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<document ID-DOI="10.11646/zootaxa.5231.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7575308" ID-ZooBank="EBD53535-397E-4894-B0A4-9AC3CC38DF3C" approvalRequired="228" approvalRequired_for_taxonomicNames="8" approvalRequired_for_textStreams="89" approvalRequired_for_treatments="131" checkinTime="1674811458118" checkinUser="plazi" docAuthor="Lelej, Arkady S. &amp; Williams, Kevin A." docDate="2023" docId="03CF87A93122B50CFF17FA91F7573657" docLanguage="en" docName="zootaxa.5231.2.1.pdf" docOrigin="Zootaxa 5231 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nordeniella Lelej 2005" docType="treatment" docVersion="2" lastPageNumber="119" masterDocId="FFF6FFD13133B51EFF80FF97F345374A" masterDocTitle="Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)" masterLastPageNumber="140" masterPageNumber="101" pageNumber="118" updateTime="1674812169880" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lelej, Arkady S.</mods:namePart>
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<mods:affiliation>Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia. lelej @ biosoil. ru; https: // orcid. org / 0000 - 0001 - 7501 - 0981</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Williams, Kevin A.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6073-1070</mods:nameIdentifier>
<mods:affiliation>Plant Pest Diagnostic Center, California Department of Food &amp; Agriculture, 3294 Meadowview Road, Sacramento, CA 95832, USA. Kevin. Williams @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 6073 - 1070</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-27</mods:number>
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<mods:detail type="volume">
<mods:number>5231</mods:number>
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<paragraph blockId="17.[151,483,1285,1349]" box="[151,483,1285,1312]" pageId="17" pageNumber="118">
<heading bold="true" box="[151,483,1285,1312]" fontSize="11" level="1" pageId="17" pageNumber="118" reason="1">
<emphasis bold="true" box="[151,483,1285,1312]" pageId="17" pageNumber="118">
14.
<taxonomicName authority="Lelej, 2005" authorityName="Lelej" authorityYear="2005" box="[195,483,1285,1312]" class="Insecta" family="Mutillidae" genus="Nordeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[195,338,1285,1312]" italics="true" pageId="17" pageNumber="118">Nordeniella</emphasis>
<bibRefCitation author="Lelej, A. S." box="[346,483,1285,1312]" pageId="17" pageNumber="118" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej, 2005</bibRefCitation>
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</emphasis>
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</paragraph>
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<paragraph blockId="17.[151,483,1285,1349]" box="[151,295,1322,1349]" pageId="17" pageNumber="118">
(
<figureCitation box="[159,287,1322,1349]" captionStart="FIGURES 4552" captionStartId="20.[152,270,1865,1889]" captionTargetBox="[151,1436,182,1838]" captionTargetId="figure-16@20.[151,1436,181,1841]" captionTargetPageId="20" captionText="FIGURES 4552. 4548. Nordeniella thermophila (Turner, 1911): 45. ♀, dorsal habitus, 46. ♀, lateral hdbitus, 47. ♁, lateral habitus, 48. ♁, head, face view. 49. Nuristanilla kabakovi Lelej in Lelej et Kabakov, 1980, ♀, holotype, lateral habitus. 5052. Paglianotilla hogenesi Lelej in Lelej et van Harten, 2006, ♁, paratype, dorsal habitus (50), lateral habitus (51), head, face view (52). (Photographs by K. Williams and V. Loktionov (49, 50))." figureDoi="http://doi.org/10.5281/zenodo.7575320" httpUri="https://zenodo.org/record/7575320/files/figure.png" pageId="17" pageNumber="118">Figs 4548</figureCitation>
)
</paragraph>
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<paragraph blockId="17.[151,1437,1383,1438]" pageId="17" pageNumber="118">
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<treatmentCitation author="Lelej, A. S." box="[151,435,1383,1408]" page="48" pageId="17" pageNumber="118" year="2005">
<taxonomicName authority="Lelej, 2005: 48" authorityName="Lelej" authorityPageNumber="48" authorityYear="2005" box="[151,435,1383,1408]" class="Insecta" family="Mutillidae" genus="Nordeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis box="[151,271,1384,1408]" italics="true" pageId="17" pageNumber="118">Nordeniella</emphasis>
<bibRefCitation author="Lelej, A. S." box="[278,435,1383,1408]" pageId="17" pageNumber="118" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej, 2005: 48</bibRefCitation>
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, 145, 149, 185, ♁,
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;
<treatmentCitation author="Lelej, A. S. &amp; Brothers, D. J." box="[661,932,1384,1408]" page="40" pageId="17" pageNumber="118" year="2008">
<bibRefCitation author="Lelej, A. S. &amp; Brothers, D. J." box="[661,932,1384,1408]" pageId="17" pageNumber="118" pagination="1 - 79" refId="ref27054" refString="Lelej, A. S. &amp; Brothers, D. J. (2008) The genus-group names of Mutillidae (Hymenoptera) and their type species, with a new genus, new name, new synonymies, new combinations and lectotypifications. Zootaxa, 1889 (1), 1 - 79. https: // doi. org / 10.11646 / zootaxa. 1889.1.1" type="journal article" year="2008">Lelej &amp; Brothers 2008: 40</bibRefCitation>
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;
<treatmentCitation author="Brothers, D. J. &amp; Lelej, A. S." box="[943,1212,1383,1408]" page="95" pageId="17" pageNumber="118" year="2017">
<bibRefCitation author="Brothers, D. J. &amp; Lelej, A. S." box="[943,1212,1383,1408]" pageId="17" pageNumber="118" pagination="1 - 97" refId="ref25501" refString="Brothers, D. J. &amp; Lelej, A. S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1 - 97. https: // doi. org / 10.3897 / jhr. 60.20091" type="journal article" year="2017">Brothers &amp; Lelej 2017: 95</bibRefCitation>
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, ♁,
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;
<treatmentCitation ID-DOI="http://doi.org/10.5281/zenodo.5923575" ID-Zenodo-Dep="5923575" author="Williams, K. A. &amp; Lelej, A. S. &amp; Okayasu, J. &amp; Borkent, C. J. &amp; Malee, R. &amp; Thoawan, K. &amp; Thaochan, N." httpUri="http://treatment.plazi.org/id/03DCF504FFE09869FF3928624829C6F4" page="22" pageId="17" pageNumber="118" year="2019">
<bibRefCitation author="Williams, K. A. &amp; Lelej, A. S. &amp; Okayasu, J. &amp; Borkent, C. J. &amp; Malee, R. &amp; Thoawan, K. &amp; Thaochan, N." pageId="17" pageNumber="118" pagination="1 - 69" refId="ref30512" refString="Williams, K. A., Lelej, A. S., Okayasu, J., Borkent, C. J., Malee, R., Thoawan, K. &amp; Thaochan, N. (2019) The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia. Zootaxa, 4602 (1), 1 - 69. https: // doi. org / 10.11646 / zootaxa. 4602.1.1" type="journal article" year="2019">
Williams
<emphasis box="[1385,1437,1384,1408]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
2019: 22
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,
<specimenCount box="[301,321,1415,1438]" pageId="17" pageNumber="118" type="female"></specimenCount>
;
<treatmentCitation author="Pagliano, G. &amp; Brothers, D. J. &amp; Cambra, R. &amp; Lelej, A. S. &amp; Lo Cascio, P. &amp; Matteini Palmerini, M. &amp; Scaramozzino, P. L. &amp; Williams, K. A. &amp; Romano, M." box="[334,591,1414,1438]" page="177" pageId="17" pageNumber="118" year="2020">
<bibRefCitation author="Pagliano, G. &amp; Brothers, D. J. &amp; Cambra, R. &amp; Lelej, A. S. &amp; Lo Cascio, P. &amp; Matteini Palmerini, M. &amp; Scaramozzino, P. L. &amp; Williams, K. A. &amp; Romano, M." box="[334,591,1414,1438]" pageId="17" pageNumber="118" pagination="5 - 425" refId="ref28648" refString="Pagliano, G., Brothers, D. J., Cambra, R., Lelej, A. S., Lo Cascio, P., Matteini Palmerini, M., Scaramozzino, P. L., Williams, K. A. &amp; Romano, M. (2020 [&quot; 2018 &quot;]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torin o, 36 (1 - 2), 5 - 425." type="journal article" year="2020">
Pagliano
<emphasis box="[429,480,1414,1438]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
2020: 177
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;
<treatmentCitation author="Okayasu, J." box="[601,787,1414,1438]" page="57" pageId="17" pageNumber="118" year="2022">
<bibRefCitation author="Okayasu, J." box="[601,787,1414,1438]" pageId="17" pageNumber="118" pagination="56 - 63" refId="ref28328" refString="Okayasu, J. (2022) Remarkable range extension of the genus Nordeniella Lelej, 2005 (Hymenoptera: Mutillidae: Smicromyrmini): Description of a new species from Sumbawa, Indonesia. Journal of Insect Biodiversity, 32 (2), 56 - 63. https: // doi. org / 10.12976 / jib / 2022.32.2.3" type="journal article" year="2022">Okayasu 2022: 57</bibRefCitation>
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,
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;
<treatmentCitation author="Thaochan, N. &amp; Williams, K. A. &amp; Thoawan, K. &amp; Jeenthong, T. &amp; Sittichaya, W." box="[830,1097,1414,1438]" page="158" pageId="17" pageNumber="118" year="2022">
<bibRefCitation author="Thaochan, N. &amp; Williams, K. A. &amp; Thoawan, K. &amp; Jeenthong, T. &amp; Sittichaya, W." box="[830,1097,1414,1438]" pageId="17" pageNumber="118" pagination="151 - 165" refId="ref30138" refString="Thaochan, N., Williams, K. A., Thoawan, K., Jeenthong, T. &amp; Sittichaya, W. (2022) Three new species and one new country record of velvet ants (Hymenoptera, Mutillidae) from Thailand. Journal of Hymenoptera Research, 93, 151 - 165. https: // doi. org / 10.3897 / jhr. 93.94727" type="journal article" year="2022">
Thaochan
<emphasis box="[936,987,1414,1438]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
2022: 158
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,
<specimenCount box="[1108,1128,1415,1438]" pageId="17" pageNumber="118" type="female"></specimenCount>
.
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</paragraph>
</subSubSection>
<subSubSection box="[151,969,1477,1504]" pageId="17" pageNumber="118" type="type_taxon">
<paragraph blockId="17.[151,1437,1477,2008]" box="[151,969,1477,1504]" pageId="17" pageNumber="118">
<emphasis bold="true" box="[151,306,1477,1504]" pageId="17" pageNumber="118">
<typeStatus box="[151,210,1478,1504]" pageId="17" pageNumber="118">Type</typeStatus>
species.
</emphasis>
<taxonomicName authority="Turner, 1911" authorityName="Turner" authorityYear="1911" box="[313,691,1478,1504]" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="thermophila">
<emphasis box="[313,540,1478,1504]" italics="true" pageId="17" pageNumber="118">Mutilla thermophila</emphasis>
<bibRefCitation author="Turner, R. E." box="[546,691,1478,1504]" pageId="17" pageNumber="118" pagination="141 - 154" refId="ref30341" refString="Turner, R. E. (1911) New Hymenoptera from Ceylon. Mutillidae and Scoliidae. Spolia Zeylanica, 7 (27), 141 - 154." type="journal article" year="1911">Turner, 1911</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118" type="diagnosis">
<paragraph blockId="17.[151,1437,1477,2008]" pageId="17" pageNumber="118">
<emphasis bold="true" box="[199,314,1513,1539]" pageId="17" pageNumber="118">Diagnosis</emphasis>
. MALE. Head angulate behind the eyes. Mandible tridentate, without subbasal tooth beneath.Clypeus rather flattened, anterior border with two medial tubercles. Ocelli small, POD less than OOD. Scape distinctly bicarinate beneath, F1 equal or slightly longer its width and evidently shorter than F2. Pronotum angulate. Notauli and parapsids developed in posterior half. Tegula slightly projected over mesoscuto-scutellar suture. Parascutal carina weak. Scutellum simple. Mesopleuron without precoxal ridge. Propodeum not shortened. Metacoxa mesally carinate. Wing well developed or shortened (in
<typeStatus box="[677,725,1694,1720]" pageId="17" pageNumber="118">type</typeStatus>
species), slightly infuscated, stigmatic cell approximately equal to distance between base of stigmatic cell and origin of
<emphasis box="[790,821,1730,1756]" italics="true" pageId="17" pageNumber="118">RS</emphasis>
on vein
<emphasis box="[922,948,1730,1756]" italics="true" pageId="17" pageNumber="118">Sc</emphasis>
. T2 with long lateral felt lines, S2 without any trace of felt lines. T7 and S8 punctured throughout. Volsella slightly longer than penial valves, ventrally with tuft of basal setae, which no longer than cuspis. FEMALE. The female of this genus is separated from the other
<taxonomicName authorityName="Bischoff" authorityYear="1920" box="[151,328,1838,1864]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="tribe" tribe="Smicromyrmini">Smicromyrmini</taxonomicName>
by F1 subequal F2. The female is similar to that of
<taxonomicName authorityName="Lelej" authorityYear="1995" box="[914,1023,1838,1864]" class="Insecta" family="Mutillidae" genus="Ephucilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis box="[914,1023,1838,1864]" italics="true" pageId="17" pageNumber="118">Ephucilla</emphasis>
</taxonomicName>
,
<taxonomicName box="[1037,1160,1838,1864]" class="Insecta" family="Mutillidae" genus="Promecilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis box="[1037,1160,1838,1864]" italics="true" pageId="17" pageNumber="118">Promecilla</emphasis>
</taxonomicName>
, and
<taxonomicName baseAuthorityName="Lelej" baseAuthorityYear="2005" box="[1222,1317,1838,1864]" class="Insecta" family="Mutillidae" genus="Sinotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis box="[1222,1317,1838,1864]" italics="true" pageId="17" pageNumber="118">Sinotilla</emphasis>
</taxonomicName>
by having the mandible slender and the pygidial plate convex and smooth, but differs by the mesosomal lateral margins subparallel or divergent posterad (convergent in
<taxonomicName authorityName="Lelej" authorityYear="1995" box="[699,808,1910,1936]" class="Insecta" family="Mutillidae" genus="Ephucilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis box="[699,808,1910,1936]" italics="true" pageId="17" pageNumber="118">Ephucilla</emphasis>
</taxonomicName>
and
<taxonomicName box="[863,986,1910,1936]" class="Insecta" family="Mutillidae" genus="Promecilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis box="[863,986,1910,1936]" italics="true" pageId="17" pageNumber="118">Promecilla</emphasis>
</taxonomicName>
), the scutellar scale obliterated (present in
<taxonomicName authorityName="Lelej" authorityYear="1995" box="[180,289,1946,1972]" class="Insecta" family="Mutillidae" genus="Ephucilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis box="[180,289,1946,1972]" italics="true" pageId="17" pageNumber="118">Ephucilla</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Lelej" baseAuthorityYear="2005" box="[342,437,1946,1972]" class="Insecta" family="Mutillidae" genus="Sinotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis box="[342,437,1946,1972]" italics="true" pageId="17" pageNumber="118">Sinotilla</emphasis>
</taxonomicName>
), and the lateral margins of pygidial plate convergent anterad and not carinate throughout (subparallel and with weak carinae posteriorly in
<taxonomicName authorityName="Lelej" authorityYear="1995" box="[706,815,1982,2008]" class="Insecta" family="Mutillidae" genus="Ephucilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis box="[706,815,1982,2008]" italics="true" pageId="17" pageNumber="118">Ephucilla</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph blockId="18.[151,1437,150,285]" pageId="18" pageNumber="119">
<emphasis bold="true" box="[199,520,150,177]" pageId="18" pageNumber="119">Diversity and distribution.</emphasis>
Nine species (one known from both sexes, one based on males only, and seven based on females only) are recognized from
<collectingCountry box="[652,712,186,212]" name="India" pageId="18" pageNumber="119">India</collectingCountry>
,
<collectingCountry box="[724,837,186,212]" name="Sri Lanka" pageId="18" pageNumber="119">Sri Lanka</collectingCountry>
,
<collectingCountry box="[848,950,186,212]" name="Thailand" pageId="18" pageNumber="119">Thailand</collectingCountry>
, and
<collectingCountry box="[1010,1119,186,212]" name="Indonesia" pageId="18" pageNumber="119">Indonesia</collectingCountry>
(
<bibRefCitation author="Lelej, A. S." box="[1134,1259,186,213]" pageId="18" pageNumber="119" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej 2005</bibRefCitation>
;
<bibRefCitation author="Williams, K. A. &amp; Lelej, A. S. &amp; Okayasu, J. &amp; Borkent, C. J. &amp; Malee, R. &amp; Thoawan, K. &amp; Thaochan, N." pageId="18" pageNumber="119" pagination="1 - 69" refId="ref30512" refString="Williams, K. A., Lelej, A. S., Okayasu, J., Borkent, C. J., Malee, R., Thoawan, K. &amp; Thaochan, N. (2019) The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia. Zootaxa, 4602 (1), 1 - 69. https: // doi. org / 10.11646 / zootaxa. 4602.1.1" type="journal article" year="2019">
Williams
<emphasis box="[1380,1437,186,212]" italics="true" pageId="18" pageNumber="119">et al.</emphasis>
2019
</bibRefCitation>
;
<bibRefCitation author="Okayasu, J." box="[222,388,222,248]" pageId="18" pageNumber="119" pagination="56 - 63" refId="ref28328" refString="Okayasu, J. (2022) Remarkable range extension of the genus Nordeniella Lelej, 2005 (Hymenoptera: Mutillidae: Smicromyrmini): Description of a new species from Sumbawa, Indonesia. Journal of Insect Biodiversity, 32 (2), 56 - 63. https: // doi. org / 10.12976 / jib / 2022.32.2.3" type="journal article" year="2022">Okayasu 2022</bibRefCitation>
).
<emphasis bold="true" box="[199,308,258,284]" pageId="18" pageNumber="119">Remarks</emphasis>
. There are no available keys to separate the species in this genus.
</paragraph>
</subSubSection>
</treatment>
</document>