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<document id="B7C6B0D52A7D15FABA9DA9CAA3C4CD69" ID-DOI="10.1649/0010-065x-69.3.463" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106050" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175225603" checkinUser="diego" docAuthor="Filippini, V., Galante, E. &amp; Micó, E." docDate="2015" docId="03CFA3486B6CFFC8FD6AFAB459B6FBBE" docLanguage="en" docName="0010-065X-69.3.463.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-69.3.463" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Anomala strigodermoides Filippini, Galante, and Mico, new species" docType="treatment" docVersion="3" lastPageNumber="474" masterDocId="FFF6DB306B66FFC3FFBEFFE95A73FFC7" masterDocTitle="Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica" masterLastPageNumber="476" masterPageNumber="463" pageNumber="473" updateTime="1699636354092" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9BDF85773D215FB74BC0DF090DD8A105" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C213C55D6D6012DB804C38E2334B49F6">
<mods:title id="1B40FD542B5E1B0C43DDA244B91FB624">Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica</mods:title>
</mods:titleInfo>
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<mods:namePart id="68DBE107676463724938455C89F8FE2E">Filippini, V.</mods:namePart>
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<mods:namePart id="9123EA41970232CE9692C60734623BCB">Galante, E.</mods:namePart>
</mods:name>
<mods:name id="2BB1CE28ED4518CDCFEA90DD776ED46C" type="personal">
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<mods:roleTerm id="E24058189C29AA96CCFF29301741EE37">Author</mods:roleTerm>
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<mods:namePart id="565D9B70EA9C1E5082565553585F6069">Micó, E.</mods:namePart>
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<mods:typeOfResource id="B20CAF973D109C75A1AE2D7A9E8E3DCF">text</mods:typeOfResource>
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<mods:title id="217DC04F453B635C8B28CAFAD0C5F56E">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="F40452913636F04AC237D8E7DE558C4F">
<mods:date id="338FCF8700AEF8F8B8F684E9305641BA">2015</mods:date>
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<mods:number id="1767C704889C773B3F46361D6E77BC80">2015-09-30</mods:number>
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<mods:detail id="AA89BDA650F32951C76F90FF4637AB66" type="volume">
<mods:number id="7516181DCCADB5FC327F67945834080C">69</mods:number>
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<mods:number id="F732AE365246B3BE7CED52FD34923CCF">3</mods:number>
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<mods:url id="975D3A5C6CF9E03BAC3085535BB25F35">http://dx.doi.org/10.1649/0010-065x-69.3.463</mods:url>
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<mods:classification id="B1757C10A56284C54C00A69D2DA174A5">journal article</mods:classification>
<mods:identifier id="81AAE53710D08963D55C68646C4F32D1" type="DOI">10.1649/0010-065x-69.3.463</mods:identifier>
<mods:identifier id="285692DD1EDA5BAE8F05F9F531D02C56" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="8F6F671E6B87C508700CD0E42D295A7E" type="Zenodo-Dep">10106050</mods:identifier>
<mods:identifier id="BD23C4ACA4B1474183890989B952E246" type="ZooBank">9A0330AD-EEF7-4A80-8D80-1B6116BD856F</mods:identifier>
</mods:mods>
<treatment id="03CFA3486B6CFFC8FD6AFAB459B6FBBE" LSID="urn:lsid:plazi:treatment:03CFA3486B6CFFC8FD6AFAB459B6FBBE" httpUri="http://treatment.plazi.org/id/03CFA3486B6CFFC8FD6AFAB459B6FBBE" lastPageId="11" lastPageNumber="474" pageId="10" pageNumber="473">
<subSubSection id="C37C41D56B6CFFC9FD6AFAB45E5AFA4A" pageId="10" pageNumber="473" type="nomenclature">
<paragraph id="8BD9125E6B6CFFC9FD6AFAB45E5AFA4A" blockId="10.[724,1176,1373,1448]" pageId="10" pageNumber="473">
<heading id="D091A5326B6CFFC9FD6AFAB45E5AFA4A" centered="true" fontSize="10" level="2" pageId="10" pageNumber="473" reason="2">
<emphasis id="B912CE4C6B6CFFC9FD6AFAB45E5AFA4A" bold="true" pageId="10" pageNumber="473">
<taxonomicName id="4C6669DD6B6CFFC9FD6AFAB459D5FA4A" authority="Filippini, Galante, and Mico" authorityName="Filippini, Galante, and Mico" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="strigodermoides" status="sp. nov.">
<emphasis id="B912CE4C6B6CFFC9FD6AFAB459A2FAB4" bold="true" box="[724,977,1373,1395]" italics="true" pageId="10" pageNumber="473">Anomala strigodermoides</emphasis>
Filippini, Galante, and Micó
</taxonomicName>
,
<taxonomicNameLabel id="A22173376B6CFFC9FC0DFA915E5AFA4A" box="[947,1065,1400,1421]" pageId="10" pageNumber="473" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6CFFC9FCFEFA7A5E58FA6F" box="[832,1067,1427,1448]" pageId="10" pageNumber="473" type="description">
<paragraph id="8BD9125E6B6CFFC9FCFEFA7A5E58FA6F" blockId="10.[724,1176,1373,1448]" box="[832,1067,1427,1448]" pageId="10" pageNumber="473">
(
<figureCitation id="135D0EDB6B6CFFC9FCF9FA7A59FDFA6F" box="[839,910,1427,1448]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="10" pageNumber="473">Figs. 7</figureCitation>
,
<figureCitation id="135D0EDB6B6CFFC9FC24FA7A59C7FA6F" box="[922,948,1427,1448]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="10" pageNumber="473">15</figureCitation>
,
<figureCitation id="135D0EDB6B6CFFC9FC7EFA7A59A9FA6F" box="[960,986,1427,1448]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="10" pageNumber="473">23</figureCitation>
,
<figureCitation id="135D0EDB6B6CFFC9FC58FA7A5E73FA6F" box="[998,1024,1427,1448]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="10" pageNumber="473">31</figureCitation>
,
<figureCitation id="135D0EDB6B6CFFC9FBB2FA7A5E56FA6F" box="[1036,1061,1427,1448]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="10" pageNumber="473">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6CFFC8FD65FA5459B3FE56" lastPageId="11" lastPageNumber="474" pageId="10" pageNumber="473" type="materials_examined">
<paragraph id="8BD9125E6B6CFFC9FD65FA545EDAF906" blockId="10.[707,1194,1469,1730]" pageId="10" pageNumber="473">
<emphasis id="B912CE4C6B6CFFC9FD65FA545901FA15" bold="true" box="[731,882,1469,1490]" pageId="10" pageNumber="473">Type Material.</emphasis>
<materialsCitation id="3B0E18036B6CFFC9FCC1FA54597EF990" collectingDate="2006-03-24" collectingDateMax="2006-03-26" collectingDateMin="2006-03-24" collectionCode="MNCR" collectorName="B. Gamboa &amp; M. Moraga &amp; J. A. Azofeifa. Tp. Luz." country="Costa Rica" elevation="850" location="Alajuela" pageId="10" pageNumber="473" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="holotype">
<typeStatus id="54DDACFC6B6CFFC9FCC1FA5459ADFA15" box="[895,990,1469,1490]" pageId="10" pageNumber="473" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D60D9D76B6CFFC9FC55FA54598EFA14" box="[1003,1021,1469,1491]" pageId="10" pageNumber="473" type="male">♂</specimenCount>
&quot;
<collectingCountry id="F37152CE6B6CFFC9FBB0FA545ED5FA15" box="[1038,1190,1469,1490]" name="Costa Rica" pageId="10" pageNumber="473">COSTA RICA</collectingCountry>
. Prov.
<collectingRegion id="49A2DCBC6B6CFFC9FCBCFA31592FFA2A" box="[770,860,1496,1517]" country="Costa Rica" name="Alajuela" pageId="10" pageNumber="473">Alajuela</collectingRegion>
. Bijagua. Albergue Heliconias.
<quantity id="4C9EBFBB6B6CFFC9FD7DFA1B595EF9C0" box="[707,813,1522,1543]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="10" pageNumber="473" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
<elevation id="004BF56D6B6CFFC9FD7DFA1B595EF9C0" box="[707,813,1522,1543]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="10" pageNumber="473" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900m</elevation>
</quantity>
.
<date id="FFD8349E6B6CFFC9FC84FA1B5989F9C0" box="[826,1018,1522,1544]" pageId="10" pageNumber="473" value="2006-03-24" valueMax="2006-03-26" valueMin="2006-03-24">
<collectingDate id="EF9CCD766B6CFFC9FC84FA1B5989F9C0" box="[826,1018,1522,1544]" pageId="10" pageNumber="473" value="2006-03-24" valueMax="2006-03-26" valueMin="2006-03-24">24–26 MAR 2006</collectingDate>
</date>
.
<collectorName id="269377886B6CFFC9FBB9FA1B5E0CF9C0" box="[1031,1151,1522,1543]" pageId="10" pageNumber="473">B. Gamboa</collectorName>
,
<collectorName id="269377886B6CFFC9FB33FA1B5962F9E5" pageId="10" pageNumber="473">M. Moraga</collectorName>
,
<collectorName id="269377886B6CFFC9FCA3F9E45E61F9E5" box="[797,1042,1549,1570]" pageId="10" pageNumber="473">J. A. Azofeifa. Tp. Luz.</collectorName>
L_N_299800_ 423800 #85743 / INB0004012910&quot; deposited at
<collectionCode id="ED778A9B6B6CFFC9FD7DF9AB597EF990" box="[707,781,1602,1623]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="10" pageNumber="473">MNCR</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B0E18036B6CFFC9FCA6F9AB5ED6F960" collectingDate="2006-04-24" collectorName="B. Gamboa &amp; M. Moraga &amp; E. Navarro. Tp. Luz." country="Costa Rica" elevation="950" location="Cartago" pageId="10" pageNumber="473" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="paratype">
<typeStatus id="54DDACFC6B6CFFC9FCA6F9AB5906F990" box="[792,885,1602,1623]" pageId="10" pageNumber="473" type="paratype">Paratypes</typeStatus>
(6):
<specimenCount id="9D60D9D76B6CFFC9FC17F9AB59B4F99F" box="[937,967,1602,1624]" pageId="10" pageNumber="473" type="female">1♀</specimenCount>
&quot;
<collectingCountry id="F37152CE6B6CFFC9FC69F9AB5E18F990" box="[983,1131,1602,1623]" name="Costa Rica" pageId="10" pageNumber="473">COSTA RICA</collectingCountry>
. Prov.
<collectingRegion id="49A2DCBC6B6CFFC9FD7DF9B45978F9B5" box="[707,779,1629,1650]" country="Costa Rica" name="Cartago" pageId="10" pageNumber="473">Cartago</collectingRegion>
. Reserva Biol. El Copal.
<quantity id="4C9EBFBB6B6CFFC9FC5AF9B45E25F9B5" box="[996,1110,1628,1650]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="10" pageNumber="473" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">
<elevation id="004BF56D6B6CFFC9FC5AF9B45E25F9B5" box="[996,1110,1628,1650]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="10" pageNumber="473" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900–1000 m</elevation>
</quantity>
.
<date id="FFD8349E6B6CFFC9FBE0F9B45886F94B" pageId="10" pageNumber="473" value="2006-04-24">
<collectingDate id="EF9CCD766B6CFFC9FBE0F9B45886F94B" pageId="10" pageNumber="473" value="2006-04-24">24 ABR 2006</collectingDate>
</date>
.
<collectorName id="269377886B6CFFC9FD43F99E5919F94B" box="[765,874,1655,1676]" pageId="10" pageNumber="473">B. Gamboa</collectorName>
,
<collectorName id="269377886B6CFFC9FCCDF99E59ADF94B" box="[883,990,1655,1676]" pageId="10" pageNumber="473">M. Moraga</collectorName>
,
<collectorName id="269377886B6CFFC9FC56F99E5ED9F94B" box="[1000,1194,1655,1676]" pageId="10" pageNumber="473">E. Navarro. Tp. Luz.</collectorName>
L_N_196916_563695 #86066 / INB0004012683&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B6CFFC9FD7DF9455EDAF906" box="[707,1193,1708,1730]" country="Costa Rica" location="Alajuela" pageId="10" pageNumber="473" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="paratype">
<specimenCount id="9D60D9D76B6CFFC9FD7DF9455893F905" box="[707,736,1708,1730]" pageId="10" pageNumber="473" type="female">1♀</specimenCount>
&quot;
<collectingCountry id="F37152CE6B6CFFC9FD55F9455906F906" box="[747,885,1708,1729]" name="Costa Rica" pageId="10" pageNumber="473">COSTA RICA</collectingCountry>
.Prov.
<collectingRegion id="49A2DCBC6B6CFFC9FC10F945598DF906" box="[942,1022,1708,1729]" country="Costa Rica" name="Alajuela" pageId="10" pageNumber="473">Alajuela</collectingRegion>
.Bijagua.Albergue
</materialsCitation>
</paragraph>
<paragraph id="8BD9125E6B6DFFC8FF29FF2B5BE3FD72" blockId="11.[151,639,194,1729]" pageId="11" pageNumber="474">
Heliconias.
<quantity id="4C9EBFBB6B6DFFC8FF41FF2B5B2CFF10" box="[255,351,194,215]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="11" pageNumber="474" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
800-
<specimenCount id="9D60D9D76B6DFFC8FE96FF2B5B2CFF10" box="[296,351,194,215]" pageId="11" pageNumber="474" type="generic">900m</specimenCount>
</quantity>
.
<date id="FFD8349E6B6DFFC8FED8FF2B5878FF10" box="[358,523,194,216]" pageId="11" pageNumber="474" value="2006-03-24" valueMax="2006-03-26" valueMin="2006-03-24">24–26 MAR 2006</date>
.B. Gamboa, M. Moraga, J. A. Azofeifa. Tp. Luz. L_N_299800_ 423800 #85743 / INB0004012911&quot;;
<materialsCitation id="3B0E18036B6DFFC8FDBFFF1E5B48FEB0" collectingDate="2006-03-24" collectingDateMax="2006-03-26" collectingDateMin="2006-03-24" collectorName="B. Gamboa &amp; J. A. Azofeifa. Tp. Luz." country="Costa Rica" elevation="850" location="Prov." municipality="Bijagua. Albergue Heliconias." pageId="11" pageNumber="474" specimenCount="901" specimenCount-female="1" stateProvince="Alajuela">
<specimenCount id="9D60D9D76B6DFFC8FDBFFF1E586CFECA" box="[513,543,247,269]" pageId="11" pageNumber="474" type="female">1♀</specimenCount>
&quot;
<collectingCountry id="F37152CE6B6DFFC8FD90FF1E5AA2FEE0" name="Costa Rica" pageId="11" pageNumber="474">COSTA RICA</collectingCountry>
.
<location id="8EB944856B6DFFC8FF64FEFB5B78FEE0" LSID="urn:lsid:plazi:treatment:03CFA3486B6CFFC8FD6AFAB459B6FBBE:8EB944856B6DFFC8FF64FEFB5B78FEE0" box="[218,267,274,295]" country="Costa Rica" municipality="Bijagua. Albergue Heliconias." name="Prov." pageId="11" pageNumber="474" stateProvince="Alajuela">Prov.</location>
<collectingRegion id="49A2DCBC6B6DFFC8FEAEFEFB5B13FEE0" box="[272,352,274,295]" country="Costa Rica" name="Alajuela" pageId="11" pageNumber="474">Alajuela</collectingRegion>
.
<collectingMunicipality id="6BBD88246B6DFFC8FED6FEFB580CFEE0" box="[360,639,274,295]" pageId="11" pageNumber="474">Bijagua. Albergue Heliconias.</collectingMunicipality>
<quantity id="4C9EBFBB6B6DFFC8FF29FEC55A87FE86" box="[151,244,300,321]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="11" pageNumber="474" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
<elevation id="004BF56D6B6DFFC8FF29FEC55A87FE86" box="[151,244,300,321]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="11" pageNumber="474" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
800-
<specimenCount id="9D60D9D76B6DFFC8FF01FEC55A87FE86" box="[191,244,300,321]" pageId="11" pageNumber="474" type="generic">900m</specimenCount>
</elevation>
</quantity>
.
<date id="FFD8349E6B6DFFC8FF42FEC55BD3FE86" box="[252,416,300,322]" pageId="11" pageNumber="474" value="2006-03-24" valueMax="2006-03-26" valueMin="2006-03-24">
<collectingDate id="EF9CCD766B6DFFC8FF42FEC55BD3FE86" box="[252,416,300,322]" pageId="11" pageNumber="474" value="2006-03-24" valueMax="2006-03-26" valueMin="2006-03-24">24–26 MAR 2006</collectingDate>
</date>
.
<collectorName id="269377886B6DFFC8FE19FEC5587EFE86" box="[423,525,300,321]" pageId="11" pageNumber="474">B. Gamboa</collectorName>
, M. Moraga,
<collectorName id="269377886B6DFFC8FF29FEAE5B15FE9B" box="[151,358,327,348]" pageId="11" pageNumber="474">J. A. Azofeifa. Tp. Luz.</collectorName>
L_N_299800_423800 #85743 / INB0004012912&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B6DFFC8FEFCFE8B5B20FE26" collectingDate="2008-05-05" collectionCode="CEUA" collectorName="Guatuso. P. N. Volcan Tenorio. Punto &amp; J. A. Azofeifa. Tp de Luz" country="Costa Rica" elevation="800" location="Prov." municipality="Rio Buenavista." pageId="11" pageNumber="474" specimenCode="CEUA00106207" specimenCount="801" specimenCount-female="1" stateProvince="Alajuela">
<specimenCount id="9D60D9D76B6DFFC8FEFCFE8B5B13FEB0" box="[322,352,353,375]" pageId="11" pageNumber="474" type="female">1♀</specimenCount>
&quot;
<collectingCountry id="F37152CE6B6DFFC8FED5FE8B5B80FEB0" box="[363,499,354,375]" name="Costa Rica" pageId="11" pageNumber="474">COSTA RICA</collectingCountry>
.
<location id="8EB944856B6DFFC8FE45FE8B5859FEB0" LSID="urn:lsid:plazi:treatment:03CFA3486B6CFFC8FD6AFAB459B6FBBE:8EB944856B6DFFC8FE45FE8B5859FEB0" box="[507,554,354,375]" country="Costa Rica" municipality="Rio Buenavista." name="Prov." pageId="11" pageNumber="474" stateProvince="Alajuela">Prov.</location>
<collectingRegion id="49A2DCBC6B6DFFC8FD93FE8B5808FEB0" box="[557,635,354,375]" country="Costa Rica" name="Alajuela" pageId="11" pageNumber="474">Alajuela</collectingRegion>
.
<collectorName id="269377886B6DFFC8FF29FE95587BFE56" box="[151,520,380,401]" pageId="11" pageNumber="474">Guatuso. P.N. Volcán Tenorio. Punto</collectorName>
1: Catarata
<collectingMunicipality id="6BBD88246B6DFFC8FF29FE7E5B58FE6B" box="[151,299,407,428]" pageId="11" pageNumber="474">Río Buenavista.</collectingMunicipality>
<specimenCount id="9D60D9D76B6DFFC8FE8EFE7E5B18FE6B" box="[304,363,407,428]" pageId="11" pageNumber="474" type="generic">
<quantity id="4C9EBFBB6B6DFFC8FE8EFE7E5B14FE6B" box="[304,359,407,428]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="474" unit="m" value="800.0">
<elevation id="004BF56D6B6DFFC8FE8EFE7E5B14FE6B" box="[304,359,407,428]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="474" unit="m" value="800.0">800m</elevation>
</quantity>
.
</specimenCount>
<date id="FFD8349E6B6DFFC8FECEFE7E5B9DFE6B" box="[368,494,407,428]" pageId="11" pageNumber="474" value="2008-05-05">
<collectingDate id="EF9CCD766B6DFFC8FECEFE7E5B9DFE6B" box="[368,494,407,428]" pageId="11" pageNumber="474" value="2008-05-05">5 MAY 2008</collectingDate>
</date>
.
<collectorName id="269377886B6DFFC8FE49FE7E5B7BFE01" pageId="11" pageNumber="474">J. A. Azofeifa. Tp de Luz</collectorName>
1. L_N_298380_427850 #93735 /
<specimenCode id="DBC0BA256B6DFFC8FF29FE255B38FE26" box="[151,331,460,481]" collectionCode="CEUA" pageId="11" pageNumber="474">CEUA00106207</specimenCode>
&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B6DFFC8FEDAFE255B27FDF7" collectingDate="2010-04-15" collectionCode="CEUA" collectorName="P. N. Volcan Tenorio. Catarata Rio Buenavista. &amp; L. M. Zumbado &amp; J. A. Azofeifa." country="Costa Rica" elevation="780" location="Alajuela" pageId="11" pageNumber="474" specimenCode="CEUA00106208" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela">
<specimenCount id="9D60D9D76B6DFFC8FEDAFE255BF0FE26" box="[356,387,459,481]" pageId="11" pageNumber="474" type="male">1♂</specimenCount>
&quot;
<collectorName id="269377886B6DFFC8FE27FE255BEBFE3C" pageId="11" pageNumber="474">P.N. Volcán Tenorio. Catarata Río Buenavista.</collectorName>
<collectingRegion id="49A2DCBC6B6DFFC8FE1CFE0F5B8BFE3C" box="[418,504,486,507]" country="Costa Rica" name="Alajuela" pageId="11" pageNumber="474">Alajuela</collectingRegion>
,
<collectingCountry id="F37152CE6B6DFFC8FDB9FE0F5809FE3C" box="[519,634,486,507]" name="Costa Rica" pageId="11" pageNumber="474">Costa Rica</collectingCountry>
.
<quantity id="4C9EBFBB6B6DFFC8FF29FDE85ABCFDD1" box="[151,207,513,534]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="11" pageNumber="474" unit="m" value="780.0">
<elevation id="004BF56D6B6DFFC8FF29FDE85ABCFDD1" box="[151,207,513,534]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="11" pageNumber="474" unit="m" value="780.0">780m</elevation>
</quantity>
<date id="FFD8349E6B6DFFC8FF6DFDE85B35FDD1" box="[211,326,513,534]" pageId="11" pageNumber="474" value="2010-04-15">
<collectingDate id="EF9CCD766B6DFFC8FF6DFDE85B35FDD1" box="[211,326,513,534]" pageId="11" pageNumber="474" value="2010-04-15">15/IV/2010</collectingDate>
</date>
.
<collectorName id="269377886B6DFFC8FEF0FDE85B9BFDD1" box="[334,488,513,534]" pageId="11" pageNumber="474">L. M. Zumbado</collectorName>
,
<collectorName id="269377886B6DFFC8FE4EFDE85800FDD1" box="[496,627,513,534]" pageId="11" pageNumber="474">J.A.Azofeifa.</collectorName>
/
<specimenCode id="DBC0BA256B6DFFC8FF29FDF25B3FFDF7" box="[151,332,539,560]" collectionCode="CEUA" pageId="11" pageNumber="474">CEUA00106208</specimenCode>
&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B6DFFC8FEDAFDF25A8EFD5C" collectingDate="2010-04-15" collectionCode="MNCR" collectorName="P. N. Volcan Tenorio. Catarata Rio Buenavista." country="Costa Rica" elevation="780" location="Alajuela" pageId="11" pageNumber="474" specimenCode="CEUA00106209" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="paratype">
<specimenCount id="9D60D9D76B6DFFC8FEDAFDF25BF0FDF6" box="[356,387,539,561]" pageId="11" pageNumber="474" type="male">1♂</specimenCount>
&quot;
<collectorName id="269377886B6DFFC8FE27FDF25BEBFD8C" pageId="11" pageNumber="474">P.N. Volcán Tenorio. Catarata Río Buenavista.</collectorName>
<collectingRegion id="49A2DCBC6B6DFFC8FE1CFDDF5B8AFD8C" box="[418,505,566,587]" country="Costa Rica" name="Alajuela" pageId="11" pageNumber="474">Alajuela</collectingRegion>
,
<collectingCountry id="F37152CE6B6DFFC8FDB9FDDF5808FD8C" box="[519,635,566,587]" name="Costa Rica" pageId="11" pageNumber="474">Costa Rica</collectingCountry>
.
<quantity id="4C9EBFBB6B6DFFC8FF29FDB85ABFFDA1" box="[151,204,593,614]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="11" pageNumber="474" unit="m" value="780.0">
<elevation id="004BF56D6B6DFFC8FF29FDB85ABFFDA1" box="[151,204,593,614]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="11" pageNumber="474" unit="m" value="780.0">780m</elevation>
</quantity>
<date id="FFD8349E6B6DFFC8FF6BFDB85B32FDA1" box="[213,321,593,614]" pageId="11" pageNumber="474" value="2010-04-15">
<collectingDate id="EF9CCD766B6DFFC8FF6BFDB85B32FDA1" box="[213,321,593,614]" pageId="11" pageNumber="474" value="2010-04-15">15/IV/2010</collectingDate>
</date>
. L. M. Zumbado, J.A.Azofeifa. /
<specimenCode id="DBC0BA256B6DFFC8FF29FD825B33FD47" box="[151,320,619,640]" collectionCode="CEUA" pageId="11" pageNumber="474">CEUA00106209</specimenCode>
&quot;. Three
<typeStatus id="54DDACFC6B6DFFC8FE2BFD825B82FD47" box="[405,497,619,640]" pageId="11" pageNumber="474" type="paratype">paratypes</typeStatus>
are deposited at
<collectionCode id="ED778A9B6B6DFFC8FF0DFD6F5A8EFD5C" box="[179,253,646,667]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="11" pageNumber="474">MNCR</collectionCode>
</materialsCitation>
, and
<materialsCitation id="3B0E18036B6DFFC8FE87FD6F5BFFFD72" country="Costa Rica" location="Universidad de Alicante" pageId="11" pageNumber="474" specimenCount="1" stateProvince="Alajuela" typeStatus="paratype">
<specimenCount id="9D60D9D76B6DFFC8FE87FD6F5BA2FD5C" box="[313,465,646,667]" pageId="11" pageNumber="474" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
are deposited at
<location id="8EB944856B6DFFC8FF29FD495BFFFD72" LSID="urn:lsid:plazi:treatment:03CFA3486B6CFFC8FD6AFAB459B6FBBE:8EB944856B6DFFC8FF29FD495BFFFD72" box="[151,396,672,693]" country="Costa Rica" name="Universidad de Alicante" pageId="11" pageNumber="474" stateProvince="Alajuela">Universidad de Alicante</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD9125E6B6DFFC8FF11FD52590EFE86" blockId="11.[151,639,194,1729]" lastBlockId="11.[685,1172,194,1145]" pageId="11" pageNumber="474">
<emphasis id="B912CE4C6B6DFFC8FF11FD525B60FD17" bold="true" box="[175,275,699,720]" pageId="11" pageNumber="474">
<typeStatus id="54DDACFC6B6DFFC8FF11FD525B7DFD17" box="[175,270,699,720]" pageId="11" pageNumber="474" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="135D0EDB6B6DFFC8FEE3FD525BE5FD17" box="[349,406,699,720]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="11" pageNumber="474">Fig. 7</figureCitation>
). Body elongate. Length
<quantity id="4C9EBFBB6B6DFFC8FF29FD3C5A80FD2D" box="[151,243,725,746]" metricMagnitude="-3" metricUnit="m" metricValue="9.04" pageId="11" pageNumber="474" unit="mm" value="9.04">9.04 mm</quantity>
, width
<quantity id="4C9EBFBB6B6DFFC8FEFCFD3C5BEDFD2D" box="[322,414,725,746]" metricMagnitude="-3" metricUnit="m" metricValue="4.89" pageId="11" pageNumber="474" unit="mm" value="4.89">4.89 mm</quantity>
. Head, pronotum, and scutellum metallic green; elytra ochre with 3 pairs of brown maculae along basal margin and a brown, wavy, median, transverse band. Rows of punctures lightly pigmented. Entire surface with green luster. Pygidium, legs, and venter brown with green luster.
<emphasis id="B912CE4C6B6DFFC8FF67FC9C5B66FC4D" bold="true" box="[217,277,885,906]" pageId="11" pageNumber="474">Head:</emphasis>
Clypeus trapezoidal, surface densely punctate-reticulate, flat. Anterior margin straight, ratio width/length 1.9. Frons densely punctate, flat. Ocular canthus long and thin, with acute apex. Ratio of interocular width/width of eye 2.76. Ratio of antennal funiculus/club 0.64.
<emphasis id="B912CE4C6B6DFFC8FE73FC13584AFBC8" bold="true" box="[461,569,1018,1039]" pageId="11" pageNumber="474">Pronotum:</emphasis>
Trapezoidal, long, narrower than elytra. Width 1.26 times length. Lateral margins angulated at 1/3 of pronotal length from anterior margins. Anterior angles acute, posterior angles obtuse. Basal margin thin, sinuate, complete. Surface irregular, with coalescing punctures.
<emphasis id="B912CE4C6B6DFFC8FE91FB705BEEFB69" bold="true" box="[303,413,1177,1198]" pageId="11" pageNumber="474">Scutellum:</emphasis>
Subtriangular, surface sparsely punctate. Ratio width/length 1.24.
<emphasis id="B912CE4C6B6DFFC8FD89FB5D580EFB0E" bold="true" box="[567,637,1204,1225]" pageId="11" pageNumber="474">Elytra:</emphasis>
With striae defined by regular rows of punctures. Surface covered with secondary punctures. Marginal membrane complete, very thin.
<emphasis id="B912CE4C6B6DFFC8FDAAFAEA580EFADF" bold="true" box="[532,637,1283,1304]" pageId="11" pageNumber="474">Pygidium:</emphasis>
Finely granulated, with short setae on disc and long setae at hind margin. Ratio width/length 1.87.
<emphasis id="B912CE4C6B6DFFC8FF6CFABA5B68FAAF" bold="true" box="[210,283,1363,1384]" pageId="11" pageNumber="474">Venter:</emphasis>
Space between mesocoxae narrow, slightly convex. Mesometasternal suture welldefined at base of mesocoxae. Abdominal sternites with 2–3 rows of setae and 3–5 rows of punctures per sternite. Last sternite strigate, with apical margin thin and curved, protruding membrane thin.
<emphasis id="B912CE4C6B6DFFC8FF6EFA1B5B79F9C0" bold="true" box="[208,266,1522,1543]" pageId="11" pageNumber="474">Legs:</emphasis>
Protibia bidentate (
<figureCitation id="135D0EDB6B6DFFC8FE62FA1B5859F9C0" box="[476,554,1522,1543]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="11" pageNumber="474">Fig. 15</figureCitation>
); apical tooth long and curved, basal tooth above internal apex of protibia, triangular, acute. Metatibia slen- der, fusiform. Ratio length/width 2.72. First external carina well-developed. Surface punctate above second external carina and rugose below. External protarsal claw curved; internal claw bifurcate, with upper branch of same length and 2/3 width of the lower one; inferior margin with a sharp angle near base.
<emphasis id="B912CE4C6B6DFFC8FD5AFF34590AFF35" bold="true" box="[740,889,221,242]" pageId="11" pageNumber="474">Male genitalia:</emphasis>
Parameres with squared apex in lateral view (
<figureCitation id="135D0EDB6B6DFFC8FCEDFF1E59D3FECB" box="[851,928,247,268]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="11" pageNumber="474">Fig. 23</figureCitation>
). Endophallus (
<figureCitation id="135D0EDB6B6DFFC8FBFFFF1E5EFFFECB" box="[1089,1164,247,268]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="11" pageNumber="474">Fig. 31</figureCitation>
) with a wide sacculus, ejaculatory duct opening at basal lateral inflation.
</paragraph>
<paragraph id="8BD9125E6B6DFFC8FD7BFEAE59B3FE56" blockId="11.[685,1172,194,1145]" pageId="11" pageNumber="474">
<emphasis id="B912CE4C6B6DFFC8FD7BFEAE5965FE9B" bold="true" box="[709,790,327,348]" pageId="11" pageNumber="474">Female.</emphasis>
Similar to male except protibia with wider apical tooth; internal protarsal claw narrower with curved inferior margin.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6DFFC8FD7BFE7E5E54FD72" pageId="11" pageNumber="474" type="description">
<paragraph id="8BD9125E6B6DFFC8FD7BFE7E5E54FD72" blockId="11.[685,1172,194,1145]" pageId="11" pageNumber="474">
<emphasis id="B912CE4C6B6DFFC8FD7BFE7E5959FE6B" bold="true" box="[709,810,407,428]" pageId="11" pageNumber="474">Variation.</emphasis>
Six
<typeStatus id="54DDACFC6B6DFFC8FCE2FE7E59C9FE6B" box="[860,954,407,428]" pageId="11" pageNumber="474" type="paratype">paratypes</typeStatus>
. Elytra with or without green luster; maculae on elytra vary from reddish to dark brown, may also be present on apex; venter varies from reddish to brown. Body length
<quantity id="4C9EBFBB6B6DFFC8FD13FDE8594AFDD1" box="[685,825,512,534]" metricMagnitude="-3" metricUnit="m" metricValue="8.905" metricValueMax="9.64" metricValueMin="8.17" pageId="11" pageNumber="474" unit="mm" value="8.905" valueMax="9.64" valueMin="8.17">8.17–9.64 mm</quantity>
, body width
<quantity id="4C9EBFBB6B6DFFC8FC0BFDE85E32FDD1" box="[949,1089,512,534]" metricMagnitude="-3" metricUnit="m" metricValue="4.654999999999999" metricValueMax="4.89" metricValueMin="4.42" pageId="11" pageNumber="474" unit="mm" value="4.654999999999999" valueMax="4.89" valueMin="4.42">4.42–4.89 mm</quantity>
. Clypeal width/length ratio 1.89–2.08. Ratio of interocular width/width of eye 2.26–3.04. Ratio of antennal funiculus/club 0.60–0.74. Pronotal width/length ratio 1.26–1.32. Scutellar width/length ratio 1.14–1.33. Pygidial width/length ratio 1.87 (
<emphasis id="B912CE4C6B6DFFC8FBEFFD6C5E2EFD5C" box="[1105,1117,645,667]" italics="true" pageId="11" pageNumber="474">n</emphasis>
= 1). Metatibial width/length ratio 2.49–2.82.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6DFFC8FD7BFD52598FFCC2" pageId="11" pageNumber="474" type="etymology">
<paragraph id="8BD9125E6B6DFFC8FD7BFD52598FFCC2" blockId="11.[685,1172,194,1145]" pageId="11" pageNumber="474">
<emphasis id="B912CE4C6B6DFFC8FD7BFD52594BFD17" bold="true" box="[709,824,699,720]" pageId="11" pageNumber="474">Etymology.</emphasis>
From the name of genus
<taxonomicName id="4C6669DD6B6DFFC8FB9AFD535EE7FD17" box="[1060,1172,698,720]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="B912CE4C6B6DFFC8FB9AFD535EE7FD17" box="[1060,1172,698,720]" italics="true" pageId="11" pageNumber="474">Strigoderma</emphasis>
</taxonomicName>
and suffix -oides, for its general resemblance with species of
<taxonomicName id="4C6669DD6B6DFFC8FCADFD19598FFCC2" authority="Burmeister." authorityName="Burmeister." box="[787,1020,752,774]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="B912CE4C6B6DFFC8FCADFD1959FBFCC1" box="[787,904,752,774]" italics="true" pageId="11" pageNumber="474">Strigoderma</emphasis>
Burmeister.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6DFFC8FD7BFCE35974FCA8" pageId="11" pageNumber="474" type="distribution">
<paragraph id="8BD9125E6B6DFFC8FD7BFCE35974FCA8" blockId="11.[685,1172,194,1145]" pageId="11" pageNumber="474">
<emphasis id="B912CE4C6B6DFFC8FD7BFCE3593BFCD8" bold="true" box="[709,840,778,799]" pageId="11" pageNumber="474">Distribution.</emphasis>
This species is known from the
<collectingRegion id="49A2DCBC6B6DFFC8FD13FCCC5957FCFD" box="[685,804,805,826]" country="Costa Rica" name="Guanacaste" pageId="11" pageNumber="474">Guanacaste</collectingRegion>
and Talamanca mountain ranges of
<collectingCountry id="F37152CE6B6DFFC8FD73FCA95933FC92" box="[717,832,832,853]" name="Costa Rica" pageId="11" pageNumber="474">Costa Rica</collectingCountry>
, from
<quantity id="4C9EBFBB6B6DFFC8FC34FCA95E5DFC92" box="[906,1070,832,853]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.5" metricValueMin="7.5" pageId="11" pageNumber="474" unit="m" value="900.0" valueMax="1050.0" valueMin="750.0">750 to 1,050 m</quantity>
elevation (
<figureCitation id="135D0EDB6B6DFFC8FD0AFCB35888FCA8" box="[692,763,858,879]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="11" pageNumber="474">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6DFFC8FD7BFC9C59B6FBBE" pageId="11" pageNumber="474" type="diagnosis">
<paragraph id="8BD9125E6B6DFFC8FD7BFC9C59B6FBBE" blockId="11.[685,1172,194,1145]" pageId="11" pageNumber="474">
<emphasis id="B912CE4C6B6DFFC8FD7BFC9C595DFC4D" bold="true" box="[709,814,885,906]" pageId="11" pageNumber="474">Diagnosis.</emphasis>
The long and narrow pronotum with a coarse surface readily separates this species from other Neotropical
<taxonomicName id="4C6669DD6B6DFFC8FCDEFC4059C5FC78" box="[864,950,937,959]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="B912CE4C6B6DFFC8FCDEFC4059C5FC78" box="[864,950,937,959]" italics="true" pageId="11" pageNumber="474">Anomala</emphasis>
</taxonomicName>
. Another species with a relatively narrow pronotum is
<taxonomicName id="4C6669DD6B6DFFC8FC56FC2D59E0FC33" authority="Ohaus, 1908" authorityName="Ohaus" authorityYear="1908" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="474" phylum="Arthropoda" rank="species" species="repressa">
<emphasis id="B912CE4C6B6DFFC8FC56FC2D5EE0FC1D" box="[1000,1171,964,986]" italics="true" pageId="11" pageNumber="474">Anomala repressa</emphasis>
Ohaus, 1908 (
<collectingCountry id="F37152CE6B6DFFC8FC82FC3659F8FC33" box="[828,907,991,1012]" name="Mexico" pageId="11" pageNumber="474">Mexico</collectingCountry>
)
</taxonomicName>
, which is of similar size. However, in
<taxonomicName id="4C6669DD6B6DFFC8FC8CFC1059D1FBC8" authorityName="Ohaus" authorityYear="1908" box="[818,930,1017,1039]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="474" phylum="Arthropoda" rank="species" species="repressa">
<emphasis id="B912CE4C6B6DFFC8FC8CFC1059D1FBC8" box="[818,930,1017,1039]" italics="true" pageId="11" pageNumber="474">A. repressa</emphasis>
</taxonomicName>
, the pronotum is wider than long, with a sinuate margin, the entire surface is covered with setae, and the elytra have a peculiar pattern with two light-colored maculae on a black to reddish basal color.
</paragraph>
</subSubSection>
</treatment>
</document>
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