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<document id="EDCDB624447418EE5E9D8A9BF0303A0E" ID-DOI="10.1649/0010-065x-69.3.463" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106050" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175225603" checkinUser="diego" docAuthor="Filippini, V., Galante, E. &amp; Micó, E." docDate="2015" docId="03CFA3486B67FFC1FF1EFBBA5EE0F9C0" docLanguage="en" docName="0010-065X-69.3.463.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-69.3.463" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Anomala coffea Filippini, Galante, and Mico, new species" docType="treatment" docVersion="3" lastPageNumber="465" masterDocId="FFF6DB306B66FFC3FFBEFFE95A73FFC7" masterDocTitle="Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica" masterLastPageNumber="476" masterPageNumber="463" pageNumber="464" updateTime="1699636354092" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="4ED8934A51F468F7E8851F3A59543241" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5505C69AE7551BACA0AC9779A1379F31">
<mods:title id="72973DF13E1E2DBABCD9CBDF2CEB8216">Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica</mods:title>
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<mods:namePart id="637412F87E2953226290A6170B583202">Filippini, V.</mods:namePart>
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<mods:namePart id="40FC13D78F017D94A767206E16C14775">Galante, E.</mods:namePart>
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<mods:roleTerm id="7EC5560A8321554079A9B19576BE830D">Author</mods:roleTerm>
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<mods:namePart id="AE438C86B042260CFFE4A6DF210BD958">Micó, E.</mods:namePart>
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<mods:title id="B30A4505A2EF75A08996515A0D069B7E">The Coleopterists Bulletin</mods:title>
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<mods:date id="53E964CDA2830DB01F6AA44E89C5F86B">2015</mods:date>
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<mods:number id="937DAAF9BC0859A75CF1CCFD3822CB68">2015-09-30</mods:number>
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<mods:number id="C03D5A060FF20A8B08988DD086949602">69</mods:number>
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<mods:number id="B0FBD040A817F4E378346A51A5204A15">3</mods:number>
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<mods:classification id="76C05A78BCCC708723CC6CFD70C2A47C">journal article</mods:classification>
<mods:identifier id="D51A19BE86A197B44CCA8D9BE4993110" type="DOI">10.1649/0010-065x-69.3.463</mods:identifier>
<mods:identifier id="3E0BA00DCD6E1922ACADD595D185BED7" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="1D91A128434E16A1AC13607E755450D2" type="Zenodo-Dep">10106050</mods:identifier>
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<treatment id="03CFA3486B67FFC1FF1EFBBA5EE0F9C0" LSID="urn:lsid:plazi:treatment:03CFA3486B67FFC1FF1EFBBA5EE0F9C0" httpUri="http://treatment.plazi.org/id/03CFA3486B67FFC1FF1EFBBA5EE0F9C0" lastPageId="2" lastPageNumber="465" pageId="1" pageNumber="464">
<subSubSection id="C37C41D56B67FFC2FF1EFBBA5BB6FB44" pageId="1" pageNumber="464" type="nomenclature">
<paragraph id="8BD9125E6B67FFC2FF1EFBBA5BB6FB44" blockId="1.[160,629,1107,1182]" pageId="1" pageNumber="464">
<heading id="D091A5326B67FFC2FF1EFBBA5BB6FB44" centered="true" fontSize="10" level="2" pageId="1" pageNumber="464" reason="2">
<emphasis id="B912CE4C6B67FFC2FF1EFBBA5BB6FB44" bold="true" pageId="1" pageNumber="464">
<taxonomicName id="4C6669DD6B67FFC2FF1EFBBA5803FBAE" authority="Filippini, Galante, and Mico" authorityName="Filippini, Galante, and Mico" box="[160,624,1107,1129]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="species" species="coffea" status="sp. nov.">
<emphasis id="B912CE4C6B67FFC2FF1EFBBA5B4DFBAE" bold="true" box="[160,318,1107,1129]" italics="true" pageId="1" pageNumber="464">Anomala coffea</emphasis>
Filippini, Galante, and Micó
</taxonomicName>
,
<taxonomicNameLabel id="A22173376B67FFC2FEF1FB875BB6FB44" box="[335,453,1134,1155]" pageId="1" pageNumber="464" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B67FFC2FEA5FB605B89FB59" box="[283,506,1161,1182]" pageId="1" pageNumber="464" type="description">
<paragraph id="8BD9125E6B67FFC2FEA5FB605B89FB59" blockId="1.[160,629,1107,1182]" box="[283,506,1161,1182]" pageId="1" pageNumber="464">
(
<figureCitation id="135D0EDB6B67FFC2FE9CFB605B14FB59" box="[290,359,1161,1182]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="1" pageNumber="464">Figs. 1</figureCitation>
,
<figureCitation id="135D0EDB6B67FFC2FECBFB605BF0FB59" box="[373,387,1161,1182]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="1" pageNumber="464">9</figureCitation>
,
<figureCitation id="135D0EDB6B67FFC2FE31FB605BDAFB59" box="[399,425,1161,1182]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="1" pageNumber="464">17</figureCitation>
,
<figureCitation id="135D0EDB6B67FFC2FE0AFB605BBDFB59" box="[436,462,1161,1182]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="1" pageNumber="464">25</figureCitation>
,
<figureCitation id="135D0EDB6B67FFC2FE64FB605B80FB59" box="[474,499,1161,1182]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="1" pageNumber="464">34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B67FFC2FF11FB5D5E0DF9FB" pageId="1" pageNumber="464" type="materials_examined">
<paragraph id="8BD9125E6B67FFC2FF11FB5D580DF906" blockId="1.[151,639,1203,1729]" pageId="1" pageNumber="464">
<emphasis id="B912CE4C6B67FFC2FF11FB5D5B32FB0E" bold="true" box="[175,321,1204,1225]" pageId="1" pageNumber="464">Type Material.</emphasis>
<materialsCitation id="3B0E18036B67FFC2FEF0FB5D5B1AFAF4" collectingDate="1991-08-08" collectingDateMax="1991-08-24" collectingDateMin="1991-08-08" collectionCode="MNCR" collectorName="C. Moraga" country="Costa Rica" elevation="700" location="Sta. Cecilia" pageId="1" pageNumber="464" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54DDACFC6B67FFC2FEF0FB5D5BDBFB0E" box="[334,424,1204,1225]" pageId="1" pageNumber="464" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D60D9D76B67FFC2FE0FFB5A5BB0FB0E" box="[433,451,1203,1225]" pageId="1" pageNumber="464" type="male">♂</specimenCount>
&quot;Est. Pitilla,
<quantity id="4C9EBFBB6B67FFC2FD83FB5D5809FB0E" box="[573,634,1204,1225]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">
<elevation id="004BF56D6B67FFC2FD83FB5D5809FB0E" box="[573,634,1204,1225]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<quantity id="4C9EBFBB6B67FFC2FF29FB275ABCFB24" box="[151,207,1230,1251]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="464" unit="km" value="9.0">9 km</quantity>
S
<location id="8EB944856B67FFC2FF4BFB275B01FB24" LSID="urn:lsid:plazi:treatment:03CFA3486B67FFC1FF1EFBBA5EE0F9C0:8EB944856B67FFC2FF4BFB275B01FB24" box="[245,370,1230,1251]" country="Costa Rica" name="Sta. Cecilia" pageId="1" pageNumber="464" stateProvince="Guanacaste">Sta. Cecilia</location>
, P. N.
<collectingRegion id="49A2DCBC6B67FFC2FE7DFB27584AFB24" box="[451,569,1230,1251]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
, Prov.
<collectingRegion id="49A2DCBC6B67FFC2FF29FB005B7DFB39" box="[151,270,1257,1278]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
,
<collectingCountry id="F37152CE6B67FFC2FEA5FB005BFEFB39" box="[283,397,1257,1278]" name="Costa Rica" pageId="1" pageNumber="464">Costa Rica</collectingCountry>
,
<collectorName id="269377886B67FFC2FE24FB005874FB39" box="[410,519,1257,1278]" pageId="1" pageNumber="464">C. Moraga</collectorName>
<date id="FFD8349E6B67FFC2FDAEFB005ABBFADF" pageId="1" pageNumber="464" value="1991-08-08" valueMax="1991-08-24" valueMin="1991-08-08">
<collectingDate id="EF9CCD766B67FFC2FDAEFB005ABBFADF" pageId="1" pageNumber="464" value="1991-08-08" valueMax="1991-08-24" valueMin="1991-08-08">8 a 24 ago 1991</collectingDate>
</date>
, L-N 330200,380200 / INBIOCRI000409517&quot; deposited at
<collectionCode id="ED778A9B6B67FFC2FEA3FAF75B1AFAF4" box="[285,361,1310,1331]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="1" pageNumber="464">MNCR</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B0E18036B67FFC2FECBFAF75809FA44" collectingDate="1988-07" collectorName="Km S. Santa Cecilia" country="Costa Rica" elevation="700" latitude="10.990556" location="Estac. Pitilla" longLatPrecision="21" longitude="-85.42777" pageId="1" pageNumber="464" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54DDACFC6B67FFC2FECBFAF75BA4FAF4" box="[373,471,1310,1331]" pageId="1" pageNumber="464" type="paratype">Paratypes</typeStatus>
(6):
<specimenCount id="9D60D9D76B67FFC2FDB0FAF7585EFAF4" box="[526,557,1309,1331]" pageId="1" pageNumber="464" type="female">1♀</specimenCount>
&quot;
<location id="8EB944856B67FFC2FD81FAF75AA5FA8A" LSID="urn:lsid:plazi:treatment:03CFA3486B67FFC1FF1EFBBA5EE0F9C0:8EB944856B67FFC2FD81FAF75AA5FA8A" country="Costa Rica" latitude="10.990556" longLatPrecision="21" longitude="-85.42777" name="Estac. Pitilla" pageId="1" pageNumber="464">Estac. Pitilla</location>
,
<quantity id="4C9EBFBB6B67FFC2FF5AFAD15B54FA8A" box="[228,295,1336,1357]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">
<elevation id="004BF56D6B67FFC2FF5AFAD15B54FA8A" box="[228,295,1336,1357]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">700 m</elevation>
</quantity>
, 9
<collectorName id="269377886B67FFC2FEF7FAD1586DFA8A" box="[329,542,1336,1357]" pageId="1" pageNumber="464">Km S. Santa Cecilia</collectorName>
, Guanac. Pr.
