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<document id="8E8219EBF202B511125B1CD1725F0BC5" ID-DOI="10.11646/zootaxa.5023.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5225402" ID-ZooBank="F6CB0172-C3B9-4E9F-ACBF-2BC006F588F5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629423656372" checkinUser="plazi" docAuthor="Ito, Tomiko" docDate="2021" docId="03CF878F4F2AFFFE68A81FCFFD2EFE10" docLanguage="en" docName="zootaxa.5023.1.2.pdf" docOrigin="Zootaxa 5023 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophila miyanoura Ito 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="51" masterDocId="FFF6FFF74F2FFFF9683F1A21FFE8F819" masterDocTitle="The genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) in Yakushima Island, northern Ryukyu, southwestern Japan" masterLastPageNumber="58" masterPageNumber="44" pageNumber="49" updateTime="1699072484263" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="43BC384AB78FEEB11F09BDF0F1231024">The genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) in Yakushima Island, northern Ryukyu, southwestern Japan</mods:title>
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<mods:date id="DD94E74E1B8CEC224714C175C9F1E2E8">2021</mods:date>
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<subSubSection id="C37C65124F2AFFFC68A81FCFFDC9FE10" box="[151,545,1518,1545]" pageId="5" pageNumber="49" type="nomenclature">
<paragraph id="8BD936994F2AFFFC68A81FCFFDC9FE10" blockId="5.[151,545,1518,1582]" box="[151,545,1518,1545]" pageId="5" pageNumber="49">
<heading id="D09181F54F2AFFFC68A81FCFFDC9FE10" bold="true" box="[151,545,1518,1545]" fontSize="11" level="1" pageId="5" pageNumber="49" reason="1">
<emphasis id="B912EA8B4F2AFFFC68A81FCFFDC9FE10" bold="true" box="[151,545,1518,1545]" pageId="5" pageNumber="49">
<taxonomicName id="4C664D1A4F2AFFFC68A81FCFFE54FE10" authority="Ito, 2021" authorityName="Ito" authorityYear="2021" box="[151,444,1518,1545]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="miyanoura" status="sp. nov.">
<emphasis id="B912EA8B4F2AFFFC68A81FCFFE54FE10" bold="true" box="[151,444,1518,1545]" italics="true" pageId="5" pageNumber="49">Rhyacophila miyanoura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157F04F2AFFFC69FC1FCEFDC9FE10" box="[451,545,1519,1545]" pageId="5" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37C65124F2AFFFC68A81C32FEF5FE37" box="[151,285,1555,1582]" pageId="5" pageNumber="49" type="description">
<paragraph id="8BD936994F2AFFFC68A81C32FEF5FE37" blockId="5.[151,545,1518,1582]" box="[151,285,1555,1582]" pageId="5" pageNumber="49">
(
<figureCitation id="135D2A1C4F2AFFFC68A01C32FF00FE34" box="[159,232,1555,1581]" captionStart="FIGURE 4" captionStartId="6.[151,250,1775,1799]" captionTargetBox="[174,1436,194,1745]" captionTargetId="figure-16@6.[151,1436,179,1750]" captionTargetPageId="6" captionText="FIGURE 4. Rhyacophila miyanoura sp. nov. 4A4F, male: 4A, genitalia, left lateral; 4B, same, caudal; 4C, apicodorsal lobe of segment IX and phallic apparatus, left lateral; 4D, apicodorsal lobe of segment IX and preanal appendages, ventral; 4E, apicodorsal lobe of segment IX, preanal appendages, and right inferior appendage, dorsal; 4F, segment IX and left inferior appendage, ventral. 4G4J, female: 4G, abdominal segments VIIIXI, left lateral; 4H, segment VIII, dorsal; 4I, same, ventral; 4J, vaginal apparatus, ventral.Abbreviations: VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ad l = apicodorsal lobe; ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band; b inf app = basal segment of inferior appendage (paired); do pr = dorsal process; end = endotheca; ph = phallotheca; pre app = preanal appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.5225414" httpUri="https://zenodo.org/record/5225414/files/figure.png" pageId="5" pageNumber="49">Figs 4</figureCitation>
,
<figureCitation id="135D2A1C4F2AFFFC68CB1C32FEFEFE37" box="[244,278,1555,1582]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[232,1361,183,1919]" captionTargetId="figure-16@12.[225,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Adult collection sites of seven species of Rhyacophila on Yakushima. Solid circle = 14 adults collected; open circle = more than 5 adults collected." figureDoi="http://doi.org/10.5281/zenodo.5225422" httpUri="https://zenodo.org/record/5225422/files/figure.png" pageId="5" pageNumber="49">6C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C65124F2AFFFE68A81C7BFCD8FC08" lastPageId="7" lastPageNumber="51" pageId="5" pageNumber="49" type="diagnosis">
<paragraph id="8BD936994F2AFFFC68A81C7BFB2FFFC4" blockId="5.[151,1436,1626,2013]" pageId="5" pageNumber="49">
<emphasis id="B912EA8B4F2AFFFC68A81C7BFEE2FE6D" bold="true" box="[151,266,1626,1652]" pageId="5" pageNumber="49">Diagnosis</emphasis>
. This new species belongs to the
<taxonomicName id="4C664D1A4F2AFFFC6AB01C7AFCE7FE6D" authorityName="Tsuda" authorityYear="1940" box="[655,783,1627,1652]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="yosiiana">
<emphasis id="B912EA8B4F2AFFFC6AB01C7AFCE7FE6D" box="[655,783,1627,1652]" italics="true" pageId="5" pageNumber="49">R. yosiiana</emphasis>
</taxonomicName>
Species Group (see Discussion), and the male resembles those of
<taxonomicName id="4C664D1A4F2AFFFC68C81C5EFDECFE80" authority="Kuranishi 1997" authorityName="Kuranishi" authorityYear="1997" box="[247,516,1663,1689]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="B912EA8B4F2AFFFC68C81C5EFEA4FE81" box="[247,332,1663,1688]" italics="true" pageId="5" pageNumber="49">R. satoi</emphasis>
<bibRefCitation id="EFF74B684F2AFFFC696B1C5EFDECFE80" author="Kuranishi, R. B." box="[340,516,1663,1689]" pageId="5" pageNumber="49" pagination="265 - 269" refId="ref9139" refString="Kuranishi, R. B. (1997) The genus Rhyacophila of the Ryukyu Archipelago, part I (Trichoptera: Rhyacophilidae). In: Holzenthal, R. W. &amp; Flint, O. S. Jr. (Eds.), Proceedings of the 8 th International Symposium on Trichoptera. Ohio Biological Survey, Columbus, Ohio, pp. 265 - 269." type="book chapter" year="1997">Kuranishi 1997</bibRefCitation>
</taxonomicName>
, inhabiting central Ryukyu, southern
<collectingCountry id="F37176094F2AFFFC6B981C5EFC02FE80" box="[935,1002,1663,1689]" name="Japan" pageId="5" pageNumber="49">Japan</collectingCountry>
, and
<taxonomicName id="4C664D1A4F2AFFFC6C1B1C5EFAC5FE80" authority="Tsuda 1940" authorityName="Tsuda" authorityYear="1940" box="[1060,1325,1663,1689]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="yosiiana">
<emphasis id="B912EA8B4F2AFFFC6C1B1C5EFBDDFE81" box="[1060,1077,1663,1688]" italics="true" pageId="5" pageNumber="49">R</emphasis>
.
<emphasis id="B912EA8B4F2AFFFC6C7C1C5EFB4AFE81" box="[1091,1186,1663,1688]" italics="true" pageId="5" pageNumber="49">yosiiana</emphasis>
<bibRefCitation id="EFF74B684F2AFFFC6C961C5EFAC5FE80" author="Tsuda, M." box="[1193,1325,1663,1689]" pageId="5" pageNumber="49" pagination="119 - 135" refId="ref9796" refString="Tsuda, M. (1940) Zur Kenntnis der Japanischen Rhyacophilinen (Rhyacophilidae, Trichoptera). Annotationes Zoologicae Japonenses, 19, 119 - 135. [in German]" type="journal article" year="1940">Tsuda 1940</bibRefCitation>
</taxonomicName>
, found on Honshu, central
<collectingCountry id="F37176094F2AFFFC69701C82FE67FEA4" box="[335,399,1699,1725]" name="Japan" pageId="5" pageNumber="49">Japan</collectingCountry>
(
<bibRefCitation id="EFF74B684F2AFFFC69A21C82FD67FEA4" author="Taira, A. &amp; Nozaki, T." box="[413,655,1699,1725]" pageId="5" pageNumber="49" refId="ref9647" refString="Taira, A. &amp; Nozaki, T. (2021) A catalogue of Japanese Trichoptera family Rhyacophilidae Stephens. Available from: http: // tobikera. eco. coocan. jp / catalog / Rhyacophilidae. html (accessed 14 February 2021)" type="url" year="2021">Taira &amp; Nozaki 2021</bibRefCitation>
), in having a scoop-shaped apicodorsal lobe of segment IX and a very small phallic apparatus without parameres (
<bibRefCitation id="EFF74B684F2AFFFC6AB41CE6FCA8FEF8" author="Kuranishi, R. B." box="[651,832,1735,1761]" pageId="5" pageNumber="49" pagination="265 - 269" refId="ref9139" refString="Kuranishi, R. B. (1997) The genus Rhyacophila of the Ryukyu Archipelago, part I (Trichoptera: Rhyacophilidae). In: Holzenthal, R. W. &amp; Flint, O. S. Jr. (Eds.), Proceedings of the 8 th International Symposium on Trichoptera. Ohio Biological Survey, Columbus, Ohio, pp. 265 - 269." type="book chapter" year="1997">Kuranishi 1997</bibRefCitation>
;
<bibRefCitation id="EFF74B684F2AFFFC6B701CE6FC0DFEF8" author="Hattori, T." box="[847,997,1734,1761]" pageId="5" pageNumber="49" pagination="414 - 437" refId="ref8732" refString="Hattori, T. (2005) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations. Tokai University Press, Kanagawa, pp. 414 - 437. [in Japanese]" type="book chapter" year="2005">Hattori 2005</bibRefCitation>
). The male of
<taxonomicName id="4C664D1A4F2AFFFC6CB01CE6FAC2FEF9" authorityName="Ito" authorityYear="2021" box="[1167,1322,1735,1760]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="miyanoura">
<emphasis id="B912EA8B4F2AFFFC6CB01CE6FAC2FEF9" box="[1167,1322,1735,1760]" italics="true" pageId="5" pageNumber="49">R. miyanoura</emphasis>
</taxonomicName>
is clearly discriminated from those of
<taxonomicName id="4C664D1A4F2AFFFC69DE1CCAFDD2FF1D" authorityName="Kuranishi" authorityYear="1997" box="[481,570,1771,1796]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="B912EA8B4F2AFFFC69DE1CCAFDD2FF1D" box="[481,570,1771,1796]" italics="true" pageId="5" pageNumber="49">R. satoi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C664D1A4F2AFFFC6A481CCAFD10FF1D" authorityName="Tsuda" authorityYear="1940" box="[631,760,1771,1796]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="yosiiana">
<emphasis id="B912EA8B4F2AFFFC6A481CCAFD10FF1D" box="[631,760,1771,1796]" italics="true" pageId="5" pageNumber="49">R. yosiiana</emphasis>
</taxonomicName>
as follows: The venter of segment IX has a large bilobed projection in the middle, the ventroposterior margin of the apicodorsal lobe of segment IX is irregularly serrated, and the apical segment of each inferior appendage is deeply bifurcate in
<taxonomicName id="4C664D1A4F2AFFFC6BF31D12FB8DFF55" authorityName="Ito" authorityYear="2021" box="[972,1125,1843,1868]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="miyanoura">
<emphasis id="B912EA8B4F2AFFFC6BF31D12FB8DFF55" box="[972,1125,1843,1868]" italics="true" pageId="5" pageNumber="49">R. miyanoura</emphasis>
</taxonomicName>
; whereas in
<taxonomicName id="4C664D1A4F2AFFFC6CC91D12FAA4FF55" authorityName="Kuranishi" authorityYear="1997" box="[1270,1356,1843,1868]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="B912EA8B4F2AFFFC6CC91D12FAA4FF55" box="[1270,1356,1843,1868]" italics="true" pageId="5" pageNumber="49">R. satoi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C664D1A4F2AFFFC6DBB1D12FF1EFF69" authorityName="Tsuda" authorityYear="1940" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="yosiiana">
<emphasis id="B912EA8B4F2AFFFC6DBB1D12FF1EFF69" italics="true" pageId="5" pageNumber="49">R. yosiiana</emphasis>
</taxonomicName>
the venter of segment IX is short, the ventroposterior margin of the apicodorsal lobe of segment IX (segment X of
<bibRefCitation id="EFF74B684F2AFFFC68E81D5AFE6CFF8C" author="Kuranishi, R. B." box="[215,388,1915,1941]" pageId="5" pageNumber="49" pagination="265 - 269" refId="ref9139" refString="Kuranishi, R. B. (1997) The genus Rhyacophila of the Ryukyu Archipelago, part I (Trichoptera: Rhyacophilidae). In: Holzenthal, R. W. &amp; Flint, O. S. Jr. (Eds.), Proceedings of the 8 th International Symposium on Trichoptera. Ohio Biological Survey, Columbus, Ohio, pp. 265 - 269." type="book chapter" year="1997">Kuranishi 1997</bibRefCitation>
) is round, and the apical segment of each inferior appendage is not bifurcate. The female of
<taxonomicName id="4C664D1A4F2AFFFC6DBB1D5AFEF8FFA1" authorityName="Ito" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="miyanoura">
<emphasis id="B912EA8B4F2AFFFC6DBB1D5AFEF8FFA1" italics="true" pageId="5" pageNumber="49">R. miyanoura</emphasis>
</taxonomicName>
also resembles that of
<taxonomicName id="4C664D1A4F2AFFFC6A2E1DBEFD8EFFA1" authorityName="Kuranishi" authorityYear="1997" box="[529,614,1951,1976]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="B912EA8B4F2AFFFC6A2E1DBEFD8EFFA1" box="[529,614,1951,1976]" italics="true" pageId="5" pageNumber="49">R. satoi</emphasis>
</taxonomicName>
in the shape of the short segment VIII and the vaginal apparatus having a bilobed posterior margin (
<bibRefCitation id="EFF74B684F2AFFFC69851DE2FD85FFC4" author="Kuranishi, R. B." box="[442,621,1987,2013]" pageId="5" pageNumber="49" pagination="265 - 269" refId="ref9139" refString="Kuranishi, R. B. (1997) The genus Rhyacophila of the Ryukyu Archipelago, part I (Trichoptera: Rhyacophilidae). In: Holzenthal, R. W. &amp; Flint, O. S. Jr. (Eds.), Proceedings of the 8 th International Symposium on Trichoptera. Ohio Biological Survey, Columbus, Ohio, pp. 265 - 269." type="book chapter" year="1997">Kuranishi 1997</bibRefCitation>
); they cannot be distinguished from each other easily.
</paragraph>
<caption id="DF1966114F29FFFF68A81CCEFB28FFF9" ID-DOI="http://doi.org/10.5281/zenodo.5225414" ID-Zenodo-Dep="5225414" httpUri="https://zenodo.org/record/5225414/files/figure.png" pageId="6" pageNumber="50" startId="6.[151,250,1775,1799]" targetBox="[174,1436,194,1745]" targetPageId="6">
<paragraph id="8BD936994F29FFFF68A81CCEFB28FFF9" blockId="6.[151,1437,1775,2016]" pageId="6" pageNumber="50">
<emphasis id="B912EA8B4F29FFFF68A81CCEFEE5FF11" bold="true" box="[151,269,1775,1800]" pageId="6" pageNumber="50">FIGURE 4</emphasis>
.
<taxonomicName id="4C664D1A4F29FFFF69251CD1FDE4FF1E" authority="Ito, 2021" authorityName="Ito" authorityYear="2021" box="[282,524,1776,1800]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="50" phylum="Arthropoda" rank="species" species="miyanoura" status="sp. nov.">
<emphasis id="B912EA8B4F29FFFF69251CD1FDE4FF1E" box="[282,524,1776,1800]" italics="true" pageId="6" pageNumber="50">Rhyacophila miyanoura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157F04F29FFFF6A2D1CD1FD8BFF1E" box="[530,611,1776,1800]" pageId="6" pageNumber="50" rank="species">
<emphasis id="B912EA8B4F29FFFF6A2D1CD1FDB5FF11" bold="true" box="[530,605,1776,1800]" pageId="6" pageNumber="50">sp. nov</emphasis>
.
</taxonomicNameLabel>
4A4F, male: 4A, genitalia, left lateral; 4B, same, caudal; 4C, apicodorsal lobe of segment IX and phallic apparatus, left lateral; 4D, apicodorsal lobe of segment IX and preanal appendages, ventral; 4E, apicodorsal lobe of segment IX, preanal appendages, and right inferior appendage, dorsal; 4F, segment IX and left inferior appendage, ventral. 4G4J, female: 4G, abdominal segments VIIIXI, left lateral; 4H, segment VIII, dorsal; 4I, same, ventral; 4J, vaginal apparatus, ventral. Abbreviations: VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ad l = apicodorsal lobe; ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band; b inf app = basal segment of inferior appendage (paired); do pr = dorsal process; end = endotheca; ph = phallotheca; pre app = preanal appendage (paired).
</paragraph>
</caption>
<paragraph id="8BD936994F28FFFE68F81AB7FBD0F9B4" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="51">
<emphasis id="B912EA8B4F28FFFE68F81AB7FEE3F8A9" bold="true" box="[199,267,150,176]" pageId="7" pageNumber="51">Adult</emphasis>
. Lengths of forewings and hind wings: Each forewing 6.210.0 mm long (mean
<quantity id="4C9E9B7C4F28FFFE6C901AB6FAE1F8A8" box="[1199,1289,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="7" pageNumber="51" unit="mm" value="7.2">7.2 mm</quantity>
, n = 18) and each hind wing
<quantity id="4C9E9B7C4F28FFFE69731A9BFE3BF8CC" box="[332,467,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="8.9" metricValueMin="5.6" pageId="7" pageNumber="51" unit="mm" value="7.25" valueMax="8.9" valueMin="5.6">5.68.9 mm</quantity>
(mean
<quantity id="4C9E9B7C4F28FFFE6A161A9BFD69F8CC" box="[553,641,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="7" pageNumber="51" unit="mm" value="6.5">6.5 mm</quantity>
, n = 18) in males, 7.09.0 mm (mean
<quantity id="4C9E9B7C4F28FFFE6C0C1A9AFB64F8CC" box="[1075,1164,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="51" unit="mm" value="7.6">7.6 mm</quantity>
, n = 9) and 6.68.0 mm (mean 7.0 mm, n = 9) in females, respectively. Head dark brown, warts light brown with brown setae, antennae brown, scapes thicker and longer than other segments; palpi brown. Thorax dark brown dorsally. Legs brown with brown spurs. Wings brown with darker veins and fulvous pterostigma, few whitish dots on forewings of both sexes. Abdomen with brown tergites and light brown sternites, dark pigments scattered dorsally; scent glands of sternite V opening on antero-angle mounds; small mid ventral process on sternite VII (or VI and VII) in male and on sternite VI in female, process on sternite VI of male often very small or completely absent.
</paragraph>
<paragraph id="8BD936994F28FFFE68F81B96FED8FAB0" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="51">
<emphasis id="B912EA8B4F28FFFE68F81B96FE9AF9C9" bold="true" box="[199,370,438,465]" pageId="7" pageNumber="51">Male genitalia</emphasis>
(
<figureCitation id="135D2A1C4F28FFFE69BE1B96FDE5F9C8" box="[385,525,439,465]" captionStart="FIGURE 4" captionStartId="6.[151,250,1775,1799]" captionTargetBox="[174,1436,194,1745]" captionTargetId="figure-16@6.[151,1436,179,1750]" captionTargetPageId="6" captionText="FIGURE 4. Rhyacophila miyanoura sp. nov. 4A4F, male: 4A, genitalia, left lateral; 4B, same, caudal; 4C, apicodorsal lobe of segment IX and phallic apparatus, left lateral; 4D, apicodorsal lobe of segment IX and preanal appendages, ventral; 4E, apicodorsal lobe of segment IX, preanal appendages, and right inferior appendage, dorsal; 4F, segment IX and left inferior appendage, ventral. 4G4J, female: 4G, abdominal segments VIIIXI, left lateral; 4H, segment VIII, dorsal; 4I, same, ventral; 4J, vaginal apparatus, ventral.Abbreviations: VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ad l = apicodorsal lobe; ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band; b inf app = basal segment of inferior appendage (paired); do pr = dorsal process; end = endotheca; ph = phallotheca; pre app = preanal appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.5225414" httpUri="https://zenodo.org/record/5225414/files/figure.png" pageId="7" pageNumber="51">Figs 4A4F</figureCitation>
). Segment IX (IX) in lateral view slightly convex dorsally with large apicodorsal lobe (ad l), venter with pair of large posteroventral projections; apicodorsal lobe scoop-shaped, in ventral view with apicomesal incision posteriorly, convex lateral margins, irregularly serrate posteroventrally. Preanal appendages (pre app) fused with basal half of apicodorsal lobe of segment IX, broadened laterally in dorsal and ventral views. Segment X (X) near basoventral edges of preanal appendages, semi-membranous, invisible in lateral view, short and wide in caudal view. Anal sclerites (an sc) small, round, weakly sclerotized; apical band (ap ba) short, semimembranous.
</paragraph>
<paragraph id="8BD936994F28FFFE68F81892FD23FB0C" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="51">Phallic apparatus very small, not exceeding basal 1/4 of basal segment of inferior appendage (b inf app). Phallotheca (ph) membranous with semi-membranous dorsal process (do pr); retracted aedeagus (ae) slender, surrounded by endotheca (end); parameres absent.</paragraph>
<paragraph id="8BD936994F28FFFE68F8193EFDBFFB98" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="51">Inferior appendages large, basal segment (b inf app) almost parallel sided with oblique caudal margins in lateral and dorsal views, apical segment (a inf app) bifurcate at apical 1/3, dorsal branch shorter than ventral one, both branches round and spiny apicomesally.</paragraph>
<paragraph id="8BD936994F28FFFE68F819AAFCD8FC08" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="51">
<emphasis id="B912EA8B4F28FFFE68F819AAFE66FBBD" bold="true" box="[199,398,906,933]" pageId="7" pageNumber="51">Female genitalia</emphasis>
(
<figureCitation id="135D2A1C4F28FFFE69A119AAFDCDFBBC" box="[414,549,907,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1775,1799]" captionTargetBox="[174,1436,194,1745]" captionTargetId="figure-16@6.[151,1436,179,1750]" captionTargetPageId="6" captionText="FIGURE 4. Rhyacophila miyanoura sp. nov. 4A4F, male: 4A, genitalia, left lateral; 4B, same, caudal; 4C, apicodorsal lobe of segment IX and phallic apparatus, left lateral; 4D, apicodorsal lobe of segment IX and preanal appendages, ventral; 4E, apicodorsal lobe of segment IX, preanal appendages, and right inferior appendage, dorsal; 4F, segment IX and left inferior appendage, ventral. 4G4J, female: 4G, abdominal segments VIIIXI, left lateral; 4H, segment VIII, dorsal; 4I, same, ventral; 4J, vaginal apparatus, ventral.Abbreviations: VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ad l = apicodorsal lobe; ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band; b inf app = basal segment of inferior appendage (paired); do pr = dorsal process; end = endotheca; ph = phallotheca; pre app = preanal appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.5225414" httpUri="https://zenodo.org/record/5225414/files/figure.png" pageId="7" pageNumber="51">Figs 4G4J</figureCitation>
). Segment VIII (VIII) short, length 0.8 times as long as basal width, posterior margins slightly convex in lateral view, subquadrate with anterior and posterior margins slightly incised in dorsal view, in ventral view roundish with large posterior incision. Vaginal apparatus slightly sclerotized, slender anteriorly, gradually thickened posteriorly with bilobed apical margin.
</paragraph>
</subSubSection>
<subSubSection id="C37C65124F28FFFE68F81E3AFC43FD4C" pageId="7" pageNumber="51" type="materials_examined">
<paragraph id="8BD936994F28FFFE68F81E3AFD6BFC40" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="51">
<materialsCitation id="3B0E3CC44F28FFFE68F81E3AFD6BFC40" collectingDate="2006-05-10" collectionCode="TI" country="Japan" county="Miyanoura-gawa" elevation="60" location="Male" municipality="Yakushima Island" pageId="7" pageNumber="51" specimenCode="CBM-ZI 0159998" specimenCount="1" typeStatus="holotype">
<emphasis id="B912EA8B4F28FFFE68F81E3AFEDCFC2C" bold="true" box="[199,308,1051,1077]" pageId="7" pageNumber="51">
<typeStatus id="54DD883B4F28FFFE68F81E3AFEDCFC2C" box="[199,308,1051,1077]" pageId="7" pageNumber="51" type="holotype">Holotype</typeStatus>
</emphasis>
.
<location id="8EB960424F28FFFE697B1E3AFE68FC2C" LSID="urn:lsid:plazi:treatment:03CF878F4F2AFFFE68A81FCFFD2EFE10:8EB960424F28FFFE697B1E3AFE68FC2C" box="[324,384,1051,1077]" country="Japan" county="Miyanoura-gawa" municipality="Yakushima Island" name="Male" pageId="7" pageNumber="51">Male</location>
,
<emphasis id="B912EA8B4F28FFFE69B21E3AFE3EFC2C" bold="true" box="[397,470,1051,1077]" pageId="7" pageNumber="51">
<collectingCountry id="F37176094F28FFFE69B21E3AFE3EFC2C" box="[397,470,1051,1077]" name="Japan" pageId="7" pageNumber="51">Japan</collectingCountry>
</emphasis>
,
<emphasis id="B912EA8B4F28FFFE69D91E3BFD57FC2C" bold="true" box="[486,703,1050,1077]" pageId="7" pageNumber="51">
<collectingMunicipality id="6BBDACE34F28FFFE69D91E3BFD57FC2C" box="[486,703,1050,1077]" pageId="7" pageNumber="51">Yakushima Island</collectingMunicipality>
</emphasis>
,
<collectingCounty id="62B84E154F28FFFE6AF11E3AFC78FC2C" box="[718,912,1051,1077]" pageId="7" pageNumber="51">Miyanoura-gawa</collectingCounty>
, small tributary (30.41˚N, 130.53˚E,
<elevation id="004BD1AA4F28FFFE6D071E3AFF24FC43" metricMagnitude="1" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="7" pageNumber="51" unit="m" value="60.0" valueMax="80.0" valueMin="40.0">
<quantity id="4C9E9B7C4F28FFFE6D071E3AFA74FC2C" box="[1336,1436,1051,1077]" metricMagnitude="1" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="7" pageNumber="51" unit="m" value="60.0" valueMax="80.0" valueMin="40.0">4080 m</quantity>
a.s.l.
</elevation>
),
<date id="FFD810594F28FFFE68DC1E1EFEBCFC40" box="[227,340,1087,1113]" pageId="7" pageNumber="51" value="2006-05-10">
<collectingDate id="EF9CE9B14F28FFFE68DC1E1EFEBCFC40" box="[227,340,1087,1113]" pageId="7" pageNumber="51" value="2006-05-10">10.v.2006</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F28FFFE69611E1EFE93FC40" box="[350,379,1087,1113]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="7" pageNumber="51">TI</collectionCode>
, S (
<specimenCode id="DBC09EE24F28FFFE699A1E1EFD9FFC40" box="[421,631,1086,1113]" collectionCode="CBM-ZI" pageId="7" pageNumber="51">CBM-ZI 0159998</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD936994F28FFFE68F81E42FE1EFD14" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="51">
<materialsCitation id="3B0E3CC44F28FFFE68F81E42FD50FC64" box="[199,696,1122,1149]" collectionCode="SPMN-IS" location="Yakushima Island" pageId="7" pageNumber="51" specimenCount="1" typeStatus="paratype">
<emphasis id="B912EA8B4F28FFFE68F81E42FED7FC64" bold="true" box="[199,319,1123,1149]" pageId="7" pageNumber="51">
<typeStatus id="54DD883B4F28FFFE68F81E42FED7FC64" box="[199,319,1123,1149]" pageId="7" pageNumber="51" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<collectionCode id="ED77AE5C4F28FFFE696D1E42FE2FFC64" box="[338,455,1123,1149]" pageId="7" pageNumber="51">SPMN-IS</collectionCode>
).
<emphasis id="B912EA8B4F28FFFE69E21E43FD50FC64" bold="true" box="[477,696,1122,1149]" pageId="7" pageNumber="51">
<location id="8EB960424F28FFFE69E21E43FD50FC64" LSID="urn:lsid:plazi:treatment:03CF878F4F2AFFFE68A81FCFFD2EFE10:8EB960424F28FFFE69E21E43FD50FC64" box="[477,696,1122,1149]" name="Yakushima Island" pageId="7" pageNumber="51">Yakushima Island</location>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B0E3CC44F28FFFE6AF41E43FB56FC64" box="[715,1214,1122,1149]" collectingDate="2006-05-10" collectionCode="TI" country="Japan" county="Miyanoura-gawa" elevation="60" location="Male" municipality="Yakushima Island" pageId="7" pageNumber="51" specimenCount="7" specimenCount-female="2" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="9D60FD104F28FFFE6AF41E43FCCFFC64" box="[715,807,1122,1149]" pageId="7" pageNumber="51" type="male">5 males</specimenCount>
,
<specimenCount id="9D60FD104F28FFFE6B081E42FC41FC64" box="[823,937,1123,1149]" pageId="7" pageNumber="51" type="female">2 females</specimenCount>
, same data as holotype
</materialsCitation>
;
<specimenCount id="9D60FD104F28FFFE6CF21E42FAF7FC64" box="[1229,1311,1123,1149]" pageId="7" pageNumber="51" type="male">1 male</specimenCount>
, Shirataniunsuikyo,
<date id="FFD810594F28FFFE69321EA6FE6DFCB8" box="[269,389,1159,1185]" pageId="7" pageNumber="51" value="2003-04-24">24.iv.2003</date>
, A. Ishizuka;
<materialsCitation id="3B0E3CC44F28FFFE6A1D1EA7FBDBFCB8" box="[546,1075,1158,1185]" collectingDate="2006-05-09" collectionCode="TI" country="Japan" county="Miyanoura-gawa" elevation="60" location="Male" municipality="Yakushima Island" pageId="7" pageNumber="51" specimenCount="7" specimenCount-female="2" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="9D60FD104F28FFFE6A1D1EA7FD94FCB8" box="[546,636,1158,1185]" pageId="7" pageNumber="51" type="male">5 males</specimenCount>
,
<specimenCount id="9D60FD104F28FFFE6AB61EA6FD11FCB8" box="[649,761,1159,1185]" pageId="7" pageNumber="51" type="female">2 females</specimenCount>
, same locality,
<date id="FFD810594F28FFFE6B981EA6FBE2FCB8" box="[935,1034,1159,1185]" pageId="7" pageNumber="51" value="2006-05-09">
<collectingDate id="EF9CE9B14F28FFFE6B981EA6FBE2FCB8" box="[935,1034,1159,1185]" pageId="7" pageNumber="51" value="2006-05-09">9.v.2006</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F28FFFE6C291EA6FBDBFCB8" box="[1046,1075,1159,1185]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="7" pageNumber="51">TI</collectionCode>
</materialsCitation>
, S;
<materialsCitation id="3B0E3CC44F28FFFE6C601EA6FD4CFCDC" collectingDate="2007-06-12" collectionCode="TI" country="Japan" pageId="7" pageNumber="51" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D60FD104F28FFFE6C601EA6FB46FCB8" box="[1119,1198,1159,1185]" pageId="7" pageNumber="51" type="male">1 male</specimenCount>
,
<specimenCount id="9D60FD104F28FFFE6C851EA6FAF7FCB8" box="[1210,1311,1159,1185]" pageId="7" pageNumber="51" type="female">1 female</specimenCount>
, Yakusugiland, Ara-kawa, Seiryu-bashi,
<date id="FFD810594F28FFFE69C31E8AFD9EFCDC" box="[508,630,1195,1221]" pageId="7" pageNumber="51" value="2007-06-12">
<collectingDate id="EF9CE9B14F28FFFE69C31E8AFD9EFCDC" box="[508,630,1195,1221]" pageId="7" pageNumber="51" value="2007-06-12">12.vi.2007</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F28FFFE6AB81E8AFD4CFCDC" box="[647,676,1195,1221]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="7" pageNumber="51">TI</collectionCode>
</materialsCitation>
, S;
<materialsCitation id="3B0E3CC44F28FFFE6AE61E8AFED1FCF0" collectingDate="2007-06-12" collectionCode="TI" country="Japan" pageId="7" pageNumber="51" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D60FD104F28FFFE6AE61E8AFCDFFCDC" box="[729,823,1195,1221]" pageId="7" pageNumber="51" type="male">2 males</specimenCount>
,
<specimenCount id="9D60FD104F28FFFE6B771E8AFC54FCDC" box="[840,956,1195,1221]" pageId="7" pageNumber="51" type="female">3 females</specimenCount>
, Yakusugi-land, Ara-kawa, small stream,
<date id="FFD810594F28FFFE68A81EEEFEF9FCF0" box="[151,273,1231,1257]" pageId="7" pageNumber="51" value="2007-06-12">
<collectingDate id="EF9CE9B14F28FFFE68A81EEEFEF9FCF0" box="[151,273,1231,1257]" pageId="7" pageNumber="51" value="2007-06-12">12.vi.2007</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F28FFFE69231EEEFED1FCF0" box="[284,313,1231,1257]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="7" pageNumber="51">TI</collectionCode>
</materialsCitation>
, S, L;
<materialsCitation id="3B0E3CC44F28FFFE69401EEEFBAFFCF0" box="[383,1095,1231,1257]" collectingDate="2006-05-08" collectionCode="TI" country="Japan" pageId="7" pageNumber="51" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D60FD104F28FFFE69401EEEFE0AFCF0" box="[383,482,1231,1257]" pageId="7" pageNumber="51" type="female">1 female</specimenCount>
, Miyanoura, Kue-gawa, Biwatsubo-bashi,
<date id="FFD810594F28FFFE6B821EEEFBC8FCF0" box="[957,1056,1231,1257]" pageId="7" pageNumber="51" value="2006-05-08">
<collectingDate id="EF9CE9B14F28FFFE6B821EEEFBC8FCF0" box="[957,1056,1231,1257]" pageId="7" pageNumber="51" value="2006-05-08">8.v.2006</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F28FFFE6C151EEEFBAFFCF0" box="[1066,1095,1231,1257]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="7" pageNumber="51">TI</collectionCode>
</materialsCitation>
, L;
<specimenCount id="9D60FD104F28FFFE6C4E1EEEFB56FCF0" box="[1137,1214,1231,1257]" pageId="7" pageNumber="51" type="male">1 male</specimenCount>
, Onoaida, Onoaidagyoko,
<date id="FFD810594F28FFFE68D41ED2FE8BFD14" box="[235,355,1267,1293]" pageId="7" pageNumber="51" value="2003-04-17">17.iv.2003</date>
, A. Ishizuka.
</paragraph>
<paragraph id="8BD936994F28FFFE68F81F37FC43FD4C" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="51">
<emphasis id="B912EA8B4F28FFFE68F81F37FE67FD29" bold="true" box="[199,399,1302,1328]" pageId="7" pageNumber="51">Other specimens</emphasis>
.
<emphasis id="B912EA8B4F28FFFE69A21F37FD9DFD28" bold="true" box="[413,629,1302,1329]" pageId="7" pageNumber="51">Yakushima Island</emphasis>
:
<materialsCitation id="3B0E3CC44F28FFFE6ABB1F36FF2AFD4C" collectingDate="2006-09-25" collectingDateMax="2006-10-23" collectingDateMin="2006-09-25" collectionCode="T" elevation="1200" pageId="7" pageNumber="51" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D60FD104F28FFFE6ABB1F36FD3BFD28" box="[644,723,1303,1329]" pageId="7" pageNumber="51" type="male">1 male</specimenCount>
, Ara-kawa,
<quantity id="4C9E9B7C4F28FFFE6B651F36FC59FD28" box="[858,945,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="51" unit="m" value="1200.0">
<elevation id="004BD1AA4F28FFFE6B651F36FC59FD28" box="[858,945,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="51" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="FFD810594F28FFFE6B821F37FB9EFD28" box="[957,1142,1302,1329]" pageId="7" pageNumber="51" value="2006-09-25" valueMax="2006-10-23" valueMin="2006-09-25">
<collectingDate id="EF9CE9B14F28FFFE6B821F37FB9EFD28" box="[957,1142,1302,1329]" pageId="7" pageNumber="51" value="2006-09-25" valueMax="2006-10-23" valueMin="2006-09-25">25.ix23.x.2006</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F28FFFE6CBD1F36FB7CFD28" box="[1154,1172,1303,1329]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="51">T</collectionCode>
. Yamauchi
<emphasis id="B912EA8B4F28FFFE6D291F36FAAFFD28" box="[1302,1351,1303,1329]" italics="true" pageId="7" pageNumber="51">et al</emphasis>
., M (
<collectionCode id="ED77AE5C4F28FFFE6DBA1F36FA70FD28" box="[1413,1432,1303,1329]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="51">T</collectionCode>
. Ito)
</materialsCitation>
;
<materialsCitation id="3B0E3CC44F28FFFE68F11F1AFC4FFD4C" box="[206,935,1339,1365]" collectingDate="2007-06-04" collectingDateMax="2007-06-07" collectingDateMin="2007-06-04" collectionCode="T" elevation="1200" pageId="7" pageNumber="51" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D60FD104F28FFFE68F11F1AFEF4FD4C" box="[206,284,1339,1365]" pageId="7" pageNumber="51" type="male">1 male</specimenCount>
, same locality,
<date id="FFD810594F28FFFE69F71F1AFDB9FD4C" box="[456,593,1339,1365]" pageId="7" pageNumber="51" value="2007-06-04" valueMax="2007-06-07" valueMin="2007-06-04">
<collectingDate id="EF9CE9B14F28FFFE69F71F1AFDB9FD4C" box="[456,593,1339,1365]" pageId="7" pageNumber="51" value="2007-06-04" valueMax="2007-06-07" valueMin="2007-06-04">47.vi.2007</collectingDate>
</date>
,
<collectionCode id="ED77AE5C4F28FFFE6A631F1AFD86FD4C" box="[604,622,1339,1365]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="51">T</collectionCode>
. Yamauchi
<emphasis id="B912EA8B4F28FFFE6AD01F1AFCC8FD4C" box="[751,800,1339,1365]" italics="true" pageId="7" pageNumber="51">et al</emphasis>
., M (
<collectionCode id="ED77AE5C4F28FFFE6B621F1AFC98FD4C" box="[861,880,1339,1365]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="51">T</collectionCode>
. Ito)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37C65124F28FFFE68F81F7FFE33FDDB" pageId="7" pageNumber="51" type="distribution">
<paragraph id="8BD936994F28FFFE68F81F7FFDADFD63" blockId="7.[151,1437,150,1545]" box="[199,581,1374,1402]" pageId="7" pageNumber="51">
<emphasis id="B912EA8B4F28FFFE68F81F7FFEB1FD61" bold="true" box="[199,345,1374,1400]" pageId="7" pageNumber="51">Distribution</emphasis>
.
<collectingCountry id="F37176094F28FFFE69581F7EFE4FFD60" box="[359,423,1375,1401]" name="Japan" pageId="7" pageNumber="51">Japan</collectingCountry>
(Yakushima).
</paragraph>
<paragraph id="8BD936994F28FFFE68F81FA3FE33FDDB" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="51">
<emphasis id="B912EA8B4F28FFFE68F81FA3FECBFD85" bold="true" box="[199,291,1410,1436]" pageId="7" pageNumber="51">Habitat</emphasis>
. Adults of this species were collected beside fast streams with stony bottoms ranging in elevation from
<quantity id="4C9E9B7C4F28FFFE68A81F86FEC3FDD8" box="[151,299,1447,1473]" metricMagnitude="2" metricUnit="m" metricValue="5.6" metricValueMax="11.0" metricValueMin="0.2" pageId="7" pageNumber="51" unit="m" value="560.0" valueMax="1100.0" valueMin="20.0">20 to 1100 m</quantity>
a.s.l. (
<figureCitation id="135D2A1C4F28FFFE694A1F86FE26FDDB" box="[373,462,1447,1474]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[232,1361,183,1919]" captionTargetId="figure-16@12.[225,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Adult collection sites of seven species of Rhyacophila on Yakushima. Solid circle = 14 adults collected; open circle = more than 5 adults collected." figureDoi="http://doi.org/10.5281/zenodo.5225422" httpUri="https://zenodo.org/record/5225422/files/figure.png" pageId="7" pageNumber="51">Fig. 6C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C65124F28FFFE68F81FEAFD2EFE10" pageId="7" pageNumber="51" type="etymology">
<paragraph id="8BD936994F28FFFE68F81FEAFAADFDFC" blockId="7.[151,1437,150,1545]" box="[199,1349,1483,1509]" pageId="7" pageNumber="51">
<emphasis id="B912EA8B4F28FFFE68F81FEAFEA0FDFC" bold="true" box="[199,328,1483,1509]" pageId="7" pageNumber="51">Etymology</emphasis>
. The name “
<taxonomicName id="4C664D1A4F28FFFE69E71FEAFDB9FDFD" authorityName="Ito" authorityYear="2021" box="[472,593,1483,1508]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="miyanoura">
<emphasis id="B912EA8B4F28FFFE69E71FEAFDB9FDFD" box="[472,593,1483,1508]" italics="true" pageId="7" pageNumber="51">miyanoura</emphasis>
</taxonomicName>
” is a noun in apposition, coined from the name of the
<typeStatus id="54DD883B4F28FFFE6C8B1FEAFB0CFDFC" box="[1204,1252,1483,1509]" pageId="7" pageNumber="51">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BD936994F28FFFE68F81FCEFD2EFE10" blockId="7.[151,1437,150,1545]" box="[199,710,1519,1545]" pageId="7" pageNumber="51">
<emphasis id="B912EA8B4F28FFFE68F81FCEFE94FE10" bold="true" box="[199,380,1519,1545]" pageId="7" pageNumber="51">Japanese name</emphasis>
. Miyanoura-nagare-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>