245 lines
35 KiB
XML
245 lines
35 KiB
XML
<document id="59B36CC74765F40055099CE61DE7E889" ID-DOI="10.5852/ejt.2021.742.1291" ID-GBIF-Dataset="62f9dc57-bade-4fbc-8af7-f92ff107593c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4655404" ID-ZooBank="23424EC3-8440-4C72-AD9E-C1EC3087F7CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617262825393" checkinUser="plazi" docAuthor="De Oliveira, Luiz Fernando M. & Brescovit, Antonio Domingos" docDate="2021" docId="03CF87ADFFB06634FDEB93AAFD52FE9D" docLanguage="en" docName="ejt-742_1-77.pdf" docOrigin="European Journal of Taxonomy 742" docSource="http://dx.doi.org/10.5852/ejt.2021.742.1291" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tafana humahuaca De Oliveira & Brescovit 2021, sp. nov." docType="treatment" docUuid="16D23428-F7B9-48A6-93EE-95B3AE8F97DC" docUuidSource="ZooBank" docVersion="10" lastPageNumber="27" masterDocId="FFF6FFD5FFA8662EFF99965EFF99FFCF" masterDocTitle="Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="25" updateTime="1701999878271" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DC0778D3EAAF05391ED7E2ABAA110403">Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species</mods:title>
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<mods:roleTerm id="C98193C0A01AF63AEE3718271E572B02">Author</mods:roleTerm>
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<mods:namePart id="D60B4FFEDF29FA359D504B84E759ADB2">De Oliveira, Luiz Fernando M.</mods:namePart>
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<mods:affiliation id="4EDAC54602F6982E3FE5F451538CE058">Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil. & Instituto de Biociências, Universidade de São Paulo, Rua do Matão, 321, 05508 - 090, São Paulo, SP, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="99D8C18525A036E60A3EEE521CB7FC75" type="email">luiz.moura.lfm@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="5F8C2E76A41C7084A86DA28A0911A936">Author</mods:roleTerm>
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<mods:namePart id="A30261607BFE2A16A0790ED7708D1214">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="26B2A6500297B8757864FCEF74064406">Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="E6012B7DADC0B78C8AA2A8EDB3381BA0" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:typeOfResource id="A0AA514C78F4FB090779DC5B04684A4B">text</mods:typeOfResource>
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<mods:title id="A9E6F7F3A94A17B1072779496F4C00C9">European Journal of Taxonomy</mods:title>
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<mods:part id="483B9B588F12051F7704304DB99A8BB6">
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<mods:date id="4F304DF9125623A8C559F9D957BF6997">2021</mods:date>
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<mods:number id="5DBE25C960EFCC495EA08DDB05D55DF7">2021-03-31</mods:number>
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<mods:url id="C29CD91D094B6ABA1C92E930AB3DFAEF">http://dx.doi.org/10.5852/ejt.2021.742.1291</mods:url>
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<treatment id="03CF87ADFFB06634FDEB93AAFD52FE9D" ID-DOI="http://doi.org/10.5281/zenodo.4658743" ID-GBIF-Taxon="180784538" ID-Zenodo-Dep="4658743" LSID="urn:lsid:zoobank.org:act:16D23428-F7B9-48A6-93EE-95B3AE8F97DC" httpUri="http://treatment.plazi.org/id/03CF87ADFFB06634FDEB93AAFD52FE9D" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
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<subSubSection id="C37C6530FFB06636FDEB93AAFC5BF9C0" box="[626,962,1524,1551]" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="8BD936BBFFB06636FDEB93AAFC5BF9C0" blockId="24.[366,1222,1524,1622]" box="[626,962,1524,1551]" pageId="24" pageNumber="25">
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<heading id="D09181D7FFB06636FDEB93AAFC5BF9C0" box="[626,962,1524,1551]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