<collectingCountry id="F37152CE6B67FFC2FF03FABA5B20FAAF" box="[189,339,1363,1384]" name="Costa Rica" pageId="1" pageNumber="464">COSTA RICA</collectingCountry>
<date id="FFD8349E6B67FFC2FEE3FABA5BC4FAAF" box="[349,439,1363,1384]" pageId="1" pageNumber="464" value="1988-07">
<collectingDate id="EF9CCD766B67FFC2FEE3FABA5BC4FAAF" box="[349,439,1363,1384]" pageId="1" pageNumber="464" value="1988-07">Jul 1988</collectingDate>
</date>
GNP Biodiversity Survey
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,
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/ INB0003315377&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B67FFC2FF29FA615809FA15" country="Costa Rica" elevation="700" location="Est. Pitilla" pageId="1" pageNumber="464" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D60D9D76B67FFC2FF29FA615AC6FA59" box="[151,181,1416,1438]" pageId="1" pageNumber="464" type="male">1♂</specimenCount>
&quot;
<location id="8EB944856B67FFC2FF78FA615B58FA5A" LSID="urn:lsid:plazi:treatment:03CFA3486B67FFC1FF1EFBBA5EE0F9C0:8EB944856B67FFC2FF78FA615B58FA5A" box="[198,299,1416,1437]" country="Costa Rica" name="Est. Pitilla" pageId="1" pageNumber="464" stateProvince="Guanacaste">Est. Pitilla</location>
,
<quantity id="4C9EBFBB6B67FFC2FE89FA615B0BFA5A" box="[311,376,1416,1437]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">
<elevation id="004BF56D6B67FFC2FE89FA615B0BFA5A" box="[311,376,1416,1437]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<locationDeviation id="9A6B0B7C6B67FFC2FE3DFA615831FA5A" box="[387,578,1416,1437]" location="Sta. Cecilia" pageId="1" pageNumber="464">
<quantity id="4C9EBFBB6B67FFC2FE3DFA615BC6FA5A" box="[387,437,1416,1437]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="464" unit="km" value="9.0">9 km</quantity>
S Sta. Cecilia
</locationDeviation>
, P. N.
<collectingRegion id="49A2DCBC6B67FFC2FF29FA4A5B71FA7F" box="[151,258,1443,1464]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
, Prov. Guan.
<collectingCountry id="F37152CE6B67FFC2FEC0FA4A5874FA7F" box="[382,519,1443,1464]" name="Costa Rica" pageId="1" pageNumber="464">COSTA RICA</collectingCountry>
, Tp Malaise, 1990. L N 330200,380200 / INBIOCRI000304012&quot;
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;
<materialsCitation id="3B0E18036B67FFC2FF29FA315B02F9FB" collectingDate="1992-05-02" collectingDateMax="1992-05-15" collectingDateMin="1992-05-02" collectorName="C. Moraga" country="Costa Rica" elevation="700" location="Est. Pitilla" municipality="Prov." pageId="1" pageNumber="464" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D60D9D76B67FFC2FF29FA315AC6FA2A" box="[151,181,1495,1517]" pageId="1" pageNumber="464" type="male">1♂</specimenCount>
&quot;
<location id="8EB944856B67FFC2FF78FA315B58FA2A" LSID="urn:lsid:plazi:treatment:03CFA3486B67FFC1FF1EFBBA5EE0F9C0:8EB944856B67FFC2FF78FA315B58FA2A" box="[198,299,1496,1517]" country="Costa Rica" municipality="Prov." name="Est. Pitilla" pageId="1" pageNumber="464" stateProvince="Guanacaste">Est. Pitilla</location>
,
<quantity id="4C9EBFBB6B67FFC2FE89FA315B0BFA2A" box="[311,376,1496,1517]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">
<elevation id="004BF56D6B67FFC2FE89FA315B0BFA2A" box="[311,376,1496,1517]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<locationDeviation id="9A6B0B7C6B67FFC2FE3DFA315831FA2A" box="[387,578,1496,1517]" location="Sta. Cecilia" pageId="1" pageNumber="464">
<quantity id="4C9EBFBB6B67FFC2FE3DFA315BC6FA2A" box="[387,437,1496,1517]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="464" unit="km" value="9.0">9 km</quantity>
S Sta. Cecilia
</locationDeviation>
, P. N.
<collectingRegion id="49A2DCBC6B67FFC2FF29FA1B5B66F9C0" box="[151,277,1522,1543]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
,
<collectingMunicipality id="6BBD88246B67FFC2FE96FA1B5B13F9C0" box="[296,352,1522,1543]" pageId="1" pageNumber="464">Prov.</collectingMunicipality>
<collectingRegion id="49A2DCBC6B67FFC2FED3FA1B5B98F9C0" box="[365,491,1522,1543]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
,
<collectingCountry id="F37152CE6B67FFC2FE43FA1B580AF9C0" box="[509,633,1522,1543]" name="Costa Rica" pageId="1" pageNumber="464">Costa Rica</collectingCountry>
,
<collectorName id="269377886B67FFC2FF29F9E45A8DF9E5" box="[151,254,1549,1570]" pageId="1" pageNumber="464">C. Moraga</collectorName>
,
<date id="FFD8349E6B67FFC2FEB6F9E45BDBF9E5" box="[264,424,1549,1570]" pageId="1" pageNumber="464" value="1992-05-02" valueMax="1992-05-15" valueMin="1992-05-02">
<collectingDate id="EF9CCD766B67FFC2FEB6F9E45BDBF9E5" box="[264,424,1549,1570]" pageId="1" pageNumber="464" value="1992-05-02" valueMax="1992-05-15" valueMin="1992-05-02">2 a 15 may 1992</collectingDate>
</date>
, L N 330200,380200 / INBIOCRI000405396&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B67FFC2FEC7F9CE5853F94B" collectingDate="1992-03-31" collectingDateMax="1992-04-29" collectingDateMin="1992-03-31" collectorName="K. Taylor" country="Costa Rica" elevation="700" location="Est. Pitilla" municipality="Prov." pageId="1" pageNumber="464" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D60D9D76B67FFC2FEC7F9CE5BE4F9FA" box="[377,407,1575,1597]" pageId="1" pageNumber="464" type="male">1♂</specimenCount>
&quot;
<location id="8EB944856B67FFC2FE1DF9CE5877F9FB" LSID="urn:lsid:plazi:treatment:03CFA3486B67FFC1FF1EFBBA5EE0F9C0:8EB944856B67FFC2FE1DF9CE5877F9FB" box="[419,516,1575,1596]" country="Costa Rica" municipality="Prov." name="Est. Pitilla" pageId="1" pageNumber="464" stateProvince="Guanacaste">Est. Pitilla</location>
,
<quantity id="4C9EBFBB6B67FFC2FDB5F9CE5834F9FB" box="[523,583,1575,1596]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">
<elevation id="004BF56D6B67FFC2FDB5F9CE5834F9FB" box="[523,583,1575,1596]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<locationDeviation id="9A6B0B7C6B67FFC2FDF1F9CE5B68F990" location="Sta. Cecilia" pageId="1" pageNumber="464">
<quantity id="4C9EBFBB6B67FFC2FDF1F9CE580DF9FB" box="[591,638,1575,1596]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="464" unit="km" value="9.0">9 km</quantity>
S Sta. Cecilia
</locationDeviation>
, P. N.
<collectingRegion id="49A2DCBC6B67FFC2FEE3F9AB5BB9F990" box="[349,458,1602,1623]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
,
<collectingMunicipality id="6BBD88246B67FFC2FE6BF9AB5874F990" box="[469,519,1602,1623]" pageId="1" pageNumber="464">Prov.</collectingMunicipality>
<collectingRegion id="49A2DCBC6B67FFC2FDB3F9AB5809F990" box="[525,634,1602,1623]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
,
<collectingCountry id="F37152CE6B67FFC2FF29F9B45B72F9B5" box="[151,257,1629,1650]" name="Costa Rica" pageId="1" pageNumber="464">Costa Rica</collectingCountry>
,
<collectorName id="269377886B67FFC2FEB2F9B45B1BF9B5" box="[268,360,1629,1650]" pageId="1" pageNumber="464">K. Taylor</collectorName>
,
<date id="FFD8349E6B67FFC2FECDF9B45830F9B5" box="[371,579,1629,1650]" pageId="1" pageNumber="464" value="1992-03-31" valueMax="1992-04-29" valueMin="1992-03-31">
<collectingDate id="EF9CCD766B67FFC2FECDF9B45830F9B5" box="[371,579,1629,1650]" pageId="1" pageNumber="464" value="1992-03-31" valueMax="1992-04-29" valueMin="1992-03-31">31 mar - 29 abr 1992</collectingDate>
</date>
, L- N 330200,380200 / INBIOCRI000523773&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B67FFC2FD95F99E580DF906" collectorName="Prov. Guan. &amp; C. Moraga &amp; Ene" country="Costa Rica" elevation="700" location="Est. Pitilla" pageId="1" pageNumber="464" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D60D9D76B67FFC2FD95F99E583AF94A" box="[555,585,1655,1677]" pageId="1" pageNumber="464" type="female">1♀</specimenCount>
&quot;
<location id="8EB944856B67FFC2FDE7F99E5ABDF960" LSID="urn:lsid:plazi:treatment:03CFA3486B67FFC1FF1EFBBA5EE0F9C0:8EB944856B67FFC2FDE7F99E5ABDF960" country="Costa Rica" name="Est. Pitilla" pageId="1" pageNumber="464" stateProvince="Guanacaste">Est. Pitilla</location>
,
<quantity id="4C9EBFBB6B67FFC2FF66F97B5B66F960" box="[216,277,1682,1703]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">
<elevation id="004BF56D6B67FFC2FF66F97B5B66F960" box="[216,277,1682,1703]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<locationDeviation id="9A6B0B7C6B67FFC2FEA1F97B5BA0F960" box="[287,467,1682,1703]" location="Sta. Cecilia" pageId="1" pageNumber="464">
<quantity id="4C9EBFBB6B67FFC2FEA1F97B5B3CF960" box="[287,335,1682,1703]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="464" unit="km" value="9.0">9 km</quantity>
S Sta. Cecilia
</locationDeviation>
, P. N.
<collectingRegion id="49A2DCBC6B67FFC2FDB1F97B5809F960" box="[527,634,1682,1703]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
,
<collectorName id="269377886B67FFC2FF29F9455B7FF906" box="[151,268,1708,1729]" pageId="1" pageNumber="464">Prov. Guan.</collectorName>
<collectingCountry id="F37152CE6B67FFC2FEADF9455BD0F906" box="[275,419,1708,1729]" name="Costa Rica" pageId="1" pageNumber="464">COSTA RICA</collectingCountry>
,
<collectorName id="269377886B67FFC2FE11F945586AF906" box="[431,537,1708,1729]" pageId="1" pageNumber="464">C. Moraga</collectorName>
, 6-28
<collectorName id="269377886B67FFC2FDE7F945580DF906" box="[601,638,1708,1729]" pageId="1" pageNumber="464">Ene</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BD9125E6B67FFC2FD13FF2B59EBFEB1" blockId="1.[685,1173,194,1730]" pageId="1" pageNumber="464">
<materialsCitation id="3B0E18036B67FFC2FD13FF2B5EFBFF10" box="[685,1160,194,215]" collectionCode="CEUA" pageId="1" pageNumber="464" specimenCode="CEUA00106195" specimenCount="1">
1992, L- N 330200,380200 /
<specimenCode id="DBC0BA256B67FFC2FC63FF2B5EFBFF10" box="[989,1160,194,215]" collectionCode="CEUA" pageId="1" pageNumber="464">CEUA00106195</specimenCode>
</materialsCitation>
&quot;;
<materialsCitation id="3B0E18036B67FFC2FD13FF345966FE9B" collectingDate="1992-10-22" collectingDateMax="1992-11-02" collectingDateMin="1992-10-22" collectionCode="MNCR" collectorName="C. Moraga" country="Costa Rica" elevation="700" location="Sta. Cecilia" pageId="1" pageNumber="464" specimenCode="CEUA00106194" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D60D9D76B67FFC2FD13FF3458BFFF35" box="[685,716,220,242]" pageId="1" pageNumber="464" type="male">1♂</specimenCount>
&quot;Est. Pitilla,
<quantity id="4C9EBFBB6B67FFC2FCEAFF3459E6FF35" box="[852,917,221,242]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">
<elevation id="004BF56D6B67FFC2FCEAFF3459E6FF35" box="[852,917,221,242]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="464" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<quantity id="4C9EBFBB6B67FFC2FC1CFF3459BEFF35" box="[930,973,221,242]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="464" unit="km" value="9.0">9km</quantity>
S
<location id="8EB944856B67FFC2FC55FF345E2DFF35" LSID="urn:lsid:plazi:treatment:03CFA3486B67FFC1FF1EFBBA5EE0F9C0:8EB944856B67FFC2FC55FF345E2DFF35" box="[1003,1118,221,242]" country="Costa Rica" name="Sta. Cecilia" pageId="1" pageNumber="464" stateProvince="Guanacaste">Sta. Cecilia</location>
, P.N.
<collectingRegion id="49A2DCBC6B67FFC2FD13FF1E596EFECB" box="[685,797,247,268]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
, Prov.
<collectingRegion id="49A2DCBC6B67FFC2FCDDFF1E59A0FECB" box="[867,979,247,268]" country="Costa Rica" name="Guanacaste" pageId="1" pageNumber="464">Guanacaste</collectingRegion>
,
<collectingCountry id="F37152CE6B67FFC2FC60FF1E5E38FECB" box="[990,1099,247,268]" name="Costa Rica" pageId="1" pageNumber="464">Costa Rica</collectingCountry>
,
<date id="FFD8349E6B67FFC2FBE8FF1E594BFEE0" pageId="1" pageNumber="464" value="1992-10-22" valueMax="1992-11-02" valueMin="1992-10-22">
<collectingDate id="EF9CCD766B67FFC2FBE8FF1E594BFEE0" pageId="1" pageNumber="464" value="1992-10-22" valueMax="1992-11-02" valueMin="1992-10-22">22 oct a 2 nov 1992</collectingDate>
</date>
,
<collectorName id="269377886B67FFC2FCFAFEFB59C3FEE0" box="[836,944,274,295]" pageId="1" pageNumber="464">C. Moraga</collectorName>
L-N 330200,380200 /
<specimenCode id="DBC0BA256B67FFC2FD13FEC55929FE86" box="[685,858,300,321]" collectionCode="CEUA" pageId="1" pageNumber="464">CEUA00106194</specimenCode>
&quot;. Four
<typeStatus id="54DDACFC6B67FFC2FC18FEC55E76FE86" box="[934,1029,300,321]" pageId="1" pageNumber="464" type="paratype">paratypes</typeStatus>
are deposited at
<collectionCode id="ED778A9B6B67FFC2FD77FEAE5966FE9B" box="[713,789,327,348]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="1" pageNumber="464">MNCR</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B0E18036B67FFC2FC9DFEAE59E7FEB1" country="Costa Rica" location="Universidad de Alicante" pageId="1" pageNumber="464" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
and
<specimenCount id="9D60D9D76B67FFC2FCEFFEAE5990FE9B" box="[849,995,327,348]" pageId="1" pageNumber="464" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
are deposited at
<location id="8EB944856B67FFC2FD13FE8859E7FEB1" LSID="urn:lsid:plazi:treatment:03CFA3486B67FFC1FF1EFBBA5EE0F9C0:8EB944856B67FFC2FD13FE8859E7FEB1" box="[685,916,353,374]" country="Costa Rica" name="Universidad de Alicante" pageId="1" pageNumber="464" stateProvince="Guanacaste">Universidad de Alicante</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD9125E6B67FFC2FD7BFE955E25F9C0" blockId="1.[685,1173,194,1730]" pageId="1" pageNumber="464">
<emphasis id="B912CE4C6B67FFC2FD7BFE95595AFE56" bold="true" box="[709,809,380,401]" pageId="1" pageNumber="464">
<typeStatus id="54DDACFC6B67FFC2FD7BFE955957FE56" box="[709,804,380,401]" pageId="1" pageNumber="464" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="135D0EDB6B67FFC2FCC4FE9559CAFE56" box="[890,953,380,401]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="1" pageNumber="464">Fig. 1</figureCitation>
). Body oval. Length
<quantity id="4C9EBFBB6B67FFC2FD13FE7E5978FE6B" box="[685,779,407,428]" metricMagnitude="-3" metricUnit="m" metricValue="8.99" pageId="1" pageNumber="464" unit="mm" value="8.99">8.99 mm</quantity>
, width
<quantity id="4C9EBFBB6B67FFC2FCE2FE7E59C9FE6B" box="[860,954,407,428]" metricMagnitude="-3" metricUnit="m" metricValue="5.19" pageId="1" pageNumber="464" unit="mm" value="5.19">5.19 mm</quantity>
. Head and pronotum dark brown. Scutellum reddish brown. Elytra dark brown but lighter than pronotum. Pygidium ochre. Legs orange on femur, reddish brown on tibia and tarsus. Venter orange on sternum, abdominal sternites ochre.
<emphasis id="B912CE4C6B67FFC2FC8EFDF25901FDF7" bold="true" box="[816,882,539,560]" pageId="1" pageNumber="464">Head:</emphasis>
Clypeus trapezoidal, short, surface densely punctate-reticulate, convex; anterior margin curved, clypeus thick in frontal view; width/length ratio 2.50. Frons sparsely punctate, flat. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/width of eye 2.84. Ratio antennal funiculus/club 0.74.
<emphasis id="B912CE4C6B67FFC2FC58FD525E29FD17" bold="true" box="[998,1114,699,720]" pageId="1" pageNumber="464">Pronotum:</emphasis>
Subtrapezoidal. Ratio width/length 1.69. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles acute, posterior angles obtuse. Basal margin sinuate, subapical bead obliterated at middle. Surface with deep, dense punctures.
<emphasis id="B912CE4C6B67FFC2FCA8FCB359F2FCA8" bold="true" box="[790,897,858,879]" pageId="1" pageNumber="464">Scutellum:</emphasis>
Subtriangular, with rounded sides, and acute apex. Ratio width/length 1.19. Surface sparsely punctate.
<emphasis id="B912CE4C6B67FFC2FC34FC6659A3FC63" bold="true" box="[906,976,911,932]" pageId="1" pageNumber="464">Elytra:</emphasis>
With costae defined by regular rows of punctures. Subsutural interstice with 1–2 rows of sparse punctures, 2
<superScript id="7C13BF166B67FFC2FBA1FC565E42FC0A" attach="none" box="[1055,1073,959,973]" fontSize="6" pageId="1" pageNumber="464">nd</superScript>
–4
<superScript id="7C13BF166B67FFC2FBF4FC565E2BFC0A" attach="left" box="[1098,1112,959,973]" fontSize="6" pageId="1" pageNumber="464">th</superScript>
interstices with 1 irregular, secondary stria.
<emphasis id="B912CE4C6B67FFC2FB92FC365EE7FC33" bold="true" box="[1068,1172,991,1012]" pageId="1" pageNumber="464">Pygidium:</emphasis>
Sparsely punctate with long setae on posterior margin. Ratio width/length 1.77.
<emphasis id="B912CE4C6B67FFC2FBBAFBFD5E3CFBEE" bold="true" box="[1028,1103,1044,1065]" pageId="1" pageNumber="464">Venter:</emphasis>
Space between mesocoxae narrow, flat. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae and 4–5 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane thin.
<emphasis id="B912CE4C6B67FFC2FD61FB5D5965FB0E" bold="true" box="[735,790,1204,1225]" pageId="1" pageNumber="464">Legs:</emphasis>
Protibia (
<figureCitation id="135D0EDB6B67FFC2FCC8FB5D59C3FB0E" box="[886,944,1204,1225]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="1" pageNumber="464">Fig. 9</figureCitation>
) tridentate, apical tooth long and curved, middle tooth above internal apex of protibia, triangular in shape, acute, basal tooth sharp. Metatibia stout, fusiform; ratio length/width 2.81. First external carina with row of sparse setae, surface punctate. External protarsal claw strongly curved, internal claw bifurcate, with upper branch slightly shorter and 2/3 width of the lower one, inferior margin curved.
<emphasis id="B912CE4C6B67FFC2FC2DFA615E59FA5A" bold="true" box="[915,1066,1416,1437]" pageId="1" pageNumber="464">Male genitalia:</emphasis>
Parameres slender (
<figureCitation id="135D0EDB6B67FFC2FD41FA4A5935FA7F" box="[767,838,1443,1464]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="1" pageNumber="464">Fig. 17</figureCitation>
), with squared, protruding apex in lateral view, ventral margin sinuate. Endophallus (
<figureCitation id="135D0EDB6B67FFC2FD0AFA315884FA2A" box="[692,759,1496,1517]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="1" pageNumber="464">Fig. 25</figureCitation>
) with 1 long, glabrous sacculus. Ejaculatory duct opening frontally on a ventral inflation.
</paragraph>
<paragraph id="8BD9125E6B67FFC2FD7BF9E45E0DF9FB" blockId="1.[685,1173,194,1730]" pageId="1" pageNumber="464">
<emphasis id="B912CE4C6B67FFC2FD7BF9E45965F9E5" bold="true" box="[709,790,1549,1570]" pageId="1" pageNumber="464">Female.</emphasis>
Similar to male except protibia (
<figureCitation id="135D0EDB6B67FFC2FBEAF9E45EFFF9E5" box="[1108,1164,1549,1570]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="1" pageNumber="464">Fig. 9</figureCitation>
) with wider teeth; last sternite with curved apex.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B67FFC1FD7BF9AB5BCDFA15" lastPageId="2" lastPageNumber="465" pageId="1" pageNumber="464" type="description">
<paragraph id="8BD9125E6B67FFC2FD7BF9AB5EE0F906" blockId="1.[685,1173,194,1730]" pageId="1" pageNumber="464">
<emphasis id="B912CE4C6B67FFC2FD7BF9AB5959F990" bold="true" box="[709,810,1602,1623]" pageId="1" pageNumber="464">Variation.</emphasis>
Six
<typeStatus id="54DDACFC6B67FFC2FCE1F9AB59CEF990" box="[863,957,1602,1623]" pageId="1" pageNumber="464" type="paratype">paratypes</typeStatus>
. Dorsal surface varies from dark reddish brown to dark brown; pygidium and venter from orange to ochre. Body length
<quantity id="4C9EBFBB6B67FFC2FD13F97B5932F960" box="[685,833,1681,1703]" metricMagnitude="-3" metricUnit="m" metricValue="9.58" metricValueMax="10.17" metricValueMin="8.99" pageId="1" pageNumber="464" unit="mm" value="9.58" valueMax="10.17" valueMin="8.99">8.99–10.17 mm</quantity>
, body width
<quantity id="4C9EBFBB6B67FFC2FC04F97B5E30F960" box="[954,1091,1681,1703]" metricMagnitude="-3" metricUnit="m" metricValue="5.475" metricValueMax="5.76" metricValueMin="5.19" pageId="1" pageNumber="464" unit="mm" value="5.475" valueMax="5.76" valueMin="5.19">5.19–5.76 mm</quantity>
. Clypeal width/length ratio 2.50–2.93. Ratio of interocular
</paragraph>
<caption id="DF1942D66B64FFC1FF7BFB135EDAFACA" ID-DOI="http://doi.org/10.5281/zenodo.10106052" ID-Zenodo-Dep="10106052" box="[197,1193,1273,1293]" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="2" pageNumber="465" startId="2.[197,237,1274,1293]" targetBox="[193,1167,186,1250]" targetPageId="2">
<paragraph id="8BD9125E6B64FFC1FF7BFB135EDAFACA" blockId="2.[173,1193,1273,1316]" box="[197,1193,1273,1293]" pageId="2" pageNumber="465">
<emphasis id="B912CE4C6B64FFC1FF7BFB135B6CFACA" bold="true" box="[197,287,1273,1293]" pageId="2" pageNumber="465">Figs. 1–8.</emphasis>
<taxonomicName id="4C6669DD6B64FFC1FE8AFB105BCCFACA" box="[308,447,1273,1293]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B912CE4C6B64FFC1FE8AFB105B0DFACB" box="[308,382,1273,1292]" italics="true" pageId="2" pageNumber="465">Anomala</emphasis>
species
</taxonomicName>
holotypes, habitus.
<emphasis id="B912CE4C6B64FFC1FDDDFB135805FACA" bold="true" box="[611,630,1274,1293]" pageId="2" pageNumber="465">1)</emphasis>
<taxonomicName id="4C6669DD6B64FFC1FDC5FB1058B0FACB" box="[635,707,1273,1292]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="coffea">
<emphasis id="B912CE4C6B64FFC1FDC5FB1058B0FACB" box="[635,707,1273,1292]" italics="true" pageId="2" pageNumber="465">A. coffea</emphasis>
</taxonomicName>
;
<emphasis id="B912CE4C6B64FFC1FD71FB135891FACA" bold="true" box="[719,738,1274,1293]" pageId="2" pageNumber="465">2)</emphasis>
<taxonomicName id="4C6669DD6B64FFC1FD59FB105948FACB" box="[743,827,1273,1292]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="cyclops">
<emphasis id="B912CE4C6B64FFC1FD59FB105948FACB" box="[743,827,1273,1292]" italics="true" pageId="2" pageNumber="465">A. cyclops</emphasis>
</taxonomicName>
;
<emphasis id="B912CE4C6B64FFC1FCF8FB13592AFACA" bold="true" box="[838,857,1274,1293]" pageId="2" pageNumber="465">3)</emphasis>
<taxonomicName id="4C6669DD6B64FFC1FCE1FB1059D5FACB" box="[863,934,1273,1292]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="divisa">
<emphasis id="B912CE4C6B64FFC1FCE1FB1059D5FACB" box="[863,934,1273,1292]" italics="true" pageId="2" pageNumber="465">A. divisa</emphasis>
</taxonomicName>
;
<emphasis id="B912CE4C6B64FFC1FC0FFB1359B6FACA" bold="true" box="[945,965,1274,1293]" pageId="2" pageNumber="465">4)</emphasis>
<taxonomicName id="4C6669DD6B64FFC1FC74FB105E60FACB" box="[970,1043,1273,1292]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="mersa">
<emphasis id="B912CE4C6B64FFC1FC74FB105E60FACB" box="[970,1043,1273,1292]" italics="true" pageId="2" pageNumber="465">A. mersa</emphasis>
</taxonomicName>
;
<emphasis id="B912CE4C6B64FFC1FBA0FB135E42FACA" bold="true" box="[1054,1073,1274,1293]" pageId="2" pageNumber="465">5)</emphasis>
<taxonomicName id="4C6669DD6B64FFC1FB89FB105ED0FACB" box="[1079,1187,1273,1292]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="mesosticta">
<emphasis id="B912CE4C6B64FFC1FB89FB105ED0FACB" box="[1079,1187,1273,1292]" italics="true" pageId="2" pageNumber="465">A. mesosticta</emphasis>
</taxonomicName>
;
</paragraph>
</caption>
<paragraph id="8BD9125E6B64FFC1FF13FAF8594FFAE3" blockId="2.[173,1193,1273,1316]" box="[173,828,1297,1316]" pageId="2" pageNumber="465">
<emphasis id="B912CE4C6B64FFC1FF13FAF85AB3FAE3" bold="true" box="[173,192,1297,1316]" pageId="2" pageNumber="465">6)</emphasis>
<taxonomicName id="4C6669DD6B64FFC1FF79FAF85B4BFAE3" box="[199,312,1297,1316]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="robiginosa">
<emphasis id="B912CE4C6B64FFC1FF79FAF85B4BFAE3" box="[199,312,1297,1316]" italics="true" pageId="2" pageNumber="465">A. robiginosa</emphasis>
</taxonomicName>
;
<emphasis id="B912CE4C6B64FFC1FEFAFAF85B2BFAE3" bold="true" box="[324,344,1297,1316]" pageId="2" pageNumber="465">7)</emphasis>
<taxonomicName id="4C6669DD6B64FFC1FEE0FAF85B8EFAE3" box="[350,509,1297,1316]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="strigodermoides">
<emphasis id="B912CE4C6B64FFC1FEE0FAF85B8EFAE3" box="[350,509,1297,1316]" italics="true" pageId="2" pageNumber="465">A. strigodermoides</emphasis>
</taxonomicName>
;
<emphasis id="B912CE4C6B64FFC1FDB4FAF8586EFAE3" bold="true" box="[522,541,1297,1316]" pageId="2" pageNumber="465">8)</emphasis>
<taxonomicName id="4C6669DD6B64FFC1FD9AFAF858E3FAE3" box="[548,656,1297,1316]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="unilineata">
<emphasis id="B912CE4C6B64FFC1FD9AFAF858E3FAE3" box="[548,656,1297,1316]" italics="true" pageId="2" pageNumber="465">A. unilineata</emphasis>
</taxonomicName>
. Scale bar =
<quantity id="4C9EBFBB6B64FFC1FCBAFAF8594BFAE3" box="[772,824,1297,1316]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="465" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
<paragraph id="8BD9125E6B64FFC1FF13FABA5BCDFA15" blockId="2.[173,660,1362,1729]" pageId="2" pageNumber="465">width/width of eye 2.84–3.19. Ratio of antennal funiculus/club 0.73–0.79. Pronotal width/length ratio 1.69–1.72. Scutellar width/length ratio 1.13–1.33. Pygidial width/length ratio 1.68–1.82. Metatibial length/width ratio 2.60–2.81.</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B64FFC1FF7BFA31586DF9FB" pageId="2" pageNumber="465" type="etymology">
<paragraph id="8BD9125E6B64FFC1FF7BFA31586DF9FB" blockId="2.[173,660,1362,1729]" pageId="2" pageNumber="465">
<emphasis id="B912CE4C6B64FFC1FF7BFA315B44FA2A" bold="true" box="[197,311,1496,1517]" pageId="2" pageNumber="465">Etymology.</emphasis>
From the generic name of coffee plants, to be treated as a noun in apposition, for its resemblance in shape and color to toasted coffee beans, a famous Costa Rican product.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B64FFC1FF7BF9AB58E7F94B" pageId="2" pageNumber="465" type="distribution">
<paragraph id="8BD9125E6B64FFC1FF7BF9AB58E7F94B" blockId="2.[173,660,1362,1729]" pageId="2" pageNumber="465">
<emphasis id="B912CE4C6B64FFC1FF7BF9AB5B34F990" bold="true" box="[197,327,1602,1623]" pageId="2" pageNumber="465">Distribution.</emphasis>
This species is found in the humid forest on the northwest side of the
<collectingRegion id="49A2DCBC6B64FFC1FE56F9B55822F9B6" box="[488,593,1628,1649]" country="Costa Rica" name="Guanacaste" pageId="2" pageNumber="465">Guanacaste</collectingRegion>
mountain range,
<collectingCountry id="F37152CE6B64FFC1FEABF99E5B0FF94B" box="[277,380,1655,1676]" name="Costa Rica" pageId="2" pageNumber="465">Costa Rica</collectingCountry>
, at
<quantity id="4C9EBFBB6B64FFC1FE20F99E5BA9F94B" box="[414,474,1655,1676]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="465" unit="m" value="700.0">700 m</quantity>
elevation (
<figureCitation id="135D0EDB6B64FFC1FDFDF99E58FAF94B" box="[579,649,1655,1676]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="2" pageNumber="465">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B64FFC1FF7BF97B5EE0F9C0" pageId="2" pageNumber="465" type="diagnosis">
<paragraph id="8BD9125E6B64FFC1FF7BF97B5EE0F9C0" blockId="2.[173,660,1362,1729]" lastBlockId="2.[707,1193,1363,1543]" pageId="2" pageNumber="465">
<emphasis id="B912CE4C6B64FFC1FF7BF97B5B59F960" bold="true" box="[197,298,1682,1703]" pageId="2" pageNumber="465">Diagnosis.</emphasis>
This species is recognized by its small size, color homogeneously dark brown, elytra with defined costae and interstices with rows of punctures, and tridentate protibia. The small size and brown color is similar to that of species of
<taxonomicName id="4C6669DD6B64FFC1FBEAFA6E5EDAFA5A" box="[1108,1193,1415,1437]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis id="B912CE4C6B64FFC1FBEAFA6E5EDAFA5A" box="[1108,1193,1415,1437]" italics="true" pageId="2" pageNumber="465">Anomala</emphasis>
</taxonomicName>
in the subgenus
<taxonomicName id="4C6669DD6B64FFC1FCD1FA4B590BFA15" authority="Ramirez-Ponce and Moron, 2012" authorityName="Ramirez-Ponce and Moron" authorityYear="2012" class="Insecta" family="Rutelidae" genus="Bucaphallanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis id="B912CE4C6B64FFC1FCD1FA4B5E75FA7F" box="[879,1030,1442,1464]" italics="true" pageId="2" pageNumber="465">Bucaphallanus</emphasis>
Ramírez-Ponce and Morón, 2012
</taxonomicName>
, but
<taxonomicName id="4C6669DD6B64FFC1FC11FA545E78FA14" box="[943,1035,1469,1491]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="coffea">
<emphasis id="B912CE4C6B64FFC1FC11FA545E78FA14" box="[943,1035,1469,1491]" italics="true" pageId="2" pageNumber="465">A. coffea</emphasis>
</taxonomicName>
does not share any diagnostic character with any of its species and has more defined punctation on the elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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