312 lines
35 KiB
XML
312 lines
35 KiB
XML
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<mods:title id="F9CF4424514E830B5CDDDA28AD632D6F">Zebrus pallaoroi sp. nov.: a new species of goby (Actinopterygii: Gobiidae) from the Mediterranean Sea with a DNA-based phylogenetic analysis of the Gobius-lineage</mods:title>
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<mods:namePart id="F6CB131FB999EC59FA8AEE36DCE579CB">Cooper, David</mods:namePart>
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<mods:namePart id="3602C71CBF0345ADD10CBEE230A99A3A">Yamaguchi, Nobuyuki</mods:namePart>
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<mods:namePart id="CE7B0AC210195C7E35B6561C2BFAC197">Macdonald, David</mods:namePart>
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<mods:namePart id="A2E0CC60D58BD1957C3123306441B441">Nanova, Olga</mods:namePart>
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<mods:namePart id="6EE4FCA90360975837FAAB197356A106">Yudin, Viktor</mods:namePart>
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<mods:namePart id="2DA2326B2B265D1466E76DE57ACEB711">Dugmore, Andrew</mods:namePart>
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<mods:namePart id="5900B3494F26674504A68107B852F8CE">Kitchener, Andrew</mods:namePart>
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<mods:title id="8212617A369354E287405EB2AAEFCBD7">Contributions to Zoology</mods:title>
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<mods:date id="490EDB4A4C8D20381BFB03C5FBF15F14">2021</mods:date>
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<mods:number id="C27A0E7BC2BF51A7C49D6A384967C5DE">2021-03-30</mods:number>
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<mods:identifier id="6F387944FF57EC64405AA381357CDAE0" type="DOI">10.1163/18759866-bja10018</mods:identifier>
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<mods:identifier id="304DAED2E85C1597F1CC1BB135CFEEE1" type="ISSN">1875-9866</mods:identifier>
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<treatment id="03CF87C0FFEBFFB6FF459C3B8A7FFED7" LSID="urn:lsid:plazi:treatment:03CF87C0FFEBFFB6FF459C3B8A7FFED7" httpUri="http://treatment.plazi.org/id/03CF87C0FFEBFFB6FF459C3B8A7FFED7" lastPageId="17" lastPageNumber="302" pageId="16" pageNumber="301">
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<subSubSection id="C37C655DFFEBFFB7FF459C3B8F21FBBB" pageId="16" pageNumber="301" type="nomenclature">
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<paragraph id="8BD936D6FFEBFFB7FF459C3B8F21FBBB" blockId="16.[129,507,1033,1095]" pageId="16" pageNumber="301">
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<emphasis id="B912EAC4FFEBFFB7FF459C3B8FABFBD8" bold="true" box="[129,507,1033,1060]" pageId="16" pageNumber="301">
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The designation of
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<taxonomicName id="4C664D55FFEBFFB7FE999C3B8F5DFBBB" ID-CoL="7GD2B" ID-ENA="85426" authority="(Risso, 1827)" baseAuthorityName="Risso" baseAuthorityYear="1827" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
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<emphasis id="B912EAC4FFEBFFB7FE999C3B8FABFBD8" bold="true" box="[349,507,1033,1060]" italics="true" pageId="16" pageNumber="301">Zebrus zebrus</emphasis>
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<emphasis id="B912EAC4FFEBFFB7FF459C1F8F21FBBB" bold="true" box="[129,369,1069,1095]" pageId="16" pageNumber="301">
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(Risso, 1827)
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<typeStatus id="54DD8874FFEBFFB7FED69C1F8F21FBBB" box="[274,369,1069,1095]" pageId="16" pageNumber="301">neotype</typeStatus>
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</emphasis>
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</taxonomicName>
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</emphasis>
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</paragraph>
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<subSubSection id="C37C655DFFEBFFB7FF459C638A7FF9C2" pageId="16" pageNumber="301" type="description">
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<paragraph id="8BD936D6FFEBFFB7FF459C638A7FF9C2" blockId="16.[129,640,1105,1742]" lastBlockId="16.[676,1188,171,1748]" pageId="16" pageNumber="301">
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<emphasis id="B912EAC4FFEBFFB7FF459C638EA0FB97" box="[129,240,1105,1131]" italics="true" pageId="16" pageNumber="301">Rationale.</emphasis>
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Based on morphological and genetic data presented here, the genus
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<taxonomicName id="4C664D55FFEBFFB7FF459CAB8E9CFB4F" authorityName="de Buen" authorityYear="1930" box="[129,204,1177,1203]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="genus">
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<emphasis id="B912EAC4FFEBFFB7FF459CAB8E9CFB4F" box="[129,204,1177,1203]" italics="true" pageId="16" pageNumber="301">Zebrus</emphasis>
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</taxonomicName>
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includes two species. Until now, the genus was considered monotypic (
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<bibRefCitation id="EFF74B27FFEBFFB7FDF29C8F8EE5FB07" author="Miller, P. J." pageId="16" pageNumber="301" pagination="339 - 362" refId="ref16111" refString="Miller, P. J. (1977) Gobies from Rhodes and systematic features of Zebrus zebrus (Teleostei: Gobiidae). Zool. J. Linn. Soc., 60, 339 - 362." type="journal article" year="1977">Miller, 1977</bibRefCitation>
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). The original description of
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<taxonomicName id="4C664D55FFEBFFB7FDF19CD28E99FAE2" baseAuthorityName="Risso" baseAuthorityYear="1827" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
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<emphasis id="B912EAC4FFEBFFB7FDF19CD28E99FAE2" italics="true" pageId="16" pageNumber="301">Zebrus zebrus</emphasis>
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</taxonomicName>
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(decribed as
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<taxonomicName id="4C664D55FFEBFFB7FEA99D368C5BFAE2" authorityName="Risso" authorityYear="1827" box="[365,523,1284,1310]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
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<emphasis id="B912EAC4FFEBFFB7FEA99D368C5BFAE2" box="[365,523,1284,1310]" italics="true" pageId="16" pageNumber="301">Gobius zebrus</emphasis>
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</taxonomicName>
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) by Risso (1827) is only general, and does not include any morphological characteristics which could unambiguosly determine which of the two currently recognised species was included under the name. Furthermore, no original type material exists (
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<bibRefCitation id="EFF74B27FFEBFFB7FE229DEE8E98F9E7" author="Fricke, R. & Eschmeyer, W. N. & Van der Laan, R." pageId="16" pageNumber="301" refId="ref14927" refString="Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds) (2020 b) Eschmeyer's catalog of fishes: Genera, species, references. (http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp). Electronic version accessed 20 October 2020." type="url" year="2020">Fricke et al., 2020b</bibRefCitation>
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) to prove the identity of
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<taxonomicName id="4C664D55FFEBFFB7FE319E338C0BF9E6" authorityName="Risso" authorityYear="1827" box="[501,603,1536,1562]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
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<emphasis id="B912EAC4FFEBFFB7FE319E338C0BF9E6" box="[501,603,1536,1562]" italics="true" pageId="16" pageNumber="301">Z. zebrus</emphasis>
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</taxonomicName>
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by type inspection. To resolve this taxonomic problem, we here designate a
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<typeStatus id="54DD8874FFEBFFB7FE0B9E7A8C7AF99E" box="[463,554,1608,1634]" pageId="16" pageNumber="301">neotype</typeStatus>
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for this species. The designated
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<typeStatus id="54DD8874FFEBFFB7FE549E5E8FBBF97A" box="[400,491,1644,1670]" pageId="16" pageNumber="301">neotype</typeStatus>
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was selected from available specimens analysed in this study to originate as close to the
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<emphasis id="B912EAC4FFEBFFB7FE309E818C2FF931" box="[500,639,1715,1741]" italics="true" pageId="16" pageNumber="301">locus typicus</emphasis>
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(Nice,
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<collectingCountry id="F3717646FFEBFFB7FD2F989E8D69FF3A" box="[747,825,172,198]" name="France" pageId="16" pageNumber="301">France</collectingCountry>
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) as possible, and is accessible by the voucher number nmp P6V 140912 at the nmp. The
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<typeStatus id="54DD8874FFEBFFB7FCD098C68D3FFEF2" box="[788,879,244,270]" pageId="16" pageNumber="301">neotype</typeStatus>
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of
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<taxonomicName id="4C664D55FFEBFFB7FC5698C68DA4FEF1" authorityName="Risso" authorityYear="1827" box="[914,1012,243,269]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
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<emphasis id="B912EAC4FFEBFFB7FC5698C68DA4FEF1" box="[914,1012,243,269]" italics="true" pageId="16" pageNumber="301">Z. zebrus</emphasis>
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</taxonomicName>
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was designated fulfilling the qualifying conditions set by iczn (1999) as follows: The description of the second species of
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<taxonomicName id="4C664D55FFEBFFB7FC61996D8DA0FE85" authorityName="de Buen" authorityYear="1930" box="[933,1008,351,377]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="genus">
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<emphasis id="B912EAC4FFEBFFB7FC61996D8DA0FE85" box="[933,1008,351,377]" italics="true" pageId="16" pageNumber="301">Zebrus</emphasis>
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</taxonomicName>
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in the hitherto monotypic genus represented an exceptional need for the designation of a
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<typeStatus id="54DD8874FFEBFFB7FC3099958A1FFE3D" box="[1012,1103,423,449]" pageId="16" pageNumber="301">neotype</typeStatus>
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for the first
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<taxonomicName id="4C664D55FFEBFFB7FD1C99F88D2CFE18" box="[728,892,458,484]" pageId="16" pageNumber="301">
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<emphasis id="B912EAC4FFEBFFB7FD1C99F88D73FE18" box="[728,803,458,484]" italics="true" pageId="16" pageNumber="301">Zebrus</emphasis>
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species
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</taxonomicName>
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(Article 75.3, iczn, 1999). The genus
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<taxonomicName id="4C664D55FFEBFFB7FCEB99DC8D2AFDF4" authorityName="de Buen" authorityYear="1930" box="[815,890,494,520]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="genus">
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<emphasis id="B912EAC4FFEBFFB7FCEB99DC8D2AFDF4" box="[815,890,494,520]" italics="true" pageId="16" pageNumber="301">Zebrus</emphasis>
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</taxonomicName>
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represents the complex zoological problem (Article 75.2, iczn, 1999) of morphologically very similar congenerics. Therefore, in addition to the diagnosis, the designation of a
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<typeStatus id="54DD8874FFEBFFB7FCA89A4D8D97FD65" box="[876,967,639,665]" pageId="16" pageNumber="301">neotype</typeStatus>
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is needed to have the name-bearing material of this species in addition to the type material of the new species. This complex zoological problem (Article 75.2, iczn, 1999) could become even more complicated if more lineages within the
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<emphasis id="B912EAC4FFEBFFB7FD609B678DD9FC93" box="[676,905,853,879]" italics="true" pageId="16" pageNumber="301">
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<taxonomicName id="4C664D55FFEBFFB7FD609B678CBEFC93" authorityName="de Buen" authorityYear="1930" box="[676,750,853,879]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="genus">Zebrus</taxonomicName>
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/
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<taxonomicName id="4C664D55FFEBFFB7FD3F9B678DD9FC93" box="[763,905,853,879]" family="Gobiidae" genus="Millerigobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="genus">Millerigobius</taxonomicName>
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</emphasis>
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clade would be found. In that case, the name-bearing material of the present species,
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<typeStatus id="54DD8874FFEBFFB7FC5E9BAF8DAEFC4B" box="[922,1022,925,951]" pageId="16" pageNumber="301">holotype</typeStatus>
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and
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<typeStatus id="54DD8874FFEBFFB7FB869BAC8ACFFC44" box="[1090,1183,926,952]" pageId="16" pageNumber="301">neotype</typeStatus>
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, should be available for comparison with the new material. The purpose was to clarify the taxonomic status of two species (Article 75.3.1, iczn, 1999). The diagnostic characters are stated in the diagnosis (Article 75.3.1, iczn, 1999), which is sufficient to ensure the recognition of the species (Article 75.3.3, iczn, 1999). We are positive that no name bearing type specimens exist for
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<taxonomicName id="4C664D55FFEBFFB7FC1F9CD38A16FB06" authorityName="Risso" authorityYear="1827" box="[987,1094,1248,1274]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
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<emphasis id="B912EAC4FFEBFFB7FC1F9CD38A16FB06" box="[987,1094,1248,1274]" italics="true" pageId="16" pageNumber="301">Z. zebrus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EFF74B27FFEBFFB7FB9F9CD28D63FAE2" author="Fricke, R. & Eschmeyer, W. N. & Van der Laan, R." pageId="16" pageNumber="301" refId="ref14927" refString="Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds) (2020 b) Eschmeyer's catalog of fishes: Genera, species, references. (http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp). Electronic version accessed 20 October 2020." type="url" year="2020">Fricke et al., 2020b</bibRefCitation>
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||
; Article 75.3.4, iczn, 1999). The
|
||
<typeStatus id="54DD8874FFEBFFB7FD609D1B8CAFFABF" box="[676,767,1321,1347]" pageId="16" pageNumber="301">neotype</typeStatus>
|
||
fits the species redescription of
|
||
<taxonomicName id="4C664D55FFEBFFB7FB4A9D1B8CBCFA9A" authorityName="Risso" authorityYear="1827" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
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||
<emphasis id="B912EAC4FFEBFFB7FB4A9D1B8CBCFA9A" italics="true" pageId="16" pageNumber="301">Z. zebrus</emphasis>
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||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFF74B27FFEBFFB7FD3D9D7E8D2BFA9A" author="Miller, P. J." box="[761,891,1356,1382]" pageId="16" pageNumber="301" pagination="339 - 362" refId="ref16111" refString="Miller, P. J. (1977) Gobies from Rhodes and systematic features of Zebrus zebrus (Teleostei: Gobiidae). Zool. J. Linn. Soc., 60, 339 - 362." type="journal article" year="1977">Miller, 1977</bibRefCitation>
|
||
; Article 75.3.5, iczn, 1999) and the
|
||
<typeStatus id="54DD8874FFEBFFB7FD3B9D438D0AFA77" box="[767,858,1393,1419]" pageId="16" pageNumber="301">neotype</typeStatus>
|
||
was collected relatively close to the original type locality (see locality of the
|
||
<typeStatus id="54DD8874FFEBFFB7FD609D8A8CAFFA2E" box="[676,767,1464,1490]" pageId="16" pageNumber="301">neotype</typeStatus>
|
||
material, Article 75.3.6, iczn, 1999). The
|
||
<typeStatus id="54DD8874FFEBFFB7FD139DEE8D62FA0A" box="[727,818,1500,1526]" pageId="16" pageNumber="301">neotype</typeStatus>
|
||
was stored in a public scientific museum collection (see storage of the
|
||
<typeStatus id="54DD8874FFEBFFB7FB839E328AF2F9E6" box="[1095,1186,1536,1562]" pageId="16" pageNumber="301">neotype</typeStatus>
|
||
material, Article 75.3.7, iczn, 1999).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C655DFFEBFFB6FD069E7A8F5AFB4F" lastPageId="17" lastPageNumber="302" pageId="16" pageNumber="301" type="materials_examined">
|
||
<paragraph id="8BD936D6FFEBFFB7FD069E7A8DAAF932" blockId="16.[676,1188,171,1748]" pageId="16" pageNumber="301">
|
||
<materialsCitation id="3B0E3C8BFFEBFFB7FD069E7A8DAAF932" collectingDate="2011-10-14" collectionCode="R" country="France" county="Sanda" latitude="42.482105" location="Ill Gross" longLatPrecision="1" longitude="3.1369333" municipality="Banyuls sur Mer" pageId="16" pageNumber="301" specimenCode="V 140912" specimenCount="1" typeStatus="neotype">
|
||
<emphasis id="B912EAC4FFEBFFB7FD069E7A8D71F99D" box="[706,801,1608,1633]" italics="true" pageId="16" pageNumber="301">
|
||
<typeStatus id="54DD8874FFEBFFB7FD069E7A8D4CF99D" box="[706,796,1608,1633]" pageId="16" pageNumber="301">Neotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
nmp P6
|
||
<specimenCode id="DBC09EADFFEBFFB7FC749E758A7DF99E" box="[944,1069,1607,1634]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="301" type="Museum">V 140912</specimenCode>
|
||
, female, 23.25 +
|
||
<quantity id="4C9E9B33FFEBFFB7FD3B9E598D30F97A" box="[767,864,1643,1670]" metricMagnitude="-3" metricUnit="m" metricValue="6.22" pageId="16" pageNumber="301" unit="mm" value="6.22">6.22 mm</quantity>
|
||
,
|
||
<location id="8EB9600DFFEBFFB7FCAB9E5C8D81F97A" LSID="urn:lsid:plazi:treatment:03CF87C0FFEBFFB6FF459C3B8A7FFED7:8EB9600DFFEBFFB7FCAB9E5C8D81F97A" box="[879,977,1644,1677]" country="France" county="Sanda" latitude="42.482105" longLatPrecision="1" longitude="3.1369333" municipality="Banyuls sur Mer" name="Ill Gross" pageId="16" pageNumber="301">Îll Gross</location>
|
||
,
|
||
<collectingMunicipality id="6BBDACACFFEBFFB7FC249E598ACFF97A" box="[992,1183,1643,1670]" pageId="16" pageNumber="301">Banyuls sur Mer</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F3717646FFEBFFB7FD609EA28CA2F956" box="[676,754,1680,1706]" name="France" pageId="16" pageNumber="301">France</collectingCountry>
|
||
;
|
||
<geoCoordinate id="EE525011FFEBFFB7FCCE9EBD8DF3F956" box="[778,931,1679,1706]" degrees="42" direction="north" minutes="28" orientation="latitude" pageId="16" pageNumber="301" precision="1" seconds="55.58" value="42.482105">42°28’55.58”N</geoCoordinate>
|
||
<geoCoordinate id="EE525011FFEBFFB7FC739EBD8A62F955" box="[951,1074,1679,1705]" degrees="3" direction="east" minutes="8" orientation="longitude" pageId="16" pageNumber="301" precision="1" seconds="12.96" value="3.1369333">3°8’12.96”E</geoCoordinate>
|
||
, coll.
|
||
<collectionCode id="ED77AE13FFEBFFB7FB499EA28AF0F956" box="[1165,1184,1680,1706]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="301">R</collectionCode>
|
||
.
|
||
<collectingCounty id="62B84E5AFFEBFFB7FD609E878CBAF928" box="[676,746,1717,1748]" pageId="16" pageNumber="301">Šanda</collectingCounty>
|
||
,
|
||
<date id="FFD81016FFEBFFB7FD309E818DCCF931" box="[756,924,1715,1741]" pageId="16" pageNumber="301" value="2011-10-14">
|
||
<collectingDate id="EF9CE9FEFFEBFFB7FD309E818DCCF931" box="[756,924,1715,1741]" pageId="16" pageNumber="301" value="2011-10-14">14 October 2011</collectingDate>
|
||
</date>
|
||
(
|
||
<figureCitation id="135D2A53FFEBFFB7FC6E9E818DBDF932" box="[938,1005,1715,1742]" captionStart="figure 1" captionStartId="10.[128,195,1556,1578]" captionTargetBox="[223,1092,575,1537]" captionTargetId="figure-3@10.[223,1092,575,1538]" captionTargetPageId="10" captionText="figure 1 Preserved specimens.(A) Zebrus pallaoroi sp.nov., npm P6V144302, holotype, male, 31.81 + 8.51 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro. Photo by M.Kovačić. (B) Zebrus zebrus, nmp P6V 140912,neotype, female, 23.25 + 6.22mm, Îll Gross, Banyuls sur Mer, France (C) Millerigobius macrocephalus, nmp P6V 142686, juvenile of unidentified sex, 14.28 + 3.97 mm, Îll Gross, Banyuls sur Mer, France." figureDoi="http://doi.org/10.5281/zenodo.8343385" httpUri="https://zenodo.org/record/8343385/files/figure.png" pageId="16" pageNumber="301">fig. 1b</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD936D6FFEAFFB6FF4598998F5AFB4F" blockId="17.[98,610,171,1706]" pageId="17" pageNumber="302">
|
||
<emphasis id="B912EAC4FFEAFFB6FF4598998FB1FF39" box="[129,481,171,197]" italics="true" pageId="17" pageNumber="302">Additional material examined.</emphasis>
|
||
nmp P6V 142530, female, 25.21 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEBF98FD8F88FF16" box="[379,472,207,234]" metricMagnitude="-3" metricUnit="m" metricValue="6.18" pageId="17" pageNumber="302" unit="mm" value="6.18">6.18 mm</quantity>
|
||
, Sv. Marak, Krk Island,
|
||
<collectingCountry id="F3717646FFEAFFB6FEC898C18F0FFEF1" box="[268,351,243,269]" name="Croatia" pageId="17" pageNumber="302">Croatia</collectingCountry>
|
||
; nmp P6V 142592, female, 21.9 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEDA99258F2EFECE" box="[286,382,279,306]" metricMagnitude="-3" metricUnit="m" metricValue="5.58" pageId="17" pageNumber="302" unit="mm" value="5.58">5.58 mm</quantity>
|
||
, nmp P6V 142593, male, 21.84 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEE499098F23FEAA" box="[288,371,315,342]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="17" pageNumber="302" unit="mm" value="5.6">5.6 mm</quantity>
|
||
, nmp P6V 142594, male, 18.76 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEDE996D8F2AFE85" box="[282,378,351,377]" metricMagnitude="-3" metricUnit="m" metricValue="4.76" pageId="17" pageNumber="302" unit="mm" value="4.76">4.76 mm</quantity>
|
||
, nmp P6V 142595, female, 19.59 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEE799B18FD4FE61" box="[291,388,387,414]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" pageId="17" pageNumber="302" unit="mm" value="4.95">4.95 mm</quantity>
|
||
, nmp P6V 142596, female, 19.39 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEE099958FD4FE3D" box="[292,388,423,450]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" pageId="17" pageNumber="302" unit="mm" value="4.85">4.85 mm</quantity>
|
||
, nmp P6V 142597, male, 21.16 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEDC99F88F27FE19" box="[280,375,458,485]" metricMagnitude="-3" metricUnit="m" metricValue="5.58" pageId="17" pageNumber="302" unit="mm" value="5.58">5.58 mm</quantity>
|
||
, nmp P6V 142598, female, 19.83 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEE199DD8FD2FDF5" box="[293,386,495,521]" metricMagnitude="-3" metricUnit="m" metricValue="4.81" pageId="17" pageNumber="302" unit="mm" value="4.81">4.81 mm</quantity>
|
||
, nmp P6V 142599, male, 21.91 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEC09A208F34FDD1" box="[260,356,530,557]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" pageId="17" pageNumber="302" unit="mm" value="5.65">5.65 mm</quantity>
|
||
, all eight from Selce, Kvarner area,
|
||
<collectingCountry id="F3717646FFEAFFB6FEC69A048F05FDAC" box="[258,341,566,592]" name="Croatia" pageId="17" pageNumber="302">Croatia</collectingCountry>
|
||
; nmp P6V 145212, male, 27.38 +
|
||
<quantity id="4C9E9B33FFEAFFB6FF039A688F78FD89" box="[199,296,602,629]" metricMagnitude="-3" metricUnit="m" metricValue="6.84" pageId="17" pageNumber="302" unit="mm" value="6.84">6.84 mm</quantity>
|
||
, nmp P6V 145213, female, 26.48 +
|
||
<quantity id="4C9E9B33FFEAFFB6FF0C9A4C8F7AFD65" box="[200,298,638,665]" metricMagnitude="-3" metricUnit="m" metricValue="6.44" pageId="17" pageNumber="302" unit="mm" value="6.44">6.44 mm</quantity>
|
||
, nmp P6V 145214, female, 25.99 +
|
||
<quantity id="4C9E9B33FFEAFFB6FF0B9A908F61FD41" box="[207,305,674,701]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" pageId="17" pageNumber="302" unit="mm" value="6.05">6.05 mm</quantity>
|
||
, nmp P6V 145215, male, 27.11 +
|
||
<quantity id="4C9E9B33FFEAFFB6FF0C9AF48F79FD1D" box="[200,297,710,737]" metricMagnitude="-3" metricUnit="m" metricValue="6.45" pageId="17" pageNumber="302" unit="mm" value="6.45">6.45 mm</quantity>
|
||
, nmp P6V 145216, male, 27.17 +
|
||
<quantity id="4C9E9B33FFEAFFB6FF0F9AD88F7CFCF8" box="[203,300,746,773]" metricMagnitude="-3" metricUnit="m" metricValue="6.59" pageId="17" pageNumber="302" unit="mm" value="6.59">6.59 mm</quantity>
|
||
, nmp P6V 145217, male, 27.8+7.17,nmpP6V145218,male,27.52+
|
||
<quantity id="4C9E9B33FFEAFFB6FDC79B3C8C0DFCD4" box="[515,605,782,808]" metricMagnitude="-3" metricUnit="m" metricValue="7.17" pageId="17" pageNumber="302" unit="mm" value="7.17">7.17 mm</quantity>
|
||
, nmp P6V 145219, male, 25.55 +
|
||
<quantity id="4C9E9B33FFEAFFB6FE069B008C73FCB0" box="[450,547,818,844]" metricMagnitude="-3" metricUnit="m" metricValue="6.03" pageId="17" pageNumber="302" unit="mm" value="6.03">6.03 mm</quantity>
|
||
, nmp P6V 145220, male, 27.57 +
|
||
<quantity id="4C9E9B33FFEAFFB6FE5D9B648FA9FC8C" box="[409,505,854,880]" metricMagnitude="-3" metricUnit="m" metricValue="7.03" pageId="17" pageNumber="302" unit="mm" value="7.03">7.03 mm</quantity>
|
||
, all nine from Kamenjak, Krk Island,
|
||
<collectingCountry id="F3717646FFEAFFB6FE039B4B8C4AFC6F" box="[455,538,889,915]" name="Croatia" pageId="17" pageNumber="302">Croatia</collectingCountry>
|
||
; nmp P6V 145287, female, 27.36 +
|
||
<quantity id="4C9E9B33FFEAFFB6FE039BAF8C4BFC44" box="[455,539,925,952]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="17" pageNumber="302" unit="mm" value="6.4">6.4 mm</quantity>
|
||
, nmp P6V 145288, male, 25.73 +
|
||
<quantity id="4C9E9B33FFEAFFB6FE489BF38FBEFC20" box="[396,494,961,988]" metricMagnitude="-3" metricUnit="m" metricValue="6.07" pageId="17" pageNumber="302" unit="mm" value="6.07">6.07 mm</quantity>
|
||
, nmp P6V 145289, male, 25.41 +
|
||
<quantity id="4C9E9B33FFEAFFB6FE8E9BD78FCDFBFC" box="[330,413,997,1024]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="17" pageNumber="302" unit="mm" value="5.7">5.7 mm</quantity>
|
||
, nmp P6V 145290, male, 25.53 +
|
||
<quantity id="4C9E9B33FFEAFFB6FEC69C3B8F01FBD8" box="[258,337,1033,1060]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="17" pageNumber="302" unit="mm" value="6.1">6.1 mm</quantity>
|
||
, all four from Selce, Kvarner area,
|
||
<collectingCountry id="F3717646FFEAFFB6FEC09C1F8F07FBBB" box="[260,343,1069,1095]" name="Croatia" pageId="17" pageNumber="302">Croatia</collectingCountry>
|
||
; nmp P6V 89017, male, 28.12 +
|
||
<quantity id="4C9E9B33FFEAFFB6FF7C9C638F46FB97" box="[184,278,1105,1131]" metricMagnitude="-3" metricUnit="m" metricValue="6.78" pageId="17" pageNumber="302" unit="mm" value="6.78">6.78 mm</quantity>
|
||
and nmp P6V 89018, female, 21.73 +
|
||
<quantity id="4C9E9B33FFEAFFB6FF749C478F5FFB73" box="[176,271,1141,1168]" metricMagnitude="-3" metricUnit="m" metricValue="5.42" pageId="17" pageNumber="302" unit="mm" value="5.42">5.42 mm</quantity>
|
||
, both from Punta Glavina, Krk Island,
|
||
<collectingCountry id="F3717646FFEAFFB6FF769CAB8F56FB4F" box="[178,262,1177,1203]" name="Croatia" pageId="17" pageNumber="302">Croatia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C655DFFEAFFB6FF459C8F8A7FFED7" pageId="17" pageNumber="302" type="diagnosis">
|
||
<paragraph id="8BD936D6FFEAFFB6FF459C8F8A7FFED7" blockId="17.[98,610,171,1706]" lastBlockId="17.[645,1157,166,300]" pageId="17" pageNumber="302">
|
||
<emphasis id="B912EAC4FFEAFFB6FF459C8F8F31FB2B" box="[129,353,1213,1239]" italics="true" pageId="17" pageNumber="302">
|
||
Diagnosis.
|
||
<taxonomicName id="4C664D55FFEAFFB6FF389C8F8F31FB2B" authorityName="Risso" authorityYear="1827" box="[252,353,1213,1239]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="302" phylum="Chordata" rank="species" species="zebrus">Z. zebrus</taxonomicName>
|
||
</emphasis>
|
||
differs from the only congeneric species,
|
||
<taxonomicName id="4C664D55FFEAFFB6FEFE9CD38FE3FB06" authority="Cooper & Yamaguchi & Macdonald & Nanova & Yudin & Dugmore & Kitchener, 2021" authorityName="Cooper & Yamaguchi & Macdonald & Nanova & Yudin & Dugmore & Kitchener" authorityYear="2021" box="[314,435,1248,1274]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="302" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">
|
||
<emphasis id="B912EAC4FFEAFFB6FEFE9CD38FE3FB06" box="[314,435,1248,1274]" italics="true" pageId="17" pageNumber="302">Z.pallaoroi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22157BFFFEAFFB6FE7D9CD38C59FB07" box="[441,521,1249,1275]" pageId="17" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by each of the following characters: (1) snout shorter than eye, its length 0.8‒0.9 of eye diameter; (2) posterior nostril 1/4‒2/5 of anterior nostril (
|
||
<figureCitation id="135D2A53FFEAFFB6FDFB9D7E8ED4FA76" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="17" pageNumber="302">fig. 2D</figureCitation>
|
||
); (3) eye diameter is
|
||
<quantity id="4C9E9B33FFEAFFB6FEAB9D428F80FA76" box="[367,464,1392,1418]" metricMagnitude="-2" metricUnit="m" metricValue="9.143999999999998" metricValueMax="10.414" metricValueMin="7.874" pageId="17" pageNumber="302" unit="in" value="3.5999999999999996" valueMax="4.1" valueMin="3.1">3.1‒4.1 in</quantity>
|
||
head length; (4) left and right ventrolateral head ridges disconnected anteriorly by midventral flat area (
|
||
<figureCitation id="135D2A53FFEAFFB6FFAD9DEE8EE7FA0A" box="[105,183,1500,1526]" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="17" pageNumber="302">fig. 2F</figureCitation>
|
||
), (5) anterior membrane in midline depth 1/4‒1/2 of spinous ray (
|
||
<figureCitation id="135D2A53FFEAFFB6FE6C9E328FA0F9E6" box="[424,496,1536,1562]" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="17" pageNumber="302">fig. 2E</figureCitation>
|
||
); (6) head canal pores of moderate size, distance between pore
|
||
<emphasis id="B912EAC4FFEAFFB6FF5F9E7A8EF6F99D" box="[155,166,1608,1633]" italics="true" pageId="17" pageNumber="302">Ρ</emphasis>
|
||
and
|
||
<emphasis id="B912EAC4FFEAFFB6FF189E7A8EBEF99E" box="[220,238,1608,1634]" italics="true" pageId="17" pageNumber="302">Ρ1</emphasis>
|
||
about three times or more longer than pore
|
||
<emphasis id="B912EAC4FFEAFFB6FF199E5E8EBAF979" box="[221,234,1644,1669]" italics="true" pageId="17" pageNumber="302">α</emphasis>
|
||
diameter; (7) suborbital sensory papillae row
|
||
<emphasis id="B912EAC4FFEAFFB6FF329EA28F59F956" box="[246,265,1680,1706]" italics="true" pageId="17" pageNumber="302">5i</emphasis>
|
||
ends downwards distant from row
|
||
<emphasis id="B912EAC4FFEAFFB6FD7898948C9AFF3C" box="[700,714,166,192]" italics="true" pageId="17" pageNumber="302">d</emphasis>
|
||
, row
|
||
<emphasis id="B912EAC4FFEAFFB6FCD098948D76FF3C" box="[788,806,166,192]" italics="true" pageId="17" pageNumber="302">5i</emphasis>
|
||
length equal or smaller than distance between row
|
||
<emphasis id="B912EAC4FFEAFFB6FC4298F88DC8FF18" box="[902,920,202,228]" italics="true" pageId="17" pageNumber="302">5i</emphasis>
|
||
and row
|
||
<emphasis id="B912EAC4FFEAFFB6FBC398F88A45FF18" box="[1031,1045,202,228]" italics="true" pageId="17" pageNumber="302">d</emphasis>
|
||
; (8) body with 6‒9 dark vertical stripes at upper edge, broader or equal than pale interspaces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |