194 lines
21 KiB
XML
194 lines
21 KiB
XML
<document id="CE2E8706AE131090CD4EF8FE0921E777" ID-CLB-Dataset="45540" ID-DOI="10.11646/zootaxa.3814.3.2" ID-GBIF-Dataset="708ea530-4c8d-473b-8f29-1a54dea92f28" ID-ISSN="1175-5326" ID-Zenodo-Dep="231107" ID-ZooBank="5391C375-896D-4794-84A6-1846DF9E407D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460434473942" checkinUser="plazi" docAuthor="Kania, Iwona" docDate="2014" docId="03D787E6FFE5FFE7FF6BA094393BBEBA" docLanguage="en" docName="zt03814p352.pdf" docOrigin="Zootaxa 3814 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helius fossilis Kania, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="338" masterDocId="FFEEFF9EFFE1FFE2FFFCA26B3C50BD09" masterDocTitle="Subfamily Limoniinae Speiser, 1909 (Diptera, Limoniidae) from Baltic amber (Eocene): the genus Helius Lepeletier & Serville, 1828" masterLastPageNumber="352" masterPageNumber="333" pageNumber="337" updateTime="1698331894480" updateUser="ExternalLinkService">
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<mods:mods id="CDA200A66CA0B891F90CB4F933EC593A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E30E043FB5738831C98BF4D04D516BC3">
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<mods:title id="84B6DF0CC4590D295CF49EF628B2C2B9">Subfamily Limoniinae Speiser, 1909 (Diptera, Limoniidae) from Baltic amber (Eocene): the genus Helius Lepeletier & Serville, 1828</mods:title>
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</mods:titleInfo>
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<mods:name id="B16D19E8084BA54CAEA38F09E2770398" type="personal">
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<mods:role id="7A327715493B19F335E8F66632ABDCED">
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<mods:roleTerm id="291B52F742EB3AEFA98383ADAF141B75">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="05A7CED0C370735F8F82D76C0BFBCBAD">Kania, Iwona</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DFD13CD0C74B7D01BE87A05A4A43F484">text</mods:typeOfResource>
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<mods:titleInfo id="F6EB9B47EF806EEF258BEBE25C54F706">
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<mods:title id="8CFDE53B1E43B683ADEF86D86FC84BB8">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="136AE2670CAA97BF098D2BEE9F1BBC64">
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<mods:date id="42C3CD41E025635F4799A49478DD26C1">2014</mods:date>
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<mods:detail id="A552EEFE37541C83B8E7519FD2F901E8" type="volume">
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<mods:number id="F04246F88BE170517F4776189DC1EB7E">3814</mods:number>
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</mods:detail>
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<mods:detail id="FB5336E83644A235931996373D8DF517" type="issue">
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<mods:number id="BBB065A55A8F758B1346632CBABF26DB">3</mods:number>
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</mods:detail>
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<mods:extent id="DFD4EA2400C597EE412B50C287278AAC" unit="page">
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<mods:start id="CB8378D5BE4991757F1FDDEBAF1F7948">333</mods:start>
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<mods:end id="BC43BDACA163CA8F059DCDCA0BE4D13C">352</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="CFD9456E8302C961C83A0AD473F5358E">journal article</mods:classification>
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<mods:identifier id="7FC3590B7E1862BF30F5B206F584707D" type="CLB-Dataset">45540</mods:identifier>
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<mods:identifier id="E5A8F6160F4F3D25ED8BD060DE3C9CC5" type="DOI">10.11646/zootaxa.3814.3.2</mods:identifier>
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<mods:identifier id="F0C61BCAAA26AE91E759371F410D4873" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C9CD57C7EA8C1DCF641A434D651C93CA" type="ZooBank">5391C375-896D-4794-84A6-1846DF9E407D</mods:identifier>
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</mods:mods>
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<treatment id="03D787E6FFE5FFE7FF6BA094393BBEBA" ID-DOI="http://doi.org/10.5281/zenodo.6138158" ID-GBIF-Taxon="119431373" ID-Zenodo-Dep="6138158" LSID="urn:lsid:plazi:treatment:03D787E6FFE5FFE7FF6BA094393BBEBA" httpUri="http://treatment.plazi.org/id/03D787E6FFE5FFE7FF6BA094393BBEBA" lastPageId="5" lastPageNumber="338" pageId="4" pageNumber="337">
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<subSubSection id="C364657BFFE5FFE6FF6BA0943D74BE33" pageId="4" pageNumber="337" type="nomenclature">
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<paragraph id="8BC136F0FFE5FFE6FF6BA0943DF6BE10" blockId="4.[151,422,767,827]" box="[151,422,767,793]" pageId="4" pageNumber="337">
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<heading id="D089819CFFE5FFE6FF6BA0943DF6BE10" bold="true" box="[151,422,767,793]" fontSize="11" level="1" pageId="4" pageNumber="337" reason="1">
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<emphasis id="B90AEAE2FFE5FFE6FF6BA0943DF6BE10" bold="true" box="[151,422,767,793]" pageId="4" pageNumber="337">
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<taxonomicName id="4C7E4D73FFE5FFE6FF6BA0943D11BE10" box="[151,321,767,793]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="fossilis" status="sp. nov.">
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<emphasis id="B90AEAE2FFE5FFE6FF6BA0943D11BE10" bold="true" box="[151,321,767,793]" italics="true" pageId="4" pageNumber="337">Helius fossilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2395799FFE5FFE6FEB4A0943DF6BE10" box="[328,422,767,793]" pageId="4" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC136F0FFE5FFE6FF6BA1493D74BE33" blockId="4.[151,422,767,827]" box="[151,292,802,827]" pageId="4" pageNumber="337">
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(
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<figureCitation id="13452A75FFE5FFE6FF63A1493CB3BE33" box="[159,227,802,827]" captionStart="FIGURE 3" captionStartId="4.[151,250,1975,1997]" captionTargetBox="[285,1298,976,1957]" captionTargetId="figure@4.[278,1307,975,1964]" captionTargetPageId="4" captionText="FIGURE 3. Helius fossilis sp. nov. No. 587 (RS 64) (male); A, head; B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231110/files/figure.png" pageId="4" pageNumber="337">Figs 3</figureCitation>
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A–C)
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</paragraph>
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</subSubSection>
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<subSubSection id="C364657BFFE5FFE6FF6BA1083D9FBEA9" pageId="4" pageNumber="337" type="materials_examined">
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<paragraph id="8BC136F0FFE5FFE6FF6BA1083D9FBEA9" blockId="4.[151,1436,867,928]" pageId="4" pageNumber="337">
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<emphasis id="B90AEAE2FFE5FFE6FF6BA1083DD1BE75" bold="true" box="[151,385,867,892]" pageId="4" pageNumber="337">Material examined.</emphasis>
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<typeStatus id="54C58852FFE5FFE6FE74A1083DA4BE75" box="[392,500,867,892]" pageId="4" pageNumber="337" type="holotype">Holotype</typeStatus>
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: No. 587 (RS64) (male) (
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<collectionCode id="ED6FAE35FFE5FFE6FCEEA10F3F39BE75" box="[786,873,868,892]" httpUri="http://grbio.org/cool/3q1q-apm2" name="Staatliches Museum fuer Naturkunde" pageId="4" pageNumber="337">SNMG</collectionCode>
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); additional material: No. 1690 (female) Coll. Ch. and H. W. Hoffeins (
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<collectionCode id="ED6FAE35FFE5FFE6FE83A1E33D93BEA9" box="[383,451,904,928]" pageId="4" pageNumber="337">SDEI</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF016678FFE5FFE6FF6BA5DC3E1CBAF9" httpUri="https://zenodo.org/record/231110/files/figure.png" pageId="4" pageNumber="337" targetBox="[285,1298,976,1957]" targetPageId="4">
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<paragraph id="8BC136F0FFE5FFE6FF6BA5DC3E1CBAF9" blockId="4.[151,1435,1975,2032]" pageId="4" pageNumber="337">
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<emphasis id="B90AEAE2FFE5FFE6FF6BA5DC3E53BAC4" bold="true" box="[151,515,1975,1998]" pageId="4" pageNumber="337">
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FIGURE 3.
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<taxonomicName id="4C7E4D73FFE5FFE6FEE5A5DC3DFCBAC7" box="[281,428,1975,1998]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="fossilis" status="sp. nov.">
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<emphasis id="B90AEAE2FFE5FFE6FEE5A5DC3DFCBAC7" bold="true" box="[281,428,1975,1998]" italics="true" pageId="4" pageNumber="337">Helius fossilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2395799FFE5FFE6FE4FA5D33E53BAC4" box="[435,515,1976,1997]" pageId="4" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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No. 587 (RS64) (male); A, head; B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation.
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</paragraph>
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</caption>
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<subSubSection id="C364657BFFE4FFE7FF3BA2FC3856BDB9" box="[199,1030,151,176]" pageId="5" pageNumber="338" type="etymology">
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<paragraph id="8BC136F0FFE4FFE7FF3BA2FC3856BDB9" blockId="5.[151,1437,151,952]" box="[199,1030,151,176]" pageId="5" pageNumber="338">
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<emphasis id="B90AEAE2FFE4FFE7FF3BA2FC3D1EBDB9" bold="true" box="[199,334,151,176]" pageId="5" pageNumber="338">Etymology.</emphasis>
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From Latin
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<taxonomicName id="4C7E4D73FFE4FFE7FE26A2F33E78BDB9" box="[474,552,152,176]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="fossilis">
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<emphasis id="B90AEAE2FFE4FFE7FE26A2F33E78BDB9" box="[474,552,152,176]" italics="true" pageId="5" pageNumber="338">fossilis</emphasis>
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</taxonomicName>
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, fossil. The species epithet is an adjective.
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</paragraph>
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</subSubSection>
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<subSubSection id="C364657BFFE4FFE7FF3BA2D13902BC11" pageId="5" pageNumber="338" type="diagnosis">
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<paragraph id="8BC136F0FFE4FFE7FF3BA2D13902BC11" blockId="5.[151,1437,151,952]" pageId="5" pageNumber="338">
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<emphasis id="B90AEAE2FFE4FFE7FF3BA2D13D11BDDA" bold="true" box="[199,321,186,211]" pageId="5" pageNumber="338">Diagnosis.</emphasis>
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Rostrum very elongated, approximately three times as long as the head, longer than palpus, longer than antenna; palpus 0.33x the length of the rostrum; the last palpal segment approximately half the length of all the preceding segments together; vein r-m rather long, well expressed, cross-vein m-cu in 1/2 of d-cell base.
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</paragraph>
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</subSubSection>
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<subSubSection id="C364657BFFE4FFE7FF3BA34938ECBFEA" pageId="5" pageNumber="338" type="description">
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<paragraph id="8BC136F0FFE4FFE7FF3BA3493843BC32" blockId="5.[151,1437,151,952]" box="[199,1043,290,315]" pageId="5" pageNumber="338">
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<emphasis id="B90AEAE2FFE4FFE7FF3BA3493D09BC32" bold="true" box="[199,345,290,315]" pageId="5" pageNumber="338">Description.</emphasis>
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Body (
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<figureCitation id="13452A75FFE4FFE7FE57A3493DAABC32" box="[427,506,290,315]" captionStart="FIGURE 11" captionStartId="14.[151,250,1881,1903]" captionTargetBox="[201,1391,193,1853]" captionTargetId="figure@14.[192,1394,193,1860]" captionTargetPageId="14" captionText="FIGURE 11. Morphology of Helius from Baltic amber. A – B H. fossilis sp. nov., No. 587 (RS 64) (male), holotype, A. the body, lateral view; B. head, lateral view C. H. hoffeinsorum sp. nov., No. 1462 (male), hypopygium; D – E H. formosus Krzemiński, 1993, No. 225 (female), holotype, D. antennae, E. wing. Abbreviation of male hypopygium: gx—gonocoxite, ing—inner gonostylus, oug—outer gonostylus, aed—aedeagus, p—paramere." httpUri="https://zenodo.org/record/231118/files/figure.png" pageId="5" pageNumber="338">Fig. 11</figureCitation>
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A): the length of specimen
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<quantity id="4C869B15FFE4FFE7FCD0A3493FC0BC33" box="[812,912,290,315]" metricMagnitude="-3" metricUnit="m" metricValue="4.36" pageId="5" pageNumber="338" unit="mm" value="4.36">4.36 mm</quantity>
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||
(
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||
<typeStatus id="54C58852FFE4FFE7FC62A3493857BC32" box="[926,1031,290,315]" pageId="5" pageNumber="338" type="holotype">holotype</typeStatus>
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).
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</paragraph>
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<paragraph id="8BC136F0FFE4FFE7FF3BA32E39CABCCF" blockId="5.[151,1437,151,952]" pageId="5" pageNumber="338">
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Head (
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<figureCitation id="13452A75FFE4FFE7FEEEA32E3D0ABC57" box="[274,346,325,350]" captionStart="FIGURE 3" captionStartId="4.[151,250,1975,1997]" captionTargetBox="[285,1298,976,1957]" captionTargetId="figure@4.[278,1307,975,1964]" captionTargetPageId="4" captionText="FIGURE 3. Helius fossilis sp. nov. No. 587 (RS 64) (male); A, head; B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231110/files/figure.png" pageId="5" pageNumber="338">Figs 3</figureCitation>
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A, 11B): head width
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<quantity id="4C869B15FFE4FFE7FDB7A32D3EE4BC54" box="[587,692,326,350]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="338" unit="mm" value="0.48">0.48 mm</quantity>
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; rostrum length
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<quantity id="4C869B15FFE4FFE7FC93A32E3F86BC54" box="[879,982,325,350]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="5" pageNumber="338" unit="mm" value="1.68">1.68 mm</quantity>
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, very elongated (
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<figureCitation id="13452A75FFE4FFE7FB60A32E388FBC57" box="[1180,1247,325,350]" captionStart="FIGURE 3" captionStartId="4.[151,250,1975,1997]" captionTargetBox="[285,1298,976,1957]" captionTargetId="figure@4.[278,1307,975,1964]" captionTargetPageId="4" captionText="FIGURE 3. Helius fossilis sp. nov. No. 587 (RS 64) (male); A, head; B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231110/files/figure.png" pageId="5" pageNumber="338">Fig. 3</figureCitation>
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B); antenna
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<quantity id="4C869B15FFE4FFE7FA97A32D3C93BC76" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="5" pageNumber="338" unit="mm" value="1.13">1.13 mm</quantity>
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long (
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<typeStatus id="54C58852FFE4FFE7FEEEA30C3D2ABC89" box="[274,378,359,384]" pageId="5" pageNumber="338" type="holotype">holotype</typeStatus>
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), 16-segmented, shorter than rostrum; scape cylindrical, large; pedicel large, its distal part conspicuously widened; first flagellomere barrel-shaped, flagellomeres 2–16 cylindrical, rather short, with four elongated setae each; setae almost the same length or slightly longer than the length of the segments bearing them.
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</paragraph>
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<paragraph id="8BC136F0FFE4FFE7FF3BA3A43F54BF27" blockId="5.[151,1437,151,952]" pageId="5" pageNumber="338">
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Palpus (
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||
<figureCitation id="13452A75FFE4FFE7FEDEA3A43D36BCE1" box="[290,358,463,488]" captionStart="FIGURE 3" captionStartId="4.[151,250,1975,1997]" captionTargetBox="[285,1298,976,1957]" captionTargetId="figure@4.[278,1307,975,1964]" captionTargetPageId="4" captionText="FIGURE 3. Helius fossilis sp. nov. No. 587 (RS 64) (male); A, head; B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231110/files/figure.png" pageId="5" pageNumber="338">Fig. 3</figureCitation>
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A):
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||
<quantity id="4C869B15FFE4FFE7FE68A3A43DAABCEE" box="[404,506,463,488]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="338" unit="mm" value="0.53">0.53 mm</quantity>
|
||
long (
|
||
<typeStatus id="54C58852FFE4FFE7FDB8A3A43EFCBCE1" box="[580,684,463,488]" pageId="5" pageNumber="338" type="holotype">holotype</typeStatus>
|
||
); the length of last palpal segment
|
||
<quantity id="4C869B15FFE4FFE7FBC1A3BB38F4BCEE" box="[1085,1188,464,488]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="338" unit="mm" value="0.22">0.22 mm</quantity>
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||
, penultimate
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||
<quantity id="4C869B15FFE4FFE7FAC2A3BB39C8BCEE" box="[1342,1432,464,488]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="338" unit="mm" value="0.1">0.1 mm</quantity>
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||
, the second segment
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||
<quantity id="4C869B15FFE4FFE7FE80A3983D83BF03" box="[380,467,499,523]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="338" unit="mm" value="0.1">0.1 mm</quantity>
|
||
(
|
||
<typeStatus id="54C58852FFE4FFE7FE1FA3993E1BBF02" box="[483,587,498,523]" pageId="5" pageNumber="338" type="holotype">holotype</typeStatus>
|
||
); the last palpal segment approximately half the length of all the preceding together, equal the length of two penultimate segments.
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||
</paragraph>
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||
<paragraph id="8BC136F0FFE4FFE7FF3BA05C393FBFB7" blockId="5.[151,1437,151,952]" pageId="5" pageNumber="338">
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||
Wing (
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||
<figureCitation id="13452A75FFE4FFE7FEEFA05C3D05BF59" box="[275,341,567,592]" captionStart="FIGURE 3" captionStartId="4.[151,250,1975,1997]" captionTargetBox="[285,1298,976,1957]" captionTargetId="figure@4.[278,1307,975,1964]" captionTargetPageId="4" captionText="FIGURE 3. Helius fossilis sp. nov. No. 587 (RS 64) (male); A, head; B, relation between the length of palpus (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231110/files/figure.png" pageId="5" pageNumber="338">Fig. 3</figureCitation>
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C): wing
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<quantity id="4C869B15FFE4FFE7FE43A0533E73BF46" box="[447,547,568,592]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="338" unit="mm" value="4.9">4.90 mm</quantity>
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length and
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||
<quantity id="4C869B15FFE4FFE7FD59A0533EABBF46" box="[677,763,568,592]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="338" unit="mm" value="1.4">1.4 mm</quantity>
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wide (
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||
<typeStatus id="54C58852FFE4FFE7FCBBA05C3FFFBF59" box="[839,943,567,592]" pageId="5" pageNumber="338" type="holotype">holotype</typeStatus>
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||
); pterostigma absent; vein Sc not elongated, ending before midlength of Rs; cross-vein sc-r behind the end of Sc; R1 ending at the level of 1/3 of R2+3+4 length; r- r (R2) atrophied; R2+3+4 elongated, straight; d-cell closed, very broad and long; M3 twice the length of d-cell, M3 longer than half the length of R5; cross-vein m-cu in 1/2 of d-cell base; A1 and A2 elongated, distinctly straight.
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</paragraph>
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<paragraph id="8BC136F0FFE4FFE7FF3BA0A038ECBFEA" blockId="5.[151,1437,151,952]" box="[199,1212,715,740]" pageId="5" pageNumber="338">
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Hypopygium:
|
||
<quantity id="4C869B15FFE4FFE7FE95A0A03D9DBFEA" box="[361,461,715,740]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="5" pageNumber="338" unit="mm" value="0.36">0.36 mm</quantity>
|
||
long, not well visible; gonocoxite elongated and relatively narrow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364657BFFE4FFE7FF3BA086393BBEBA" pageId="5" pageNumber="338" type="discussion">
|
||
<paragraph id="8BC136F0FFE4FFE7FF3BA086393BBEBA" blockId="5.[151,1437,151,952]" pageId="5" pageNumber="338">
|
||
<emphasis id="B90AEAE2FFE4FFE7FF3BA0863D69BE0F" bold="true" box="[199,313,749,774]" pageId="5" pageNumber="338">Remarks.</emphasis>
|
||
The species
|
||
<taxonomicName id="4C7E4D73FFE4FFE7FE37A0853E69BE0C" box="[459,569,749,773]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="fossilis" status="sp. nov.">
|
||
<emphasis id="B90AEAE2FFE4FFE7FE37A0853E69BE0C" box="[459,569,749,773]" italics="true" pageId="5" pageNumber="338">H. fossilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2395799FFE4FFE7FDBEA0863ECABE0F" box="[578,666,749,774]" pageId="5" pageNumber="338" rank="species">
|
||
<emphasis id="B90AEAE2FFE4FFE7FDBEA0863ECABE0F" bold="true" box="[578,666,749,774]" pageId="5" pageNumber="338">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
distinctly differ from most of other species of this genus from Baltic amber in very elongated rostrum, which characterizes also
|
||
<taxonomicName id="4C7E4D73FFE4FFE7FCD9A17A3FD9BE21" box="[805,905,785,808]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="mutus">
|
||
<emphasis id="B90AEAE2FFE4FFE7FCD9A17A3FD9BE21" box="[805,905,785,808]" italics="true" pageId="5" pageNumber="338">H. mutus</emphasis>
|
||
</taxonomicName>
|
||
. In both species rostrum is shorter than half the body length. They differ in following features: in
|
||
<taxonomicName id="4C7E4D73FFE4FFE7FD48A15F3F46BE42" box="[692,790,820,843]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="mutus">
|
||
<emphasis id="B90AEAE2FFE4FFE7FD48A15F3F46BE42" box="[692,790,820,843]" italics="true" pageId="5" pageNumber="338">H. mutus</emphasis>
|
||
</taxonomicName>
|
||
antenna is about half the length of rostrum, palpus is about half the length of antenna, in
|
||
<taxonomicName id="4C7E4D73FFE4FFE7FE20A13D3E19BE64" box="[476,585,853,877]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="fossilis">
|
||
<emphasis id="B90AEAE2FFE4FFE7FE20A13D3E19BE64" box="[476,585,853,877]" italics="true" pageId="5" pageNumber="338">H. fossilis</emphasis>
|
||
</taxonomicName>
|
||
antenna is much longer than half the length of rostrum, palpus is longer than half the antenna length. These species differ also in wing venation details. In contrast to
|
||
<taxonomicName id="4C7E4D73FFE4FFE7FB6AA11238ADBE99" box="[1174,1277,889,912]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="mutus">
|
||
<emphasis id="B90AEAE2FFE4FFE7FB6AA11238ADBE99" box="[1174,1277,889,912]" italics="true" pageId="5" pageNumber="338">H. mutus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E4D73FFE4FFE7FAF2A11239D0BE99" box="[1294,1408,888,912]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="fossilis">
|
||
<emphasis id="B90AEAE2FFE4FFE7FAF2A11239D0BE99" box="[1294,1408,888,912]" italics="true" pageId="5" pageNumber="338">H. fossilis</emphasis>
|
||
</taxonomicName>
|
||
is characterized by wing without pterostigma, vein R1 ending in 1/3 of R2+3+4 (opposite 1/4 of its length in
|
||
<taxonomicName id="4C7E4D73FFE4FFE7FB04A1F7390BBEBA" box="[1272,1371,924,947]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="mutus">
|
||
<emphasis id="B90AEAE2FFE4FFE7FB04A1F7390BBEBA" box="[1272,1371,924,947]" italics="true" pageId="5" pageNumber="338">H. mutus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |