153 lines
18 KiB
XML
153 lines
18 KiB
XML
<document id="1C9B215A4C54F5E20FA3F28D19433A0C" ID-DOI="10.5281/zenodo.4665621" ID-ISSN="1638-9395" ID-Zenodo-Dep="4665621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617712844033" checkinUser="felipe" docAuthor="Venczel, Márton & Stiuc, Emanoil" docDate="2008" docId="03D08790FFD6FFFB5656A5EEFB3A54E9" docLanguage="en" docName="Geodiversitas.30.4.731-763.pdf" docOrigin="Geodiversitas 30 (4)" docStyle="DocumentStyle:9DCE1246056655C2F9CBE9822E831E4E.3:Geodiversitas.2000-2008.journal_article.type3" docStyleId="9DCE1246056655C2F9CBE9822E831E4E" docStyleName="Geodiversitas.2000-2008.journal_article.type3" docStyleVersion="3" docTitle="Pelophylax Fitzinger 1843" docType="treatment" docVersion="3" lastPageNumber="750" masterDocId="FFE9FFE8FFC5FFE85703A736FFA55105" masterDocTitle="Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania" masterLastPageNumber="763" masterPageNumber="731" pageNumber="750" updateTime="1698901057390" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods id="01F19C8ADEFC55CB35896F0260C05C25" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="003798198F4787D60AF290F4A9E563FF">
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<mods:title id="0E189EC76EDCBA729B9FD245592F059A">Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania</mods:title>
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</mods:titleInfo>
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<mods:name id="FD707949E5CF5866C20781ACA990C303" type="personal">
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<mods:role id="842B46B91688E752BEBEBED06FB2B0AE">
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<mods:roleTerm id="FAE2F2CAEDC777B078242EF72DA78843">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="232EF3EAE8E84F6E8B1A76A9C8E2F22D">Venczel, Márton</mods:namePart>
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<mods:affiliation id="BF1381E105A3C8610097EEDBF15F0245">Tării ‚ Crişurilor Museum, B-dul Dacia 1 - 3, RO- 410464 Oradea (Romania) mvenczel @ rdslink. ro</mods:affiliation>
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<mods:nameIdentifier id="E1F243B6A009743A4F567E9D6E374060" type="email">mvenczel@rdslink.ro</mods:nameIdentifier>
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</mods:name>
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<mods:name id="AAE6C9C33C7677C14BFCFC5A11213998" type="personal">
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<mods:role id="878307535CA7B768CD65FC0B079283A2">
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<mods:roleTerm id="354565448849DF8F15FD7A169D7B4F83">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7746F567A2908B06865C64984ADD256E">Stiuc, Emanoil</mods:namePart>
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<mods:affiliation id="DCF4C4AD7FC6CA9CB87D9F5A12AB27B5">Speleological Institute “ Emil Racovit, ă ”, Calea 13 Septembrie nr. 13, RO- 050711 Bucureşti (Romania) stiucaemil @ yahoo. com.</mods:affiliation>
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<mods:nameIdentifier id="05D248576ED518D8E642C0C5CEBC46F6" type="email">stiucaemil@yahoo.com</mods:nameIdentifier>
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<mods:typeOfResource id="85A568B514365522693D707A01B82604">text</mods:typeOfResource>
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<mods:title id="A9688F2B4F20D7BD6FF83020A41708A8">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="663D89642472DD4EDADE9324F32A2B5A">
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<mods:date id="F6D69FF1F1170EE175E4666C9DD48A3F">2008</mods:date>
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<mods:detail id="647F98465A7A469F73B05DCBEBA2B262" type="volume">
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<mods:number id="8A6894626685ADF12235EFFCD393E02F">30</mods:number>
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</mods:detail>
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<mods:number id="D5ABC014CDEBE7F844C0F5A9F1BF120A">4</mods:number>
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</mods:detail>
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<mods:start id="871851753AAE54799601876AEE4BDC6A">731</mods:start>
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<mods:identifier id="A2FC68974D6496C888EC750389B6D6BD" type="DOI">10.5281/zenodo.4665621</mods:identifier>
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<mods:identifier id="6A2ACA4206463DDA46A2FE574E84D111" type="ISSN">1638-9395</mods:identifier>
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<treatment id="03D08790FFD6FFFB5656A5EEFB3A54E9" LSID="urn:lsid:plazi:treatment:03D08790FFD6FFFB5656A5EEFB3A54E9" httpUri="http://treatment.plazi.org/id/03D08790FFD6FFFB5656A5EEFB3A54E9" lastPageNumber="750" pageId="19" pageNumber="750">
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<subSubSection id="C363650DFFD6FFFB5656A5EEFE4F5211" pageId="19" pageNumber="750" type="nomenclature">
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<paragraph id="8BC63686FFD6FFFB5656A5EEFE4F5211" blockId="19.[341,494,728,788]" pageId="19" pageNumber="750">
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<heading id="D08E81EAFFD6FFFB5656A5EEFE4F5211" centered="true" fontSize="11" level="2" pageId="19" pageNumber="750" reason="2">
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<taxonomicName id="4C794D05FFD6FFFB5656A5EEFE6C53F6" ID-CoL="6KDF" authorityName="Fitzinger" authorityYear="1843" box="[341,457,728,755]" class="Amphibia" family="Ranidae" genus="Pelophylax" kingdom="Animalia" order="Anura" pageId="19" pageNumber="750" phylum="Chordata" rank="genus">
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<emphasis id="B90DEA94FFD6FFFB5656A5EEFE6C53F6" bold="true" box="[341,457,728,755]" italics="true" pageId="19" pageNumber="750">Pelophylax</emphasis>
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</taxonomicName>
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sp. (
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<figureCitation id="13422A03FFD6FFFB5662A5CFFE0E5211" box="[353,427,761,788]" captionStart="FIG" captionStartId="18.[714,725,1008,1025]" captionTargetBox="[738,1228,254,963]" captionTargetId="figure-21@18.[715,1245,233,977]" captionTargetPageId="18" captionText="FIG. 11. — A-C, Hyla cf.arborea; A, ilium (ISER Tt-0411/1);B, same (ISER Tt-0411/2); C, prearticular (ISER Tt-0410);D-G, Pelophylax sp.; D, presacral vertebra (ISER Tt-0421/1); E, humerus (ISER Tt- 0422/1); F, ilium (ISER Tt-0423/1); G, same (ISER Tt-0423/2);A, B, G, left lateral views; C, D, dorsal views; E, ventral view; F, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665655" httpUri="https://zenodo.org/record/4665655/files/figure.png" pageId="19" pageNumber="750">Fig. 11</figureCitation>
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D-G)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C363650DFFD6FFFB5794A404FDF0529C" pageId="19" pageNumber="750" type="materials_examined">
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<paragraph id="8BC63686FFD6FFFB5794A404FDF0529C" blockId="19.[149,686,818,921]" pageId="19" pageNumber="750">
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MATERIAL EXAMINED. —Two premaxillae(
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<collectionCode id="ED68AE43FFD6FFFB5515A404FDEC524F" box="[534,585,818,842]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="19" pageNumber="750">ISER</collectionCode>
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Tt-0420/1 and 2), one presacral vertebra (
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<collectionCode id="ED68AE43FFD6FFFB56C9A47AFDA55261" box="[458,512,844,868]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="19" pageNumber="750">ISER</collectionCode>
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Tt-0421/1), one sacral vertebra (
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<collectionCode id="ED68AE43FFD6FFFB5631A451FECD527A" box="[306,360,871,895]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="19" pageNumber="750">ISER</collectionCode>
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Tt-0421/2), three humeri (
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<collectionCode id="ED68AE43FFD6FFFB557BA451FD0B527A" box="[632,686,871,895]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="19" pageNumber="750">ISER</collectionCode>
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Tt-0422/1-3), seven ilia (
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<collectionCode id="ED68AE43FFD6FFFB568DA4B7FE61529C" box="[398,452,897,921]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="19" pageNumber="750">ISER</collectionCode>
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Tt-0423/1-7).
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</paragraph>
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</subSubSection>
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<subSubSection id="C363650DFFD6FFFB5794A481FBC25514" pageId="19" pageNumber="750" type="description">
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<paragraph id="8BC63686FFD6FFFB5794A481FE8D52CA" blockId="19.[149,684,951,1199]" box="[151,296,951,977]" pageId="19" pageNumber="750">DESCRIPTION</paragraph>
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<paragraph id="8BC63686FFD6FFFB5794A4E0FEA252F5" blockId="19.[149,684,951,1199]" box="[151,263,982,1008]" pageId="19" pageNumber="750">
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<emphasis id="B90DEA94FFD6FFFB5794A4E0FEA252F5" box="[151,263,982,1008]" italics="true" pageId="19" pageNumber="750">Premaxilla</emphasis>
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</paragraph>
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<paragraph id="8BC63686FFD6FFFB5794A4C0FECE55AA" blockId="19.[149,684,951,1199]" pageId="19" pageNumber="750">
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The anterior surface is smooth and flat with a welldefined lateral crest. The tip of the alary process in both specimens is broken off. The
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<emphasis id="B90DEA94FFD6FFFB56EDA303FD09554A" box="[494,684,1077,1103]" italics="true" pageId="19" pageNumber="750">lamina horizontalis</emphasis>
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is well developed, but the
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<emphasis id="B90DEA94FFD6FFFB56B4A360FD9A556A" box="[439,575,1109,1135]" italics="true" pageId="19" pageNumber="750">pars palatina</emphasis>
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is broken off. The number of preserved tooth positions is 11 and 12 respectively.
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</paragraph>
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<paragraph id="8BC63686FFD6FFFB5794A3E2FEE955EB" blockId="19.[149,686,1236,1516]" box="[151,332,1236,1262]" pageId="19" pageNumber="750">
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<emphasis id="B90DEA94FFD6FFFB5794A3E2FEE955EB" box="[151,332,1236,1262]" italics="true" pageId="19" pageNumber="750">Presacral vertebra</emphasis>
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</paragraph>
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<paragraph id="8BC63686FFD6FFFB5794A3C5FEFF54E9" blockId="19.[149,686,1236,1516]" pageId="19" pageNumber="750">
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The centrum is procoelous and relatively short with the condyle flattened dorsoventrally. The neural arch is of non-imbricate
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<typeStatus id="54C28824FFD6FFFB56A3A205FE6B544B" box="[416,462,1331,1358]" pageId="19" pageNumber="750">type</typeStatus>
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and provided with a low but distinct neural crest. The only preserved transverse process is straight and tends to widen distally (
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<figureCitation id="13422A03FFD6FFFB57F0A2A4FEF254A8" box="[243,343,1426,1453]" captionStart="FIG" captionStartId="18.[714,725,1008,1025]" captionTargetBox="[738,1228,254,963]" captionTargetId="figure-21@18.[715,1245,233,977]" captionTargetPageId="18" captionText="FIG. 11. — A-C, Hyla cf.arborea; A, ilium (ISER Tt-0411/1);B, same (ISER Tt-0411/2); C, prearticular (ISER Tt-0410);D-G, Pelophylax sp.; D, presacral vertebra (ISER Tt-0421/1); E, humerus (ISER Tt- 0422/1); F, ilium (ISER Tt-0423/1); G, same (ISER Tt-0423/2);A, B, G, left lateral views; C, D, dorsal views; E, ventral view; F, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665655" httpUri="https://zenodo.org/record/4665655/files/figure.png" pageId="19" pageNumber="750">Fig. 11D</figureCitation>
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). Based on the latter character it is assumed that the specimen represents the third presacral vertebra.
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</paragraph>
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<paragraph id="8BC63686FFD6FFFB5794A126FE95572F" blockId="19.[149,685,1552,1706]" box="[151,304,1552,1578]" pageId="19" pageNumber="750">
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<emphasis id="B90DEA94FFD6FFFB5794A126FE95572F" box="[151,304,1552,1578]" italics="true" pageId="19" pageNumber="750">Sacral vertebra</emphasis>
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</paragraph>
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<paragraph id="8BC63686FFD6FFFB5794A106FD0857AF" blockId="19.[149,685,1552,1706]" pageId="19" pageNumber="750">The centrum is procoelous and displays two posterior condyles for the articulation with the urostyle. The neural arch is extremely short and provided with a low keel extending laterally to the dorsal</paragraph>
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<paragraph id="8BC63686FFD6FFFB579AA199FC32503D" blockId="19.[153,686,1711,1738]" lastBlockId="19.[714,1248,254,312]" pageId="19" pageNumber="750">surfaces of the transverse processes. Distally the latter structures are broken off but were presumably rather slender.</paragraph>
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<paragraph id="8BC63686FFD6FFFB55C9A66BFC8F5073" blockId="19.[712,1248,349,566]" box="[714,810,349,374]" pageId="19" pageNumber="750">
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<emphasis id="B90DEA94FFD6FFFB55C9A66BFC8F5073" box="[714,810,349,374]" italics="true" pageId="19" pageNumber="750">Humerus</emphasis>
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</paragraph>
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<paragraph id="8BC63686FFD6FFFB55C9A64AFB1D5333" blockId="19.[712,1248,349,566]" pageId="19" pageNumber="750">
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Only distal parts of humeri are preserved. The distal humeral ball is round and there is a well-developed ventral cubital fossa (
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<figureCitation id="13422A03FFD6FFFB54A7A68AFBA750D2" box="[932,1026,444,471]" captionStart="FIG" captionStartId="18.[714,725,1008,1025]" captionTargetBox="[738,1228,254,963]" captionTargetId="figure-21@18.[715,1245,233,977]" captionTargetPageId="18" captionText="FIG. 11. — A-C, Hyla cf.arborea; A, ilium (ISER Tt-0411/1);B, same (ISER Tt-0411/2); C, prearticular (ISER Tt-0410);D-G, Pelophylax sp.; D, presacral vertebra (ISER Tt-0421/1); E, humerus (ISER Tt- 0422/1); F, ilium (ISER Tt-0423/1); G, same (ISER Tt-0423/2);A, B, G, left lateral views; C, D, dorsal views; E, ventral view; F, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665655" httpUri="https://zenodo.org/record/4665655/files/figure.png" pageId="19" pageNumber="750">Fig. 11E</figureCitation>
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). In ISER Tt-0422/1 the
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<emphasis id="B90DEA94FFD6FFFB55F3A6EDFC1250F0" box="[752,951,475,501]" italics="true" pageId="19" pageNumber="750">epicondylus medialis</emphasis>
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as well as the
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<emphasis id="B90DEA94FFD6FFFB5343A6EAFB6950F0" box="[1088,1228,475,501]" italics="true" pageId="19" pageNumber="750">crista medialis</emphasis>
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is distinctly larger than in ISER Tt-0422/2.The latter two attributes may be related to sexual dimorphism.
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</paragraph>
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<paragraph id="8BC63686FFD6FFFB55C9A56CFCA15371" blockId="19.[712,1249,602,1041]" box="[714,772,602,628]" pageId="19" pageNumber="750">
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<emphasis id="B90DEA94FFD6FFFB55C9A56CFCA15371" box="[714,772,602,628]" italics="true" pageId="19" pageNumber="750">Ilium</emphasis>
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</paragraph>
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<paragraph id="8BC63686FFD6FFFB55C9A54CFBC25514" blockId="19.[712,1249,602,1041]" pageId="19" pageNumber="750">
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Based on the morphology of ISER Tt-0423/1, the supraacetabular and the preacetabular regions are roughly of equal height. The acetabulum is nearly circular with the acetabular rim projecting above the preacetabular region and the preacetabular fossa is absent or extremely shallow (
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<figureCitation id="13422A03FFD6FFFB531DA42EFB3A5236" box="[1054,1183,792,819]" captionStart="FIG" captionStartId="18.[714,725,1008,1025]" captionTargetBox="[738,1228,254,963]" captionTargetId="figure-21@18.[715,1245,233,977]" captionTargetPageId="18" captionText="FIG. 11. — A-C, Hyla cf.arborea; A, ilium (ISER Tt-0411/1);B, same (ISER Tt-0411/2); C, prearticular (ISER Tt-0410);D-G, Pelophylax sp.; D, presacral vertebra (ISER Tt-0421/1); E, humerus (ISER Tt- 0422/1); F, ilium (ISER Tt-0423/1); G, same (ISER Tt-0423/2);A, B, G, left lateral views; C, D, dorsal views; E, ventral view; F, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665655" httpUri="https://zenodo.org/record/4665655/files/figure.png" pageId="19" pageNumber="750">Fig. 11F, G</figureCitation>
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). The tuber superior is high and positioned anterodorsally to the acetabulum. In ISER Tt-0423/2 the tuber superior is damaged but its ventral margin is produced in a distinct overhang delimiting dorsally the
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<emphasis id="B90DEA94FFD6FFFB55F7A480FC4852D5" box="[756,1005,950,976]" italics="true" pageId="19" pageNumber="750">fossula tuberis superioris</emphasis>
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. The iliac crest is thin and it was presumably high, but in all the available specimens its dorsal margin is lacking.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C363650DFFD6FFFB55C9A300FB3A54E9" pageId="19" pageNumber="750" type="discussion">
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<paragraph id="8BC63686FFD6FFFB55C9A300FBF0544B" blockId="19.[714,1250,1078,1358]" pageId="19" pageNumber="750">
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REMARKS All the above remains show a close morphological resemblance with those of the genus
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<taxonomicName id="4C794D05FFD6FFFB536DA343FC3655AA" authority="(Frost et al. 2006)" baseAuthorityName="Frost" baseAuthorityYear="2006" class="Amphibia" family="Ranidae" genus="Pelophylax" kingdom="Animalia" order="Anura" pageId="19" pageNumber="750" phylum="Chordata" rank="genus">
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<emphasis id="B90DEA94FFD6FFFB536DA343FB7A558A" box="[1134,1247,1141,1167]" italics="true" pageId="19" pageNumber="750">Pelophylax</emphasis>
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(Frost
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<emphasis id="B90DEA94FFD6FFFB5411A3A3FCE255AB" box="[786,839,1172,1198]" italics="true" pageId="19" pageNumber="750">et al.</emphasis>
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2006)
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</taxonomicName>
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. The variation observed in the shape and thickness of the ventral margin of the tuber superior is also reminiscent of specimens described from the late middle Miocene (MN 7) of Mátraszőlős,
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<collectingCountry id="F36E7616FFD6FFFB547CA225FC47542B" box="[895,994,1299,1326]" name="Hungary" pageId="19" pageNumber="750">Hungary</collectingCountry>
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under the name
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<taxonomicName id="4C794D05FFD6FFFB53A4A222FC835448" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="19" pageNumber="750" phylum="Chordata" rank="species" species="esculenta">
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<emphasis id="B90DEA94FFD6FFFB53A4A222FC835448" italics="true" pageId="19" pageNumber="750">Rana esculenta</emphasis>
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</taxonomicName>
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synklepton (
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<bibRefCitation id="EFE84B77FFD6FFFB54B2A205FBEC544B" author="VENCZEL M." box="[945,1097,1331,1358]" pageId="19" pageNumber="750" pagination="151 - 174" refId="ref18344" refString="VENCZEL M. 2004. - Middle Miocene anurans from the Carpathian Basin. Palaeontographica abt. A. 271: 151 - 174." type="journal article" year="2004">Venczel 2004</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BC63686FFD6FFFB5414A2A4FB3A54E9" blockId="19.[778,1183,1426,1548]" pageId="19" pageNumber="750">
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Class
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<taxonomicName id="4C794D05FFD6FFFB5456A2A4FB3654A8" authority="McCartney, 1802" authorityName="McCartney" authorityYear="1802" box="[853,1171,1426,1453]" class="Reptilia" kingdom="Animalia" pageId="19" pageNumber="750" phylum="Chordata" rank="class">REPTILIA McCartney, 1802</taxonomicName>
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Order
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<taxonomicName id="4C794D05FFD6FFFB545EA287FB3354C9" authority="Merrem, 1820" authorityName="Merrem" authorityYear="1820" box="[861,1174,1457,1484]" class="Reptilia" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="750" phylum="Chordata" rank="order">SQUAMATA Merrem, 1820</taxonomicName>
|
||
Suborder
|
||
<taxonomicName id="4C794D05FFD6FFFB5477A2E7FB3A54E9" authority="Owen, 1842" authorityName="Owen" authorityYear="1842" box="[884,1183,1489,1516]" class="Reptilia" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="750" phylum="Chordata" rank="subOrder" subOrder="Lacertilia">LACERTILIA Owen, 1842</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |