286 lines
35 KiB
XML
286 lines
35 KiB
XML
<document id="F9E7F46A6FF745EDCC7FD5CB4B50F90D" ID-CLB-Dataset="24794" ID-DOI="10.11646/zootaxa.4701.1.4" ID-GBIF-Dataset="ca29e500-63fd-4f87-b24b-c41cdfa2cf1f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3557825" ID-ZooBank="338B950C-1590-4E98-8254-83DD1AD39259" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575043589075" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03CF87DF697EFFE115C2AD31FC95FAF0" docLanguage="en" docName="zootaxa.4701.1.4.pdf" docOrigin="Zootaxa 4701 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Veraphis loebli Jałoszyński 2019, sp. n." docType="treatment" docVersion="7" lastPageNumber="64" masterDocId="FFF6FFA7697FFFE31555A80CFFA6FFA7" masterDocTitle="Four new species and new records of Veraphis Casey in Japan (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="64" masterPageNumber="54" pageNumber="54" updateTime="1698769603799" updateUser="ExternalLinkService">
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<mods:mods id="915814FEEEA2FCF57AC0A58DD62AC9EB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FC241D2B6309E52BF82D9F0BA9B1668E">
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<mods:title id="CC0A2E29025E7EF6436C2E6C22F90929">Four new species and new records of Veraphis Casey in Japan (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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</mods:titleInfo>
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<mods:name id="4B4D685FF8BDF6A99B85560F400D80A2" type="personal">
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<mods:role id="2976013B9DEA6C117E95B81457724E37">
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<mods:roleTerm id="8CC746317BCC9D8A2E44E7CAAB8CE9DF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="97F38D5F90D7E689E94B3AAC7CBFF962">Jałoszyński, Paweł</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0CE5D4BC93C7DBADF672D4D90C510466">text</mods:typeOfResource>
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<mods:titleInfo id="AE9CBC5C67ECFF86922BBA2780308A43">
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<mods:title id="DEE7726C3836F75F25FB81F7EC701DB9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6482FCBF5D6F7A5ECC07929CFC1C306C">
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<mods:date id="8B6E388202A92BB21BE783430AAE57B1">2019</mods:date>
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<mods:detail id="302863BC3633FFF08703FACC9947455F" type="pubDate">
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<mods:number id="3F13BDF48F3670CC646E604E30F302E6">2019-11-26</mods:number>
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</mods:detail>
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<mods:detail id="E4C4A6EE94F91073CF12FFAAAAF57C69" type="volume">
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<mods:number id="4358E68AEA705AE611E9AC6622737E62">4701</mods:number>
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</mods:detail>
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<mods:detail id="2A2D59132C054DFC5B33413D32045693" type="issue">
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<mods:number id="DE8D939A80BE02E4D4F5F71A26F3DF4A">1</mods:number>
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<mods:start id="DDB479EA4A03AEB7D54D91DC7E442623">54</mods:start>
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<mods:end id="2A3F7168428A5A1D42A47E8634EE8A04">64</mods:end>
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<mods:classification id="8F2C79D5844B8F9DB4B83822DDE9842F">journal article</mods:classification>
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<mods:identifier id="7F5627C98E2C8A78F04CDE69AA33CD2F" type="CLB-Dataset">24794</mods:identifier>
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<mods:identifier id="0B3C92BEA261D74A69074309EDF601FA" type="DOI">10.11646/zootaxa.4701.1.4</mods:identifier>
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<mods:identifier id="B261EA9880D975EDF5FAE4ECCEA93894" type="GBIF-Dataset">ca29e500-63fd-4f87-b24b-c41cdfa2cf1f</mods:identifier>
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<mods:identifier id="E766CABC535DA4F2DA31517FFA261807" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="38D2209581CE3DD216D10CD5D9D36B02" type="Zenodo-Dep">3557825</mods:identifier>
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<mods:identifier id="B196A390C8238B3971778DF34C1A0A9A" type="ZooBank">338B950C-1590-4E98-8254-83DD1AD39259</mods:identifier>
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</mods:mods>
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<treatment id="03CF87DF697EFFE115C2AD31FC95FAF0" ID-GBIF-Taxon="160605358" LSID="urn:lsid:plazi:treatment:03CF87DF697EFFE115C2AD31FC95FAF0" httpUri="http://treatment.plazi.org/id/03CF87DF697EFFE115C2AD31FC95FAF0" lastPageId="2" pageId="1">
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<subSubSection id="C37C6542697EFFE215C2AD31FE33FAF0" box="[151,405,1341,1368]" pageId="1" type="nomenclature">
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<paragraph id="8BD936C9697EFFE215C2AD31FE33FAF0" blockId="1.[151,463,1341,1403]" box="[151,405,1341,1368]" pageId="1">
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<heading id="D09181A5697EFFE215C2AD31FE33FAF0" bold="true" box="[151,405,1341,1368]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="B912EADB697EFFE215C2AD31FE33FAF0" bold="true" box="[151,405,1341,1368]" pageId="1">
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<taxonomicName id="4C664D4A697EFFE215C2AD31FEEDFAFF" ID-CoL="8YC6Y" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,331,1341,1368]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
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<emphasis id="B912EADB697EFFE215C2AD31FEEDFAFF" bold="true" box="[151,331,1341,1368]" italics="true" pageId="1">Veraphis loebli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157A0697EFFE21406AD31FE33FAF0" box="[339,405,1341,1367]" pageId="1" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C37C6542697EFFE215C2AD6DFE69FADC" box="[151,463,1376,1403]" pageId="1" type="description">
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<paragraph id="8BD936C9697EFFE215C2AD6DFE69FADC" blockId="1.[151,463,1341,1403]" box="[151,463,1376,1403]" pageId="1">
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(
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<figureCitation id="135D2A4C697EFFE215CAAD6DFF40FADC" box="[159,230,1377,1403]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1575,1600]" captionTargetBox="[155,1426,854,1517]" captionTargetId="figure@4.[151,1436,841,1526]" captionTargetPageId="4" captionText="FIGURES 1–4. Dorsal habitus of males. Veraphis loebli sp. n. (1); V. mutsuensis sp. n. (2); V. aomoriensis sp. n. (3); V. sakaii sp. n. (4)." figureDoi="http://doi.org/10.5281/zenodo.3557827" httpUri="https://zenodo.org/record/3557827/files/figure.png" pageId="1">Figs 1</figureCitation>
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,
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<figureCitation id="135D2A4C697EFFE215A1AD6CFE86FADC" box="[244,288,1376,1403]" captionStart="FIGURES 5–12" captionStartId="5.[151,264,1966,1991]" captionTargetBox="[184,1424,314,1942]" captionTargetId="figure@5.[151,1436,301,1946]" captionTargetPageId="5" captionText="FIGURES 5–12. Aedeagus in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views. Veraphis loebli sp. n. (5–6); V. mutsuensis sp. n. (7–8); V. aomoriensis sp. n. (9–10); V. sakaii sp. n. (11–12)." figureDoi="http://doi.org/10.5281/zenodo.3557829" httpUri="https://zenodo.org/record/3557829/files/figure.png" pageId="1">5–6</figureCitation>
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,
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<figureCitation id="135D2A4C697EFFE21479AD6DFED3FADC" box="[300,373,1377,1403]" captionStart="FIGURES 13–20" captionStartId="7.[151,264,1920,1945]" captionTargetBox="[176,1432,248,1881]" captionTargetId="figure@7.[151,1436,229,1901]" captionTargetPageId="7" captionText="FIGURES 13–20. Aedeagus in ventral (13, 15, 17, 19) and lateral (14, 16, 18, 20) views. Veraphis loebli sp. n. (13–14); V. mutsuensis sp. n. (15–16); V. aomoriensis sp. n. (17–18); V. sakaii sp. n. (19–20)." figureDoi="http://doi.org/10.5281/zenodo.3557831" httpUri="https://zenodo.org/record/3557831/files/figure.png" pageId="1">13–14</figureCitation>
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,
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<figureCitation id="135D2A4C697EFFE214D5AD6DFE38FADC" box="[384,414,1377,1403]" captionStart="FIGURES 21–24" captionStartId="8.[151,264,1456,1481]" captionTargetBox="[169,1427,432,1413]" captionTargetId="figure@8.[151,1436,404,1436]" captionTargetPageId="8" captionText="FIGURES 21–24. Fore leg of male. Veraphis loebli sp. n. (21); V. mutsuensis sp. n. (22); V. aomoriensis sp. n. (23); V. sakaii sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3557833" httpUri="https://zenodo.org/record/3557833/files/figure.png" pageId="1">21</figureCitation>
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,
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<figureCitation id="135D2A4C697EFFE214FFAD6CFE61FADC" box="[426,455,1376,1403]" captionStart="FIGURE 25" captionStartId="9.[151,250,1702,1727]" captionTargetBox="[151,1436,696,1668]" captionTargetId="figure@9.[151,1436,692,1683]" captionTargetPageId="9" captionText="FIGURE 25. Distribution of Veraphis in Japan." figureDoi="http://doi.org/10.5281/zenodo.3557835" httpUri="https://zenodo.org/record/3557835/files/figure.png" pageId="1">25</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C6542697EFFE215C2ADA5FE04F9AC" pageId="1" type="materials_examined">
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<paragraph id="8BD936C9697EFFE215C2ADA5FE04F9AC" blockId="1.[151,1437,1448,2015]" pageId="1">
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<emphasis id="B912EADB697EFFE215C2ADA5FEE2FA64" bold="true" box="[151,324,1448,1475]" pageId="1">Type material.</emphasis>
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<materialsCitation id="3B0E3C94697EFFE2141FADA5FE04F9AC" ID-GBIF-Occurrence="2460169393" collectionCode="MHNG" collectorName="P. Jaloszynski" country="Japan" elevation="1350" location="Honshu" pageId="1" specimenCount="1" specimenCount-male="1" stateProvince="Tochigi" typeStatus="holotype">
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<emphasis id="B912EADB697EFFE2141FADA5FCE2FA64" bold="true" box="[330,836,1448,1475]" pageId="1">
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<typeStatus id="54DD886B697EFFE2141FADA5FE1DFA64" box="[330,443,1449,1475]" pageId="1" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3717659697EFFE21492ADA4FD87FA64" box="[455,545,1448,1475]" name="Japan" pageId="1">JAPAN</collectingCountry>
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(
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<location id="8EB96012697EFFE2177BADA4FD29FA64" LSID="urn:lsid:plazi:treatment:03CF87DF697EFFE115C2AD31FC95FAF0:8EB96012697EFFE2177BADA4FD29FA64" box="[558,655,1448,1475]" country="Japan" name="Honshu" pageId="1" stateProvince="Tochigi">Honshu</location>
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,
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<collectingRegion id="49A2F82B697EFFE217CFADA4FC91FA64" box="[666,823,1448,1475]" country="Japan" name="Tochigi" pageId="1">Tochigi Pref.</collectingRegion>
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):
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</emphasis>
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<specimenCount id="9D60FD40697EFFE2161FADA5FCC7FA64" box="[842,865,1449,1475]" pageId="1" type="male">♂</specimenCount>
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, two labels: “JAPAN
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<collectingRegion id="49A2F82B697EFFE21107ADA5FB63FA64" box="[1106,1221,1449,1475]" country="Japan" name="Tochigi" pageId="1">TOCHIGI</collectingRegion>
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/ Nikko Nat. Park / Chuzenji
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<quantity id="4C9E9B2C697EFFE21456ADC0FEFEFA40" box="[259,344,1484,1511]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="1" unit="m" value="1350.0">
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||
<elevation id="004BD1FA697EFFE21456ADC0FEFEFA40" box="[259,344,1484,1511]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="1" unit="m" value="1350.0">1350 m</elevation>
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</quantity>
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/ Löbl 14.7.980” [white, printed], “
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<taxonomicName id="4C664D4A697EFFE217B0ADC1FCC0FA40" authorityName="Casey" authorityYear="1897" box="[741,870,1485,1511]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B912EADB697EFFE217B0ADC1FCC0FA40" box="[741,870,1485,1511]" italics="true" pageId="1">VERAPHIS</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C664D4A697EFFE2162EADC1FC1DFA40" authorityName="Jałoszyński" authorityYear="2019" box="[891,955,1485,1511]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="loebli">
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<emphasis id="B912EADB697EFFE2162EADC1FC1DFA40" box="[891,955,1485,1511]" italics="true" pageId="1">loebli</emphasis>
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</taxonomicName>
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m. /
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<collectorName id="2693531F697EFFE216A6ADC1FB37FA40" box="[1011,1169,1485,1511]" pageId="1">P. Jałoszyński</collectorName>
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, 2019 /
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<typeStatus id="54DD886B697EFFE211BCADC1FA35FA40" box="[1257,1427,1485,1511]" pageId="1" type="holotype">HOLOTYPUS</typeStatus>
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” [red, printed] (
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<collectionCode id="ED77AE0C697EFFE2146FADFDFE30F9AC" box="[314,406,1521,1547]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" type="Museum">MHNG</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C6542697EFFE21592AE18FB27F93C" pageId="1" type="diagnosis">
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<paragraph id="8BD936C9697EFFE21592AE18FB27F93C" blockId="1.[151,1437,1448,2015]" pageId="1">
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<emphasis id="B912EADB697EFFE21592AE18FEE4F988" bold="true" box="[199,322,1556,1583]" pageId="1">Diagnosis.</emphasis>
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Male protrochanter unmodified; male protibia with subapical fin-like projection about as broad as 2/3 of tibial width; median lobe of aedeagus in lateral view strongly curved; in ventral view with shallowly concave walls in submedian and subapical regions; ventral diaphragm shorter than 1/3 of median lobe, close to basal margin of aedeagus; apical margin shallowly concave; parameres in lateral view strongly curved.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C6542697EFFE11592AEA8FE38FD54" lastPageId="2" pageId="1" type="description">
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<paragraph id="8BD936C9697EFFE21592AEA8FDCAF944" blockId="1.[151,1437,1448,2015]" pageId="1">
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<emphasis id="B912EADB697EFFE21592AEA8FEFFF918" bold="true" box="[199,345,1700,1727]" pageId="1">Description.</emphasis>
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BL
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<quantity id="4C9E9B2C697EFFE214DFAEA9FE57F918" box="[394,497,1701,1727]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="1" unit="mm" value="1.23">1.23 mm</quantity>
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. Body of male (
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<figureCitation id="135D2A4C697EFFE217F1AEA9FD42F918" box="[676,740,1701,1727]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1575,1600]" captionTargetBox="[155,1426,854,1517]" captionTargetId="figure@4.[151,1436,841,1526]" captionTargetPageId="4" captionText="FIGURES 1–4. Dorsal habitus of males. Veraphis loebli sp. n. (1); V. mutsuensis sp. n. (2); V. aomoriensis sp. n. (3); V. sakaii sp. n. (4)." figureDoi="http://doi.org/10.5281/zenodo.3557827" httpUri="https://zenodo.org/record/3557827/files/figure.png" pageId="1">Fig. 1</figureCitation>
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) flattened, slender, moderately dark brown, covered with yel- lowish vestiture, antennae slightly lighter.
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</paragraph>
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<paragraph id="8BD936C9697EFFE21592AEE1FAC9F878" blockId="1.[151,1437,1448,2015]" pageId="1">
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Head broadest at large, moderately convex eyes, HL
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<quantity id="4C9E9B2C697EFFE21646AEE0FCDEF8A0" box="[787,888,1772,1799]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" unit="mm" value="0.15">0.15 mm</quantity>
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, HW
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<quantity id="4C9E9B2C697EFFE216EDAEE1FBB8F8A0" box="[952,1054,1773,1799]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="1" unit="mm" value="0.23">0.23 mm</quantity>
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||
; vertex with pair of small postero- median pits, each prolonged by shallow but distinct longitudinal groove extending anterolaterad to posteromesal margin of weakly elevated supraantennal tubercle; area between grooves distinctly flattened and depressed in rela- tion to convex sides of vertex and frons. Punctures on head dorsum fine and shallow, inconspicuous; setae short, sparse and suberect. Antennae slender, with distinctly demarcated trimerous club, AnL
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<quantity id="4C9E9B2C697EFFE2110AAF70FB60F830" box="[1119,1222,1916,1943]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="1" unit="mm" value="0.45">0.45 mm</quantity>
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; antennomeres I–II distinctly elongate; III distinctly transverse; IV about as long as broad; V indistinctly elongate; VI as long as broad; VII–X each slightly transverse (VIII narrower than VII), XI indistinctly elongate and slightly narrower than X.
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</paragraph>
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<paragraph id="8BD936C9697DFFE11592A895FCE5FEC0" blockId="2.[151,1437,152,1367]" pageId="2">
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Pronotum nearly semielliptical, broadest near anterior third; PL
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||
<quantity id="4C9E9B2C697DFFE116DDA895FC4BFF14" box="[904,1005,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" unit="mm" value="0.3">0.30 mm</quantity>
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, PW
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<quantity id="4C9E9B2C697DFFE11173A894FB2AFF14" box="[1062,1164,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="2" unit="mm" value="0.35">0.35 mm</quantity>
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||
; anterior margin broadly and evenly rounded; lateral margins strongly rounded in anterior third, behind middle nearly straight and distinctly convergent towards obtuse and blunt hind angles; posterior margin indistinctly bisinuate. Pronotal base with small and shallow median antebasal pit located in distinct transverse groove which laterally does not reach lateral pair of elongate and deep impressions; lateral pronotal margins narrowly carinate in posterior 2/3. Punctures on pronotal disc fine and inconspicuous; setae short and sparse, suberect.
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</paragraph>
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<paragraph id="8BD936C9697DFFE11592A97DFA9DFE74" blockId="2.[151,1437,152,1367]" pageId="2">
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Elytra slightly more convex than pronotum, together oval, broadest slightly in front of middle; EL
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<quantity id="4C9E9B2C697DFFE11057A97DFAC1FE2C" box="[1282,1383,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="2" unit="mm" value="0.63">0.63 mm</quantity>
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, EW
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<quantity id="4C9E9B2C697DFFE115C2A999FF5BFE08" box="[151,253,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="2" unit="mm" value="0.48">0.48 mm</quantity>
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, EI 1.32; humeral calli well-marked, elongate. Surface of elytra less glossy than that of pronotum, covered with fine, superficial, inconspicuous punctures and setae similar to those on pronotum but slightly thicker.
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</paragraph>
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<paragraph id="8BD936C9697DFFE11592A9D1FE0CFDBC" blockId="2.[151,1437,152,1367]" pageId="2">
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Legs moderately long and slender; protrochanter unmodified, protibia with conspicuously large subapical finlike projection (
|
||
<figureCitation id="135D2A4C697DFFE1141CAA0DFE3AFDBC" box="[329,412,513,539]" captionStart="FIGURES 21–24" captionStartId="8.[151,264,1456,1481]" captionTargetBox="[169,1427,432,1413]" captionTargetId="figure@8.[151,1436,404,1436]" captionTargetPageId="8" captionText="FIGURES 21–24. Fore leg of male. Veraphis loebli sp. n. (21); V. mutsuensis sp. n. (22); V. aomoriensis sp. n. (23); V. sakaii sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3557833" httpUri="https://zenodo.org/record/3557833/files/figure.png" pageId="2">Fig. 21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD936C9697DFFE11592AA29FC0FFD68" blockId="2.[151,1437,152,1367]" pageId="2">
|
||
Aedeagus (
|
||
<figureCitation id="135D2A4C697DFFE11411AA29FE01FD98" box="[324,423,548,575]" captionStart="FIGURES 5–12" captionStartId="5.[151,264,1966,1991]" captionTargetBox="[184,1424,314,1942]" captionTargetId="figure@5.[151,1436,301,1946]" captionTargetPageId="5" captionText="FIGURES 5–12. Aedeagus in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views. Veraphis loebli sp. n. (5–6); V. mutsuensis sp. n. (7–8); V. aomoriensis sp. n. (9–10); V. sakaii sp. n. (11–12)." figureDoi="http://doi.org/10.5281/zenodo.3557829" httpUri="https://zenodo.org/record/3557829/files/figure.png" pageId="2">Figs 5–6</figureCitation>
|
||
,
|
||
<figureCitation id="135D2A4C697DFFE114E6AA29FE5CFD98" box="[435,506,549,575]" captionStart="FIGURES 13–20" captionStartId="7.[151,264,1920,1945]" captionTargetBox="[176,1432,248,1881]" captionTargetId="figure@7.[151,1436,229,1901]" captionTargetPageId="7" captionText="FIGURES 13–20. Aedeagus in ventral (13, 15, 17, 19) and lateral (14, 16, 18, 20) views. Veraphis loebli sp. n. (13–14); V. mutsuensis sp. n. (15–16); V. aomoriensis sp. n. (17–18); V. sakaii sp. n. (19–20)." figureDoi="http://doi.org/10.5281/zenodo.3557831" httpUri="https://zenodo.org/record/3557831/files/figure.png" pageId="2">13–14</figureCitation>
|
||
) elongate and slender; AeL
|
||
<quantity id="4C9E9B2C697DFFE11665AA29FC30FD98" box="[816,918,549,575]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="2" unit="mm" value="0.28">0.28 mm</quantity>
|
||
; in ventral view median lobe broadly and shal- lowly constricted in sub-basal region, with subtrapezoidal, elongate apical portion, with lateral margins shallowly concave in subapical area; apical margin shallowly concave; ventral diaphragm small, shorter than 1/3 of median lobe and with its proximal margin close to base of aedeagus; in lateral view median lobe strongly curved, also para- meres strongly curved, each with three apical setae of various lengths.
|
||
</paragraph>
|
||
<paragraph id="8BD936C9697DFFE11592AAD5FE38FD54" blockId="2.[151,1437,152,1367]" box="[199,414,729,755]" pageId="2">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C6542697DFFE11592AAF0FD76FCB0" box="[199,720,764,791]" pageId="2" type="distribution">
|
||
<paragraph id="8BD936C9697DFFE11592AAF0FD76FCB0" blockId="2.[151,1437,152,1367]" box="[199,720,764,791]" pageId="2">
|
||
<emphasis id="B912EADB697DFFE11592AAF0FEC7FCB0" bold="true" box="[199,353,764,791]" pageId="2">Distribution.</emphasis>
|
||
<collectingCountry id="F3717659697DFFE11432AAF1FE0CFCB0" box="[359,426,765,791]" name="Japan" pageId="2">Japan</collectingCountry>
|
||
, central Honshu (
|
||
<figureCitation id="135D2A4C697DFFE1173AAAF1FD65FCB0" box="[623,707,764,791]" captionStart="FIGURE 25" captionStartId="9.[151,250,1702,1727]" captionTargetBox="[151,1436,696,1668]" captionTargetId="figure@9.[151,1436,692,1683]" captionTargetPageId="9" captionText="FIGURE 25. Distribution of Veraphis in Japan." figureDoi="http://doi.org/10.5281/zenodo.3557835" httpUri="https://zenodo.org/record/3557835/files/figure.png" pageId="2">Fig. 25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C6542697DFFE11592AB2DFB1BFC9C" box="[199,1213,801,827]" pageId="2" type="etymology">
|
||
<paragraph id="8BD936C9697DFFE11592AB2DFB1BFC9C" blockId="2.[151,1437,152,1367]" box="[199,1213,801,827]" pageId="2">
|
||
<emphasis id="B912EADB697DFFE11592AB2DFEE8FC9C" bold="true" box="[199,334,801,827]" pageId="2">Etymology.</emphasis>
|
||
This species is dedicated to Ivan Löbl, the collector of the
|
||
<typeStatus id="54DD886B697DFFE116B6AB2DFBE3FC9C" box="[995,1093,801,827]" pageId="2" type="holotype">holotype</typeStatus>
|
||
specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C6542697DFFE11592AB49FC95FAF0" pageId="2" type="discussion">
|
||
<paragraph id="8BD936C9697DFFE11592AB49FC95FAF0" blockId="2.[151,1437,152,1367]" pageId="2">
|
||
<emphasis id="B912EADB697DFFE11592AB49FE9DFCF8" bold="true" box="[199,315,837,863]" pageId="2">Remarks.</emphasis>
|
||
<taxonomicName id="4C664D4A697DFFE11411AB49FE56FCF8" authorityName="Jałoszyński" authorityYear="2019" box="[324,496,837,863]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B912EADB697DFFE11411AB49FE56FCF8" box="[324,496,837,863]" italics="true" pageId="2">Veraphis loebli</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C664D4A697DFFE117F1AB49FCA7FCF8" authorityName="Jaloszynski & Hoshina" authorityYear="2005" box="[676,769,837,863]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="sawadai">
|
||
<emphasis id="B912EADB697DFFE117F1AB49FCA7FCF8" box="[676,769,837,863]" italics="true" pageId="2">sawadai</emphasis>
|
||
</taxonomicName>
|
||
group of species (
|
||
<bibRefCitation id="EFF74B38697DFFE1168DAB49FA80FCF8" author="Jaloszynski, P. & Hoshina, H." box="[984,1318,836,863]" pageId="2" pagination="101 - 136" refId="ref5900" refString="Jaloszynski, P. & Hoshina, H. (2005) Revision of Palearctic species of Veraphis Casey (Coleoptera, Scydmaenidae). Entomological Review of Japan, 60, 101 - 136." type="journal article" year="2005">Jałoszyński & Hoshina 2005</bibRefCitation>
|
||
), which is characterized by an unmodified antennomere XI in both sexes, unmodified male protrochanters, male protibiae with a subapical fin-like projection, the metaventrite in males with a longitudinal median impression, and the ae- deagus lacking apical setae, in most species with an apical emargination of the median lobe (with one exception,
|
||
<taxonomicName id="4C664D4A697DFFE115C2ABDAFE9AFC48" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,316,982,1007]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="aomoriensis" status="sp. nov.">
|
||
<emphasis id="B912EADB697DFFE115C2ABDAFE9AFC48" box="[151,316,982,1007]" italics="true" pageId="2">V. aomoriensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912EADB697DFFE11416ABD9FE25FC48" bold="true" box="[323,387,981,1007]" pageId="2">
|
||
<taxonomicNameLabel id="A22157A0697DFFE11416ABD9FE25FC48" box="[323,387,981,1007]" pageId="2" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
described below). This group includes
|
||
<taxonomicName id="4C664D4A697DFFE1166BABDAFAACFC48" authority="Jaloszynski & Hoshina, 2005" authorityName="Jaloszynski & Hoshina" authorityYear="2005" box="[830,1290,980,1007]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="sawadai">
|
||
<emphasis id="B912EADB697DFFE1166BABDAFC10FC48" box="[830,950,981,1007]" italics="true" pageId="2">V. sawadai</emphasis>
|
||
<bibRefCitation id="EFF74B38697DFFE116EBABD9FAACFC48" author="Jaloszynski, P. & Hoshina, H." box="[958,1290,980,1007]" pageId="2" pagination="101 - 136" refId="ref5900" refString="Jaloszynski, P. & Hoshina, H. (2005) Revision of Palearctic species of Veraphis Casey (Coleoptera, Scydmaenidae). Entomological Review of Japan, 60, 101 - 136." type="journal article" year="2005">Jałoszyński & Hoshina, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C664D4A697DFFE11040ABDAFE47FBB4" authority="Jaloszynski & Hoshina, 2005" authorityName="Jaloszynski & Hoshina" authorityYear="2005" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="kurbatovi">
|
||
<emphasis id="B912EADB697DFFE11040ABDAFA3AFC48" box="[1301,1436,981,1007]" italics="true" pageId="2">V. kurbatovi</emphasis>
|
||
<bibRefCitation id="EFF74B38697DFFE115C2ABF5FE47FBB4" author="Jaloszynski, P. & Hoshina, H." box="[151,481,1016,1043]" pageId="2" pagination="101 - 136" refId="ref5900" refString="Jaloszynski, P. & Hoshina, H. (2005) Revision of Palearctic species of Veraphis Casey (Coleoptera, Scydmaenidae). Entomological Review of Japan, 60, 101 - 136." type="journal article" year="2005">Jałoszyński & Hoshina, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, and three species described in the present paper:
|
||
<taxonomicName id="4C664D4A697DFFE11159ABF6FBC1FBB4" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[1036,1127,1017,1043]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
|
||
<emphasis id="B912EADB697DFFE11159ABF6FBC1FBB4" box="[1036,1127,1017,1043]" italics="true" pageId="2">V. loebli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912EADB697DFFE1113BABF5FB0BFBB4" bold="true" box="[1134,1197,1017,1043]" pageId="2">
|
||
<taxonomicNameLabel id="A22157A0697DFFE1113BABF5FB0BFBB4" box="[1134,1197,1017,1043]" pageId="2" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C664D4A697DFFE111EFABF6FAF6FBB4" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[1210,1360,1018,1043]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="mutsuensis" status="sp. nov.">
|
||
<emphasis id="B912EADB697DFFE111EFABF6FAF6FBB4" box="[1210,1360,1018,1043]" italics="true" pageId="2">V. mutsuensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912EADB697DFFE11002ABF5FA30FBB4" bold="true" box="[1367,1430,1017,1043]" pageId="2">
|
||
<taxonomicNameLabel id="A22157A0697DFFE11002ABF5FA30FBB4" box="[1367,1430,1017,1043]" pageId="2" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C664D4A697DFFE11590AC12FECFFB90" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[197,361,1054,1079]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="aomoriensis" status="sp. nov.">
|
||
<emphasis id="B912EADB697DFFE11590AC12FECFFB90" box="[197,361,1054,1079]" italics="true" pageId="2">V. aomoriensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912EADB697DFFE1143AAC11FE0BFB90" bold="true" box="[367,429,1053,1079]" pageId="2">
|
||
<taxonomicNameLabel id="A22157A0697DFFE1143AAC11FE0BFB90" box="[367,429,1053,1079]" pageId="2" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
External differences between males of these species are rather elusive; the body length and proportions of body parts slightly differ, but only examination of the aedeagus allows for unambiguous determination. The aedeagus of
|
||
<taxonomicName id="4C664D4A697DFFE114DFAC6AFE42FBD8" authorityName="Jałoszyński" authorityYear="2019" box="[394,484,1125,1151]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B912EADB697DFFE114DFAC6AFE42FBD8" box="[394,484,1125,1151]" italics="true" pageId="2">V. loebli</emphasis>
|
||
</taxonomicName>
|
||
differs from those of
|
||
<taxonomicName id="4C664D4A697DFFE11787AC6AFCEFFBD8" authorityName="Jaloszynski & Hoshina" authorityYear="2005" box="[722,841,1125,1151]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="sawadai">
|
||
<emphasis id="B912EADB697DFFE11787AC6AFCEFFBD8" box="[722,841,1125,1151]" italics="true" pageId="2">V. sawadai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C664D4A697DFFE11628AC6AFBA4FBD8" authorityName="Jaloszynski & Hoshina" authorityYear="2005" box="[893,1026,1125,1151]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="kurbatovi">
|
||
<emphasis id="B912EADB697DFFE11628AC6AFBA4FBD8" box="[893,1026,1125,1151]" italics="true" pageId="2">V. kurbatovi</emphasis>
|
||
</taxonomicName>
|
||
in (among others) distinctly concave lateral margins in the subapical region (convex in
|
||
<taxonomicName id="4C664D4A697DFFE11797AC86FC9FFB04" authorityName="Jaloszynski & Hoshina" authorityYear="2005" box="[706,825,1161,1187]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="sawadai">
|
||
<emphasis id="B912EADB697DFFE11797AC86FC9FFB04" box="[706,825,1161,1187]" italics="true" pageId="2">V. sawadai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C664D4A697DFFE1163BAC86FC52FB04" authorityName="Jaloszynski & Hoshina" authorityYear="2005" box="[878,1012,1161,1187]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="kurbatovi">
|
||
<emphasis id="B912EADB697DFFE1163BAC86FC52FB04" box="[878,1012,1161,1187]" italics="true" pageId="2">V. kurbatovi</emphasis>
|
||
</taxonomicName>
|
||
). This character, however, is the same in
|
||
<taxonomicName id="4C664D4A697DFFE115E1ACA2FEA9FB60" authorityName="Jałoszyński" authorityYear="2019" box="[180,271,1197,1223]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B912EADB697DFFE115E1ACA2FEA9FB60" box="[180,271,1197,1223]" italics="true" pageId="2">V. loebli</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C664D4A697DFFE11449ACA2FE14FB60" authorityName="Jałoszyński" authorityYear="2019" box="[284,434,1198,1223]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="mutsuensis">
|
||
<emphasis id="B912EADB697DFFE11449ACA2FE14FB60" box="[284,434,1198,1223]" italics="true" pageId="2">V. mutsuensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C664D4A697DFFE114BDACA2FD2AFB60" authorityName="Jałoszyński" authorityYear="2019" box="[488,652,1198,1223]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="aomoriensis">
|
||
<emphasis id="B912EADB697DFFE114BDACA2FD2AFB60" box="[488,652,1198,1223]" italics="true" pageId="2">V. aomoriensis</emphasis>
|
||
</taxonomicName>
|
||
. Among these three species,
|
||
<taxonomicName id="4C664D4A697DFFE1169AACA2FBD5FB60" authorityName="Jałoszyński" authorityYear="2019" box="[975,1139,1198,1223]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="aomoriensis">
|
||
<emphasis id="B912EADB697DFFE1169AACA2FBD5FB60" box="[975,1139,1198,1223]" italics="true" pageId="2">V. aomoriensis</emphasis>
|
||
</taxonomicName>
|
||
is most easily distinguishable by a rounded, and not emarginate, apex of the median lobe. The aedeagi of
|
||
<taxonomicName id="4C664D4A697DFFE11144ACDEFBCAFB4C" authorityName="Jałoszyński" authorityYear="2019" box="[1041,1132,1233,1259]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B912EADB697DFFE11144ACDEFBCAFB4C" box="[1041,1132,1233,1259]" italics="true" pageId="2">V. loebli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C664D4A697DFFE111F4ACDEFA91FB4C" authorityName="Jałoszyński" authorityYear="2019" box="[1185,1335,1234,1259]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="mutsuensis">
|
||
<emphasis id="B912EADB697DFFE111F4ACDEFA91FB4C" box="[1185,1335,1234,1259]" italics="true" pageId="2">V. mutsuensis</emphasis>
|
||
</taxonomicName>
|
||
are most similar; the aedeagus of
|
||
<taxonomicName id="4C664D4A697DFFE114FBACFAFDACFAA8" authorityName="Jałoszyński" authorityYear="2019" box="[430,522,1269,1295]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B912EADB697DFFE114FBACFAFDACFAA8" box="[430,522,1269,1295]" italics="true" pageId="2">V. loebli</emphasis>
|
||
</taxonomicName>
|
||
is more curved in lateral view; it has a much smaller ventral diaphragm that oc- cupies the sub-basal 1/3 of the ventral wall (nearly half in
|
||
<taxonomicName id="4C664D4A697DFFE11676AD16FC1FFA94" authorityName="Jałoszyński" authorityYear="2019" box="[803,953,1306,1331]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="mutsuensis">
|
||
<emphasis id="B912EADB697DFFE11676AD16FC1FFA94" box="[803,953,1306,1331]" italics="true" pageId="2">V. mutsuensis</emphasis>
|
||
</taxonomicName>
|
||
); and the apical portion of the median lobe in ventral view is longer, with more strongly concave sides.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |