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<document id="990759333BD08F7B13909F21F3D29013" ID="10.11646/zootaxa.4242.1.8SLASH10438" ID-CLB-Dataset="36313" ID-DOI="10.11646/zootaxa.4242.1.8" ID-GBIF-Dataset="ba343a99-6ace-4ebf-8d07-5ac585a63d39" ID-ISSN="1175-5326" ID-Zenodo-Dep="376068" ID-ZooBank="D18EDA2A-92BD-49A9-95A6-DA3F3CAC4336" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489051298620" checkinUser="plazi" docAuthor="Mvogo Ndongo, Pierre A., Schubart, Christoph D., Rintelen, Thomas Von, Tamesse, Joseph L. &amp; Cumberlidge, Neil" docDate="2017" docId="03CF007BFFCFC42A7FB62F5D1466FB69" docLanguage="en" docName="zootaxa.4242.1.8SLASH10438.pdf" docOrigin="Zootaxa 4242 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sudanonautes tiko Mvogo Ndongo, Schubart &amp; Cumberlidge, n. sp." docType="treatment" docVersion="9" lastPageNumber="171" masterDocId="FFF67803FFCDC4207F2129111051FF81" masterDocTitle="Morphological and molecular evidence for a new species of freshwater crab of the genus Sudanonautes Bott, 1955 (Brachyura: Potamoidea: Potamonautidae) from Cameroon, with notes on its ecology" masterLastPageNumber="173" masterPageNumber="161" pageNumber="163" updateTime="1698437774838" updateUser="ExternalLinkService">
<mods:mods id="33655D92E55108A05D182BEBEE6E8EBF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3C6ABDA0DD728FF2EF70F29C250FB1FD">
<mods:title id="A69C7DF6E73FE81109ECC1BB5B5010C6">Morphological and molecular evidence for a new species of freshwater crab of the genus Sudanonautes Bott, 1955 (Brachyura: Potamoidea: Potamonautidae) from Cameroon, with notes on its ecology</mods:title>
</mods:titleInfo>
<mods:name id="16A146C1A7D74AB5321246FB9D66F7C0" type="personal">
<mods:role id="B9905F5128B65BE5C55126F7CD09F00E">
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<mods:namePart id="D3B09CB2B5E8ECABB1F07ACD8B5656BB">Mvogo Ndongo, Pierre A.</mods:namePart>
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<mods:name id="D76AF1762E22D72051594F7432F23CBF" type="personal">
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<mods:namePart id="FE307623FC1DA53564AEB0EE2A67C6D2">Schubart, Christoph D.</mods:namePart>
</mods:name>
<mods:name id="25C6105D444099FD385E4579C1183E35" type="personal">
<mods:role id="F50C8018E535FFECB67D72F6ABDA53F3">
<mods:roleTerm id="2AFE978F2021D532371E2BB85B07BCFE">Author</mods:roleTerm>
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<mods:namePart id="7EA52EFA8D6D92D28DD0C515D21B3D3B">Rintelen, Thomas Von</mods:namePart>
</mods:name>
<mods:name id="256054E2AC9DE43E506AB25A6836BACD" type="personal">
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<mods:roleTerm id="53448F33CF29F60175F6C8F84E5B8B0A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A20C59AB76ED2F7DEBD32E73CD887A0C">Tamesse, Joseph L.</mods:namePart>
</mods:name>
<mods:name id="48D97397B084715983E3A734F48CA32D" type="personal">
<mods:role id="B4256F797DF446A0D3F9AEEF6B1272A7">
<mods:roleTerm id="A4641611FCC99D7D13217AEBFC1CFA16">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C446A4ED4BF9E8A69F98479E3026EB7E">Cumberlidge, Neil</mods:namePart>
</mods:name>
<mods:typeOfResource id="887B5A06831F5B2BEE1E45592F5E3176">text</mods:typeOfResource>
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<mods:title id="8B8B974EFBF76ECC6CF7F8BA05B41D24">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D9EA11734D7D02CBDC38C3BF691245BA">
<mods:date id="6039CDE0A71E424E3E85260D4E1E4398">2017</mods:date>
<mods:detail id="B688946B4B48203D293596B77C808221" type="volume">
<mods:number id="C4D9D7B9B11ADA14AC50BEA21B134C27">4242</mods:number>
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<mods:identifier id="040AE762DD303F73BF48ED503E08FC79" type="CLB-Dataset">36313</mods:identifier>
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<treatment id="03CF007BFFCFC42A7FB62F5D1466FB69" ID-DOI="http://doi.org/10.5281/zenodo.5697098" ID-GBIF-Taxon="127721674" ID-Zenodo-Dep="5697098" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69" httpUri="http://treatment.plazi.org/id/03CF007BFFCFC42A7FB62F5D1466FB69" lastPageId="10" lastPageNumber="171" pageId="2" pageNumber="163">
<subSubSection id="C37CE2E6FFCFC4227FB62F5D126BF906" pageId="2" pageNumber="163" type="nomenclature">
<paragraph id="8BD9B16DFFCFC4227FB62F5D1387F9E7" blockId="2.[151,982,1612,1707]" box="[151,982,1612,1638]" pageId="2" pageNumber="163">
<heading id="D0910601FFCFC4227FB62F5D1387F9E7" bold="true" box="[151,982,1612,1638]" fontSize="11" level="1" pageId="2" pageNumber="163" reason="1">
<emphasis id="B9126D7FFFCFC4227FB62F5D1387F9E7" bold="true" box="[151,982,1612,1638]" pageId="2" pageNumber="163">
<taxonomicName id="4C66CAEEFFCFC4227FB62F5D13DDF9E7" ID-CoL="7B2Z6" authority="Mvogo Ndongo, Schubart &amp; Cumberlidge" authorityName="Mvogo Ndongo, Schubart &amp; Cumberlidge" box="[151,908,1612,1638]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFCFC4227FB62F5D1128F9E7" bold="true" box="[151,377,1612,1638]" italics="true" pageId="2" pageNumber="163">Sudanonautes tiko</emphasis>
Mvogo Ndongo, Schubart &amp; Cumberlidge
</taxonomicName>
<taxonomicNameLabel id="A221D004FFCFC4227CB52F5D1387F9E7" box="[916,982,1612,1638]" pageId="2" pageNumber="163" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD9B16DFFCFC4227FB62F7E126BF906" blockId="2.[151,982,1612,1707]" box="[151,570,1646,1671]" pageId="2" pageNumber="163">
Common name:
<taxonomicName id="4C66CAEEFFCFC4227E702F7F11D4F906" box="[337,389,1646,1671]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
freshwater crab
</paragraph>
</subSubSection>
<subSubSection id="C37CE2E6FFCFC4227FB62F831164F92A" box="[151,309,1682,1707]" pageId="2" pageNumber="163" type="description">
<paragraph id="8BD9B16DFFCFC4227FB62F831164F92A" blockId="2.[151,982,1612,1707]" box="[151,309,1682,1707]" pageId="2" pageNumber="163">
(
<figureCitation id="135DADE8FFCFC4227FBE2F83115BF92A" box="[159,266,1682,1707]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[151,250,1882,1904]" captionStartId-1="5.[151,250,1915,1937]" captionStartId-2="6.[151,250,1233,1255]" captionStartId-3="7.[151,250,1055,1077]" captionTargetBox-0="[235,1349,203,1854]" captionTargetBox-1="[241,1315,203,1890]" captionTargetBox-2="[237,1350,197,1207]" captionTargetBox-3="[235,1349,520,1030]" captionTargetId-0="figure@4.[228,1359,195,1858]" captionTargetId-1="figure@5.[233,1328,193,1899]" captionTargetId-2="figure@6.[231,1356,193,1212]" captionTargetId-3="figure@7.[233,1354,517,1033]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Sudanonautes tiko n. sp. from Cameroon. Habitus in dorsal view: a, holotype, adult male (CW 36.2 mm) ZMB Crust. 29628, Tamba Forest, Tiko (colour in ethanol); b, paratype, adult male (CW 35.4 mm) ZMB Crust. 29629 from Edea (colour in life). Scale bar equals 16 mm." captionText-1="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." captionText-2="FIGURE 3. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, left G 1 ventral view; b: left G 1 dorsal view; c, lleft G 2 ventral view. Scale bar equals 2 mm (a, b, c)." captionText-3="FIGURE 4. Habitat of Sudanonautes tiko n. sp. at Edea, Cameroon." httpUri-0="https://zenodo.org/record/376069/files/figure.png" httpUri-1="https://zenodo.org/record/376070/files/figure.png" httpUri-2="https://zenodo.org/record/376071/files/figure.png" httpUri-3="https://zenodo.org/record/376072/files/figure.png" pageId="2" pageNumber="163">Figs. 14</figureCitation>
,
<figureCitation id="135DADE8FFCFC4227E372F831177F92A" box="[278,294,1682,1707]" captionStart="FIGURE 6" captionStartId="9.[151,250,1367,1389]" captionTargetBox="[233,1354,411,1340]" captionTargetId="figure@9.[225,1362,409,1346]" captionTargetPageId="9" captionText="FIGURE 6. Whole animal dorsal views showing living color of three species of Sudanonautes from Cameroon. a, S. africanus adult female, CW 87.4 mm, from Tiko (LZUY- 03); b, S. aubryi subadult male, CW 35.8 mm, from Edea, Bolekoa, Littoral Region (LZUY- 06); c, S. floweri adult male, CW 35.4 mm, from Yabassi, Parc des Princes, Littoral Region (LZUY- 09); d, S. floweri adult female, CW 53.9 mm, from Tiko, Tamba forest, Southwest Region (LZUY- 11). Scale bar equals 72 mm (for a), 32 mm (for b), 32 mm (for c), 45 mm (for d)." httpUri="https://zenodo.org/record/376074/files/figure.png" pageId="2" pageNumber="163">6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37CE2E6FFCFC4237FB62FCB13A4FB91" lastPageId="3" lastPageNumber="164" pageId="2" pageNumber="163" type="materials_examined">
<paragraph id="8BD9B16DFFCFC4227FB62FCB1402F802" blockId="2.[151,1437,1754,2031]" pageId="2" pageNumber="163">
<emphasis id="B9126D7FFFCFC4227FB62FCB1224F972" bold="true" box="[151,629,1754,1779]" pageId="2" pageNumber="163">
Type material examined.
<materialsCitation id="3B0EBB30FFCFC4227EE52FCB1224F972" ID-GBIF-Occurrence="1438485818" box="[452,629,1754,1779]" country="Cameroon" location="Tamba Forest" pageId="2" pageNumber="163" specimenCount="1">
<collectingCountry id="F371F1FDFFCFC4227EE52FCB1221F972" box="[452,624,1754,1779]" name="Cameroon" pageId="2" pageNumber="163">CAMEROON</collectingCountry>
.
</materialsCitation>
</emphasis>
<taxonomicName id="4C66CAEEFFCFC4227D5A2FCB12E3F972" box="[635,690,1754,1779]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
,
<location id="8EB9E7B6FFCFC4227D9D2FCB1305F973" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69:8EB9E7B6FFCFC4227D9D2FCB1305F973" box="[700,852,1754,1779]" country="Cameroon" name="Tamba Forest" pageId="2" pageNumber="163">Tamba Forest</location>
,
<materialsCitation id="3B0EBB30FFCFC4227C432FCB1423F8BA" ID-GBIF-Occurrence="1438485828" collectingDate="2015-08-22" collectionCode="ZMB, DNA" country="Switzerland" elevation="100" latitude="4.1805553" location="Southwest Region" pageId="2" pageNumber="163" specimenCount="2" specimenCount-adult="1" specimenCount-male="1" stateProvince="Southwest Region" typeStatus="holotype">
<collectingRegion id="49A27F8FFFCFC4227C432FCB147FF972" box="[866,1070,1754,1779]" country="Cameroon" name="South-West" pageId="2" pageNumber="163">Southwest Region</collectingRegion>
(
<geoCoordinate id="EE52D7AAFFCFC4227B1C2FCA1481F972" box="[1085,1232,1754,1779]" direction="north" orientation="latitude" pageId="2" pageNumber="163" precision="15" value="4.1805553">04° 10' 50''N</geoCoordinate>
, 09° 23' 40'E)
<quantity id="4C9E1C88FFCFC4227A532FCA10FCF897" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="163" unit="m" value="100.0">
<elevation id="004B565EFFCFC4227A532FCA10FCF897" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="163" unit="m" value="100.0">100 m</elevation>
</quantity>
asl,
<specimenCount id="9D607AE4FFCFC4227FC52FEE1163F896" box="[228,306,1790,1815]" pageId="2" pageNumber="163" type="adult">1 adult</specimenCount>
<specimenCount id="9D607AE4FFCFC4227E1B2FEE111EF897" box="[314,335,1791,1814]" pageId="2" pageNumber="163" type="male"></specimenCount>
<typeStatus id="54DD0FCFFFCFC4227E762FEF11E8F896" box="[343,441,1790,1815]" pageId="2" pageNumber="163" type="holotype">holotype</typeStatus>
, CW
<quantity id="4C9E1C88FFCFC4227EDF2FEF1234F897" box="[510,613,1790,1815]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="2" pageNumber="163" unit="mm" value="36.2">36.2 mm</quantity>
, CL
<quantity id="4C9E1C88FFCFC4227DBE2FEE1357F897" box="[671,774,1791,1815]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="2" pageNumber="163" unit="mm" value="24.8">24.8 mm</quantity>
,
<collectingCountry id="F371F1FDFFCFC4227C322FEE136BF896" box="[787,826,1791,1815]" name="Switzerland" pageId="2" pageNumber="163">CH</collectingCountry>
<quantity id="4C9E1C88FFCFC4227C632FEE13FBF897" box="[834,938,1791,1815]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="2" pageNumber="163" unit="mm" value="12.2">12.2 mm</quantity>
, FW
<quantity id="4C9E1C88FFCFC4227CC82FEE1400F897" box="[1001,1105,1791,1815]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="163" unit="mm" value="10.4">10.4 mm</quantity>
,
<date id="FFD897ADFFCFC4227B7F2FEE14ABF896" box="[1118,1274,1790,1815]" pageId="2" pageNumber="163" value="2015-08-22">
<collectingDate id="EF9C6E45FFCFC4227B7F2FEE14ABF896" box="[1118,1274,1790,1815]" pageId="2" pageNumber="163" value="2015-08-22">22 Aug. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCFC4227A062FEE156FF896" box="[1319,1342,1791,1815]" country="Nigeria" name="Rivers" pageId="2" pageNumber="163">A</collectingRegion>
. Mvogo Ndongo) (
<collectionCode id="ED7729A8FFCFC4227E2F2E32111DF8BA" box="[270,332,1827,1851]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="163">ZMB</collectionCode>
Crust. 29628;
<collectionCode id="ED7729A8FFCFC4227D202E32126FF8BA" box="[513,574,1827,1851]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="163">DNA</collectionCode>
extraction code T262-11). Edea, Mbus Michon
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCFC4227BA32E331402F802" ID-GBIF-Occurrence="1438485826" collectingDate="2015-07-10" collectionCode="ZMB, DNA" country="Switzerland" elevation="137" location="Littoral Region" pageId="2" pageNumber="163" specimenCount="1" specimenCount-male="1" stateProvince="Littoral Region" typeStatus="paratype">
<collectingRegion id="49A27F8FFFCFC4227BA32E331562F8BA" box="[1154,1331,1826,1851]" country="Cameroon" name="Littoral" pageId="2" pageNumber="163">
<collectingRegion id="49A27F8FFFCFC4227BA32E331487F8BA" box="[1154,1238,1826,1851]" country="Equatorial Guinea" name="Litoral" pageId="2" pageNumber="163">Littoral</collectingRegion>
Region
</collectingRegion>
(03° 43' 146''N, 010° 7' 734''E),
<quantity id="4C9E1C88FFCFC4227E872E5611BFF8DF" box="[422,494,1863,1887]" metricMagnitude="2" metricUnit="m" metricValue="1.3699999999999999" pageId="2" pageNumber="163" unit="m" value="137.0">
<elevation id="004B565EFFCFC4227E872E5611BFF8DF" box="[422,494,1863,1887]" metricMagnitude="2" metricUnit="m" metricValue="1.3699999999999999" pageId="2" pageNumber="163" unit="m" value="137.0">137 m</elevation>
</quantity>
asl, adult
<typeStatus id="54DD0FCFFFCFC4227D472E561297F8DE" box="[614,710,1863,1887]" pageId="2" pageNumber="163" type="paratype">paratype</typeStatus>
<specimenCount id="9D607AE4FFCFC4227DEE2E5612B7F8DF" box="[719,742,1863,1886]" pageId="2" pageNumber="163" type="male"></specimenCount>
, CW
<quantity id="4C9E1C88FFCFC4227C092E5713C1F8DF" box="[808,912,1862,1887]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" pageId="2" pageNumber="163" unit="mm" value="35.4">35.4 mm</quantity>
, CL
<quantity id="4C9E1C88FFCFC4227CE32E571479F8DF" box="[962,1064,1862,1887]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="2" pageNumber="163" unit="mm" value="25.1">25.1 mm</quantity>
,
<collectingCountry id="F371F1FDFFCFC4227B142E56140DF8DE" box="[1077,1116,1863,1887]" name="Switzerland" pageId="2" pageNumber="163">CH</collectingCountry>
12.0 mm, FW
<quantity id="4C9E1C88FFCFC4227A2D2E561522F8DF" box="[1292,1395,1863,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="2" pageNumber="163" unit="mm" value="10.3">10.3 mm</quantity>
,
<date id="FFD897ADFFCFC4227AA12E5610ACF802" pageId="2" pageNumber="163" value="2015-07-10">
<collectingDate id="EF9C6E45FFCFC4227AA12E5610ACF802" pageId="2" pageNumber="163" value="2015-07-10">10 Jul. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCFC4227E092E7A116EF802" box="[296,319,1899,1923]" country="Nigeria" name="Rivers" pageId="2" pageNumber="163">A</collectingRegion>
. Mvogo Ndongo) (
<collectionCode id="ED7729A8FFCFC4227D342E7A1202F802" box="[533,595,1899,1923]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="163">ZMB</collectionCode>
Crust. 29629;
<collectionCode id="ED7729A8FFCFC4227DD82E7A1367F802" box="[761,822,1899,1923]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="163">DNA</collectionCode>
extraction code T262-7).
</materialsCitation>
</paragraph>
<paragraph id="8BD9B16DFFCFC4237FE62E9F13CDFF55" blockId="2.[151,1437,1754,2031]" lastBlockId="3.[151,1437,151,1999]" lastPageId="3" lastPageNumber="164" pageId="2" pageNumber="163">
<emphasis id="B9126D7FFFCFC4227FE62E9F1257F826" bold="true" box="[199,518,1934,1959]" pageId="2" pageNumber="163">Other material examined.</emphasis>
CAMEROON. Edea, Mbus Michon,
<materialsCitation id="3B0EBB30FFCFC4227CE72E9F1388F86E" ID-GBIF-Occurrence="1438485833" collectingDate="2015-07-09" collectionCode="DNA" country="Switzerland" elevation="137" location="Bwenga forest" pageId="2" pageNumber="163" specimenCount="1" specimenCount-male="1" stateProvince="Littoral Region">
<collectingRegion id="49A27F8FFFCFC4227CE72E9F1426F826" box="[966,1143,1934,1959]" country="Cameroon" name="Littoral" pageId="2" pageNumber="163">
<collectingRegion id="49A27F8FFFCFC4227CE72E9F144BF826" box="[966,1050,1934,1959]" country="Equatorial Guinea" name="Litoral" pageId="2" pageNumber="163">Littoral</collectingRegion>
Region
</collectingRegion>
(03° 43' 146''N, 010° 7' 734''E)
<quantity id="4C9E1C88FFCFC4227FCC2EA21164F84B" box="[237,309,1971,1995]" metricMagnitude="2" metricUnit="m" metricValue="1.3699999999999999" pageId="2" pageNumber="163" unit="m" value="137.0">
<elevation id="004B565EFFCFC4227FCC2EA21164F84B" box="[237,309,1971,1995]" metricMagnitude="2" metricUnit="m" metricValue="1.3699999999999999" pageId="2" pageNumber="163" unit="m" value="137.0">137 m</elevation>
</quantity>
asl, adult
<specimenCount id="9D607AE4FFCFC4227E8D2EA21192F84B" box="[428,451,1971,1994]" pageId="2" pageNumber="163" type="male"></specimenCount>
, CW
<quantity id="4C9E1C88FFCFC4227D242EA3122BF84B" box="[517,634,1970,1995]" metricMagnitude="-2" metricUnit="m" metricValue="2.869" pageId="2" pageNumber="163" unit="mm" value="28.69">28.69 mm</quantity>
, CL
<quantity id="4C9E1C88FFCFC4227D922EA21379F84B" box="[691,808,1971,1995]" metricMagnitude="-2" metricUnit="m" metricValue="2.071" pageId="2" pageNumber="163" unit="mm" value="20.71">20.71 mm</quantity>
,
<collectingCountry id="F371F1FDFFCFC4227C152EA2130AF84A" box="[820,859,1971,1995]" name="Switzerland" pageId="2" pageNumber="163">CH</collectingCountry>
<quantity id="4C9E1C88FFCFC4227C422EA3139AF84B" box="[867,971,1970,1995]" metricMagnitude="-3" metricUnit="m" metricValue="9.65" pageId="2" pageNumber="163" unit="mm" value="9.65">9.65 mm</quantity>
, FW
<quantity id="4C9E1C88FFCFC4227B282EA31420F84B" box="[1033,1137,1970,1995]" metricMagnitude="-3" metricUnit="m" metricValue="9.16" pageId="2" pageNumber="163" unit="mm" value="9.16">9.16 mm</quantity>
,
<date id="FFD897ADFFCFC4227B5C2EA214AAF84A" box="[1149,1275,1970,1995]" pageId="2" pageNumber="163" value="2015-07-09">
<collectingDate id="EF9C6E45FFCFC4227B5C2EA214AAF84A" box="[1149,1275,1970,1995]" pageId="2" pageNumber="163" value="2015-07-09">9 Jul. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCFC4227A092EA2156EF84A" box="[1320,1343,1971,1995]" country="Nigeria" name="Rivers" pageId="2" pageNumber="163">A</collectingRegion>
. Mvogo Ndongo) (LZUY-01;
<collectionCode id="ED7729A8FFCFC4227EA82EC61197F86E" box="[393,454,2007,2031]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="163">DNA</collectionCode>
extraction code T262-8);
<taxonomicName id="4C66CAEEFFCFC4227DCE2EC71377F86E" box="[751,806,2006,2031]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
,
<location id="8EB9E7B6FFCFC4227C132EC61388F86E" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69:8EB9E7B6FFCFC4227C132EC61388F86E" box="[818,985,2006,2031]" country="Switzerland" name="Bwenga forest" pageId="2" pageNumber="163" stateProvince="Littoral Region">Bwenga forest</location>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCFC4237CC42EC713CDFF55" ID-GBIF-Occurrence="1438485827" collectingDate="2015-08-24" collectionCode="RMNH, CRUST, DNA" country="Switzerland" elevation="38" lastPageId="3" lastPageNumber="164" latitude="4.0477777" location="Southwest Region" pageId="2" pageNumber="163" specimenCount="1" specimenCount-male="1" stateProvince="Southwest Region">
<collectingRegion id="49A27F8FFFCFC4227CC42EC714E2F86E" box="[997,1203,2006,2031]" country="Cameroon" name="South-West" pageId="2" pageNumber="163">Southwest Region</collectingRegion>
(
<geoCoordinate id="EE52D7AAFFCFC4227BE52EC6150BF86E" box="[1220,1370,2006,2031]" direction="north" orientation="latitude" pageId="2" pageNumber="163" precision="15" value="4.0477777">04° 02' 52''N</geoCoordinate>
, 009° 19' 22'E),
<quantity id="4C9E1C88FFCEC4237E2A29891114FF2E" box="[267,325,152,176]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="164" unit="m" value="38.0">
<elevation id="004B565EFFCEC4237E2A29891114FF2E" box="[267,325,152,176]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="164" unit="m" value="38.0">38 m</elevation>
</quantity>
asl, adult
<specimenCount id="9D607AE4FFCEC4237E9D29891182FF2E" box="[444,467,152,175]" pageId="3" pageNumber="164" type="male"></specimenCount>
, CW
<quantity id="4C9E1C88FFCEC4237D372986122CFF2E" box="[534,637,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="3.36" pageId="3" pageNumber="164" unit="mm" value="33.6">33.6 mm</quantity>
, CL
<quantity id="4C9E1C88FFCEC4237D972989134FFF2E" box="[694,798,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="3" pageNumber="164" unit="mm" value="23.1">23.1 mm</quantity>
,
<collectingCountry id="F371F1FDFFCEC4237C0A29891303FF31" box="[811,850,152,176]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237C7B29891390FF2E" box="[858,961,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="3" pageNumber="164" unit="mm" value="10.1">10.1 mm</quantity>
, FW
<quantity id="4C9E1C88FFCEC4237B2029861403FF31" box="[1025,1106,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="164" unit="mm" value="8.5">8.5mm</quantity>
,
<date id="FFD897ADFFCEC4237B7F298914ABFF31" box="[1118,1274,151,176]" pageId="3" pageNumber="164" value="2015-08-24">
<collectingDate id="EF9C6E45FFCEC4237B7F298914ABFF31" box="[1118,1274,151,176]" pageId="3" pageNumber="164" value="2015-08-24">24 Aug. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237A092989156EFF31" box="[1320,1343,152,176]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (
<collectionCode id="ED7729A8FFCEC4237E2829AD1132FF54" box="[265,355,188,213]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="164">RMNH</collectionCode>
.
<collectionCode id="ED7729A8FFCEC4237E4729AD1194FF54" box="[358,453,188,213]" pageId="3" pageNumber="164">CRUST</collectionCode>
.D.57073;
<collectionCode id="ED7729A8FFCEC4237D1429AD1223FF55" box="[565,626,188,212]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="164">DNA</collectionCode>
extraction code T273-12).
</materialsCitation>
</paragraph>
<paragraph id="8BD9B16DFFCEC4237FE629CE13A4FB91" blockId="3.[151,1437,151,1999]" pageId="3" pageNumber="164">
<emphasis id="B9126D7FFFCEC4237FE629CE11BDFF79" bold="true" box="[199,492,223,248]" italics="true" pageId="3" pageNumber="164">
<taxonomicName id="4C66CAEEFFCEC4237FE629CE11B9FF79" box="[199,488,223,248]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="africanus">Sudanonautes africanus</taxonomicName>
.
</emphasis>
<materialsCitation id="3B0EBB30FFCEC4237ED929F112DEFE9D" ID-GBIF-Occurrence="1438485830" country="Cameroon" latitude="4.0477777" location="Bwenga forest" pageId="3" pageNumber="164" specimenCount="1" specimenCount-male="1" stateProvince="Southwest Region">
<collectingCountry id="F371F1FDFFCEC4237ED929F112CDFF79" box="[504,668,224,248]" name="Cameroon" pageId="3" pageNumber="164">CAMEROON</collectingCountry>
.
<taxonomicName id="4C66CAEEFFCEC4237D8B29CE12B0FF79" box="[682,737,223,248]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
,
<location id="8EB9E7B6FFCEC4237DD129F113CBFF79" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69:8EB9E7B6FFCEC4237DD129F113CBFF79" box="[752,922,223,248]" country="Cameroon" latitude="4.0477777" name="Bwenga forest" pageId="3" pageNumber="164" stateProvince="Southwest Region">Bwenga forest</location>
,
<collectingRegion id="49A27F8FFFCEC4237C8B29CE142AFF79" box="[938,1147,223,248]" country="Cameroon" name="South-West" pageId="3" pageNumber="164">Southwest Region</collectingRegion>
(
<geoCoordinate id="EE52D7AAFFCEC4237BAF29F1157AFF79" box="[1166,1323,223,248]" direction="north" orientation="latitude" pageId="3" pageNumber="164" precision="15" value="4.0477777">04° 02' 52''N</geoCoordinate>
, 009° 19' 22'E), adult
<specimenCount id="9D607AE4FFCEC4237E042814116DFE9D" box="[293,316,261,284]" pageId="3" pageNumber="164" type="male"></specimenCount>
, CW
<quantity id="4C9E1C88FFCEC4237EA3281511BBFE9D" box="[386,490,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="9.28" pageId="3" pageNumber="164" unit="mm" value="92.8">92.8 mm</quantity>
, CL
<quantity id="4C9E1C88FFCEC4237D07281512DEFE9D" box="[550,655,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="6.859999999999999" pageId="3" pageNumber="164" unit="mm" value="68.6">68.6 mm</quantity>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237DBF2815110DFEC1" ID-GBIF-Occurrence="1438485819" collectingDate="2015-08-24" country="Switzerland" location="A" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237DBF28151294FE9D" box="[670,709,260,284]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237DEF28151366FE9D" box="[718,823,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="3" pageNumber="164" unit="mm" value="27.6">27.6 mm</quantity>
, FW
<quantity id="4C9E1C88FFCEC4237C58281513B1FE9D" box="[889,992,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="3" pageNumber="164" unit="mm" value="25.9">25.9 mm</quantity>
<date id="FFD897ADFFCEC4237CCB281514D8FE9C" box="[1002,1161,260,285]" pageId="3" pageNumber="164" value="2015-08-24">
<collectingDate id="EF9C6E45FFCEC4237CCB281514D8FE9C" box="[1002,1161,260,285]" pageId="3" pageNumber="164" value="2015-08-24">24 Aug. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237B9828151481FE9D" box="[1209,1232,260,284]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-02);
<taxonomicName id="4C66CAEEFFCEC4237E042836110DFEC1" box="[293,348,295,320]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237E46283615C6FEBE" ID-GBIF-Occurrence="1438485831" box="[359,1431,295,320]" country="Switzerland" elevation="37" latitude="4.0477777" location="Southwest Region" pageId="3" pageNumber="164" specimenCount="1" specimenCount-female="1" stateProvince="Southwest Region">
<collectingRegion id="49A27F8FFFCEC4237E4628361265FEC1" box="[359,564,295,320]" country="Cameroon" name="South-West" pageId="3" pageNumber="164">Southwest Region</collectingRegion>
(
<geoCoordinate id="EE52D7AAFFCEC4237D6228391289FEC1" box="[579,728,295,320]" direction="north" orientation="latitude" pageId="3" pageNumber="164" precision="15" value="4.0477777">04° 02' 52''N</geoCoordinate>
, 009° 19' 22'E),
<quantity id="4C9E1C88FFCEC4237CB02839139BFEBE" box="[913,970,296,320]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="164" unit="m" value="37.0">
<elevation id="004B565EFFCEC4237CB02839139BFEBE" box="[913,970,296,320]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="164" unit="m" value="37.0">37 m</elevation>
</quantity>
asl, adult
<specimenCount id="9D607AE4FFCEC4237B1F28391402FEBE" box="[1086,1107,296,319]" pageId="3" pageNumber="164" type="female"></specimenCount>
, CW
<quantity id="4C9E1C88FFCEC4237BB4283914AAFEBE" box="[1173,1275,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="8.74" pageId="3" pageNumber="164" unit="mm" value="87.4">87.4 mm</quantity>
, CL 66.0 mm
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237FB6285D11C3FE06" ID-GBIF-Occurrence="1438485825" collectingDate="2015-08-26" collectionCode="DNA" country="Switzerland" location="A" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237FB6285D10EFFEE5" box="[151,190,332,356]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237FE6285D1161FEE5" box="[199,304,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="3" pageNumber="164" unit="mm" value="26.9">26.9 mm</quantity>
, FW
<quantity id="4C9E1C88FFCEC4237E53285D118AFEE5" box="[370,475,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="3" pageNumber="164" unit="mm" value="23.5">23.5 mm</quantity>
,
<date id="FFD897ADFFCEC4237EC8285D12D9FEE4" box="[489,648,332,357]" pageId="3" pageNumber="164" value="2015-08-26">
<collectingDate id="EF9C6E45FFCEC4237EC8285D12D9FEE4" box="[489,648,332,357]" pageId="3" pageNumber="164" value="2015-08-26">26 Aug. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237D99285D129EFEE5" box="[696,719,332,356]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-03;
<collectionCode id="ED7729A8FFCEC4237B0D285D1438FEE5" box="[1068,1129,332,356]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="164">DNA</collectionCode>
extraction code T273-10); Edea, Bolekoa stream
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237EBF287E1134FE2D" ID-GBIF-Occurrence="1438485838" country="Switzerland" elevation="115" location="Littoral Region" pageId="3" pageNumber="164" specimenCount="3" specimenCount-male="1" specimenCount-subadult="2" stateProvince="Littoral Region">
<collectingRegion id="49A27F8FFFCEC4237EBF287E121AFE09" box="[414,587,367,392]" country="Cameroon" name="Littoral" pageId="3" pageNumber="164">
<collectingRegion id="49A27F8FFFCEC4237EBF287E11A3FE09" box="[414,498,367,392]" country="Equatorial Guinea" name="Litoral" pageId="3" pageNumber="164">Littoral</collectingRegion>
Region
</collectingRegion>
(03° 51' 288''N; 010° 4' 110''E),
<quantity id="4C9E1C88FFCEC4237CE5287E145AFE06" box="[964,1035,367,392]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="164" unit="m" value="115.0">
<elevation id="004B565EFFCEC4237CE5287E145AFE06" box="[964,1035,367,392]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="164" unit="m" value="115.0">115 m</elevation>
</quantity>
asl,
<specimenCount id="9D607AE4FFCEC4237B60286114E6FE09" box="[1089,1207,367,392]" pageId="3" pageNumber="164" type="subadult">2 subadult</specimenCount>
<specimenCount id="9D607AE4FFCEC4237B9E28611487FE06" box="[1215,1238,368,391]" pageId="3" pageNumber="164" type="male"></specimenCount>
, largest CW
<quantity id="4C9E1C88FFCEC4237A4A28611094FE2D" metricMagnitude="-2" metricUnit="m" metricValue="7.83" pageId="3" pageNumber="164" unit="mm" value="78.3">78.3 mm</quantity>
, CL
<quantity id="4C9E1C88FFCEC4237FDF28851134FE2D" box="[254,357,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="5.82" pageId="3" pageNumber="164" unit="mm" value="58.2">58.2 mm</quantity>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237E532885121DFE4E" ID-GBIF-Occurrence="1438485832" collectingDate="2013-11-26" collectionCode="DNA" country="Switzerland" location="A" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237E53288511C8FE2D" box="[370,409,404,428]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237E8028851259FE2D" box="[417,520,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="3" pageNumber="164" unit="mm" value="22.5">22.5 mm</quantity>
, FW
<quantity id="4C9E1C88FFCEC4237D69288512FEFE2D" box="[584,687,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="3" pageNumber="164" unit="mm" value="21.2">21.2 mm</quantity>
,
<date id="FFD897ADFFCEC4237D9A28851304FE2D" box="[699,853,404,429]" pageId="3" pageNumber="164" value="2013-11-26">
<collectingDate id="EF9C6E45FFCEC4237D9A28851304FE2D" box="[699,853,404,429]" pageId="3" pageNumber="164" value="2013-11-26">26 Nov. 2013</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237CA2288513CBFE2D" box="[899,922,404,428]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo (LZUY-04,
<collectionCode id="ED7729A8FFCEC4237BC628851575FE2D" box="[1255,1316,404,428]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="164">DNA</collectionCode>
extraction code R1028-1); Edea, Bolekoa stream
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237D7828A61270FE75" ID-GBIF-Occurrence="1438485829" country="Switzerland" elevation="115" location="Littoral Region" pageId="3" pageNumber="164" specimenCount="3" specimenCount-female="1" specimenCount-subadult="2" stateProvince="Littoral Region">
<collectingRegion id="49A27F8FFFCEC4237D7828A61356FE51" box="[601,775,439,464]" country="Cameroon" name="Littoral" pageId="3" pageNumber="164">
<collectingRegion id="49A27F8FFFCEC4237D7828A612FCFE51" box="[601,685,439,464]" country="Equatorial Guinea" name="Litoral" pageId="3" pageNumber="164">Littoral</collectingRegion>
Region
</collectingRegion>
(03° 51' 288''N; 010° 4' 110''E),
<quantity id="4C9E1C88FFCEC4237BA328A6149BFE4E" box="[1154,1226,439,464]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="164" unit="m" value="115.0">
<elevation id="004B565EFFCEC4237BA328A6149BFE4E" box="[1154,1226,439,464]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="164" unit="m" value="115.0">115 m</elevation>
</quantity>
asl,
<specimenCount id="9D607AE4FFCEC4237A2328A91529FE51" box="[1282,1400,439,464]" pageId="3" pageNumber="164" type="subadult">2 subadult</specimenCount>
<specimenCount id="9D607AE4FFCEC4237AA128A915C4FE4E" box="[1408,1429,440,463]" pageId="3" pageNumber="164" type="female"></specimenCount>
, largest CW
<quantity id="4C9E1C88FFCEC4237E3C28CD11D2FE75" box="[285,387,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="6.26" pageId="3" pageNumber="164" unit="mm" value="62.6">62.6 mm</quantity>
, CL
<quantity id="4C9E1C88FFCEC4237E9A28CD1270FE75" box="[443,545,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="4.72" pageId="3" pageNumber="164" unit="mm" value="47.2">47.2 mm</quantity>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237D0C28CD1357FD99" ID-GBIF-Occurrence="1438485835" collectingDate="2015-08-26" collectionCode="DNA" country="Switzerland" location="A" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237D0C28CD1205FE75" box="[557,596,476,500]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237D7D28CD1293FE75" box="[604,706,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="3" pageNumber="164" unit="mm" value="18.7">18.7 mm</quantity>
, FW
<quantity id="4C9E1C88FFCEC4237C2128CD1337FE75" box="[768,870,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="3" pageNumber="164" unit="mm" value="16.2">16.2 mm</quantity>
,
<date id="FFD897ADFFCEC4237C5328CD145DFE74" box="[882,1036,476,501]" pageId="3" pageNumber="164" value="2015-08-26">
<collectingDate id="EF9C6E45FFCEC4237C5328CD145DFE74" box="[882,1036,476,501]" pageId="3" pageNumber="164" value="2015-08-26">26 Aug. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237B1928CD141EFE75" box="[1080,1103,476,500]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-05;
<collectionCode id="ED7729A8FFCEC4237FB62B111085FD99" box="[151,212,512,536]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="164">DNA</collectionCode>
extraction code T263-4).
<taxonomicName id="4C66CAEEFFCEC4237D2828EE1357FD99" box="[521,774,511,536]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B9126D7FFFCEC4237D2828EE1357FD99" bold="true" box="[521,774,511,536]" italics="true" pageId="3" pageNumber="164">Sudanonautes aubryi</emphasis>
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="3B0EBB30FFCEC4237C372B1113A4FDBD" ID-GBIF-Occurrence="1438485823" country="Cameroon" elevation="115" location="Bolekoa" municipality="Edea" pageId="3" pageNumber="164" specimenCount="1" specimenCount-male="1" stateProvince="Littoral Region">
<collectingCountry id="F371F1FDFFCEC4237C372B1113EBFD99" box="[790,954,512,536]" name="Cameroon" pageId="3" pageNumber="164">CAMEROON</collectingCountry>
.
<collectingMunicipality id="6BBD2B17FFCEC4237CE828EE1455FD99" box="[969,1028,511,536]" pageId="3" pageNumber="164">Edea</collectingMunicipality>
,
<location id="8EB9E7B6FFCEC4237B3228EE1422FD99" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69:8EB9E7B6FFCEC4237B3228EE1422FD99" box="[1043,1139,511,536]" country="Cameroon" municipality="Edea" name="Bolekoa" pageId="3" pageNumber="164" stateProvince="Littoral Region">Bolekoa</location>
,
<collectingRegion id="49A27F8FFFCEC4237BA228EE1565FD99" box="[1155,1332,511,536]" country="Cameroon" name="Littoral" pageId="3" pageNumber="164">
<collectingRegion id="49A27F8FFFCEC4237BA228EE1486FD99" box="[1155,1239,511,536]" country="Equatorial Guinea" name="Litoral" pageId="3" pageNumber="164">Littoral</collectingRegion>
Region
</collectingRegion>
(03° 51' 288''N, 010° 4' 110''E),
<quantity id="4C9E1C88FFCEC4237EBE2B3511B4FDBD" box="[415,485,548,573]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="164" unit="m" value="115.0">
<elevation id="004B565EFFCEC4237EBE2B3511B4FDBD" box="[415,485,548,573]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="164" unit="m" value="115.0">115 m</elevation>
</quantity>
asl, subadult
<specimenCount id="9D607AE4FFCEC4237D5E2B3412C7FDBD" box="[639,662,549,572]" pageId="3" pageNumber="164" type="male"></specimenCount>
, CW
<quantity id="4C9E1C88FFCEC4237DF72B35131BFDBD" box="[726,842,548,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="3" pageNumber="164" unit="mm" value="35.8">35.80 mm</quantity>
, CL
<quantity id="4C9E1C88FFCEC4237CA02B3513A4FDBD" box="[897,1013,548,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.505" pageId="3" pageNumber="164" unit="mm" value="25.05">25.05 mm</quantity>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237B202B3514D3FDE1" ID-GBIF-Occurrence="1438485822" collectingDate="2014-07-07" collectionCode="DNA" country="Switzerland" location="A" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237B202B351479FDBD" box="[1025,1064,548,572]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237B0E2B3514F2FDBD" box="[1071,1187,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.2029999999999998" pageId="3" pageNumber="164" unit="mm" value="12.03">12.03 mm</quantity>
, FW
<quantity id="4C9E1C88FFCEC4237BFE2B351502FDBD" box="[1247,1363,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.031" pageId="3" pageNumber="164" unit="mm" value="10.31">10.31 mm</quantity>
,
<date id="FFD897ADFFCEC4237A7E2B34109EFDE1" pageId="3" pageNumber="164" value="2014-07-07">
<collectingDate id="EF9C6E45FFCEC4237A7E2B34109EFDE1" pageId="3" pageNumber="164" value="2014-07-07">7 Jul. 2014</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237FDA2B591143FDE1" box="[251,274,584,608]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-06;
<collectionCode id="ED7729A8FFCEC4237D492B5912F4FDE1" box="[616,677,584,608]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="164">DNA</collectionCode>
extraction code R1026-9); Edea, Bolekoa
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237BAF2B561386FD05" ID-GBIF-Occurrence="1438485839" country="Switzerland" elevation="115" location="Littoral Region" pageId="3" pageNumber="164" specimenCount="1" specimenCount-female="1" stateProvince="Littoral Region">
<collectingRegion id="49A27F8FFFCEC4237BAF2B56156AFDE1" box="[1166,1339,583,608]" country="Cameroon" name="Littoral" pageId="3" pageNumber="164">
<collectingRegion id="49A27F8FFFCEC4237BAF2B5614B3FDE1" box="[1166,1250,583,608]" country="Equatorial Guinea" name="Litoral" pageId="3" pageNumber="164">Littoral</collectingRegion>
Region
</collectingRegion>
(03° 51' 288''N, 010° 4' 110'' E),
<quantity id="4C9E1C88FFCEC4237E822B7D11B9FD05" box="[419,488,620,645]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="164" unit="m" value="115.0">
<elevation id="004B565EFFCEC4237E822B7D11B9FD05" box="[419,488,620,645]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="164" unit="m" value="115.0">115 m</elevation>
</quantity>
asl, subadult
<specimenCount id="9D607AE4FFCEC4237DA12B7C12C4FD05" box="[640,661,621,644]" pageId="3" pageNumber="164" type="female"></specimenCount>
, CW 34.0 mm, CL
<quantity id="4C9E1C88FFCEC4237C502B7D1386FD05" box="[881,983,620,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="3" pageNumber="164" unit="mm" value="24.5">24.5 mm</quantity>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237CC02B7D13AAFD29" ID-GBIF-Occurrence="1438485834" collectingDate="2015-08-27" collectionCode="DNA" country="Switzerland" location="A" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237CC02B7D1459FD05" box="[993,1032,620,644]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237B2E2B7D1437FD05" box="[1039,1126,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="3" pageNumber="164" unit="mm" value="9.9">9.9 mm</quantity>
, FW
<quantity id="4C9E1C88FFCEC4237B832B7D14A8FD05" box="[1186,1273,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="3" pageNumber="164" unit="mm" value="9.2">9.2 mm</quantity>
,
<date id="FFD897ADFFCEC4237A252B7D15CDFD04" box="[1284,1436,620,645]" pageId="3" pageNumber="164" value="2015-08-27">
<collectingDate id="EF9C6E45FFCEC4237A252B7D15CDFD04" box="[1284,1436,620,645]" pageId="3" pageNumber="164" value="2015-08-27">27 Aug. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237F9B2B811080FD29" box="[186,209,656,680]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-07;
<collectionCode id="ED7729A8FFCEC4237D002B81120FFD29" box="[545,606,656,680]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="164">DNA</collectionCode>
extraction code T273-5); Didzangué
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237B272B9E14B7FD29" ID-GBIF-Occurrence="1438485841" box="[1030,1254,655,680]" country="Iceland" location="Lake" pageId="3" pageNumber="164" specimenCount="1" stateProvince="Lake">
<collectingCountry id="F371F1FDFFCEC4237B272B9E141BFD29" box="[1030,1098,655,680]" name="Iceland" pageId="3" pageNumber="164">Island</collectingCountry>
of
<collectingRegion id="49A27F8FFFCEC4237B4E2B9E14F6FD29" box="[1135,1191,655,680]" country="South Sudan" name="Lakes" pageId="3" pageNumber="164">Lake</collectingRegion>
Ossa
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237BD02B9E14CBFD4D" ID-GBIF-Occurrence="1438485820" country="Iceland" elevation="90" latitude="3.8159833" location="Littoral Region" longLatPrecision="1" longitude="10.05515" pageId="3" pageNumber="164" specimenCount="3" specimenCount-male="1" specimenCount-subadult="2" stateProvince="Littoral Region">
<collectingRegion id="49A27F8FFFCEC4237BD02B9E15CDFD29" box="[1265,1436,655,680]" country="Cameroon" name="Littoral" pageId="3" pageNumber="164">
<collectingRegion id="49A27F8FFFCEC4237BD02B9E1514FD29" box="[1265,1349,655,680]" country="Equatorial Guinea" name="Litoral" pageId="3" pageNumber="164">Littoral</collectingRegion>
Region
</collectingRegion>
(
<geoCoordinate id="EE52D7AAFFCEC4237FBE2BA51167FD4D" box="[159,310,692,717]" direction="north" orientation="latitude" pageId="3" pageNumber="164" precision="1" value="3.8159833">03°48.959' N</geoCoordinate>
,
<geoCoordinate id="EE52D7AAFFCEC4237E1B2BA5118EFD4D" box="[314,479,692,716]" direction="east" orientation="longitude" pageId="3" pageNumber="164" precision="1" value="10.05515">010°03.309' E</geoCoordinate>
),
<quantity id="4C9E1C88FFCEC4237ED52BA5127FFD4D" box="[500,558,692,716]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="164" unit="m" value="90.0">
<elevation id="004B565EFFCEC4237ED52BA5127FFD4D" box="[500,558,692,716]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="164" unit="m" value="90.0">90 m</elevation>
</quantity>
asl,
<specimenCount id="9D607AE4FFCEC4237D422BA51289FD4C" box="[611,728,692,717]" pageId="3" pageNumber="164" type="subadult">2 subadult</specimenCount>
<specimenCount id="9D607AE4FFCEC4237DFE2BA412A7FD4D" box="[735,758,693,716]" pageId="3" pageNumber="164" type="male"></specimenCount>
s, largest CW
<quantity id="4C9E1C88FFCEC4237CB42BA513AAFD4D" box="[917,1019,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="3.27" pageId="3" pageNumber="164" unit="mm" value="32.7">32.7 mm</quantity>
, CL
<quantity id="4C9E1C88FFCEC4237B122BA514CBFD4D" box="[1075,1178,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="3" pageNumber="164" unit="mm" value="23.8">23.8 mm</quantity>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237B872BA510BAFC95" ID-GBIF-Occurrence="1438485836" collectingDate="2015-08-28" collectionCode="DNA" country="Switzerland" location="A" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237B872BA5149CFD4D" box="[1190,1229,692,716]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
10.0 mm, FW
<quantity id="4C9E1C88FFCEC4237A582BA51094FD6E" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="3" pageNumber="164" unit="mm" value="8.6">8.6 mm</quantity>
,
<date id="FFD897ADFFCEC4237FF12BC91139FD71" box="[208,360,727,752]" pageId="3" pageNumber="164" value="2015-08-28">
<collectingDate id="EF9C6E45FFCEC4237FF12BC91139FD71" box="[208,360,727,752]" pageId="3" pageNumber="164" value="2015-08-28">28 Aug. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237EB22BC911FBFD71" box="[403,426,728,752]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-08;
<collectionCode id="ED7729A8FFCEC4237DDA2BC91369FD71" box="[763,824,728,752]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="164">DNA</collectionCode>
extraction code: T262-9 and T262-10).
<taxonomicName id="4C66CAEEFFCEC4237BD62BC610BAFC95" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFCEC4237BD62BC610BAFC95" bold="true" italics="true" pageId="3" pageNumber="164">Sudanonautes floweri</emphasis>
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="3B0EBB30FFCEC4237FD62BED125CFCB6" ID-GBIF-Occurrence="1438485821" country="Cameroon" county="Yabassi" elevation="255" latitude="4.4724336" location="Princes" longLatPrecision="1" longitude="9.957216" municipality="Parc" pageId="3" pageNumber="164" specimenCount="1" specimenCount-male="1" stateProvince="Littoral Region">
<collectingCountry id="F371F1FDFFCEC4237FD62BED11CAFC95" box="[247,411,764,788]" name="Cameroon" pageId="3" pageNumber="164">CAMEROON</collectingCountry>
.
<collectingCounty id="62B8C9E1FFCEC4237E872BED11AEFC94" box="[422,511,764,789]" pageId="3" pageNumber="164">Yabassi</collectingCounty>
,
<collectingMunicipality id="6BBD2B17FFCEC4237D2A2BEC126CFC95" box="[523,573,765,788]" pageId="3" pageNumber="164">Parc</collectingMunicipality>
des
<location id="8EB9E7B6FFCEC4237D522BED1299FC94" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69:8EB9E7B6FFCEC4237D522BED1299FC94" box="[627,712,764,789]" country="Cameroon" county="Yabassi" latitude="4.4724336" longLatPrecision="1" longitude="9.957216" municipality="Parc" name="Princes" pageId="3" pageNumber="164" stateProvince="Littoral Region">Princes</location>
,
<collectingRegion id="49A27F8FFFCEC4237DF52BED13D0FC94" box="[724,897,764,789]" country="Cameroon" name="Littoral" pageId="3" pageNumber="164">
<collectingRegion id="49A27F8FFFCEC4237DF52BED1379FC94" box="[724,808,764,789]" country="Equatorial Guinea" name="Litoral" pageId="3" pageNumber="164">Littoral</collectingRegion>
Region
</collectingRegion>
(
<geoCoordinate id="EE52D7AAFFCEC4237CB02BED1479FC94" box="[913,1064,764,789]" direction="north" orientation="latitude" pageId="3" pageNumber="164" precision="1" value="4.4724336">04°28.346''N</geoCoordinate>
,
<geoCoordinate id="EE52D7AAFFCEC4237B152BED1480FC94" box="[1076,1233,764,789]" direction="east" orientation="longitude" pageId="3" pageNumber="164" precision="1" value="9.957216">009°57.433'E</geoCoordinate>
),
<quantity id="4C9E1C88FFCEC4237BC42BED157FFC95" box="[1253,1326,764,789]" metricMagnitude="2" metricUnit="m" metricValue="2.55" pageId="3" pageNumber="164" unit="m" value="255.0">
<elevation id="004B565EFFCEC4237BC42BED157FFC95" box="[1253,1326,764,789]" metricMagnitude="2" metricUnit="m" metricValue="2.55" pageId="3" pageNumber="164" unit="m" value="255.0">255 m</elevation>
</quantity>
asl, adult
<specimenCount id="9D607AE4FFCEC4237FB62A3110FFFCB6" box="[151,174,800,823]" pageId="3" pageNumber="164" type="male"></specimenCount>
, CW
<quantity id="4C9E1C88FFCEC4237FCF2A0E1132FCB6" box="[238,355,799,824]" metricMagnitude="-2" metricUnit="m" metricValue="3.538" pageId="3" pageNumber="164" unit="mm" value="35.38">35.38 mm</quantity>
, CL
<quantity id="4C9E1C88FFCEC4237EB82A31125CFCB6" box="[409,525,800,824]" metricMagnitude="-2" metricUnit="m" metricValue="2.493" pageId="3" pageNumber="164" unit="mm" value="24.93">24.93 mm</quantity>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237D382A3111E5FCDC" ID-GBIF-Occurrence="1438485842" collectingDate="2015-08-30" country="Switzerland" county="Yabassi" location="Princes" municipality="Parc" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237D382A311211FCB9" box="[537,576,800,824]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237D692A3112EDFCB6" box="[584,700,800,824]" metricMagnitude="-2" metricUnit="m" metricValue="1.439" pageId="3" pageNumber="164" unit="mm" value="14.39">14.39 mm</quantity>
, FW
<quantity id="4C9E1C88FFCEC4237DD82A31130EFCB6" box="[761,863,800,824]" metricMagnitude="-3" metricUnit="m" metricValue="7.73" pageId="3" pageNumber="164" unit="mm" value="7.73">7.73 mm</quantity>
,
<date id="FFD897ADFFCEC4237C4A2A311455FCB9" box="[875,1028,799,824]" pageId="3" pageNumber="164" value="2015-08-30">
<collectingDate id="EF9C6E45FFCEC4237C4A2A311455FCB9" box="[875,1028,799,824]" pageId="3" pageNumber="164" value="2015-08-30">30 Aug. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237B112A311416FCB9" box="[1072,1095,800,824]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-09);
<collectingCounty id="62B8C9E1FFCEC4237FB62A5510A1FCDC" box="[151,240,836,861]" pageId="3" pageNumber="164">Yabassi</collectingCounty>
,
<collectingMunicipality id="6BBD2B17FFCEC4237FDB2A54117DFCDD" box="[250,300,837,860]" pageId="3" pageNumber="164">Parc</collectingMunicipality>
des
<location id="8EB9E7B6FFCEC4237E7F2A5511E5FCDC" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69:8EB9E7B6FFCEC4237E7F2A5511E5FCDC" box="[350,436,836,861]" country="Switzerland" county="Yabassi" municipality="Parc" name="Princes" pageId="3" pageNumber="164" stateProvince="A">Princes</location>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237E9F2A551094FCFE" ID-GBIF-Occurrence="1438485837" country="Switzerland" elevation="255" latitude="4.4724336" location="Littoral Region" longLatPrecision="1" longitude="9.957216" pageId="3" pageNumber="164" specimenCount="1" specimenCount-female="1" stateProvince="Littoral Region">
<collectingRegion id="49A27F8FFFCEC4237E9F2A551238FCDC" box="[446,617,836,861]" country="Cameroon" name="Littoral" pageId="3" pageNumber="164">
<collectingRegion id="49A27F8FFFCEC4237E9F2A551243FCDC" box="[446,530,836,861]" country="Equatorial Guinea" name="Litoral" pageId="3" pageNumber="164">Littoral</collectingRegion>
Region
</collectingRegion>
(
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;
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),
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asl, adult
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, CW
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, CL
<quantity id="4C9E1C88FFCEC4237A4A2A551094FCFE" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="3" pageNumber="164" unit="mm" value="26.3">26.3 mm</quantity>
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,
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<collectingCountry id="F371F1FDFFCEC4237FF02A7910A9FC01" box="[209,248,872,896]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
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, FW
<quantity id="4C9E1C88FFCEC4237E842A7911AFFCFE" box="[421,510,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="3" pageNumber="164" unit="mm" value="9.1">9.1 mm</quantity>
,
<date id="FFD897ADFFCEC4237D2A2A7912F7FC01" box="[523,678,871,896]" pageId="3" pageNumber="164" value="2015-08-30">
<collectingDate id="EF9C6E45FFCEC4237D2A2A7912F7FC01" box="[523,678,871,896]" pageId="3" pageNumber="164" value="2015-08-30">30 Aug. 2015</collectingDate>
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(P.
<collectingRegion id="49A27F8FFFCEC4237DF32A7912B8FC01" box="[722,745,872,896]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-10;
<collectionCode id="ED7729A8FFCEC4237B612A79142CFC01" box="[1088,1149,872,896]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="164">DNA</collectionCode>
extraction code T262-3);
<taxonomicName id="4C66CAEEFFCEC4237FB62A9D109FFC24" box="[151,206,908,933]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
,
<location id="8EB9E7B6FFCEC4237FFB2A9D1120FC24" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69:8EB9E7B6FFCEC4237FFB2A9D1120FC24" box="[218,369,908,933]" country="Switzerland" name="Tamba forest" pageId="3" pageNumber="164" stateProvince="A">Tamba forest</location>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237E5C2A9D1094FC46" ID-GBIF-Occurrence="1438485840" country="Switzerland" elevation="100" latitude="4.1805553" location="Southwest Region" pageId="3" pageNumber="164" specimenCount="1" specimenCount-female="1" stateProvince="Southwest Region">
<collectingRegion id="49A27F8FFFCEC4237E5C2A9D121BFC24" box="[381,586,908,933]" country="Cameroon" name="South-West" pageId="3" pageNumber="164">Southwest Region</collectingRegion>
(
<geoCoordinate id="EE52D7AAFFCEC4237D7A2A9D12A0FC24" box="[603,753,908,933]" direction="north" orientation="latitude" pageId="3" pageNumber="164" precision="15" value="4.1805553">04° 10' 50''N</geoCoordinate>
, 009° 23' 40'E),
<quantity id="4C9E1C88FFCEC4237C852A9D13BFFC25" box="[932,1006,908,932]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="164" unit="m" value="100.0">
<elevation id="004B565EFFCEC4237C852A9D13BFFC25" box="[932,1006,908,932]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="164" unit="m" value="100.0">100 m</elevation>
</quantity>
asl, adult
<specimenCount id="9D607AE4FFCEC4237B452A9C1428FC25" box="[1124,1145,909,932]" pageId="3" pageNumber="164" type="female"></specimenCount>
, CW
<quantity id="4C9E1C88FFCEC4237B9F2A9D1562FC25" box="[1214,1331,908,933]" metricMagnitude="-2" metricUnit="m" metricValue="5.388" pageId="3" pageNumber="164" unit="mm" value="53.88">53.88 mm</quantity>
, CL 38.0 mm
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237FF42AA11548FC49" ID-GBIF-Occurrence="1438485846" box="[213,1305,943,968]" collectingDate="2015-09-03" country="Switzerland" location="Tamba forest" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237FF42AA110ADFC49" box="[213,252,944,968]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237E272AA11121FC46" box="[262,368,944,968]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="3" pageNumber="164" unit="mm" value="20.4">20.4 mm</quantity>
, FW 11.7,
<date id="FFD897ADFFCEC4237ED62AA112D5FC49" box="[503,644,943,968]" pageId="3" pageNumber="164" value="2015-09-03">
<collectingDate id="EF9C6E45FFCEC4237ED62AA112D5FC49" box="[503,644,943,968]" pageId="3" pageNumber="164" value="2015-09-03">3 Sep. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237D962AA1129FFC49" box="[695,718,944,968]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-11);
<taxonomicName id="4C66CAEEFFCEC4237B182ABE1421FC49" box="[1081,1136,943,968]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
,
<location id="8EB9E7B6FFCEC4237B5E2ABE1548FC49" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69:8EB9E7B6FFCEC4237B5E2ABE1548FC49" box="[1151,1305,943,968]" country="Switzerland" name="Tamba forest" pageId="3" pageNumber="164" stateProvince="A">Tamba forest</location>
</materialsCitation>
,
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<collectingRegion id="49A27F8FFFCEC4237A092ABE10B9FC6C" country="Cameroon" name="South-West" pageId="3" pageNumber="164">Southwest Region</collectingRegion>
(
<geoCoordinate id="EE52D7AAFFCEC4237FD62AC511DBFC6C" box="[247,394,980,1005]" direction="north" orientation="latitude" pageId="3" pageNumber="164" precision="15" value="4.1805553">04° 10' 50''N</geoCoordinate>
, 009° 23' 40'E),
<elevation id="004B565EFFCEC4237D632AC512D9FC6D" box="[578,648,980,1004]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="164" unit="m" value="100.0">
<quantity id="4C9E1C88FFCEC4237D632AC512D9FC6D" box="[578,648,980,1004]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="164" unit="m" value="100.0">100 m</quantity>
</elevation>
asl, adult
<specimenCount id="9D607AE4FFCEC4237DDB2AC41340FC6D" box="[762,785,981,1004]" pageId="3" pageNumber="164" type="male"></specimenCount>
, CW
<quantity id="4C9E1C88FFCEC4237C712AC513E7FC6D" box="[848,950,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="3.6399999999999997" pageId="3" pageNumber="164" unit="mm" value="36.4">36.4 mm</quantity>
, CL
<quantity id="4C9E1C88FFCEC4237CCA2AC51400FC6D" box="[1003,1105,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" pageId="3" pageNumber="164" unit="mm" value="25.2">25.2 mm</quantity>
</materialsCitation>
,
<materialsCitation id="3B0EBB30FFCEC4237B7D2AC513A4FB91" ID-GBIF-Occurrence="1438485843" collectingDate="2015-09-03" collectionCode="DNA" country="Switzerland" location="A" pageId="3" pageNumber="164" specimenCount="1" stateProvince="A">
<collectingCountry id="F371F1FDFFCEC4237B7D2AC514D2FC6D" box="[1116,1155,980,1004]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
<quantity id="4C9E1C88FFCEC4237BAB2AC514A1FC6D" box="[1162,1264,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="3" pageNumber="164" unit="mm" value="12.9">12.9 mm</quantity>
, FW
<quantity id="4C9E1C88FFCEC4237A0D2AC515D2FC6D" box="[1324,1411,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="3" pageNumber="164" unit="mm" value="8.9">8.9 mm</quantity>
,
<date id="FFD897ADFFCEC4237AAF2AC51156FB91" pageId="3" pageNumber="164" value="2015-09-03">
<collectingDate id="EF9C6E45FFCEC4237AAF2AC51156FB91" pageId="3" pageNumber="164" value="2015-09-03">3 Sep. 2015</collectingDate>
</date>
(P.
<collectingRegion id="49A27F8FFFCEC4237E132AE91118FB91" box="[306,329,1016,1040]" country="Nigeria" name="Rivers" pageId="3" pageNumber="164">A</collectingRegion>
. Mvogo Ndongo) (LZUY-12;
<collectionCode id="ED7729A8FFCEC4237DBA2AE91289FB91" box="[667,728,1016,1040]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="164">DNA</collectionCode>
extraction code T262-5).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37CE2E6FFCEC4237FE62D0D11BFFA3B" pageId="3" pageNumber="164" type="diagnosis">
<paragraph id="8BD9B16DFFCEC4237FE62D0D11BFFA3B" blockId="3.[151,1437,151,1999]" pageId="3" pageNumber="164">
<emphasis id="B9126D7FFFCEC4237FE62D0D116BFBB4" bold="true" box="[199,314,1052,1077]" pageId="3" pageNumber="164">Diagnosis</emphasis>
. Carapace subovoid; postfrontal crest distinct, completely crossing carapace, meeting epibranchial teeth; exorbital tooth low, blunt; exorbital tooth reduced to granule; intermediate tooth between exorbital, epibranchial teeth present, reduced to granule (
<figureCitation id="135DADE8FFCEC4237DE52D721358FBFD" box="[708,777,1123,1148]" captionStart="FIGURE 1" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[235,1349,203,1854]" captionTargetId="figure@4.[228,1359,195,1858]" captionTargetPageId="4" captionText="FIGURE 1. Sudanonautes tiko n. sp. from Cameroon. Habitus in dorsal view: a, holotype, adult male (CW 36.2 mm) ZMB Crust. 29628, Tamba Forest, Tiko (colour in ethanol); b, paratype, adult male (CW 35.4 mm) ZMB Crust. 29629 from Edea (colour in life). Scale bar equals 16 mm." httpUri="https://zenodo.org/record/376069/files/figure.png" pageId="3" pageNumber="164">Fig. 1</figureCitation>
a,b, 2a,b). Vertical sulcus on carapace sidewall meeting anterolateral margin at epibranchial tooth (
<figureCitation id="135DADE8FFCEC4237D5F2D971295FB1E" box="[638,708,1158,1183]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
a,b). Third maxilliped ischium with distinct vertical sulcus (
<figureCitation id="135DADE8FFCEC4237A4E2D9710F6FB43" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
d). S3/s4 reduced to 2 short notches on lateral margins; margins of s1s4 distinctly thickened, raised; anterior margin of sternoabdominal cavity low, not raised (
<figureCitation id="135DADE8FFCEC4237DF52DDC1348FB67" box="[724,793,1229,1254]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
c). Fingers of male major cheliped slim, elongated, fixed finger (pollex) with 2 large teeth proximally, movable finger (dactylus) arched, with single large tooth one-third distance from base, enclosing oval interspace when closed (
<figureCitation id="135DADE8FFCEC4237C132C051324FAAC" box="[818,885,1300,1325]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
f); cheliped carpus with 2 acute teeth, distal tooth larger than proximal (
<figureCitation id="135DADE8FFCEC4237EAB2C261198FAD1" box="[394,457,1335,1360]" captionStart="FIGURE 1" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[235,1349,203,1854]" captionTargetId="figure@4.[228,1359,195,1858]" captionTargetPageId="4" captionText="FIGURE 1. Sudanonautes tiko n. sp. from Cameroon. Habitus in dorsal view: a, holotype, adult male (CW 36.2 mm) ZMB Crust. 29628, Tamba Forest, Tiko (colour in ethanol); b, paratype, adult male (CW 35.4 mm) ZMB Crust. 29629 from Edea (colour in life). Scale bar equals 16 mm." httpUri="https://zenodo.org/record/376069/files/figure.png" pageId="3" pageNumber="164">Fig. 1</figureCitation>
a,b). G1 subterminal article slim; G1 terminal article long, one third length of gonopod, proximal G1 terminal article straight basally, distal half curving sharply outward, widened medially, evenly tapering to narrow, pointed tip (
<figureCitation id="135DADE8FFCEC4237D302C6F1209FA16" box="[529,600,1406,1431]" captionStart="FIGURE 3" captionStartId="6.[151,250,1233,1255]" captionTargetBox="[237,1350,197,1207]" captionTargetId="figure@6.[231,1356,193,1212]" captionTargetPageId="6" captionText="FIGURE 3. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, left G 1 ventral view; b: left G 1 dorsal view; c, lleft G 2 ventral view. Scale bar equals 2 mm (a, b, c)." httpUri="https://zenodo.org/record/376071/files/figure.png" pageId="3" pageNumber="164">Fig. 3</figureCitation>
a,b). Terminal article G2 extremely short, only one-fifteenth as long as subterminal segment (
<figureCitation id="135DADE8FFCEC4237EAE2CB31183FA3B" box="[399,466,1442,1467]" captionStart="FIGURE 3" captionStartId="6.[151,250,1233,1255]" captionTargetBox="[237,1350,197,1207]" captionTargetId="figure@6.[231,1356,193,1212]" captionTargetPageId="6" captionText="FIGURE 3. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, left G 1 ventral view; b: left G 1 dorsal view; c, lleft G 2 ventral view. Scale bar equals 2 mm (a, b, c)." httpUri="https://zenodo.org/record/376071/files/figure.png" pageId="3" pageNumber="164">Fig. 3</figureCitation>
c).
</paragraph>
</subSubSection>
<subSubSection id="C37CE2E6FFCEC4267FE62CD412D9F84F" lastPageId="6" lastPageNumber="167" pageId="3" pageNumber="164" type="description">
<paragraph id="8BD9B16DFFCEC4237FE62CD414E1F84E" blockId="3.[151,1437,151,1999]" pageId="3" pageNumber="164">
<emphasis id="B9126D7FFFCEC4237FE62CD41100FA5F" bold="true" box="[199,337,1477,1502]" pageId="3" pageNumber="164">Description</emphasis>
.
<emphasis id="B9126D7FFFCEC4237E412CD41243FA5F" bold="true" box="[352,530,1477,1502]" pageId="3" pageNumber="164">
Male
<typeStatus id="54DD0FCFFFCEC4237E852CD4125CFA5F" box="[420,525,1477,1502]" pageId="3" pageNumber="164" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace subovoid, widest at anterior third (CW/CL 1.46), high (
<collectingCountry id="F371F1FDFFCEC4237BDF2CD71574FA5F" box="[1278,1317,1478,1502]" name="Switzerland" pageId="3" pageNumber="164">CH</collectingCountry>
/CL 0.49), front broad (FW/CW 0. 29), deflexed, anterior margin slightly concave medially; postfrontal crest distinct, complete, consisting of fused epigastric, postorbital crests, divided medially by shallow median sulcus forked posteriorly, each lateral end curving back to meet epibranchial tooth; urogastric, cardiac, cervical grooves all distinct; exorbital tooth short, blunt; anterolateral margin between exorbital, epibranchial teeth curving slightly outward, with small granule-sized intermediate tooth; anterolateral margin smooth posterior to epibranchial tooth (
<figureCitation id="135DADE8FFCEC4237FBE2F8B10BCF932" box="[159,237,1690,1715]" captionStart="FIGURE 1" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[235,1349,203,1854]" captionTargetId="figure@4.[228,1359,195,1858]" captionTargetPageId="4" captionText="FIGURE 1. Sudanonautes tiko n. sp. from Cameroon. Habitus in dorsal view: a, holotype, adult male (CW 36.2 mm) ZMB Crust. 29628, Tamba Forest, Tiko (colour in ethanol); b, paratype, adult male (CW 35.4 mm) ZMB Crust. 29629 from Edea (colour in life). Scale bar equals 16 mm." httpUri="https://zenodo.org/record/376069/files/figure.png" pageId="3" pageNumber="164">Figs. 1</figureCitation>
a,b, 2a,b); suborbital margin smooth; epistomial median tooth large, triangular, edges lined by granules (
<figureCitation id="135DADE8FFCEC4237FBE2FAC10B7F957" box="[159,230,1725,1750]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
c). Carapace sidewall with 2 sutures, 1 longitudinal, 1 vertical, dividing sidewall into 3 parts (
<figureCitation id="135DADE8FFCEC4237A042FAC153DF957" box="[1317,1388,1725,1750]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
a,b); longitudinal suture dividing suborbital, subhepatic regions from pterygostomial region, beginning at respiratory opening, curving backward across side wall; short vertical suture dividing suborbital region from subhepatic region; additional suture beginning at base of minute epibranchial tooth, curving forward under intermediate tooth, then curving sharply down meeting longitudinal suture, marked by row of granules. S2/s3, completely crossing sternum; s3/s4 incomplete, reduced to one small notch at each side (
<figureCitation id="135DADE8FFCEC4237CAF2E7E1383F809" box="[910,978,1903,1928]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
c). Third maxillipeds filling entire buccal cavern except for transversely oval efferent respiratory openings visible at superior lateral corners (
<figureCitation id="135DADE8FFCEC4237A162E8315D1F82A" box="[1335,1408,1938,1963]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
c); exopod with well-developed slender flagellum; ischium with distinct vertical sulcus (
<figureCitation id="135DADE8FFCEC4237B702EA714C4F84E" box="[1105,1173,1974,1999]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
c).
</paragraph>
<caption id="DF19E1E5FFC9C4247FB62E4B1261F82F" httpUri="https://zenodo.org/record/376069/files/figure.png" pageId="4" pageNumber="165" targetBox="[235,1349,203,1854]" targetPageId="4">
<paragraph id="8BD9B16DFFC9C4247FB62E4B1261F82F" blockId="4.[151,1436,1882,1966]" pageId="4" pageNumber="165">
<emphasis id="B9126D7FFFC9C4247FB62E4B115FF8EE" bold="true" box="[151,270,1882,1904]" pageId="4" pageNumber="165">FIGURE 1</emphasis>
.
<taxonomicName id="4C66CAEEFFC9C4247E3D2E4B1189F8F1" box="[284,472,1882,1904]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="165" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC9C4247E3D2E4B1189F8F1" box="[284,472,1882,1904]" italics="true" pageId="4" pageNumber="165">Sudanonautes tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFC9C4247EC12E4A124BF8F1" box="[480,538,1883,1904]" pageId="4" pageNumber="165" rank="species">
<emphasis id="B9126D7FFFC9C4247EC12E4A1245F8F1" bold="true" box="[480,532,1883,1904]" pageId="4" pageNumber="165">n. sp</emphasis>
.
</taxonomicNameLabel>
from Cameroon. Habitus in dorsal view: a, holotype, adult male (CW 36.2 mm) ZMB Crust. 29628, Tamba Forest,
<taxonomicName id="4C66CAEEFFC9C4247EE52E6811A2F80E" box="[452,499,1913,1935]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
(colour in ethanol); b, paratype, adult male (CW 35.4 mm) ZMB Crust. 29629 from Edea (colour in life). Scale bar equals 16 mm.
</paragraph>
</caption>
<caption id="DF19E1E5FFC8C4257FB62E6A123DF86D" httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="5" pageNumber="166" targetBox="[241,1315,203,1890]" targetPageId="5">
<paragraph id="8BD9B16DFFC8C4257FB62E6A123DF86D" blockId="5.[151,1436,1915,2028]" pageId="5" pageNumber="166">
<emphasis id="B9126D7FFFC8C4257FB62E6A1142F811" bold="true" box="[151,275,1915,1937]" pageId="5" pageNumber="166">FIGURE 2.</emphasis>
<taxonomicName id="4C66CAEEFFC8C4257E3A2E6A1186F810" box="[283,471,1915,1937]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC8C4257E3A2E6A1186F810" box="[283,471,1915,1937]" italics="true" pageId="5" pageNumber="166">Sudanonautes tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFC8C4257EFF2E6A1248F810" box="[478,537,1915,1937]" pageId="5" pageNumber="166" rank="species">
<emphasis id="B9126D7FFFC8C4257EFF2E6A1242F811" bold="true" box="[478,531,1915,1936]" pageId="5" pageNumber="166">n. sp</emphasis>
.
</taxonomicNameLabel>
holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus,dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f).
</paragraph>
</caption>
<caption id="DF19E1E5FFCBC4267FB62DC0131FFA87" httpUri="https://zenodo.org/record/376071/files/figure.png" pageId="6" pageNumber="167" targetBox="[237,1350,197,1207]" targetPageId="6">
<paragraph id="8BD9B16DFFCBC4267FB62DC0131FFA87" blockId="6.[151,1435,1233,1286]" pageId="6" pageNumber="167">
<emphasis id="B9126D7FFFCBC4267FB62DC01143FB67" bold="true" box="[151,274,1233,1255]" pageId="6" pageNumber="167">FIGURE 3.</emphasis>
<taxonomicName id="4C66CAEEFFCBC4267E392DC01183FB66" box="[280,466,1233,1255]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFCBC4267E392DC01183FB66" box="[280,466,1233,1255]" italics="true" pageId="6" pageNumber="167">Sudanonautes tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFCBC4267EF62DC31241FB66" box="[471,528,1234,1255]" pageId="6" pageNumber="167" rank="species">
<emphasis id="B9126D7FFFCBC4267EF62DC3125BFB66" bold="true" box="[471,522,1234,1255]" pageId="6" pageNumber="167">n. sp</emphasis>
.
</taxonomicNameLabel>
holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, left G1 ventral view; b: left G1 dorsal view; c, lleft G2 ventral view. Scale bar equals 2 mm (a, b, c).
</paragraph>
</caption>
<paragraph id="8BD9B16DFFCBC4267FE62C3C13E6F9C3" blockId="6.[151,1437,1325,2034]" pageId="6" pageNumber="167">
Chelipeds of adult male unequal, right (major) cheliped (propodal length
<quantity id="4C9E1C88FFCBC4267B292C3C143EFAC7" box="[1032,1135,1325,1350]" metricMagnitude="-2" metricUnit="m" metricValue="5.33" pageId="6" pageNumber="167" unit="mm" value="53.3">53.3 mm</quantity>
) 1.3 times longer than left cheliped (propodal length
<quantity id="4C9E1C88FFCBC4267EE32C43127AFAEB" box="[450,555,1362,1386]" metricMagnitude="-2" metricUnit="m" metricValue="4.279999999999999" pageId="6" pageNumber="167" unit="mm" value="42.8">42.8 mm</quantity>
). Fingers slim, elongated; movable finger (dactylus) arched, with single large tooth one-third distance from base, enclosing oval interspace when closed; fixed finger with 2 large teeth proximally. Minor cheliped with occluding margins of fingers lined by row of small teeth (
<figureCitation id="135DADE8FFCBC4267BAA2C88149BFA33" box="[1163,1226,1433,1458]" captionStart="FIGURE 1" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[235,1349,203,1854]" captionTargetId="figure@4.[228,1359,195,1858]" captionTargetPageId="4" captionText="FIGURE 1. Sudanonautes tiko n. sp. from Cameroon. Habitus in dorsal view: a, holotype, adult male (CW 36.2 mm) ZMB Crust. 29628, Tamba Forest, Tiko (colour in ethanol); b, paratype, adult male (CW 35.4 mm) ZMB Crust. 29629 from Edea (colour in life). Scale bar equals 16 mm." httpUri="https://zenodo.org/record/376069/files/figure.png" pageId="6" pageNumber="167">Fig. 1</figureCitation>
a,b); carpus with 2 sharp teeth, distal tooth larger than proximal; inferior margins of merus bearing series of small sharp teeth and a larger distal edge tooth (
<figureCitation id="135DADE8FFCBC4267E972CF011AFFA7B" box="[438,510,1505,1530]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="6" pageNumber="167">Fig. 2</figureCitation>
e), distal tooth sharp, superior surface of merus smooth (
<figureCitation id="135DADE8FFCBC4267BB92CF014B1FA7B" box="[1176,1248,1505,1530]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="6" pageNumber="167">Fig. 2</figureCitation>
e). Walking legs (pereiopods 25) moderately slender, posterior margins of propodi serrated with small blunt teeth, dactyli tapering, each bearing rows of downward-pointing large sharp spines (
<figureCitation id="135DADE8FFCBC4267C622F3813D3F9C3" box="[835,898,1577,1602]" captionStart="FIGURE 1" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[235,1349,203,1854]" captionTargetId="figure@4.[228,1359,195,1858]" captionTargetPageId="4" captionText="FIGURE 1. Sudanonautes tiko n. sp. from Cameroon. Habitus in dorsal view: a, holotype, adult male (CW 36.2 mm) ZMB Crust. 29628, Tamba Forest, Tiko (colour in ethanol); b, paratype, adult male (CW 35.4 mm) ZMB Crust. 29629 from Edea (colour in life). Scale bar equals 16 mm." httpUri="https://zenodo.org/record/376069/files/figure.png" pageId="6" pageNumber="167">Fig. 1</figureCitation>
a,b).
</paragraph>
<paragraph id="8BD9B16DFFCBC4267FE62F5C111AF8BF" blockId="6.[151,1437,1325,2034]" pageId="6" pageNumber="167">
Male pleon (abdomen) broadly triangular with straight edges (
<figureCitation id="135DADE8FFCBC4267CB32F5C1388F9E7" box="[914,985,1613,1638]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[241,1315,203,1890]" captionTargetId="figure@5.[233,1328,193,1899]" captionTargetPageId="5" captionText="FIGURE 2. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, frontal view of carapace; b, lateral view of carapace sidewall; c, thoracic sternite; d, left third maxilliped frontal view; e, superior view of the merus of the left cheliped showing distal tooth; f, major cheliped showing propodus, dactylus, and carpus. Scale bar equals 10 mm (a), 5 mm (b), 6 mm (c), 1 mm (d), 5 mm (e), 13 mm (f)." httpUri="https://zenodo.org/record/376070/files/figure.png" pageId="6" pageNumber="167">Fig. 2</figureCitation>
c). G1 terminal article long (about onethird length of total length), proximal half straight, distal half curving sharply outward, widened medially, with short setae, evenly tapering to narrow, pointed tip (
<figureCitation id="135DADE8FFCBC4267DF52F841347F92F" box="[724,790,1685,1710]" captionStart="FIGURE 3" captionStartId="6.[151,250,1233,1255]" captionTargetBox="[237,1350,197,1207]" captionTargetId="figure@6.[231,1356,193,1212]" captionTargetPageId="6" captionText="FIGURE 3. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, left G 1 ventral view; b: left G 1 dorsal view; c, lleft G 2 ventral view. Scale bar equals 2 mm (a, b, c)." httpUri="https://zenodo.org/record/376071/files/figure.png" pageId="6" pageNumber="167">Fig. 3</figureCitation>
a,b); G2 shorter than G1 reaching only to G1 subterminal segment/terminal article junction; terminal article extremely short, only one-fifteenth as long as subterminal segment; G2 subterminal segment widest at base, then tapering sharply inward, forming long, thin, pointed, upright process supporting short terminal segment; rounded collar at junction between terminal article and subterminal article (
<figureCitation id="135DADE8FFCBC4267FCD2E34117EF8BF" box="[236,303,1829,1854]" captionStart="FIGURE 3" captionStartId="6.[151,250,1233,1255]" captionTargetBox="[237,1350,197,1207]" captionTargetId="figure@6.[231,1356,193,1212]" captionTargetPageId="6" captionText="FIGURE 3. Sudanonautes tiko n. sp. holotype adult male (CW 36.2 mm) ZMB Crust. 29628. a, left G 1 ventral view; b: left G 1 dorsal view; c, lleft G 2 ventral view. Scale bar equals 2 mm (a, b, c)." httpUri="https://zenodo.org/record/376071/files/figure.png" pageId="6" pageNumber="167">Fig. 3</figureCitation>
c).
</paragraph>
<paragraph id="8BD9B16DFFCBC4267FE62E58134FF8E0" blockId="6.[151,1437,1325,2034]" box="[199,798,1865,1890]" pageId="6" pageNumber="167">
<emphasis id="B9126D7FFFCBC4267FE62E5810AFF8E3" bold="true" box="[199,254,1865,1890]" pageId="6" pageNumber="167">Size.</emphasis>
<collectingRegion id="49A27F8FFFCBC4267E242E5B1148F8E3" box="[261,281,1866,1890]" country="Nigeria" name="Rivers" pageId="6" pageNumber="167">A</collectingRegion>
medium-sized species, adult at CW
<quantity id="4C9E1C88FFCBC4267D922E58134BF8E0" box="[691,794,1865,1890]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="6" pageNumber="167" unit="mm" value="36.2">36.2 mm</quantity>
.
</paragraph>
<paragraph id="8BD9B16DFFCBC4267FE62E7C12D9F84F" blockId="6.[151,1437,1325,2034]" pageId="6" pageNumber="167">
<emphasis id="B9126D7FFFCBC4267FE62E7C1108F807" bold="true" box="[199,345,1901,1926]" pageId="6" pageNumber="167">Color in life</emphasis>
.
<typeStatus id="54DD0FCFFFCBC4267E492E7C1181F807" box="[360,464,1901,1926]" pageId="6" pageNumber="167" type="holotype">Holotype</typeStatus>
from
<taxonomicName id="4C66CAEEFFCBC4267D392E7C121DF807" box="[536,588,1901,1926]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
(
<figureCitation id="135DADE8FFCBC4267D7D2E7C12CCF807" box="[604,669,1901,1926]" captionStart="FIGURE 1" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[235,1349,203,1854]" captionTargetId="figure@4.[228,1359,195,1858]" captionTargetPageId="4" captionText="FIGURE 1. Sudanonautes tiko n. sp. from Cameroon. Habitus in dorsal view: a, holotype, adult male (CW 36.2 mm) ZMB Crust. 29628, Tamba Forest, Tiko (colour in ethanol); b, paratype, adult male (CW 35.4 mm) ZMB Crust. 29629 from Edea (colour in life). Scale bar equals 16 mm." httpUri="https://zenodo.org/record/376069/files/figure.png" pageId="6" pageNumber="167">Fig. 1</figureCitation>
a) dorsal carapace and appendages dark brown, green in specimens from Edea (
<figureCitation id="135DADE8FFCBC4267E3C2E80110FF82B" box="[285,350,1937,1962]" captionStart="FIGURE 1" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[235,1349,203,1854]" captionTargetId="figure@4.[228,1359,195,1858]" captionTargetPageId="4" captionText="FIGURE 1. Sudanonautes tiko n. sp. from Cameroon. Habitus in dorsal view: a, holotype, adult male (CW 36.2 mm) ZMB Crust. 29628, Tamba Forest, Tiko (colour in ethanol); b, paratype, adult male (CW 35.4 mm) ZMB Crust. 29629 from Edea (colour in life). Scale bar equals 16 mm." httpUri="https://zenodo.org/record/376069/files/figure.png" pageId="6" pageNumber="167">Fig. 1</figureCitation>
b); articulations between carpus and merus of cheliped and ambulatory legs pale yellow-brown. Preserved specimens uniformly light brown.
</paragraph>
</subSubSection>
<subSubSection id="C37CE2E6FFCBC4267FE62EC815C0F873" box="[199,1425,2009,2034]" pageId="6" pageNumber="167" type="materials_examined">
<paragraph id="8BD9B16DFFCBC4267FE62EC815C0F873" blockId="6.[151,1437,1325,2034]" box="[199,1425,2009,2034]" pageId="6" pageNumber="167">
<materialsCitation id="3B0EBB30FFCBC4267FE62EC815C0F873" ID-GBIF-Occurrence="1438485845" box="[199,1425,2009,2034]" country="Cameroon" elevation="100" latitude="4.1805553" location="Tamba Forest" pageId="6" pageNumber="167" specimenCount="1" stateProvince="Southwest Region" typeStatus="holotype">
<emphasis id="B9126D7FFFCBC4267FE62EC81134F873" bold="true" box="[199,357,2009,2034]" pageId="6" pageNumber="167">
<typeStatus id="54DD0FCFFFCBC4267FE62EC81153F873" box="[199,258,2009,2034]" pageId="6" pageNumber="167">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F371F1FDFFCBC4267E4D2ECB11B5F873" box="[364,484,2010,2034]" name="Cameroon" pageId="6" pageNumber="167">Cameroon</collectingCountry>
,
<location id="8EB9E7B6FFCBC4267ECE2EC812DBF870" LSID="urn:lsid:plazi:treatment:03CF007BFFCFC42A7FB62F5D1466FB69:8EB9E7B6FFCBC4267ECE2EC812DBF870" box="[495,650,2009,2034]" country="Cameroon" latitude="4.1805553" name="Tamba Forest" pageId="6" pageNumber="167" stateProvince="Southwest Region">Tamba Forest</location>
,
<taxonomicName id="4C66CAEEFFCBC4267DB72EC8129CF873" box="[662,717,2009,2034]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
, in the
<collectingRegion id="49A27F8FFFCBC4267C3C2EC813BBF873" box="[797,1002,2009,2034]" country="Cameroon" name="South-West" pageId="6" pageNumber="167">Southwest Region</collectingRegion>
(
<geoCoordinate id="EE52D7AAFFCBC4267CD82EC814D1F873" box="[1017,1152,2009,2034]" direction="north" orientation="latitude" pageId="6" pageNumber="167" precision="15" value="4.1805553">04°10'50''N</geoCoordinate>
, 09°23'40'E),
<quantity id="4C9E1C88FFCBC4267A3D2ECB1532F870" box="[1308,1379,2010,2034]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="167" unit="m" value="100.0">
<elevation id="004B565EFFCBC4267A3D2ECB1532F870" box="[1308,1379,2010,2034]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="167" unit="m" value="100.0">100 m</elevation>
</quantity>
asl.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37CE2E6FFCAC4277FE6298612AAFE4E" pageId="7" pageNumber="168" type="etymology">
<paragraph id="8BD9B16DFFCAC4277FE629861238FF54" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="168">
<emphasis id="B9126D7FFFCAC4277FE629861117FF31" bold="true" box="[199,326,151,176]" pageId="7" pageNumber="168">Etymology</emphasis>
. The new species is named for
<taxonomicName id="4C66CAEEFFCAC4277D93298612B7FF31" box="[690,742,151,176]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
in southwestern
<collectingCountry id="F371F1FDFFCAC4277C8829891471FF31" box="[937,1056,152,176]" name="Cameroon" pageId="7" pageNumber="168">Cameroon</collectingCountry>
, where it was first collected. The species name
<taxonomicName id="4C66CAEEFFCAC4277E1329AD110DFF55" box="[306,348,188,212]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko">
<emphasis id="B9126D7FFFCAC4277E1329AD110DFF55" box="[306,348,188,212]" italics="true" pageId="7" pageNumber="168">tiko</emphasis>
</taxonomicName>
is a noun in apposition.
</paragraph>
<paragraph id="8BD9B16DFFCAC4277FE629CE12AAFE4E" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="168">
<emphasis id="B9126D7FFFCAC4277FE629CE117BFF79" bold="true" box="[199,298,223,248]" pageId="7" pageNumber="168">Habitat.</emphasis>
<taxonomicName id="4C66CAEEFFCAC4277E1329F11250FF79" box="[306,513,224,248]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFCAC4277E1329F11250FF79" box="[306,513,224,248]" italics="true" pageId="7" pageNumber="168">Sudanonautes tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFCAC4277D2829CE1218FF79" bold="true" box="[521,585,223,248]" pageId="7" pageNumber="168">
<taxonomicNameLabel id="A221D004FFCAC4277D2829CE1218FF79" box="[521,585,223,248]" pageId="7" pageNumber="168" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is restricted to the humid area of the coastal rain forest of Cameroon.
<taxonomicName id="4C66CAEEFFCAC4277A4929CE15CDFF79" box="[1384,1436,223,248]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
(the type locality) and Edea both lie in the coastal zone of Cameroon which is one of the wettest places in Africa, with an annual average rainfall of more than
<quantity id="4C9E1C88FFCAC4277DA8283612ACFEBE" box="[649,765,295,320]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="168" unit="mm" value="5000.0">5,000 mm</quantity>
, which includes Mount Cameroon (
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asl), the highest mountain in West and Central Africa (
<bibRefCitation id="EFF7CC9CFFCAC4277D7C285D12A1FEE5" author="Molua" box="[605,752,332,357]" pageId="7" pageNumber="168" refString="Molua, E. L. (2009) An empirical assessment of the impact of climate change on smallholder agriculture in Cameroon. Global and Planetary Change, 67, 205 - 208." type="journal article" year="2009">Molua 2009</bibRefCitation>
). At Edea,
<taxonomicName id="4C66CAEEFFCAC4277C59285D1392FEE5" box="[888,963,332,356]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFCAC4277C59285D1392FEE5" box="[888,963,332,356]" italics="true" pageId="7" pageNumber="168">S. tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFCAC4277CEF285D1443FEE4" bold="true" box="[974,1042,332,357]" pageId="7" pageNumber="168">
<taxonomicNameLabel id="A221D004FFCAC4277CEF285D1443FEE4" box="[974,1042,332,357]" pageId="7" pageNumber="168" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
was found under small stones in wetlands near the
<taxonomicName id="4C66CAEEFFCAC4277E4A286111EDFE09" box="[363,444,368,392]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="sanaga">Sanaga</taxonomicName>
River (
<figureCitation id="135DADE8FFCAC4277D37287E120DFE09" box="[534,604,367,392]" captionStart="FIGURE 4" captionStartId="7.[151,250,1055,1077]" captionTargetBox="[235,1349,520,1030]" captionTargetId="figure@7.[233,1354,517,1033]" captionTargetPageId="7" captionText="FIGURE 4. Habitat of Sudanonautes tiko n. sp. at Edea, Cameroon." httpUri="https://zenodo.org/record/376072/files/figure.png" pageId="7" pageNumber="168">Fig. 4</figureCitation>
) and not in burrows, while at
<taxonomicName id="4C66CAEEFFCAC4277CE0287E13A4FE09" box="[961,1013,367,392]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
the new species was found in small streams under small rocks. At the
<taxonomicName id="4C66CAEEFFCAC4277D3528851219FE2C" box="[532,584,404,429]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
locality,
<taxonomicName id="4C66CAEEFFCAC4277D8F288512A5FE2D" box="[686,756,404,428]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFCAC4277D8F288512A5FE2D" box="[686,756,404,428]" italics="true" pageId="7" pageNumber="168">S. tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFCAC4277DDA28851368FE2C" bold="true" box="[763,825,404,429]" pageId="7" pageNumber="168">
<taxonomicNameLabel id="A221D004FFCAC4277DDA28851368FE2C" box="[763,825,404,429]" pageId="7" pageNumber="168" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
occurs sympatrically in the streams with other species of freshwater crabs such as
<taxonomicName id="4C66CAEEFFCAC4277EEC28A91202FE51" box="[461,595,440,464]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFCAC4277EEC28A91202FE51" box="[461,595,440,464]" italics="true" pageId="7" pageNumber="168">S. africanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C66CAEEFFCAC4277DA828A912A5FE51" box="[649,756,440,464]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFCAC4277DA828A912A5FE51" box="[649,756,440,464]" italics="true" pageId="7" pageNumber="168">S. floweri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF19E1E5FFCAC4277FB62D0E131FFBB4" box="[151,846,1055,1077]" httpUri="https://zenodo.org/record/376072/files/figure.png" pageId="7" pageNumber="168" targetBox="[235,1349,520,1030]" targetPageId="7">
<paragraph id="8BD9B16DFFCAC4277FB62D0E131FFBB4" blockId="7.[151,846,1055,1077]" box="[151,846,1055,1077]" pageId="7" pageNumber="168">
<emphasis id="B9126D7FFFCAC4277FB62D0E1143FBB5" bold="true" box="[151,274,1055,1077]" pageId="7" pageNumber="168">FIGURE 4.</emphasis>
Habitat of
<taxonomicName id="4C66CAEEFFCAC4277EA42D0E1211FBB4" box="[389,576,1055,1077]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFCAC4277EA42D0E1211FBB4" box="[389,576,1055,1077]" italics="true" pageId="7" pageNumber="168">Sudanonautes tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFCAC4277D672D0E122EFBB5" bold="true" box="[582,639,1055,1076]" pageId="7" pageNumber="168">
<taxonomicNameLabel id="A221D004FFCAC4277D672D0E122EFBB5" box="[582,639,1055,1076]" pageId="7" pageNumber="168" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
at Edea, Cameroon.
</paragraph>
</caption>
<subSubSection id="C37CE2E6FFCAC4277FE62D4D12EBFA5C" pageId="7" pageNumber="168" type="reference_group">
<paragraph id="8BD9B16DFFCAC4277FE62D4D12EBFA5C" blockId="7.[151,1436,1116,2005]" pageId="7" pageNumber="168">
<taxonomicName id="4C66CAEEFFCAC4277FE62D4D11C7FBF5" ID-CoL="7B2Z6" box="[199,406,1116,1140]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFCAC4277FE62D4D11C7FBF5" box="[199,406,1116,1140]" italics="true" pageId="7" pageNumber="168">Sudanonautes tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFCAC4277EBC2D4D118DFBF5" box="[413,476,1116,1141]" pageId="7" pageNumber="168" rank="species">
<emphasis id="B9126D7FFFCAC4277EBC2D4D1187FBF4" bold="true" box="[413,470,1116,1141]" pageId="7" pageNumber="168">n. sp</emphasis>
.
</taxonomicNameLabel>
may be threatened by human impacts on its habitat. The water in the streams at
<taxonomicName id="4C66CAEEFFCAC4277A492D4D15CDFBF4" ID-CoL="7B2Z6" box="[1384,1436,1116,1141]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
was found to be quite acidic (between
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4.06.5) and at a lower
<collectingCountry id="F371F1FDFFCAC4277C882D90139AFB19" box="[937,971,1153,1176]" name="Philippines" pageId="7" pageNumber="168">pH</collectingCountry>
than would normally be expected for a tropical freshwater ecosystem (where the majority of aquatic organisms prefer water with a
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between 6.59.0 (Robertson-Brayan 2004)). The lower
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range of the water at
<taxonomicName id="4C66CAEEFFCAC4277C462DD913CAFB60" ID-CoL="7B2Z6" box="[871,923,1224,1249]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
may be attributed to anthropogenic pollution from human wastes. Changes in key physical and chemical parameters at the landscape scale (
<bibRefCitation id="EFF7CC9CFFCAC4277BEA2DFD15C6FA84" author="Walsh" box="[1227,1431,1260,1285]" pageId="7" pageNumber="168" refString="Walsh, J., Anisimov, O., Hagen, J. O., Jakobsson, T., Oerlemans, T., Prowse, T. D., Romanovsky, V., Savelieva, N. &amp; Serreze, M. (2005) Chapter 6. Crysophere and hydrology. In: Arctic Climate Impact Assessment. Cambridge University Press, Cambridge, pp. 183 - 242." type="book chapter" year="2005">
Walsh
<emphasis id="B9126D7FFFCAC4277A3B2DFC151AFA85" box="[1306,1355,1260,1284]" italics="true" pageId="7" pageNumber="168">et al</emphasis>
. 2005
</bibRefCitation>
;
<bibRefCitation id="EFF7CC9CFFCAC4277FB62C0111C4FAA8" author="Callaghan" box="[151,405,1296,1321]" pageId="7" pageNumber="168" refString="Callaghan, T. V., Bjorn, L. O., Chapin, F. S. III., Chernov, Y., Christensen, T. R., Huntley, B., Ims, R. &amp; Johansson, M. (2005) Chapter 7. Arctic tundra and polar desert ecosystems. In: Arctic Climate Impact Assessment. Cambridge University Press, Cambridge, pp. 244 - 335." type="book chapter" year="2005">
Callaghan
<emphasis id="B9126D7FFFCAC4277E322C001119FAA9" box="[275,328,1296,1320]" italics="true" pageId="7" pageNumber="168">et al</emphasis>
. 2005
</bibRefCitation>
;
<bibRefCitation id="EFF7CC9CFFCAC4277E852C0012D0FAA8" author="Prowse" box="[420,641,1296,1321]" pageId="7" pageNumber="168" refString="Prowse, T. D., Wrona, F. J., Reist, J. D., Hobbie, J. E., Levesque, L. M. J. &amp; Vincent, W. F. (2006) Historical changes in arctic freshwater ecosystems. Ambio, 35, 339 - 346." type="journal article" year="2006">
Prowse
<emphasis id="B9126D7FFFCAC4277D212C001265FAA9" box="[512,564,1296,1320]" italics="true" pageId="7" pageNumber="168">et al</emphasis>
. 2006
</bibRefCitation>
) are likely to affect the species richness and biodiversity freshwater ecosystems and to alter the food webs and the levels of primary and secondary productivity. In contrast, the
<collectingCountry id="F371F1FDFFCAC4277A7A2C24152CFACD" box="[1371,1405,1333,1356]" name="Philippines" pageId="7" pageNumber="168">pH</collectingCountry>
of the stream at Edea was less acidic and was in the expected
<collectingCountry id="F371F1FDFFCAC4277C692C48133BFAF1" box="[840,874,1369,1392]" name="Philippines" pageId="7" pageNumber="168">pH</collectingCountry>
range (6.59.0). The populations of
<taxonomicName id="4C66CAEEFFCAC4277A352C48150DFAF1" ID-CoL="7B2Z6" box="[1300,1372,1368,1392]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko">
<emphasis id="B9126D7FFFCAC4277A352C48150DFAF1" box="[1300,1372,1368,1392]" italics="true" pageId="7" pageNumber="168">S. tiko</emphasis>
</taxonomicName>
from Edea, however, may be threatened by the intensive and destructive agricultural practices that are already impacting the condition of the
<taxonomicName id="4C66CAEEFFCAC4277E5F2CB0119EFA38" box="[382,463,1441,1465]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="sanaga">Sanaga</taxonomicName>
<collectingRegion id="49A27F8FFFCAC4277EF92CB11249FA38" box="[472,536,1440,1465]" country="Nigeria" name="Rivers" pageId="7" pageNumber="168">River</collectingRegion>
, which have the potential to extirpate local populations of the freshwater crabs that depend on the ecosystem for their existence.
</paragraph>
</subSubSection>
<subSubSection id="C37CE2E6FFCAC4277FE62CF91156F830" pageId="7" pageNumber="168" type="distribution">
<paragraph id="8BD9B16DFFCAC4277FE62CF91156F830" blockId="7.[151,1436,1116,2005]" pageId="7" pageNumber="168">
<emphasis id="B9126D7FFFCAC4277FE62CF91131F980" bold="true" box="[199,352,1512,1537]" pageId="7" pageNumber="168">Distribution.</emphasis>
<taxonomicName id="4C66CAEEFFCAC4277E492CF91266F981" box="[360,567,1512,1536]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFCAC4277E492CF91266F981" box="[360,567,1512,1536]" italics="true" pageId="7" pageNumber="168">Sudanonautes tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFCAC4277D1E2CF9122FF981" box="[575,638,1512,1537]" pageId="7" pageNumber="168" rank="species">
<emphasis id="B9126D7FFFCAC4277D1E2CF91226F980" bold="true" box="[575,631,1512,1537]" pageId="7" pageNumber="168">n. sp</emphasis>
.
</taxonomicNameLabel>
, is so far known only from lowland tropical rainforest in the
<collectingRegion id="49A27F8FFFCAC4277A162CF915CDF980" box="[1335,1436,1512,1537]" country="Ghana" name="Northern" pageId="7" pageNumber="168">Northern</collectingRegion>
and Southern gulfs of
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drainage ecoregions where the species is associated with two major rivers close to where they drain into the ocean. The localities where this is known to occur are
<taxonomicName id="4C66CAEEFFCAC4277B0B2F211430F9C8" box="[1066,1121,1584,1609]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
, Tamba Forest and Bwenga Forest in the Mungo
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Basin, close to Mount
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,
<collectingRegion id="49A27F8FFFCAC4277C652F451445F9EC" box="[836,1044,1620,1645]" country="Cameroon" name="South-West" pageId="7" pageNumber="168">Southwest Region</collectingRegion>
, and Edea in the
<collectingRegion id="49A27F8FFFCAC4277BF32F45152FF9EC" box="[1234,1406,1620,1645]" country="Cameroon" name="Littoral" pageId="7" pageNumber="168">
<collectingRegion id="49A27F8FFFCAC4277BF32F451577F9EC" box="[1234,1318,1620,1645]" country="Equatorial Guinea" name="Litoral" pageId="7" pageNumber="168">Littoral</collectingRegion>
Region
</collectingRegion>
of
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in the
<taxonomicName id="4C66CAEEFFCAC4277E7A2F6811FDF910" box="[347,428,1657,1681]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="sanaga">Sanaga</taxonomicName>
<collectingRegion id="49A27F8FFFCAC4277E952F6911A3F910" box="[436,498,1656,1681]" country="Nigeria" name="Rivers" pageId="7" pageNumber="168">River</collectingRegion>
drainage close to the main river. The Mungo and
<taxonomicName id="4C66CAEEFFCAC4277B0A2F68142DF910" box="[1067,1148,1657,1681]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="sanaga">Sanaga</taxonomicName>
rivers both drain into the Atlantic Ocean in the Bay of
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, and the above localities are all in tropical rainforest. Three freshwater ecoregions are represented in southern part of
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where this study was carried out (
<bibRefCitation id="EFF7CC9CFFCAC4277B962FD115C9F958" author="Thieme" box="[1207,1432,1728,1753]" pageId="7" pageNumber="168" refString="Thieme, M. L., Abell, R., Stiassny, M. L. J., Skelton, P., Lehner, B., Teugels, G. G., Dinerstein, E., Kamdem Toham, A., Burgess, N. &amp; Olson, D. (2005) Freshwater ecoregions of Africa and Madagascar: a conservation assessment. Island Press, Washington DC, pp. 1 - 483." type="book chapter" year="2005">
Thieme
<emphasis id="B9126D7FFFCAC4277A392FD01502F959" box="[1304,1363,1728,1752]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2005
</bibRefCitation>
;
<bibRefCitation id="EFF7CC9CFFCAC4277FB62FF51104F97C" author="Abell" box="[151,341,1764,1789]" pageId="7" pageNumber="168" refString="Abell, R., Thieme, M. L., Revenga, C., Bryer, M., Kottelat, M., Bogutskaya, N., Coad, B., Mandrak, N., Contreras Balderas, S., Bussing, W., Stiassny, M. L. J., Skelton, P., Allen, G. R ,, Unmack, P., Naseka, A., Ng, R., Sindorf, N., Robertson, J., Armijo, E., Higgins, J. V., Heibel, T. J., Wikramanayake, E., Olson, D., Lopez, H. L., Reis, R. E., Lundberg, J. G., Sabaj Perez, M. H. &amp; Petry, P. (2008) Freshwater Ecoregions of the World: A new map of biogeographic units for freshwater biodiversity conservation. BioScience, 58, 403 - 414." type="journal article" year="2008">
Abell
<emphasis id="B9126D7FFFCAC4277FFD2FF41145F97D" box="[220,276,1764,1788]" italics="true" pageId="7" pageNumber="168">et al.</emphasis>
2008
</bibRefCitation>
), and all have endemic species of freshwater crabs. These ecoregions are: (1) the
<collectingRegion id="49A27F8FFFCAC4277BFE2FF51515F97C" box="[1247,1348,1764,1789]" country="Ghana" name="Northern" pageId="7" pageNumber="168">Northern</collectingRegion>
Gulf of
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drainage in
<collectingCountry id="F371F1FDFFCAC4277E502E1811B7F8A0" box="[369,486,1801,1825]" name="Cameroon" pageId="7" pageNumber="168">Cameroon</collectingCountry>
(where
<taxonomicName id="4C66CAEEFFCAC4277D612E1812D4F8A1" box="[576,645,1800,1824]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFCAC4277D612E1812D4F8A1" box="[576,645,1800,1824]" italics="true" pageId="7" pageNumber="168">S. tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFCAC4277DAD2E19129BF8A1" box="[652,714,1800,1825]" pageId="7" pageNumber="168" rank="species">
<emphasis id="B9126D7FFFCAC4277DAD2E191292F8A0" bold="true" box="[652,707,1800,1825]" pageId="7" pageNumber="168">n. sp</emphasis>
.
</taxonomicNameLabel>
is found with
<taxonomicName id="4C66CAEEFFCAC4277C482E181458F8A1" box="[873,1033,1800,1824]" class="Malacostraca" family="Potamonautidae" genus="Louisea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="B9126D7FFFCAC4277C482E181458F8A1" box="[873,1033,1800,1824]" italics="true" pageId="7" pageNumber="168">Louisea balssi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C66CAEEFFCAC4277B362E18151AF8A1" box="[1047,1355,1800,1824]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="mambilorum">
<emphasis id="B9126D7FFFCAC4277B362E18151AF8A1" box="[1047,1355,1800,1824]" italics="true" pageId="7" pageNumber="168">Potamonemus mambilorum</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C66CAEEFFCAC4277AA62E18115CF8C5" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="orthostylis">
<emphasis id="B9126D7FFFCAC4277AA62E18115CF8C5" italics="true" pageId="7" pageNumber="168">S. orthostylis</emphasis>
</taxonomicName>
); (2) the Southern Gulf of
<collectingCountry id="F371F1FDFFCAC4277D762E3D12F9F8C4" box="[599,680,1836,1861]" name="Guinea-Bissau" pageId="7" pageNumber="168">Guinea</collectingCountry>
drainage in southern
<collectingCountry id="F371F1FDFFCAC4277C8C2E3C1473F8C4" box="[941,1058,1837,1861]" name="Cameroon" pageId="7" pageNumber="168">Cameroon</collectingCountry>
(where
<taxonomicName id="4C66CAEEFFCAC4277BA72E3C1480F8C5" box="[1158,1233,1836,1860]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="tiko">
<emphasis id="B9126D7FFFCAC4277BA72E3C1480F8C5" box="[1158,1233,1836,1860]" italics="true" pageId="7" pageNumber="168">S. tiko</emphasis>
</taxonomicName>
is found with
<taxonomicName id="4C66CAEEFFCAC4277AA42E3C10A8F8E9" class="Malacostraca" family="Potamonautidae" genus="Louisea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="edeaensi">
<emphasis id="B9126D7FFFCAC4277AA42E3C10A8F8E9" italics="true" pageId="7" pageNumber="168">L. edeaensi</emphasis>
</taxonomicName>
s); (3) and the
<taxonomicName id="4C66CAEEFFCAC4277EBD2E4111BFF8E8" box="[412,494,1872,1897]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="169" phylum="Arthropoda" rank="species" species="sangha">Sangha</taxonomicName>
ecoregion, which includes the rivers and streams of the lowland forests of southeast
<collectingCountry id="F371F1FDFFCAC4277FB62E64115EF80C" box="[151,271,1909,1933]" name="Cameroon" pageId="7" pageNumber="168">Cameroon</collectingCountry>
,
<collectingCountry id="F371F1FDFFCAC4277E012E64113CF80C" box="[288,365,1909,1933]" name="Democratic Republic of the Congo" pageId="7" pageNumber="168">
<collectingRegion id="49A27F8FFFCAC4277E012E64113CF80C" box="[288,365,1909,1933]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="7" pageNumber="168">Congo</collectingRegion>
</collectingCountry>
, and western
<collectingCountry id="F371F1FDFFCAC4277D3A2E651315F80C" box="[539,836,1908,1933]" name="Central African Republic" pageId="7" pageNumber="168">
<collectingRegion id="49A27F8FFFCAC4277D3A2E65123FF80C" box="[539,622,1908,1933]" country="Togo" name="Centrale" pageId="7" pageNumber="168">Central</collectingRegion>
African Republic
</collectingCountry>
(where
<taxonomicName id="4C66CAEEFFCAC4277C8C2E641470F80D" box="[941,1057,1908,1932]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="sangha">
<emphasis id="B9126D7FFFCAC4277C8C2E641470F80D" box="[941,1057,1908,1932]" italics="true" pageId="7" pageNumber="168">S. sangha</emphasis>
</taxonomicName>
and
<taxonomicName id="4C66CAEEFFCAC4277B452E64153AF80D" box="[1124,1387,1909,1932]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="regnieri">
<emphasis id="B9126D7FFFCAC4277B452E64153AF80D" box="[1124,1387,1909,1932]" italics="true" pageId="7" pageNumber="168">Potamonautes regnieri</emphasis>
</taxonomicName>
are endemic).
</paragraph>
</subSubSection>
<subSubSection id="C37CE2E6FFCAC42A7FE62EAD1533FE2C" lastPageId="10" lastPageNumber="171" pageId="7" pageNumber="168" type="discussion">
<paragraph id="8BD9B16DFFCAC4287FE62EAD11D3FD05" blockId="7.[151,1436,1116,2005]" lastBlockId="8.[151,1437,151,896]" lastPageId="8" lastPageNumber="169" pageId="7" pageNumber="168">
<emphasis id="B9126D7FFFCAC4277FE62EAD1165F854" bold="true" box="[199,308,1980,2005]" pageId="7" pageNumber="168">Remarks</emphasis>
. The new species is assigned to
<taxonomicName id="4C66CAEEFFCAC4277D872EAD1312F855" box="[678,835,1980,2004]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B9126D7FFFCAC4277D872EAD1312F855" box="[678,835,1980,2004]" italics="true" pageId="7" pageNumber="168">Sudanonautes</emphasis>
</taxonomicName>
because it conforms to the genus in every diagnostic aspect (
<bibRefCitation id="EFF7CC9CFFC5C4287FCF29891135FF31" author="Bott" box="[238,356,151,176]" pageId="8" pageNumber="169" refString="Bott, R. (1955) Die Suswasserkrabben von Afrika (Crust., Decap.) und ihre Stammesgeschichte. Annales du Musee du Congo belge (Tervuren, Belgique) C-Zoologie, I, (3, 3), 209 - 352." type="journal article" year="1955">Bott 1955</bibRefCitation>
;
<bibRefCitation id="EFF7CC9CFFC5C4287E522986121AFF31" author="Cumberlidge" box="[371,587,151,176]" pageId="8" pageNumber="169" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
).
<taxonomicName id="4C66CAEEFFC5C4287D4129891361FF31" box="[608,816,152,176]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287D4129891361FF31" box="[608,816,152,176]" italics="true" pageId="8" pageNumber="169">Sudanonautes tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFC5C4287C1B2986132DFF31" bold="true" box="[826,892,151,176]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="A221D004FFC5C4287C1B2986132DFF31" box="[826,892,151,176]" pageId="8" pageNumber="169" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from all other congeneric species by the following characters: the G1 terminal article of
<taxonomicName id="4C66CAEEFFC5C4287C4629AD13FEFF55" box="[871,943,188,212]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko">
<emphasis id="B9126D7FFFC5C4287C4629AD13FEFF55" box="[871,943,188,212]" italics="true" pageId="8" pageNumber="169">S. tiko</emphasis>
</taxonomicName>
is conspicuously long (about one-third the length of the entire gonopod), whereas in all other species of this genus the G1 terminal article is only about onequarter of the length of the entire gonopod. Other distinctions between
<taxonomicName id="4C66CAEEFFC5C4287CFE28151478FE9D" box="[991,1065,260,284]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287CFE28151478FE9D" box="[991,1065,260,284]" italics="true" pageId="8" pageNumber="169">S. tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFC5C4287B1528151426FE9C" bold="true" box="[1076,1143,260,285]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="A221D004FFC5C4287B1528151426FE9C" box="[1076,1143,260,285]" pageId="8" pageNumber="169" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and its congeners are as follows: the new species can be distinguished from
<taxonomicName id="4C66CAEEFFC5C4287DC52839133DFEC1" box="[740,876,296,320]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC5C4287DC52839133DFEC1" box="[740,876,296,320]" italics="true" pageId="8" pageNumber="169">S. africanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C66CAEEFFC5C4287C5A2839145EFEC1" box="[891,1039,296,320]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="chavanesii">
<emphasis id="B9126D7FFFC5C4287C5A2839145EFEC1" box="[891,1039,296,320]" italics="true" pageId="8" pageNumber="169">S. chavanesii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C66CAEEFFC5C4287B01283914E8FEC1" box="[1056,1209,296,320]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="faradjensis">
<emphasis id="B9126D7FFFC5C4287B01283914E8FEC1" box="[1056,1209,296,320]" italics="true" pageId="8" pageNumber="169">S. faradjensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C66CAEEFFC5C4287BDB28391539FEC1" box="[1274,1384,296,320]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="sangha">
<emphasis id="B9126D7FFFC5C4287BDB28391539FEC1" box="[1274,1384,296,320]" italics="true" pageId="8" pageNumber="169">S. sangha</emphasis>
</taxonomicName>
(the four large-bodied species in this genus that are found in
<collectingCountry id="F371F1FDFFC5C4287C3E285D13C5FEE5" box="[799,916,332,356]" name="Cameroon" pageId="8" pageNumber="169">Cameroon</collectingCountry>
and
<collectingRegion id="49A27F8FFFC5C4287CEF285D1471FEE4" box="[974,1056,332,357]" country="Togo" name="Centrale" pageId="8" pageNumber="169">Central</collectingRegion>
Africa) by its medium body size with a pubertal moult around CW
<quantity id="4C9E1C88FFC5C4287D35287E1233FE06" box="[532,610,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="169" unit="mm" value="35.0">35 mm</quantity>
(vs. a pubertal moult at CW
<quantity id="4C9E1C88FFC5C4287C82287E13A3FE06" box="[931,1010,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="169" unit="mm" value="50.0">50 mm</quantity>
or greater in these other four species).
<taxonomicName id="4C66CAEEFFC5C4287FB628851139FE2D" box="[151,360,404,428]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287FB628851139FE2D" box="[151,360,404,428]" italics="true" pageId="8" pageNumber="169">Sudanonautes tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFC5C4287E52288511E4FE2C" bold="true" box="[371,437,404,429]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="A221D004FFC5C4287E52288511E4FE2C" box="[371,437,404,429]" pageId="8" pageNumber="169" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C66CAEEFFC5C4287DD72885130FFE2D" box="[758,862,404,428]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B9126D7FFFC5C4287DD72885130FFE2D" box="[758,862,404,428]" italics="true" pageId="8" pageNumber="169">S. aubryi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C66CAEEFFC5C4287CBD28851465FE2D" box="[924,1076,404,428]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9126D7FFFC5C4287CBD28851465FE2D" box="[924,1076,404,428]" italics="true" pageId="8" pageNumber="169">S. granulatus</emphasis>
</taxonomicName>
on the basis of the size of the intermediate tooth on the anterolateral margin, which is reduced to a small granule in
<taxonomicName id="4C66CAEEFFC5C4287B5928A91490FE51" box="[1144,1217,440,464]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287B5928A91490FE51" box="[1144,1217,440,464]" italics="true" pageId="8" pageNumber="169">S. tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFC5C4287BEB28A6155AFE4E" box="[1226,1291,439,464]" pageId="8" pageNumber="169" rank="species">
<emphasis id="B9126D7FFFC5C4287BEB28A61555FE51" bold="true" box="[1226,1284,439,464]" pageId="8" pageNumber="169">n. sp</emphasis>
.
</taxonomicNameLabel>
(vs. a large, triangular tooth in these two species).
<taxonomicName id="4C66CAEEFFC5C4287D6228CD1340FE75" box="[579,785,476,500]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287D6228CD1340FE75" box="[579,785,476,500]" italics="true" pageId="8" pageNumber="169">Sudanonautes tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFC5C4287C3828CD1309FE74" bold="true" box="[793,856,476,501]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="A221D004FFC5C4287C3828CD1309FE74" box="[793,856,476,501]" pageId="8" pageNumber="169" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C66CAEEFFC5C4287BA828CD14A8FE75" box="[1161,1273,476,500]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="monodi">
<emphasis id="B9126D7FFFC5C4287BA828CD14A8FE75" box="[1161,1273,476,500]" italics="true" pageId="8" pageNumber="169">S. monodi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C66CAEEFFC5C4287A1128CD15CAFE75" box="[1328,1435,476,500]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC5C4287A1128CD15CAFE75" box="[1328,1435,476,500]" italics="true" pageId="8" pageNumber="169">S. floweri</emphasis>
</taxonomicName>
by the width of the subterminal segment of G1, which is slim in
<taxonomicName id="4C66CAEEFFC5C4287C552B1113EAFD99" box="[884,955,512,536]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287C552B1113EAFD99" box="[884,955,512,536]" italics="true" pageId="8" pageNumber="169">S. tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFC5C4287CE228EE1452FD96" box="[963,1027,511,536]" pageId="8" pageNumber="169" rank="species">
<emphasis id="B9126D7FFFC5C4287CE228EE13ADFD99" bold="true" box="[963,1020,511,536]" pageId="8" pageNumber="169">n. sp</emphasis>
.
</taxonomicNameLabel>
(vs. distinctly widened in these two species). Finally,
<taxonomicName id="4C66CAEEFFC5C4287E432B3511FAFDBD" box="[354,427,548,572]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287E432B3511FAFDBD" box="[354,427,548,572]" italics="true" pageId="8" pageNumber="169">S. tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFC5C4287E942B3511A9FDBD" box="[437,504,548,573]" pageId="8" pageNumber="169" rank="species">
<emphasis id="B9126D7FFFC5C4287E942B3511A1FDBC" bold="true" box="[437,496,548,573]" pageId="8" pageNumber="169">n. sp</emphasis>
.
</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4C66CAEEFFC5C4287C1B2B35138EFDBD" box="[826,991,548,572]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="kagoroensis">
<emphasis id="B9126D7FFFC5C4287C1B2B35138EFDBD" box="[826,991,548,572]" italics="true" pageId="8" pageNumber="169">S. kagoroensis</emphasis>
</taxonomicName>
from central
<taxonomicName id="4C66CAEEFFC5C4287BA32B351487FDBC" box="[1154,1238,548,573]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="161" phylum="Arthropoda" rank="species" species="nigeria">
<collectingCountry id="F371F1FDFFC5C4287BA32B351487FDBC" box="[1154,1238,548,573]" name="Nigeria" pageId="8" pageNumber="169">Nigeria</collectingCountry>
</taxonomicName>
by the distinctly thickened outer margins of sternal segment
<emphasis id="B9126D7FFFC5C4287DA12B5812DAFDE1" box="[640,651,585,608]" italics="true" pageId="8" pageNumber="169">s</emphasis>
<quantity id="4C9E1C88FFC5C4287DAA2B5912E7FDE1" box="[651,694,583,608]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="169" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C66CAEEFFC5C4287D9C2B591352FDE1" box="[701,771,584,608]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287D9C2B591352FDE1" box="[701,771,584,608]" italics="true" pageId="8" pageNumber="169">S. tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFC5C4287C2B2B56131BFDDE" box="[778,842,583,608]" pageId="8" pageNumber="169" rank="species">
<emphasis id="B9126D7FFFC5C4287C2B2B561313FDE1" bold="true" box="[778,834,583,608]" pageId="8" pageNumber="169">n. sp</emphasis>
.
</taxonomicNameLabel>
(vs. smooth and not thickened in
<taxonomicName id="4C66CAEEFFC5C4287BE42B591539FDE1" box="[1221,1384,584,608]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="kagoroensis">
<emphasis id="B9126D7FFFC5C4287BE42B591539FDE1" box="[1221,1384,584,608]" italics="true" pageId="8" pageNumber="169">S. kagoroensis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFF7CC9CFFC5C4287FB62B7D113AFD05" author="Cumberlidge" box="[151,363,620,645]" pageId="8" pageNumber="169" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
)).
</paragraph>
<paragraph id="8BD9B16DFFC5C4287FE62B9E117FFC01" blockId="8.[151,1437,151,896]" pageId="8" pageNumber="169">
<emphasis id="B9126D7FFFC5C4287FE62B9E11FEFD29" bold="true" box="[199,431,655,680]" pageId="8" pageNumber="169">Molecular analysis.</emphasis>
The phylogenetic tree with confidence values according to BI and ML derived from 1475 basepairs of combined COI and 16S RNA mitochondrial DNA sequences (
<figureCitation id="135DADE8FFC5C4287B002BA51438FD4C" box="[1057,1129,692,717]" captionStart="FIGURE 5" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[168,1406,960,1930]" captionTargetId="figure@8.[155,1418,949,1936]" captionTargetPageId="8" captionText="FIGURE 5. BI / ML tree topology for the freshwater crab taxa from Cameroon included in this study derived from mtDNA sequences corresponding to two loci (partial 16 S rRNA and COI genes, 1475 basepairs). BI and ML statistical values on the nodes indicate posterior probabilities and bootstrap support, respectively (%)." httpUri="https://zenodo.org/record/376073/files/figure.png" pageId="8" pageNumber="169">Fig. 5</figureCitation>
) recovered all species of
<taxonomicName id="4C66CAEEFFC5C4287FB62BC91165FD71" box="[151,308,728,752]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B9126D7FFFC5C4287FB62BC91165FD71" box="[151,308,728,752]" italics="true" pageId="8" pageNumber="169">Sudanonautes</emphasis>
</taxonomicName>
as a single clade, but with relatively low confidence values (BI and ML values of 64/43 at this node, respectively). Each of the included Cameroonian species (i.e.,
<taxonomicName id="4C66CAEEFFC5C4287C792BED138FFC95" box="[856,990,764,788]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC5C4287C792BED138FFC95" box="[856,990,764,788]" italics="true" pageId="8" pageNumber="169">S. africanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C66CAEEFFC5C4287CCD2BED1400FC95" box="[1004,1105,764,788]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B9126D7FFFC5C4287CCD2BED1400FC95" box="[1004,1105,764,788]" italics="true" pageId="8" pageNumber="169">S. aubryi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C66CAEEFFC5C4287B7F2BED1498FC95" box="[1118,1225,764,788]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC5C4287B7F2BED1498FC95" box="[1118,1225,764,788]" italics="true" pageId="8" pageNumber="169">S. floweri</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C66CAEEFFC5C4287A262BED151CFC95" box="[1287,1357,764,788]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287A262BED151CFC95" box="[1287,1357,764,788]" italics="true" pageId="8" pageNumber="169">S. tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFC5C4287A752BED15C0FC95" box="[1364,1425,764,789]" pageId="8" pageNumber="169" rank="species">
<emphasis id="B9126D7FFFC5C4287A752BED15DDFC94" bold="true" box="[1364,1420,764,789]" pageId="8" pageNumber="169">n. sp</emphasis>
.
</taxonomicNameLabel>
) were all found to be well-supported independent lineages.
<taxonomicName id="4C66CAEEFFC5C4287C172A311457FCB9" box="[822,1030,800,824]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC5C4287C172A311457FCB9" box="[822,1030,800,824]" italics="true" pageId="8" pageNumber="169">Sudanonautes tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFC5C4287B2E2A0E1401FCB6" box="[1039,1104,799,824]" pageId="8" pageNumber="169" rank="species">
<emphasis id="B9126D7FFFC5C4287B2E2A0E1418FCB9" bold="true" box="[1039,1097,799,824]" pageId="8" pageNumber="169">n. sp</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C66CAEEFFC5C4287BAA2A3114A6FCB9" box="[1163,1271,800,824]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC5C4287BAA2A3114A6FCB9" box="[1163,1271,800,824]" italics="true" pageId="8" pageNumber="169">S. floweri</emphasis>
</taxonomicName>
were grouped together as sister species by a node with strong support (BI/ML: 100/96), as were
<taxonomicName id="4C66CAEEFFC5C4287B612A551496FCDD" box="[1088,1223,836,860]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC5C4287B612A551496FCDD" box="[1088,1223,836,860]" italics="true" pageId="8" pageNumber="169">S. africanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C66CAEEFFC5C4287A212A551537FCDD" box="[1280,1382,836,860]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B9126D7FFFC5C4287A212A551537FCDD" box="[1280,1382,836,860]" italics="true" pageId="8" pageNumber="169">S. aubryi</emphasis>
</taxonomicName>
(BI/ ML: 100/95).
</paragraph>
<caption id="DF19E1E5FFC5C4287FB62E8913F8F86A" httpUri="https://zenodo.org/record/376073/files/figure.png" pageId="8" pageNumber="169" targetBox="[168,1406,960,1930]" targetPageId="8">
<paragraph id="8BD9B16DFFC5C4287FB62E8913F8F86A" blockId="8.[151,1436,1944,2027]" pageId="8" pageNumber="169">
<emphasis id="B9126D7FFFC5C4287FB62E891144F82C" bold="true" box="[151,277,1944,1966]" pageId="8" pageNumber="169">FIGURE 5.</emphasis>
BI/ML tree topology for the freshwater crab taxa from Cameroon included in this study derived from mtDNA sequences corresponding to two loci (partial 16S rRNA and COI genes, 1475 basepairs). BI and ML statistical values on the nodes indicate posterior probabilities and bootstrap support, respectively (%).
</paragraph>
</caption>
<paragraph id="8BD9B16DFFC4C4297FE62986110AFEE4" blockId="9.[151,1436,151,357]" pageId="9" pageNumber="170">
The phylogenetic tree offers insights into the large amount of genetic diversity within each of the species of
<taxonomicName id="4C66CAEEFFC4C4297FB629AD1165FF55" box="[151,308,188,212]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus">
<emphasis id="B9126D7FFFC4C4297FB629AD1165FF55" box="[151,308,188,212]" italics="true" pageId="9" pageNumber="170">Sudanonautes</emphasis>
</taxonomicName>
from the coastal zone of
<collectingCountry id="F371F1FDFFC4C4297D4429AD128CFF55" box="[613,733,188,212]" name="Cameroon" pageId="9" pageNumber="170">Cameroon</collectingCountry>
. For example, populations of
<taxonomicName id="4C66CAEEFFC4C4297B1B29AD14D5FF55" box="[1082,1156,188,212]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC4C4297B1B29AD14D5FF55" box="[1082,1156,188,212]" italics="true" pageId="9" pageNumber="170">S. tiko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D004FFC4C4297BAF29AD1481FF55" box="[1166,1232,188,213]" pageId="9" pageNumber="170" rank="species">
<emphasis id="B9126D7FFFC4C4297BAF29AD1499FF54" bold="true" box="[1166,1224,188,213]" pageId="9" pageNumber="170">n. sp</emphasis>
.
</taxonomicNameLabel>
from
<taxonomicName id="4C66CAEEFFC4C4297A3A29AD151EFF54" box="[1307,1359,188,213]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
in the
<collectingRegion id="49A27F8FFFC4C4297FB629CE1137FF79" box="[151,358,223,248]" country="Cameroon" name="South-West" pageId="9" pageNumber="170">Southwest Region</collectingRegion>
form a separate subgroup within the clade from populations of this species from Edea in the
<collectingRegion id="49A27F8FFFC4C4297FB62815111DFE9C" box="[151,332,260,285]" country="Cameroon" name="Littoral" pageId="9" pageNumber="170">
<collectingRegion id="49A27F8FFFC4C4297FB6281510BAFE9C" box="[151,235,260,285]" country="Equatorial Guinea" name="Litoral" pageId="9" pageNumber="170">Littoral</collectingRegion>
Region
</collectingRegion>
.
<collectingRegion id="49A27F8FFFC4C4297E7D28151121FE9D" box="[348,368,260,284]" country="Nigeria" name="Rivers" pageId="9" pageNumber="170">A</collectingRegion>
considerable amount of genetic diversification was also found between populations of
<taxonomicName id="4C66CAEEFFC4C4297AA628151150FEC1" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC4C4297AA628151150FEC1" italics="true" pageId="9" pageNumber="170">S. africanus</emphasis>
</taxonomicName>
from
<taxonomicName id="4C66CAEEFFC4C4297E6428361128FEC1" box="[325,377,295,320]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
(T273-10) and Edea (T263-4). This species is nevertheless supported as a monophyletic taxon (BI/ML: 100/65).
</paragraph>
<caption id="DF19E1E5FFC4C4297FB62C461213FA69" httpUri="https://zenodo.org/record/376074/files/figure.png" pageId="9" pageNumber="170" targetBox="[233,1354,411,1340]" targetPageId="9">
<paragraph id="8BD9B16DFFC4C4297FB62C461213FA69" blockId="9.[151,1436,1367,1512]" pageId="9" pageNumber="170">
<emphasis id="B9126D7FFFC4C4297FB62C461143FAED" bold="true" box="[151,274,1367,1389]" pageId="9" pageNumber="170">FIGURE 6.</emphasis>
Whole animal dorsal views showing living color of three species of
<taxonomicName id="4C66CAEEFFC4C4297CE82C461406FAEC" box="[969,1111,1367,1389]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="genus">
<emphasis id="B9126D7FFFC4C4297CE82C461406FAEC" box="[969,1111,1367,1389]" italics="true" pageId="9" pageNumber="170">Sudanonautes</emphasis>
</taxonomicName>
from Cameroon. a,
<taxonomicName id="4C66CAEEFFC4C4297A022C4915CDFAEC" box="[1315,1436,1367,1389]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC4C4297A022C4915CDFAEC" box="[1315,1436,1367,1389]" italics="true" pageId="9" pageNumber="170">S. africanus</emphasis>
</taxonomicName>
adult female, CW 87.4 mm, from
<taxonomicName id="4C66CAEEFFC4C4297EDA2C67127BFA0D" box="[507,554,1398,1420]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
(LZUY-03); b,
<taxonomicName id="4C66CAEEFFC4C4297DF02C66137FFA0D" box="[721,814,1398,1420]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B9126D7FFFC4C4297DF02C66137FFA0D" box="[721,814,1398,1420]" italics="true" pageId="9" pageNumber="170">S. aubryi</emphasis>
</taxonomicName>
subadult male, CW 35.8 mm, from Edea, Bolekoa, Littoral Region (LZUY-06); c,
<taxonomicName id="4C66CAEEFFC4C4297EA32C8411B4FA2B" box="[386,485,1428,1450]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC4C4297EA32C8411B4FA2B" box="[386,485,1428,1450]" italics="true" pageId="9" pageNumber="170">S. floweri</emphasis>
</taxonomicName>
adult male, CW 35.4 mm, from Yabassi, Parc des Princes, Littoral Region (LZUY-09); d,
<taxonomicName id="4C66CAEEFFC4C4297AA82C84108EFA48" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC4C4297AA82C84108EFA48" italics="true" pageId="9" pageNumber="170">S. floweri</emphasis>
</taxonomicName>
adult female, CW 53.9 mm, from
<taxonomicName id="4C66CAEEFFC4C4297D182CA2123BFA48" box="[569,618,1459,1481]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
, Tamba forest, Southwest Region (LZUY-11). Scale bar equals 72 mm (for a), 32 mm (for b), 32 mm (for c), 45 mm (for d).
</paragraph>
</caption>
<paragraph id="8BD9B16DFFC4C42A7FE62F051533FE2C" blockId="9.[151,1437,1556,2014]" lastBlockId="10.[151,1437,151,1256]" lastPageId="10" lastPageNumber="171" pageId="9" pageNumber="170">
<emphasis id="B9126D7FFFC4C4297FE62F0511BEF9AC" bold="true" box="[199,495,1556,1581]" pageId="9" pageNumber="170">Ecological Comparisons.</emphasis>
<taxonomicName id="4C66CAEEFFC4C4297ED42F041355F9AC" box="[501,772,1557,1581]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC4C4297ED42F041355F9AC" box="[501,772,1557,1581]" italics="true" pageId="9" pageNumber="170">Sudanonautes africanus</emphasis>
</taxonomicName>
(
<figureCitation id="135DADE8FFC4C4297C322F051304F9AC" box="[787,853,1556,1581]" captionStart="FIGURE 6" captionStartId="9.[151,250,1367,1389]" captionTargetBox="[233,1354,411,1340]" captionTargetId="figure@9.[225,1362,409,1346]" captionTargetPageId="9" captionText="FIGURE 6. Whole animal dorsal views showing living color of three species of Sudanonautes from Cameroon. a, S. africanus adult female, CW 87.4 mm, from Tiko (LZUY- 03); b, S. aubryi subadult male, CW 35.8 mm, from Edea, Bolekoa, Littoral Region (LZUY- 06); c, S. floweri adult male, CW 35.4 mm, from Yabassi, Parc des Princes, Littoral Region (LZUY- 09); d, S. floweri adult female, CW 53.9 mm, from Tiko, Tamba forest, Southwest Region (LZUY- 11). Scale bar equals 72 mm (for a), 32 mm (for b), 32 mm (for c), 45 mm (for d)." httpUri="https://zenodo.org/record/376074/files/figure.png" pageId="9" pageNumber="170">Fig. 6</figureCitation>
a) occurs in the coastal rainforest regions of
<taxonomicName id="4C66CAEEFFC4C4297A692F0515CDF9AC" box="[1352,1436,1556,1581]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="161" phylum="Arthropoda" rank="species" species="nigeria">
<collectingCountry id="F371F1FDFFC4C4297A692F0515CDF9AC" box="[1352,1436,1556,1581]" name="Nigeria" pageId="9" pageNumber="170">Nigeria</collectingCountry>
</taxonomicName>
and
<collectingRegion id="49A27F8FFFC4C4297FE72F291149F9D0" box="[198,280,1592,1617]" country="Togo" name="Centrale" pageId="9" pageNumber="170">Central</collectingRegion>
Africa (
<bibRefCitation id="EFF7CC9CFFC4C4297E552F291204F9D0" author="Cumberlidge" box="[372,597,1592,1617]" pageId="9" pageNumber="170" refString="Cumberlidge, N. (1995 c) Redescription of the African fresh-water crab Sudanonautes africanus (A. Milne-Edwards, 1869) (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 15, 588 - 598." type="journal article" year="1995" yearSuffix="c">Cumberlidge 1995c</bibRefCitation>
,
<bibRefCitation id="EFF7CC9CFFC4C4297D402F2812CBF9D0" author="Cumberlidge" box="[609,666,1593,1617]" pageId="9" pageNumber="170" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">1999</bibRefCitation>
) and was collected during the present study from rainforest localities along the coastal region between Mount
<collectingCountry id="F371F1FDFFC4C4297D7E2F4C1285F9F4" box="[607,724,1629,1653]" name="Cameroon" pageId="9" pageNumber="170">Cameroon</collectingCountry>
and the border with
<collectingCountry id="F371F1FDFFC4C4297C9D2F4C14D8F9F7" box="[956,1161,1629,1654]" name="Equatorial Guinea" pageId="9" pageNumber="170">Equatorial Guinea</collectingCountry>
(Campo, Kribi, Bipindi,
<collectingRegion id="49A27F8FFFC4C4297FB62F91109EF918" box="[151,207,1664,1689]" country="South Sudan" name="Lakes" pageId="9" pageNumber="170">Lake</collectingRegion>
Ossa, Edea, Yabassi,
<taxonomicName id="4C66CAEEFFC4C4297EFD2F911244F918" box="[476,533,1664,1689]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
). This species has also been collected from a number of other localities in
<collectingCountry id="F371F1FDFFC4C4297FB62FB4115DF93C" box="[151,268,1701,1725]" name="Cameroon" pageId="9" pageNumber="170">Cameroon</collectingCountry>
(
<collectingRegion id="49A27F8FFFC4C4297E3D2FB41107F93F" box="[284,342,1701,1726]" country="South Sudan" name="Lakes" pageId="9" pageNumber="170">Lake</collectingRegion>
Barombi, Kumba,
<collectingRegion id="49A27F8FFFC4C4297D152FB412D1F93F" box="[564,640,1701,1726]" country="Cameroon" name="West" pageId="9" pageNumber="170">Limbe</collectingRegion>
, Yaounde, Enongal, and Ebolowa;
<bibRefCitation id="EFF7CC9CFFC4C4297B332FB414BBF93C" author="Cumberlidge" box="[1042,1258,1701,1726]" pageId="9" pageNumber="170" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
).
<taxonomicName id="4C66CAEEFFC4C4297BDE2FB41150F960" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC4C4297BDE2FB41150F960" italics="true" pageId="9" pageNumber="170">Sudanonautes africanus</emphasis>
</taxonomicName>
was the commonest freshwater crab collected during this study and its abundance and large size make it a popular food item for the local population. This species occurs in a range of permanent aquatic habitats from large rivers to small streams (
<bibRefCitation id="EFF7CC9CFFC4C4297EC52E0112EDF8A8" author="Cumberlidge" box="[484,700,1808,1833]" pageId="9" pageNumber="170" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
). Male and female sub-adults and juveniles were captured during the day either by hand from under stones in small streams or from large rivers downstream of the collection site, if they escaped in the turbid waters created when the rocks were turned. Some were trapped at night entangled in nylon fishing nets set bank-to-bank across the stream and baited with cassava. Others were collected in a sieve passed through the water. These methods tend to capture only juveniles, because the larger subadults and adults of
<taxonomicName id="4C66CAEEFFC4C4297FB62ED4114EF85C" box="[151,287,1989,2013]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC4C4297FB62ED4114EF85C" box="[151,287,1989,2013]" italics="true" pageId="9" pageNumber="170">S. africanus</emphasis>
</taxonomicName>
are semiterrestrial, live in
<quantity id="4C9E1C88FFC4C4297D782ED412D7F85C" box="[601,646,1989,2013]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="170" unit="m" value="1.0">1 m</quantity>
deep burrows close to streams, and need to be collected by hand or caught at night in pitfall traps (
<bibRefCitation id="EFF7CC9CFFC7C42A7ED72986129FFF31" author="Cumberlidge" box="[502,718,151,176]" pageId="10" pageNumber="171" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
). Adult males and females of
<taxonomicName id="4C66CAEEFFC7C42A7B03298914F8FF31" box="[1058,1193,152,176]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC7C42A7B03298914F8FF31" box="[1058,1193,152,176]" italics="true" pageId="10" pageNumber="171">S. africanus</emphasis>
</taxonomicName>
dig separate burrows and do not co-inhabit a single hole. The burrows dug by adult males were further away from the water than those of adult females and subadults of both sexes. The distribution of the size classes in the population indicates that large adult specimens of
<taxonomicName id="4C66CAEEFFC7C42A7E4F281511A5FE9D" box="[366,500,260,284]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC7C42A7E4F281511A5FE9D" box="[366,500,260,284]" italics="true" pageId="10" pageNumber="171">S. africanus</emphasis>
</taxonomicName>
are less abundant than the more numerous smaller individuals (
<bibRefCitation id="EFF7CC9CFFC7C42A7B98281515C1FE9D" author="Cumberlidge" box="[1209,1424,260,285]" pageId="10" pageNumber="171" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
), so that the capture of reproductive adults causes the greatest loss to the population. In addition, juveniles and subadults of
<taxonomicName id="4C66CAEEFFC7C42A7E0B285D11E3FEE5" box="[298,434,332,356]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC7C42A7E0B285D11E3FEE5" box="[298,434,332,356]" italics="true" pageId="10" pageNumber="171">S. africanus</emphasis>
</taxonomicName>
living in the coastal zone of
<collectingCountry id="F371F1FDFFC7C42A7C27285D132AFEE5" box="[774,891,332,356]" name="Cameroon" pageId="10" pageNumber="171">Cameroon</collectingCountry>
are heavily exploited as a food source by local human populations. These food harvesting activities constitute a serious threat to the long-term existence of this species in this part of
<collectingCountry id="F371F1FDFFC7C42A7EAD28851255FE2D" box="[396,516,404,428]" name="Cameroon" pageId="10" pageNumber="171">Cameroon</collectingCountry>
. The carapace, claws and walking legs of
<taxonomicName id="4C66CAEEFFC7C42A7CFA28851431FE2D" box="[987,1120,404,428]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC7C42A7CFA28851431FE2D" box="[987,1120,404,428]" italics="true" pageId="10" pageNumber="171">S. africanus</emphasis>
</taxonomicName>
are dark brown in life.
</paragraph>
</subSubSection>
<subSubSection id="C37CE2E6FFC7C42A7FE628A91466FB69" pageId="10" pageNumber="171" type="reference_group">
<paragraph id="8BD9B16DFFC7C42A7FE628A91095FCB9" blockId="10.[151,1437,151,1256]" pageId="10" pageNumber="171">
<taxonomicName id="4C66CAEEFFC7C42A7FE628A911E9FE51" ID-CoL="7B2YW" box="[199,440,440,464]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B9126D7FFFC7C42A7FE628A911E9FE51" box="[199,440,440,464]" italics="true" pageId="10" pageNumber="171">Sudanonautes aubryi</emphasis>
</taxonomicName>
(
<figureCitation id="135DADE8FFC7C42A7EEC28A61242FE51" box="[461,531,439,464]" captionStart="FIGURE 6" captionStartId="9.[151,250,1367,1389]" captionTargetBox="[233,1354,411,1340]" captionTargetId="figure@9.[225,1362,409,1346]" captionTargetPageId="9" captionText="FIGURE 6. Whole animal dorsal views showing living color of three species of Sudanonautes from Cameroon. a, S. africanus adult female, CW 87.4 mm, from Tiko (LZUY- 03); b, S. aubryi subadult male, CW 35.8 mm, from Edea, Bolekoa, Littoral Region (LZUY- 06); c, S. floweri adult male, CW 35.4 mm, from Yabassi, Parc des Princes, Littoral Region (LZUY- 09); d, S. floweri adult female, CW 53.9 mm, from Tiko, Tamba forest, Southwest Region (LZUY- 11). Scale bar equals 72 mm (for a), 32 mm (for b), 32 mm (for c), 45 mm (for d)." httpUri="https://zenodo.org/record/376074/files/figure.png" pageId="10" pageNumber="171">Fig. 6</figureCitation>
b) is found both in the Lower Guinea and Upper Guinea forest zones in the rainforest and moist guinea savannas in Côte-d'Ivoire, Ghana, Togo, Benin,
<taxonomicName id="4C66CAEEFFC7C42A7CC428CD146DFE74" ID-CoL="7B2Z4" box="[997,1084,476,501]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="nigeria">Nigeria</taxonomicName>
, Cameroon, Equatorial Guinea, and Gabon (
<bibRefCitation id="EFF7CC9CFFC7C42A7E0B28EE1257FD99" author="Cumberlidge" box="[298,518,511,536]" pageId="10" pageNumber="171" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
). In Cameroon, this species was collected from rainforest localities along the coastal region near Edea.
<taxonomicName id="4C66CAEEFFC7C42A7E982B3512F6FDBD" ID-CoL="7B2YW" box="[441,679,548,572]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B9126D7FFFC7C42A7E982B3512F6FDBD" box="[441,679,548,572]" italics="true" pageId="10" pageNumber="171">Sudanonautes aubryi</emphasis>
</taxonomicName>
has also been collected from the following localities in Cameroon: Lake Barombi, Kumba, Limbe, Yaounde, Bipindi, Enongal, and Ebolowa, and in the Mfiende, Enyumu, and Lokoundié Rivers (
<bibRefCitation id="EFF7CC9CFFC7C42A7E532B7D121AFD05" author="Cumberlidge" box="[370,587,620,645]" pageId="10" pageNumber="171" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
). All size classes of
<taxonomicName id="4C66CAEEFFC7C42A7C192B7D13CEFD05" ID-CoL="7B2YW" box="[824,927,620,644]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B9126D7FFFC7C42A7C192B7D13CEFD05" box="[824,927,620,644]" italics="true" pageId="10" pageNumber="171">S. aubryi</emphasis>
</taxonomicName>
were collected from streams and rivers, and none were collected on land or in burrows near the water, and this species was sympatric with juveniles of
<emphasis id="B9126D7FFFC7C42A7AA62B811251FD4D" italics="true" pageId="10" pageNumber="171">
<taxonomicName id="4C66CAEEFFC7C42A7AA62B811154FD4D" ID-CoL="7B2ZV" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="africanus">S. africanus</taxonomicName>
.
<taxonomicName id="4C66CAEEFFC7C42A7E302BA51251FD4D" ID-CoL="7B2YW" box="[273,512,692,716]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="aubryi">Sudanonautes aubryi</taxonomicName>
</emphasis>
was collected either by hand or with a nylon fishnet or dipnet from under rocks and stones in the stream bed, and was only found in this study in two localities near Edea (Bolekoa stream and Bedimet Island in Lake Ossa). The carapace, claws and walking legs of
<taxonomicName id="4C66CAEEFFC7C42A7C9E2BED1475FC95" ID-CoL="7B2YW" box="[959,1060,764,788]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="aubryi">
<emphasis id="B9126D7FFFC7C42A7C9E2BED1475FC95" box="[959,1060,764,788]" italics="true" pageId="10" pageNumber="171">S. aubryi</emphasis>
</taxonomicName>
are generally yellowish brown in life.
</paragraph>
<paragraph id="8BD9B16DFFC7C42A7FE62A551466FB69" blockId="10.[151,1437,151,1256]" pageId="10" pageNumber="171">
<taxonomicName id="4C66CAEEFFC7C42A7FE62A5511EDFCDD" ID-CoL="7B2YY" box="[199,444,836,860]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC7C42A7FE62A5511EDFCDD" box="[199,444,836,860]" italics="true" pageId="10" pageNumber="171">Sudanonautes floweri</emphasis>
</taxonomicName>
(
<figureCitation id="135DADE8FFC7C42A7EEC2A551240FCDC" box="[461,529,836,861]" captionStart="FIGURE 6" captionStartId="9.[151,250,1367,1389]" captionTargetBox="[233,1354,411,1340]" captionTargetId="figure@9.[225,1362,409,1346]" captionTargetPageId="9" captionText="FIGURE 6. Whole animal dorsal views showing living color of three species of Sudanonautes from Cameroon. a, S. africanus adult female, CW 87.4 mm, from Tiko (LZUY- 03); b, S. aubryi subadult male, CW 35.8 mm, from Edea, Bolekoa, Littoral Region (LZUY- 06); c, S. floweri adult male, CW 35.4 mm, from Yabassi, Parc des Princes, Littoral Region (LZUY- 09); d, S. floweri adult female, CW 53.9 mm, from Tiko, Tamba forest, Southwest Region (LZUY- 11). Scale bar equals 72 mm (for a), 32 mm (for b), 32 mm (for c), 45 mm (for d)." httpUri="https://zenodo.org/record/376074/files/figure.png" pageId="10" pageNumber="171">Fig. 6</figureCitation>
c,d) is found in the moister regions of the woodland and guinea savanna zones from central
<taxonomicName id="4C66CAEEFFC7C42A7E162A7611DAFC01" ID-CoL="7B2Z4" box="[311,395,871,896]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="nigeria">Nigeria</taxonomicName>
to South Sudan and northern Uganda and from humid tropical rainforest habitats in Cameroon, Bioko, Central African Republic, D. R. Congo, Congo, Gabon, Cabinda Angola, and Equatorial Guinea (
<bibRefCitation id="EFF7CC9CFFC7C42A7FBF2ABE112AFC49" author="Cumberlidge" box="[158,379,943,968]" pageId="10" pageNumber="171" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
). In Cameroon, this species was collected only in rainforest habitats at Yabassi in south Cameroon, and at
<taxonomicName id="4C66CAEEFFC7C42A7E4B2AC511CFFC6C" ID-CoL="7B2Z6" box="[362,414,980,1005]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
near Mount Cameroon, but it has also been recorded by others from Limbe and Bibundi west of Mount Cameroon (
<bibRefCitation id="EFF7CC9CFFC7C42A7EF32AE612FDFB91" author="Cumberlidge" box="[466,684,1015,1040]" pageId="10" pageNumber="171" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
).
<taxonomicName id="4C66CAEEFFC7C42A7DE22AE913E8FB91" ID-CoL="7B2YY" box="[707,953,1016,1040]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC7C42A7DE22AE913E8FB91" box="[707,953,1016,1040]" italics="true" pageId="10" pageNumber="171">Sudanonautes floweri</emphasis>
</taxonomicName>
is found under rocks in shallow streams, rivers, and ponds, in burrows near waterways, and on land next some distance from the nearest water (
<bibRefCitation id="EFF7CC9CFFC7C42A7A292D0D1082FBD9" author="Cumberlidge" pageId="10" pageNumber="171" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. In: Faune et Flore tropicales. Fol. 35. IRD, Paris, pp. 1 - 382." type="journal article" year="1999">Cumberlidge 1999</bibRefCitation>
). At Yabassi,
<taxonomicName id="4C66CAEEFFC7C42A7E4E2D51118AFBD9" ID-CoL="7B2YY" box="[367,475,1088,1112]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC7C42A7E4E2D51118AFBD9" box="[367,475,1088,1112]" italics="true" pageId="10" pageNumber="171">S. floweri</emphasis>
</taxonomicName>
digs burrows near streams, whereas at
<taxonomicName id="4C66CAEEFFC7C42A7CB82D2E139CFBD9" ID-CoL="7B2Z6" box="[921,973,1087,1112]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
it is found in the stream together with
<taxonomicName id="4C66CAEEFFC7C42A7AA62D511150FBFD" ID-CoL="7B2ZV" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC7C42A7AA62D511150FBFD" italics="true" pageId="10" pageNumber="171">S. africanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C66CAEEFFC7C42A7E162D75112CFBFD" ID-CoL="7B2Z6" box="[311,381,1124,1148]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC7C42A7E162D75112CFBFD" box="[311,381,1124,1148]" italics="true" pageId="10" pageNumber="171">S. tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFC7C42A7EA52D751192FBFC" bold="true" box="[388,451,1124,1149]" pageId="10" pageNumber="171">
<taxonomicNameLabel id="A221D004FFC7C42A7EA52D751192FBFC" box="[388,451,1124,1149]" pageId="10" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. It is noteworthy that males and females of
<taxonomicName id="4C66CAEEFFC7C42A7C8E2D75144BFBFD" ID-CoL="7B2YY" box="[943,1050,1124,1148]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC7C42A7C8E2D75144BFBFD" box="[943,1050,1124,1148]" italics="true" pageId="10" pageNumber="171">S. floweri</emphasis>
</taxonomicName>
usually share the same burrow (in contrast to
<taxonomicName id="4C66CAEEFFC7C42A7E332D9911C6FB21" ID-CoL="7B2ZV" box="[274,407,1160,1184]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC7C42A7E332D9911C6FB21" box="[274,407,1160,1184]" italics="true" pageId="10" pageNumber="171">S. africanus</emphasis>
</taxonomicName>
, where adult males and females dig separate holes). The carapace of
<taxonomicName id="4C66CAEEFFC7C42A7BB82D991555FB21" ID-CoL="7B2YY" box="[1177,1284,1160,1184]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC7C42A7BB82D991555FB21" box="[1177,1284,1160,1184]" italics="true" pageId="10" pageNumber="171">S. floweri</emphasis>
</taxonomicName>
is either dark brown, yellowish, or purple, and the claws and walking legs are pale yellow in life. At Tamba forest in
<taxonomicName id="4C66CAEEFFC7C42A7A6B2DBD152FFB44" ID-CoL="7B2Z6" box="[1354,1406,1196,1221]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">Tiko</taxonomicName>
<taxonomicName id="4C66CAEEFFC7C42A7AA62DBD10B7FB69" ID-CoL="7B2YY" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="B9126D7FFFC7C42A7AA62DBD10B7FB69" italics="true" pageId="10" pageNumber="171">S. floweri</emphasis>
</taxonomicName>
is sympatric (living in the same stream) with
<taxonomicName id="4C66CAEEFFC7C42A7DC92DC1137FFB69" ID-CoL="7B2Z6" box="[744,814,1232,1256]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="tiko" status="sp. nov.">
<emphasis id="B9126D7FFFC7C42A7DC92DC1137FFB69" box="[744,814,1232,1256]" italics="true" pageId="10" pageNumber="171">S. tiko</emphasis>
</taxonomicName>
<emphasis id="B9126D7FFFC7C42A7C142DDE1325FB69" bold="true" box="[821,884,1231,1256]" pageId="10" pageNumber="171">
<taxonomicNameLabel id="A221D004FFC7C42A7C142DDE1325FB69" box="[821,884,1231,1256]" pageId="10" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C66CAEEFFC7C42A7C8A2DC11461FB69" ID-CoL="7B2ZV" box="[939,1072,1232,1256]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="171" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B9126D7FFFC7C42A7C8A2DC11461FB69" box="[939,1072,1232,1256]" italics="true" pageId="10" pageNumber="171">S. africanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>