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<document id="990C2F9C18992A24471475A3654D25A7" ID-CLB-Dataset="57492" ID-DOI="10.5852/ejt.2023.885.2183" ID-GBIF-Dataset="f1935cd8-83c6-45fd-aac5-8c6659a934aa" ID-ISSN="2118-9773" ID-Zenodo-Dep="8177540" ID-ZooBank="A928AB6A-0029-4CEA-B941-C2238643835A" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201302250" checkinUser="felipe" docAuthor="Arriaga-Jiménez, Alfonsina, Zunino, Mario, Halffter, Gonzalo, Escobar-Hernández, Fernando &amp; Rossini, Michele" docDate="2023" docId="03CF324D04701F47FE6CF220FC80FBF9" docLanguage="en" docName="ejt-885_1-20.pdf" docOrigin="European Journal of Taxonomy 885" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ceratotrupes bolivari Halffter &amp; Martinez 1962" docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="FFF64A3504761F4FFF97F328FF93FFF7" masterDocTitle="A taxonomic review of the endemic Mexican genus Ceratotrupes Jekel, 1865 (Coleoptera: Geotrupidae: Ceratotrupini), with the description of an unexpected new species from the Sierra Madre del Sur" masterLastPageNumber="20" masterPageNumber="1" pageNumber="7" updateTime="1699544954118" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="861A4C76BF38D38B69F95E37E4ABEFD0">A taxonomic review of the endemic Mexican genus Ceratotrupes Jekel, 1865 (Coleoptera: Geotrupidae: Ceratotrupini), with the description of an unexpected new species from the Sierra Madre del Sur</mods:title>
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<mods:namePart id="0A2D460D788FCD48B2D7A5487B856E48">Arriaga-Jiménez, Alfonsina</mods:namePart>
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<mods:affiliation id="3B399ADDC7728C17E1EE1223E00D7FF3">Insect Ecology Lab, School of Environmental and Rural Sciences, University of New England, Armidale NSW 2351, Australia.</mods:affiliation>
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<mods:namePart id="B3D11A7BB41FCAFFDF5E609AACA1CE13">Zunino, Mario</mods:namePart>
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<mods:affiliation id="9D9FB0D3FF174E5BF6AAB75374F3BAB2">Instituto de Ecología A. C., Invited researcher, Carretera antigua a Coatepec 351, Xalapa 91070, Veracruz, Mexico.</mods:affiliation>
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<mods:namePart id="584D9C1E68F8939A234C039D38ED1F28">Halffter, Gonzalo</mods:namePart>
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<mods:affiliation id="D16E72DDF66FB6699F920678F187D7AD">Deceased 11 March 2022. Former address: Aldama Street 17, Centro, 91500 Coatepec, Veracruz, Mexico.</mods:affiliation>
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<mods:namePart id="85176DB900C64650BDC42969358301AB">Escobar-Hernández, Fernando</mods:namePart>
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<mods:affiliation id="093B20512C00F258EE920199A21A136D">Red de Ecoetología, Instituto de Ecología, INECOL A. C. Xalapa, México, Carretera antigua a Coatepec 351, El Haya, Xalapa 91070, Veracruz, Mexico.</mods:affiliation>
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<mods:affiliation id="D24F38E788DEF70F202E95907B8D468C">Finnish Museum of Natural History (LUOMUS), University of Helsinki, Pohjoinen Rautatiekatu 13, Helsinki, 00014, Finland. &amp; Department of Agronomy, Food, Natural resources, Animals and Environment (DAFNAE), University of Padova, Via dell'Università 16, 35020, Legnaro (PD), Italy.</mods:affiliation>
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<taxonomicName id="4C66F8D804701F49FE6CF220FBAAFED5" ID-CoL="SQJ4" authority="Halffter &amp; Martinez, 1962" authorityName="Halffter &amp; Martinez" authorityYear="1962" box="[507,1081,264,291]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B9125F4904701F49FE6CF220FD6FFED4" bold="true" box="[507,764,264,291]" italics="true" pageId="6" pageNumber="7">Ceratotrupes bolivari</emphasis>
<bibRefCitation id="EFF7FEAA04701F49FC94F220FBAAFED5" author="Halffter G. &amp; Martinez A." box="[771,1081,264,291]" pageId="6" pageNumber="7" pagination="145 - 159" refId="ref9786" refString="Halffter G. &amp; Martinez A. 1962. Monografia del genero Ceratotrupes Jekel (Coleopt., Scarab., Geotrup.). Ciencia 11 (4): 145 - 159." type="journal article" year="1962">Halffter &amp; Martínez, 1962</bibRefCitation>
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<figureCitation id="135D9FDE04701F49FD60F204FCAFFEB1" box="[759,828,300,326]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[240,1358,467,1881]" captionTargetId="figure-61@7.[189,1398,458,1891]" captionTargetPageId="7" captionText="Fig. 2. Ceratotrupes bolivari Halffter &amp; Martínez, 1962 (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Detail of the elliptical groove of the last antennal lamella. D–E. Aedeagus in dorsal (D) and lateral view (E)." figureDoi="http://doi.org/10.5281/zenodo.8177544" httpUri="https://zenodo.org/record/8177544/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
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<taxonomicName id="4C66F8D804701F49FF2AF246FCA3FE7F" ID-CoL="SQJ4" authority="Halffter &amp; Martinez 1962" authorityName="Halffter &amp; Martinez" authorityPageNumber="153" authorityYear="1962" box="[189,816,365,392]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B9125F4904701F49FF2AF246FE2FFE70" box="[189,444,365,392]" italics="true" pageId="6" pageNumber="7">Ceratotrupes bolivari</emphasis>
<bibRefCitation id="EFF7FEAA04701F49FE54F245FCA3FE7F" author="Halffter G. &amp; Martinez A." box="[451,816,365,392]" pageId="6" pageNumber="7" pagination="145 - 159" refId="ref9786" refString="Halffter G. &amp; Martinez A. 1962. Monografia del genero Ceratotrupes Jekel (Coleopt., Scarab., Geotrup.). Ciencia 11 (4): 145 - 159." type="journal article" year="1962">Halffter &amp; Martínez 1962: 153</bibRefCitation>
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<taxonomicName id="4C66F8D804701F49FF2AF287FE2DFE3F" ID-CoL="SQJ4" authorityName="Halffter &amp; Martinez" authorityYear="1962" box="[189,446,430,457]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B9125F4904701F49FF2AF287FE2DFE3F" box="[189,446,430,457]" italics="true" pageId="6" pageNumber="7">Ceratotrupes bolivari</emphasis>
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–
<treatmentCitation id="0AC7A54A04701F49FE49F286FD24FE3F" author="Howden H. F." box="[478,695,430,456]" page="46" pageId="6" pageNumber="7" year="1964">
<bibRefCitation id="EFF7FEAA04701F49FE49F286FD24FE3F" author="Howden H. F." box="[478,695,430,456]" pageId="6" pageNumber="7" pagination="5 - 91" refId="ref9941" refString="Howden H. F. 1964. The Geotrupinae of North and Central America. The Memoirs of the Entomological Society of Canada 96 (S 39): 5 - 91. https: // doi. org / 10.4039 / entm 9639 fv" type="journal article" year="1964">Howden 1964: 46</bibRefCitation>
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;
<treatmentCitation id="0AC7A54A04701F49FD53F287FCD3FE3E" author="Howden H. F." box="[708,832,430,457]" page="569" pageId="6" pageNumber="7" year="1974">
<bibRefCitation id="EFF7FEAA04701F49FD53F287FCD3FE3E" author="Howden H. F." box="[708,832,430,457]" pageId="6" pageNumber="7" pagination="567 - 575" refId="ref9986" refString="Howden H. F. 1974. Additional records and descriptions of North and Central American Geotrupinae (Coleoptera, Scarabaeidae). Canadian Journal of Zoology 52 (5): 567 - 575. https: // doi. org / 10.1139 / z 74 - 073" type="journal article" year="1974">1974: 569</bibRefCitation>
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;
<treatmentCitation id="0AC7A54A04701F49FCDAF287FC5BFE3E" author="Howden H. F." box="[845,968,431,457]" page="100" pageId="6" pageNumber="7" year="2003">
<bibRefCitation id="EFF7FEAA04701F49FCDAF287FC5BFE3E" author="Howden H. F." box="[845,968,431,457]" pageId="6" pageNumber="7" pagination="95 - 108" refId="ref10098" refString="Howden H. F. 2003. Subfamilia Geotrupinae. In: Moron M. A. (ed.) Atlas de los Escarabajos de Mexico. Coleoptera: Lamellicornia Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae: 95 - 108. Argania, Barcelona." type="book chapter" year="2003">2003: 100</bibRefCitation>
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. —
<treatmentCitation id="0AC7A54A04701F49FC6CF286FB47FE3E" author="Zunino M." box="[1019,1236,430,457]" page="109" pageId="6" pageNumber="7" year="1984">
<bibRefCitation id="EFF7FEAA04701F49FC6CF286FB47FE3E" author="Zunino M." box="[1019,1236,430,457]" pageId="6" pageNumber="7" pagination="9 - 162" refId="ref11069" refString="Zunino M. 1984. Sistematica generica dei Geotrupinae filogenesi della sottofamiglia e considerazioni biogeografiche. Bollettino del Museo regionale di Scienze naturali 2 (1): 9 - 162." type="journal article" year="1984">Zunino 1984: 109</bibRefCitation>
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. —
<treatmentCitation id="0AC7A54A04701F49FA90F287FE59FE1B" author="Moron M. A. &amp; Deloya C." page="216" pageId="6" pageNumber="7" year="1991">
<bibRefCitation id="EFF7FEAA04701F49FA90F287FE59FE1B" author="Moron M. A. &amp; Deloya C." pageId="6" pageNumber="7" pagination="209 - 283" refId="ref10299" refString="Moron M. A. &amp; Deloya C. 1991. Los coleopteros lamelicornios de la reserva de la biosfera ' La Michilia', Durango, Mexico. Folia entomologica mexicana 81: 209 - 283." type="journal article" year="1991">Morón &amp; Deloya 1991: 216</bibRefCitation>
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. —
<treatmentCitation id="0AC7A54A04701F49FD96F2FAFC0CFE1B" author="Navarrete-Heredia J. L. &amp; Delgado L. &amp; Lopez H. E. F." box="[513,927,466,492]" page="49" pageId="6" pageNumber="7" year="2001">
<bibRefCitation id="EFF7FEAA04701F49FD96F2FAFC0CFE1B" author="Navarrete-Heredia J. L. &amp; Delgado L. &amp; Lopez H. E. F." box="[513,927,466,492]" pageId="6" pageNumber="7" pagination="37 - 93" refId="ref10336" refString="Navarrete-Heredia J. L., Delgado L. &amp; Lopez H. E. F. 2001. Coleoptera Sacarabaeoidea de Jalisco, Mexico. Dugesiana 8 (1): 37 - 93." type="journal article" year="2001">
Navarrete-Heredia
<emphasis id="B9125F4904701F49FD71F2FBFC8EFE1B" box="[742,797,466,492]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2001: 49
</bibRefCitation>
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. —
<treatmentCitation id="0AC7A54A04701F49FC41F2FAFAD9FE1B" author="Delgado L. &amp; Marquez J." box="[982,1354,466,492]" page="84" pageId="6" pageNumber="7" year="2006">
<bibRefCitation id="EFF7FEAA04701F49FC41F2FAFAD9FE1B" author="Delgado L. &amp; Marquez J." box="[982,1354,466,492]" pageId="6" pageNumber="7" pagination="57 - 108" refId="ref9470" refString="Delgado L. &amp; Marquez J. 2006. Estado del conocimiento y conservacion de los coleopteros Scarabaeoidea (Insecta) del estado de Hidalgo, Mexico. Acta zoologica mexicana 22 (2): 57 - 108. https: // doi. org / 10.21829 / azm. 2006.2222015" type="journal article" year="2006">Delgado &amp; Márquez 2006: 84</bibRefCitation>
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<treatmentCitation id="0AC7A54A04701F49FF7DF2DDFD9FFDF8" author="Halffter G. &amp; Verdu J. R. &amp; Marquez J. &amp; Moreno C. E." box="[234,524,501,527]" page="284" pageId="6" pageNumber="7" year="2008">
<bibRefCitation id="EFF7FEAA04701F49FF7DF2DDFD9FFDF8" author="Halffter G. &amp; Verdu J. R. &amp; Marquez J. &amp; Moreno C. E." box="[234,524,501,527]" pageId="6" pageNumber="7" pagination="273 - 322" refId="ref9821" refString="Halffter G., Verdu J. R., Marquez J. &amp; Moreno C. E. 2008. Biogeographical analysis of Scarabaeinae and Geotrupinae along a transect in central Mexico (Coleoptera, Scarabaeoidea). Fragmenta Entomologica 40 (2): 273 - 322. https: // doi. org / 10.4081 / fe. 2008.99" type="journal article" year="2008">
Halffter
<emphasis id="B9125F4904701F49FED9F2DEFE11FDF8" box="[334,386,501,527]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2008: 284
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<treatmentCitation id="0AC7A54A04701F49FDAAF2DDFD74FDF8" author="Smith A. B. T." box="[573,743,501,527]" page="4" pageId="6" pageNumber="7" year="2009">
<bibRefCitation id="EFF7FEAA04701F49FDAAF2DDFD74FDF8" author="Smith A. B. T." box="[573,743,501,527]" pageId="6" pageNumber="7" refId="ref10728" refString="Smith A. B. T. 2009. Checklist and Nomenclatural Authority File of the Scarabaeoidea of the Nearctic Realm. Version 4. Available from https: // unsm-ento. unl. edu / SSSA / Nearctic-Scarabs 4. pdf [accessed 30 Apr. 2021]." type="url" year="2009">Smith 2009: 4</bibRefCitation>
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<bibRefCitation id="EFF7FEAA04701F49FC8FF2DDFB1DFDF8" author="Trotta-Moreu N. &amp; Lobo J. M." box="[792,1166,501,527]" pageId="6" pageNumber="7" pagination="42 - 49" refId="ref10835" refString="Trotta-Moreu N. &amp; Lobo J. M. 2010. Deriving the species richness distribution of Geotrupinae (Coleoptera: Scarabaeoidea) in Mexico from the overlap of individual model predictions. Environmental Entomology 39 (1): 42 - 49. https: // doi. org / 10.1603 / EN 08179" type="journal article" year="2010">Trotta-Moreu &amp; Lobo 2010: 46</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AC7A54A04701F49FB28F2DDFE37FDC4" author="Garcia de Jesus S. &amp; Moron M. &amp; Aragon A." page="151" pageId="6" pageNumber="7" year="2013">
<bibRefCitation id="EFF7FEAA04701F49FB28F2DDFE37FDC4" author="Garcia de Jesus S. &amp; Moron M. &amp; Aragon A." pageId="6" pageNumber="7" pagination="147 - 168" refId="ref9617" refString="Garcia de Jesus S., Moron M. &amp; Aragon A. 2013. Region del Metlalcueyetl (La Malinche). In: Moron M. (ed.) Fauna de Escarabajos del Estado de Puebla: 147 - 168." type="book chapter" year="2013">
García de Jesús
<emphasis id="B9125F4904701F49FF7CF131FE8DFDC5" box="[235,286,536,562]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2013: 151
</bibRefCitation>
</treatmentCitation>
, 158–159, 164–167. —
<treatmentCitation id="0AC7A54A04701F49FD28F130FB15FDC5" author="Perez-Villamares J. C. &amp; Jimenez-Sanchez E. &amp; Padilla-Ramirez J." box="[703,1158,536,563]" page="446" pageId="6" pageNumber="7" year="2016">
<bibRefCitation id="EFF7FEAA04701F49FD28F130FB15FDC5" author="Perez-Villamares J. C. &amp; Jimenez-Sanchez E. &amp; Padilla-Ramirez J." box="[703,1158,536,563]" pageId="6" pageNumber="7" pagination="443 - 450" refId="ref10435" refString="Perez-Villamares J. C., Jimenez-Sanchez E. &amp; Padilla-Ramirez J. 2016. Escarabajos atraidos a la carrona (Coleoptera: Scarabaeidae, Geotrupidae, Hybosoridae, Trogidae y Silphidae) en las canadas de Coatepec Harinas, Estado de Mexico, Mexico. Revista mexicana de Biodiversidad 87 (2): 443 - 450. https: // doi. org / 10.1016 / j. rmb. 2016.03.005" type="journal article" year="2016">
Pérez-Villamares
<emphasis id="B9125F4904701F49FC18F131FC50FDC5" box="[911,963,536,562]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2016: 446–448
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AC7A54A04701F49FB24F130FE14FDA2" author="Arriaga-Jimenez A. &amp; Ros M. &amp; Halffter G." page="8" pageId="6" pageNumber="7" year="2018">
<bibRefCitation id="EFF7FEAA04701F49FB24F130FE14FDA2" author="Arriaga-Jimenez A. &amp; Ros M. &amp; Halffter G." pageId="6" pageNumber="7" pagination="4468" refId="ref8897" refString="Arriaga-Jimenez A., Ros M. &amp; Halffter G. 2018. High variability of dung beetle diversity patterns at four mountains of the Trans-Mexican Volcanic Belt. PeerJ 6: e 4468. https: // doi. org / 10.7717 / peerj. 4468" type="journal article" year="2018">
Arriaga-Jiménez
<emphasis id="B9125F4904701F49FF7DF114FE8DFDA2" box="[234,286,571,597]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2018: 8
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC7A54A04701F49FE02F113FD9CFDA2" author="Arriaga-Jimenez A. &amp; Kohlmann B. &amp; Cruz-Garcia B. J." box="[405,527,571,597]" page="498" pageId="6" pageNumber="7" year="2020">
<bibRefCitation id="EFF7FEAA04701F49FE02F113FD9CFDA2" author="Arriaga-Jimenez A. &amp; Kohlmann B. &amp; Cruz-Garcia B. J." box="[405,527,571,597]" pageId="6" pageNumber="7" pagination="493 - 508" refId="ref8944" refString="Arriaga-Jimenez A., Kohlmann B. &amp; Cruz-Garcia B. J. 2020. A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae). Acta Entomologica Musei Nationalis Pragae 60 (2): 493 - 508. https: // doi. org / 10.37520 / aemnp. 2020.032" type="journal article" year="2020">2020: 498</bibRefCitation>
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. —
<treatmentCitation id="0AC7A54A04701F49FDABF113FCDDFDA1" author="Alvarado F. &amp; Salomao R. P. &amp; Hernandez-Rivera A. &amp; de Araujo Lira A. F." box="[572,846,571,598]" page="5" pageId="6" pageNumber="7" year="2020">
<bibRefCitation id="EFF7FEAA04701F49FDABF113FCDDFDA1" author="Alvarado F. &amp; Salomao R. P. &amp; Hernandez-Rivera A. &amp; de Araujo Lira A. F." box="[572,846,571,598]" pageId="6" pageNumber="7" pagination="103533" refId="ref8833" refString="Alvarado F., Salomao R. P., Hernandez-Rivera A. &amp; de Araujo Lira A. F. 2020. Different responses of dung beetle diversity and feeding guilds from natural and disturbed habitats across a subtropical elevational gradient. Acta Oecologica 104: e 103533. https: // doi. org / 10.1016 / j. actao. 2020.103533" type="journal article" year="2020">
Alvarado
<emphasis id="B9125F4904701F49FD26F114FD77FDA2" box="[689,740,571,597]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2020: 5
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37CD0D004701F49FF2AF154FD79FCFB" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BD9835B04701F49FF2AF154FEA5FD61" blockId="6.[189,310,636,662]" box="[189,310,636,662]" pageId="6" pageNumber="7">
<heading id="D091343704701F49FF2AF154FEA5FD61" bold="true" box="[189,310,636,662]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B9125F4904701F49FF2AF154FEA5FD61" bold="true" box="[189,310,636,662]" pageId="6" pageNumber="7">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD9835B04701F49FF2AF183FD79FCFB" blockId="6.[188,1398,683,780]" pageId="6" pageNumber="7">
Males and females of
<taxonomicName id="4C66F8D804701F49FE2BF184FDA8FD32" authorityName="Halffter &amp; Martinez" authorityYear="1962" box="[444,571,683,710]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B9125F4904701F49FE2BF184FDA8FD32" box="[444,571,683,710]" italics="true" pageId="6" pageNumber="7">C. bolivari</emphasis>
</taxonomicName>
have lateral edges of pronotum and anterior angles completely curved; apical antennal lamella with semi-elliptical groove (
<figureCitation id="135D9FDE04701F49FCB7F1E6FCEDFD1E" box="[800,894,718,745]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[240,1358,467,1881]" captionTargetId="figure-61@7.[189,1398,458,1891]" captionTargetPageId="7" captionText="Fig. 2. Ceratotrupes bolivari Halffter &amp; Martínez, 1962 (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Detail of the elliptical groove of the last antennal lamella. D–E. Aedeagus in dorsal (D) and lateral view (E)." figureDoi="http://doi.org/10.5281/zenodo.8177544" httpUri="https://zenodo.org/record/8177544/files/figure.png" pageId="6" pageNumber="7">Fig. 2C</figureCitation>
); anteromedial pronotal projection of male very long, almost reaching tip of cephalic horn.
</paragraph>
</subSubSection>
<subSubSection id="C37CD0D004701F49FF2AF01BFBBAFA2E" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BD9835B04701F49FF2AF01BFE7FFCBA" blockId="6.[189,492,819,845]" box="[189,492,819,845]" pageId="6" pageNumber="7">
<heading id="D091343704701F49FF2AF01BFE7FFCBA" bold="true" box="[189,492,819,845]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B9125F4904701F49FF2AF01BFE7FFCBA" bold="true" box="[189,492,819,845]" pageId="6" pageNumber="7">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD9835B04701F49FF7DF04AFAA5FB61" blockId="6.[189,1399,866,1174]" pageId="6" pageNumber="7">
<materialsCitation id="3B0E890604701F49FF7DF04AFC33FC35" ID-GBIF-Occurrence="4164095331" collectingDate="1953-09-20" collectionCode="CMNC" country="Mexico" elevation="2400" location="Morelos Huizilac" pageId="6" pageNumber="7" specimenCode="CMNC 00030977" specimenCount="1" specimenCount-male="1" stateProvince="Morelos" typeStatus="paratype">
<emphasis id="B9125F4904701F49FF7DF04AFEF4FC8B" bold="true" box="[234,359,866,892]" pageId="6" pageNumber="7">
<typeStatus id="54DD3DF904701F49FF7DF04AFEF4FC8B" box="[234,359,866,892]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F371C3CB04701F49FF2AF0ADFEA2FC68" box="[189,305,901,927]" name="Mexico" pageId="6" pageNumber="7">MEXICO</collectingCountry>
–
<emphasis id="B9125F4904701F49FEC3F0ADFE29FC68" bold="true" box="[340,442,901,927]" pageId="6" pageNumber="7">
<collectingRegion id="49A24DB904701F49FEC3F0ADFE29FC68" box="[340,442,901,927]" country="Mexico" name="Morelos" pageId="6" pageNumber="7">Morelos</collectingRegion>
</emphasis>
•
<specimenCount id="9D6048D204701F49FE4EF0ADFD98FC68" box="[473,523,901,927]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
; “
<location id="8EB9D58004701F49FDB2F0ADFD64FC68" LSID="urn:lsid:plazi:treatment:03CF324D04701F47FE6CF220FC80FBF9:8EB9D58004701F49FDB2F0ADFD64FC68" box="[549,759,901,927]" country="Mexico" name="Morelos Huizilac" pageId="6" pageNumber="7" stateProvince="Morelos">Morelos Huizilac</location>
<elevation id="004B646804701F49FC95F0ADFC0CFC68" box="[770,927,901,927]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="7" unit="m" value="2400.0">
<quantity id="4C9E2EBE04701F49FC95F0ADFCCDFC68" box="[770,862,901,927]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="7" unit="m" value="2400.0">2400 m</quantity>
a.s.l.
</elevation>
/
<date id="FFD8A59B04701F49FC2BF0ADFBCFFC57" box="[956,1116,901,928]" pageId="6" pageNumber="7" value="1953-09-20">
<collectingDate id="EF9C5C7304701F49FC2BF0ADFBCFFC57" box="[956,1116,901,928]" pageId="6" pageNumber="7" value="1953-09-20">20 Sep. 1953</collectingDate>
</date>
S. Bolivar / Paratype /
<taxonomicName id="4C66F8D804701F49FF2AF080FD27FC35" authority="Halffter &amp; Martinez" authorityName="Halffter &amp; Martinez" authorityYear="1962" box="[189,692,936,962]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bolivari">Ceratotrupes bolivari Halffter &amp; Martínez</taxonomicName>
”;
<specimenCode id="DBC02B2004701F49FD5BF080FC33FC35" box="[716,928,936,962]" collectionCode="CMNC" pageId="6" pageNumber="7">CMNC 00030977</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B0E890604701F49FC2BF080FE02FBFE" ID-GBIF-Occurrence="4164095324" collectingDate="1953-09-18" collectionCode="CMNC" collectorName="S. Bolivar, H. &amp; A. Howden" country="Mexico" elevation="2500" location="La Venta" pageId="6" pageNumber="7" specimenCode="CMNC 00030981" specimenCount="1" specimenCount-male="1" stateProvince="Morelos" typeStatus="paratype">
<specimenCount id="9D6048D204701F49FC2BF080FC7EFC34" box="[956,1005,936,963]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
; “La Venta
<elevation id="004B646804701F49FBECF080FA87FC35" box="[1147,1300,936,963]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="7" unit="m" value="2500.0">
<quantity id="4C9E2EBE04701F49FBECF080FB46FC34" box="[1147,1237,936,963]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="7" unit="m" value="2500.0">2500 m</quantity>
a.s.l.
</elevation>
<date id="FFD8A59B04701F49FA8AF080FF6BFC11" pageId="6" pageNumber="7" value="1953-09-18">
<collectingDate id="EF9C5C7304701F49FA8AF080FF6BFC11" pageId="6" pageNumber="7" value="1953-09-18">18 Sep. 1953</collectingDate>
</date>
/ S. Bolivar, H. &amp; A. Howden /
<typeStatus id="54DD3DF904701F49FDE2F0E4FD4FFC11" box="[629,732,972,998]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
/
<taxonomicName id="4C66F8D804701F49FD64F0E3FB73FC12" authority="Halffter &amp; Martinez" authorityName="Halffter &amp; Martinez" authorityYear="1962" box="[755,1248,971,997]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bolivari">Ceratotrupes bolivari Halffter &amp; Martínez</taxonomicName>
Collection”;
<specimenCode id="DBC02B2004701F49FF2AF0C7FE02FBFE" box="[189,401,1007,1033]" collectionCode="CMNC" pageId="6" pageNumber="7">CMNC 00030981</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B0E890604701F49FE3BF0C7FC11FBDB" ID-GBIF-Occurrence="4164095344" collectingDate="1952-11-20" collectionCode="CMNC" collectorName="J. Hendrichs" country="Mexico" location="Rio Frio" municipality="Valle Grande" pageId="6" pageNumber="7" specimenCode="CMNC 00030982" specimenCount="1" specimenCount-male="1" stateProvince="Morelos" typeStatus="paratype">
<specimenCount id="9D6048D204701F49FE3BF0C7FE4FFBFE" box="[428,476,1007,1033]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
; “
<collectingMunicipality id="6BBD192104701F49FE63F0C6FD07FBFF" box="[500,660,1006,1032]" pageId="6" pageNumber="7">Valle Grande</collectingMunicipality>
,
<location id="8EB9D58004701F49FD36F0C6FC96FBFF" LSID="urn:lsid:plazi:treatment:03CF324D04701F47FE6CF220FC80FBF9:8EB9D58004701F49FD36F0C6FC96FBFF" box="[673,773,1006,1032]" country="Mexico" municipality="Valle Grande" name="Rio Frio" pageId="6" pageNumber="7" stateProvince="Morelos">Rio Frio</location>
<date id="FFD8A59B04701F49FC9AF0C7FC3EFBFE" box="[781,941,1006,1033]" pageId="6" pageNumber="7" value="1952-11-20">
<collectingDate id="EF9C5C7304701F49FC9AF0C7FC3EFBFE" box="[781,941,1006,1033]" pageId="6" pageNumber="7" value="1952-11-20">20 Nov. 1952</collectingDate>
</date>
<collectorName id="2693E68D04701F49FC22F0C7FBDAFBFF" box="[949,1097,1006,1033]" pageId="6" pageNumber="7">J. Hendrichs</collectorName>
/
<typeStatus id="54DD3DF904701F49FBF5F0C7FB5AFBFE" box="[1122,1225,1007,1033]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
/
<taxonomicName id="4C66F8D804701F49FB76F0C7FD81FBDB" authority="Halffter &amp; Martinez" authorityName="Halffter &amp; Martinez" authorityYear="1962" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bolivari">Ceratotrupes bolivari Halffter &amp; Martínez</taxonomicName>
det. 1960”;
<specimenCode id="DBC02B2004701F49FD3DF73AFC11FBDB" box="[682,898,1042,1068]" collectionCode="CMNC" pageId="6" pageNumber="7">CMNC 00030982</specimenCode>
</materialsCitation>
. –
<materialsCitation id="3B0E890604701F49FC3DF73AFCEDFB85" ID-GBIF-Occurrence="4164095323" collectingDate="1950-09-22" collectionCode="CMNC" collectorName="Ray F. Smith &amp; Howden" country="Mexico" location="El Salto" municipality="La Campana" pageId="6" pageNumber="7" specimenCode="CMNC 00030984" specimenCount="1" specimenCount-male="1" stateProvince="Durango" typeStatus="paratype">
<emphasis id="B9125F4904701F49FC3DF73AFB8AFBDB" bold="true" box="[938,1049,1042,1068]" pageId="6" pageNumber="7">
<collectingRegion id="49A24DB904701F49FC3DF73AFB8AFBDB" box="[938,1049,1042,1068]" country="Mexico" name="Durango" pageId="6" pageNumber="7">Durango</collectingRegion>
</emphasis>
•
<specimenCount id="9D6048D204701F49FBA3F73AFBF6FBDB" box="[1076,1125,1042,1068]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
; “
<collectingMunicipality id="6BBD192104701F49FBE8F73AFA8FFBDB" box="[1151,1308,1042,1068]" pageId="6" pageNumber="7">La Campana</collectingMunicipality>
,
<quantity id="4C9E2EBE04701F49FABEF73AFAE4FBDB" box="[1321,1399,1042,1068]" metricMagnitude="4" metricUnit="m" metricValue="3.0577536" pageId="6" pageNumber="7" unit="mi" value="19.0">19 mi.</quantity>
W. of
<location id="8EB9D58004701F49FE92F71DFEF1FBB8" LSID="urn:lsid:plazi:treatment:03CF324D04701F47FE6CF220FC80FBF9:8EB9D58004701F49FE92F71DFEF1FBB8" box="[261,354,1077,1103]" country="Mexico" municipality="La Campana" name="El Salto" pageId="6" pageNumber="7" stateProvince="Durango">El Salto</location>
<date id="FFD8A59B04701F49FEFDF71DFE6EFBA7" box="[362,509,1077,1104]" pageId="6" pageNumber="7" value="1950-09-22">
<collectingDate id="EF9C5C7304701F49FEFDF71DFE6EFBA7" box="[362,509,1077,1104]" pageId="6" pageNumber="7" value="1950-09-22">22 Sep 1950</collectingDate>
</date>
<collectorName id="2693E68D04701F49FD93F71DFD0DFBB8" box="[516,670,1077,1103]" pageId="6" pageNumber="7">Ray F. Smith</collectorName>
/
<collectorName id="2693E68D04701F49FD22F71DFC8BFBB8" box="[693,792,1077,1103]" pageId="6" pageNumber="7">Howden</collectorName>
coll. ex.
<collectorName id="2693E68D04701F49FC14F71DFB82FBB8" box="[899,1041,1077,1103]" pageId="6" pageNumber="7">A. Martínez</collectorName>
coll. / Paratype /
<taxonomicName id="4C66F8D804701F49FB76F71DFE8BFB85" authority="Arriaga-Jiménez &amp; Zunino &amp; Halffter &amp; Escobar-Hernández &amp; Rossini, 2023" authorityName="Arriaga-Jiménez &amp; Zunino &amp; Halffter &amp; Escobar-Hernández &amp; Rossini" authorityYear="2023" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bolivari" status="sp. nov.">Ceratotrupes bolivari</taxonomicName>
<taxonomicNameLabel id="A221E23204701F49FE88F771FEDDFB84" box="[287,334,1113,1139]" pageId="6" pageNumber="7" rank="species">sp.n</taxonomicNameLabel>
(male) Halffter &amp; Martínez”;
<specimenCode id="DBC02B2004701F49FD3AF770FCEDFB85" box="[685,894,1112,1138]" collectionCode="CMNC" pageId="6" pageNumber="7">CMNC 00030984</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B0E890604701F49FC01F770FAA1FB61" ID-GBIF-Occurrence="4164095352" collectingDate="1953-09-19" collectionCode="CMNC" collectorName="Halffter &amp; Martínez" country="Mexico" elevation="2950" location="Rio Frio" pageId="6" pageNumber="7" specimenCode="CMNC 00030985" specimenCount="1" specimenCount-male="1" stateProvince="Durango" typeStatus="paratype">
<specimenCount id="9D6048D204701F49FC01F770FC57FB84" box="[918,964,1112,1139]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
; “
<location id="8EB9D58004701F49FC4DF770FBADFB85" LSID="urn:lsid:plazi:treatment:03CF324D04701F47FE6CF220FC80FBF9:8EB9D58004701F49FC4DF770FBADFB85" box="[986,1086,1112,1138]" country="Mexico" name="Rio Frio" pageId="6" pageNumber="7" stateProvince="Durango">Rio Frio</location>
<elevation id="004B646804701F49FBD3F770FB4AFB85" box="[1092,1241,1112,1139]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="6" pageNumber="7" unit="m" value="2950.0">
<quantity id="4C9E2EBE04701F49FBD3F770FB0FFB84" box="[1092,1180,1112,1139]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="6" pageNumber="7" unit="m" value="2950.0">2950 m</quantity>
a.s.l.
</elevation>
<date id="FFD8A59B04701F49FB48F770FAE4FB84" box="[1247,1399,1112,1139]" pageId="6" pageNumber="7" value="1953-09-19">
<collectingDate id="EF9C5C7304701F49FB48F770FAE4FB84" box="[1247,1399,1112,1139]" pageId="6" pageNumber="7" value="1953-09-19">19 Sep. 1953</collectingDate>
</date>
S. Bolivar / Paratype /
<taxonomicName id="4C66F8D804701F49FE5FF753FD53FB62" authority="Arriaga-Jiménez &amp; Zunino &amp; Halffter &amp; Escobar-Hernández &amp; Rossini, 2023" authorityName="Arriaga-Jiménez &amp; Zunino &amp; Halffter &amp; Escobar-Hernández &amp; Rossini" authorityYear="2023" box="[456,704,1147,1173]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bolivari" status="sp. nov.">Ceratotrupes bolivari</taxonomicName>
<taxonomicNameLabel id="A221E23204701F49FD5FF754FD6BFB61" box="[712,760,1148,1174]" pageId="6" pageNumber="7" rank="species">sp.n</taxonomicNameLabel>
(male) Halffter &amp; Martínez”;
<specimenCode id="DBC02B2004701F49FBCCF753FAA1FB61" box="[1115,1330,1147,1174]" collectionCode="CMNC" pageId="6" pageNumber="7">CMNC 00030985</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD9835B04701F49FF2AF794FDA7FB21" blockId="6.[189,564,1212,1238]" box="[189,564,1212,1238]" pageId="6" pageNumber="7">
<heading id="D091343704701F49FF2AF794FDA7FB21" bold="true" box="[189,564,1212,1238]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B9125F4904701F49FF2AF794FDA7FB21" bold="true" box="[189,564,1212,1238]" pageId="6" pageNumber="7">Additional material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD9835B04701F49FF2AF7C3FBBAFA2E" blockId="6.[189,1399,1259,1497]" pageId="6" pageNumber="7">
<materialsCitation id="3B0E890604701F49FF2AF7C3FC35FADE" ID-GBIF-Occurrence="4164095327" collectingDate="1987-07-17" collectingDateMax="1987-08-09" collectingDateMin="1987-07-17" collectionCode="CEMT" collectorName="S. Anduaga" country="Mexico" location="Reserva La Michilía" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Durango">
<collectingCountry id="F371C3CB04701F49FF2AF7C3FEA2FAF2" box="[189,305,1259,1285]" name="Mexico" pageId="6" pageNumber="7">MEXICO</collectingCountry>
–
<emphasis id="B9125F4904701F49FED8F7C3FE2DFAF2" bold="true" box="[335,446,1259,1285]" pageId="6" pageNumber="7">
<collectingRegion id="49A24DB904701F49FED8F7C3FE2DFAF2" box="[335,446,1259,1285]" country="Mexico" name="Durango" pageId="6" pageNumber="7">Durango</collectingRegion>
</emphasis>
•
<specimenCount id="9D6048D204701F49FE40F7C3FD95FAF1" box="[471,518,1259,1286]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
; “
<location id="8EB9D58004701F49FD8AF7C3FDEDFAF2" LSID="urn:lsid:plazi:treatment:03CF324D04701F47FE6CF220FC80FBF9:8EB9D58004701F49FD8AF7C3FDEDFAF2" box="[541,638,1259,1285]" country="Mexico" name="Reserva" pageId="6" pageNumber="7" stateProvince="Durango">Reserva</location>
‘La Michilía’
<date id="FFD8A59B04701F49FCBEF7C3FB84FAF2" box="[809,1047,1259,1285]" pageId="6" pageNumber="7" value="1987-07-17" valueMax="1987-08-09" valueMin="1987-07-17">
<collectingDate id="EF9C5C7304701F49FCBEF7C3FB84FAF2" box="[809,1047,1259,1285]" pageId="6" pageNumber="7" value="1987-07-17" valueMax="1987-08-09" valueMin="1987-07-17">17 Jul.–9 Aug. 1987</collectingDate>
</date>
bosque de pino-encino [pine-oak forest], trampa NTP 80 [NTP80 trap] /
<collectorName id="2693E68D04701F49FD24F627FCACFADF" box="[691,831,1294,1321]" pageId="6" pageNumber="7">S. Anduaga</collectorName>
”;
<collectionCode id="ED771B9E04701F49FCC2F627FC35FADE" box="[853,934,1295,1321]" pageId="6" pageNumber="7">CEMT</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0E890604701F49FC2AF627FCC3FABB" ID-GBIF-Occurrence="4164095319" collectingDate="1971-06-17" collectingDateMax="1971-06-19" collectingDateMin="1971-06-17" collectionCode="CMNC" collectorName="H. F. Howden" country="Mexico" location="El Salto." pageId="6" pageNumber="7" specimenCode="CMNC 00030989" specimenCount="1" specimenCount-female="1" stateProvince="Durango">
<specimenCount id="9D6048D204701F49FC2AF627FC7AFADE" box="[957,1001,1295,1321]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
; “
<quantity id="4C9E2EBE04701F49FB95F627FBDCFADF" box="[1026,1103,1294,1321]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="6" pageNumber="7" unit="mi" value="20.0">20 mi.</quantity>
E of
<location id="8EB9D58004701F49FB1BF626FB62FADF" LSID="urn:lsid:plazi:treatment:03CF324D04701F47FE6CF220FC80FBF9:8EB9D58004701F49FB1BF626FB62FADF" box="[1164,1265,1294,1320]" country="Mexico" name="El Salto." pageId="6" pageNumber="7" stateProvince="Durango">El Salto.</location>
8000’ a.s.l.
<date id="FFD8A59B04701F49FF2AF61AFEEDFABB" box="[189,382,1330,1356]" pageId="6" pageNumber="7" value="1971-06-17" valueMax="1971-06-19" valueMin="1971-06-17">
<collectingDate id="EF9C5C7304701F49FF2AF61AFEEDFABB" box="[189,382,1330,1356]" pageId="6" pageNumber="7" value="1971-06-17" valueMax="1971-06-19" valueMin="1971-06-17">17–19 Jun. 1971</collectingDate>
</date>
/
<collectorName id="2693E68D04701F49FE05F61AFDBEFABB" box="[402,557,1330,1356]" pageId="6" pageNumber="7">H.F. Howden</collectorName>
leg.”;
<specimenCode id="DBC02B2004701F49FDECF61AFCC3FABB" box="[635,848,1330,1356]" collectionCode="CMNC" pageId="6" pageNumber="7">CMNC 00030989</specimenCode>
</materialsCitation>
. –
<materialsCitation id="3B0E890604701F49FCF9F619FDACFA98" ID-GBIF-Occurrence="4164095337" collectingDate="1994-07-10" collectionCode="CEMT" collectorName="G. Nogueira" country="Mexico" elevation="1900" location="Loberas" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Sinaloa">
<emphasis id="B9125F4904701F49FCF9F619FC58FABC" bold="true" box="[878,971,1329,1355]" pageId="6" pageNumber="7">
<collectingRegion id="49A24DB904701F49FCF9F619FC58FABC" box="[878,971,1329,1355]" country="Mexico" name="Sinaloa" pageId="6" pageNumber="7">Sinaloa</collectingRegion>
</emphasis>
•
<specimenCount id="9D6048D204701F49FC74F61AFB9CFABB" box="[995,1039,1330,1356]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
; “
<location id="8EB9D58004701F49FBB0F61AFB1BFABB" LSID="urn:lsid:plazi:treatment:03CF324D04701F47FE6CF220FC80FBF9:8EB9D58004701F49FBB0F61AFB1BFABB" box="[1063,1160,1330,1356]" country="Mexico" name="Loberas" pageId="6" pageNumber="7" stateProvince="Sinaloa">Loberas</location>
<elevation id="004B646804701F49FB19F61AFAB0FABB" box="[1166,1315,1330,1356]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="7" unit="m" value="1900.0">
<quantity id="4C9E2EBE04701F49FB19F61AFB75FABB" box="[1166,1254,1330,1356]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="7" unit="m" value="1900.0">1900 m</quantity>
a.s.l.
</elevation>
<date id="FFD8A59B04701F49FABEF61AFF6BFA98" pageId="6" pageNumber="7" value="1994-07-10">
<collectingDate id="EF9C5C7304701F49FABEF61AFF6BFA98" pageId="6" pageNumber="7" value="1994-07-10">10 Jul. 1994</collectingDate>
</date>
/
<collectorName id="2693E68D04701F49FE99F67DFE0CFA98" box="[270,415,1365,1391]" pageId="6" pageNumber="7">G. Nogueira</collectorName>
leg.”;
<collectionCode id="ED771B9E04701F49FE79F67DFDACFA98" box="[494,575,1365,1391]" pageId="6" pageNumber="7">CEMT</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0E890604701F49FDC0F67DFECAFA65" ID-GBIF-Occurrence="4164095302" collectingDate="1994-08-06" collectionCode="CEMT" collectorName="D. Curoe" country="Mexico" elevation="1990" location="Loberas" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Sinaloa">
<specimenCount id="9D6048D204701F49FDC0F67DFD17FA98" box="[599,644,1365,1391]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
; “
<location id="8EB9D58004701F49FD0AF67DFD6DFA98" LSID="urn:lsid:plazi:treatment:03CF324D04701F47FE6CF220FC80FBF9:8EB9D58004701F49FD0AF67DFD6DFA98" box="[669,766,1365,1391]" country="Mexico" name="Loberas" pageId="6" pageNumber="7" stateProvince="Sinaloa">Loberas</location>
<elevation id="004B646804701F49FC92F67DFC0FFA98" box="[773,924,1365,1391]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="6" pageNumber="7" unit="m" value="1990.0">
<quantity id="4C9E2EBE04701F49FC92F67DFCCDFA98" box="[773,862,1365,1391]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="6" pageNumber="7" unit="m" value="1990.0">1990 m</quantity>
a.s.l.
</elevation>
<date id="FFD8A59B04701F49FC35F67DFBA0FA98" box="[930,1075,1365,1391]" pageId="6" pageNumber="7" value="1994-08-06">
<collectingDate id="EF9C5C7304701F49FC35F67DFBA0FA98" box="[930,1075,1365,1391]" pageId="6" pageNumber="7" value="1994-08-06">6 Aug. 1994</collectingDate>
</date>
luz Hg [Hg light]
<collectorName id="2693E68D04701F49FA9CF67DFAE4FA98" box="[1291,1399,1365,1391]" pageId="6" pageNumber="7">D. Curoe</collectorName>
leg.”;
<collectionCode id="ED771B9E04701F49FE91F650FECAFA65" box="[262,345,1400,1426]" pageId="6" pageNumber="7">CEMT</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B0E890604701F49FEE8F650FB83FA42" ID-GBIF-Occurrence="4164095333" collectingDate="1996-07-17" collectingDateMax="1996-07-18" collectingDateMin="1996-07-17" collectionCode="CEMT" collectorName="W.D. Edmonds and P Reyes" country="Mexico" elevation="1600" location="3 km NW of Yecora" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Sonora">
<emphasis id="B9125F4904701F49FEE8F650FE4AFA65" bold="true" box="[383,473,1400,1426]" pageId="6" pageNumber="7">
<collectingRegion id="49A24DB904701F49FEE8F650FE4AFA65" box="[383,473,1400,1426]" country="Mexico" name="Sonora" pageId="6" pageNumber="7">Sonora</collectingRegion>
</emphasis>
•
<specimenCount id="9D6048D204701F49FE64F650FDB7FA64" box="[499,548,1400,1427]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
; “
<location id="8EB9D58004701F49FDA8F650FCA4FA65" LSID="urn:lsid:plazi:treatment:03CF324D04701F47FE6CF220FC80FBF9:8EB9D58004701F49FDA8F650FCA4FA65" box="[575,823,1400,1427]" country="Mexico" name="3 km NW of Yecora" pageId="6" pageNumber="7" stateProvince="Sonora">
<quantity id="4C9E2EBE04701F49FDA8F650FDEFFA65" box="[575,636,1400,1426]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="km" value="3.0">3 km</quantity>
NW of Yécora
</location>
, on hwy 16
<elevation id="004B646804701F49FC5BF650FBF5FA65" box="[972,1126,1400,1427]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="7" unit="m" value="1600.0">
<quantity id="4C9E2EBE04701F49FC5BF650FBB5FA64" box="[972,1062,1400,1427]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="7" unit="m" value="1600.0">1600 m</quantity>
a.s.l.
</elevation>
<date id="FFD8A59B04701F49FBF9F650FAA2FA65" box="[1134,1329,1400,1426]" pageId="6" pageNumber="7" value="1996-07-17" valueMax="1996-07-18" valueMin="1996-07-17">
<collectingDate id="EF9C5C7304701F49FBF9F650FAA2FA65" box="[1134,1329,1400,1426]" pageId="6" pageNumber="7" value="1996-07-17" valueMax="1996-07-18" valueMin="1996-07-17">17–18 Jul. 1996</collectingDate>
</date>
. Oak, pine, juniper forest. human feces / W.D. Edmonds and P Reyes”;
<collectionCode id="ED771B9E04701F49FC28F6B3FB83FA42" box="[959,1040,1435,1461]" pageId="6" pageNumber="7">CEMT</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0E890604701F49FBBDF6B3FE28FA2E" ID-GBIF-Occurrence="4164095311" collectingDate="1996-07-17" collectingDateMax="1996-07-18" collectingDateMin="1996-07-17" collectionCode="CEMT" collectorName="W.D. Edmonds and P Reyes" country="Mexico" elevation="1600" location="3 km NW of Yecora" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Sonora">
<specimenCount id="9D6048D204701F49FBBDF6B3FBCAFA41" box="[1066,1113,1435,1462]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED771B9E04701F49FEFDF697FE28FA2E" box="[362,443,1471,1497]" pageId="6" pageNumber="7">CEMT</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0E890604701F49FE43F697FBB6FA2E" ID-GBIF-Occurrence="4164095334" box="[468,1061,1470,1497]" collectingDate="1996-07-17" collectingDateMax="1996-07-18" collectingDateMin="1996-07-17" collectionCode="CEMT" collectorName="W.D. Edmonds and P Reyes" country="Mexico" elevation="1600" location="3 km NW of Yecora" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Sonora">
<specimenCount id="9D6048D204701F49FE43F697FD92FA2E" box="[468,513,1471,1497]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED771B9E04701F49FC45F697FBB6FA2E" box="[978,1061,1471,1497]" pageId="6" pageNumber="7">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37CD0D004701F47FF2AF528FC06FDB4" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="8BD9835B04701F49FF2AF528FE6EF9ED" blockId="6.[189,509,1536,1562]" box="[189,509,1536,1562]" pageId="6" pageNumber="7">
<heading id="D091343704701F49FF2AF528FE6EF9ED" bold="true" box="[189,509,1536,1562]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B9125F4904701F49FF2AF528FE6EF9ED" bold="true" box="[189,509,1536,1562]" pageId="6" pageNumber="7">Redescription of the male</emphasis>
</heading>
</paragraph>
<paragraph id="8BD9835B04701F49FF2AF507FEA2F978" blockId="6.[188,1399,1582,1679]" pageId="6" pageNumber="7">
<smallCapsWord id="8D3F158704701F49FF2AF507FE8BF9B0" baselines="1602,1602" box="[189,280,1583,1609]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="6" pageNumber="7">COLOUR</smallCapsWord>
. Body black, sometimes with purplish to bluish tinges near pronotal edges, and on elytral striation; dorsally, shining, pronotal tegument sometimes silky; antennal segments brown, antennal club brownish.
</paragraph>
<paragraph id="8BD9835B04701F49FF2AF59FFC02F8AA" blockId="6.[188,1399,1718,1886]" pageId="6" pageNumber="7">
<smallCapsWord id="8D3F158704701F49FF2AF59FFF6FF938" baselines="1738,1738" box="[189,252,1719,1745]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="6" pageNumber="7">HEAD</smallCapsWord>
. Clypeus obtusely triangular, longitudinally crossed by a medial, high and triangular horn, projected forward, with apex curved upward; from posterior side, horn with medial, shallow depression; horn slightly narrow basally and thicker near apex, tip narrowly rounded; clypeal punctation dense and superficial; genae squared; frons smooth, unpunctured, with two oblique ridges along inner side of eyes. Apical antennal lamella with semi-elliptical groove (
<figureCitation id="135D9FDE04701F49FCB0F46BFC17F8AA" box="[807,900,1859,1885]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[240,1358,467,1881]" captionTargetId="figure-61@7.[189,1398,458,1891]" captionTargetPageId="7" captionText="Fig. 2. Ceratotrupes bolivari Halffter &amp; Martínez, 1962 (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Detail of the elliptical groove of the last antennal lamella. D–E. Aedeagus in dorsal (D) and lateral view (E)." figureDoi="http://doi.org/10.5281/zenodo.8177544" httpUri="https://zenodo.org/record/8177544/files/figure.png" pageId="6" pageNumber="7">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8BD9835B04701F48FF2AF4ADFCB1FE7A" blockId="6.[187,1398,1924,2021]" lastBlockId="7.[187,1399,265,397]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<smallCapsWord id="8D3F158704701F49FF2AF4ADFE8AF86A" baselines="1944,1944" box="[189,281,1925,1951]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="6" pageNumber="7">THORAX</smallCapsWord>
. In lateral view, pronotum weakly convex; in dorsal view, posterior edge of pronotum completely margined, lateral edges evenly curved from posterior to anterior angles; fine punctures concentrated on midline of pronotal projection, isolated punctures close to lateral and anterior edges; in lateral view, anteromedial projection dorsally slightly depressed; in dorsal view, basal region of projection with sides convergent, apical region with parallel or slightly divergent sides; apex of projection widely emarginated at middle. Elytral interstriae slightly convex, unpunctured. Ventral side of thoracic sclerites (i.e., pro-, meso- and metasternum) with long, yellowish setae.
</paragraph>
<caption id="DF19D3D304711F48FF2AF4A0FE3EF81F" ID-DOI="http://doi.org/10.5281/zenodo.8177544" ID-Zenodo-Dep="8177544" httpUri="https://zenodo.org/record/8177544/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1928,1954]" targetBox="[240,1358,467,1881]" targetPageId="7">
<paragraph id="8BD9835B04711F48FF2AF4A0FE3EF81F" blockId="7.[188,1399,1928,2024]" pageId="7" pageNumber="8">
<emphasis id="B9125F4904711F48FF2AF4A0FE98F855" bold="true" box="[189,267,1928,1954]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
<taxonomicName id="4C66F8D804711F48FE86F4A0FCD9F855" authority="Halffter &amp; Martinez, 1962" authorityName="Halffter &amp; Martinez" authorityYear="1962" box="[273,842,1928,1954]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B9125F4904711F48FE86F4A0FD83F855" box="[273,528,1928,1954]" italics="true" pageId="7" pageNumber="8">Ceratotrupes bolivari</emphasis>
<bibRefCitation id="EFF7FEAA04711F48FD81F4A0FCD9F855" author="Halffter G. &amp; Martinez A." box="[534,842,1928,1954]" pageId="7" pageNumber="8" pagination="145 - 159" refId="ref9786" refString="Halffter G. &amp; Martinez A. 1962. Monografia del genero Ceratotrupes Jekel (Coleopt., Scarab., Geotrup.). Ciencia 11 (4): 145 - 159." type="journal article" year="1962">Halffter &amp; Martínez, 1962</bibRefCitation>
</taxonomicName>
(CMNC).
<emphasis id="B9125F4904711F48FC5FF4A0FB93F855" bold="true" box="[968,1024,1928,1954]" pageId="7" pageNumber="8">A–B</emphasis>
. Dorsal habitus of male (A) and female (B).
<emphasis id="B9125F4904711F48FEDCF483FEF3F832" bold="true" box="[331,352,1963,1989]" pageId="7" pageNumber="8">C</emphasis>
. Detail of the elliptical groove of the last antennal lamella.
<emphasis id="B9125F4904711F48FB89F483FBC6F832" bold="true" box="[1054,1109,1963,1989]" pageId="7" pageNumber="8">D–E</emphasis>
. Aedeagus in dorsal (D) and lateral view (E).
</paragraph>
</caption>
<paragraph id="8BD9835B047E1F47FF2AF221FC1EFE9D" blockId="8.[189,1398,265,362]" pageId="8" pageNumber="9">
<smallCapsWord id="8D3F1587047E1F47FF2AF221FF65FED6" baselines="284,284" box="[189,246,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="8" pageNumber="9">LEGS</smallCapsWord>
. Protibiae robust, with six external teeth, ventral side with a longitudinal, crenulated carina; apical spur long and slender; meso- and metatibiae respectively with 3 and 4 entire, transversal carinae on posterior side; femora unmodified, with long yellowish setae.
</paragraph>
<paragraph id="8BD9835B047E1F47FF2AF2B8FE23FE5D" blockId="8.[189,432,400,426]" box="[189,432,400,426]" pageId="8" pageNumber="9">
<heading id="D0913437047E1F47FF2AF2B8FE23FE5D" bold="true" box="[189,432,400,426]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B9125F49047E1F47FF2AF2B8FE23FE5D" bold="true" box="[189,432,400,426]" pageId="8" pageNumber="9">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="8BD9835B047E1F47FF2AF297FC06FDB4" blockId="8.[189,1399,447,579]" pageId="8" pageNumber="9">Female specimens have clypeal horn distinctly lower; from above, clypeal horn triangular-shaped; pronotal punctation simple, shallower and sparse on disc, stronger and denser near lateral and anterior edges; anteromedial pronotal protuberance callus-like and transverse, depressed on superior side and close to anterior pronotal edge; anterior angles simply curved.</paragraph>
</subSubSection>
<subSubSection id="C37CD0D0047E1F47FF2AF142FC80FBF9" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="8BD9835B047E1F47FF2AF142FD9AFD73" blockId="8.[189,521,618,644]" box="[189,521,618,644]" pageId="8" pageNumber="9">
<heading id="D0913437047E1F47FF2AF142FD9AFD73" bold="true" box="[189,521,618,644]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B9125F49047E1F47FF2AF142FD9AFD73" bold="true" box="[189,521,618,644]" pageId="8" pageNumber="9">Ecology and biogeography</emphasis>
</heading>
</paragraph>
<paragraph id="8BD9835B047E1F47FF2AF1B1FD37FD0E" blockId="8.[189,1399,665,762]" pageId="8" pageNumber="9">
Among the members of the genus,
<taxonomicName id="4C66F8D8047E1F47FDFCF1B1FD7EFD44" authorityName="Halffter &amp; Martinez" authorityYear="1962" box="[619,749,665,691]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B9125F49047E1F47FDFCF1B1FD7EFD44" box="[619,749,665,691]" italics="true" pageId="8" pageNumber="9">C. bolivari</emphasis>
</taxonomicName>
is the species with the widest distribution (
<figureCitation id="135D9FDE047E1F47FA96F1B1FAD8FD44" box="[1281,1355,665,691]" captionStart="Fig" captionStartId="15.[189,232,1216,1242]" captionTargetBox="[208,1383,278,1155]" captionTargetId="figure-249@15.[189,1398,263,1170]" captionTargetPageId="15" captionText="Fig. 6. Geographic distribution of the genus Ceratotrupes Jekel, 1865." figureDoi="http://doi.org/10.5281/zenodo.8177554" httpUri="https://zenodo.org/record/8177554/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
). It inhabits pine, oak, or mixed pine-oak forests, and open areas with alpine pastures between 900 and
<quantity id="4C9E2EBE047E1F47FF2AF1F7FE85FD0D" box="[189,278,735,762]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="m" value="3000.0">3000 m</quantity>
a.s.l. (
<bibRefCitation id="EFF7FEAA047E1F47FEF4F1F7FD04FD0E" author="Halffter G. &amp; Martinez A." box="[355,663,735,761]" pageId="8" pageNumber="9" pagination="145 - 159" refId="ref9786" refString="Halffter G. &amp; Martinez A. 1962. Monografia del genero Ceratotrupes Jekel (Coleopt., Scarab., Geotrup.). Ciencia 11 (4): 145 - 159." type="journal article" year="1962">Halffter &amp; Martínez 1962</bibRefCitation>
).
</paragraph>
<paragraph id="8BD9835B047E1F47FF2AF009FC80FBF9" blockId="8.[189,1399,801,1038]" pageId="8" pageNumber="9">
The geographic range of this species encompasses the Sierra Madre Occidental. Its northernmost limit of distribution is located in the Mesa de Huracán with records from Yepachic,
<collectingRegion id="49A24DB9047E1F47FBC0F06CFB45FCA9" box="[1111,1238,836,862]" country="Mexico" name="Chihuahua" pageId="8" pageNumber="9">Chihuahua</collectingRegion>
(unpublished data, material collected by P. Reyes-Castillo and W.D. Edmonds in 1996 and deposited in MNHN and IEXA), while the southernmost records come from the Sierra de Tapalpa,
<collectingRegion id="49A24DB9047E1F47FBF5F0A2FB25FC53" box="[1122,1206,906,932]" country="Mexico" name="Jalisco" pageId="8" pageNumber="9">Jalisco</collectingRegion>
. To the east, the distribution of
<taxonomicName id="4C66F8D8047E1F47FEFAF086FE7EFC30" authorityName="Halffter &amp; Martinez" authorityYear="1962" box="[365,493,941,968]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B9125F49047E1F47FEFAF086FE7EFC30" box="[365,493,941,968]" italics="true" pageId="8" pageNumber="9">C. bolivari</emphasis>
</taxonomicName>
is limited by the Trans-Mexican Volcanic Belt, and specifically by the Cofre de Perote. To date, the easternmost records are a series of specimens collected in Omitlán de Juárez (Sierra de Pachuca,
<collectingRegion id="49A24DB9047E1F47FE3FF0DCFD9BFBF9" box="[424,520,1012,1038]" country="Mexico" name="Hidalgo" pageId="8" pageNumber="9">Hidalgo</collectingRegion>
) (
<bibRefCitation id="EFF7FEAA047E1F47FD88F0DCFC94FBF9" author="Halffter G. &amp; Verdu J. R. &amp; Marquez J. &amp; Moreno C. E." box="[543,775,1012,1038]" pageId="8" pageNumber="9" pagination="273 - 322" refId="ref9821" refString="Halffter G., Verdu J. R., Marquez J. &amp; Moreno C. E. 2008. Biogeographical analysis of Scarabaeinae and Geotrupinae along a transect in central Mexico (Coleoptera, Scarabaeoidea). Fragmenta Entomologica 40 (2): 273 - 322. https: // doi. org / 10.4081 / fe. 2008.99" type="journal article" year="2008">
Halffter
<emphasis id="B9125F49047E1F47FD12F0DDFD2CFBF9" box="[645,703,1012,1038]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2008
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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