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<document id="BA994A6EDDAC7A1F0338275F4FC3E7E5" ID-DOI="10.11646/zootaxa.5023.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5225402" ID-ZooBank="F6CB0172-C3B9-4E9F-ACBF-2BC006F588F5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629423656372" checkinUser="plazi" docAuthor="Ito, Tomiko" docDate="2021" docId="03CF878F4F2EFFFA68A81E44FD2FFD84" docLanguage="en" docName="zootaxa.5023.1.2.pdf" docOrigin="Zootaxa 5023 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophila yakushima Ito 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="47" masterDocId="FFF6FFF74F2FFFF9683F1A21FFE8F819" masterDocTitle="The genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) in Yakushima Island, northern Ryukyu, southwestern Japan" masterLastPageNumber="58" masterPageNumber="44" pageNumber="45" updateTime="1699072484263" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="AA5CBD916470EF60CAC0AFC5C9AEFFF5">The genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) in Yakushima Island, northern Ryukyu, southwestern Japan</mods:title>
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<mods:namePart id="11B3C3C846A94B4905D8701646AED1C0">Ito, Tomiko</mods:namePart>
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<paragraph id="8BD936994F2EFFF868A81E44FDC9FC99" blockId="1.[151,545,1125,1189]" box="[151,545,1125,1152]" pageId="1" pageNumber="45">
<heading id="D09181F54F2EFFF868A81E44FDC9FC99" bold="true" box="[151,545,1125,1152]" fontSize="11" level="1" pageId="1" pageNumber="45" reason="1">
<emphasis id="B912EA8B4F2EFFF868A81E44FDC9FC99" bold="true" box="[151,545,1125,1152]" pageId="1" pageNumber="45">
<taxonomicName id="4C664D1A4F2EFFF868A81E44FE54FC99" authority="Ito, 2021" authorityName="Ito" authorityYear="2021" box="[151,444,1125,1152]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="yakushima" status="sp. nov.">
<emphasis id="B912EA8B4F2EFFF868A81E44FE54FC99" bold="true" box="[151,444,1125,1152]" italics="true" pageId="1" pageNumber="45">Rhyacophila yakushima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157F04F2EFFF869FC1E47FDC9FC99" box="[451,545,1126,1152]" pageId="1" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C37C65124F2EFFF868A81EABFD2DFCFF" pageId="1" pageNumber="45" type="description">
<paragraph id="8BD936994F2EFFF868A81EABFEF7FCBC" blockId="1.[151,545,1125,1189]" box="[151,287,1162,1189]" pageId="1" pageNumber="45">
(
<figureCitation id="135D2A1C4F2EFFF868A01EABFF00FCBD" box="[159,232,1162,1188]" captionStart="FIGURE 2" captionStartId="2.[151,250,1777,1801]" captionTargetBox="[210,1371,200,1741]" captionTargetId="figure-16@2.[201,1386,181,1753]" captionTargetPageId="2" captionText="FIGURE 2. Rhyacophila yakushima sp. nov. 2A2E, male: 2A, genitalia, left lateral; 2B, same, dorsal; 2C, same, caudal, omitting inferior appendages; 2D, phallic apparatus, ventral, with enlarged figure of apical part of a paramere; 2E, right inferior appendage, dorso-mesal.2F2J, female: 2F, abdominal segments VIIIXI, left lateral; 2G, same, ventral; 2H, abdominal segment VIII, dorsal; 2I, vaginal apparatus, left lateral; 2J, same, ventral. Abbreviations: VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ad l = apicodorsal lobe; ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band; b inf app = basal segment of inferior appendage (paired); do pr = dorsal process; pa = paramere (paired); ph = phallotheca; pre app = preanal appendage; pp = posterior process; ps = processus spermathecae." figureDoi="http://doi.org/10.5281/zenodo.5225406" httpUri="https://zenodo.org/record/5225406/files/figure.png" pageId="1" pageNumber="45">Figs 2</figureCitation>
,
<figureCitation id="135D2A1C4F2EFFF868CB1EABFEFFFCBC" box="[244,279,1162,1189]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[232,1361,183,1919]" captionTargetId="figure-16@12.[225,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Adult collection sites of seven species of Rhyacophila on Yakushima. Solid circle = 14 adults collected; open circle = more than 5 adults collected." figureDoi="http://doi.org/10.5281/zenodo.5225422" httpUri="https://zenodo.org/record/5225422/files/figure.png" pageId="1" pageNumber="45">6A</figureCitation>
)
</paragraph>
<paragraph id="8BD936994F2EFFF868A81EEFFD2DFCFF" blockId="1.[151,709,1230,1255]" box="[151,709,1230,1255]" pageId="1" pageNumber="45">
<taxonomicName id="4C664D1A4F2EFFF868A81EEFFEB9FCFE" box="[151,337,1230,1255]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="undefined-2">
<emphasis id="B912EA8B4F2EFFF868A81EEFFEFFFCFF" box="[151,279,1230,1254]" italics="true" pageId="1" pageNumber="45">Rhyacophila</emphasis>
sp. 2
</taxonomicName>
:
<bibRefCitation id="EFF74B684F2EFFF869641EEFFDF5FCFE" author="Kuhara, N. &amp; Ito, T." box="[347,541,1230,1255]" pageId="1" pageNumber="45" pagination="11 - 20" refId="ref9051" refString="Kuhara, N. &amp; Ito, T. (2017) Caddisflies (Trichoptera) of Yakushima, northern Ryukyus, southwestern Japan. Biology of Inland Waters, 31, 11 - 20. [in Japanese with English abstract]" type="journal article" year="2017">Kuhara &amp; Ito 2017</bibRefCitation>
, 16, Yakushima.
</paragraph>
</subSubSection>
<subSubSection id="C37C65124F2EFFFA68A81F35FEB7FB20" lastPageId="3" lastPageNumber="47" pageId="1" pageNumber="45" type="diagnosis">
<paragraph id="8BD936994F2EFFF868A81F35FB8CFE32" blockId="1.[151,1437,1300,2011]" pageId="1" pageNumber="45">
<emphasis id="B912EA8B4F2EFFF868A81F35FEE2FD37" bold="true" box="[151,266,1300,1326]" pageId="1" pageNumber="45">Diagnosis</emphasis>
. This new species belongs to the
<taxonomicName id="4C664D1A4F2EFFF86AAC1F34FCE3FD37" authorityName="KJ Morton" authorityYear="1900" box="[659,779,1301,1326]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="anatina">
<emphasis id="B912EA8B4F2EFFF86AAC1F34FCE3FD37" box="[659,779,1301,1326]" italics="true" pageId="1" pageNumber="45">R. anatina</emphasis>
</taxonomicName>
Species Group and the male resembles that of
<taxonomicName id="4C664D1A4F2EFFF86D111F34FEF7FD4A" authority="Tsuda 1940" authorityName="Tsuda" authorityYear="1940" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="pacata">
<emphasis id="B912EA8B4F2EFFF86D111F34FA74FD37" box="[1326,1436,1301,1326]" italics="true" pageId="1" pageNumber="45">R. pacata</emphasis>
<bibRefCitation id="EFF74B684F2EFFF868A81F18FEF7FD4A" author="Tsuda, M." box="[151,287,1337,1363]" pageId="1" pageNumber="45" pagination="119 - 135" refId="ref9796" refString="Tsuda, M. (1940) Zur Kenntnis der Japanischen Rhyacophilinen (Rhyacophilidae, Trichoptera). Annotationes Zoologicae Japonenses, 19, 119 - 135. [in German]" type="journal article" year="1940">Tsuda 1940</bibRefCitation>
</taxonomicName>
, found in
<collectingCountry id="F37176094F2EFFF869A61F18FE31FD4A" box="[409,473,1337,1363]" name="Japan" pageId="1" pageNumber="45">Japan</collectingCountry>
(Honshu), in having large preanal appendages and a long and thick basal segment and short sock-like apical segment of each inferior appendage (
<bibRefCitation id="EFF74B684F2EFFF86B551F7CFC1AFD6E" author="Tsuda, M." box="[874,1010,1373,1399]" pageId="1" pageNumber="45" pagination="119 - 135" refId="ref9796" refString="Tsuda, M. (1940) Zur Kenntnis der Japanischen Rhyacophilinen (Rhyacophilidae, Trichoptera). Annotationes Zoologicae Japonenses, 19, 119 - 135. [in German]" type="journal article" year="1940">Tsuda 1940</bibRefCitation>
;
<bibRefCitation id="EFF74B684F2EFFF86BC01F7CFB91FD6E" author="Ross, H. H." box="[1023,1145,1372,1399]" pageId="1" pageNumber="45" refId="ref9465" refString="Ross, H. H. (1956) Evolution and Classification of Mountain Caddisflies. University of Illinois Press, Urbana, Illinois, 213 pp." type="book" year="1956">Ross 1956</bibRefCitation>
;
<bibRefCitation id="EFF74B684F2EFFF86CB81F7CFAF4FD6E" author="Hattori, T." box="[1159,1308,1372,1399]" pageId="1" pageNumber="45" pagination="414 - 437" refId="ref8732" refString="Hattori, T. (2005) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations. Tokai University Press, Kanagawa, pp. 414 - 437. [in Japanese]" type="book chapter" year="2005">Hattori 2005</bibRefCitation>
). However, the male of
<taxonomicName id="4C664D1A4F2EFFF8691D1FA0FE54FD82" authorityName="Ito" authorityYear="2021" box="[290,444,1409,1435]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="yakushima">
<emphasis id="B912EA8B4F2EFFF8691D1FA0FE54FD82" box="[290,444,1409,1435]" italics="true" pageId="1" pageNumber="45">R. yakushima</emphasis>
</taxonomicName>
is clearly distinguished from
<taxonomicName id="4C664D1A4F2EFFF86B2B1FA0FC69FD83" authorityName="Tsuda" authorityYear="1940" box="[788,897,1409,1434]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="pacata">
<emphasis id="B912EA8B4F2EFFF86B2B1FA0FC69FD83" box="[788,897,1409,1434]" italics="true" pageId="1" pageNumber="45">R. pacata</emphasis>
</taxonomicName>
as follows: The preanal appendages are nearly rectangular in lateral view and the apicodorsal lobe of segment IX is present in
<taxonomicName id="4C664D1A4F2EFFF86BC11F84FB7BFDA6" authorityName="Ito" authorityYear="2021" box="[1022,1171,1445,1471]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="yakushima">
<emphasis id="B912EA8B4F2EFFF86BC11F84FB7BFDA6" box="[1022,1171,1445,1471]" italics="true" pageId="1" pageNumber="45">R. yakushima</emphasis>
</taxonomicName>
; but the appendages are long and triangular in lateral view and the lobe of segment IX is undeveloped in
<taxonomicName id="4C664D1A4F2EFFF86C1F1FE8FB64FDFB" authorityName="Tsuda" authorityYear="1940" box="[1056,1164,1481,1506]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="pacata">
<emphasis id="B912EA8B4F2EFFF86C1F1FE8FB64FDFB" box="[1056,1164,1481,1506]" italics="true" pageId="1" pageNumber="45">R. pacata</emphasis>
</taxonomicName>
. The female of this new species is also similar to that of
<taxonomicName id="4C664D1A4F2EFFF86A3D1FCCFCF1FE1E" authority="(Hattori 2005)" baseAuthorityName="Hattori" baseAuthorityYear="2005" box="[514,793,1516,1543]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="pacata">
<emphasis id="B912EA8B4F2EFFF86A3D1FCCFD85FE1F" box="[514,621,1517,1542]" italics="true" pageId="1" pageNumber="45">R. pacata</emphasis>
(
<bibRefCitation id="EFF74B684F2EFFF86A431FCCFCF9FE1E" author="Hattori, T." box="[636,785,1516,1543]" pageId="1" pageNumber="45" pagination="414 - 437" refId="ref8732" refString="Hattori, T. (2005) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations. Tokai University Press, Kanagawa, pp. 414 - 437. [in Japanese]" type="book chapter" year="2005">Hattori 2005</bibRefCitation>
)
</taxonomicName>
, but clearly discriminated from the latter by the presence of paired subquadrate sclerites at the base of the intersegmental membranes of VIIIX.
</paragraph>
<paragraph id="8BD936994F2EFFF868F81C15FDEDFF76" blockId="1.[151,1437,1300,2011]" pageId="1" pageNumber="45">
<emphasis id="B912EA8B4F2EFFF868F81C15FEE3FE57" bold="true" box="[199,267,1588,1614]" pageId="1" pageNumber="45">Adult</emphasis>
. Lengths of forewings and hind wings: Each forewing
<quantity id="4C9E9B7C4F2EFFF86BB21C14FBFFFE56" box="[909,1047,1589,1615]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.2" metricValueMin="6.8" pageId="1" pageNumber="45" unit="mm" value="8.0" valueMax="9.2" valueMin="6.8">6.89.2 mm</quantity>
long (mean
<quantity id="4C9E9B7C4F2EFFF86C941C14FAEEFE56" box="[1195,1286,1589,1615]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="45" unit="mm" value="8.1">8.1 mm</quantity>
, n = 21) and each hind wing 6.0
<quantity id="4C9E9B7C4F2EFFF869431C78FE39FE6A" box="[380,465,1625,1651]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="1" pageNumber="45" unit="mm" value="8.3">8.3 mm</quantity>
(mean
<quantity id="4C9E9B7C4F2EFFF86A1A1C78FD95FE6A" box="[549,637,1625,1651]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="45" unit="mm" value="7.1">7.1 mm</quantity>
, n = 21) in males, 8.0
<quantity id="4C9E9B7C4F2EFFF86BBF1C78FC0BFE6A" box="[896,995,1625,1651]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="45" unit="mm" value="10.2">10.2 mm</quantity>
(mean
<quantity id="4C9E9B7C4F2EFFF86C081C79FB67FE6A" box="[1079,1167,1624,1651]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="45" unit="mm" value="9.5">9.5 mm</quantity>
, n = 8) and 7.39.0 mm (mean
<quantity id="4C9E9B7C4F2EFFF868D91C5CFEA9FE8E" box="[230,321,1661,1687]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="45" unit="mm" value="8.1">8.1 mm</quantity>
, n = 8) in females, respectively. Head dark brown, warts light brown with brown setae, antennae dark brown, scapes thicker and longer than other segments; palpi brown. Thorax dark brown dorsally. Legs brown with dark brown spurs. Wings brown with darker veins and fulvous pterostigma, whitish dots scattered on apical 2/3 of forewings in both sexes. Abdomen with dark brown tergites and light brown sternites, dark pigments scattered dorsally; scent glands of sternite V open on antero-angle mounds well encircled by sutures; small mid-ventral process on sternite VII (or VI and VII) in male and on sternite VI in female, process on sternite VI of male often very small or completely absent.
</paragraph>
<paragraph id="8BD936994F2EFFFA68F81D58FC12F958" blockId="1.[151,1437,1300,2011]" lastBlockId="3.[151,1437,151,1437]" lastPageId="3" lastPageNumber="47" pageId="1" pageNumber="45">
<emphasis id="B912EA8B4F2EFFF868F81D58FE9DFF8B" bold="true" box="[199,373,1912,1939]" pageId="1" pageNumber="45">Male genitalia</emphasis>
(
<figureCitation id="135D2A1C4F2EFFF869B91D58FDFDFF8A" box="[390,533,1913,1939]" captionStart="FIGURE 2" captionStartId="2.[151,250,1777,1801]" captionTargetBox="[210,1371,200,1741]" captionTargetId="figure-16@2.[201,1386,181,1753]" captionTargetPageId="2" captionText="FIGURE 2. Rhyacophila yakushima sp. nov. 2A2E, male: 2A, genitalia, left lateral; 2B, same, dorsal; 2C, same, caudal, omitting inferior appendages; 2D, phallic apparatus, ventral, with enlarged figure of apical part of a paramere; 2E, right inferior appendage, dorso-mesal.2F2J, female: 2F, abdominal segments VIIIXI, left lateral; 2G, same, ventral; 2H, abdominal segment VIII, dorsal; 2I, vaginal apparatus, left lateral; 2J, same, ventral. Abbreviations: VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ad l = apicodorsal lobe; ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band; b inf app = basal segment of inferior appendage (paired); do pr = dorsal process; pa = paramere (paired); ph = phallotheca; pre app = preanal appendage; pp = posterior process; ps = processus spermathecae." figureDoi="http://doi.org/10.5281/zenodo.5225406" httpUri="https://zenodo.org/record/5225406/files/figure.png" pageId="1" pageNumber="45">Figs 2A2E</figureCitation>
). In lateral view segment IX (IX) short ventrally, long dorsally, dorsal margin about 5 times as long as ventral margin, apicodorsal lobe (ad l) almost 2/3 as long as lower margin of basal segment of inferior appendage (b inf app); in dorsal view anterior margin of segment IX roundly incised, apicodorsal lobe deeply incised at apex. Preanal appendages (pre app) large, slightly longer than apicodorsal lobe, in lateral view rectangular and slightly tapered posteriorly, round apically; in dorsal view slightly narrowing apically with nearly V-shaped divergence apically. Segment X (X) little reduced, below basomesal edges of preanal appendages, slightly sclerotized, connected with anal sclerites (an sc). Anal sclerites heavily sclerotized, bowl-shaped, covered with spinules. Apical band (ap ba) small, semi-membranous, round in lateral view.
</paragraph>
<caption id="DF1966114F2DFFFB68A81CD0FB8CFFF8" ID-DOI="http://doi.org/10.5281/zenodo.5225406" ID-Zenodo-Dep="5225406" httpUri="https://zenodo.org/record/5225406/files/figure.png" pageId="2" pageNumber="46" startId="2.[151,250,1777,1801]" targetBox="[210,1371,200,1741]" targetPageId="2">
<paragraph id="8BD936994F2DFFFB68A81CD0FB8CFFF8" blockId="2.[151,1436,1777,2017]" pageId="2" pageNumber="46">
<emphasis id="B912EA8B4F2DFFFB68A81CD0FEF8FF10" bold="true" box="[151,272,1777,1801]" pageId="2" pageNumber="46">FIGURE 2</emphasis>
.
<taxonomicName id="4C664D1A4F2DFFFB69201CD0FDFCFF10" authority="Ito, 2021" authorityName="Ito" authorityYear="2021" box="[287,532,1777,1801]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="46" phylum="Arthropoda" rank="species" species="yakushima" status="sp. nov.">
<emphasis id="B912EA8B4F2DFFFB69201CD0FDFCFF10" box="[287,532,1777,1801]" italics="true" pageId="2" pageNumber="46">Rhyacophila yakushima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157F04F2DFFFB6A211CD0FD9BFF10" box="[542,627,1777,1801]" pageId="2" pageNumber="46" rank="species">
<emphasis id="B912EA8B4F2DFFFB6A211CD0FD84FF10" bold="true" box="[542,620,1777,1801]" pageId="2" pageNumber="46">sp. nov</emphasis>
.
</taxonomicNameLabel>
2A2E, male: 2A, genitalia, left lateral; 2B, same, dorsal; 2C, same, caudal, omitting inferior appendages; 2D, phallic apparatus, ventral, with enlarged figure of apical part of a paramere; 2E, right inferior appendage, dorso-mesal. 2F2J, female: 2F, abdominal segments VIIIXI, left lateral; 2G, same, ventral; 2H, abdominal segment VIII, dorsal; 2I, vaginal apparatus, left lateral; 2J, same, ventral. Abbreviations: VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ad l = apicodorsal lobe; ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band; b inf app = basal segment of inferior appendage (paired); do pr = dorsal process; pa = paramere (paired); ph = phallotheca; pre app = preanal appendage; pp = posterior process; ps = processus spermathecae.
</paragraph>
</caption>
<paragraph id="8BD936994F2CFFFA68F81B6AFDFDF9C8" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="47">Phallic apparatus complex; phallotheca (ph) in lateral view somewhat rectangular; dorsal process (do pr) elongate, in ventral and lateral views almost parallel-sided with abruptly tapered and subacute apex; aedeagus (ae) slender, sub-truncate apically in lateral and ventral views; pair of parameres (pa) slender with numerous spindles on apical half, shorter than aedeagus.</paragraph>
<paragraph id="8BD936994F2CFFFA68F81BFAFD68FA78" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="47">Basal segment of each inferior appendage (b inf app) long, in lateral view with dorsal and ventral margins curved caudad and parallel to each other, anterior margin oblique, posterior margin subtruncate, in dorsal view appendages curved mesad; apical segment of each inferior appendage (a inf app) in lateral view somewhat socklike, mesal sub-marginal part with spinules.</paragraph>
<paragraph id="8BD936994F2CFFFA68F8184AFEB7FB20" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="47">
<emphasis id="B912EA8B4F2CFFFA68F8184AFE65FA9D" bold="true" box="[199,397,618,645]" pageId="3" pageNumber="47">Female genitalia</emphasis>
(
<figureCitation id="135D2A1C4F2CFFFA69A3184AFDF7FA9C" box="[412,543,619,645]" captionStart="FIGURE 2" captionStartId="2.[151,250,1777,1801]" captionTargetBox="[210,1371,200,1741]" captionTargetId="figure-16@2.[201,1386,181,1753]" captionTargetPageId="2" captionText="FIGURE 2. Rhyacophila yakushima sp. nov. 2A2E, male: 2A, genitalia, left lateral; 2B, same, dorsal; 2C, same, caudal, omitting inferior appendages; 2D, phallic apparatus, ventral, with enlarged figure of apical part of a paramere; 2E, right inferior appendage, dorso-mesal.2F2J, female: 2F, abdominal segments VIIIXI, left lateral; 2G, same, ventral; 2H, abdominal segment VIII, dorsal; 2I, vaginal apparatus, left lateral; 2J, same, ventral. Abbreviations: VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ad l = apicodorsal lobe; ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band; b inf app = basal segment of inferior appendage (paired); do pr = dorsal process; pa = paramere (paired); ph = phallotheca; pre app = preanal appendage; pp = posterior process; ps = processus spermathecae." figureDoi="http://doi.org/10.5281/zenodo.5225406" httpUri="https://zenodo.org/record/5225406/files/figure.png" pageId="3" pageNumber="47">Figs 2F2J</figureCitation>
). Segment VIII (VIII) large, length as long as basal width, gradually narrowing posteriorly in lateral view, parallel sided in dorsal and ventral views; posterior margin with shallow and wide incision dorsally and deep round incision ventrally. Small subquadrate sclerites at base of intersegmental membranes of segments VIIIX ventrally. Vaginal apparatus long; processus spermathecae (ps) round in ventral view, semimembranous; posterior process (pp) heavily sclerotized, long, tapered, 2.5 times as long as processus spermathecae, subacute apically.
</paragraph>
</subSubSection>
<subSubSection id="C37C65124F2CFFFA68F81962FE97FCF3" pageId="3" pageNumber="47" type="materials_examined">
<paragraph id="8BD936994F2CFFFA68F81962FD41FB98" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="47">
<materialsCitation id="3B0E3CC44F2CFFFA68F81962FD41FB98" collectingDate="2007-06-12" collectionCode="TI" country="Japan" county="Ara-kawa" elevation="1100" location="Male" municipality="Yakusugi-land" pageId="3" pageNumber="47" specimenCode="CBM-ZI 0159996" specimenCount="1" typeStatus="holotype">
<emphasis id="B912EA8B4F2CFFFA68F81962FEDCFB44" bold="true" box="[199,308,835,861]" pageId="3" pageNumber="47">
<typeStatus id="54DD883B4F2CFFFA68F81962FEDCFB44" box="[199,308,835,861]" pageId="3" pageNumber="47" type="holotype">Holotype</typeStatus>
</emphasis>
.
<location id="8EB960424F2CFFFA697E1962FE95FB44" LSID="urn:lsid:plazi:treatment:03CF878F4F2EFFFA68A81E44FD2FFD84:8EB960424F2CFFFA697E1962FE95FB44" box="[321,381,835,861]" country="Japan" county="Ara-kawa" municipality="Yakusugi-land" name="Male" pageId="3" pageNumber="47">Male</location>
,
<emphasis id="B912EA8B4F2CFFFA69B71962FE39FB44" bold="true" box="[392,465,835,861]" pageId="3" pageNumber="47">
<collectingCountry id="F37176094F2CFFFA69B71962FE39FB44" box="[392,465,835,861]" name="Japan" pageId="3" pageNumber="47">Japan</collectingCountry>
</emphasis>
,
<emphasis id="B912EA8B4F2CFFFA69E11963FD5DFB44" bold="true" box="[478,693,834,861]" pageId="3" pageNumber="47">
<location id="8EB960424F2CFFFA69E11963FD5DFB44" LSID="urn:lsid:plazi:treatment:03CF878F4F2EFFFA68A81E44FD2FFD84:8EB960424F2CFFFA69E11963FD5DFB44" box="[478,693,834,861]" country="Japan" county="Ara-kawa" municipality="Yakusugi-land" name="Yakushima Island" pageId="3" pageNumber="47">Yakushima Island</location>
</emphasis>
,
<collectingMunicipality id="6BBDACE34F2CFFFA6AFF1962FC8DFB44" box="[704,869,835,861]" pageId="3" pageNumber="47">Yakusugi-land</collectingMunicipality>
,
<collectingCounty id="62B84E154F2CFFFA6B511962FC37FB44" box="[878,991,835,861]" pageId="3" pageNumber="47">Ara-kawa</collectingCounty>
, a tributary (30.302˚N, 130.540˚E,
<elevation id="004BD1AA4F2CFFFA6D5A1962FF01FB9B" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="47" unit="m" value="1100.0">
<quantity id="4C9E9B7C4F2CFFFA6D5A1962FF45FB98" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="47" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
),
<date id="FFD810594F2CFFFA68C01946FE92FB98" box="[255,378,871,897]" pageId="3" pageNumber="47" value="2007-06-12">
<collectingDate id="EF9CE9B14F2CFFFA68C01946FE92FB98" box="[255,378,871,897]" pageId="3" pageNumber="47" value="2007-06-12">12.vi.2007</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA69BA1946FE4AFB98" box="[389,418,871,897]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="3" pageNumber="47">TI</collectionCode>
, S (
<specimenCode id="DBC09EE24F2CFFFA69F41946FD75FB98" box="[459,669,870,897]" collectionCode="CBM-ZI" pageId="3" pageNumber="47">CBM-ZI 0159996</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD936994F2CFFFA68F819AAFCF3FC64" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="47">
<materialsCitation id="3B0E3CC44F2CFFFA68F819AAFD43FBBC" box="[199,683,906,933]" collectionCode="SPMN-IS" location="Yakushima Island" pageId="3" pageNumber="47" specimenCount="1" typeStatus="paratype">
<emphasis id="B912EA8B4F2CFFFA68F819AAFED7FBBC" bold="true" box="[199,319,907,933]" pageId="3" pageNumber="47">
<typeStatus id="54DD883B4F2CFFFA68F819AAFED7FBBC" box="[199,319,907,933]" pageId="3" pageNumber="47" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<collectionCode id="ED77AE5C4F2CFFFA697219AAFE2AFBBC" box="[333,450,907,933]" pageId="3" pageNumber="47">SPMN-IS</collectionCode>
).
<emphasis id="B912EA8B4F2CFFFA69EC19ABFD43FBBC" bold="true" box="[467,683,906,933]" pageId="3" pageNumber="47">
<location id="8EB960424F2CFFFA69EC19ABFD43FBBC" LSID="urn:lsid:plazi:treatment:03CF878F4F2EFFFA68A81E44FD2FFD84:8EB960424F2CFFFA69EC19ABFD43FBBC" box="[467,683,906,933]" name="Yakushima Island" pageId="3" pageNumber="47">Yakushima Island</location>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B0E3CC44F2CFFFA6A8619AAFB67FBBC" box="[697,1167,907,933]" collectingDate="2007-06-12" collectionCode="TI" country="Japan" county="Ara-kawa" elevation="1100" location="Male" municipality="Yakusugi-land" pageId="3" pageNumber="47" specimenCount="12" specimenCount-female="6" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="9D60FD104F2CFFFA6A8619AAFCFAFBBC" box="[697,786,907,933]" pageId="3" pageNumber="47" type="male">6 males</specimenCount>
,
<specimenCount id="9D60FD104F2CFFFA6B2219AAFC63FBBC" box="[797,907,907,933]" pageId="3" pageNumber="47" type="female">6 females</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="3B0E3CC44F2CFFFA6CA519AAFE7CFBD0" collectingDate="2007-10-16" collectionCode="TI" country="Japan" pageId="3" pageNumber="47" specimenCount="8" specimenCount-female="1" specimenCount-male="7" typeStatus="paratype">
<specimenCount id="9D60FD104F2CFFFA6CA519AAFB1BFBBC" box="[1178,1267,907,933]" pageId="3" pageNumber="47" type="male">7 males</specimenCount>
,
<specimenCount id="9D60FD104F2CFFFA6CC119AAFA89FBBC" box="[1278,1377,907,933]" pageId="3" pageNumber="47" type="female">1 female</specimenCount>
, type locality,
<date id="FFD810594F2CFFFA68C6198EFE83FBD0" box="[249,363,943,969]" pageId="3" pageNumber="47" value="2007-10-16">
<collectingDate id="EF9CE9B14F2CFFFA68C6198EFE83FBD0" box="[249,363,943,969]" pageId="3" pageNumber="47" value="2007-10-16">16.x.2007</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA6948198EFE7CFBD0" box="[375,404,943,969]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="3" pageNumber="47">TI</collectionCode>
</materialsCitation>
, S;
<materialsCitation id="3B0E3CC44F2CFFFA69FE198EFF5CFBF4" collectingDate="2007-06-12" collectionCode="TI" country="Japan" pageId="3" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D60FD104F2CFFFA69FE198EFDF8FBD0" box="[449,528,943,969]" pageId="3" pageNumber="47" type="male">1 male</specimenCount>
, Yakusugi-land, Ara-kawa, different tributary from the type locality,
<date id="FFD810594F2CFFFA6D22198EFA70FBD0" box="[1309,1432,943,969]" pageId="3" pageNumber="47" value="2007-06-12">
<collectingDate id="EF9CE9B14F2CFFFA6D22198EFA70FBD0" box="[1309,1432,943,969]" pageId="3" pageNumber="47" value="2007-06-12">12.vi.2007</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA68A819F2FF5CFBF4" box="[151,180,979,1005]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="3" pageNumber="47">TI</collectionCode>
</materialsCitation>
, S;
<materialsCitation id="3B0E3CC44F2CFFFA68DD19F2FC59FBF4" box="[226,945,979,1005]" collectingDate="2007-06-12" collectionCode="TI" country="Japan" pageId="3" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D60FD104F2CFFFA68DD19F2FEDAFBF4" box="[226,306,979,1005]" pageId="3" pageNumber="47" type="male">1 male</specimenCount>
, Yakusugi-land, Ara-kawa, small stream,
<date id="FFD810594F2CFFFA6B3219F2FC6FFBF4" box="[781,903,979,1005]" pageId="3" pageNumber="47" value="2007-06-12">
<collectingDate id="EF9CE9B14F2CFFFA6B3219F2FC6FFBF4" box="[781,903,979,1005]" pageId="3" pageNumber="47" value="2007-06-12">12.vi.2007</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA6BAB19F2FC59FBF4" box="[916,945,979,1005]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="3" pageNumber="47">TI</collectionCode>
</materialsCitation>
, S;
<materialsCitation id="3B0E3CC44F2CFFFA6BE119F2FDCEFC08" collectingDate="2007-10-16" collectionCode="TI" country="Japan" pageId="3" pageNumber="47" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D60FD104F2CFFFA6BE119F2FBD1FBF4" box="[990,1081,979,1005]" pageId="3" pageNumber="47" type="male">2 males</specimenCount>
,
<specimenCount id="9D60FD104F2CFFFA6C7819F2FB44FBF4" box="[1095,1196,979,1005]" pageId="3" pageNumber="47" type="female">1 female</specimenCount>
, Yakusugi-land, Arakawa, Seiryu-bashi,
<date id="FFD810594F2CFFFA69B919D6FE10FC08" box="[390,504,1015,1041]" pageId="3" pageNumber="47" value="2007-10-16">
<collectingDate id="EF9CE9B14F2CFFFA69B919D6FE10FC08" box="[390,504,1015,1041]" pageId="3" pageNumber="47" value="2007-10-16">16.x.2007</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA6A3619D6FDCEFC08" box="[521,550,1015,1041]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="3" pageNumber="47">TI</collectionCode>
</materialsCitation>
, S;
<specimenCount id="9D60FD104F2CFFFA6A6519D6FD45FC08" box="[602,685,1015,1041]" pageId="3" pageNumber="47" type="male">1 male</specimenCount>
, Shiratani-unsuikyo,
<date id="FFD810594F2CFFFA6B9919D6FBF6FC08" box="[934,1054,1015,1041]" pageId="3" pageNumber="47" value="2003-04-24">24.iv.2003</date>
, A. Ishizuka, M;
<materialsCitation id="3B0E3CC44F2CFFFA6CCB19D6FE6AFC2C" collectingDate="2006-05-09" collectionCode="TI" country="Japan" pageId="3" pageNumber="47" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D60FD104F2CFFFA6CCB19D6FABAFC08" box="[1268,1362,1015,1041]" pageId="3" pageNumber="47" type="male">3 males</specimenCount>
, same locality,
<date id="FFD810594F2CFFFA68C81E3AFEB2FC2C" box="[247,346,1051,1077]" pageId="3" pageNumber="47" value="2006-05-09">
<collectingDate id="EF9CE9B14F2CFFFA68C81E3AFEB2FC2C" box="[247,346,1051,1077]" pageId="3" pageNumber="47" value="2006-05-09">9.v.2006</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA695A1E3AFE6AFC2C" box="[357,386,1051,1077]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="3" pageNumber="47">TI</collectionCode>
</materialsCitation>
, S;
<materialsCitation id="3B0E3CC44F2CFFFA69931E3AFBE8FC2C" box="[428,1024,1051,1077]" collectingDate="2006-05-08" collectionCode="TI" country="Japan" pageId="3" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D60FD104F2CFFFA69931E3AFE12FC2C" box="[428,506,1051,1077]" pageId="3" pageNumber="47" type="male">1 male</specimenCount>
, Shiratani-gawa, Shiratani-bashi,
<date id="FFD810594F2CFFFA6B4A1E3AFC30FC2C" box="[885,984,1051,1077]" pageId="3" pageNumber="47" value="2006-05-08">
<collectingDate id="EF9CE9B14F2CFFFA6B4A1E3AFC30FC2C" box="[885,984,1051,1077]" pageId="3" pageNumber="47" value="2006-05-08">8.v.2006</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA6BDC1E3AFBE8FC2C" box="[995,1024,1051,1077]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="3" pageNumber="47">TI</collectionCode>
</materialsCitation>
, P;
<materialsCitation id="3B0E3CC44F2CFFFA6C151E3AFE1CFC40" collectingDate="2006-05-08" collectionCode="TI" country="Japan" pageId="3" pageNumber="47" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D60FD104F2CFFFA6C151E3AFB66FC2C" box="[1066,1166,1051,1077]" pageId="3" pageNumber="47" type="female">1 female</specimenCount>
, Miyanoura, Kue-gawa, Biwatsubo-bashi,
<date id="FFD810594F2CFFFA695A1E1EFE21FC40" box="[357,457,1087,1113]" pageId="3" pageNumber="47" value="2006-05-08">
<collectingDate id="EF9CE9B14F2CFFFA695A1E1EFE21FC40" box="[357,457,1087,1113]" pageId="3" pageNumber="47" value="2006-05-08">8.v.2006</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA69E81E1EFE1CFC40" box="[471,500,1087,1113]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="3" pageNumber="47">TI</collectionCode>
</materialsCitation>
, L;
<materialsCitation id="3B0E3CC44F2CFFFA6A171E1EFAB4FC40" box="[552,1372,1087,1113]" collectingDate="2007-10-17" collectionCode="TI" country="Japan" pageId="3" pageNumber="47" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D60FD104F2CFFFA6A171E1EFD6DFC40" box="[552,645,1087,1113]" pageId="3" pageNumber="47" type="male">2 males</specimenCount>
,
<specimenCount id="9D60FD104F2CFFFA6AAB1E1EFD14FC40" box="[660,764,1087,1113]" pageId="3" pageNumber="47" type="female">1 female</specimenCount>
, Anbo-rindo, Harumaki-rokugo-bashi,
<date id="FFD810594F2CFFFA6C811E1EFAD8FC40" box="[1214,1328,1087,1113]" pageId="3" pageNumber="47" value="2007-10-17">
<collectingDate id="EF9CE9B14F2CFFFA6C811E1EFAD8FC40" box="[1214,1328,1087,1113]" pageId="3" pageNumber="47" value="2007-10-17">17.x.2007</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA6D001E1EFAB4FC40" box="[1343,1372,1087,1113]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="3" pageNumber="47">TI</collectionCode>
</materialsCitation>
, S;
<specimenCount id="9D60FD104F2CFFFA6DB11E1EFF33FC64" pageId="3" pageNumber="47" type="male">2 males</specimenCount>
, Onoaida, Onoaida-gyoko,
<date id="FFD810594F2CFFFA6A2F1E42FD60FC64" box="[528,648,1123,1149]" pageId="3" pageNumber="47" value="2003-04-24">24.iv.2003</date>
, A. Ishizuka.
</paragraph>
<paragraph id="8BD936994F2CFFFA68F81EA7FE97FCF3" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="47">
<emphasis id="B912EA8B4F2CFFFA68F81EA7FE79FCB9" bold="true" box="[199,401,1158,1184]" pageId="3" pageNumber="47">Other specimens</emphasis>
.
<emphasis id="B912EA8B4F2CFFFA699D1EA7FD94FCB8" bold="true" box="[418,636,1158,1185]" pageId="3" pageNumber="47">Yakushima Island</emphasis>
:
<materialsCitation id="3B0E3CC44F2CFFFA6AB11EA6FAD0FCB8" box="[654,1336,1159,1185]" collectingDate="1992-07-12" collectionCode="T" collectorName="Ogata &amp; Ito" pageId="3" pageNumber="47" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D60FD104F2CFFFA6AB11EA6FD1CFCB8" box="[654,756,1159,1185]" pageId="3" pageNumber="47" type="female">1 female</specimenCount>
, Shiratani-unsuikyo,
<date id="FFD810594F2CFFFA6BD71EA6FB82FCB8" box="[1000,1130,1159,1185]" pageId="3" pageNumber="47" value="1992-07-12">
<collectingDate id="EF9CE9B14F2CFFFA6BD71EA6FB82FCB8" box="[1000,1130,1159,1185]" pageId="3" pageNumber="47" value="1992-07-12">12.vii.1992</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F2CFFFA6C471EA6FB62FCB8" box="[1144,1162,1159,1185]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="47">T</collectionCode>
.
<collectorName id="2693534F4F2CFFFA6CA71EA6FB33FCB8" box="[1176,1243,1159,1185]" pageId="3" pageNumber="47">Ogata</collectorName>
(
<collectionCode id="ED77AE5C4F2CFFFA6CD21EA6FAE8FCB8" box="[1261,1280,1159,1185]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="47">T</collectionCode>
.
<collectorName id="2693534F4F2CFFFA6D321EA6FAD8FCB8" box="[1293,1328,1159,1185]" pageId="3" pageNumber="47">Ito</collectorName>
)
</materialsCitation>
;
<specimenCount id="9D60FD104F2CFFFA6D781EA6FA70FCB8" box="[1351,1432,1159,1185]" pageId="3" pageNumber="47" type="male">1 male</specimenCount>
, Kurio, Kuromi-gawa,
<date id="FFD810594F2CFFFA69AD1E8AFE1EFCDC" box="[402,502,1195,1221]" pageId="3" pageNumber="47" value="1996-10-02">2.x.1996</date>
, M. Sato (N. Kuhara);
<specimenCount id="9D60FD104F2CFFFA6AC31E8AFCA3FCDC" box="[764,843,1195,1221]" pageId="3" pageNumber="47" type="male">1 male</specimenCount>
, Yoshida-gawa, Yoshida-bashi,
<quantity id="4C9E9B7C4F2CFFFA6C8E1E8AFB05FCDC" box="[1201,1261,1195,1221]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="47" unit="m" value="40.0">40 m</quantity>
,
<date id="FFD810594F2CFFFA6CC51E8BFA9DFCDC" box="[1274,1397,1194,1221]" pageId="3" pageNumber="47" value="2003-09-25">25.ix.2003</date>
, N. Kuhara (N. Kuhara).
</paragraph>
</subSubSection>
<subSubSection id="C37C65124F2CFFFA68F81ED3FBB5FD4F" pageId="3" pageNumber="47" type="distribution">
<paragraph id="8BD936994F2CFFFA68F81ED3FDADFD17" blockId="3.[151,1437,151,1437]" box="[199,581,1266,1294]" pageId="3" pageNumber="47">
<emphasis id="B912EA8B4F2CFFFA68F81ED3FEB1FD15" bold="true" box="[199,345,1266,1292]" pageId="3" pageNumber="47">Distribution</emphasis>
.
<collectingCountry id="F37176094F2CFFFA69581ED2FE4FFD14" box="[359,423,1267,1293]" name="Japan" pageId="3" pageNumber="47">Japan</collectingCountry>
(Yakushima).
</paragraph>
<paragraph id="8BD936994F2CFFFA68F81F37FBB5FD4F" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="47">
<emphasis id="B912EA8B4F2CFFFA68F81F37FECBFD29" bold="true" box="[199,291,1302,1328]" pageId="3" pageNumber="47">Habitat</emphasis>
. Adults of this species were collected beside fast flowing streams with stony bottoms. They were abundant at
<quantity id="4C9E9B7C4F2CFFFA691F1F1AFE2BFD4C" box="[288,451,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="3" pageNumber="47" unit="m" value="950.0" valueMax="1200.0" valueMin="700.0">700 to 1200 m</quantity>
a.s.l., and rare at sites lower than
<quantity id="4C9E9B7C4F2CFFFA6B7F1F1BFC91FD4C" box="[832,889,1338,1365]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="47" unit="m" value="50.0">50 m</quantity>
elevation (
<figureCitation id="135D2A1C4F2CFFFA6BCA1F1AFBA7FD4F" box="[1013,1103,1339,1366]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[232,1361,183,1919]" captionTargetId="figure-16@12.[225,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Adult collection sites of seven species of Rhyacophila on Yakushima. Solid circle = 14 adults collected; open circle = more than 5 adults collected." figureDoi="http://doi.org/10.5281/zenodo.5225422" httpUri="https://zenodo.org/record/5225422/files/figure.png" pageId="3" pageNumber="47">Fig. 6A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C65124F2CFFFA68F81F7EFD2FFD84" pageId="3" pageNumber="47" type="etymology">
<paragraph id="8BD936994F2CFFFA68F81F7EFB15FD60" blockId="3.[151,1437,151,1437]" box="[199,1277,1375,1401]" pageId="3" pageNumber="47">
<emphasis id="B912EA8B4F2CFFFA68F81F7EFEA0FD60" bold="true" box="[199,328,1375,1401]" pageId="3" pageNumber="47">Etymology</emphasis>
. The name “
<taxonomicName id="4C664D1A4F2CFFFA69E71F7EFDB8FD60" authorityName="Ito" authorityYear="2021" box="[472,592,1375,1401]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="47" phylum="Arthropoda" rank="species" species="yakushima">
<emphasis id="B912EA8B4F2CFFFA69E71F7EFDB8FD60" box="[472,592,1375,1401]" italics="true" pageId="3" pageNumber="47">yakushima</emphasis>
</taxonomicName>
” is a noun in apposition, coined from the name of the island.
</paragraph>
<paragraph id="8BD936994F2CFFFA68F81FA2FD2FFD84" blockId="3.[151,1437,151,1437]" box="[199,711,1411,1437]" pageId="3" pageNumber="47">
<emphasis id="B912EA8B4F2CFFFA68F81FA2FE94FD84" bold="true" box="[199,380,1411,1437]" pageId="3" pageNumber="47">Japanese name</emphasis>
. Yakushima-nagare-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>