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<taxonomicName id="4C664D38FFB06636FDEB93AAFCF9F9C0" ID-CoL="98Z2Z" authority="De Oliveira & Brescovit, 2021" authorityName="De Oliveira & Brescovit" authorityYear="2021" box="[626,864,1524,1551]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="humahuaca" status="sp. nov.">
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<emphasis id="B912EAA9FFB06636FDEB93AAFCF9F9C0" bold="true" box="[626,864,1524,1551]" italics="true" pageId="24" pageNumber="25">Tafana humahuaca</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157D2FFB06636FCFE93ABFC5BF9C0" box="[871,962,1525,1551]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37C6530FFB06636FEF79046FCEEF999" pageId="24" pageNumber="25" type="description">
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<paragraph id="8BD936BBFFB06636FEF79046FB5FF9FC" blockId="24.[366,1222,1524,1622]" box="[366,1222,1560,1587]" pageId="24" pageNumber="25">
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<uri id="FFF73AB9FFB06636FEF79046FB5FF9FC" box="[366,1222,1560,1587]" pageId="24" pageNumber="25">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC00C6EFFB06636FD099046FB5FF9FC" box="[656,1222,1560,1587]" pageId="24" pageNumber="25">16D23428-F7B9-48A6-93EE-95B3AE8F97DC</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BD936BBFFB06636FD259065FCEEF999" blockId="24.[366,1222,1524,1622]" box="[700,887,1595,1622]" pageId="24" pageNumber="25">
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<figureCitation id="135D2A3EFFB06636FD259065FC8CF999" box="[700,789,1595,1622]" captionStart="Fig" captionStartId="26.[189,232,1784,1810]" captionTargetBox="[189,1398,469,1754]" captionTargetId="figure-149@26.[189,1398,469,1754]" captionTargetPageId="26" captionText="Fig. 15. Tafana humahuaca sp. nov. (A, C–E =holotype, ♂, MACN-Ar 41787; B, F–G=paratype, ♀, MACN-Ar 41788). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A= atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T =tegulum; VTP= ventral tegular process. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655442" httpUri="https://zenodo.org/record/4655442/files/figure.png" pageId="24" pageNumber="25">Figs 15</figureCitation>
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,
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<figureCitation id="135D2A3EFFB06636FCBB9062FCD7F999" box="[802,846,1596,1622]" captionStart="Fig" captionStartId="56.[189,232,1682,1708]" captionTargetBox="[308,1279,265,1652]" captionTargetId="figure-164@56.[308,1279,265,1652]" captionTargetPageId="56" captionText="Fig. 32. Female genitalia, digested. A. Arachosia praesignis (Keyserling, 1891) (Amaurobioidinae) (IBSP 87975). B. Xiruana gracilipes (Keyserling, 1891) (Anyphaeninae) (IBSP 166120). C. Aysha prospera Keyserling, 1891 (IBSP 19783). D. Anyphaena accentuata (Walckenaer, 1802) (MCN). E. Patrera fulvastra Simon, 1903 (MCN 24208). F. Tafana riveti Simon, 1903 (MCZ). G. Tafana straminea (L. Koch, 1866) (AMNH). H. Tafana quelchi (Pocock, 1895) (IBSP 32131). I. Tafana kunturmarqa sp. nov. (ICN-Ar). J. Tafana humahuaca sp. nov. (MACN-Ar 41788). K. Tafana pastaza sp. nov. (AMNH). L. Tafana nevada sp. nov. (AMNH). Abbreviations: CD =copulatory ducts; CO= copulatory opening; FD = fertilization ducts; H =hood; S1= primary spermathecae; S2 =secondary spermathecae. Asterisks to globose membranous extension of proximal copulatory ducts. Scale bars: A–B, D–I, K–L=0.25 mm; C=0.3 mm; J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655507" httpUri="https://zenodo.org/record/4655507/files/figure.png" pageId="24" pageNumber="25">32J</figureCitation>
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,
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<figureCitation id="135D2A3EFFB06636FCC39065FCEEF999" box="[858,887,1595,1622]" captionStart="Fig" captionStartId="59.[189,232,1997,2023]" captionTargetBox="[222,1366,265,1955]" captionTargetId="figure-20@59.[222,1366,265,1955]" captionTargetPageId="59" captionText="Fig. 35. Distribution map of species of Tafana." figureDoi="http://doi.org/10.5281/zenodo.4655519" httpUri="https://zenodo.org/record/4655519/files/figure.png" pageId="24" pageNumber="25">35</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C6530FFB06636FF249023FAEEF89A" pageId="24" pageNumber="25" type="diagnosis">
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||
<paragraph id="8BD936BBFFB06636FF249023FEAFF958" blockId="24.[189,310,1661,1687]" box="[189,310,1661,1687]" pageId="24" pageNumber="25">
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<heading id="D09181D7FFB06636FF249023FEAFF958" bold="true" box="[189,310,1661,1687]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
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<emphasis id="B912EAA9FFB06636FF249023FEAFF958" bold="true" box="[189,310,1661,1687]" pageId="24" pageNumber="25">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD936BBFFB06636FF2490F2FAEEF89A" blockId="24.[188,1400,1708,1878]" pageId="24" pageNumber="25">
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Males of
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<taxonomicName id="4C664D38FFB06636FEA890F3FD8AF909" authority="De Oliveira & Brescovit, 2021" authorityName="De Oliveira & Brescovit" authorityYear="2021" box="[305,531,1708,1735]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="humahuaca" status="sp. nov.">
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<emphasis id="B912EAA9FFB06636FEA890F3FD8AF909" box="[305,531,1708,1735]" italics="true" pageId="24" pageNumber="25">Tafana humahuaca</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157D2FFB06636FD8390F3FDEFF908" box="[538,630,1709,1735]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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resemble those of
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<taxonomicName id="4C664D38FFB06636FCC590F3FC21F909" authorityName="Simon" authorityYear="1903" box="[860,952,1709,1734]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="riveti">
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<emphasis id="B912EAA9FFB06636FCC590F3FC21F909" box="[860,952,1709,1734]" italics="true" pageId="24" pageNumber="25">T. riveti</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C664D38FFB06636FC6D90F3FBF1F909" authority="De Oliveira & Brescovit, 2021" authorityName="De Oliveira & Brescovit" authorityYear="2021" box="[1012,1128,1708,1734]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nevada" status="sp. nov.">
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<emphasis id="B912EAA9FFB06636FC6D90F3FBF1F909" box="[1012,1128,1708,1734]" italics="true" pageId="24" pageNumber="25">T. nevada</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157D2FFB06636FBF690F3FB52F908" box="[1135,1227,1709,1735]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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by having the short embolus, but differ by having the apophysis at the base of the embolus truncated, extremely short and with excavation between the base of the embolus and the apophysis (
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<figureCitation id="135D2A3EFFB06636FB9E90AAFB0BF8C0" box="[1031,1170,1780,1807]" captionStart="Fig" captionStartId="26.[189,232,1784,1810]" captionTargetBox="[189,1398,469,1754]" captionTargetId="figure-149@26.[189,1398,469,1754]" captionTargetPageId="26" captionText="Fig. 15. Tafana humahuaca sp. nov. (A, C–E =holotype, ♂, MACN-Ar 41787; B, F–G=paratype, ♀, MACN-Ar 41788). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A= atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T =tegulum; VTP= ventral tegular process. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655442" httpUri="https://zenodo.org/record/4655442/files/figure.png" pageId="24" pageNumber="25">Fig. 15C–E</figureCitation>
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). Females resemble those of
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<taxonomicName id="4C664D38FFB06636FEBA9146FE56F8FE" authority="De Oliveira & Brescovit, 2021" authorityName="De Oliveira & Brescovit" authorityYear="2021" box="[291,463,1815,1842]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nevada" status="sp. nov.">
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<emphasis id="B912EAA9FFB06636FEBA9146FE56F8FE" box="[291,463,1815,1842]" italics="true" pageId="24" pageNumber="25">Tafana nevada</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157D2FFB06636FE4E9146FDA8F8FD" box="[471,561,1816,1842]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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by having the epigynal plate, but differ by having atrium narrow in the anterior region, small hood with lateral lobes extremely curved in the anterior region (
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<figureCitation id="135D2A3EFFB06636FB3A9165FAA8F899" box="[1187,1329,1851,1878]" captionStart="Fig" captionStartId="26.[189,232,1784,1810]" captionTargetBox="[189,1398,469,1754]" captionTargetId="figure-149@26.[189,1398,469,1754]" captionTargetPageId="26" captionText="Fig. 15. Tafana humahuaca sp. nov. (A, C–E =holotype, ♂, MACN-Ar 41787; B, F–G=paratype, ♀, MACN-Ar 41788). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A= atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T =tegulum; VTP= ventral tegular process. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655442" httpUri="https://zenodo.org/record/4655442/files/figure.png" pageId="24" pageNumber="25">Figs 15F–G</figureCitation>
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,
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<figureCitation id="135D2A3EFFB06636FAA49165FAF3F89A" box="[1341,1386,1851,1877]" captionStart="Fig" captionStartId="56.[189,232,1682,1708]" captionTargetBox="[308,1279,265,1652]" captionTargetId="figure-164@56.[308,1279,265,1652]" captionTargetPageId="56" captionText="Fig. 32. Female genitalia, digested. A. Arachosia praesignis (Keyserling, 1891) (Amaurobioidinae) (IBSP 87975). B. Xiruana gracilipes (Keyserling, 1891) (Anyphaeninae) (IBSP 166120). C. Aysha prospera Keyserling, 1891 (IBSP 19783). D. Anyphaena accentuata (Walckenaer, 1802) (MCN). E. Patrera fulvastra Simon, 1903 (MCN 24208). F. Tafana riveti Simon, 1903 (MCZ). G. Tafana straminea (L. Koch, 1866) (AMNH). H. Tafana quelchi (Pocock, 1895) (IBSP 32131). I. Tafana kunturmarqa sp. nov. (ICN-Ar). J. Tafana humahuaca sp. nov. (MACN-Ar 41788). K. Tafana pastaza sp. nov. (AMNH). L. Tafana nevada sp. nov. (AMNH). Abbreviations: CD =copulatory ducts; CO= copulatory opening; FD = fertilization ducts; H =hood; S1= primary spermathecae; S2 =secondary spermathecae. Asterisks to globose membranous extension of proximal copulatory ducts. Scale bars: A–B, D–I, K–L=0.25 mm; C=0.3 mm; J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655507" httpUri="https://zenodo.org/record/4655507/files/figure.png" pageId="24" pageNumber="25">32J</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C6530FFB06636FF249123FDC5F827" pageId="24" pageNumber="25" type="etymology">
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<paragraph id="8BD936BBFFB06636FF249123FEDDF858" blockId="24.[189,324,1917,1943]" box="[189,324,1917,1943]" pageId="24" pageNumber="25">
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<heading id="D09181D7FFB06636FF249123FEDDF858" bold="true" box="[189,324,1917,1943]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="B912EAA9FFB06636FF249123FEDDF858" bold="true" box="[189,324,1917,1943]" pageId="24" pageNumber="25">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB06636FF2491F5FDC5F827" blockId="24.[189,1399,1963,2025]" pageId="24" pageNumber="25">
|
||
The specific name, a noun in apposition, is a tribute to ‘Quebrada de Humahuaca’, a deep canyon located in
|
||
<collectingRegion id="49A2F859FFB06636FF429190FE14F827" box="[219,397,1998,2024]" country="Argentina" name="Jujuy" pageId="24" pageNumber="25">Jujuy Province</collectingRegion>
|
||
, the
|
||
<typeStatus id="54DD8819FFB06636FE5D9191FE6EF826" box="[452,503,1999,2025]" pageId="24" pageNumber="25">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C6530FFB16637FF249756FB57FDC7" pageId="25" pageNumber="26" type="materials_examined">
|
||
<paragraph id="8BD936BBFFB16637FF249756FE33FEED" blockId="25.[189,426,264,290]" box="[189,426,264,290]" pageId="25" pageNumber="26">
|
||
<heading id="D09181D7FFB16637FF249756FE33FEED" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="B912EAA9FFB16637FF249756FE33FEED" bold="true" box="[189,426,264,290]" pageId="25" pageNumber="26">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF739764FDC5FE50" blockId="25.[189,1399,314,415]" pageId="25" pageNumber="26">
|
||
<materialsCitation id="3B0E3CE6FFB16637FF739764FDC5FE50" ID-GBIF-Occurrence="3066856314" collectingDate="1969-10" collectionCode="MACN-Ar" collectorName="G. Williner" country="Argentina" latitude="-24.30861" location="Santa Barbara" longLatPrecision="21" longitude="-64.66028" municipality="Santa Barbara" pageId="25" pageNumber="26" specimenCode="MACN-Ar 41787" specimenCount="1" specimenCount-male="1" stateProvince="Jujuy Province" typeStatus="holotype">
|
||
<emphasis id="B912EAA9FFB16637FF739764FEC5FE9B" bold="true" box="[234,348,314,340]" pageId="25" pageNumber="26">
|
||
<typeStatus id="54DD8819FFB16637FF739764FEC5FE9B" box="[234,348,314,340]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F371762BFFB16637FF249701FEF1FEB6" box="[189,360,351,377]" name="Argentina" pageId="25" pageNumber="26">ARGENTINA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D60FD32FFB16637FE089701FE50FEB5" box="[401,457,351,378]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49A2F859FFB16637FE469701FD03FEB6" box="[479,666,351,377]" country="Argentina" name="Jujuy" pageId="25" pageNumber="26">Jujuy Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BBDACC1FFB16637FD379701FCF8FEB6" box="[686,865,351,377]" pageId="25" pageNumber="26">Santa Bárbara</collectingMunicipality>
|
||
; [
|
||
<geoCoordinate id="EE52507CFFB16637FCE59700FB81FEB6" box="[892,1048,350,378]" degrees="24" direction="south" minutes="18" orientation="latitude" pageId="25" pageNumber="26" precision="15" seconds="31" value="-24.30861">24°18′31″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE52507CFFB16637FBB79700FB4CFEB5" box="[1070,1237,350,378]" degrees="64" direction="west" minutes="39" orientation="longitude" pageId="25" pageNumber="26" precision="15" seconds="37" value="-64.66028">64°39′37″ W</geoCoordinate>
|
||
];
|
||
<date id="FFD8107BFFB16637FB6B9701FAEBFEB6" box="[1266,1394,351,377]" pageId="25" pageNumber="26" value="1969-10">
|
||
<collectingDate id="EF9CE993FFB16637FB6B9701FAEBFEB6" box="[1266,1394,351,377]" pageId="25" pageNumber="26" value="1969-10">Oct. 1969</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2693536DFFB16637FF2497DBFED8FE50" box="[189,321,389,415]" pageId="25" pageNumber="26">G. Williner</collectorName>
|
||
leg.;
|
||
<specimenCode id="DBC09EC0FFB16637FE1A97DBFDC1FE50" box="[387,600,389,415]" collectionCode="MACN-Ar" pageId="25" pageNumber="26">MACN-Ar 41787</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF739797FB57FDC7" blockId="25.[189,1230,457,521]" pageId="25" pageNumber="26">
|
||
<materialsCitation id="3B0E3CE6FFB16637FF739797FB50FDC7" ID-GBIF-Occurrence="3066856334" collectingDate="1962-10" collectionCode="MACN-Ar" collectorName="G. Williner" country="Argentina" latitude="-24.30861" location="ARGENTINA" longLatPrecision="21" longitude="-64.66028" pageId="25" pageNumber="26" specimenCode="MACN-Ar 41788" specimenCount="1" specimenCount-female="1" stateProvince="Jujuy" typeStatus="paratype">
|
||
<emphasis id="B912EAA9FFB16637FF739797FEC5FE2C" bold="true" box="[234,348,457,483]" pageId="25" pageNumber="26">
|
||
<typeStatus id="54DD8819FFB16637FF739797FEC5FE2C" box="[234,348,457,483]" pageId="25" pageNumber="26" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F371762BFFB16637FF2497B0FEF1FDC7" box="[189,360,494,520]" name="Argentina" pageId="25" pageNumber="26">ARGENTINA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D60FD32FFB16637FEE697B0FE35FDC6" box="[383,428,494,521]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<date id="FFD8107BFFB16637FCE897B0FC71FDC7" box="[881,1000,494,520]" pageId="25" pageNumber="26" value="1962-10">
|
||
<collectingDate id="EF9CE993FFB16637FCE897B0FC71FDC7" box="[881,1000,494,520]" pageId="25" pageNumber="26" value="1962-10">Oct. 1962</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="DBC09EC0FFB16637FC6D97B0FB50FDC7" box="[1012,1225,494,520]" collectionCode="MACN-Ar" pageId="25" pageNumber="26">MACN-Ar 41788</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C6530FFB16637FF24946CFAEEF827" pageId="25" pageNumber="26" type="description">
|
||
<paragraph id="8BD936BBFFB16637FF24946CFED7FD83" blockId="25.[189,334,562,588]" box="[189,334,562,588]" pageId="25" pageNumber="26">
|
||
<heading id="D09181D7FFB16637FF24946CFED7FD83" bold="true" box="[189,334,562,588]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="B912EAA9FFB16637FF24946CFED7FD83" bold="true" box="[189,334,562,588]" pageId="25" pageNumber="26">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF73943DFD15FDB2" blockId="25.[187,1399,611,713]" box="[234,652,611,637]" pageId="25" pageNumber="26">
|
||
<emphasis id="B912EAA9FFB16637FF73943DFEB3FDB2" bold="true" box="[234,298,611,637]" pageId="25" pageNumber="26">Male</emphasis>
|
||
(
|
||
<typeStatus id="54DD8819FFB16637FEA0943DFE3DFDB2" box="[313,420,611,637]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
, MACN-Ar 41787)
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF2494D7FAC3FD06" blockId="25.[187,1399,611,713]" pageId="25" pageNumber="26">
|
||
COLOR. Exemplar probably discolored. Carapace orange (
|
||
<figureCitation id="135D2A3EFFB16637FCC994D7FC25FD6B" box="[848,956,649,676]" captionStart="Fig" captionStartId="26.[189,232,1784,1810]" captionTargetBox="[189,1398,469,1754]" captionTargetId="figure-149@26.[189,1398,469,1754]" captionTargetPageId="26" captionText="Fig. 15. Tafana humahuaca sp. nov. (A, C–E =holotype, ♂, MACN-Ar 41787; B, F–G=paratype, ♀, MACN-Ar 41788). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A= atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T =tegulum; VTP= ventral tegular process. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655442" httpUri="https://zenodo.org/record/4655442/files/figure.png" pageId="25" pageNumber="26">Fig. 15A</figureCitation>
|
||
). Labium, sternum and endites yellow. Chelicerae yellow. Legs I–III yellow with spots on the articles. Abdomen with dark spots (
|
||
<figureCitation id="135D2A3EFFB16637FB7994F0FAD4FD06" box="[1248,1357,686,713]" captionStart="Fig" captionStartId="26.[189,232,1784,1810]" captionTargetBox="[189,1398,469,1754]" captionTargetId="figure-149@26.[189,1398,469,1754]" captionTargetPageId="26" captionText="Fig. 15. Tafana humahuaca sp. nov. (A, C–E =holotype, ♂, MACN-Ar 41787; B, F–G=paratype, ♀, MACN-Ar 41788). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A= atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T =tegulum; VTP= ventral tegular process. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655442" httpUri="https://zenodo.org/record/4655442/files/figure.png" pageId="25" pageNumber="26">Fig. 15A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF2494ADFE4DFCB2" blockId="25.[188,1399,754,893]" pageId="25" pageNumber="26">MEASUREMENTS. Total length 5.3, carapace length 2.3, width 1.7. Clypeus height 0.06. Eye diameters and interdistances: AME 0.06, ALE 0.1, PME 0.1, PLE 0.08; AME–AME 0.02, AME–ALE 0.02, PME– PME 0.1, PME–PLE 0.08, ALE–PLE 0.02. Chelicerae 0.8 long, with five promarginal teeth and four retromarginal denticles.</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF2495F6FD02FC28" blockId="25.[185,1399,935,1000]" pageId="25" pageNumber="26">LEG MEASUREMENTS. I—lost; II—femur 1.7/ patella 0.73 /tibia 1.7/ metatarsus 1.6 /tarsus 0.84 / total 6.57; III—1.6 / 0.7/ 1.4 /1.4 / 0.7/ 5.8; IV—lost.</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF24924FFB61FBE4" blockId="25.[189,1272,1041,1067]" box="[189,1272,1041,1067]" pageId="25" pageNumber="26">LEG SPINATION. II—tibia v2-1-2, p1-1-0; III—tibia v1-2-0, p0-1-0, metatarsus v2-1-0, p0-1-0.</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF24920BFDD5FB2E" blockId="25.[188,1399,1109,1249]" pageId="25" pageNumber="26">
|
||
PALP. Retrolateral tibial apophysis conical with apex curved; cymbium with retrolateral projection more rounded; embolus short; ventral tegular process near the median apophysis and sclerotized; sperm duct in vertical position with secondary looping reaching the apex of the tegulum; median apophysis slender and curved apically (
|
||
<figureCitation id="135D2A3EFFB16637FE2B9298FDA6FB2E" box="[434,575,1222,1249]" captionStart="Fig" captionStartId="26.[189,232,1784,1810]" captionTargetBox="[189,1398,469,1754]" captionTargetId="figure-149@26.[189,1398,469,1754]" captionTargetPageId="26" captionText="Fig. 15. Tafana humahuaca sp. nov. (A, C–E =holotype, ♂, MACN-Ar 41787; B, F–G=paratype, ♀, MACN-Ar 41788). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A= atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T =tegulum; VTP= ventral tegular process. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655442" httpUri="https://zenodo.org/record/4655442/files/figure.png" pageId="25" pageNumber="26">Fig. 15C–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF249354FAEEFAEB" blockId="25.[189,1399,1290,1316]" box="[189,1399,1290,1316]" pageId="25" pageNumber="26">ABDOMEN. Length 2.7, epigastric furrow 0.6 from tracheal spiracle, spiracle 1.0 from base of spinnerets.</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF739310FD3FFAA7" blockId="25.[189,1222,1358,1422]" box="[234,678,1358,1384]" pageId="25" pageNumber="26">
|
||
<emphasis id="B912EAA9FFB16637FF739310FEDCFAA7" bold="true" box="[234,325,1358,1384]" pageId="25" pageNumber="26">Female</emphasis>
|
||
(
|
||
<typeStatus id="54DD8819FFB16637FECC9310FE26FAA7" box="[341,447,1358,1384]" pageId="25" pageNumber="26" type="paratype">paratype</typeStatus>
|
||
, MACN-Ar 41788)
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF24932DFB5FFA41" blockId="25.[189,1222,1358,1422]" box="[189,1222,1395,1422]" pageId="25" pageNumber="26">
|
||
COLOR. Exemplar probably discolored. Coloration as in males of this species (
|
||
<figureCitation id="135D2A3EFFB16637FBD7932DFB21FA41" box="[1102,1208,1395,1422]" captionStart="Fig" captionStartId="26.[189,232,1784,1810]" captionTargetBox="[189,1398,469,1754]" captionTargetId="figure-149@26.[189,1398,469,1754]" captionTargetPageId="26" captionText="Fig. 15. Tafana humahuaca sp. nov. (A, C–E =holotype, ♂, MACN-Ar 41787; B, F–G=paratype, ♀, MACN-Ar 41788). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A= atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T =tegulum; VTP= ventral tegular process. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655442" httpUri="https://zenodo.org/record/4655442/files/figure.png" pageId="25" pageNumber="26">Fig. 15B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF2493E6FE4DF98D" blockId="25.[189,1399,1463,1602]" pageId="25" pageNumber="26">MEASUREMENTS. Total length 3.2, carapace length 2.4, width 1.3. Clypeus height 0.06. Eye diameters and interdistances: AME 0.08, ALE 0.1, PME 0.1, PLE 0.1; AME–AME 0.04, AME–ALE 0.04, PME– PME 0.16, PME–PLE 0.12, ALE–PLE 0.04. Chelicerae 1.1 long, with four promarginal teeth and five retromarginal denticles.</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF249033FAB6F963" blockId="25.[189,1398,1644,1708]" pageId="25" pageNumber="26">LEG MEASUREMENTS. I—femur 1.6 /patella 0.8/ tibia 1.4/ metatarsus 1.3/ tarsus 0.8 /total 5.9; II— 1.9/ 0.8/ 1.1 /1.15 / 0.6 /5.55; III—1.3/ 0.63 / 0.8/ 1.2/ 0.5/ 4.43; IV—1.9 / 0.8/ 1.5/ 2.1/ 0.65 / 6.95.</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF249088FD53F8DA" blockId="25.[189,1399,1749,1814]" pageId="25" pageNumber="26">LEG SPINATION. I—tibia v2-2-0, metatarsus v2-0-0; II—tibia v1-1-0; III—tibia v1-2-2, p1-1-0, r1-1-0, metatarsus v2-1-2; IV—tibia v1-2-2, p1-1-1.</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF24911EFB0CF86A" blockId="25.[189,1399,1855,1957]" pageId="25" pageNumber="26">
|
||
EPIGYNUM. Hood semicircular and sclerotized; short atrium; lateral lobes over copulatory ducts. Internally, copulatory ducts narrow; primary spermathecae large and oval; secondary spermathecae above the primary spermathecae and rounded; fertilization ducts long (
|
||
<figureCitation id="135D2A3EFFB16637FC6591D4FB10F86A" box="[1020,1161,1930,1957]" captionStart="Fig" captionStartId="26.[189,232,1784,1810]" captionTargetBox="[189,1398,469,1754]" captionTargetId="figure-149@26.[189,1398,469,1754]" captionTargetPageId="26" captionText="Fig. 15. Tafana humahuaca sp. nov. (A, C–E =holotype, ♂, MACN-Ar 41787; B, F–G=paratype, ♀, MACN-Ar 41788). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A= atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T =tegulum; VTP= ventral tegular process. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655442" httpUri="https://zenodo.org/record/4655442/files/figure.png" pageId="25" pageNumber="26">Fig. 15F–G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB16637FF249191FAEEF827" blockId="25.[189,1399,1998,2025]" box="[189,1399,1998,2025]" pageId="25" pageNumber="26">ABDOMEN. Length 3.2, epigastric furrow 0.7 from tracheal spiracle, spiracle 1.3 from base of spinnerets.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C6530FFB26634FF249756FD52FE9D" box="[189,715,264,338]" pageId="26" pageNumber="27" type="distribution">
|
||
<paragraph id="8BD936BBFFB26634FF249756FECFFEED" blockId="26.[189,342,264,290]" box="[189,342,264,290]" pageId="26" pageNumber="27">
|
||
<heading id="D09181D7FFB26634FF249756FECFFEED" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
|
||
<emphasis id="B912EAA9FFB26634FF249756FECFFEED" bold="true" box="[189,342,264,290]" pageId="26" pageNumber="27">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFFB26634FF249766FD52FE9D" blockId="26.[189,715,311,338]" box="[189,715,311,338]" pageId="26" pageNumber="27">
|
||
Known only from
|
||
<collectingRegion id="49A2F859FFB26634FE0E9769FE41FE9E" box="[407,472,311,337]" country="Argentina" name="Jujuy" pageId="26" pageNumber="27">Jujuy</collectingRegion>
|
||
,
|
||
<collectingCountry id="F371762BFFB26634FE789769FDCFFE9E" box="[481,598,311,337]" name="Argentina" pageId="26" pageNumber="27">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="135D2A3EFFB26634FDFF9769FD27FE9D" box="[614,702,311,338]" captionStart="Fig" captionStartId="59.[189,232,1997,2023]" captionTargetBox="[222,1366,265,1955]" captionTargetId="figure-20@59.[222,1366,265,1955]" captionTargetPageId="59" captionText="Fig. 35. Distribution map of species of Tafana." figureDoi="http://doi.org/10.5281/zenodo.4655519" httpUri="https://zenodo.org/record/4655519/files/figure.png" pageId="26" pageNumber="27">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |