>
: No. 7895 (male), (
<collectionCode id="ED6FAE35FFE9FFEAFD02A6553F30B95F" box="[766,864,1086,1110]" pageId="8" pageNumber="341">MEPAS</collectionCode>
); additional material: No. MB.J. 224 (male), Coll. Kühl (
<collectionCode id="ED6FAE35FFE9FFEAFF22A6093D65B973" box="[222,309,1122,1146]" pageId="8" pageNumber="341">ZMHB</collectionCode>
); No. K-4265, Z-253 (male), No. K1696, Z-4557 (male) (
<collectionCode id="ED6FAE35FFE9FFEAFC46A6093845B973" box="[954,1045,1122,1146]" httpUri="http://grbio.org/cool/2qt9-gmq1" name="Universitat Gottingen, Geologisches-Palaatologisches Museum" pageId="8" pageNumber="341">GMUG</collectionCode>
), No. MP/3342 (male) (
<collectionCode id="ED6FAE35FFE9FFEAFADCA609390FB973" box="[1312,1375,1122,1146]" httpUri="http://grbio.org/cool/hv18-23j2" name="Institute of Systematic Zoology" pageId="8" pageNumber="341">ISZP</collectionCode>
).
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF3BA6EE3EA5B803" blockId="8.[151,1437,617,1949]" pageId="8" pageNumber="341">
<emphasis id="B90AEAE2FFE9FFEAFF3BA6EE3D80B997" bold="true" box="[199,464,1157,1182]" pageId="8" pageNumber="341">Diagnostic characters.</emphasis>
Rostrum approximately 1.2x the length of the head (
<figureCitation id="13452A75FFE9FFEAFBDDA6EE3835B997" box="[1057,1125,1157,1182]" captionStart="FIGURE 6" captionStartId="9.[151,250,1277,1299]" captionTargetBox="[329,1261,200,1245]" captionTargetId="figure@9.[322,1267,193,1252]" captionTargetPageId="9" captionText="FIGURE 6. Helius minutus Loew, 1850, No. MB. J. 225 (male); A, head (after Krzemiński 1993); B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231113/files/figure.png" pageId="8" pageNumber="341">Fig. 6</figureCitation>
B), approximately equal the length of palpus, or palpus slightly longer, rostrum shorter than antenna; antenna more than twice longer than the rostrum; outer gonostylus short, straight, with two broad spikes apically; vein r-m rather long, well expressed, cross-vein m-cu almost in proximal 1/3 of d-cell base.
</paragraph>
</subSubSection>
<subSubSection id="C364657BFFE9FFEAFF3BA77E397FBBEF" pageId="8" pageNumber="341" type="description">
<paragraph id="8BC136F0FFE9FFEAFF3BA77E3873B824" blockId="8.[151,1437,617,1949]" box="[199,1059,1301,1326]" pageId="8" pageNumber="341">
<emphasis id="B90AEAE2FFE9FFEAFF3BA77E3D24B827" bold="true" box="[199,372,1301,1326]" pageId="8" pageNumber="341">Redescription.</emphasis>
Body: the length of specimens approximately
<quantity id="4C869B15FFE9FFEAFC7BA77E384FB824" box="[903,1055,1301,1326]" metricMagnitude="-3" metricUnit="m" metricValue="4.365" metricValueMax="5.7" metricValueMin="3.03" pageId="8" pageNumber="341" unit="mm" value="4.365" valueMax="5.7" valueMin="3.03">3.03–5.7 mm</quantity>
.
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF3BA7523940B8B4" blockId="8.[151,1437,617,1949]" pageId="8" pageNumber="341">
Head (
<figureCitation id="13452A75FFE9FFEAFEEFA7523D06B85B" box="[275,342,1337,1362]" captionStart="FIGURE 6" captionStartId="9.[151,250,1277,1299]" captionTargetBox="[329,1261,200,1245]" captionTargetId="figure@9.[322,1267,193,1252]" captionTargetPageId="9" captionText="FIGURE 6. Helius minutus Loew, 1850, No. MB. J. 225 (male); A, head (after Krzemiński 1993); B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231113/files/figure.png" pageId="8" pageNumber="341">Fig. 6</figureCitation>
A, 10D): rostrum
<quantity id="4C869B15FFE9FFEAFDDFA7523E97B858" box="[547,711,1337,1362]" metricMagnitude="-4" metricUnit="m" metricValue="6.85" metricValueMax="7.3" metricValueMin="6.4" pageId="8" pageNumber="341" unit="mm" value="0.685" valueMax="0.73" valueMin="0.64">0.64–0.73 mm</quantity>
long (
<typeStatus id="54C58852FFE9FFEAFCEDA7513F3FB85B" box="[785,879,1338,1362]" pageId="8" pageNumber="341" type="neotype">neotype</typeStatus>
), approximately 1.2x the length of the head (
<figureCitation id="13452A75FFE9FFEAFA93A7523CF6B87F" captionStart="FIGURE 6" captionStartId="9.[151,250,1277,1299]" captionTargetBox="[329,1261,200,1245]" captionTargetId="figure@9.[322,1267,193,1252]" captionTargetPageId="9" captionText="FIGURE 6. Helius minutus Loew, 1850, No. MB. J. 225 (male); A, head (after Krzemiński 1993); B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231113/files/figure.png" pageId="8" pageNumber="341">Fig. 6</figureCitation>
B); antenna 16-segmented,
<quantity id="4C869B15FFE9FFEAFE20A7363E11B87C" box="[476,577,1373,1398]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="341" unit="mm" value="1.15">1.15 mm</quantity>
long, more than twice longer than rostrum; scape cylindrical, large; pedicel barrel-shaped, broad; first flagellomere short, broad, flagellomeres 2–16 cylindrical with four elongated setae (
<figureCitation id="13452A75FFE9FFEAFA93A7EA3CE5B8B7" captionStart="FIGURE 10" captionStartId="13.[151,250,1878,1900]" captionTargetBox="[191,1395,193,1850]" captionTargetId="figure@13.[189,1398,193,1857]" captionTargetPageId="13" captionText="FIGURE 10. Morphology of Helius from Baltic amber. A H. hoffeinsorum sp. nov., No. 1462 (male), holotype, the body, lateral view; B. H. similis sp. nov., No. 3413 (male) head and thorax latero-ventral view, C – D. H. minutus Loew, 1850, No. 7895 (male), neotype, C. antennae, palpi, rostrum, D. head." httpUri="https://zenodo.org/record/231117/files/figure.png" pageId="8" pageNumber="341">Fig. 10</figureCitation>
C); 1.5–2 times than the segments bearing them; last flagellomere with two elongated setae at apex.
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF3BA7A238C6BB0F" blockId="8.[151,1437,617,1949]" pageId="8" pageNumber="341">
Palpus (
<figureCitation id="13452A75FFE9FFEAFEDDA7A23D34B8EB" box="[289,356,1481,1506]" captionStart="FIGURE 6" captionStartId="9.[151,250,1277,1299]" captionTargetBox="[329,1261,200,1245]" captionTargetId="figure@9.[322,1267,193,1252]" captionTargetPageId="9" captionText="FIGURE 6. Helius minutus Loew, 1850, No. MB. J. 225 (male); A, head (after Krzemiński 1993); B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231113/files/figure.png" pageId="8" pageNumber="341">Fig. 6</figureCitation>
A, 10C): almost as long as the rostrum or slightly longer, the last palpal segment
<quantity id="4C869B15FFE9FFEAFB04A7A1390CB8E8" box="[1272,1372,1482,1506]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="341" unit="mm" value="0.22">0.22 mm</quantity>
long, almost equal two preceding segments together; first to third palpal segments
<quantity id="4C869B15FFE9FFEAFC08A7853807BB0C" box="[1012,1111,1518,1542]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="341" unit="mm" value="0.11">0.11 mm</quantity>
long.
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF3BA47A3968BB95" blockId="8.[151,1437,617,1949]" pageId="8" pageNumber="341">
Wing (
<figureCitation id="13452A75FFE9FFEAFEE9A47A3D0BBB23" box="[277,347,1553,1578]" captionStart="FIGURE 6" captionStartId="9.[151,250,1277,1299]" captionTargetBox="[329,1261,200,1245]" captionTargetId="figure@9.[322,1267,193,1252]" captionTargetPageId="9" captionText="FIGURE 6. Helius minutus Loew, 1850, No. MB. J. 225 (male); A, head (after Krzemiński 1993); B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231113/files/figure.png" pageId="8" pageNumber="341">Fig. 6</figureCitation>
C): wing
<quantity id="4C869B15FFE9FFEAFE3BA4793E01BB20" box="[455,593,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="8" pageNumber="341" unit="mm" value="4.6" valueMax="4.8" valueMin="4.4">4.4–4.8 mm</quantity>
length and
<quantity id="4C869B15FFE9FFEAFD26A4793F10BB20" box="[730,832,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="8" pageNumber="341" unit="mm" value="1.01">1.01 mm</quantity>
wide (
<typeStatus id="54C58852FFE9FFEAFC6CA4793FBFBB23" box="[912,1007,1554,1578]" pageId="8" pageNumber="341" type="neotype">neotype</typeStatus>
); pterostigma present, large and oval; vein Sc ending opposite 1/3 the length of Rs; cross-vein sc-r at the end of Sc; R1 ending opposite 1/4 of R2+3+4 length; r-r (R2) atrophied; R2+3+4 elongated, straight; d-cell elongated, closed; M3 longer than d-cell, M3 longer than half the length of R5; cross-vein m-cu almost in proximal 1/3 of d-cell base; A1 and A2 elongated, straight.
</paragraph>
<paragraph id="8BC136F0FFE9FFEAFF3BA4C2397FBBEF" blockId="8.[151,1437,617,1949]" pageId="8" pageNumber="341">Hypopygium: outer gonostylus short, broad, with two broad spikes apically; inner gonostylus rather short, lobe-shaped; gonocoxite basally equipped with large, broad process terminating into narrow, acute spine.</paragraph>
</subSubSection>
<subSubSection id="C364657BFFE9FFEAFF3BA49A3CB5BA94" pageId="8" pageNumber="341" type="discussion">
<paragraph id="8BC136F0FFE9FFEAFF3BA49A3CB5BA94" blockId="8.[151,1437,617,1949]" pageId="8" pageNumber="341">
<emphasis id="B90AEAE2FFE9FFEAFF3BA49A3D6BBA03" bold="true" box="[199,315,1777,1802]" pageId="8" pageNumber="341">Remarks.</emphasis>
Antenna is 16-segmented like in other species of the genus
<taxonomicName id="4C7E4D73FFE9FFEAFC11A4993866BA03" box="[1005,1078,1778,1802]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAE2FFE9FFEAFC11A4993866BA03" box="[1005,1078,1778,1802]" italics="true" pageId="8" pageNumber="341">Helius</emphasis>
</taxonomicName>
from Baltic amber, but the last flegellomere is comparatively small, distinctly shorter than the rest. The wing venation in specimens No. K-4265, Z-253 differs from the other specimens of this species by following features: Sc is ended opposite midlength of Rs, R1 is ended opposite midlength of R2+3+4, d-cell is very short and broad and cross-vein m-cu occur in half of d-cell length.
</paragraph>
</subSubSection>
</treatment>
</document>
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